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<mods:title id="1F582643C1BE2C72102D500FEAF7ECB1">Revision of the genus Tityopsis Armas 1974 (Scorpiones Buthidae) Part 1 General updates and description of four new species</mods:title>
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<taxonomicName id="4C745B8EFFD6390C2DEB74E7FB18FEB1" ID-CoL="5797J" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[1019,1204,234,259]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">Tityopsis sheylae</taxonomicName>
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(
<figureCitation id="134F3C88FFD6390C2DA57507FBB0FE90" box="[949,1052,267,291]" captionStart="Figure 2" captionStartId="4.[113,181,849,871]" captionTargetBox="[113,1474,234,838]" captionTargetId="figure-658@4.[113,1474,234,838]" captionTargetPageId="4" captionText="Figure 2. Western Cuba, showing the complete geographical distribution of the genus Tityopsis (white symbols) and overlaid, that of the four species of Tityopsis treated herein (red symbols, complemented with habitus of their holotypes): T. mulata sp. n. (star), T. canizaresorum sp. n. (rhombs, the southernmost is the type locality), T. pumila sp. n. (triangle) and T. sheylae sp. n. (square). Image frame size, 500 x 250 km." figureDoi="http://doi.org/10.5281/zenodo.4648297" httpUri="https://zenodo.org/record/4648297/files/figure.png" pageId="27" pageNumber="26">Figures 2</figureCitation>
,
<figureCitation id="134F3C88FFD6390C2A387507FBC2FE90" box="[1064,1134,267,291]" captionStart-0="Figures 23–24" captionStart-1="Figures 25–26" captionStart-2="Figures 27–28" captionStart-3="Figures 29–30" captionStart-4="Figures 31–32" captionStartId-0="29.[115,192,2018,2040]" captionStartId-1="30.[113,190,1982,2004]" captionStartId-2="31.[115,192,2011,2033]" captionStartId-3="32.[116,193,1436,1458]" captionStartId-4="33.[113,190,1986,2008]" captionTargetBox-0="[205,1391,240,2005]" captionTargetBox-1="[158,1454,285,1902]" captionTargetBox-2="[113,1474,240,1986]" captionTargetBox-3="[113,1474,234,1429]" captionTargetBox-4="[125,1459,280,1967]" captionTargetId-0="figure-7@29.[113,1474,231,2009]" captionTargetId-1="figure-13@30.[113,1474,234,1972]" captionTargetId-2="figure-9@31.[113,1476,234,1986]" captionTargetId-3="figure-14@32.[113,1474,234,1429]" captionTargetId-4="figure-9@33.[113,1474,230,1979]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="32" captionTargetPageId-4="33" captionText-0="Figures 23–24. Tityopsis sheylae sp. n., male holotype (23) and female paratopotype (24), habitus: a) dorsal; b) ventral. Scale bar = 10 mm." captionText-1="Figures 25–26. Tityopsis sheylae sp. n., male holotype (25) and female paratopotype (26): a) pedipalp, dorsal view; b) pedipalp chela, external view; c) pedipalp chela, ventral view; d) telson, lateral view." captionText-2="Figures 27–28. Tityopsis sheylae sp. n., male holotype (27) and female paratopotype (28): a) chelicerae and carapace; b) tergites." captionText-3="Figures 29–30. Tityopsis sheylae sp. n., male holotype (29) and female paratopotype (30): a) sternopectinal region; b) sternite V." captionText-4="Figures 31–32. Tityopsis sheylae sp. n., male holotype (31) and female paratopotype (32), metasoma and telson: a) dorsal, b) lateral, and c) ventral views." figureDoi-0="http://doi.org/10.5281/zenodo.4648321" figureDoi-1="http://doi.org/10.5281/zenodo.4648323" figureDoi-2="http://doi.org/10.5281/zenodo.4648325" figureDoi-3="http://doi.org/10.5281/zenodo.4648327" figureDoi-4="http://doi.org/10.5281/zenodo.4648329" httpUri-0="https://zenodo.org/record/4648321/files/figure.png" httpUri-1="https://zenodo.org/record/4648323/files/figure.png" httpUri-2="https://zenodo.org/record/4648325/files/figure.png" httpUri-3="https://zenodo.org/record/4648327/files/figure.png" httpUri-4="https://zenodo.org/record/4648329/files/figure.png" pageId="27" pageNumber="26">23–32</figureCitation>
,
<figureCitation id="134F3C88FFD6390C2A687507FB39FE90" box="[1144,1173,267,291]" captionStart="Figure 36" captionStartId="37.[113,181,1923,1945]" captionTargetBox="[113,1474,249,1820]" captionTargetId="figure-8@37.[113,1474,249,1820]" captionTargetPageId="37" captionText="Figure 36. Tityopsis sheylae sp. n., photographed alive in its natural habitat unless otherwise noted: a) male holotype; b) small-sized male paratopotype; c) female paratopotype; d) captive female paratopotype, with mixed litter of instars 1–2; e) juvenile female paratopotype; f) just-molted, small-sized adult male paratopotype, being predated upon by the pholcid Ciboneya nuriae (the spider is not visible because while taking the photographs, it fled from the camera’s flash and hid in the crevice behind the scorpion). Photos courtesy Sheyla Yong (a–b)." figureDoi="http://doi.org/10.5281/zenodo.4648335" httpUri="https://zenodo.org/record/4648335/files/figure.png" pageId="27" pageNumber="26">36</figureCitation>
,
<figureCitation id="134F3C88FFD6390C2AB07507FB11FE90" box="[1184,1213,267,291]" captionStart="Figures 39–40" captionStartId="39.[113,190,1347,1369]" captionTargetBox="[113,1474,249,1338]" captionTargetId="figure-349@39.[113,1474,249,1338]" captionTargetPageId="39" captionText="Figures 39–40: Type localities of Tityopsis canizaresorum sp. n. (39) and T. sheylae sp. n. (40), a) habitat, b) microhabitat (photo courtesy Sheyla Yong)." figureDoi="http://doi.org/10.5281/zenodo.4648340" httpUri="https://zenodo.org/record/4648340/files/figure.png" pageId="27" pageNumber="26">40</figureCitation>
,
<tableCitation id="C6F615B6FFD6390C2AD87507FA91FE90" box="[1224,1341,267,291]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="28.[115,171,1108,1129]" captionStartId-1="34.[114,170,598,619]" captionTargetBox-1="[124,1452,250,581]" captionTargetId-1="graphics-728@34.[119,1470,238,585]" captionText-0="Table 3. Measurements (mm) of four adult types of Tityopsis sheylae sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 4. Pectinal tooth count variation in the four species of the genus Tityopsis treated in this paper. Abbreviations: number of pectines (N)," httpUri-1="http://table.plazi.org/id/DF0B7085FFEF39352E62765AFE5BFD3B" pageId="27" pageNumber="26" tableUuid-1="DF0B7085FFEF39352E62765AFE5BFD3B">Tables 3–4</tableCitation>
)
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.
<collectorName id="268145DBFFD6390C2AED7687FACDFD10" box="[1277,1377,651,675]" pageId="27" pageNumber="26">Teruel, S</collectorName>
. Yong &amp;
<collectionCode id="ED65B8C8FFD6390C2D2076A7FCEEFD71" box="[816,834,683,706]" name="Tavera, Department of Geology and Geophysics" pageId="27" pageNumber="26">T</collectionCode>
. M.
<collectorName id="268145DBFFD6390C2D6976A7FC47FD70" box="[889,1003,683,707]" pageId="27" pageNumber="26">Rodríguez</collectorName>
,
<specimenCount id="9D72EB84FFD6390C2DE976A7FBB6FD70" box="[1017,1050,683,707]" pageId="27" pageNumber="26" type="male">1♂</specimenCount>
(
<typeStatus id="54CF9EAFFFD6390C2A3C76A7FB23FD70" box="[1068,1167,683,707]" pageId="27" pageNumber="26" type="holotype">holotype</typeStatus>
), RTO,
</materialsCitation>
<materialsCitation id="3B1C2A50FFD6390C2AF976A7FC43FCB0" ID-GBIF-Occurrence="3113711322" collectingDate="2019-04-03" collectingDateMax="2018-01-05" collectingDateMin="2017-05-12" collectionCode="TYPE, SPECIMENS" collectorName="Teruel, S" country="Cuba" county="San Jose de Las Lajas" elevation="190" location="Tapaste" municipality="Jaruco Municipalities" pageId="27" pageNumber="26" specimenCount="1" specimenCount-female="12" specimenCount-male="1" stateProvince="Mayabeque" typeStatus="paratype">
<date id="FFCA06CDFFD6390C2AF976A7FA12FD70" box="[1257,1470,683,707]" pageId="27" pageNumber="26" value="2014-12-13">
<collectingDate id="EF8EFF25FFD6390C2AF976A7FA12FD70" box="[1257,1470,683,707]" pageId="27" pageNumber="26" value="2014-12-13">13 December 2014</collectingDate>
</date>
, leg.
<collectionCode id="ED65B8C8FFD6390C2D7076C7FCDEFD51" box="[864,882,715,738]" name="Tavera, Department of Geology and Geophysics" pageId="27" pageNumber="26">T</collectionCode>
. M.
<collectorName id="268145DBFFD6390C2DB476C7FBBFFD50" box="[932,1043,715,739]" pageId="27" pageNumber="26">Rodríguez</collectorName>
&amp; A.
<collectorName id="268145DBFFD6390C2A4A76C7FB66FD50" box="[1114,1226,715,739]" pageId="27" pageNumber="26">Longueira</collectorName>
,
<specimenCount id="9D72EB84FFD6390C2AC676C7FB5BFD50" box="[1238,1271,715,739]" pageId="27" pageNumber="26" type="male">3♂</specimenCount>
<specimenCount id="9D72EB84FFD6390C2AE776C7FABFFD50" box="[1271,1299,715,739]" pageId="27" pageNumber="26" type="female">7♀</specimenCount>
<specimenCount id="9D72EB84FFD6390C2B0376C7FA9AFD50" box="[1299,1334,715,739]" pageId="27" pageNumber="26" type="male">2♂</specimenCount>
juvs.
<specimenCount id="9D72EB84FFD6390C2B7F76C7FA25FD50" box="[1391,1417,715,739]" pageId="27" pageNumber="26" type="female">3♀</specimenCount>
juvs. (
<typeStatus id="54CF9EAFFFD6390C2D2876E7FC08FCB0" box="[824,932,747,771]" pageId="27" pageNumber="26" type="paratype">paratypes</typeStatus>
), RTO,
</materialsCitation>
<materialsCitation id="3B1C2A50FFD6390C2DE476E7FBC9FC90" ID-GBIF-Occurrence="3113711327" collectingDate="2019-04-03" collectingDateMax="2018-01-05" collectingDateMin="2017-05-12" collectionCode="TYPE, SPECIMENS" collectorName="Teruel, S" country="Cuba" county="San Jose de Las Lajas" elevation="190" location="Tapaste" municipality="Jaruco Municipalities" pageId="27" pageNumber="26" specimenCount="1" specimenCount-female="12" specimenCount-male="1" stateProvince="Mayabeque" typeStatus="paratype">
<date id="FFCA06CDFFD6390C2DE476E7FB2FFCB0" box="[1012,1155,747,771]" pageId="27" pageNumber="26" value="2017-05-12">
<collectingDate id="EF8EFF25FFD6390C2DE476E7FB2FFCB0" box="[1012,1155,747,771]" pageId="27" pageNumber="26" value="2017-05-12">12 May 2017</collectingDate>
</date>
, leg.
<collectorName id="268145DBFFD6390C2AAA76E7FAA1FCB0" box="[1210,1293,747,771]" pageId="27" pageNumber="26">S. Yong</collectorName>
&amp;
<collectorName id="268145DBFFD6390C2B3D76E7FA12FCB0" box="[1325,1470,747,771]" pageId="27" pageNumber="26">L. Forcelledo</collectorName>
,
<specimenCount id="9D72EB84FFD6390C2D207707FCC9FC90" box="[816,869,779,803]" pageId="27" pageNumber="26" type="female">1♀1♀</specimenCount>
juv. (
<typeStatus id="54CF9EAFFFD6390C2DBB7707FBBBFC90" box="[939,1047,779,803]" pageId="27" pageNumber="26" type="paratype">paratypes</typeStatus>
), RTO,
</materialsCitation>
<materialsCitation id="3B1C2A50FFD6390C2A7E7707FCC6FCD0" ID-GBIF-Occurrence="3113711313" collectingDate="2019-04-03" collectingDateMax="2018-01-05" collectingDateMin="2017-05-12" collectionCode="TYPE, SPECIMENS" collectorName="Teruel, S" country="Cuba" county="San Jose de Las Lajas" elevation="190" location="Tapaste" municipality="Jaruco Municipalities" pageId="27" pageNumber="26" specimenCount="1" specimenCount-female="12" specimenCount-male="1" stateProvince="Mayabeque" typeStatus="paratype">
<date id="FFCA06CDFFD6390C2A7E7707FAB5FC90" box="[1134,1305,779,803]" pageId="27" pageNumber="26" value="2018-01-05">
<collectingDate id="EF8EFF25FFD6390C2A7E7707FAB5FC90" box="[1134,1305,779,803]" pageId="27" pageNumber="26" value="2018-01-05">5 January 2018</collectingDate>
</date>
, leg.
<collectionCode id="ED65B8C8FFD6390C2B467707FAC7FC91" box="[1366,1387,779,802]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="268145DBFFD6390C2B687707FA13FC90" box="[1400,1471,779,803]" pageId="27" pageNumber="26">Teruel</collectorName>
, S.
<collectorName id="268145DBFFD6390C2D5D7727FC2AFCF0" box="[845,902,811,835]" pageId="27" pageNumber="26">Yong</collectorName>
&amp; L.
<collectorName id="268145DBFFD6390C2DD87727FB91FCF0" box="[968,1085,811,835]" pageId="27" pageNumber="26">Forcelledo</collectorName>
,
<specimenCount id="9D72EB84FFD6390C2A587727FBC7FCF0" box="[1096,1131,811,835]" pageId="27" pageNumber="26" type="male">5♂</specimenCount>
<specimenCount id="9D72EB84FFD6390C2A7B7727FB2BFCF0" box="[1131,1159,811,835]" pageId="27" pageNumber="26" type="female">9♀</specimenCount>
<specimenCount id="9D72EB84FFD6390C2A987727FB07FCF0" box="[1160,1195,811,835]" pageId="27" pageNumber="26" type="male">2♂</specimenCount>
juvs.
<specimenCount id="9D72EB84FFD6390C2AF67727FAACFCF0" box="[1254,1280,811,835]" pageId="27" pageNumber="26" type="female">3♀</specimenCount>
juvs. (
<typeStatus id="54CF9EAFFFD6390C2B5B7727FA1BFCF0" box="[1355,1463,811,835]" pageId="27" pageNumber="26" type="paratype">paratypes</typeStatus>
), RTO,
</materialsCitation>
<materialsCitation id="3B1C2A50FFD6390C2D607747FB9AFC30" ID-GBIF-Occurrence="3113711304" collectingDate="2019-04-03" collectingDateMax="2018-01-05" collectingDateMin="2017-05-12" collectionCode="TYPE, SPECIMENS" collectorName="Teruel, S" country="Cuba" county="San Jose de Las Lajas" elevation="190" location="Tapaste" municipality="Jaruco Municipalities" pageId="27" pageNumber="26" specimenCount="1" specimenCount-female="12" specimenCount-male="1" stateProvince="Mayabeque" typeStatus="paratype">
<date id="FFCA06CDFFD6390C2D607747FC5BFCD0" box="[880,1015,843,867]" pageId="27" pageNumber="26" value="2019-04-03">
<collectingDate id="EF8EFF25FFD6390C2D607747FC5BFCD0" box="[880,1015,843,867]" pageId="27" pageNumber="26" value="2019-04-03">3 April 2019</collectingDate>
</date>
, leg.
<collectionCode id="ED65B8C8FFD6390C2A3E7747FBEFFCD1" box="[1070,1091,843,866]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="26">R</collectionCode>
.
<collectorName id="268145DBFFD6390C2A5B7747FB00FCD0" box="[1099,1196,843,867]" pageId="27" pageNumber="26">Teruel, S</collectorName>
. Yong &amp;
<collectionCode id="ED65B8C8FFD6390C2B1E7747FA8CFCD1" box="[1294,1312,843,866]" name="Tavera, Department of Geology and Geophysics" pageId="27" pageNumber="26">T</collectionCode>
. M.
<collectorName id="268145DBFFD6390C2B5C7747FA12FCD0" box="[1356,1470,843,867]" pageId="27" pageNumber="26">Rodríguez</collectorName>
,
<specimenCount id="9D72EB84FFD6390C2D207767FCF9FC30" box="[816,853,875,899]" pageId="27" pageNumber="26" type="male">2♂</specimenCount>
<specimenCount id="9D72EB84FFD6390C2D457767FCDFFC30" box="[853,883,875,899]" pageId="27" pageNumber="26" type="female">6♀</specimenCount>
(
<typeStatus id="54CF9EAFFFD6390C2D917767FC41FC30" box="[897,1005,875,899]" pageId="27" pageNumber="26" type="paratype">paratypes</typeStatus>
), RTO
</materialsCitation>
.
</paragraph>
<paragraph id="8BCB200DFFD6390C2D2077A7FC70FBD0" blockId="27.[816,1474,938,1315]" pageId="27" pageNumber="26">
<materialsCitation id="3B1C2A50FFD6390C2D2077A7FC74FBD0" ID-GBIF-Occurrence="3113711319" collectingDate="1995-12-23" collectingDateMax="1999-09-27" collectingDateMin="1995-12-23" collectionCode="OTHER, MATERIAL, EXAMINED" collectorName="A. Perez &amp; E. Fonseca" country="Cuba" elevation="190" location="San Jose de Las Lajas" pageId="27" pageNumber="26" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mayabeque">
<collectionCode id="ED65B8C8FFD6390C2D2077A7FBC1FC70" box="[816,1133,939,963]" pageId="27" pageNumber="26">OTHER MATERIAL EXAMINED</collectionCode>
.
<emphasis id="B900FC1FFFD6390C2A9177A6FB13FC71" bold="true" box="[1153,1215,938,962]" pageId="27" pageNumber="26">
<collectingCountry id="F363609DFFD6390C2A9177A6FB13FC71" box="[1153,1215,938,962]" name="Cuba" pageId="27" pageNumber="26">Cuba</collectingCountry>
</emphasis>
,
<collectingRegion id="49B0EEEFFFD6390C2AC377A7FA13FC70" box="[1235,1471,939,963]" country="Cuba" name="Mayabeque" pageId="27" pageNumber="26">Mayabeque Province</collectingRegion>
, border between
<location id="8EAB76D6FFD6390C2DED77C7FA84FC50" LSID="urn:lsid:plazi:treatment:03DD911BFFD639302DEB74E7FC70F87A:8EAB76D6FFD6390C2DED77C7FA84FC50" box="[1021,1320,971,995]" country="Cuba" name="San Jose de Las Lajas" pageId="27" pageNumber="26" stateProvince="Mayabeque">San José de Las Lajas</location>
and Jaruco Municipalities, Tapaste, Escaleras de Jaruco, La Jaula, Cueva del Indio,
<elevation id="0059C73EFFD6390C2DB47007FB97FB90" box="[932,1083,1035,1059]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="27" pageNumber="26" unit="m" value="190.0">
<quantity id="4C8C8DE8FFD6390C2DB47007FC47FB91" box="[932,1003,1035,1059]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="27" pageNumber="26" unit="m" value="190.0">190 m</quantity>
a. s. l.
</elevation>
,
<date id="FFCA06CDFFD6390C2A5A7007FAB3FB90" box="[1098,1311,1035,1059]" pageId="27" pageNumber="26" value="1995-12-23">
<collectingDate id="EF8EFF25FFD6390C2A5A7007FAB3FB90" box="[1098,1311,1035,1059]" pageId="27" pageNumber="26" value="1995-12-23">23 December 1995</collectingDate>
</date>
, leg.
<collectorName id="268145DBFFD6390C2B4C7007FA12FB90" box="[1372,1470,1035,1059]" pageId="27" pageNumber="26">A. Pérez</collectorName>
,
<specimenCount id="9D72EB84FFD6390C2D207027FCFEFBF0" box="[816,850,1067,1091]" pageId="27" pageNumber="26" type="male">1♂</specimenCount>
(topotype), IES,
<date id="FFCA06CDFFD6390C2A1D7027FB4DFBF0" box="[1037,1249,1067,1091]" pageId="27" pageNumber="26" value="1999-09-27">
<collectingDate id="EF8EFF25FFD6390C2A1D7027FB4DFBF0" box="[1037,1249,1067,1091]" pageId="27" pageNumber="26" value="1999-09-27">27 September 1999</collectingDate>
</date>
, leg.
<collectorName id="268145DBFFD6390C2B0C7027FA39FBF1" box="[1308,1429,1067,1090]" pageId="27" pageNumber="26">E. Fonseca</collectorName>
,
<specimenCount id="9D72EB84FFD6390C2BB17027FA6EFBF0" box="[1441,1474,1067,1091]" pageId="27" pageNumber="26" type="female">1♀</specimenCount>
(topotype), IES
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E7386FFD6390C2D207067FCD4FA90" pageId="27" pageNumber="26" type="description">
<paragraph id="8BCB200DFFD6390C2D207067FCD4FA90" blockId="27.[816,1474,938,1315]" pageId="27" pageNumber="26">
<emphasis id="B900FC1FFFD6390C2D207067FCCBFB30" bold="true" box="[816,871,1131,1155]" pageId="27" pageNumber="26">Note</emphasis>
. Both specimens were repeatedly seen by one of us (RT) between 2005 and 2011, but are currently missing from IES collection. As their whereabouts is unknown (there is no official record of a loan, destruction or loss in the collection log), both were excluded from the
<typeStatus id="54CF9EAFFFD6390C2B8270E7FA6DFAB0" box="[1426,1473,1259,1283]" pageId="27" pageNumber="26">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C36E7386FFD6390C2D207147FB55F9B0" pageId="27" pageNumber="26" type="etymology">
<paragraph id="8BCB200DFFD6390C2D207147FB55F9B0" blockId="27.[816,1475,1355,1539]" pageId="27" pageNumber="26">
ETYMOLOGY. The selected epithet is a Latinized matronym honoring the senior author’s wife Sheyla Yong, who enthusiastically assisted us all along the present revision, both in the field (she actually collected several of the
<typeStatus id="54CF9EAFFFD6390C2B8471A7FA6EFA70" box="[1428,1474,1451,1475]" pageId="27" pageNumber="26">type</typeStatus>
specimens) and in the lab (part of the data gathering and processing, as well as photographic work).
</paragraph>
</subSubSection>
<subSubSection id="C36E7386FFD6390B2D207227FB6FFAA8" lastPageId="28" lastPageNumber="27" pageId="27" pageNumber="26" type="diagnosis">
<paragraph id="8BCB200DFFD6390B2D207227FB6FFAA8" blockId="27.[816,1475,1579,2019]" lastBlockId="28.[816,1474,1187,1308]" lastPageId="28" lastPageNumber="27" pageId="27" pageNumber="26">
DIAGNOSIS. Adult size moderately large for the genus (
<quantity id="4C8C8DE8FFD6390C2B887227FCD3F9D1" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="27" pageNumber="26" unit="mm" value="25.5" valueMax="27.0" valueMin="24.0">24– 27 mm</quantity>
in males,
<quantity id="4C8C8DE8FFD6390C2DE77247FBC3F9D1" box="[1015,1135,1611,1635]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="27" pageNumber="26" unit="mm" value="33.0" valueMax="34.0" valueMin="32.0">32–34 mm</quantity>
in females). Adult coloration: base light yellowish brown, slightly darker on metasomal segments IV–V and slightly paler on legs, venter and telson; dark patterns essentially absent, except for faint infuscation in carapace (mostly around eyes) and metasomal segments II–IV (in some specimens only, mostly females). Juvenile coloration: base pale yellowish brown, paler on pedipalp chelae, legs, venter and telson; dark patterns essentially absent, except for diffuse gray to dark brown reticulation in chelicerae, carapace (along anterior margin and around eyes) and metasomal segments II–V (mostly on posterior part and around carinae); tergites with posterior margins translucent whitish; pedipalps fingers infuscate. Pedipalps remarkably elongate and robust for the genus (length/width ratio of femur, patella and chela: 2.94–3.11, 2.58–2.65 and
<quantity id="4C8C8DE8FFD1390B2E6170EFFF50FB48" box="[113,252,1251,1275]" metricMagnitude="-2" metricUnit="m" metricValue="9.0932" metricValueMax="9.2964" metricValueMin="8.89" pageId="28" pageNumber="27" unit="in" value="3.58" valueMax="3.66" valueMin="3.5">3.50–3.66 in</quantity>
males, 2.79–2.96, 2.64–2.71 and
<quantity id="4C8C8DE8FFD1390B2C6870EFFCAFFB48" box="[632,771,1251,1275]" metricMagnitude="-2" metricUnit="m" metricValue="8.9916" metricValueMax="9.0678" metricValueMin="8.9154" pageId="28" pageNumber="27" unit="in" value="3.54" valueMax="3.57" valueMin="3.51">3.51–3.57 in</quantity>
females); chelae with manus incrassate oval and very slightly longer in males (length/width ratio:
<quantity id="4C8C8DE8FFD1390B2C0F712FFD03FA88" box="[543,687,1315,1339]" metricMagnitude="-2" metricUnit="m" metricValue="3.8862" metricValueMax="3.9624" metricValueMin="3.81" pageId="28" pageNumber="27" unit="in" value="1.53" valueMax="1.56" valueMin="1.5">1.50–1.56 in</quantity>
males,
<quantity id="4C8C8DE8FFD1390B2E61714FFF50FAE8" box="[113,252,1347,1371]" metricMagnitude="-2" metricUnit="m" metricValue="3.7973" metricValueMax="3.8862" metricValueMin="3.7084" pageId="28" pageNumber="27" unit="in" value="1.495" valueMax="1.53" valueMin="1.46">1.46–1.53 in</quantity>
females) and with fingers very long (movable finger/manus length ratio:
<quantity id="4C8C8DE8FFD1390B2F87716FFD8DFAC8" box="[407,545,1379,1403]" metricMagnitude="-2" metricUnit="m" metricValue="3.3909" metricValueMax="3.4036" metricValueMin="3.3782" pageId="28" pageNumber="27" unit="in" value="1.335" valueMax="1.34" valueMin="1.33">1.33–1.34 in</quantity>
males,
<quantity id="4C8C8DE8FFD1390B2C69716FFCAFFAC8" box="[633,771,1379,1403]" metricMagnitude="-2" metricUnit="m" metricValue="3.4671" metricValueMax="3.5559999999999996" metricValueMin="3.3782" pageId="28" pageNumber="27" unit="in" value="1.365" valueMax="1.4" valueMin="1.33">1.33–1.40 in</quantity>
females), fixed/movable fingers with 12–13/12–13 principal rows of denticles (mode 12/12). Tegument of carapace, tergites, sternites, metasoma and telson sexually dimorphic between adults: matt in male
<emphasis id="B900FC1FFFD1390B2FC971E8FE5AFA48" box="[473,502,1508,1531]" italics="true" pageId="28" pageNumber="27">vs.</emphasis>
with a satin sheen in females. Carapace and tergites with many medium-sized to coarse granules scattered (similar in both sexes). Pectines with 13–16 teeth in males,
<quantity id="4C8C8DE8FFD1390B2FB2724FFDA8F9E8" box="[418,516,1603,1627]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="28" pageNumber="27" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
females (mode 14 and 13–14, respectively); basal middle lamella slightly enlarged and teardrop-shaped in males, greatly enlarged and widely oval in females. Sternite V with median smooth patch widely cordiform and about as long as wide in both sexes, very large in males and moderately large in females; lateral smooth patches oval, oblique and bulky in both sexes, moderately large in males and small in females. Metasoma slightly elongate in both sexes (length/width ratio of segments I–V: 1.12–1.16, 1.42–1.44, 1.54–1.58, 1.76–1.78 and
<quantity id="4C8C8DE8FFD1390B2C86736FFF2AF828" metricMagnitude="-2" metricUnit="m" metricValue="5.6769" metricValueMax="5.715" metricValueMin="5.6388" pageId="28" pageNumber="27" unit="in" value="2.235" valueMax="2.25" valueMin="2.22">2.22–2.25 in</quantity>
males
<emphasis id="B900FC1FFFD1390B2EC47388FF5DF828" box="[212,241,1924,1947]" italics="true" pageId="28" pageNumber="27">vs.</emphasis>
1.14–1.15, 1.43–1.45, 1.54–1.64, 1.70–1.71 and
<quantity id="4C8C8DE8FFD1390B2E6173AFFF50F808" box="[113,252,1955,1979]" metricMagnitude="-2" metricUnit="m" metricValue="5.6642" metricValueMax="5.6896" metricValueMin="5.6388" pageId="28" pageNumber="27" unit="in" value="2.23" valueMax="2.24" valueMin="2.22">2.22–2.24 in</quantity>
females), with 10/10/8/8/7 complete to almost complete, coarsely serrate to serrato-crenulate carinae; lateral inframedians on II obsolete only on basal one-sixth (males) to one-fifth (females); intercarinal tegument of segment V with many small (males) to medium-sized (females) conical granules. Telson with vesicle less globose than standard for the genus, subaculear tubercle small.
</paragraph>
</subSubSection>
<paragraph id="8BCB200DFFD1390B2C1474FAFACBFB8E" pageId="28" pageNumber="27">
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<tr id="3544224FFFD1C6E82E6A74FAFA08FEBD" box="[122,1444,246,270]" gridrow="0" pageId="28" pageNumber="27" rowspan-0="1" rowspan-1="1">
<th id="76954B33FFD1C6E82C1174FAFD15FEBD" box="[513,697,246,270]" gridcol="2" gridrow="0" pageId="28" pageNumber="27">
<taxonomicName id="4C745B8EFFD1390B2C1474FAFDDEFEBD" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[516,626,246,270]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFD1390B2C1474FAFDDEFEBD" bold="true" box="[516,626,246,270]" italics="true" pageId="28" pageNumber="27">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFD1390B2C6D74FAFD15FEBD" box="[637,697,246,270]" pageId="28" pageNumber="27" rank="species">
<emphasis id="B900FC1FFFD1390B2C6D74FAFD3AFEBD" bold="true" box="[637,662,246,270]" pageId="28" pageNumber="27">sp</emphasis>
.
<emphasis id="B900FC1FFFD1390B2CB374FAFD1EFEBD" bold="true" box="[675,690,246,270]" pageId="28" pageNumber="27">n</emphasis>
.
</taxonomicNameLabel>
</th>
<th id="76954B33FFD1C6E82CE974FAFC1EFEBD" box="[761,946,246,270]" gridcol="3" gridrow="0" pageId="28" pageNumber="27">
<taxonomicName id="4C745B8EFFD1390B2CED74FAFCC7FEBD" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[765,875,246,270]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFD1390B2CED74FAFCC7FEBD" bold="true" box="[765,875,246,270]" italics="true" pageId="28" pageNumber="27">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFD1390B2D6674FAFC1EFEBD" box="[886,946,246,270]" pageId="28" pageNumber="27" rank="species">
<emphasis id="B900FC1FFFD1390B2D6674FAFC23FEBD" bold="true" box="[886,911,246,270]" pageId="28" pageNumber="27">sp</emphasis>
.
<emphasis id="B900FC1FFFD1390B2D8C74FAFC07FEBD" bold="true" box="[924,939,246,270]" pageId="28" pageNumber="27">n</emphasis>
.
</taxonomicNameLabel>
</th>
<th id="76954B33FFD1C6E82DE274FAFB07FEBD" box="[1010,1195,246,270]" gridcol="4" gridrow="0" pageId="28" pageNumber="27">
<taxonomicName id="4C745B8EFFD1390B2DE674FAFBCFFEBD" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[1014,1123,246,270]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFD1390B2DE674FAFBCFFEBD" bold="true" box="[1014,1123,246,270]" italics="true" pageId="28" pageNumber="27">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFD1390B2A7F74FAFB07FEBD" box="[1135,1195,246,270]" pageId="28" pageNumber="27" rank="species">
<emphasis id="B900FC1FFFD1390B2A7F74FAFB24FEBD" bold="true" box="[1135,1160,246,270]" pageId="28" pageNumber="27">sp</emphasis>
.
<emphasis id="B900FC1FFFD1390B2A8574FAFB08FEBD" bold="true" box="[1173,1188,246,270]" pageId="28" pageNumber="27">n</emphasis>
.
</taxonomicNameLabel>
</th>
<th id="76954B33FFD1C6E82AFB74FAFA08FEBD" box="[1259,1444,246,270]" gridcol="5" gridrow="0" pageId="28" pageNumber="27">
<taxonomicName id="4C745B8EFFD1390B2AFF74FAFAF0FEBD" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[1263,1372,246,270]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFD1390B2AFF74FAFAF0FEBD" bold="true" box="[1263,1372,246,270]" italics="true" pageId="28" pageNumber="27">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFD1390B2B7874FAFA08FEBD" box="[1384,1444,246,270]" pageId="28" pageNumber="27" rank="species">
<emphasis id="B900FC1FFFD1390B2B7874FAFA2DFEBD" bold="true" box="[1384,1409,246,270]" pageId="28" pageNumber="27">sp</emphasis>
.
<emphasis id="B900FC1FFFD1390B2B9E74FAFA31FEBD" bold="true" box="[1422,1437,246,270]" pageId="28" pageNumber="27">n</emphasis>
.
</taxonomicNameLabel>
</th>
</tr>
<tr id="3544224FFFD1C6E82E6A7512FA08FE8A" box="[122,1444,286,313]" gridrow="1" pageId="28" pageNumber="27" rowspan-1="1">
<th id="76954B33FFD1C6E82E6A7512FEEAFE8A" box="[122,326,286,313]" gridcol="0" gridrow="1" pageId="28" pageNumber="27">
<emphasis id="B900FC1FFFD1390B2E6A752CFEEAFE8B" bold="true" box="[122,326,288,312]" pageId="28" pageNumber="27">Dimensions (MM)</emphasis>
</th>
<td id="76954B33FFD1C6E82C117512FD15FE8A" box="[513,697,286,313]" gridcol="2" gridrow="1" pageId="28" pageNumber="27">
<emphasis id="B900FC1FFFD1390B2C317512FD30FE8A" bold="true" box="[545,668,286,313]" pageId="28" pageNumber="27">♂ holotype</emphasis>
</td>
<td id="76954B33FFD1C6E82CE97512FC1EFE8A" box="[761,946,286,313]" gridcol="3" gridrow="1" pageId="28" pageNumber="27">
<emphasis id="B900FC1FFFD1390B2CEE7512FC1CFE8A" bold="true" box="[766,944,286,313]" pageId="28" pageNumber="27">♂ paratopotype</emphasis>
</td>
<td id="76954B33FFD1C6E82DE27512FB07FE8A" box="[1010,1195,286,313]" gridcol="4" gridrow="1" pageId="28" pageNumber="27">
<emphasis id="B900FC1FFFD1390B2DE77512FB04FE8A" bold="true" box="[1015,1192,286,313]" pageId="28" pageNumber="27">♀ paratopotype</emphasis>
</td>
<td id="76954B33FFD1C6E82AFB7512FA08FE8A" box="[1259,1444,286,313]" gridcol="5" gridrow="1" pageId="28" pageNumber="27">
<emphasis id="B900FC1FFFD1390B2AE07512FA0DFE8A" bold="true" box="[1264,1441,286,313]" pageId="28" pageNumber="27">♀ paratopotype</emphasis>
</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7545FA08FED2" box="[122,1444,329,353]" gridrow="2" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7545FEEAFED2" box="[122,326,329,353]" gridcol="0" gridrow="2" pageId="28" pageNumber="27">Carapace</th>
<td id="76954B33FFD1C6E82F7E7545FE7FFED2" box="[366,467,329,353]" gridcol="1" gridrow="2" pageId="28" pageNumber="27">L / W</td>
<td id="76954B33FFD1C6E82C117545FD15FED2" box="[513,697,329,353]" gridcol="2" gridrow="2" pageId="28" pageNumber="27">3.15 / 3.40</td>
<td id="76954B33FFD1C6E82CE97545FC1EFED2" box="[761,946,329,353]" gridcol="3" gridrow="2" pageId="28" pageNumber="27">2.90 / 3.10</td>
<td id="76954B33FFD1C6E82DE27545FB07FED2" box="[1010,1195,329,353]" gridcol="4" gridrow="2" pageId="28" pageNumber="27">3.87 / 4.10</td>
<td id="76954B33FFD1C6E82AFB7545FA08FED2" box="[1259,1444,329,353]" gridcol="5" gridrow="2" pageId="28" pageNumber="27">4.07 / 4.25</td>
</tr>
<tr id="3544224FFFD1C6E82E6A757DFA08FE3A" box="[122,1444,369,393]" gridrow="3" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A757DFEEAFE3A" box="[122,326,369,393]" gridcol="0" gridrow="3" pageId="28" pageNumber="27">Mesosoma</th>
<td id="76954B33FFD1C6E82F7E757DFE7FFE3A" box="[366,467,369,393]" gridcol="1" gridrow="3" pageId="28" pageNumber="27">L</td>
<td id="76954B33FFD1C6E82C11757DFD15FE3A" box="[513,697,369,393]" gridcol="2" gridrow="3" pageId="28" pageNumber="27">6.90</td>
<td id="76954B33FFD1C6E82CE9757DFC1EFE3A" box="[761,946,369,393]" gridcol="3" gridrow="3" pageId="28" pageNumber="27">5.65</td>
<td id="76954B33FFD1C6E82DE2757DFB07FE3A" box="[1010,1195,369,393]" gridcol="4" gridrow="3" pageId="28" pageNumber="27">8.70</td>
<td id="76954B33FFD1C6E82AFB757DFA08FE3A" box="[1259,1444,369,393]" gridcol="5" gridrow="3" pageId="28" pageNumber="27">9.75</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7596FA08FE00" box="[122,1444,410,435]" gridrow="4" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7596FEEAFE00" box="[122,326,410,435]" gridcol="0" gridrow="4" pageId="28" pageNumber="27">Tergite VII</th>
<td id="76954B33FFD1C6E82F7E7596FE7FFE00" box="[366,467,410,435]" gridcol="1" gridrow="4" pageId="28" pageNumber="27">L / W</td>
<td id="76954B33FFD1C6E82C117596FD15FE00" box="[513,697,410,435]" gridcol="2" gridrow="4" pageId="28" pageNumber="27">2.05 / 3.20</td>
<td id="76954B33FFD1C6E82CE97596FC1EFE00" box="[761,946,410,435]" gridcol="3" gridrow="4" pageId="28" pageNumber="27">1.80 / 2.85</td>
<td id="76954B33FFD1C6E82DE27596FB07FE00" box="[1010,1195,410,435]" gridcol="4" gridrow="4" pageId="28" pageNumber="27">2.70 / 4.32</td>
<td id="76954B33FFD1C6E82AFB7596FA08FE00" box="[1259,1444,410,435]" gridcol="5" gridrow="4" pageId="28" pageNumber="27">2.67 / 4.40</td>
</tr>
<tr id="3544224FFFD1C6E82E6A75CEFA08FE68" box="[122,1444,450,475]" gridrow="5" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A75CEFEEAFE68" box="[122,326,450,475]" gridcol="0" gridrow="5" pageId="28" pageNumber="27">Metasoma</th>
<td id="76954B33FFD1C6E82F7E75CEFE7FFE68" box="[366,467,450,475]" gridcol="1" gridrow="5" pageId="28" pageNumber="27">L</td>
<td id="76954B33FFD1C6E82C1175CEFD15FE68" box="[513,697,450,475]" gridcol="2" gridrow="5" pageId="28" pageNumber="27">13.31</td>
<td id="76954B33FFD1C6E82CE975CEFC1EFE68" box="[761,946,450,475]" gridcol="3" gridrow="5" pageId="28" pageNumber="27">12.08</td>
<td id="76954B33FFD1C6E82DE275CEFB07FE68" box="[1010,1195,450,475]" gridcol="4" gridrow="5" pageId="28" pageNumber="27">15.44</td>
<td id="76954B33FFD1C6E82AFB75CEFA08FE68" box="[1259,1444,450,475]" gridcol="5" gridrow="5" pageId="28" pageNumber="27">15.94</td>
</tr>
<tr id="3544224FFFD1C6E82E6A75E7FA08FDB7" box="[122,1444,491,516]" gridrow="6" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A75E7FEEAFDB7" box="[122,326,491,516]" gridcol="0" gridrow="6" pageId="28" pageNumber="27">Segment I</th>
<td id="76954B33FFD1C6E82F7E75E7FE7FFDB7" box="[366,467,491,516]" gridcol="1" gridrow="6" pageId="28" pageNumber="27">L / W / D</td>
<td id="76954B33FFD1C6E82C1175E7FD15FDB7" box="[513,697,491,516]" gridcol="2" gridrow="6" pageId="28" pageNumber="27">2.05 / 1.77 / 1.60</td>
<td id="76954B33FFD1C6E82CE975E7FC1EFDB7" box="[761,946,491,516]" gridcol="3" gridrow="6" pageId="28" pageNumber="27">1.80 / 1.61 / 1.45</td>
<td id="76954B33FFD1C6E82DE275E7FB07FDB7" box="[1010,1195,491,516]" gridcol="4" gridrow="6" pageId="28" pageNumber="27">2.35 / 2.05 / 1.75</td>
<td id="76954B33FFD1C6E82AFB75E7FA08FDB7" box="[1259,1444,491,516]" gridcol="5" gridrow="6" pageId="28" pageNumber="27">2.52 / 2.20 / 1.87</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7618FA08FD9E" box="[122,1444,532,557]" gridrow="7" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7618FEEAFD9E" box="[122,326,532,557]" gridcol="0" gridrow="7" pageId="28" pageNumber="27">Segment II</th>
<td id="76954B33FFD1C6E82F7E7618FE7FFD9E" box="[366,467,532,557]" gridcol="1" gridrow="7" pageId="28" pageNumber="27">L / W / D</td>
<td id="76954B33FFD1C6E82C117618FD15FD9E" box="[513,697,532,557]" gridcol="2" gridrow="7" pageId="28" pageNumber="27">2.45 / 1.70 / 1.50</td>
<td id="76954B33FFD1C6E82CE97618FC1EFD9E" box="[761,946,532,557]" gridcol="3" gridrow="7" pageId="28" pageNumber="27">2.25 / 1.58 / 1.40</td>
<td id="76954B33FFD1C6E82DE27618FB07FD9E" box="[1010,1195,532,557]" gridcol="4" gridrow="7" pageId="28" pageNumber="27">2.82 / 1.95 / 1.70</td>
<td id="76954B33FFD1C6E82AFB7618FA08FD9E" box="[1259,1444,532,557]" gridcol="5" gridrow="7" pageId="28" pageNumber="27">2.95 / 2.07 / 1.77</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7630FA08FDE6" box="[122,1444,572,597]" gridrow="8" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7630FEEAFDE6" box="[122,326,572,597]" gridcol="0" gridrow="8" pageId="28" pageNumber="27">Segment III</th>
<td id="76954B33FFD1C6E82F7E7630FE7FFDE6" box="[366,467,572,597]" gridcol="1" gridrow="8" pageId="28" pageNumber="27">L / W / D</td>
<td id="76954B33FFD1C6E82C117630FD15FDE6" box="[513,697,572,597]" gridcol="2" gridrow="8" pageId="28" pageNumber="27">2.62 / 1.70 / 1.47</td>
<td id="76954B33FFD1C6E82CE97630FC1EFDE6" box="[761,946,572,597]" gridcol="3" gridrow="8" pageId="28" pageNumber="27">2.45 / 1.55 / 1.40</td>
<td id="76954B33FFD1C6E82DE27630FB07FDE6" box="[1010,1195,572,597]" gridcol="4" gridrow="8" pageId="28" pageNumber="27">3.15 / 1.92 / 1.70</td>
<td id="76954B33FFD1C6E82AFB7630FA08FDE6" box="[1259,1444,572,597]" gridcol="5" gridrow="8" pageId="28" pageNumber="27">3.15 / 2.05 / 1.75</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7669FA08FDCD" box="[122,1444,613,638]" gridrow="9" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7669FEEAFDCD" box="[122,326,613,638]" gridcol="0" gridrow="9" pageId="28" pageNumber="27">Segment IV</th>
<td id="76954B33FFD1C6E82F7E7669FE7FFDCD" box="[366,467,613,638]" gridcol="1" gridrow="9" pageId="28" pageNumber="27">L / W / D</td>
<td id="76954B33FFD1C6E82C117669FD15FDCD" box="[513,697,613,638]" gridcol="2" gridrow="9" pageId="28" pageNumber="27">2.82 / 1.60 / 1.42</td>
<td id="76954B33FFD1C6E82CE97669FC1EFDCD" box="[761,946,613,638]" gridcol="3" gridrow="9" pageId="28" pageNumber="27">2.58 / 1.45 / 1.35</td>
<td id="76954B33FFD1C6E82DE27669FB07FDCD" box="[1010,1195,613,638]" gridcol="4" gridrow="9" pageId="28" pageNumber="27">3.20 / 1.87 / 1.65</td>
<td id="76954B33FFD1C6E82AFB7669FA08FDCD" box="[1259,1444,613,638]" gridcol="5" gridrow="9" pageId="28" pageNumber="27">3.32 / 1.95 / 1.75</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7682FA08FD14" box="[122,1444,654,679]" gridrow="10" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7682FEEAFD14" box="[122,326,654,679]" gridcol="0" gridrow="10" pageId="28" pageNumber="27">Segment V</th>
<td id="76954B33FFD1C6E82F7E7682FE7FFD14" box="[366,467,654,679]" gridcol="1" gridrow="10" pageId="28" pageNumber="27">L / W / D</td>
<td id="76954B33FFD1C6E82C117682FD15FD14" box="[513,697,654,679]" gridcol="2" gridrow="10" pageId="28" pageNumber="27">3.37 / 1.50 / 1.45</td>
<td id="76954B33FFD1C6E82CE97682FC1EFD14" box="[761,946,654,679]" gridcol="3" gridrow="10" pageId="28" pageNumber="27">3.00 / 1.35 / 1.30</td>
<td id="76954B33FFD1C6E82DE27682FB07FD14" box="[1010,1195,654,679]" gridcol="4" gridrow="10" pageId="28" pageNumber="27">3.92 / 1.75 / 1.62</td>
<td id="76954B33FFD1C6E82AFB7682FA08FD14" box="[1259,1444,654,679]" gridcol="5" gridrow="10" pageId="28" pageNumber="27">4.00 / 1.80 / 1.70</td>
</tr>
<tr id="3544224FFFD1C6E82E6A76BBFA08FD7C" box="[122,1444,695,719]" gridrow="11" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A76BBFEEAFD7C" box="[122,326,695,719]" gridcol="0" gridrow="11" pageId="28" pageNumber="27">Telson</th>
<td id="76954B33FFD1C6E82F7E76BBFE7FFD7C" box="[366,467,695,719]" gridcol="1" gridrow="11" pageId="28" pageNumber="27">L</td>
<td id="76954B33FFD1C6E82C1176BBFD15FD7C" box="[513,697,695,719]" gridcol="2" gridrow="11" pageId="28" pageNumber="27">3.40</td>
<td id="76954B33FFD1C6E82CE976BBFC1EFD7C" box="[761,946,695,719]" gridcol="3" gridrow="11" pageId="28" pageNumber="27">2.99</td>
<td id="76954B33FFD1C6E82DE276BBFB07FD7C" box="[1010,1195,695,719]" gridcol="4" gridrow="11" pageId="28" pageNumber="27">4.10</td>
<td id="76954B33FFD1C6E82AFB76BBFA08FD7C" box="[1259,1444,695,719]" gridcol="5" gridrow="11" pageId="28" pageNumber="27">4.24</td>
</tr>
<tr id="3544224FFFD1C6E82E6A76D3FA08FD4B" box="[122,1444,735,760]" gridrow="12" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A76D3FEEAFD4B" box="[122,326,735,760]" gridcol="0" gridrow="12" pageId="28" pageNumber="27">Vesicle</th>
<td id="76954B33FFD1C6E82F7E76D3FE7FFD4B" box="[366,467,735,760]" gridcol="1" gridrow="12" pageId="28" pageNumber="27">L / W / D</td>
<td id="76954B33FFD1C6E82C1176D3FD15FD4B" box="[513,697,735,760]" gridcol="2" gridrow="12" pageId="28" pageNumber="27">1.95 / 1.32 / 1.30</td>
<td id="76954B33FFD1C6E82CE976D3FC1EFD4B" box="[761,946,735,760]" gridcol="3" gridrow="12" pageId="28" pageNumber="27">1.65 / 1.16 / 1.16</td>
<td id="76954B33FFD1C6E82DE276D3FB07FD4B" box="[1010,1195,735,760]" gridcol="4" gridrow="12" pageId="28" pageNumber="27">2.35 / 1.60 / 1.57</td>
<td id="76954B33FFD1C6E82AFB76D3FA08FD4B" box="[1259,1444,735,760]" gridcol="5" gridrow="12" pageId="28" pageNumber="27">2.37 / 1.65 / 1.62</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7704FA08FC93" box="[122,1444,776,800]" gridrow="13" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7704FEEAFC93" box="[122,326,776,800]" gridcol="0" gridrow="13" pageId="28" pageNumber="27">Aculeus</th>
<td id="76954B33FFD1C6E82F7E7704FE7FFC93" box="[366,467,776,800]" gridcol="1" gridrow="13" pageId="28" pageNumber="27">L</td>
<td id="76954B33FFD1C6E82C117704FD15FC93" box="[513,697,776,800]" gridcol="2" gridrow="13" pageId="28" pageNumber="27">1.45</td>
<td id="76954B33FFD1C6E82CE97704FC1EFC93" box="[761,946,776,800]" gridcol="3" gridrow="13" pageId="28" pageNumber="27">1.34</td>
<td id="76954B33FFD1C6E82DE27704FB07FC93" box="[1010,1195,776,800]" gridcol="4" gridrow="13" pageId="28" pageNumber="27">1.75</td>
<td id="76954B33FFD1C6E82AFB7704FA08FC93" box="[1259,1444,776,800]" gridcol="5" gridrow="13" pageId="28" pageNumber="27">1.87</td>
</tr>
<tr id="3544224FFFD1C6E82E6A773CFA08FCFA" box="[122,1444,816,841]" gridrow="14" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A773CFEEAFCFA" box="[122,326,816,841]" gridcol="0" gridrow="14" pageId="28" pageNumber="27">Pedipalp</th>
<td id="76954B33FFD1C6E82F7E773CFE7FFCFA" box="[366,467,816,841]" gridcol="1" gridrow="14" pageId="28" pageNumber="27">L</td>
<td id="76954B33FFD1C6E82C11773CFD15FCFA" box="[513,697,816,841]" gridcol="2" gridrow="14" pageId="28" pageNumber="27">12.17</td>
<td id="76954B33FFD1C6E82CE9773CFC1EFCFA" box="[761,946,816,841]" gridcol="3" gridrow="14" pageId="28" pageNumber="27">11.00</td>
<td id="76954B33FFD1C6E82DE2773CFB07FCFA" box="[1010,1195,816,841]" gridcol="4" gridrow="14" pageId="28" pageNumber="27">14.64</td>
<td id="76954B33FFD1C6E82AFB773CFA08FCFA" box="[1259,1444,816,841]" gridcol="5" gridrow="14" pageId="28" pageNumber="27">14.95</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7755FA08FCC1" box="[122,1444,857,882]" gridrow="15" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7755FEEAFCC1" box="[122,326,857,882]" gridcol="0" gridrow="15" pageId="28" pageNumber="27">Femur</th>
<td id="76954B33FFD1C6E82F7E7755FE7FFCC1" box="[366,467,857,882]" gridcol="1" gridrow="15" pageId="28" pageNumber="27">L / W</td>
<td id="76954B33FFD1C6E82C117755FD15FCC1" box="[513,697,857,882]" gridcol="2" gridrow="15" pageId="28" pageNumber="27">2.95 / 0.95</td>
<td id="76954B33FFD1C6E82CE97755FC1EFCC1" box="[761,946,857,882]" gridcol="3" gridrow="15" pageId="28" pageNumber="27">2.65 / 0.90</td>
<td id="76954B33FFD1C6E82DE27755FB07FCC1" box="[1010,1195,857,882]" gridcol="4" gridrow="15" pageId="28" pageNumber="27">3.40 / 1.22</td>
<td id="76954B33FFD1C6E82AFB7755FA08FCC1" box="[1259,1444,857,882]" gridcol="5" gridrow="15" pageId="28" pageNumber="27">3.55 / 1.20</td>
</tr>
<tr id="3544224FFFD1C6E82E6A778DFA08FC29" box="[122,1444,897,922]" gridrow="16" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A778DFEEAFC29" box="[122,326,897,922]" gridcol="0" gridrow="16" pageId="28" pageNumber="27">Patella</th>
<td id="76954B33FFD1C6E82F7E778DFE7FFC29" box="[366,467,897,922]" gridcol="1" gridrow="16" pageId="28" pageNumber="27">L / W</td>
<td id="76954B33FFD1C6E82C11778DFD15FC29" box="[513,697,897,922]" gridcol="2" gridrow="16" pageId="28" pageNumber="27">3.37 / 1.27</td>
<td id="76954B33FFD1C6E82CE9778DFC1EFC29" box="[761,946,897,922]" gridcol="3" gridrow="16" pageId="28" pageNumber="27">3.10 / 1.20</td>
<td id="76954B33FFD1C6E82DE2778DFB07FC29" box="[1010,1195,897,922]" gridcol="4" gridrow="16" pageId="28" pageNumber="27">4.10 / 1.55</td>
<td id="76954B33FFD1C6E82AFB778DFA08FC29" box="[1259,1444,897,922]" gridcol="5" gridrow="16" pageId="28" pageNumber="27">4.20 / 1.55</td>
</tr>
<tr id="3544224FFFD1C6E82E6A77A6FA08FC70" box="[122,1444,938,963]" gridrow="17" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A77A6FEEAFC70" box="[122,326,938,963]" gridcol="0" gridrow="17" pageId="28" pageNumber="27">Chela</th>
<td id="76954B33FFD1C6E82F7E77A6FE7FFC70" box="[366,467,938,963]" gridcol="1" gridrow="17" pageId="28" pageNumber="27">L</td>
<td id="76954B33FFD1C6E82C1177A6FD15FC70" box="[513,697,938,963]" gridcol="2" gridrow="17" pageId="28" pageNumber="27">5.85</td>
<td id="76954B33FFD1C6E82CE977A6FC1EFC70" box="[761,946,938,963]" gridcol="3" gridrow="17" pageId="28" pageNumber="27">5.25</td>
<td id="76954B33FFD1C6E82DE277A6FB07FC70" box="[1010,1195,938,963]" gridcol="4" gridrow="17" pageId="28" pageNumber="27">7.14</td>
<td id="76954B33FFD1C6E82AFB77A6FA08FC70" box="[1259,1444,938,963]" gridcol="5" gridrow="17" pageId="28" pageNumber="27">7.20</td>
</tr>
<tr id="3544224FFFD1C6E82E6A77DFFA08FC5F" box="[122,1444,979,1004]" gridrow="18" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A77DFFEEAFC5F" box="[122,326,979,1004]" gridcol="0" gridrow="18" pageId="28" pageNumber="27">Manus</th>
<td id="76954B33FFD1C6E82F7E77DFFE7FFC5F" box="[366,467,979,1004]" gridcol="1" gridrow="18" pageId="28" pageNumber="27">W / D</td>
<td id="76954B33FFD1C6E82C1177DFFD15FC5F" box="[513,697,979,1004]" gridcol="2" gridrow="18" pageId="28" pageNumber="27">2.50 / 1.60 / 1.52</td>
<td id="76954B33FFD1C6E82CE977DFFC1EFC5F" box="[761,946,979,1004]" gridcol="3" gridrow="18" pageId="28" pageNumber="27">2.25 / 1.50 / 1.44</td>
<td id="76954B33FFD1C6E82DE277DFFB07FC5F" box="[1010,1195,979,1004]" gridcol="4" gridrow="18" pageId="28" pageNumber="27">3.07 / 2.00 / 1.87</td>
<td id="76954B33FFD1C6E82AFB77DFFA08FC5F" box="[1259,1444,979,1004]" gridcol="5" gridrow="18" pageId="28" pageNumber="27">3.00 / 2.05 / 1.80</td>
</tr>
<tr id="3544224FFFD1C6E82E6A77F7FA08FBA7" box="[122,1444,1019,1044]" gridrow="19" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A77F7FEEAFBA7" box="[122,326,1019,1044]" gridcol="0" gridrow="19" pageId="28" pageNumber="27">Movable finger</th>
<td id="76954B33FFD1C6E82F7E77F7FE7FFBA7" box="[366,467,1019,1044]" gridcol="1" gridrow="19" pageId="28" pageNumber="27">L</td>
<td id="76954B33FFD1C6E82C1177F7FD15FBA7" box="[513,697,1019,1044]" gridcol="2" gridrow="19" pageId="28" pageNumber="27">3.35</td>
<td id="76954B33FFD1C6E82CE977F7FC1EFBA7" box="[761,946,1019,1044]" gridcol="3" gridrow="19" pageId="28" pageNumber="27">3.00</td>
<td id="76954B33FFD1C6E82DE277F7FB07FBA7" box="[1010,1195,1019,1044]" gridcol="4" gridrow="19" pageId="28" pageNumber="27">4.07</td>
<td id="76954B33FFD1C6E82AFB77F7FA08FBA7" box="[1259,1444,1019,1044]" gridcol="5" gridrow="19" pageId="28" pageNumber="27">4.20</td>
</tr>
<tr id="3544224FFFD1C6E82E6A7029FA08FB8E" box="[122,1444,1061,1085]" gridrow="20" pageId="28" pageNumber="27">
<th id="76954B33FFD1C6E82E6A7029FEEAFB8E" box="[122,326,1061,1085]" gridcol="0" gridrow="20" pageId="28" pageNumber="27">Total</th>
<td id="76954B33FFD1C6E82F7E7029FE7FFB8E" box="[366,467,1061,1085]" gridcol="1" gridrow="20" pageId="28" pageNumber="27">L</td>
<td id="76954B33FFD1C6E82C117029FD15FB8E" box="[513,697,1061,1085]" gridcol="2" gridrow="20" pageId="28" pageNumber="27">26.76</td>
<td id="76954B33FFD1C6E82CE97029FC1EFB8E" box="[761,946,1061,1085]" gridcol="3" gridrow="20" pageId="28" pageNumber="27">23.62</td>
<td id="76954B33FFD1C6E82DE27029FB07FB8E" box="[1010,1195,1061,1085]" gridcol="4" gridrow="20" pageId="28" pageNumber="27">32.11</td>
<td id="76954B33FFD1C6E82AFB7029FA08FB8E" box="[1259,1444,1061,1085]" gridcol="5" gridrow="20" pageId="28" pageNumber="27">34.00</td>
</tr>
</table>
</paragraph>
<caption id="DF0B7085FFD1390B2E637058FE39FB34" pageId="28" pageNumber="27">
<paragraph id="8BCB200DFFD1390B2E637058FE39FB34" blockId="28.[115,1451,1108,1159]" pageId="28" pageNumber="27">
<emphasis id="B900FC1FFFD1390B2E637058FF11FBDA" bold="true" box="[115,189,1108,1129]" pageId="28" pageNumber="27">Table 3</emphasis>
. Measurements (mm) of four adult types of
<taxonomicName id="4C745B8EFFD1390B2C777058FCA6FBDA" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[615,778,1108,1129]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFD1390B2C777058FCA6FBDA" box="[615,778,1108,1129]" italics="true" pageId="28" pageNumber="27">Tityopsis sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFD1390B2D007058FCEAFBDA" box="[784,838,1108,1129]" pageId="28" pageNumber="27" rank="species">
<emphasis id="B900FC1FFFD1390B2D007058FC8BFBDA" bold="true" box="[784,807,1108,1129]" pageId="28" pageNumber="27">sp</emphasis>
.
<emphasis id="B900FC1FFFD1390B2D237058FCECFBDA" bold="true" box="[819,832,1108,1129]" pageId="28" pageNumber="27">n</emphasis>
.
</taxonomicNameLabel>
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
</paragraph>
</caption>
<subSubSection id="C36E7386FFD139302D20714FFDD1F85A" lastPageId="39" lastPageNumber="38" pageId="28" pageNumber="27" type="description">
<paragraph id="8BCB200DFFD1390B2D20714FFC70FAC8" blockId="28.[816,1475,1347,2043]" pageId="28" pageNumber="27">
DESCRIPTION (adult male
<typeStatus id="54CF9EAFFFD1390B2A50714FFB0DFAE8" box="[1088,1185,1347,1371]" pageId="28" pageNumber="27" type="holotype">holotype</typeStatus>
;
<figureCitation id="134F3C88FFD1390B2ABC714FFB5AFAE8" box="[1196,1270,1347,1371]" captionStart="Figure 2" captionStartId="4.[113,181,849,871]" captionTargetBox="[113,1474,234,838]" captionTargetId="figure-658@4.[113,1474,234,838]" captionTargetPageId="4" captionText="Figure 2. Western Cuba, showing the complete geographical distribution of the genus Tityopsis (white symbols) and overlaid, that of the four species of Tityopsis treated herein (red symbols, complemented with habitus of their holotypes): T. mulata sp. n. (star), T. canizaresorum sp. n. (rhombs, the southernmost is the type locality), T. pumila sp. n. (triangle) and T. sheylae sp. n. (square). Image frame size, 500 x 250 km." figureDoi="http://doi.org/10.5281/zenodo.4648297" httpUri="https://zenodo.org/record/4648297/files/figure.png" pageId="28" pageNumber="27">Figs. 2</figureCitation>
,
<figureCitation id="134F3C88FFD1390B2B11714FFAB2FAE8" box="[1281,1310,1347,1371]" captionStart="Figures 23–24" captionStartId="29.[115,192,2018,2040]" captionTargetBox="[205,1391,240,2005]" captionTargetId="figure-7@29.[113,1474,231,2009]" captionTargetPageId="29" captionText="Figures 23–24. Tityopsis sheylae sp. n., male holotype (23) and female paratopotype (24), habitus: a) dorsal; b) ventral. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648321" httpUri="https://zenodo.org/record/4648321/files/figure.png" pageId="28" pageNumber="27">23</figureCitation>
,
<figureCitation id="134F3C88FFD1390B2B39714FFAEAFAE8" box="[1321,1350,1347,1371]" captionStart="Figures 25–26" captionStartId="30.[113,190,1982,2004]" captionTargetBox="[158,1454,285,1902]" captionTargetId="figure-13@30.[113,1474,234,1972]" captionTargetPageId="30" captionText="Figures 25–26. Tityopsis sheylae sp. n., male holotype (25) and female paratopotype (26): a) pedipalp, dorsal view; b) pedipalp chela, external view; c) pedipalp chela, ventral view; d) telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4648323" httpUri="https://zenodo.org/record/4648323/files/figure.png" pageId="28" pageNumber="27">25</figureCitation>
,
<figureCitation id="134F3C88FFD1390B2B41714FFAC2FAE8" box="[1361,1390,1347,1371]" captionStart="Figures 27–28" captionStartId="31.[115,192,2011,2033]" captionTargetBox="[113,1474,240,1986]" captionTargetId="figure-9@31.[113,1476,234,1986]" captionTargetPageId="31" captionText="Figures 27–28. Tityopsis sheylae sp. n., male holotype (27) and female paratopotype (28): a) chelicerae and carapace; b) tergites." figureDoi="http://doi.org/10.5281/zenodo.4648325" httpUri="https://zenodo.org/record/4648325/files/figure.png" pageId="28" pageNumber="27">27</figureCitation>
,
<figureCitation id="134F3C88FFD1390B2B69714FFA3AFAE8" box="[1401,1430,1347,1371]" captionStart="Figures 29–30" captionStartId="32.[116,193,1436,1458]" captionTargetBox="[113,1474,234,1429]" captionTargetId="figure-14@32.[113,1474,234,1429]" captionTargetPageId="32" captionText="Figures 29–30. Tityopsis sheylae sp. n., male holotype (29) and female paratopotype (30): a) sternopectinal region; b) sternite V." figureDoi="http://doi.org/10.5281/zenodo.4648327" httpUri="https://zenodo.org/record/4648327/files/figure.png" pageId="28" pageNumber="27">29</figureCitation>
,
<figureCitation id="134F3C88FFD1390B2BB1714FFA12FAE8" box="[1441,1470,1347,1371]" captionStart="Figures 31–32" captionStartId="33.[113,190,1986,2008]" captionTargetBox="[125,1459,280,1967]" captionTargetId="figure-9@33.[113,1474,230,1979]" captionTargetPageId="33" captionText="Figures 31–32. Tityopsis sheylae sp. n., male holotype (31) and female paratopotype (32), metasoma and telson: a) dorsal, b) lateral, and c) ventral views." figureDoi="http://doi.org/10.5281/zenodo.4648329" httpUri="https://zenodo.org/record/4648329/files/figure.png" pageId="28" pageNumber="27">31</figureCitation>
,
<figureCitation id="134F3C88FFD1390B2D20716FFCF6FAC8" box="[816,858,1379,1403]" captionStart="Figure 36" captionStartId="37.[113,181,1923,1945]" captionTargetBox="[113,1474,249,1820]" captionTargetId="figure-8@37.[113,1474,249,1820]" captionTargetPageId="37" captionText="Figure 36. Tityopsis sheylae sp. n., photographed alive in its natural habitat unless otherwise noted: a) male holotype; b) small-sized male paratopotype; c) female paratopotype; d) captive female paratopotype, with mixed litter of instars 1–2; e) juvenile female paratopotype; f) just-molted, small-sized adult male paratopotype, being predated upon by the pholcid Ciboneya nuriae (the spider is not visible because while taking the photographs, it fled from the camera’s flash and hid in the crevice behind the scorpion). Photos courtesy Sheyla Yong (a–b)." figureDoi="http://doi.org/10.5281/zenodo.4648335" httpUri="https://zenodo.org/record/4648335/files/figure.png" pageId="28" pageNumber="27">36a</figureCitation>
;
<tableCitation id="C6F615B6FFD1390B2D74716FFC7CFAC8" box="[868,976,1379,1403]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="28.[115,171,1108,1129]" captionStartId-1="34.[114,170,598,619]" captionTargetBox-1="[124,1452,250,581]" captionTargetId-1="graphics-728@34.[119,1470,238,585]" captionText-0="Table 3. Measurements (mm) of four adult types of Tityopsis sheylae sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 4. Pectinal tooth count variation in the four species of the genus Tityopsis treated in this paper. Abbreviations: number of pectines (N)," httpUri-1="http://table.plazi.org/id/DF0B7085FFEF39352E62765AFE5BFD3B" pageId="28" pageNumber="27" tableUuid-1="DF0B7085FFEF39352E62765AFE5BFD3B">Tabs. 3–4</tableCitation>
).
</paragraph>
<paragraph id="8BCB200DFFD1390B2D20718FFBA0F848" blockId="28.[816,1475,1347,2043]" pageId="28" pageNumber="27">
<emphasis id="B900FC1FFFD1390B2D20718FFC01FA28" bold="true" box="[816,941,1411,1435]" pageId="28" pageNumber="27">Coloration</emphasis>
(
<figureCitation id="134F3C88FFD1390B2DD5718FFB84FA28" box="[965,1064,1411,1435]" captionStart="Figures 23–24" captionStartId="29.[115,192,2018,2040]" captionTargetBox="[205,1391,240,2005]" captionTargetId="figure-7@29.[113,1474,231,2009]" captionTargetPageId="29" captionText="Figures 23–24. Tityopsis sheylae sp. n., male holotype (23) and female paratopotype (24), habitus: a) dorsal; b) ventral. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648321" httpUri="https://zenodo.org/record/4648321/files/figure.png" pageId="28" pageNumber="27">Figs. 23</figureCitation>
,
<figureCitation id="134F3C88FFD1390B2A2C718FFBC9FA28" box="[1084,1125,1411,1435]" captionStart="Figure 36" captionStartId="37.[113,181,1923,1945]" captionTargetBox="[113,1474,249,1820]" captionTargetId="figure-8@37.[113,1474,249,1820]" captionTargetPageId="37" captionText="Figure 36. Tityopsis sheylae sp. n., photographed alive in its natural habitat unless otherwise noted: a) male holotype; b) small-sized male paratopotype; c) female paratopotype; d) captive female paratopotype, with mixed litter of instars 1–2; e) juvenile female paratopotype; f) just-molted, small-sized adult male paratopotype, being predated upon by the pholcid Ciboneya nuriae (the spider is not visible because while taking the photographs, it fled from the camera’s flash and hid in the crevice behind the scorpion). Photos courtesy Sheyla Yong (a–b)." figureDoi="http://doi.org/10.5281/zenodo.4648335" httpUri="https://zenodo.org/record/4648335/files/figure.png" pageId="28" pageNumber="27">36a</figureCitation>
) base light yellowish brown, slightly darker on metasomal segments IV–V (medium brown) and slightly paler on legs and venter. Chelicerae manus very sparsely reticulate with brown (more densely distally); fingers very faintly and irregularly infuscate. Pedipalps immaculate, with carinae and fingers slightly darker due to heavier sclerotization; fingers infuscate. Carapace largely immaculate, with faint symmetrical infuscation around all three eye groups; eyes and ocular tubercles black. Tergites immaculate, with carinae of VII slightly darker due to heavier sclerotization. Pectines immaculate pale yellowish brown, with basal portion and basal plate progressively darker due to heavier sclerotization. Sternites immaculate; V with median and lateral smooth patches bright white. Legs immaculate, with carinae slightly darker due to heavier sclerotization. Metasoma with segments IV–V slightly darker (medium brown), with carinae darker due to heavier sclerotization and without any annulated pattern; all surfaces immaculate. Telson immaculate, with distal half of aculeus dark reddish brown.
</paragraph>
<caption id="DF0B7085FFD0390A2E6373EEFA14F84B" ID-DOI="http://doi.org/10.5281/zenodo.4648321" ID-Zenodo-Dep="4648321" box="[115,1464,2018,2041]" httpUri="https://zenodo.org/record/4648321/files/figure.png" pageId="29" pageNumber="28" startId="29.[115,192,2018,2040]" targetBox="[205,1391,240,2005]" targetPageId="29">
<paragraph id="8BCB200DFFD0390A2E6373EEFA14F84B" blockId="29.[115,1464,2018,2041]" box="[115,1464,2018,2041]" pageId="29" pageNumber="28">
<emphasis id="B900FC1FFFD0390A2E6373EEFEAEF844" bold="true" box="[115,258,2018,2040]" pageId="29" pageNumber="28">Figures 23–24</emphasis>
.
<taxonomicName id="4C745B8EFFD0390A2F1E73EFFE1FF84B" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[270,435,2019,2040]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFD0390A2F1E73EFFE1FF84B" box="[270,435,2019,2040]" italics="true" pageId="29" pageNumber="28">Tityopsis sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFD0390A2FA973EFFE42F84B" box="[441,494,2019,2040]" pageId="29" pageNumber="28" rank="species">
<emphasis id="B900FC1FFFD0390A2FA973EFFE7CF84B" bold="true" box="[441,464,2019,2040]" pageId="29" pageNumber="28">sp</emphasis>
.
<emphasis id="B900FC1FFFD0390A2FCB73EFFE44F84B" bold="true" box="[475,488,2019,2040]" pageId="29" pageNumber="28">n</emphasis>
.
</taxonomicNameLabel>
, male holotype (23) and female paratopotype (24), habitus:
<emphasis id="B900FC1FFFD0390A2A2073EEFBE8F84B" bold="true" box="[1072,1092,2018,2040]" pageId="29" pageNumber="28">a)</emphasis>
dorsal;
<emphasis id="B900FC1FFFD0390A2A8273EEFB0BF84B" bold="true" box="[1170,1191,2018,2040]" pageId="29" pageNumber="28">b)</emphasis>
ventral. Scale bar = 10 mm.
</paragraph>
</caption>
<caption id="DF0B7085FFD339092E6173B2FD15F842" ID-DOI="http://doi.org/10.5281/zenodo.4648323" ID-Zenodo-Dep="4648323" httpUri="https://zenodo.org/record/4648323/files/figure.png" pageId="30" pageNumber="29" startId="30.[113,190,1982,2004]" targetBox="[158,1454,285,1902]" targetPageId="30">
<paragraph id="8BCB200DFFD339092E6173B2FD15F842" blockId="30.[113,1471,1982,2033]" pageId="30" pageNumber="29">
<emphasis id="B900FC1FFFD339092E6173B2FEACF860" bold="true" box="[113,256,1982,2004]" pageId="30" pageNumber="29">Figures 25–26</emphasis>
.
<taxonomicName id="4C745B8EFFD339092F1B73B3FE03F860" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[267,431,1982,2004]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFD339092F1B73B3FE03F860" box="[267,431,1982,2004]" italics="true" pageId="30" pageNumber="29">Tityopsis sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFD339092FA473B2FE44F867" box="[436,488,1982,2004]" pageId="30" pageNumber="29" rank="species">
<emphasis id="B900FC1FFFD339092FA473B2FE67F860" bold="true" box="[436,459,1982,2003]" pageId="30" pageNumber="29">sp</emphasis>
.
<emphasis id="B900FC1FFFD339092FC573B2FE4EF860" bold="true" box="[469,482,1982,2003]" pageId="30" pageNumber="29">n</emphasis>
.
</taxonomicNameLabel>
, male holotype (25) and female paratopotype (26):
<emphasis id="B900FC1FFFD339092DC273B2FC49F867" bold="true" box="[978,997,1982,2004]" pageId="30" pageNumber="29">a)</emphasis>
pedipalp, dorsal view;
<emphasis id="B900FC1FFFD339092AD273B2FB74F867" bold="true" box="[1218,1240,1982,2004]" pageId="30" pageNumber="29">b)</emphasis>
pedipalp chela, external view;
<emphasis id="B900FC1FFFD339092EBD73D7FF13F842" bold="true" box="[173,191,2011,2033]" pageId="30" pageNumber="29">c)</emphasis>
pedipalp chela, ventral view;
<emphasis id="B900FC1FFFD339092FF173D7FE5AF842" bold="true" box="[481,502,2011,2033]" pageId="30" pageNumber="29">d)</emphasis>
telson, lateral view.
</paragraph>
</caption>
<caption id="DF0B7085FFD239082E6373D7FAFFF842" ID-DOI="http://doi.org/10.5281/zenodo.4648325" ID-Zenodo-Dep="4648325" box="[115,1363,2011,2033]" httpUri="https://zenodo.org/record/4648325/files/figure.png" pageId="31" pageNumber="30" startId="31.[115,192,2011,2033]" targetBox="[113,1474,240,1986]" targetPageId="31">
<paragraph id="8BCB200DFFD239082E6373D7FAFFF842" blockId="31.[115,1363,2011,2033]" box="[115,1363,2011,2033]" pageId="31" pageNumber="30">
<emphasis id="B900FC1FFFD239082E6373D7FEAEF843" bold="true" box="[115,258,2011,2033]" pageId="31" pageNumber="30">Figures 27–28</emphasis>
.
<taxonomicName id="4C745B8EFFD239082F1E73D0FE1FF843" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[270,435,2011,2033]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFD239082F1E73D0FE1FF843" box="[270,435,2011,2033]" italics="true" pageId="31" pageNumber="30">Tityopsis sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFD239082FA973D7FE42F843" box="[441,494,2011,2032]" pageId="31" pageNumber="30" rank="species">
<emphasis id="B900FC1FFFD239082FA973D7FE7CF843" bold="true" box="[441,464,2011,2032]" pageId="31" pageNumber="30">sp</emphasis>
.
<emphasis id="B900FC1FFFD239082FCB73D7FE44F843" bold="true" box="[475,488,2011,2032]" pageId="31" pageNumber="30">n</emphasis>
.
</taxonomicNameLabel>
, male holotype (27) and female paratopotype (28):
<emphasis id="B900FC1FFFD239082DCF73D7FC5FF842" bold="true" box="[991,1011,2011,2033]" pageId="31" pageNumber="30">a)</emphasis>
chelicerae and carapace;
<emphasis id="B900FC1FFFD239082AFA73D7FB53F842" bold="true" box="[1258,1279,2011,2033]" pageId="31" pageNumber="30">b)</emphasis>
tergites.
</paragraph>
</caption>
<caption id="DF0B7085FFED39372E647190FAFDFA01" ID-DOI="http://doi.org/10.5281/zenodo.4648327" ID-Zenodo-Dep="4648327" box="[116,1361,1436,1459]" httpUri="https://zenodo.org/record/4648327/files/figure.png" pageId="32" pageNumber="31" startId="32.[116,193,1436,1458]" targetBox="[113,1474,234,1429]" targetPageId="32">
<paragraph id="8BCB200DFFED39372E647190FAFDFA01" blockId="32.[116,1361,1436,1459]" box="[116,1361,1436,1459]" pageId="32" pageNumber="31">
<emphasis id="B900FC1FFFED39372E647190FEAFFA02" bold="true" box="[116,259,1436,1458]" pageId="32" pageNumber="31">Figures 29–30</emphasis>
.
<taxonomicName id="4C745B8EFFED39372F1F7191FE18FA01" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[271,436,1437,1458]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFED39372F1F7191FE18FA01" box="[271,436,1437,1458]" italics="true" pageId="32" pageNumber="31">Tityopsis sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFED39372FAA7191FE43FA01" box="[442,495,1437,1458]" pageId="32" pageNumber="31" rank="species">
<emphasis id="B900FC1FFFED39372FAA7191FE7DFA01" bold="true" box="[442,465,1437,1458]" pageId="32" pageNumber="31">sp</emphasis>
.
<emphasis id="B900FC1FFFED39372FCD7191FE46FA01" bold="true" box="[477,490,1437,1458]" pageId="32" pageNumber="31">n</emphasis>
.
</taxonomicNameLabel>
, male holotype (29) and female paratopotype (30):
<emphasis id="B900FC1FFFED39372DF07190FC58FA01" bold="true" box="[992,1012,1436,1458]" pageId="32" pageNumber="31">a)</emphasis>
sternopectinal region;
<emphasis id="B900FC1FFFED39372AC17190FB4AFA01" bold="true" box="[1233,1254,1436,1458]" pageId="32" pageNumber="31">b)</emphasis>
sternite V.
</paragraph>
</caption>
<paragraph id="8BCB200DFFED39372E6171C6FD8CF912" blockId="32.[113,772,1482,2050]" pageId="32" pageNumber="31">
<emphasis id="B900FC1FFFED39372E6171C6FF46FA51" bold="true" box="[113,234,1482,1506]" pageId="32" pageNumber="31">Chelicerae</emphasis>
(
<figureCitation id="134F3C88FFED39372EED71C6FEF5FA51" box="[253,345,1482,1506]" captionStart="Figures 27–28" captionStartId="31.[115,192,2011,2033]" captionTargetBox="[113,1474,240,1986]" captionTargetId="figure-9@31.[113,1476,234,1986]" captionTargetPageId="31" captionText="Figures 27–28. Tityopsis sheylae sp. n., male holotype (27) and female paratopotype (28): a) chelicerae and carapace; b) tergites." figureDoi="http://doi.org/10.5281/zenodo.4648325" httpUri="https://zenodo.org/record/4648325/files/figure.png" pageId="32" pageNumber="31">Fig. 27a</figureCitation>
). With dentition typical for the genus, teeth standard-sized and sharp. Tegument glossy but with minute granulation scattered, dorsodistal portion of manus with coarse, glossy granules irregularly arranged transversally around dorsodistal depression. Setation very dense ventrally, but essentially lacking dorsally, except for five rigid macrosetae around dorsodistal depression of manus.
</paragraph>
<paragraph id="8BCB200DFFED39372E6172A6FA83F951" blockId="32.[113,772,1482,2050]" lastBlockId="32.[816,1474,1482,2050]" pageId="32" pageNumber="31">
<emphasis id="B900FC1FFFED39372E6172A6FF4CF971" bold="true" box="[113,224,1706,1730]" pageId="32" pageNumber="31">Pedipalps</emphasis>
(
<figureCitation id="134F3C88FFED39372EE172A6FECAF971" box="[241,358,1706,1730]" captionStart="Figures 25–26" captionStartId="30.[113,190,1982,2004]" captionTargetBox="[158,1454,285,1902]" captionTargetId="figure-13@30.[113,1474,234,1972]" captionTargetPageId="30" captionText="Figures 25–26. Tityopsis sheylae sp. n., male holotype (25) and female paratopotype (26): a) pedipalp, dorsal view; b) pedipalp chela, external view; c) pedipalp chela, ventral view; d) telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4648323" httpUri="https://zenodo.org/record/4648323/files/figure.png" pageId="32" pageNumber="31">Fig. 25a–c</figureCitation>
). Remarkably enlarged for the genus, i.e., all segments longer and chelae also heavier than standard. Femur essentially straight (very slightly bent inwards distally), with all carinae very strong, coarsely denticulate to serrate; intercarinal tegument matt, very densely and irregularly granulose; internal (
<emphasis id="B900FC1FFFED39372FC07347FE7BF8D1" box="[464,471,1867,1890]" italics="true" pageId="32" pageNumber="31">i</emphasis>
) trichobothria surrounding a large, irregularly triangular spur. Patella straight, with all carinae very strong, coarsely granulose to serrato-crenulate; intercarinal tegument matt, very finely and densely granulose, internally with many conical denticles. Chela with manus incrassate oval (1.56 times longer than wide), much wider than patella (ratio 1.26) and with the basal half slightly widest, all carinae strong, coarsely serrate to serrato-crenulate, intercarinal tegument matt, very finely and densely granulose on all surfaces; fingers very long for the genus (movable finger 1.34 times longer than underhand) and evenly curved, fixed finger with 12/12 principal rows of denticles, movable finger with 12/12 plus an apical subrow of four denticles and a large internal accessory denticle (large terminal denticle not included), basal lobe/notch combination absent.
</paragraph>
<paragraph id="8BCB200DFFED39352D2072E6FCA8FC06" blockId="32.[816,1474,1482,2050]" lastBlockId="34.[113,772,669,2037]" lastPageId="34" lastPageNumber="33" pageId="32" pageNumber="31">
<emphasis id="B900FC1FFFED39372D2072E6FC32F8B1" bold="true" box="[816,926,1770,1794]" pageId="32" pageNumber="31">Carapace</emphasis>
(
<figureCitation id="134F3C88FFED39372DA072E6FBA0F8B1" box="[944,1036,1770,1794]" captionStart="Figures 27–28" captionStartId="31.[115,192,2011,2033]" captionTargetBox="[113,1474,240,1986]" captionTargetId="figure-9@31.[113,1476,234,1986]" captionTargetPageId="31" captionText="Figures 27–28. Tityopsis sheylae sp. n., male holotype (27) and female paratopotype (28): a) chelicerae and carapace; b) tergites." figureDoi="http://doi.org/10.5281/zenodo.4648325" httpUri="https://zenodo.org/record/4648325/files/figure.png" pageId="32" pageNumber="31">Fig. 27a</figureCitation>
). Wider than long (length/width ratio = 0.93). Anterior margin widely V-shaped, with two pairs of dark macrosetae. Carinae: anterior medians anteriorly divergent and formed by coarse, isolated granules and irregularly fused to the superciliaries, which are composed of smaller and much closer granules; posterior medians straight, parallel and formed by medium-sized granules; other carinae absent or indistinct from general granulation. Furrows: anterior marginal, anterior median, median ocular, lateral oculars, central transverse, central median, posterior median and posterior marginal all fused, wide and moderately deep; posterior laterals long, oblique, wide and moderately deep, almost fused to posterior marginal; posterior transverse narrow and moderately deep; other furrows absent or indistinct. Tegument matt, very densely and irregularly but finely granulose, with many medium-sized, rough granules scattered. Median eyes relatively large and separated by about one ocular diameter; lateral eyes noticeably smaller and moderately exposed beyond anterolateral margin.
</paragraph>
<caption id="DF0B7085FFEC39362E6173CEFEBEF846" ID-DOI="http://doi.org/10.5281/zenodo.4648329" ID-Zenodo-Dep="4648329" httpUri="https://zenodo.org/record/4648329/files/figure.png" pageId="33" pageNumber="32" startId="33.[113,190,1986,2008]" targetBox="[125,1459,280,1967]" targetPageId="33">
<paragraph id="8BCB200DFFEC39362E6173CEFEBEF846" blockId="33.[113,1478,1986,2037]" pageId="33" pageNumber="32">
<emphasis id="B900FC1FFFEC39362E6173CEFEAFF864" bold="true" box="[113,259,1986,2008]" pageId="33" pageNumber="32">Figures 31–32</emphasis>
.
<taxonomicName id="4C745B8EFFEC39362F0073CFFE1BF864" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[272,439,1986,2008]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEC39362F0073CFFE1BF864" box="[272,439,1986,2008]" italics="true" pageId="33" pageNumber="32">Tityopsis sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEC39362FAF73CFFE5AF86B" box="[447,502,1987,2008]" pageId="33" pageNumber="32" rank="species">
<emphasis id="B900FC1FFFEC39362FAF73CFFE7AF86B" bold="true" box="[447,470,1987,2008]" pageId="33" pageNumber="32">sp</emphasis>
.
<emphasis id="B900FC1FFFEC39362FF373CFFE5CF86B" bold="true" box="[483,496,1987,2008]" pageId="33" pageNumber="32">n</emphasis>
.
</taxonomicNameLabel>
, male holotype (31) and female paratopotype (32), metasoma and telson:
<emphasis id="B900FC1FFFEC39362AC173CEFB49F86B" bold="true" box="[1233,1253,1986,2008]" pageId="33" pageNumber="32">a)</emphasis>
dorsal,
<emphasis id="B900FC1FFFEC39362B2673CEFAE7F86B" bold="true" box="[1334,1355,1986,2008]" pageId="33" pageNumber="32">b)</emphasis>
lateral, and
<emphasis id="B900FC1FFFEC39362E6173D3FF28F846" bold="true" box="[113,132,2015,2037]" pageId="33" pageNumber="32">c)</emphasis>
ventral views.
</paragraph>
</caption>
<paragraph id="8BCB200DFFEF39352E6C752BFA00FDF0" pageId="34" pageNumber="33">
<table id="F974D2ADFFEFC6E82E6C74F6FA00FDF6" box="[124,1452,250,581]" gridcols="11" gridrows="5" pageId="34" pageNumber="33">
<tr id="3544224FFFEFC6E82E6C74F6FA00FEF2" box="[124,1452,250,321]" gridrow="0" pageId="34" pageNumber="33">
<th id="76954B33FFEFC6E82E6C74F6FE6FFEF2" box="[124,451,250,321]" gridcol="0" gridrow="0" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352E6C752BFF67FEF2" bold="true" box="[124,203,295,321]" pageId="34" pageNumber="33">Species</emphasis>
</th>
<th id="76954B33FFEFC6E82C0574F6FD97FEF2" box="[533,571,250,321]" gridcol="1" gridrow="0" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352C05752BFD97FEF2" bold="true" box="[533,571,295,321]" pageId="34" pageNumber="33">Sex</emphasis>
</th>
<th id="76954B33FFEFC6E82C6B74F6FD39FEF2" box="[635,661,250,321]" gridcol="2" gridrow="0" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352C90752BFD3FFEF2" bold="true" box="[640,659,295,321]" pageId="34" pageNumber="33">N</emphasis>
</th>
<th id="76954B33FFEFC6E82CCD74F6FD5BFEF2" box="[733,759,250,321]" gridcol="3" gridrow="0" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352CCD752BFD5BFEF2" bold="true" box="[733,759,295,321]" pageId="34" pageNumber="33">11</emphasis>
</th>
<th id="76954B33FFEFC6E82D2C74F6FBD5FEF2" box="[828,1145,250,321]" colspan="4" colspanRight="3" gridcol="4" gridrow="0" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352D7E74F6FBD5FEF2" bold="true" pageId="34" pageNumber="33">Pectinal tooth count 12 13 14 15</emphasis>
</th>
<th id="76954B33FFEFC6E82AAF74F6FB75FEF2" box="[1215,1241,250,321]" gridcol="8" gridrow="0" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352AAF752BFB75FEF2" bold="true" box="[1215,1241,295,321]" pageId="34" pageNumber="33">16</emphasis>
</th>
<th id="76954B33FFEFC6E82B1E74F6FAE4FEF2" box="[1294,1352,250,321]" gridcol="9" gridrow="0" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352B04752BFAE9FEF2" bold="true" box="[1300,1349,295,321]" pageId="34" pageNumber="33">Ave.</emphasis>
</th>
<th id="76954B33FFEFC6E82B7B74F6FA00FEF2" box="[1387,1452,250,321]" gridcol="10" gridrow="0" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352B6D752BFA32FEF2" bold="true" box="[1405,1438,295,321]" pageId="34" pageNumber="33">SD</emphasis>
</th>
</tr>
<tr id="3544224FFFEFC6E82E6C7543FA00FE2A" box="[124,1452,335,409]" gridrow="1" pageId="34" pageNumber="33" rowspan-3="1">
<th id="76954B33FFEFC6E82E6C7543FE6FFE2A" box="[124,451,335,409]" gridcol="0" gridrow="1" pageId="34" pageNumber="33">
<taxonomicName id="4C745B8EFFEF39352E6C755CFE84FEDA" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[124,296,335,361]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="mulata" status="sp. nov.">
<emphasis id="B900FC1FFFEF39352E6C755CFE84FEDA" box="[124,296,335,361]" italics="true" pageId="34" pageNumber="33">Tityopsis mulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEF39352F23755CFEC0FED9" box="[307,364,336,362]" pageId="34" pageNumber="33" rank="species">
<emphasis id="B900FC1FFFEF39352F23755CFEE7FED9" bold="true" box="[307,331,336,362]" pageId="34" pageNumber="33">sp</emphasis>
.
<emphasis id="B900FC1FFFEF39352F48755CFECAFED9" bold="true" box="[344,358,336,362]" pageId="34" pageNumber="33">n</emphasis>
.
</taxonomicNameLabel>
</th>
<td id="76954B33FFEFC6E82C057543FD97FE2A" box="[533,571,335,409]" gridcol="1" gridrow="1" pageId="34" pageNumber="33">♂ ♀</td>
<td id="76954B33FFEFC6E82C6B7543FD39FE2A" box="[635,661,335,409]" gridcol="2" gridrow="1" pageId="34" pageNumber="33">8 12</td>
<td id="76954B33FFEFC6E82D2C7543FCFAFE2A" box="[828,854,335,409]" gridcol="4" gridrow="1" pageId="34" pageNumber="33">1</td>
<td id="76954B33FFEFC6E82D8D7543FC1BFE2A" box="[925,951,335,409]" gridcol="5" gridrow="1" pageId="34" pageNumber="33">3</td>
<td id="76954B33FFEFC6E82DED7543FBBBFE2A" box="[1021,1047,335,409]" gridcol="6" gridrow="1" pageId="34" pageNumber="33">3 5</td>
<td id="76954B33FFEFC6E82A747543FBD5FE2A" box="[1124,1145,335,409]" gridcol="7" gridrow="1" pageId="34" pageNumber="33">4 3</td>
<td id="76954B33FFEFC6E82AAF7543FB75FE2A" box="[1215,1241,335,409]" gridcol="8" gridrow="1" pageId="34" pageNumber="33">1</td>
<td id="76954B33FFEFC6E82B1E7543FAE4FE2A" box="[1294,1352,335,409]" gridcol="9" gridrow="1" pageId="34" pageNumber="33">14.75 13.83</td>
<td id="76954B33FFEFC6E82B7B7543FA00FE2A" box="[1387,1452,335,409]" gridcol="10" gridrow="1" pageId="34" pageNumber="33">± 0.66 ± 0.90</td>
</tr>
<tr id="3544224FFFEFC6E82E6C75ABFA00FE77" box="[124,1452,423,452]" gridrow="2" pageId="34" pageNumber="33" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
<th id="76954B33FFEFC6E82E6C75ABFE6FFE77" box="[124,451,423,452]" gridcol="0" gridrow="2" pageId="34" pageNumber="33">
<taxonomicName id="4C745B8EFFEF39352E6C75A4FE84FE72" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[124,296,423,449]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="pumila" status="sp. nov.">
<emphasis id="B900FC1FFFEF39352E6C75A4FE84FE72" box="[124,296,423,449]" italics="true" pageId="34" pageNumber="33">Tityopsis pumila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEF39352F2975A4FEDEFE71" box="[313,370,424,450]" pageId="34" pageNumber="33" rank="species">
<emphasis id="B900FC1FFFEF39352F2975A4FEFDFE71" bold="true" box="[313,337,424,450]" pageId="34" pageNumber="33">sp</emphasis>
.
<emphasis id="B900FC1FFFEF39352F4E75A4FEC0FE71" bold="true" box="[350,364,424,450]" pageId="34" pageNumber="33">n</emphasis>
.
</taxonomicNameLabel>
</th>
<td id="76954B33FFEFC6E82C0575ABFD97FE77" box="[533,571,423,452]" gridcol="1" gridrow="2" pageId="34" pageNumber="33">♂</td>
<td id="76954B33FFEFC6E82C6B75ABFD39FE77" box="[635,661,423,452]" gridcol="2" gridrow="2" pageId="34" pageNumber="33">2</td>
<td id="76954B33FFEFC6E82DED75ABFBBBFE77" box="[1021,1047,423,452]" gridcol="6" gridrow="2" pageId="34" pageNumber="33">2</td>
<td id="76954B33FFEFC6E82B1E75ABFAE4FE77" box="[1294,1352,423,452]" gridcol="9" gridrow="2" pageId="34" pageNumber="33">14.00</td>
<td id="76954B33FFEFC6E82B7B75ABFA00FE77" box="[1387,1452,423,452]" gridcol="10" gridrow="2" pageId="34" pageNumber="33">± 0.00</td>
</tr>
<tr id="3544224FFFEFC6E82E6C75DEFA00FE5C" box="[124,1452,466,495]" gridrow="3" pageId="34" pageNumber="33" rowspan-7="1" rowspan-8="1">
<th id="76954B33FFEFC6E82E6C75DEFE6FFE5C" box="[124,451,466,495]" gridcol="0" gridrow="3" pageId="34" pageNumber="33">
<taxonomicName id="4C745B8EFFEF39352E6C75DFFED5FE5F" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[124,377,467,492]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="canizaresorum" status="sp. nov.">
<emphasis id="B900FC1FFFEF39352E6C75DFFED5FE5F" box="[124,377,467,492]" italics="true" pageId="34" pageNumber="33">Tityopsis canizaresorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEF39352F9A75DFFE6FFE5E" box="[394,451,467,493]" pageId="34" pageNumber="33" rank="species">
<emphasis id="B900FC1FFFEF39352F9A75DFFE0EFE5E" bold="true" box="[394,418,467,493]" pageId="34" pageNumber="33">sp</emphasis>
.
<emphasis id="B900FC1FFFEF39352FBF75DFFE11FE5E" bold="true" box="[431,445,467,493]" pageId="34" pageNumber="33">n</emphasis>
.
</taxonomicNameLabel>
</th>
<td id="76954B33FFEFC6E82C0575DEFD97FE5C" box="[533,571,466,495]" gridcol="1" gridrow="3" pageId="34" pageNumber="33">♀</td>
<td id="76954B33FFEFC6E82C6B75DEFD39FE5C" box="[635,661,466,495]" gridcol="2" gridrow="3" pageId="34" pageNumber="33">76</td>
<td id="76954B33FFEFC6E82CCD75DEFD5BFE5C" box="[733,759,466,495]" gridcol="3" gridrow="3" pageId="34" pageNumber="33">2</td>
<td id="76954B33FFEFC6E82D2C75DEFCFAFE5C" box="[828,854,466,495]" gridcol="4" gridrow="3" pageId="34" pageNumber="33">33</td>
<td id="76954B33FFEFC6E82D8D75DEFC1BFE5C" box="[925,951,466,495]" gridcol="5" gridrow="3" pageId="34" pageNumber="33">39</td>
<td id="76954B33FFEFC6E82DED75DEFBBBFE5C" box="[1021,1047,466,495]" gridcol="6" gridrow="3" pageId="34" pageNumber="33">2</td>
<td id="76954B33FFEFC6E82B1E75DEFAE4FE5C" box="[1294,1352,466,495]" gridcol="9" gridrow="3" pageId="34" pageNumber="33">12.54</td>
<td id="76954B33FFEFC6E82B7B75DEFA00FE5C" box="[1387,1452,466,495]" gridcol="10" gridrow="3" pageId="34" pageNumber="33">± 0.59</td>
</tr>
<tr id="3544224FFFEFC6E82E6C75F7FA00FDF6" box="[124,1452,507,581]" gridrow="4" pageId="34" pageNumber="33" rowspan-3="1" rowspan-4="1">
<th id="76954B33FFEFC6E82E6C75F7FE6FFDF6" box="[124,451,507,581]" gridcol="0" gridrow="4" pageId="34" pageNumber="33">
<taxonomicName id="4C745B8EFFEF39352E6C75F0FE81FDA6" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[124,301,507,533]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEF39352E6C75F0FE81FDA6" box="[124,301,507,533]" italics="true" pageId="34" pageNumber="33">Tityopsis sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEF39352F2875F0FEDDFDA5" box="[312,369,508,534]" pageId="34" pageNumber="33" rank="species">
<emphasis id="B900FC1FFFEF39352F2875F0FEFCFDA5" bold="true" box="[312,336,508,534]" pageId="34" pageNumber="33">sp</emphasis>
.
<emphasis id="B900FC1FFFEF39352F4D75F0FEC7FDA5" bold="true" box="[349,363,508,534]" pageId="34" pageNumber="33">n</emphasis>
.
</taxonomicNameLabel>
</th>
<td id="76954B33FFEFC6E82C0575F7FD97FDF6" box="[533,571,507,581]" gridcol="1" gridrow="4" pageId="34" pageNumber="33">♂ ♀</td>
<td id="76954B33FFEFC6E82C6B75F7FD39FDF6" box="[635,661,507,581]" gridcol="2" gridrow="4" pageId="34" pageNumber="33">30 52</td>
<td id="76954B33FFEFC6E82D8D75F7FC1BFDF6" box="[925,951,507,581]" gridcol="5" gridrow="4" pageId="34" pageNumber="33">2 27</td>
<td id="76954B33FFEFC6E82DED75F7FBBBFDF6" box="[1021,1047,507,581]" gridcol="6" gridrow="4" pageId="34" pageNumber="33">21 24</td>
<td id="76954B33FFEFC6E82A7475F7FBD5FDF6" box="[1124,1145,507,581]" gridcol="7" gridrow="4" pageId="34" pageNumber="33">5 1</td>
<td id="76954B33FFEFC6E82AAF75F7FB75FDF6" box="[1215,1241,507,581]" gridcol="8" gridrow="4" pageId="34" pageNumber="33">2</td>
<td id="76954B33FFEFC6E82B1E75F7FAE4FDF6" box="[1294,1352,507,581]" gridcol="9" gridrow="4" pageId="34" pageNumber="33">14.23 13.50</td>
<td id="76954B33FFEFC6E82B7B75F7FA00FDF6" box="[1387,1452,507,581]" gridcol="10" gridrow="4" pageId="34" pageNumber="33">± 0.67 ± 0.54</td>
</tr>
</table>
</paragraph>
<caption id="DF0B7085FFEF39352E62765AFE5BFD3B" ID-Table-UUID="DF0B7085FFEF39352E62765AFE5BFD3B" httpUri="http://table.plazi.org/id/DF0B7085FFEF39352E62765AFE5BFD3B" pageId="34" pageNumber="33" startId="34.[114,170,598,619]" targetBox="[124,1452,250,581]" targetIsTable="true" targetPageId="34">
<paragraph id="8BCB200DFFEF39352E62765AFE5BFD3B" blockId="34.[114,1463,597,648]" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352E62765AFF10FDD9" bold="true" box="[114,188,597,619]" pageId="34" pageNumber="33">Table 4</emphasis>
. Pectinal tooth count variation in the four species of the genus
<taxonomicName id="4C745B8EFFEF39352D0A765AFCC3FDD8" authorityName=": Armas &amp; Martin Frias" authorityYear="1998" box="[794,879,598,619]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B900FC1FFFEF39352D0A765AFCC3FDD8" box="[794,879,598,619]" italics="true" pageId="34" pageNumber="33">Tityopsis</emphasis>
</taxonomicName>
treated in this paper. Abbreviations: number of pectines (N), average (Ave.), standard deviation (SD).
</paragraph>
</caption>
<paragraph id="8BCB200DFFEF39352E6177B1FD28FBA6" blockId="34.[113,772,669,2037]" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352E6177B1FF78FC66" bold="true" box="[113,212,957,981]" pageId="34" pageNumber="33">Sternum</emphasis>
(
<figureCitation id="134F3C88FFEF39352EF677B1FEEAFC66" box="[230,326,957,981]" captionStart="Figures 29–30" captionStartId="32.[116,193,1436,1458]" captionTargetBox="[113,1474,234,1429]" captionTargetId="figure-14@32.[113,1474,234,1429]" captionTargetPageId="32" captionText="Figures 29–30. Tityopsis sheylae sp. n., male holotype (29) and female paratopotype (30): a) sternopectinal region; b) sternite V." figureDoi="http://doi.org/10.5281/zenodo.4648327" httpUri="https://zenodo.org/record/4648327/files/figure.png" pageId="34" pageNumber="33">Fig. 29a</figureCitation>
). Standard for the genus, although more pentagonal in shape, with three pairs of dark macrosetae. Tegument very densely and irregularly granulose.
</paragraph>
<paragraph id="8BCB200DFFEF39352E617011FD4FFB26" blockId="34.[113,772,669,2037]" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352E617011FEE5FB86" bold="true" box="[113,329,1053,1077]" pageId="34" pageNumber="33">Genital operculum</emphasis>
(
<figureCitation id="134F3C88FFEF39352F4A7011FE15FB86" box="[346,441,1053,1077]" captionStart="Figures 29–30" captionStartId="32.[116,193,1436,1458]" captionTargetBox="[113,1474,234,1429]" captionTargetId="figure-14@32.[113,1474,234,1429]" captionTargetPageId="32" captionText="Figures 29–30. Tityopsis sheylae sp. n., male holotype (29) and female paratopotype (30): a) sternopectinal region; b) sternite V." figureDoi="http://doi.org/10.5281/zenodo.4648327" httpUri="https://zenodo.org/record/4648327/files/figure.png" pageId="34" pageNumber="33">Fig. 29a</figureCitation>
). Standard for the genus, with three pairs of dark macrosetae. Valves roundly subtriangular and not separated medially; tegument minutely and densely granulose. Genital papillae large, thick and not protruding.
</paragraph>
<paragraph id="8BCB200DFFEF39352E617091FD20FA06" blockId="34.[113,772,669,2037]" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352E617091FF62FB06" bold="true" box="[113,206,1181,1205]" pageId="34" pageNumber="33">Pectines</emphasis>
(
<figureCitation id="134F3C88FFEF39352EF07091FE92FB06" box="[224,318,1181,1205]" captionStart="Figures 29–30" captionStartId="32.[116,193,1436,1458]" captionTargetBox="[113,1474,234,1429]" captionTargetId="figure-14@32.[113,1474,234,1429]" captionTargetPageId="32" captionText="Figures 29–30. Tityopsis sheylae sp. n., male holotype (29) and female paratopotype (30): a) sternopectinal region; b) sternite V." figureDoi="http://doi.org/10.5281/zenodo.4648327" httpUri="https://zenodo.org/record/4648327/files/figure.png" pageId="34" pageNumber="33">Fig. 29a</figureCitation>
). Moderately large for the genus (but not reaching coxa-trochanter joint of leg IV). Tooth count 14/15, teeth straight and slightly swollen. Fulcra large and bulky, paraboloid to round. Basal middle lamella slightly enlarged and teardrop-shaped. Pre-pectinal plate well visible, heavily sclerotized, very wide and short; tegument coriaceous to smooth. Basal plate normally sclerotized, about as long as wide; anterior margin with a very wide, V-shaped anteromedian notch, posterior margin convex; tegument smooth.
</paragraph>
<paragraph id="8BCB200DFFEF39352E6171B1FDE5F9E6" blockId="34.[113,772,669,2037]" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352E6171B1FF0AFA66" bold="true" box="[113,166,1469,1493]" pageId="34" pageNumber="33">Legs</emphasis>
. Slender, with all carinae strong, finely denticulate to granulose. Intercarinal tegument minutely and densely granulose, with a row of slightly coarser granules defining an additional carina (externomedian) on lateral surface of femur of all legs. Claws short and strongly curved.
</paragraph>
<paragraph id="8BCB200DFFEF39352E617251FB36FC26" blockId="34.[113,772,669,2037]" lastBlockId="34.[816,1475,669,1909]" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352E617251FF45F9C6" bold="true" box="[113,233,1629,1653]" pageId="34" pageNumber="33">Mesosoma</emphasis>
(
<figureCitation id="134F3C88FFEF39352EEB7251FEC9F9C6" box="[251,357,1629,1653]" captionStart="Figures 27–28" captionStartId="31.[115,192,2011,2033]" captionTargetBox="[113,1474,240,1986]" captionTargetId="figure-9@31.[113,1476,234,1986]" captionTargetPageId="31" captionText="Figures 27–28. Tityopsis sheylae sp. n., male holotype (27) and female paratopotype (28): a) chelicerae and carapace; b) tergites." figureDoi="http://doi.org/10.5281/zenodo.4648325" httpUri="https://zenodo.org/record/4648325/files/figure.png" pageId="34" pageNumber="33">Figs. 27b</figureCitation>
,
<figureCitation id="134F3C88FFEF39352F647251FE3EF9C6" box="[372,402,1629,1653]" captionStart="Figures 29–30" captionStartId="32.[116,193,1436,1458]" captionTargetBox="[113,1474,234,1429]" captionTargetId="figure-14@32.[113,1474,234,1429]" captionTargetPageId="32" captionText="Figures 29–30. Tityopsis sheylae sp. n., male holotype (29) and female paratopotype (30): a) sternopectinal region; b) sternite V." figureDoi="http://doi.org/10.5281/zenodo.4648327" httpUri="https://zenodo.org/record/4648327/files/figure.png" pageId="34" pageNumber="33">29</figureCitation>
). Tergites matt, very densely and irregularly but finely granulose, with many medium-sized to coarse, rough granules scattered; carinae: median longitudinal strong, serrato-crenulate to serrate and progressively longer from I–VII, submedians absent on I–VI, complete, straight and strongly serrato-denticulate on VII, laterals absent on I–VI, complete, straight and strongly serrato-denticulate on VII. Sternites matt, very densely and irregularly but finely granulose, with some dark macrosetae scattered all over; lateral margins of all sternites subgranulose to granulose; posterior margin of all sternites smooth to vestigially granulose, on III–IV vestigially bilobed, on V bilobed (due to protuberance of lateral smooth patches), on VI essentially straight, on VII widely concave; carinae: III–V acarinate, VI-VII with paired laterals and submedians which are long, straight and finely crenulate; spiracles small, narrowly oval to short slit-like; sternite V with median smooth patch very large, widely cordiform (about as long as wide), bulky and glossy, not protruding from posterior margin; lateral smooth patches moderately large, oblique, oval, bulky and glossy, moderately protruding from posterior margin.
</paragraph>
<paragraph id="8BCB200DFFEF39352D207791FB5FF9C6" blockId="34.[816,1475,669,1909]" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352D207791FC0AFC06" bold="true" box="[816,934,925,949]" pageId="34" pageNumber="33">Metasoma</emphasis>
(
<figureCitation id="134F3C88FFEF39352DAA7791FBA2FC06" box="[954,1038,925,949]" captionStart="Figures 31–32" captionStartId="33.[113,190,1986,2008]" captionTargetBox="[125,1459,280,1967]" captionTargetId="figure-9@33.[113,1474,230,1979]" captionTargetPageId="33" captionText="Figures 31–32. Tityopsis sheylae sp. n., male holotype (31) and female paratopotype (32), metasoma and telson: a) dorsal, b) lateral, and c) ventral views." figureDoi="http://doi.org/10.5281/zenodo.4648329" httpUri="https://zenodo.org/record/4648329/files/figure.png" pageId="34" pageNumber="33">Fig. 31</figureCitation>
). Slightly longer than standard for the genus (4.23 times longer than carapace) and slightly narrower distally. Segments I–II with ten complete to almost complete carinae, III–IV with eight, V with seven: dorsal laterals strong, coarsely serrate to serrato-denticulate on I–III, coarsely crenulate-denticulate on IV, absent on V but subtly indicated by a few granules; lateral supramedians strong, coarsely serrate to serrato-crenulate on I–V; lateral inframedians strong, coarsely serrato-crenulate on I–II (obsolete on basal one-sixth of the latter), absent on III–V (but indicated in each segment by an incomplete row of granules that never form a true, raised carina); ventral laterals strong, coarsely crenulate to serrato-crenulate on I–V; ventral submedians strong, coarsely serrate to serrato-crenulate on I–IV, indicated by an incomplete but raised row of coarse granules on less than basal half of V; ventral median absent on I–IV, strong, coarsely serrato-crenulate on V. Intercarinal tegument matt, very finely and densely granulose, with many medium-sized granules scattered on dorsal surface of all segments and many small, comical granules on lateral surfaces of V; dorsal furrow absent or indistinct on segments I–III, complete, narrow and moderately deep on IV–V; setation sparse, with two pairs of dark macrosetae on essentially all carinae.
</paragraph>
<paragraph id="8BCB200DFFEF39352D207271FAE5F8C6" blockId="34.[816,1475,669,1909]" pageId="34" pageNumber="33">
<emphasis id="B900FC1FFFEF39352D207271FCD5F926" bold="true" box="[816,889,1661,1685]" pageId="34" pageNumber="33">Telson</emphasis>
(
<figureCitation id="134F3C88FFEF39352D9A7271FC4BF926" box="[906,999,1661,1685]" captionStart="Figures 25–26" captionStartId="30.[113,190,1982,2004]" captionTargetBox="[158,1454,285,1902]" captionTargetId="figure-13@30.[113,1474,234,1972]" captionTargetPageId="30" captionText="Figures 25–26. Tityopsis sheylae sp. n., male holotype (25) and female paratopotype (26): a) pedipalp, dorsal view; b) pedipalp chela, external view; c) pedipalp chela, ventral view; d) telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4648323" httpUri="https://zenodo.org/record/4648323/files/figure.png" pageId="34" pageNumber="33">Fig. 25d</figureCitation>
). Vesicle less globose than standard for the genus (1.48 times longer than wide, 1.02 times wider than deep) and with several dark and whitish setae scattered; tegument coriaceous and matt, smooth even ventrally; ventral median carina very weak but coarsely subgranulose, continued into the subaculear tubercle that is small, broadly conical and unarmed (i.e., without any granules). Aculeus standard-sized, sharp, shorter than vesicle and moderately curved.
</paragraph>
<paragraph id="8BCB200DFFEF39332D207392FCA8F96F" blockId="34.[816,1474,1949,2037]" lastBlockId="36.[113,772,1347,1756]" lastPageId="36" lastPageNumber="35" pageId="34" pageNumber="33">
FEMALE (adult paratopotypes;
<figureCitation id="134F3C88FFEF39352A617391FB66F806" box="[1137,1226,1949,1973]" captionStart="Figures 23–24" captionStartId="29.[115,192,2018,2040]" captionTargetBox="[205,1391,240,2005]" captionTargetId="figure-7@29.[113,1474,231,2009]" captionTargetPageId="29" captionText="Figures 23–24. Tityopsis sheylae sp. n., male holotype (23) and female paratopotype (24), habitus: a) dorsal; b) ventral. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648321" httpUri="https://zenodo.org/record/4648321/files/figure.png" pageId="34" pageNumber="33">Figs. 24</figureCitation>
,
<figureCitation id="134F3C88FFEF39352AC67391FB5FF806" box="[1238,1267,1949,1973]" captionStart="Figures 25–26" captionStartId="30.[113,190,1982,2004]" captionTargetBox="[158,1454,285,1902]" captionTargetId="figure-13@30.[113,1474,234,1972]" captionTargetPageId="30" captionText="Figures 25–26. Tityopsis sheylae sp. n., male holotype (25) and female paratopotype (26): a) pedipalp, dorsal view; b) pedipalp chela, external view; c) pedipalp chela, ventral view; d) telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4648323" httpUri="https://zenodo.org/record/4648323/files/figure.png" pageId="34" pageNumber="33">26</figureCitation>
,
<figureCitation id="134F3C88FFEF39352AEF7391FAB7F806" box="[1279,1307,1949,1973]" captionStart="Figures 27–28" captionStartId="31.[115,192,2011,2033]" captionTargetBox="[113,1474,240,1986]" captionTargetId="figure-9@31.[113,1476,234,1986]" captionTargetPageId="31" captionText="Figures 27–28. Tityopsis sheylae sp. n., male holotype (27) and female paratopotype (28): a) chelicerae and carapace; b) tergites." figureDoi="http://doi.org/10.5281/zenodo.4648325" httpUri="https://zenodo.org/record/4648325/files/figure.png" pageId="34" pageNumber="33">28</figureCitation>
,
<figureCitation id="134F3C88FFEF39352B387391FAE9F806" box="[1320,1349,1949,1973]" captionStart="Figures 29–30" captionStartId="32.[116,193,1436,1458]" captionTargetBox="[113,1474,234,1429]" captionTargetId="figure-14@32.[113,1474,234,1429]" captionTargetPageId="32" captionText="Figures 29–30. Tityopsis sheylae sp. n., male holotype (29) and female paratopotype (30): a) sternopectinal region; b) sternite V." figureDoi="http://doi.org/10.5281/zenodo.4648327" httpUri="https://zenodo.org/record/4648327/files/figure.png" pageId="34" pageNumber="33">30</figureCitation>
,
<figureCitation id="134F3C88FFEF39352B427391FAC3F806" box="[1362,1391,1949,1973]" captionStart="Figures 31–32" captionStartId="33.[113,190,1986,2008]" captionTargetBox="[125,1459,280,1967]" captionTargetId="figure-9@33.[113,1474,230,1979]" captionTargetPageId="33" captionText="Figures 31–32. Tityopsis sheylae sp. n., male holotype (31) and female paratopotype (32), metasoma and telson: a) dorsal, b) lateral, and c) ventral views." figureDoi="http://doi.org/10.5281/zenodo.4648329" httpUri="https://zenodo.org/record/4648329/files/figure.png" pageId="34" pageNumber="33">32</figureCitation>
,
<figureCitation id="134F3C88FFEF39352B6B7391FA12F806" box="[1403,1470,1949,1973]" captionStart="Figure 36" captionStartId="37.[113,181,1923,1945]" captionTargetBox="[113,1474,249,1820]" captionTargetId="figure-8@37.[113,1474,249,1820]" captionTargetPageId="37" captionText="Figure 36. Tityopsis sheylae sp. n., photographed alive in its natural habitat unless otherwise noted: a) male holotype; b) small-sized male paratopotype; c) female paratopotype; d) captive female paratopotype, with mixed litter of instars 1–2; e) juvenile female paratopotype; f) just-molted, small-sized adult male paratopotype, being predated upon by the pholcid Ciboneya nuriae (the spider is not visible because while taking the photographs, it fled from the camera’s flash and hid in the crevice behind the scorpion). Photos courtesy Sheyla Yong (a–b)." figureDoi="http://doi.org/10.5281/zenodo.4648335" httpUri="https://zenodo.org/record/4648335/files/figure.png" pageId="34" pageNumber="33">36c–d</figureCitation>
;
<tableCitation id="C6F615B6FFEF39352D2073B1FC33F866" box="[816,927,1981,2005]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="28.[115,171,1108,1129]" captionStartId-1="34.[114,170,598,619]" captionTargetBox-1="[124,1452,250,581]" captionTargetId-1="graphics-728@34.[119,1470,238,585]" captionText-0="Table 3. Measurements (mm) of four adult types of Tityopsis sheylae sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 4. Pectinal tooth count variation in the four species of the genus Tityopsis treated in this paper. Abbreviations: number of pectines (N)," httpUri-1="http://table.plazi.org/id/DF0B7085FFEF39352E62765AFE5BFD3B" pageId="34" pageNumber="33" tableUuid-1="DF0B7085FFEF39352E62765AFE5BFD3B">Tabs. 3–4</tableCitation>
). Sexual dimorphism well-marked, differing from described male by:
<emphasis id="B900FC1FFFEF39352A1F73D1FB88F846" bold="true" box="[1039,1060,2013,2037]" pageId="34" pageNumber="33">1)</emphasis>
size conspicuously larger (32.1–34.0 mm);
<emphasis id="B900FC1FFFE939332EA0714FFF6AFAE8" bold="true" box="[176,198,1347,1371]" pageId="36" pageNumber="35">2)</emphasis>
tegument of carapace, tergites, sternites, metasoma and telson with a satin sheen;
<emphasis id="B900FC1FFFE939332F67716FFE21FAC8" bold="true" box="[375,397,1379,1403]" pageId="36" pageNumber="35">3)</emphasis>
pedipalps relatively shorter, slightly more robust (especially chela manus) and with carinae finer;
<emphasis id="B900FC1FFFE939332E6171AFFF2BFA08" bold="true" box="[113,135,1443,1467]" pageId="36" pageNumber="35">4)</emphasis>
carapace slightly wider posteriorly;
<emphasis id="B900FC1FFFE939332C1C71AFFD8EFA08" bold="true" box="[524,546,1443,1467]" pageId="36" pageNumber="35">5)</emphasis>
mesosoma relatively wider, with lateral sides markedly convex;
<emphasis id="B900FC1FFFE939332C2171CFFDEBFA68" bold="true" box="[561,583,1475,1499]" pageId="36" pageNumber="35">6)</emphasis>
pre-pectinal plate absent;
<emphasis id="B900FC1FFFE939332ED171EFFF7BFA48" bold="true" box="[193,215,1507,1531]" pageId="36" pageNumber="35">7)</emphasis>
genital papillae absent;
<emphasis id="B900FC1FFFE939332FC271EFFE44FA48" bold="true" box="[466,488,1507,1531]" pageId="36" pageNumber="35">8)</emphasis>
pectines relatively smaller, with teeth shorter, narrower and slightly fewer in number (13–15, mode 13–14), and with basal middle lamella greatly enlarged, widely oval in shape and whitish;
<emphasis id="B900FC1FFFE939332C58724FFDF1F9E8" bold="true" box="[584,605,1603,1627]" pageId="36" pageNumber="35">9)</emphasis>
sternite V with median and lateral smooth patches much smaller, less bulky and translucent;
<emphasis id="B900FC1FFFE939332F33728FFEEAF928" bold="true" box="[291,326,1667,1691]" pageId="36" pageNumber="35">10)</emphasis>
metasoma with carinae slightly stronger;
<emphasis id="B900FC1FFFE939332E6172AFFF3FF908" bold="true" box="[113,147,1699,1723]" pageId="36" pageNumber="35">11)</emphasis>
telson vesicle with many medium-sized granules scattered ventrally and ventrolaterally, and ventral median carina stronger.
</paragraph>
<caption id="DF0B7085FFEE39342E6373BBFB2BF85A" ID-DOI="http://doi.org/10.5281/zenodo.4648331" ID-Zenodo-Dep="4648331" httpUri="https://zenodo.org/record/4648331/files/figure.png" pageId="35" pageNumber="34" startId="35.[115,183,1975,1997]" targetBox="[113,1474,255,1923]" targetPageId="35">
<paragraph id="8BCB200DFFEE39342E6373BBFB2BF85A" blockId="35.[115,1473,1975,2026]" pageId="35" pageNumber="34">
<emphasis id="B900FC1FFFEE39342E6373BBFF7FF87F" bold="true" box="[115,211,1975,1997]" pageId="35" pageNumber="34">Figure 33</emphasis>
.
<taxonomicName id="4C745B8EFFEE39342ECD73B4FED6F87F" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[221,378,1975,1997]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="mulata" status="sp. nov.">
<emphasis id="B900FC1FFFEE39342ECD73B4FED6F87F" box="[221,378,1975,1997]" italics="true" pageId="35" pageNumber="34">Tityopsis mulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEE39342F6E73BBFE1DF87E" box="[382,433,1975,1997]" pageId="35" pageNumber="34" rank="species">
<emphasis id="B900FC1FFFEE39342F6E73BBFE39F87F" bold="true" box="[382,405,1975,1996]" pageId="35" pageNumber="34">sp</emphasis>
.
<emphasis id="B900FC1FFFEE39342F8F73BBFE00F87F" bold="true" box="[415,428,1975,1996]" pageId="35" pageNumber="34">n</emphasis>
.
</taxonomicNameLabel>
, photographed alive in its natural habitat unless otherwise noted:
<emphasis id="B900FC1FFFEE39342A0573BBFB85F87E" bold="true" box="[1045,1065,1975,1997]" pageId="35" pageNumber="34">a)</emphasis>
male holotype;
<emphasis id="B900FC1FFFEE39342AAE73BBFB46F87E" bold="true" box="[1214,1258,1975,1997]" pageId="35" pageNumber="34">b–c)</emphasis>
female paratopotypes;
<emphasis id="B900FC1FFFEE39342E6373D8FF32F859" bold="true" box="[115,158,2004,2026]" pageId="35" pageNumber="34">d–e)</emphasis>
captive female paratopotype, with litter of instars 1 (d) and 2 (e);
<emphasis id="B900FC1FFFEE39342D0973D8FC86F859" bold="true" box="[793,810,2004,2026]" pageId="35" pageNumber="34">f)</emphasis>
captive juvenile male paratopotype.
</paragraph>
</caption>
<caption id="DF0B7085FFE939332E6170F4FA6CFA99" ID-DOI="http://doi.org/10.5281/zenodo.4648333" ID-Zenodo-Dep="4648333" httpUri="https://zenodo.org/record/4648333/files/figure.png" pageId="36" pageNumber="35" startId="36.[113,189,1272,1294]" targetBox="[113,1474,234,1263]" targetPageId="36">
<paragraph id="8BCB200DFFE939332E6170F4FA6CFA99" blockId="36.[113,1472,1272,1323]" pageId="36" pageNumber="35">
<emphasis id="B900FC1FFFE939332E6170F4FF50FABE" bold="true" box="[113,252,1272,1294]" pageId="36" pageNumber="35">Figures 34–35</emphasis>
:
<emphasis id="B900FC1FFFE939332F1770F4FECAFABE" bold="true" box="[263,358,1272,1294]" pageId="36" pageNumber="35">Figure 34</emphasis>
.
<taxonomicName id="4C745B8EFFE939332F6070F5FE6BFABE" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[368,455,1272,1294]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="pumila" status="sp. nov.">
<emphasis id="B900FC1FFFE939332F6070F5FE6BFABE" box="[368,455,1272,1294]" italics="true" pageId="36" pageNumber="35">T. pumila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFE939332FDC70F4FE53FABD" box="[460,511,1272,1294]" pageId="36" pageNumber="35" rank="species">
<emphasis id="B900FC1FFFE939332FDC70F4FE4EFABE" bold="true" box="[460,482,1272,1293]" pageId="36" pageNumber="35">sp</emphasis>
.
<emphasis id="B900FC1FFFE939332FFD70F4FE56FABE" bold="true" box="[493,506,1272,1293]" pageId="36" pageNumber="35">n</emphasis>
.
</taxonomicNameLabel>
, male holotype photographed alive in its natural habitat (two different shots, photos courtesy Raimundo López-Silvero).
<emphasis id="B900FC1FFFE939332F197119FEC5FA99" bold="true" box="[265,361,1301,1323]" pageId="36" pageNumber="35">Figure 35</emphasis>
.
<taxonomicName id="4C745B8EFFE939332F63711AFDBBFA98" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[371,535,1302,1323]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="canizaresorum" status="sp. nov.">
<emphasis id="B900FC1FFFE939332F63711AFDBBFA98" box="[371,535,1302,1323]" italics="true" pageId="36" pageNumber="35">T. canizaresorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFE939332C0B7119FDE2FA99" box="[539,590,1301,1322]" pageId="36" pageNumber="35" rank="species">
<emphasis id="B900FC1FFFE939332C0B7119FD9DFA99" bold="true" box="[539,561,1301,1322]" pageId="36" pageNumber="35">sp</emphasis>
.
<emphasis id="B900FC1FFFE939332C2C7119FDE5FA99" bold="true" box="[572,585,1301,1322]" pageId="36" pageNumber="35">n</emphasis>
.
</taxonomicNameLabel>
, photographed alive in its natural habitat:
<emphasis id="B900FC1FFFE939332DC57119FC45FA98" bold="true" box="[981,1001,1301,1323]" pageId="36" pageNumber="35">a)</emphasis>
female holotype;
<emphasis id="B900FC1FFFE939332A807119FB08FA98" bold="true" box="[1168,1188,1301,1323]" pageId="36" pageNumber="35">b)</emphasis>
juvenile female paratopotype.
</paragraph>
</caption>
<paragraph id="8BCB200DFFE939332E617308FD75F86F" blockId="36.[113,772,1796,2044]" pageId="36" pageNumber="35">
VARIATION.The 44 paratopotypes are remarkably homogeneous for most morphological characters such as coloration, tegument sculpture, carination, setation and morphometric ratios, where variation was only minor and mostly due to eval differences (
<figureCitation id="134F3C88FFE939332E697388FF64F82F" box="[121,200,1924,1948]" captionStart="Figure 36" captionStartId="37.[113,181,1923,1945]" captionTargetBox="[113,1474,249,1820]" captionTargetId="figure-8@37.[113,1474,249,1820]" captionTargetPageId="37" captionText="Figure 36. Tityopsis sheylae sp. n., photographed alive in its natural habitat unless otherwise noted: a) male holotype; b) small-sized male paratopotype; c) female paratopotype; d) captive female paratopotype, with mixed litter of instars 1–2; e) juvenile female paratopotype; f) just-molted, small-sized adult male paratopotype, being predated upon by the pholcid Ciboneya nuriae (the spider is not visible because while taking the photographs, it fled from the camera’s flash and hid in the crevice behind the scorpion). Photos courtesy Sheyla Yong (a–b)." figureDoi="http://doi.org/10.5281/zenodo.4648335" httpUri="https://zenodo.org/record/4648335/files/figure.png" pageId="36" pageNumber="35">Fig. 36</figureCitation>
), e. g., older adults have slightly darker coloration and setation, granulation and claws worn-out to different degrees. The only noteworthy exceptions are discussed as follows.
</paragraph>
<paragraph id="8BCB200DFFE939332E6173E8FAF6FA0F" blockId="36.[113,772,1796,2044]" lastBlockId="36.[816,1475,1348,2044]" pageId="36" pageNumber="35">
Adult size (
<tableCitation id="C6F615B6FFE939332EE173E8FE9BF84F" box="[241,311,2020,2044]" captionStart="Table 3" captionStartId="28.[115,171,1108,1129]" captionText="Table 3. Measurements (mm) of four adult types of Tityopsis sheylae sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="36" pageNumber="35">Tab. 3</tableCitation>
) varied from
<quantity id="4C8C8DE8FFE939332FC173E8FDC2F848" box="[465,622,2020,2044]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" metricValueMax="2.68" metricValueMin="2.36" pageId="36" pageNumber="35" unit="mm" value="25.2" valueMax="26.8" valueMin="23.6">23.6–26.8 mm</quantity>
in males and 32.1–34.0 mm in females. There are two size classes even distinguishable to unaided eye in adult males (separated by a maximum gap of
<quantity id="4C8C8DE8FFE939332A147188FBCAFA28" box="[1028,1126,1412,1436]" metricMagnitude="-3" metricUnit="m" metricValue="3.14" pageId="36" pageNumber="35" unit="mm" value="3.14">3.14 mm</quantity>
, i.e., a 12% size difference), but only one in females (5% maximum size difference).
</paragraph>
<paragraph id="8BCB200DFFE939332D2071C8FAF9F8AF" blockId="36.[816,1475,1348,2044]" pageId="36" pageNumber="35">
Number of principal rows of denticles varied from 12–13 on both fixed and movable fingers, with a clearly defined mode of
<quantity id="4C8C8DE8FFE939332B9D71E8FA6FFA4F" box="[1421,1475,1508,1532]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="36" pageNumber="35" unit="in" value="12.0">12 in</quantity>
each case. Fixed, symmetrical counts for both pedipalps (i.e., same number of rows in all four fingers of the same individual), were the dominant occurrence by far: of
<specimenCount id="9D72EB84FFE939332AC37248FACCF9EF" box="[1235,1376,1604,1628]" pageId="36" pageNumber="35" type="generic">35 specimens</specimenCount>
analyzed (10 paratopotypes excluded due to having at least one finger broken or too damaged for reliable counting), 25 had 12/12 rows (including the
<typeStatus id="54CF9EAFFFE939332A1072A8FBF1F90F" box="[1024,1117,1700,1724]" pageId="36" pageNumber="35" type="holotype">holotype</typeStatus>
) and one had 13/13 rows (an adult female). The cases of asymmetry did not show any bias towards a defined finger, i.e., they occur in similar frequencies on fixed and movable fingers, and on left and right pedipalps.
</paragraph>
<paragraph id="8BCB200DFFE939332D207328FC01F84F" blockId="36.[816,1475,1348,2044]" pageId="36" pageNumber="35">
Pectinal tooth counts (
<tableCitation id="C6F615B6FFE939332A0D7328FBCDF88F" box="[1053,1121,1828,1852]" captionStart="Table 4" captionStartId="34.[114,170,598,619]" captionTargetBox="[124,1452,250,581]" captionTargetId="graphics-728@34.[119,1470,238,585]" captionText="Table 4. Pectinal tooth count variation in the four species of the genus Tityopsis treated in this paper. Abbreviations: number of pectines (N)," httpUri="http://table.plazi.org/id/DF0B7085FFEF39352E62765AFE5BFD3B" pageId="36" pageNumber="35" tableUuid="DF0B7085FFEF39352E62765AFE5BFD3B">Tab. 4</tableCitation>
) varied from
<quantity id="4C8C8DE8FFE939332AE37328FAFDF88F" box="[1267,1361,1828,1852]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="4.064" metricValueMin="3.302" pageId="36" pageNumber="35" unit="in" value="14.5" valueMax="16.0" valueMin="13.0">13–16 in</quantity>
males and
<quantity id="4C8C8DE8FFE939332D207348FC3DF8EF" box="[816,913,1860,1884]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="36" pageNumber="35" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
females, with clearly defined modes of 14 and 13– 14, respectively. Single-tooth asymmetry was a very common occurrence: 7 out of
<specimenCount id="9D72EB84FFE939332A027388FBDFF82F" box="[1042,1139,1924,1948]" pageId="36" pageNumber="35" type="male">15 males</specimenCount>
and 11 out of
<specimenCount id="9D72EB84FFE939332B047388FA25F82F" box="[1300,1417,1924,1948]" pageId="36" pageNumber="35" type="female">25 females</specimenCount>
(two adults of this sex had a pecten either missing or too damaged for counting); there were no specimens with two or more teeth asymmetry.
</paragraph>
<caption id="DF0B7085FFE839322E61738FFA1BF843" ID-DOI="http://doi.org/10.5281/zenodo.4648335" ID-Zenodo-Dep="4648335" httpUri="https://zenodo.org/record/4648335/files/figure.png" pageId="37" pageNumber="36" startId="37.[113,181,1923,1945]" targetBox="[113,1474,249,1820]" targetPageId="37">
<paragraph id="8BCB200DFFE839322E61738FFA1BF843" blockId="37.[113,1477,1923,2032]" pageId="37" pageNumber="36">
<emphasis id="B900FC1FFFE839322E61738FFF78F82B" bold="true" box="[113,212,1923,1945]" pageId="37" pageNumber="36">Figure 36</emphasis>
.
<taxonomicName id="4C745B8EFFE839322EF27388FE24F82A" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[226,392,1924,1945]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFE839322EF27388FE24F82A" box="[226,392,1924,1945]" italics="true" pageId="37" pageNumber="36">Tityopsis sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFE839322F9F7388FE69F82A" box="[399,453,1924,1945]" pageId="37" pageNumber="36" rank="species">
<emphasis id="B900FC1FFFE839322F9F7388FE0AF82A" bold="true" box="[399,422,1924,1945]" pageId="37" pageNumber="36">sp</emphasis>
.
<emphasis id="B900FC1FFFE839322FA37388FE6CF82A" bold="true" box="[435,448,1924,1945]" pageId="37" pageNumber="36">n</emphasis>
.
</taxonomicNameLabel>
, photographed alive in its natural habitat unless otherwise noted:
<emphasis id="B900FC1FFFE839322A5B738FFBF3F82A" bold="true" box="[1099,1119,1923,1945]" pageId="37" pageNumber="36">a)</emphasis>
male holotype;
<emphasis id="B900FC1FFFE839322AEF738FFAB8F82A" bold="true" box="[1279,1300,1923,1945]" pageId="37" pageNumber="36">b)</emphasis>
small-sized male paratopotype;
<emphasis id="B900FC1FFFE839322EEE73ACFEBCF805" bold="true" box="[254,272,1952,1974]" pageId="37" pageNumber="36">c)</emphasis>
female paratopotype;
<emphasis id="B900FC1FFFE839322FE173ACFDABF805" bold="true" box="[497,519,1952,1974]" pageId="37" pageNumber="36">d)</emphasis>
captive female paratopotype, with mixed litter of instars 1–2;
<emphasis id="B900FC1FFFE839322A6E73ACFB3CF805" bold="true" box="[1150,1168,1952,1974]" pageId="37" pageNumber="36">e)</emphasis>
juvenile female paratopotype;
<emphasis id="B900FC1FFFE839322E6173B1FF2DF860" bold="true" box="[113,129,1981,2003]" pageId="37" pageNumber="36">f)</emphasis>
just-molted, small-sized adult male paratopotype, being predated upon by the pholcid
<taxonomicName id="4C745B8EFFE839322DC473B1FBDBF860" authorityName="Huber &amp; Perez" authorityYear="2001" box="[980,1143,1981,2003]" class="Arachnida" family="Pholcidae" genus="Ciboneya" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="nuriae">
<emphasis id="B900FC1FFFE839322DC473B1FBDBF860" box="[980,1143,1981,2003]" italics="true" pageId="37" pageNumber="36">Ciboneya nuriae</emphasis>
</taxonomicName>
(the spider is not visible because while taking the photographs, it fled from the camera’s flash and hid in the crevice behind the scorpion). Photos courtesy Sheyla Yong (a–b).
</paragraph>
</caption>
<caption id="DF0B7085FFEB39312E61710CFB65FAA5" ID-DOI="http://doi.org/10.5281/zenodo.4648338" ID-Zenodo-Dep="4648338" box="[113,1225,1280,1302]" httpUri="https://zenodo.org/record/4648338/files/figure.png" pageId="38" pageNumber="37" startId="38.[113,190,1280,1302]" targetBox="[113,1474,240,1267]" targetPageId="38">
<paragraph id="8BCB200DFFEB39312E61710CFB65FAA5" blockId="38.[113,1225,1280,1302]" box="[113,1225,1280,1302]" pageId="38" pageNumber="37">
<emphasis id="B900FC1FFFEB39312E61710CFEADFAA6" bold="true" box="[113,257,1280,1302]" pageId="38" pageNumber="37">Figures 37–38</emphasis>
: Type localities of
<taxonomicName id="4C745B8EFFEB39312FAB710DFDF7FAA6" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[443,603,1280,1302]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="mulata" status="sp. nov.">
<emphasis id="B900FC1FFFEB39312FAB710DFDF7FAA6" box="[443,603,1280,1302]" italics="true" pageId="38" pageNumber="37">Tityopsis mulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEB39312C71710CFD3BFAA5" box="[609,663,1280,1302]" pageId="38" pageNumber="37" rank="species">
<emphasis id="B900FC1FFFEB39312C71710CFDD4FAA6" bold="true" box="[609,632,1280,1301]" pageId="38" pageNumber="37">sp</emphasis>
.
<emphasis id="B900FC1FFFEB39312C93710CFD3CFAA6" bold="true" box="[643,656,1280,1301]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
(37) and
<taxonomicName id="4C745B8EFFEB39312CE3710DFCFCFAA6" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[755,848,1280,1302]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="pumila" status="sp. nov.">
<emphasis id="B900FC1FFFEB39312CE3710DFCFCFAA6" box="[755,848,1280,1302]" italics="true" pageId="38" pageNumber="37">T. pumila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEB39312D45710CFC27FAA5" box="[853,907,1280,1302]" pageId="38" pageNumber="37" rank="species">
<emphasis id="B900FC1FFFEB39312D45710CFCC0FAA6" bold="true" box="[853,876,1280,1301]" pageId="38" pageNumber="37">sp</emphasis>
.
<emphasis id="B900FC1FFFEB39312D68710CFC29FAA6" bold="true" box="[888,901,1280,1301]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
(38),
<emphasis id="B900FC1FFFEB39312DD5710CFC75FAA5" bold="true" box="[965,985,1280,1302]" pageId="38" pageNumber="37">a)</emphasis>
habitat,
<emphasis id="B900FC1FFFEB39312A3D710CFBEEFAA5" bold="true" box="[1069,1090,1280,1302]" pageId="38" pageNumber="37">b)</emphasis>
microhabitat.
</paragraph>
</caption>
<paragraph id="8BCB200DFFEB39312E617139FE4FF99E" blockId="38.[113,771,1333,2029]" pageId="38" pageNumber="37">
AFFINITIES. By adults exhibiting the unusual combination of a size large for the genus (over
<quantity id="4C8C8DE8FFEB39312FDE7159FDB6FADF" box="[462,538,1365,1389]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="38" pageNumber="37" unit="mm" value="23.0">23 mm</quantity>
in males and
<quantity id="4C8C8DE8FFEB39312CA67159FCAFFADF" box="[694,771,1365,1389]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="38" pageNumber="37" unit="mm" value="32.0">32 mm</quantity>
in females) with a light yellowish, essentially immaculate coloration, this species resembles only
<taxonomicName id="4C745B8EFFEB39312C007199FD35FA1E" baseAuthorityName="Armas" baseAuthorityYear="1974" box="[528,665,1429,1453]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B900FC1FFFEB39312C007199FD35FA1E" box="[528,665,1429,1453]" italics="true" pageId="38" pageNumber="37">T. inaequalis</emphasis>
</taxonomicName>
. All other congeners (either described or not) are combined conversely, i.e., if coloration is pale, then size is small and vice versa (actually, most of them are medium-sized and reddish-colored with well-defined darker patterns).
</paragraph>
<paragraph id="8BCB200DFFEB39312EB17239FEE1F8FE" blockId="38.[113,771,1333,2029]" pageId="38" pageNumber="37">
Adults of
<taxonomicName id="4C745B8EFFEB39312F097239FE07F9FE" baseAuthorityName="Armas" baseAuthorityYear="1974" box="[281,427,1589,1613]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B900FC1FFFEB39312F097239FE07F9FE" box="[281,427,1589,1613]" italics="true" pageId="38" pageNumber="37">T. inaequalis</emphasis>
</taxonomicName>
can be easily separated from those of
<taxonomicName id="4C745B8EFFEB39312EC87259FEE6F9DE" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[216,330,1621,1645]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEB39312EC87259FEE6F9DE" box="[216,330,1621,1645]" italics="true" pageId="38" pageNumber="37">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEB39312F447259FE3AF9DF" box="[340,406,1621,1645]" pageId="38" pageNumber="37" rank="species">
<emphasis id="B900FC1FFFEB39312F447259FEC2F9DE" bold="true" box="[340,366,1621,1645]" pageId="38" pageNumber="37">sp</emphasis>
.
<emphasis id="B900FC1FFFEB39312F907259FE23F9DE" bold="true" box="[384,399,1621,1645]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
by the following characters:
<emphasis id="B900FC1FFFEB39312CFC7258FCAFF9DF" bold="true" box="[748,771,1620,1644]" pageId="38" pageNumber="37">1)</emphasis>
size slightly but consistently larger (
<quantity id="4C8C8DE8FFEB39312C067279FD3DF93F" box="[534,657,1653,1677]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="38" pageNumber="37" unit="mm" value="27.5" valueMax="29.0" valueMin="26.0">26–29 mm</quantity>
in males,
<quantity id="4C8C8DE8FFEB39312E617299FF47F91F" box="[113,235,1685,1709]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="38" pageNumber="37" unit="mm" value="34.5" valueMax="36.0" valueMin="33.0">33–36 mm</quantity>
in females);
<emphasis id="B900FC1FFFEB39312F987298FE33F91F" bold="true" box="[392,415,1684,1708]" pageId="38" pageNumber="37">2)</emphasis>
female pedipalp chela slightly shorter and more robust (length/width ratio:
<quantity id="4C8C8DE8FFEB39312C6572B8FCAFF97E" box="[629,771,1716,1741]" metricMagnitude="-2" metricUnit="m" metricValue="8.813799999999999" metricValueMax="8.9154" metricValueMin="8.7122" pageId="38" pageNumber="37" unit="in" value="3.4699999999999998" valueMax="3.51" valueMin="3.43">3.43–3.51 in</quantity>
females);
<emphasis id="B900FC1FFFEB39312EF772D8FF52F95F" bold="true" box="[231,254,1748,1772]" pageId="38" pageNumber="37">3)</emphasis>
female pectines with basal middle lamella slightly smaller and more oval in shape;
<emphasis id="B900FC1FFFEB39312C2A72F8FDFDF8BF" bold="true" box="[570,593,1780,1804]" pageId="38" pageNumber="37">4)</emphasis>
male sternite V with median smooth patch narrower, longer than wide and lanceolate in shape.
</paragraph>
<paragraph id="8BCB200DFFEB39312EB17359FE75F85E" blockId="38.[113,771,1333,2029]" pageId="38" pageNumber="37">
Besides, both species are widely allopatric in isolated orographic systems:
<taxonomicName id="4C745B8EFFEB39312F457379FE4DF83E" baseAuthorityName="Armas" baseAuthorityYear="1974" box="[341,481,1909,1933]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B900FC1FFFEB39312F457379FE4DF83E" box="[341,481,1909,1933]" italics="true" pageId="38" pageNumber="37">T. inaequalis</emphasis>
</taxonomicName>
in Sierra de Los Órganos (
<collectingRegion id="49B0EEEFFFEB39312E687399FEDCF81E" box="[120,368,1941,1965]" country="Cuba" name="Pinar del Rio" pageId="38" pageNumber="37">Pinar del Río Province</collectingRegion>
) and
<taxonomicName id="4C745B8EFFEB39312FBC7399FDBBF81E" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[428,535,1941,1965]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEB39312FBC7399FDBBF81E" box="[428,535,1941,1965]" italics="true" pageId="38" pageNumber="37">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEB39312C0E7399FDF7F81F" box="[542,603,1941,1965]" pageId="38" pageNumber="37" rank="species">
<emphasis id="B900FC1FFFEB39312C0E7399FD9BF81E" bold="true" box="[542,567,1941,1965]" pageId="38" pageNumber="37">sp</emphasis>
.
<emphasis id="B900FC1FFFEB39312C557399FDF8F81E" bold="true" box="[581,596,1941,1965]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
in Escaleras de Jaruco (
<collectingRegion id="49B0EEEFFFEB39312ED673B9FE02F87E" box="[198,430,1973,1997]" country="Cuba" name="Mayabeque" pageId="38" pageNumber="37">Mayabeque Province</collectingRegion>
), with their nearest occurrences separated by
<quantity id="4C8C8DE8FFEB39312EEE73D9FEE3F85E" box="[254,335,2005,2029]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="38" pageNumber="37" unit="km" value="170.0">170 km</quantity>
air-distance.
</paragraph>
<paragraph id="8BCB200DFFEB39312D207139FBD2F9BE" blockId="38.[816,1475,1333,2029]" pageId="38" pageNumber="37">
ECOLOGICAL NOTES.
<taxonomicName id="4C745B8EFFEB39312A077139FB2FFAFE" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[1047,1155,1333,1357]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEB39312A077139FB2FFAFE" box="[1047,1155,1333,1357]" italics="true" pageId="38" pageNumber="37">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEB39312A9C7139FB67FAFF" box="[1164,1227,1333,1357]" pageId="38" pageNumber="37" rank="species">
<emphasis id="B900FC1FFFEB39312A9C7139FB09FAFE" bold="true" box="[1164,1189,1333,1357]" pageId="38" pageNumber="37">sp</emphasis>
.
<emphasis id="B900FC1FFFEB39312AA57139FB68FAFE" bold="true" box="[1205,1220,1333,1357]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
has been found so far only inside a single cave (
<figureCitation id="134F3C88FFEB39312A717159FB1BFADE" box="[1121,1207,1365,1389]" captionStart="Figures 39–40" captionStartId="39.[113,190,1347,1369]" captionTargetBox="[113,1474,249,1338]" captionTargetId="figure-349@39.[113,1474,249,1338]" captionTargetPageId="39" captionText="Figures 39–40: Type localities of Tityopsis canizaresorum sp. n. (39) and T. sheylae sp. n. (40), a) habitat, b) microhabitat (photo courtesy Sheyla Yong)." figureDoi="http://doi.org/10.5281/zenodo.4648340" httpUri="https://zenodo.org/record/4648340/files/figure.png" pageId="38" pageNumber="37">Fig. 40</figureCitation>
). It occurs scattered all along it, from the first chamber right beyond the entrance (not completely dark, relatively cold, less humid and essentially devoid of bat guano), through the deepest corridors (in absolute darkness, very hot, with water-saturated air and densely covered by bat guano).
</paragraph>
<paragraph id="8BCB200DFFEB39312D707219FA6FF8BE" blockId="38.[816,1475,1333,2029]" pageId="38" pageNumber="37">We always used the UV-detection technique to find it and were able to notice that apparently because of the different microclimatic conditions, the scorpions changed their behavior accordingly: those from the first chamber were mostly found hidden under rocks (only one adult of each sex was found walking between the rocks, in separate collecting events), while all those living in the deep were totally or partially exposed on the walls and floor, either walking slowly or sit-and-wait hunting.</paragraph>
<paragraph id="8BCB200DFFEB39302D707319FE2DFA5A" blockId="38.[816,1475,1333,2029]" lastBlockId="39.[113,772,1425,2025]" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37">
It lives syntopically with the buthid
<taxonomicName id="4C745B8EFFEB39312AF27319FC73F8FE" authority="(Latreille, 1804)" baseAuthorityName="Latreille" baseAuthorityYear="1804" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B900FC1FFFEB39312AF27319FA6EF89E" box="[1250,1474,1813,1837]" italics="true" pageId="38" pageNumber="37">Centruroides gracilis</emphasis>
(Latreille, 1804)
</taxonomicName>
, but one species always outnumbers the other depending on the cave site. In the hottest chamber
<taxonomicName id="4C745B8EFFEB39312B277359FA31F8DE" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[1335,1437,1877,1901]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEB39312B277359FA31F8DE" box="[1335,1437,1877,1901]" italics="true" pageId="38" pageNumber="37">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEB39312BB37359FCEAF83F" pageId="38" pageNumber="37" rank="species">
<emphasis id="B900FC1FFFEB39312BB37359FA10F8DE" bold="true" box="[1443,1468,1877,1901]" pageId="38" pageNumber="37">sp</emphasis>
.
<emphasis id="B900FC1FFFEB39312D207379FC93F83E" bold="true" box="[816,831,1909,1933]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
is rare (only three specimens found so far), while hundreds of
<taxonomicName id="4C745B8EFFEB39312D207399FC33F81E" baseAuthorityName="Latreille" baseAuthorityYear="1804" box="[816,927,1941,1965]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B900FC1FFFEB39312D207399FC33F81E" box="[816,927,1941,1965]" italics="true" pageId="38" pageNumber="37">C. gracilis</emphasis>
</taxonomicName>
virtually cover all available surfaces. Conversely, in the colder chambers and corridors we have found less than 15 individuals of
<taxonomicName id="4C745B8EFFEB39312DD673D9FB98F85E" baseAuthorityName="Latreille" baseAuthorityYear="1804" box="[966,1076,2005,2029]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B900FC1FFFEB39312DD673D9FB98F85E" box="[966,1076,2005,2029]" italics="true" pageId="38" pageNumber="37">C. gracilis</emphasis>
</taxonomicName>
in total, against 42 of
<taxonomicName id="4C745B8EFFEB39312B0A73D9FA2DF85E" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[1306,1409,2005,2029]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEB39312B0A73D9FA2DF85E" box="[1306,1409,2005,2029]" italics="true" pageId="38" pageNumber="37">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEB39312B9873D9FA6EF85F" box="[1416,1474,2005,2029]" pageId="38" pageNumber="37" rank="species">
<emphasis id="B900FC1FFFEB39312B9873D9FA0DF85E" bold="true" box="[1416,1441,2005,2029]" pageId="38" pageNumber="37">sp</emphasis>
.
<emphasis id="B900FC1FFFEB39312BBD73D9FA10F85E" bold="true" box="[1453,1468,2005,2029]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
Interestingly, no cases of mutual predation have been observed yet, in spite of the coexistence and abundance of both scorpion species all along the cave.
</paragraph>
<caption id="DF0B7085FFEA39302E61714FFF56FAC5" ID-DOI="http://doi.org/10.5281/zenodo.4648340" ID-Zenodo-Dep="4648340" httpUri="https://zenodo.org/record/4648340/files/figure.png" pageId="39" pageNumber="38" startId="39.[113,190,1347,1369]" targetBox="[113,1474,249,1338]" targetPageId="39">
<paragraph id="8BCB200DFFEA39302E61714FFF56FAC5" blockId="39.[113,1471,1347,1398]" pageId="39" pageNumber="38">
<emphasis id="B900FC1FFFEA39302E61714FFEAEFAEA" bold="true" box="[113,258,1347,1369]" pageId="39" pageNumber="38">Figures 39–40</emphasis>
: Type localities of
<taxonomicName id="4C745B8EFFEA39302FAF7148FD00FAEA" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[447,684,1348,1369]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="canizaresorum" status="sp. nov.">
<emphasis id="B900FC1FFFEA39302FAF7148FD00FAEA" box="[447,684,1348,1369]" italics="true" pageId="39" pageNumber="38">Tityopsis canizaresorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEA39302CA27148FD45FAEA" box="[690,745,1348,1369]" pageId="39" pageNumber="38" rank="species">
<emphasis id="B900FC1FFFEA39302CA27148FD65FAEA" bold="true" box="[690,713,1348,1369]" pageId="39" pageNumber="38">sp</emphasis>
.
<emphasis id="B900FC1FFFEA39302CC67148FD4FFAEA" bold="true" box="[726,739,1348,1369]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
(39) and
<taxonomicName id="4C745B8EFFEA39302D587148FC06FAEA" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[840,938,1348,1369]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEA39302D587148FC06FAEA" box="[840,938,1348,1369]" italics="true" pageId="39" pageNumber="38">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEA39302DA17148FC44FAEA" box="[945,1000,1348,1369]" pageId="39" pageNumber="38" rank="species">
<emphasis id="B900FC1FFFEA39302DA17148FC64FAEA" bold="true" box="[945,968,1348,1369]" pageId="39" pageNumber="38">sp</emphasis>
.
<emphasis id="B900FC1FFFEA39302DC57148FC4EFAEA" bold="true" box="[981,994,1348,1369]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
(40),
<emphasis id="B900FC1FFFEA39302A34714FFB9BFAEA" bold="true" box="[1060,1079,1347,1369]" pageId="39" pageNumber="38">a)</emphasis>
habitat,
<emphasis id="B900FC1FFFEA39302A9C714FFB0EFAEA" bold="true" box="[1164,1186,1347,1369]" pageId="39" pageNumber="38">b)</emphasis>
microhabitat (photo courtesy Sheyla Yong).
</paragraph>
</caption>
<paragraph id="8BCB200DFFEA39302EB171FDFE93F89A" blockId="39.[113,772,1425,2025]" pageId="39" pageNumber="38">
On
<date id="FFCA06CDFFEA39302EDA71FDFEFAF9BA" box="[202,342,1521,1545]" pageId="39" pageNumber="38" value="2019-04-03">3 April 2019</date>
, we found a small-sized adult male that was being predated upon by an adult female of the spider
<taxonomicName id="4C745B8EFFEA39302E61723DFD6AF9FA" authority="Huber &amp; Perez, 2001" authorityName="Huber &amp; Perez" authorityYear="2001" box="[113,710,1585,1609]" class="Arachnida" family="Pholcidae" genus="Ciboneya" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="nuriae">
<emphasis id="B900FC1FFFEA39302E61723DFE87F9FA" box="[113,299,1585,1609]" italics="true" pageId="39" pageNumber="38">Ciboneya nuriae</emphasis>
Huber &amp; Pérez, 2001 (Pholcidae)
</taxonomicName>
. The scorpion had just molted (its exoskeleton was still soft and not completely stretched yet, see
<figureCitation id="134F3C88FFEA39302FA1727DFDA4F93A" box="[433,520,1649,1673]" captionStart="Figure 36" captionStartId="37.[113,181,1923,1945]" captionTargetBox="[113,1474,249,1820]" captionTargetId="figure-8@37.[113,1474,249,1820]" captionTargetPageId="37" captionText="Figure 36. Tityopsis sheylae sp. n., photographed alive in its natural habitat unless otherwise noted: a) male holotype; b) small-sized male paratopotype; c) female paratopotype; d) captive female paratopotype, with mixed litter of instars 1–2; e) juvenile female paratopotype; f) just-molted, small-sized adult male paratopotype, being predated upon by the pholcid Ciboneya nuriae (the spider is not visible because while taking the photographs, it fled from the camera’s flash and hid in the crevice behind the scorpion). Photos courtesy Sheyla Yong (a–b)." figureDoi="http://doi.org/10.5281/zenodo.4648335" httpUri="https://zenodo.org/record/4648335/files/figure.png" pageId="39" pageNumber="38">Fig. 36f</figureCitation>
) and was already dead about
<quantity id="4C8C8DE8FFEA39302EA5729DFF40F91A" box="[181,236,1681,1705]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="38" unit="cm" value="1.0">1 cm</quantity>
away from its exuvium, while the spider sucked it somewhere between the chelicerae and pedipalp coxae. We deduce the scorpion anchored to molt beside the web and once it came out of the exuvium, it became entangled and was then killed by the spider.
</paragraph>
<paragraph id="8BCB200DFFEA39302EB1733DFDD1F85A" blockId="39.[113,772,1425,2025]" pageId="39" pageNumber="38">According to its collecting label, the male collected on 1995 (see above in Other Material Examined section) was also found dead, entangled in a spider’s web. Unfortunately, this sample is currently missing from its repository and the identity of the spider cannot be established, but this second finding evidences that spiders are a routine predator of this scorpion.</paragraph>
</subSubSection>
<subSubSection id="C36E7386FFEA39302D20719DFB4EF95A" pageId="39" pageNumber="38" type="discussion">
<paragraph id="8BCB200DFFEA39302D20719DFB4EF95A" blockId="39.[816,1476,1425,1769]" pageId="39" pageNumber="38">
COMMENTS. The habitat outside the cave is mesophylous semicaducifolious forest on limestone karstic soil (
<figureCitation id="134F3C88FFEA39302B4871BDFA19FA7A" box="[1368,1461,1457,1481]" captionStart="Figures 39–40" captionStartId="39.[113,190,1347,1369]" captionTargetBox="[113,1474,249,1338]" captionTargetId="figure-349@39.[113,1474,249,1338]" captionTargetPageId="39" captionText="Figures 39–40: Type localities of Tityopsis canizaresorum sp. n. (39) and T. sheylae sp. n. (40), a) habitat, b) microhabitat (photo courtesy Sheyla Yong)." figureDoi="http://doi.org/10.5281/zenodo.4648340" httpUri="https://zenodo.org/record/4648340/files/figure.png" pageId="39" pageNumber="38">Fig. 40a</figureCitation>
), typical for most members of the genus. Nevertheless, repeated intensive searches conducted by us and some collaborators there and in other mogotes of this plateau, never yielded any evidences of an epigean occurrence of
<taxonomicName id="4C745B8EFFEA39302AC5723EFAECF9FA" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" box="[1237,1344,1585,1609]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEA39302AC5723EFAECF9FA" box="[1237,1344,1585,1609]" italics="true" pageId="39" pageNumber="38">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEA39302B57723DFA29F9FA" box="[1351,1413,1585,1609]" pageId="39" pageNumber="38" rank="species">
<emphasis id="B900FC1FFFEA39302B57723DFACCF9FA" bold="true" box="[1351,1376,1585,1609]" pageId="39" pageNumber="38">sp</emphasis>
.
<emphasis id="B900FC1FFFEA39302B7F723DFAD2F9FA" bold="true" box="[1391,1406,1585,1609]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
(e.g., live specimens or remains of carcasses and exuvia). Only two females tentatively referable to
<taxonomicName id="4C745B8EFFEA39302A6E727EFABEF93A" baseAuthorityName="Moreno" baseAuthorityYear="1940" box="[1150,1298,1650,1673]" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="inexpectata">
<emphasis id="B900FC1FFFEA39302A6E727EFABEF93A" box="[1150,1298,1650,1673]" italics="true" pageId="39" pageNumber="38">T. inexpectata</emphasis>
</taxonomicName>
were found, one of them from IES collection and which vouchers the Escaleras de Jaruco record by
<bibRefCitation id="EFE55DFCFFEA39302A0872BDFB7FF97A" author="ARMAS" box="[1048,1235,1713,1737]" pageId="39" pageNumber="38" pagination="1 - 15" refId="ref23103" refString="ARMAS, L. F. DE. 1974. Escorpiones del archipielago cubano. 3. Genero Tityus C. L. Koch, 1836 (Scorpionida: Buthidae). Poeyana, 135: 1 - 15." type="journal article" year="1974">Armas (1974: 5)</bibRefCitation>
. Thus, we classify
<taxonomicName id="4C745B8EFFEA39302BBE72BEFC2CF95A" authority="Teruel &amp; Rodríguez-Cabrera, 2020" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2020" class="Arachnida" family="Buthidae" genus="Tityopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="sheylae" status="sp. nov.">
<emphasis id="B900FC1FFFEA39302BBE72BEFC2CF95A" italics="true" pageId="39" pageNumber="38">T. sheylae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2334164FFEA39302D9772DDFC6FF95A" box="[903,963,1745,1769]" pageId="39" pageNumber="38" rank="species">
<emphasis id="B900FC1FFFEA39302D9772DDFC0CF95A" bold="true" box="[903,928,1745,1769]" pageId="39" pageNumber="38">sp</emphasis>
.
<emphasis id="B900FC1FFFEA39302DBE72DDFC11F95A" bold="true" box="[942,957,1745,1769]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
herein as a true troglobite.
</paragraph>
</subSubSection>
<subSubSection id="C36E7386FFEA39302D20731DFC70F87A" pageId="39" pageNumber="38" type="distribution">
<paragraph id="8BCB200DFFEA39302D20731DFC70F87A" blockId="39.[816,1474,1809,1993]" pageId="39" pageNumber="38">
DISTRIBUTION (
<figureCitation id="134F3C88FFEA39302DC2731DFBB8F89A" box="[978,1044,1809,1833]" captionStart="Figure 2" captionStartId="4.[113,181,849,871]" captionTargetBox="[113,1474,234,838]" captionTargetId="figure-658@4.[113,1474,234,838]" captionTargetPageId="4" captionText="Figure 2. Western Cuba, showing the complete geographical distribution of the genus Tityopsis (white symbols) and overlaid, that of the four species of Tityopsis treated herein (red symbols, complemented with habitus of their holotypes): T. mulata sp. n. (star), T. canizaresorum sp. n. (rhombs, the southernmost is the type locality), T. pumila sp. n. (triangle) and T. sheylae sp. n. (square). Image frame size, 500 x 250 km." figureDoi="http://doi.org/10.5281/zenodo.4648297" httpUri="https://zenodo.org/record/4648297/files/figure.png" pageId="39" pageNumber="38">Fig. 2</figureCitation>
). This species is apparently a troglobite, known only from the
<typeStatus id="54CF9EAFFFEA39302A09733EFBEBF8F9" box="[1049,1095,1842,1866]" pageId="39" pageNumber="38">type</typeStatus>
locality. Cueva del Indio is located in the southern mogotes of Escaleras de Jaruco, which is a medium-sized (15 x
<quantity id="4C8C8DE8FFEA39302A1A737DFBEFF83A" box="[1034,1091,1905,1929]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="38" unit="km" value="5.0">5 km</quantity>
), roughly east-west oriented, karstic limestone plateau, isolated in the middle of the
<collectingRegion id="49B0EEEFFFEA39302B78739DFC3AF87A" country="Cuba" name="Matanzas" pageId="39" pageNumber="38">Habana-Matanzas</collectingRegion>
Plain.
</paragraph>
</subSubSection>
</treatment>
</document>
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