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<document id="E5B8D42ACDBCA64922EA367A7C44EE8F" ID-DOI="10.5281/zenodo.273781" ID-GBIF-Dataset="381d12c0-7801-41b4-997b-48b5742e7c55" ID-ISSN="1175-5326" ID-Zenodo-Dep="273781" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459843893481" checkinUser="plazi" docAuthor="Cisneros-Heredia, Diego F. &amp; Meza-Ramos, Pa Ú L" docDate="2007" docId="038D3172FFC0E3103DA1FA8FDE40FD22" docLanguage="en" docName="zt01485p041.pdf" docOrigin="Zootaxa 1485" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Cochranella amelie Cisneros-Heredia &amp; Meza-Ramos, new species" docType="treatment" docVersion="7" lastPageNumber="37" masterDocId="FFB4490AFFC1E3143D36FF80DF67FF98" masterDocTitle="An enigmatic new species of Glassfrog (Amphibia: Anura: Centrolenidae) from the Amazonian Andean slopes of Ecuador" masterLastPageNumber="41" masterPageNumber="33" pageNumber="34" updateTime="1698224602763" updateUser="plazi">
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<mods:title id="60C9E6829164A2DFB59AEC7D006B506F">An enigmatic new species of Glassfrog (Amphibia: Anura: Centrolenidae) from the Amazonian Andean slopes of Ecuador</mods:title>
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<mods:namePart id="2BA3E295AA0D9A036854F14659F4E560">Cisneros-Heredia, Diego F.</mods:namePart>
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<mods:namePart id="A46479FEDF331D5C06AB81DB66E92EE6">Meza-Ramos, Pa Ú L</mods:namePart>
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<mods:part id="FC3D6092ADFC349195B41E5ECEB47EDF">
<mods:date id="54152BE3FF0F5049E8AE5B11DE7FFB2B">2007</mods:date>
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<mods:identifier id="8515908AE1A1C0387B6C78E7185A7A1F" type="DOI">10.5281/zenodo.273781</mods:identifier>
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<treatment id="038D3172FFC0E3103DA1FA8FDE40FD22" ID-DOI="http://doi.org/10.5281/zenodo.6243985" ID-GBIF-Taxon="119361126" ID-Zenodo-Dep="6243985" LSID="urn:lsid:plazi:treatment:038D3172FFC0E3103DA1FA8FDE40FD22" httpUri="http://treatment.plazi.org/id/038D3172FFC0E3103DA1FA8FDE40FD22" lastPageId="4" lastPageNumber="37" pageId="1" pageNumber="34">
<subSubSection id="C33ED3EFFFC0E3153DA1FA8FDF9DFAD4" pageId="1" pageNumber="34" type="nomenclature">
<paragraph id="8B9B8064FFC0E3153DA1FA8FDCA8FAB1" blockId="1.[151,975,1295,1356]" box="[151,975,1295,1321]" pageId="1" pageNumber="34">
<heading id="D0D33708FFC0E3153DA1FA8FDCA8FAB1" bold="true" box="[151,975,1295,1321]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
<emphasis id="B9505C76FFC0E3153DA1FA8FDCA8FAB1" bold="true" box="[151,975,1295,1321]" pageId="1" pageNumber="34">
<taxonomicName id="4C24FBE7FFC0E3153DA1FA8FDC54FAB1" ID-CoL="WNQP" authority="Cisneros-Heredia &amp; Meza-Ramos" authorityName="Cisneros-Heredia &amp; Meza-Ramos" box="[151,819,1295,1321]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="1" pageNumber="34" phylum="Chordata" rank="species" species="amelie" status="sp. nov.">
<emphasis id="B9505C76FFC0E3153DA1FA8FDEE3FAB1" bold="true" box="[151,388,1295,1321]" italics="true" pageId="1" pageNumber="34">Cochranella amelie</emphasis>
Cisneros-Heredia &amp; Meza-Ramos
</taxonomicName>
,
<taxonomicNameLabel id="A263E10DFFC0E3153E76FA8FDCA8FAB1" box="[832,975,1295,1321]" pageId="1" pageNumber="34" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B8064FFC0E3153DA1FAB2DF9DFAD4" blockId="1.[151,975,1295,1356]" box="[151,250,1330,1356]" pageId="1" pageNumber="34">
<figureCitation id="131F9CE1FFC0E3153DA1FAB2DF9DFAD4" box="[151,250,1330,1356]" captionStart="FIGURE 1" captionStartId="2.[151,255,967,991]" captionTargetBox="[415,1171,434,945]" captionTargetId="figure@2.[415,1171,434,945]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Cochranella amelie in life (DHMECN 3066), adult male, SVL = 18.3 mm." httpUri="https://zenodo.org/record/273782/files/figure.png" pageId="1" pageNumber="34">Figure 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33ED3EFFFC0E3153DA1FA03DA7FF98C" pageId="1" pageNumber="34" type="materials_examined">
<paragraph id="8B9B8064FFC0E3153DA1FA03DD86FA74" blockId="1.[151,1436,1410,2008]" pageId="1" pageNumber="34">
<emphasis id="B9505C76FFC0E3153DA1FA03DE64FA04" box="[151,259,1411,1436]" italics="true" pageId="1" pageNumber="34">
<typeStatus id="549F3EC6FFC0E3153DA1FA03DE64FA04" box="[151,259,1411,1436]" pageId="1" pageNumber="34" type="holotype">Holotype</typeStatus>
</emphasis>
.—DHMECN 3066 (field number
<collectionCode id="ED3518A1FFC0E3153FADFA02DDBEFA04" box="[667,729,1410,1436]" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="1" pageNumber="34">PMR</collectionCode>
073), adult male collected at Comunidad de Oglán, Cantón Arajuno, Provincia de Pastaza,
<collectingCountry id="F333C0F4FFC0E3153F3EFA2ADC73FA5C" box="[520,788,1450,1476]" name="Ecuador" pageId="1" pageNumber="34">República del Ecuador</collectingCountry>
(
<geoCoordinate id="EE10E6A3FFC0E3153E12FA2ADCDEFA5C" box="[804,953,1450,1476]" direction="south" orientation="latitude" pageId="1" pageNumber="34" precision="15" value="-1.3180555">01°1865” S</geoCoordinate>
,
<geoCoordinate id="EE10E6A3FFC0E3153EF0FA2ADB03FA5C" box="[966,1124,1450,1476]" direction="west" orientation="longitude" pageId="1" pageNumber="34" precision="15" value="-77.71139">77°4241” W</geoCoordinate>
,
<quantity id="4CDC2D81FFC0E3153946FA2ADBDDFA5C" box="[1136,1210,1450,1476]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="34" unit="m" value="600.0">600 m</quantity>
elevation),
<date id="FF9AA6A4FFC0E3153870FA2ADFB5FA74" pageId="1" pageNumber="34" value="2004-07-28">28 July 2004</date>
by Paủl Meza-Ramos and Fabricio Narváez.
</paragraph>
<paragraph id="8B9B8064FFC0E3153DF0FA7BDA7FF98C" blockId="1.[151,1436,1410,2008]" box="[198,1304,1530,1556]" pageId="1" pageNumber="34">
<emphasis id="B9505C76FFC0E3153DF0FA7BDE56F98C" box="[198,305,1531,1556]" italics="true" pageId="1" pageNumber="34">
<typeStatus id="549F3EC6FFC0E3153DF0FA7BDE56F98C" box="[198,305,1531,1556]" pageId="1" pageNumber="34" type="paratype">Paratype</typeStatus>
</emphasis>
.—DHMECN 3591 (field number
<collectionCode id="ED3518A1FFC0E3153FF0FA7ADC63F98C" box="[710,772,1530,1556]" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="1" pageNumber="34">PMR</collectionCode>
072), adult male collected with the
<typeStatus id="549F3EC6FFC0E315399CFA7ADA73F98C" box="[1194,1300,1530,1556]" pageId="1" pageNumber="34" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33ED3EFFFC0E3163DF0F9A2DBB7FB6C" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="diagnosis">
<paragraph id="8B9B8064FFC0E3163DF0F9A2DCA2FEE2" blockId="1.[151,1436,1410,2008]" lastBlockId="2.[151,1436,152,378]" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
<emphasis id="B9505C76FFC0E3153DF0F9A2DE21F9A4" bold="true" box="[198,326,1570,1596]" pageId="1" pageNumber="34">Diagnosis.</emphasis>
<taxonomicName id="4C24FBE7FFC0E3153C78F9A3DD50F9A4" box="[334,567,1571,1596]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="1" pageNumber="34" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC0E3153C78F9A3DD50F9A4" box="[334,567,1571,1596]" italics="true" pageId="1" pageNumber="34">Cochranella amelie</emphasis>
</taxonomicName>
is a small centrolenid diagnosed from all other Glassfrogs by having: (1) vomerine teeth absent; (2) snout slightly truncate in dorsal view and rounded in lateral view; nostrils slightly elevated producing a depression in the internarial area; loreal region concave; (3) tympanic annulus evident, oriented dorsolaterally with posterior inclination; very weak supratympanic fold above the tympanum; (4) dorsal skin shagreened; (5) ventral and subanal area granular, with two large, rounded, flat subanal tubercles, other anal ornamentations absent; (6) parietal peritoneum completely clear/transparent (without white pigmentation/guanophores), all visceral peritonea covered by white pigmentation; (7) liver bulbous; (8) humeral spine absent in adult males; (9) webbing on hand, basal between finger I and II, outer fingers
<date id="FF9AA6A4FFC0E31539D6F8BDDA6DF8CF" box="[1248,1290,1853,1879]" pageId="1" pageNumber="34">II 1</date>
-3
<date id="FF9AA6A4FFC0E3153805F8BDDA01F8CF" box="[1331,1382,1853,1879]" pageId="1" pageNumber="34">III 1</date>
1/2
<date id="FF9AA6A4FFC0E31538A7F8BDDFD0F81D" pageId="1" pageNumber="34">1 IV</date>
; (10) webbing on feet
<date id="FF9AA6A4FFC0E3153CFBF8EBDE97F81D" box="[461,496,1899,1925]" pageId="1" pageNumber="34">I 1</date>
-1 1/2
<date id="FF9AA6A4FFC0E3153F02F8EBDD07F81D" box="[564,608,1899,1925]" pageId="1" pageNumber="34">II 1</date>
-
<date id="FF9AA6A4FFC0E3153F43F8EBDDC8F81D" box="[629,687,1896,1925]" bridgedPair="-" pageId="1" pageNumber="34">2- III</date>
12-
<date id="FF9AA6A4FFC0E3153FC3F8EBDC4AF81D" box="[757,813,1899,1925]" pageId="1" pageNumber="34">IV 2</date>
-0+ V; (11) low non-enameled ulnar fold; low nonenameled inner tarsal fold; (12) nuptial excrescences present, type-I; concealed prepollex; (13) first finger longer than second, (14) eye diameter larger than width of disc on finger III; (15) colour in life, uniform bluish green dorsal surfaces; (16) colour in preservative, uniform lavender dorsal surfaces; (17) iris coloration in life between dark to light grey; in preservative, cream background with dense lavender punctuations especially concentrated around the midline; (18) extensive melanophores on finger IV and toes IV and V; (19) males call from the upper surfaces of leaves; advertisement call unrecorded; (20) fighting behaviour unknown; (21) onelayer egg clutches deposited on the upper surfaces of leaves next to streams; (22) tadpoles unknown; (23) snout-vent length in adult males 18.1-18.3 (n = 2); females unknown.
</paragraph>
<caption id="DF5BD0ECFFC3E3163DA1FC47DBA1FC46" box="[151,1222,967,991]" httpUri="https://zenodo.org/record/273782/files/figure.png" pageId="2" pageNumber="35" targetBox="[415,1171,434,945]" targetPageId="2">
<paragraph id="8B9B8064FFC3E3163DA1FC47DBA1FC46" blockId="2.[151,1222,967,991]" box="[151,1222,967,991]" pageId="2" pageNumber="35">
<emphasis id="B9505C76FFC3E3163DA1FC47DE7EFC47" bold="true" box="[151,281,967,991]" pageId="2" pageNumber="35">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="4C24FBE7FFC3E3163C90FC47DD1EFC46" box="[422,633,967,990]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC3E3163C90FC47DD1EFC46" box="[422,633,967,990]" italics="true" pageId="2" pageNumber="35">Cochranella amelie</emphasis>
</taxonomicName>
in life (DHMECN 3066), adult male, SVL = 18.3 mm.
</paragraph>
</caption>
<paragraph id="8B9B8064FFC3E3163DF0FBB9DBB7FB6C" blockId="2.[151,1437,1081,2035]" pageId="2" pageNumber="35">
<emphasis id="B9505C76FFC3E3163DF0FBB9DE14FBCB" bold="true" box="[198,371,1081,1107]" pageId="2" pageNumber="35">Comparisons.</emphasis>
<taxonomicName id="4C24FBE7FFC3E3163C49FBBADD0BFBCB" box="[383,620,1082,1107]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC3E3163C49FBBADD0BFBCB" box="[383,620,1082,1107]" italics="true" pageId="2" pageNumber="35">Cochranella amelie</emphasis>
</taxonomicName>
differs from all species currently placed in
<taxonomicName id="4C24FBE7FFC3E31639B9FBBADA77FBCB" box="[1167,1296,1082,1107]" class="Amphibia" family="Centrolenidae" genus="Centrolene" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B9505C76FFC3E31639B9FBBADA77FBCB" box="[1167,1296,1082,1107]" italics="true" pageId="2" pageNumber="35">Centrolene</emphasis>
</taxonomicName>
by lacking humeral spines in adult males. Further, most species of
<taxonomicName id="4C24FBE7FFC3E3163E29FBE2DCC7FBE3" box="[799,928,1122,1147]" class="Amphibia" family="Centrolenidae" genus="Centrolene" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B9505C76FFC3E3163E29FBE2DCC7FBE3" box="[799,928,1122,1147]" italics="true" pageId="2" pageNumber="35">Centrolene</emphasis>
</taxonomicName>
have the parietal peritoneum totally or partially covered by white pigmentation, with the exception of
<emphasis id="B9505C76FFC3E3163E6DFB0ADB7AFB3B" box="[859,1053,1162,1187]" italics="true" pageId="2" pageNumber="35">
Ce.
<taxonomicName id="4C24FBE7FFC3E3163EBDFB0ADB7AFB3B" box="[907,1053,1162,1187]" class="Amphibia" family="Centrolenidae" genus="Centrolene" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="mariaelenae">mariaelenae</taxonomicName>
</emphasis>
.
<taxonomicName id="4C24FBE7FFC3E316391AFB0ADA2FFB3B" box="[1068,1352,1162,1187]" class="Amphibia" family="Centrolenidae" genus="Centrolene" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="mariaelenae">
<emphasis id="B9505C76FFC3E316391AFB0ADA2FFB3B" box="[1068,1352,1162,1187]" italics="true" pageId="2" pageNumber="35">Centrolene mariaelenae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C24FBE7FFC3E3163DEEFB32DEA5FB53" box="[216,450,1202,1227]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC3E3163DEEFB32DEA5FB53" box="[216,450,1202,1227]" italics="true" pageId="2" pageNumber="35">Cochranella amelie</emphasis>
</taxonomicName>
by having a green dorsum with many small dark punctuations and scattered larger dark spots in life that turns cream lavender with dark punctuations and spots in preservative.
</paragraph>
</subSubSection>
<subSubSection id="C33ED3EFFFC3E3173DF3FA82DF90FEC5" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35" type="reference_group">
<paragraph id="8B9B8064FFC3E3163DF3FA82DAFBF877" blockId="2.[151,1437,1081,2035]" pageId="2" pageNumber="35">
<taxonomicName id="4C24FBE7FFC3E3163DF3FA82DECAFA83" ID-CoL="WNQP" box="[197,429,1282,1307]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC3E3163DF3FA82DECAFA83" box="[197,429,1282,1307]" italics="true" pageId="2" pageNumber="35">Cochranella amelie</emphasis>
</taxonomicName>
differs from most species of
<taxonomicName id="4C24FBE7FFC3E3163E33FA82DCFFFA83" ID-CoL="3RQB" box="[773,920,1282,1307]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B9505C76FFC3E3163E33FA82DCFFFA83" box="[773,920,1282,1307]" italics="true" pageId="2" pageNumber="35">Cochranella</emphasis>
</taxonomicName>
by having a completely transparent parietal peritoneum; only
<taxonomicName id="4C24FBE7FFC3E3163C5FFAAADD6CFADB" ID-CoL="WNSX" box="[361,523,1322,1347]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="pulverata">
<emphasis id="B9505C76FFC3E3163C5FFAAADD6CFADB" box="[361,523,1322,1347]" italics="true" pageId="2" pageNumber="35">Co. pulverata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C24FBE7FFC3E3163F70FAAADC63FADB" ID-CoL="WNQS" box="[582,772,1322,1347]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="antisthenesi">
<emphasis id="B9505C76FFC3E3163F70FAAADC63FADB" box="[582,772,1322,1347]" italics="true" pageId="2" pageNumber="35">Co. antisthenesi</emphasis>
</taxonomicName>
are similar in this character.
<taxonomicName id="4C24FBE7FFC3E3163968FAAADA0EFADB" ID-CoL="WNSX" box="[1118,1385,1322,1347]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="pulverata">
<emphasis id="B9505C76FFC3E3163968FAAADA0EFADB" box="[1118,1385,1322,1347]" italics="true" pageId="2" pageNumber="35">Cochranella pulverata</emphasis>
</taxonomicName>
differs by having a sloping snout in lateral view, vomerine teeth, enameled folds or tubercles on arms and tarsus, green dorsum with white spots in life, pale lavender to cream lavender dorsum in preservative, and a larger size (
<quantity id="4CDC2D81FFC3E3163DE3FA22DEE3FA24" box="[213,388,1442,1468]" metricMagnitude="-2" metricUnit="m" metricValue="2.535" metricValueMax="2.93" metricValueMin="2.1399999999999997" pageId="2" pageNumber="35" unit="mm" value="25.35" valueMax="29.3" valueMin="21.4">21.429.3 mm</quantity>
SVL in adult males
<taxonomicName id="4C24FBE7FFC3E3163F48FA22DC46FA23" ID-CoL="WNSX" box="[638,801,1442,1467]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="pulverata">
<emphasis id="B9505C76FFC3E3163F48FA22DC46FA23" box="[638,801,1442,1467]" italics="true" pageId="2" pageNumber="35">Co. pulverata</emphasis>
</taxonomicName>
; Savage 2002; Cisneros-Heredia
<emphasis id="B9505C76FFC3E3163985FA22DB88FA23" box="[1203,1263,1442,1467]" italics="true" pageId="2" pageNumber="35">et al.</emphasis>
unpubl. data).
<taxonomicName id="4C24FBE7FFC3E3163DA1FA4ADED8FA7B" ID-CoL="WNQS" box="[151,447,1482,1507]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="antisthenesi">
<emphasis id="B9505C76FFC3E3163DA1FA4ADED8FA7B" box="[151,447,1482,1507]" italics="true" pageId="2" pageNumber="35">Cochranella antisthenesi</emphasis>
</taxonomicName>
differs by having vomerine teeth, green dorsum with light spots, and a larger size (
<quantity id="4CDC2D81FFC3E3163DA9FA72DE36F994" box="[159,337,1522,1548]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" metricValueMax="2.62" metricValueMin="2.1399999999999997" pageId="2" pageNumber="35" unit="mm" value="23.799999999999997" valueMax="26.2" valueMin="21.4">21.426.2 mm</quantity>
SVL in adult males
<taxonomicName id="4C24FBE7FFC3E3163F54FA72DC42F993" ID-CoL="WNQS" box="[610,805,1522,1547]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="antisthenesi">
<emphasis id="B9505C76FFC3E3163F54FA72DC42F993" box="[610,805,1522,1547]" italics="true" pageId="2" pageNumber="35">Co. antisthenesi</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB5FD95FFC3E3163E0CFA72DBC0F994" author="Senaris" box="[826,1191,1522,1548]" pageId="2" pageNumber="41" refString="Senaris, J. C. &amp; Ayarzaguena, J. (2005) Revision taxonomica de la Familia Centrolenidae (Amphibia; Anura) de Venezuela. Publicaciones del Comite Espanol del Programa Hombre y Biosfera - Red IberoMaB de la UNESCO. Sevilla, Espana, 337 pp." type="book" year="2005">Señaris &amp; Ayarzagüena 2005</bibRefCitation>
). Species related to
<taxonomicName id="4C24FBE7FFC3E3163DA1F99ADECFF9AB" ID-CoL="WNRV" box="[151,424,1562,1587]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="B9505C76FFC3E3163DA1F99ADECFF9AB" box="[151,424,1562,1587]" italics="true" pageId="2" pageNumber="35">Cochranella granulosa</emphasis>
</taxonomicName>
(= “
<taxonomicName id="4C24FBE7FFC3E3163CE8F99ADD10F9AB" ID-CoL="WNRV" box="[478,631,1562,1587]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="B9505C76FFC3E3163CE8F99ADD10F9AB" box="[478,631,1562,1587]" italics="true" pageId="2" pageNumber="35">C. granulosa</emphasis>
</taxonomicName>
species-group”) resemble to
<taxonomicName id="4C24FBE7FFC3E3163EF9F99ADB29F9AB" ID-CoL="WNQP" box="[975,1102,1562,1587]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC3E3163EF9F99ADB29F9AB" box="[975,1102,1562,1587]" italics="true" pageId="2" pageNumber="35">Co. amelie</emphasis>
</taxonomicName>
in having guanophores over the visceral peritonea, but differ by having sloping snouts, crenulated dermal folds, vomerine teeth, and white parietal peritoneum. Other species of
<taxonomicName id="4C24FBE7FFC3E3163F79F9EADD85F91B" ID-CoL="3RQB" box="[591,738,1642,1667]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B9505C76FFC3E3163F79F9EADD85F91B" box="[591,738,1642,1667]" italics="true" pageId="2" pageNumber="35">Cochranella</emphasis>
</taxonomicName>
having white visceral peritonea are
<taxonomicName id="4C24FBE7FFC3E31639BEF9EADA03F91B" ID-CoL="WNR3" box="[1160,1380,1642,1667]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cariticommata">
<emphasis id="B9505C76FFC3E31639BEF9EADA03F91B" box="[1160,1380,1642,1667]" italics="true" pageId="2" pageNumber="35">Co. cariticommata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C24FBE7FFC3E3163844F9EADFBAF933" ID-CoL="5ZHG6" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="wileyi">
<emphasis id="B9505C76FFC3E3163844F9EADFBAF933" italics="true" pageId="2" pageNumber="35">Co. wileyi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C24FBE7FFC3E3163DDAF912DEF4F933" ID-CoL="WNQQ" box="[236,403,1682,1707]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="ametarsia">
<emphasis id="B9505C76FFC3E3163DDAF912DEF4F933" box="[236,403,1682,1707]" italics="true" pageId="2" pageNumber="35">Co. ametarsia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C24FBE7FFC3E3163C95F912DD7CF933" ID-CoL="WNSC" box="[419,539,1682,1707]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="midas">
<emphasis id="B9505C76FFC3E3163C95F912DD7CF933" box="[419,539,1682,1707]" italics="true" pageId="2" pageNumber="35">Co. midas</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C24FBE7FFC3E3163F6BF912DC47F933" ID-CoL="WNSM" box="[605,800,1682,1707]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="oyampiensis">
<emphasis id="B9505C76FFC3E3163F6BF912DC47F933" box="[605,800,1682,1707]" italics="true" pageId="2" pageNumber="35">Co. oyampiensis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C24FBE7FFC3E3163E19F912DB7FF933" ID-CoL="WNQP" box="[815,1048,1682,1707]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC3E3163E19F912DB7FF933" box="[815,1048,1682,1707]" italics="true" pageId="2" pageNumber="35">Cochranella amelie</emphasis>
</taxonomicName>
differs from all by having transparent parietal peritoneum;
<taxonomicName id="4C24FBE7FFC3E3163CEBF93ADC45F94B" ID-CoL="WNR3" box="[477,802,1722,1747]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cariticommata">
<emphasis id="B9505C76FFC3E3163CEBF93ADC45F94B" box="[477,802,1722,1747]" italics="true" pageId="2" pageNumber="35">Cochranella cariticommata</emphasis>
</taxonomicName>
further differs by having a green dorsum with yellow dots in life that turns lavender with light dots in preservative, less hand and foot webbing, lacking guanophores on the hepatic and intestinal peritonea, and by its larger body size (
<quantity id="4CDC2D81FFC3E316393CF88ADBDFF8BC" box="[1034,1208,1802,1828]" metricMagnitude="-2" metricUnit="m" metricValue="2.295" metricValueMax="2.35" metricValueMin="2.2399999999999998" pageId="2" pageNumber="35" unit="mm" value="22.95" valueMax="23.5" valueMin="22.4">22.423.5 mm</quantity>
SVL in adult male
<taxonomicName id="4C24FBE7FFC3E3163DA1F8B2DE14F8D3" ID-CoL="WNR3" box="[151,371,1842,1867]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="cariticommata">
<emphasis id="B9505C76FFC3E3163DA1F8B2DE14F8D3" box="[151,371,1842,1867]" italics="true" pageId="2" pageNumber="35">Co. cariticommata</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB5FD95FFC3E3163CB5F8B2DD64F8D4" author="Wild" box="[387,515,1842,1868]" pageId="2" pageNumber="41" refString="Wild, E. R. (1994) Two new species of Centrolenid Frogs from the Amazonian slope of the Cordillera Oriental, Ecuador. Journal of Herpetology, 28 (3), 299 - 310." type="journal article" year="1994">Wild 1994</bibRefCitation>
;
<bibRefCitation id="EFB5FD95FFC3E3163F39F8B2DC95F8D4" author="Cisneros-Heredia" box="[527,1010,1842,1868]" pageId="2" pageNumber="40" refString="Cisneros-Heredia, D. F. &amp; Yanez-Munoz, M. (2007) Amphibia, Anura, Centrolenidae, Centrolene balionotum, Centrolene geckoideum, and Cochranella cariticommata: Distribution extension, new provincial records, Ecuador. Check List, 3 (1), 39 - 42." type="journal article" year="2007">Cisneros-Heredia &amp; Yánez-Muñoz 2007</bibRefCitation>
).
<taxonomicName id="4C24FBE7FFC3E3163930F8B2DB81F8D3" ID-CoL="5ZHG6" box="[1030,1254,1842,1867]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="wileyi">
<emphasis id="B9505C76FFC3E3163930F8B2DB81F8D3" box="[1030,1254,1842,1867]" italics="true" pageId="2" pageNumber="35">Cochranella wileyi</emphasis>
</taxonomicName>
differs by having less hand and foot webbing, lacking guanophores on the hepatic and intestinal peritonea, and by its larger body size (
<quantity id="4CDC2D81FFC3E3163C23F802DEA6F804" box="[277,449,1922,1948]" metricMagnitude="-2" metricUnit="m" metricValue="2.47" metricValueMax="2.61" metricValueMin="2.33" pageId="2" pageNumber="35" unit="mm" value="24.7" valueMax="26.1" valueMin="23.3">23.326.1 mm</quantity>
SVL in adult male
<taxonomicName id="4C24FBE7FFC3E3163F91F802DC79F803" ID-CoL="5ZHG6" box="[679,798,1922,1947]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="wileyi">
<emphasis id="B9505C76FFC3E3163F91F802DC79F803" box="[679,798,1922,1947]" italics="true" pageId="2" pageNumber="35">Co. wileyi</emphasis>
</taxonomicName>
; Guayasamin
<emphasis id="B9505C76FFC3E3163EF1F802DB66F803" box="[967,1025,1922,1947]" italics="true" pageId="2" pageNumber="35">et al.</emphasis>
2006).
<taxonomicName id="4C24FBE7FFC3E316396AF802DA0DF803" ID-CoL="WNQQ" box="[1116,1386,1922,1947]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="ametarsia">
<emphasis id="B9505C76FFC3E316396AF802DA0DF803" box="[1116,1386,1922,1947]" italics="true" pageId="2" pageNumber="35">Cochranella ametarsia</emphasis>
</taxonomicName>
differs by having a green dorsum with dark spots in life that turns lavender with dark spots in preservative, guanophores covering the anterior 1/4 parietal peritoneum, and by lacking guanophores on the hepatic peritonea.
</paragraph>
<paragraph id="8B9B8064FFC2E3173DA1FF18DF90FEC5" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">
<taxonomicName id="4C24FBE7FFC2E3173DA1FF18DE1FFF29" ID-CoL="WNSC" box="[151,376,152,177]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="midas">
<emphasis id="B9505C76FFC2E3173DA1FF18DE1FFF29" box="[151,376,152,177]" italics="true" pageId="3" pageNumber="36">Cochranella midas</emphasis>
</taxonomicName>
differs by having a green dorsum with yellow spots in life that turns lavender with light spots in preservative, guanophores covering the anterior 1/3 to 1/2 parietal peritoneum, and by lacking guanophores on the hepatic peritonea.
<taxonomicName id="4C24FBE7FFC2E3173F24FF6EDC5BFE9F" ID-CoL="WNSM" box="[530,828,238,263]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="oyampiensis">
<emphasis id="B9505C76FFC2E3173F24FF6EDC5BFE9F" box="[530,828,238,263]" italics="true" pageId="3" pageNumber="36">Cochranella oyampiensis</emphasis>
</taxonomicName>
differs by having a green dorsum with dark spots in life that turns lavender with dark spots in preservative, and guanophores covering the anterior 1/4 parietal peritoneum.
</paragraph>
</subSubSection>
<subSubSection id="C33ED3EFFFC2E3103DF3FEECDD86FECA" lastPageId="4" lastPageNumber="37" pageId="3" pageNumber="36" type="description">
<paragraph id="8B9B8064FFC2E3173DF3FEECDBCCFDBD" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">
<taxonomicName id="4C24FBE7FFC2E3173DF3FEECDECBFE1D" box="[197,428,364,389]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC2E3173DF3FEECDECBFE1D" box="[197,428,364,389]" italics="true" pageId="3" pageNumber="36">Cochranella amelie</emphasis>
</taxonomicName>
differs from all species related to
<taxonomicName id="4C24FBE7FFC2E3173E0AFEECDBD8FE1D" box="[828,1215,364,389]" class="Amphibia" family="Centrolenidae" genus="Hyalinobatrachium" kingdom="Animalia" order="Anura" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="fleischmanni">
<emphasis id="B9505C76FFC2E3173E0AFEECDBD8FE1D" box="[828,1215,364,389]" italics="true" pageId="3" pageNumber="36">Hyalinobatrachium fleischmanni</emphasis>
</taxonomicName>
(= “
<taxonomicName id="4C24FBE7FFC2E31739C2FEECDE1BFE35" class="Amphibia" family="Centrolenidae" genus="Hyalinobatrachium" kingdom="Animalia" order="Anura" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="fleischmanni">
<emphasis id="B9505C76FFC2E31739C2FEECDE1BFE35" italics="true" pageId="3" pageNumber="36">Hyalinobatrachium fleischmanni</emphasis>
</taxonomicName>
species-group”) by having a uniform green dorsum in life that turns lavender in preservative, by depositing its egg clutches on the upper surfaces of leaves; and from most species by having a type-I nuptial pad.
<taxonomicName id="4C24FBE7FFC2E3173C11FE64DD69FE65" box="[295,526,484,509]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="amelie">
<emphasis id="B9505C76FFC2E3173C11FE64DD69FE65" box="[295,526,484,509]" italics="true" pageId="3" pageNumber="36">Cochranella amelie</emphasis>
</taxonomicName>
differs from all species related to
<taxonomicName id="4C24FBE7FFC2E3173EACFE64DB4AFE65" box="[922,1069,484,509]" class="Amphibia" family="Centrolenidae" genus="Hyalinobatrachium" kingdom="Animalia" order="Anura" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="parvulum">
<emphasis id="B9505C76FFC2E3173EACFE64DB4AFE65" box="[922,1069,484,509]" italics="true" pageId="3" pageNumber="36">H. parvulum</emphasis>
</taxonomicName>
(= “
<taxonomicName id="4C24FBE7FFC2E3173954FE64DFA4FDBD" class="Amphibia" family="Centrolenidae" genus="Hyalinobatrachium" kingdom="Animalia" order="Anura" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="parvulum">
<emphasis id="B9505C76FFC2E3173954FE64DFA4FDBD" italics="true" pageId="3" pageNumber="36">Hyalinobatrachium parvulum</emphasis>
</taxonomicName>
species-group”) by having a transparent urinary bladder, and lacking vomerine teeth.
</paragraph>
<paragraph id="8B9B8064FFC2E3173DF0FDB3DFB3FC45" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">
<emphasis id="B9505C76FFC2E3173DF0FDB3DD47FDD5" bold="true" box="[198,544,563,589]" pageId="3" pageNumber="36">
Description of the
<typeStatus id="549F3EC6FFC2E3173C9BFDB3DD7DFDD5" box="[429,538,563,589]" pageId="3" pageNumber="36" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male, SVL =
<quantity id="4CDC2D81FFC2E3173E27FDB3DC19FDD5" box="[785,894,563,589]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="3" pageNumber="36" unit="mm" value="18.3">18.3 mm</quantity>
. Body slender. Head distinct, wider than long, and wider than body; HW/HL = 1.14, HW/SVL = 0.36, HL/SVL = 0.32. Snout short, slightly truncate in dorsal view and rounded in lateral view, EN/HL = 0.28; nostrils slightly elevated producing a depression in the internarial area; loreal region concave; canthus rostralis rounded, fairly indistinct, a shallow platform between the canthus rostralis; lips slightly flared. Eyes large, ED/HL = 0.40, directed anterolaterally at about 48° from midline, eyes can be seen when viewed from below, interorbital area wider than eye diameter, IOD/ED = 1.61, EN/ED = 0.70, EN/IOD = 0.43. Tympanic annulus evident, oriented dorsolaterally with slight posterodorsal inclination; very weak supratympanic fold above the tympanum. Dentigerous processes of vomers absent, choanae moderately sized, elliptical, widely separated between them, very close to the margin of mouth; tongue oval, not indented posteriorly; vocal slits paired, extending from anterior base of tongue to angles of jaws.
</paragraph>
<paragraph id="8B9B8064FFC2E3173DF3FC6BDEB6FBE5" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">Skin of dorsal surfaces finely shagreened, without warts, spicules, or tubercles; ventral surfaces granular. Cloacal opening directed posteriorly at upper level of thighs; no distinct cloacal sheath; a pair of large, oval, flat tubercles on ventral surfaces of tights below vent, other anal ornamentation absent, ventral/subanal skin granular but not enameled.</paragraph>
<paragraph id="8B9B8064FFC2E3173DF3FB0BDAEAFAEB" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">
Upper arm thin, forearm robust, breadth of upper arm about half that of forearm. Humeral spine absent; low non-enameled ulnar fold present. Relative lengths of fingers III&gt; IV&gt; I&gt; II; webbing basal between finger I and II, outer fingers
<date id="FF9AA6A4FFC2E3173CFBFB5EDE9FFB60" box="[461,504,1246,1272]" pageId="3" pageNumber="36">II 1</date>
-3
<date id="FF9AA6A4FFC2E3173F12FB5EDD3FFB60" box="[548,600,1246,1272]" pageId="3" pageNumber="36">III 1</date>
1/2
<date id="FF9AA6A4FFC2E3173FB1FB5EDDDCFB60" box="[647,699,1246,1272]" pageId="3" pageNumber="36">1 IV</date>
; bulla absent; finger discs wide, between truncate to rounded; disc on third finger slightly larger than those on toes, and shorter than eye diameter, 3DW/ED = 0.52; subarticular tubercles rounded to oval but flat and almost indistinct, palmar surface granular; palmar tubercle large, rounded, flat; tenar tubercle elliptic. Concealed prepollex, unpigmented nuptial excrescences
<typeStatus id="549F3EC6FFC2E31739EBFAD9DA70FAEB" box="[1245,1303,1369,1395]" pageId="3" pageNumber="36">Type</typeStatus>
I present.
</paragraph>
<paragraph id="8B9B8064FFC2E3173DF3FA01DA67F981" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">
<collectingCountry id="F333C0F4FFC2E3173DF3FA01DE67FA03" box="[197,256,1409,1435]" name="India" pageId="3" pageNumber="36">Hind</collectingCountry>
limbs slender; TL/SVL = 0.61, FL/SVL = 0.48. Low non-enameled inner tarsal fold present; inner metatarsal tubercle elliptical, elongated; outer metatarsal tubercle indistinct. Subarticular tubercles rounded and flat; supernumerary tubercles indistinct. Webbing on foot
<date id="FF9AA6A4FFC2E3173E5AFA54DCEBFA76" box="[876,908,1492,1518]" pageId="3" pageNumber="36">I 1</date>
-1 1/2
<date id="FF9AA6A4FFC2E3173EFBFA54DC91FA76" box="[973,1014,1492,1518]" pageId="3" pageNumber="36">II 1</date>
-
<date id="FF9AA6A4FFC2E317393AFA54DB24FA76" box="[1036,1091,1489,1518]" bridgedPair="-" pageId="3" pageNumber="36">2- III</date>
12-
<date id="FF9AA6A4FFC2E31739B5FA54DBD0FA76" box="[1155,1207,1492,1518]" pageId="3" pageNumber="36">IV 2</date>
-0+ V; disc on toe I oval with a distinct globular point at the end, all other discs bluntly truncate without projections.
</paragraph>
<paragraph id="8B9B8064FFC2E3173DF0F9A7DE57F921" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">
<emphasis id="B9505C76FFC2E3173DF0F9A7DE89F9D9" bold="true" box="[198,494,1575,1601]" pageId="3" pageNumber="36">
Coloration of
<typeStatus id="549F3EC6FFC2E3173C4AF9A7DE8EF9D9" box="[380,489,1575,1601]" pageId="3" pageNumber="36" type="holotype">holotype</typeStatus>
.
</emphasis>
In life, all dorsal surfaces bluish green (between cerulean green and viridian green), whitish upper lip, yellowish cream flanks, greenish throat, transparent ventral skin, all visceral peritonea covered by guanophores. Iris between dark to light grey, and black pupil with a yellow pupillary ring. Bones green.
</paragraph>
<paragraph id="8B9B8064FFC2E3173DF0F947DB55F8C1" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">In preservative, all dorsal surfaces uniform lavender, without dark or light flecks, spots, ocelli or other marks. Upper eyelid dark lavender. Venter transparent cream. Parietal peritoneum without guanophores. Guanophores covering the pericardium and most visceral peritonea (esophageal, stomach, intestinal, hepatic, and gallbladder), except for the urinary bladder and the renal capsule that are clear.</paragraph>
<paragraph id="8B9B8064FFC2E3173DF0F8E7DD79F849" blockId="3.[151,1437,152,2001]" pageId="3" pageNumber="36">
<emphasis id="B9505C76FFC2E3173DF0F8E7DE26F819" bold="true" box="[198,321,1895,1921]" pageId="3" pageNumber="36">Variation.</emphasis>
The adult male
<typeStatus id="549F3EC6FFC2E3173F34F8E7DD00F819" box="[514,615,1895,1921]" pageId="3" pageNumber="36" type="paratype">paratype</typeStatus>
is similar to the
<typeStatus id="549F3EC6FFC2E3173E1CF8E7DCF6F819" box="[810,913,1895,1921]" pageId="3" pageNumber="36" type="holotype">holotype</typeStatus>
in all diagnostic characters, including coloration. The only pertinent difference is the renal capsule in the
<typeStatus id="549F3EC6FFC2E3173E42F80FDCBEF831" box="[884,985,1935,1961]" pageId="3" pageNumber="36" type="paratype">paratype</typeStatus>
covered by guanophores. Variation of measurements is provided below.
</paragraph>
<paragraph id="8B9B8064FFC5E3103DF0FF17DD86FECA" blockId="4.[151,1436,151,698]" pageId="4" pageNumber="37">
<emphasis id="B9505C76FFC5E3103DF0FF17DEE2FF29" bold="true" box="[198,389,151,177]" pageId="4" pageNumber="37">Measurements.</emphasis>
The measurements of the
<typeStatus id="549F3EC6FFC5E3103FF6FF18DC41FF2A" box="[704,806,152,178]" pageId="4" pageNumber="37" type="holotype">holotype</typeStatus>
and
<typeStatus id="549F3EC6FFC5E3103E54FF18DCA0FF2A" box="[866,967,152,178]" pageId="4" pageNumber="37" type="paratype">paratype</typeStatus>
(in parentheses) are: snout-vent length, 18.3 (18.1) mm; head width, 6.6 (6.6) mm; head length, 5.8 (5.7) mm; horizontal eye diameter, 2.3 (2,3) mm; inter-orbital distance, 3.7 (3.4) mm; eye-nostril distance, 1.6 (1.6) mm; internarial distance between the nostrils, 1.9 (1.7) mm; tympanum diameter, 0.7 (0.7) mm; width of disc on the third finger, 1.2 (1.1) mm; tibia length, 11.1 (11.1) mm; foot length, 8.7 (9.1) mm.
</paragraph>
</subSubSection>
<subSubSection id="C33ED3EFFFC5E3103DF0FEDFDC43FE6A" pageId="4" pageNumber="37" type="etymology">
<paragraph id="8B9B8064FFC5E3103DF0FEDFDC43FE6A" blockId="4.[151,1436,151,698]" pageId="4" pageNumber="37">
<emphasis id="B9505C76FFC5E3103DF0FEDFDE34FEE1" bold="true" box="[198,339,351,377]" pageId="4" pageNumber="37">Etymology.</emphasis>
The specific name—a noun in apposition—of this new species of Glassfrog is for
<taxonomicName id="4C24FBE7FFC5E3103808FEE0DAF0FEE2" box="[1342,1431,352,378]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="38" phylum="Chordata" rank="species" species="amelie">Amelie</taxonomicName>
, protagonist of the extraordinary movie “Le Fabuleux Destin d'Amélie Poulain”; a movie where little details play an important role in the achievement of
<emphasis id="B9505C76FFC5E3103FF2FE30DC3BFE51" box="[708,860,432,457]" italics="true" pageId="4" pageNumber="37">joie de vivre</emphasis>
; like the important role that Glassfrogs and all amphibians and reptiles play in the health of our planet.
</paragraph>
</subSubSection>
<subSubSection id="C33ED3EFFFC5E3103DF0FE7FDE40FD22" pageId="4" pageNumber="37" type="distribution">
<paragraph id="8B9B8064FFC5E3103DF0FE7FDE40FD22" blockId="4.[151,1436,151,698]" pageId="4" pageNumber="37">
<emphasis id="B9505C76FFC5E3103DF0FE7FDD39FD81" bold="true" box="[198,606,511,537]" pageId="4" pageNumber="37">Distribution and natural history.</emphasis>
The two known specimens were collected on
<taxonomicName id="4C24FBE7FFC5E310394CFD80DB89FD81" box="[1146,1262,512,537]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="4" pageNumber="37" phylum="Tracheophyta" rank="genus">
<emphasis id="B9505C76FFC5E310394CFD80DB89FD81" box="[1146,1262,512,537]" italics="true" pageId="4" pageNumber="37">Heliconia</emphasis>
</taxonomicName>
leaves (ca.
<quantity id="4CDC2D81FFC5E3103841FD80DFC9FDDA" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="37" unit="m" value="0.8">0.8 m</quantity>
above floor) over a small affluent of the Oglán River, in primary forest. Both specimens were calling on the upper surface of leaves. The
<typeStatus id="549F3EC6FFC5E3103CDEFDD0DD2AFDF2" box="[488,589,592,618]" pageId="4" pageNumber="37" type="paratype">paratype</typeStatus>
(DHMECN 3591) was found next to a single-layer egg clutch deposited on the upper surface of a leaf. Other anurans found along the stream included
<taxonomicName id="4C24FBE7FFC5E3103973FDF8DA76FD09" box="[1093,1297,632,657]" class="Amphibia" family="Hylidae" genus="Hypsiboas" kingdom="Animalia" order="Anura" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="boans">
<emphasis id="B9505C76FFC5E3103973FDF8DA76FD09" box="[1093,1297,632,657]" italics="true" pageId="4" pageNumber="37">Hypsiboas boans</emphasis>
</taxonomicName>
and
<taxonomicName id="4C24FBE7FFC5E3103878FDF8DF98FD21" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B9505C76FFC5E3103878FDF8DF98FD21" italics="true" pageId="4" pageNumber="37">Osteocephalus</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
</treatment>
</document>