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<document id="671BCFF6D1E800CC77CB0A25018DF972" ID-DOI="http://dx.doi.org/10.3897/zookeys.456.8413" ID-GBIF-Dataset="ed1098b6-164c-45ae-98bc-dca2e1c70750" ID-PMC="PMC4329402" ID-Pensoft-Pub="1313-2970-456-109" ID-PubMed="25709524" ID-ZooBank="469905581AF047749A01DFAF08B665AE" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-456-109" ModsDocOrigin="ZooKeys 456" ModsDocTitle="On the Domene species of China, with descriptions of four new species (Coleoptera, Staphylinidae)" checkinTime="1451245088058" checkinUser="pensoft" docAuthor="Feldmann, Benedikt, Peng, Zhong &amp; Li, Li-Zhen" docDate="2014" docId="DB5ED4D7BC6FA4CE9E62819857C8C92D" docLanguage="en" docName="ZooKeys 456: 109-138" docOrigin="ZooKeys 456" docSource="http://dx.doi.org/10.3897/zookeys.456.8413" docTitle="Domene (Macromene) reitteri Koch 1939" docType="treatment" docVersion="4" lastPageNumber="122" masterDocId="FF86FFB04A1DF366FF8FFFD2E54CFF8B" masterDocTitle="On the Domene species of China, with descriptions of four new species (Coleoptera, Staphylinidae)" masterLastPageNumber="138" masterPageNumber="109" pageNumber="117" updateTime="1701317721500" updateUser="plazi">
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<mods:title id="0CDCFF782BF309B45C27F6AC98AB76ED">On the Domene species of China, with descriptions of four new species (Coleoptera, Staphylinidae)</mods:title>
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<mods:namePart id="9CAC2404ACA4C7B8273C660FB61F1EA9">Feldmann, Benedikt</mods:namePart>
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<mods:namePart id="E03EF4EAF2E992D464909745D739497A">Li, Li-Zhen</mods:namePart>
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<subSubSection id="FE63DB7A07943987E9D59BCB612C0147" pageId="8" pageNumber="117" type="multiple">
<paragraph id="863887D7E7AB40C4C0791466645261AF" pageId="8" pageNumber="117">
<pageBreakToken id="B00B2C11072F00AC7293FC4B4877936E" pageId="8" pageNumber="117" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Staphylinidae
</paragraph>
</subSubSection>
<subSubSection id="55338E376627883DF7B0B9A864F8B8B5" pageId="8" pageNumber="117" type="nomenclature">
<paragraph id="9506B1DCAEAC8F18C43804D6C3352A98" pageId="8" pageNumber="117">
<taxonomicName id="7752E34B633051C47EFA8E3D282096AC" ID-CoL="3797K" authority="Koch, 1939" authorityName="Koch" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene (Macromene) reitteri" order="Coleoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="reitteri" subGenus="Macromene">Domene (Macromene) reitteri Koch, 1939</taxonomicName>
Figs 1, 7A, 8, 9
</paragraph>
</subSubSection>
<subSubSection id="67FE8450EBDE1B0CB94AF512546BC19B" pageId="9" pageNumber="118" type="reference_group">
<paragraph id="D5E3B74CED5E8ABA1CC1404F416D0389" pageId="9" pageNumber="118">
<taxonomicName id="E01AC356A0ACFBD460EEAA6ADB401740" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene (Macromene) reitteri" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="reitteri" subGenus="Macromene">
<pageBreakToken id="D07A31D2780BE0E883F8EE41AF81DAA8" pageId="9" pageNumber="118" start="start">Domene</pageBreakToken>
(Macromene) reitteri
</taxonomicName>
Koch, 1939: 161
</paragraph>
</subSubSection>
<subSubSection id="17093C8C28E62284C62CF3550A7FEE55" pageId="9" pageNumber="118" type="type material examined">
<paragraph id="3491E346EB50DAA4790122A2AE269403" pageId="9" pageNumber="118">Type material examined.</paragraph>
<paragraph id="F168FF52819CAE6CB2FA4875C39C0D34" pageId="9" pageNumber="118">
Lectotype ♂, present designation: &quot;Tienmuschan, N. W China, Rtt. / Type /
<taxonomicName id="0502B8E47B8EB8BDA1280396C89C9673" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
Koch det. C. Koch / Holotype 1956 det. Kamp / Holotypus
<taxonomicName id="305AD69FD73429D6EA82A0CAE621AD49" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
Koch /
<taxonomicName id="1FDF4D5A8C83423A5EFF269EBF43393F" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
Koch V. L. Gusarov det. 1993 / Lectotypus ♂,
<taxonomicName id="602928A6075CC78B8F0B8C0744476B01" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
Koch, desig. B. Feldmann 2010&quot; (NHMB).
</paragraph>
<paragraph id="1E1225EACB67A4A3EF47B8DFFF02BF23" pageId="9" pageNumber="118">
Paralectotypes 3 ♀♀: &quot;Tienmuschan, N. W China, Rtt. / Cotype / Paratypus
<taxonomicName id="59C6A61D1BBB178E051801907970A8C8" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
Koch /
<taxonomicName id="AAA67DF52456E4DB3C2CD0690B1DEA7F" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
Koch V. L. Gusarov det. 1993&quot;; 1 ex. (abdomen missing): &quot;Tienmuschan, N. W China, Rtt. / Cotype&quot; (NHMB).
</paragraph>
</subSubSection>
<subSubSection id="AE806EC808FFC83CF1A0BFC090B34AE0" pageId="10" pageNumber="119" type="comment">
<paragraph id="8414FA91638A510C13A4B3F1145E6CC4" pageId="10" pageNumber="119">
<pageBreakToken id="72E3793FBA9A580EA87EEED327A4D8C0" pageId="10" pageNumber="119" start="start">Comment</pageBreakToken>
.
</paragraph>
<paragraph id="5E67335123A47DA1DAB88FC96E1BA038" pageId="10" pageNumber="119">
The original description of
<taxonomicName id="5B44DDC6A1EC4C11043555A4A7CB4968" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
is based on an unspecified number of syntypes from &quot;Tienmuschan (nordwestliches [sic] China) ex coll. E. Reitter&quot; (
<bibRefCitation id="D0335F1A905CB87B97BA38F76D8DA37B" author="Koch, C" journalOrPublisher="Entomologische Blaetter" pageId="24" pageNumber="133" pagination="156 - 172" title="ueber neue und wenig bekannte palaearktische Paederinae (Col. Staph.). III." volume="35" year="1939">Koch 1939</bibRefCitation>
). Five syntypes, one male, three females and one unsexed specimen, were located in the Koch collection at the Naturhistorisches Museum Basel. The male syntype is designated as the lectotype.
</paragraph>
</subSubSection>
<subSubSection id="C97CC6807F361A0780A5C23B301267C1" lastPageId="11" lastPageNumber="120" pageId="10" pageNumber="119" type="additional material examined">
<paragraph id="660FC9D7C6D60814EFE07B9E839EFEDF" pageId="10" pageNumber="119">Additional material examined</paragraph>
<paragraph id="75D0B8943A4FB8BA9179DC8DCBC6BFE8" lastPageId="11" lastPageNumber="120" pageId="10" pageNumber="119">
(87 ♂♂, 59 ♀♀). China: Zhejiang: 11 ♂♂, 7 ♀♀, Anji County, Longwang Shan,
<geoCoordinate id="D0AD651718BD4415AF96DF148B2A48C8" direction="north" orientation="latitude" precision="15" value="30.399721">30°23'59&quot;N</geoCoordinate>
<geoCoordinate id="7D872399C39BD8EBA5546B0B30A672F8" direction="east" orientation="longitude" precision="15" value="119.44056">119°26'26&quot;E</geoCoordinate>
, 1300-1450 m, 14.V.2013, leg. Hu (SNUC); 31 ♂♂, 14 ♀♀, Longwang Shan,
<geoCoordinate id="B08F9D93BA5E18C86DE23272EB659D3A" direction="north" orientation="latitude" precision="15" value="30.407778">30°24'28&quot;N</geoCoordinate>
<geoCoordinate id="9C5DA4C3FC33274E4D1799127C8E21B6" direction="east" orientation="longitude" precision="15" value="119.44361">119°26'37&quot;E</geoCoordinate>
, 1050-1200 m, 15.V.2013, leg. Li &amp; al. (SNUC); 15 ♂♂, 7 ♀♀, Longwang Shan, Qianmutian,
<geoCoordinate id="4B9E20572A4BF31BA50BBE3D7FCF6881" direction="north" orientation="latitude" precision="925" value="30.4">30°24'N</geoCoordinate>
<geoCoordinate id="D4BCF9BAA7F9B0953500486BD05523D4" direction="east" orientation="longitude" precision="925" value="119.433334">119°26'E</geoCoordinate>
, 1050-1250 m, 08.VI.2012, leg. Yin &amp; Hu (SNUC, cAss); 1 ♂, Longwang Shan,
<geoCoordinate id="1143163545CEDAC71E7C841E633F40D3" direction="north" orientation="latitude" precision="925" value="30.4">30°24'N</geoCoordinate>
<geoCoordinate id="2DF13A611C9A6751A5935326DC7DD9EA" direction="east" orientation="longitude" precision="925" value="119.433334">119°26'E</geoCoordinate>
, 1250-1450 m, 14.V.2013, leg. Chen &amp; Pan (SNUC); 3 ♂♂, 5 ♀♀, Longwang Shan, Dongguan, 1250 m, 26.V.2009, leg. Feng &amp; al. (SNUC); 3 ♂♂, 6 ♀♀, Longwang Shan, Qianmutian, 1300 m, 24.V.2009, leg. Feng &amp; al.(SNUC); 8 ♂♂, 10 ♀♀, Longwang Shan, 950-1200 m, 25.IV.2006, leg. He (SNUC); 2 ♂♂, 3 ♀♀, Longwang Shan, Qianmutian, 1300 m, 29.V.2009, leg. Feng &amp; al.(SNUC); 1 ♀, Longwang Shan, Qianmutian, 700-1325 m, 28.VII.2011, leg. Pan (SNUC); 1 ♂, 1 ♀, Tianmu Shan, 1200-1300 m, 25.-29.VII.2011, leg. Chen (SNUC); 2 ♂♂, 2 ♀♀, Tianmu Shan, 300 m, 17.V.2006, leg. Hu &amp; Tang (SNUC); 1
<pageBreakToken id="1A603F3C4A53B6415E8D71308FE23BBB" pageId="11" pageNumber="120" start="start"></pageBreakToken>
, Tianmu Shan, 1100 m, 24.VII.2011, leg. Hu &amp; Tang (SNUC); 1 ♂, Tianmu Shan, 1500 m, 15.VIII.2010, leg. Hu (SNUC); 1 ♂, Tianmu Shan, 300-400 m, 29.V.2010, leg. Wang (SNUC); 1 ♂, East Tianmu Shan, 1050-1150 m, 13. IV.2011, leg. Peng &amp; Zhu (SNUC); 2 ♂♂, Tianmu Shan, 13.VI.2009, leg. Song (SNUC); 2 ♂♂, Tianmu Shan, 1000 m, 2.V.2009, leg. Song (SNUC); 2 ♂♂, 1 ♀, Tianmu Shan, 1500 m, 15.VIII.2010, leg. Hu (SNUC); 2 ♂♂, 1 ♀, West Tianmu Shan N.R., path to peak of immortals (&quot;Blind Alley&quot;),
<geoCoordinate id="A880F4747D701A72E171A933632AB7BB" direction="north" orientation="latitude" precision="15" value="30.34278">30°20'34&quot;N</geoCoordinate>
,
<geoCoordinate id="9025EA5D1B93281CD20A92B97A5A2DA3" direction="east" orientation="longitude" precision="15" value="119.43083">119°25'51&quot;E</geoCoordinate>
, 1100-1200 m, primary mixed forest, litter moss, sifted, 15.VI.2007, leg. Wrase (cSch, cFel).
</paragraph>
</subSubSection>
<subSubSection id="50D3318D972BB19E3756ED542AA3F185" lastPageId="13" lastPageNumber="122" pageId="11" pageNumber="120" type="redescription">
<paragraph id="CA9C5F8FFD61A6BF07F4279F02ACCE6F" pageId="11" pageNumber="120">Redescription.</paragraph>
<paragraph id="318B84F4673D380DE5C24B3ADDE7C1DD" pageId="11" pageNumber="120">Measurements (in mm) and ratios: TL 5.78-8.62, FL 4.16-4.43, HL 1.07-1.17, HW 1.05-1.11, AnL 2.78-3.05, NW 0.37-0.41, PL 1.28-1.35, PW 1.02-1.06, EL 1.07-1.13, EW 1.12-1.20, TiL 1.39-1.44, TaL 0.89-0.96, AW 1.12-1.24, AL 1.07-1.17, HL/HW 1.02-1.05, HW/PW 1.03-1.05, HL/PL 0.84-0.87, NW/HW 0.35-0.37, PL/PW 1.25-1.28, EL/PL 0.84-0.89.</paragraph>
<paragraph id="2E8443B67902661F20D25622A36D7EEA" pageId="12" pageNumber="121">
<pageBreakToken id="A42B04B07F5DABE6A07DE008CC623276" pageId="12" pageNumber="121" start="start">Habitus</pageBreakToken>
as in Figs 7A, 8A, 8D. Body blackish brown; legs with dark brown profemora and protibiae, basal halves of metafemora light brown, distal halves gradually infuscate; antennae brown to light brown.
</paragraph>
<caption id="20C2995A794285EAEF8041E131A6E2D5" pageId="12" pageNumber="121">
<paragraph id="D1C5186B8AB38DA5CC57FECAECFE520D" pageId="12" pageNumber="121">
Figure 7. Habitus of
<taxonomicName id="CE38523EFD8855E9B2B86B3B12AB2FA8" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="genus">Domene</taxonomicName>
spp., A
<taxonomicName id="99C9E1393A0A6C1D2ECB82FE7F6197D6" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
B
<taxonomicName id="2B1E208AA3E0690B1A0BEE45E9A3BDDB" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene chenae" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="chenae">Domene chenae</taxonomicName>
. Scales: 1.0 mm.
</paragraph>
</caption>
<caption id="6FB8D29D4A7214EA5F7C9D338593B5DD" pageId="12" pageNumber="121">
<paragraph id="8537FE7167F9A2E54B2339F6257E5069" pageId="12" pageNumber="121">
Figure 8.
<taxonomicName id="F44C902CA23917AB2BD05FB3B321BAF5" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
. A lectotype B male sternite
<normalizedToken id="6BD316280F0F52A592AB6011556E562E" originalValue="VIIVIII">VII-VIII</normalizedToken>
C aedeagus of lectotype in lateral view D paralectotype. Scales: A, D 1.0 mm;
<normalizedToken id="0E095EE6E1340215B7A216083827BDDA" originalValue="BC">B-C</normalizedToken>
0.5 mm.
</paragraph>
</caption>
<paragraph id="AE7D7D25CBF15F27DAC38F56021AAB2D" pageId="12" pageNumber="121">
Head orbicular, broadest across eyes; punctation (Fig. 9A) moderately coarse, weakly umbilicate, and very dense, interstices forming very narrow ridges. All antennomeres longer than broad; antennomeres
<normalizedToken id="6B0045E7B0CC0B497907F7224C87CEB5" originalValue="IVX">IV-X</normalizedToken>
of equal length; antennomere I 1.6 times, II 1.1 times, III 1.3 times, XI 1.3 times as long as IV. Maxillary palpus very slender, preapical joint 2.7-3.2 times as long as broad.
</paragraph>
<caption id="5184F02C0869E4503D1CDF71246491FD" pageId="12" pageNumber="121">
<paragraph id="A618A8D71ACCFE7408DBDF77B7A692D5" pageId="12" pageNumber="121">
Figure 9.
<taxonomicName id="25E3F383BE94B1F566CB0E93B85829E8" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
. A median dorsal portion of head B median portion of pronotum C female tergite VIII D female sternite VIII E female tergites
<normalizedToken id="7D6D3C6F1E61A82D587C59CE1A3C0936" originalValue="IXX">IX-X</normalizedToken>
. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales:
<normalizedToken id="F114876584E8FC17C6455FA9FFE9B0F5" originalValue="AB">A-B</normalizedToken>
0.2 mm;
<normalizedToken id="8E3C41036A02327C4E4441D69730B2AC" originalValue="CJ">C-J</normalizedToken>
0.5 mm.
</paragraph>
</caption>
<paragraph id="5624BDCD74C4F06ECA92662C3DD9BC09" pageId="13" pageNumber="122">
<pageBreakToken id="FF51BEC3E1051F9410F8382448E769C2" pageId="13" pageNumber="122" start="start">Pronotum</pageBreakToken>
slightly narrower than head, widest in the middle; lateral margins convex in dorsal view; punctation (Fig. 9B) somewhat coarser than that of head; midline with rudiment of a fine glossy line.
</paragraph>
<paragraph id="BA2D3459B3E6EC8321D6576EA6DF13A6" pageId="13" pageNumber="122">
Elytra without distinct longitudinal ridges; suture weakly elevated; punctation very fine, dense and uniform; hind wings reduced. Protarsomeres
<normalizedToken id="082807B52504FA905DBD18A6CEB8E772" originalValue="IIV">I-IV</normalizedToken>
dilated in both sexes.
</paragraph>
<paragraph id="CDC5D8F4D98A54725B35E1D0488D1D36" pageId="13" pageNumber="122">
Abdomen with punctation fine and dense on tergites
<normalizedToken id="1A967E8AE7C2F52BCB3EBDFE0EAA52A3" originalValue="IIIVI">III-VI</normalizedToken>
, finer and somewhat sparser on tergite VIII, posterior margin of tergite VIII weakly convex in the middle (Fig. 9C); interstices with shallow microreticulation; posterior margin of tergite VII with palisade fringe.
</paragraph>
<paragraph id="BD9652F76C5811048B7E8AE98F446915" pageId="13" pageNumber="122">
Male. Sternites
<normalizedToken id="68683361BD73E2ABA3FFE1C417E7D8AC" originalValue="IIIVI">III-VI</normalizedToken>
unmodified; sternite VII (Figs 8B, 9F) distinctly transverse, with median impression of triangular shape posteriorly, this impression with numerous distinctly modified, short and stout black setae; posterior margin distinctly concave in the middle; sternite VIII (Figs 8B, 9G) transverse, with pronounced and symmetric impression posteriorly, this impression with distinctly modified short and stout black setae, posterior excision small and U-shaped; aedeagus as in Figs 8C, 9
<normalizedToken id="94FFA0E73B7D55F1567B60A75FB98971" originalValue="HJ">H-J</normalizedToken>
, ventral process stout and apically acute; dorsal fig with large and lamellate apical portion, and with short, thin basal portion; internal sac with membranous structures.
</paragraph>
<paragraph id="50BBE10E60974A91F3E197482D533CB3" pageId="13" pageNumber="122">Female. Posterior margin of sternite VIII (Fig. 9D) broadly convex; genital segments with weakly asymmetric large and moderately sclerotized structure (Fig. 9E).</paragraph>
</subSubSection>
<subSubSection id="FCCDF3539B07B083F0F10AD95BF9B89D" pageId="13" pageNumber="122" type="comparative notes">
<paragraph id="09758E3F1769A4C93E6E87A0ABC43253" pageId="13" pageNumber="122">Comparative notes.</paragraph>
<paragraph id="01B9947403FC675E0B4C99D1F46DFCB9" pageId="13" pageNumber="122">
The fine, dense and uniform punctation of the elytra, and the similar shape and chaetotaxy of the male sternite VII and sternite VIII suggest that
<taxonomicName id="47F4545AF6F16EDE52E56E31291F6C1F" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene reitteri" order="Coleoptera" pageId="13" pageNumber="122" phylum="Arthropoda" rank="species" species="reitteri">Domene reitteri</taxonomicName>
is most closely allied to
<taxonomicName id="8770ABEB97F1BF816CE70C77C5CAE4F9" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene chenae" order="Coleoptera" pageId="13" pageNumber="122" phylum="Arthropoda" rank="species" species="chenae">Domene chenae</taxonomicName>
. It is distinguished from
<taxonomicName id="D84B8879DACBA428166E134EF22624E2" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Domene chenae" order="Coleoptera" pageId="13" pageNumber="122" phylum="Arthropoda" rank="species" species="chenae">Domene chenae</taxonomicName>
by the finer punctation of the head and pronotum, the numerous distinctly modified, short and stout black setae on the male sternite VII, the stouter ventral process of the aedeagus and by the shape of the sclerotized structure in the female genital segments.
</paragraph>
</subSubSection>
<subSubSection id="E75A72C8125BC93E7191BEE9E3A7FE94" pageId="13" pageNumber="122" type="distribution and natural history">
<paragraph id="6F82180375DFB819FAFC15E329343DBF" pageId="13" pageNumber="122">Distribution and natural history.</paragraph>
<paragraph id="A08FFA995643390F38ECDA023E444F39" pageId="13" pageNumber="122">The distribution is confined to several localities in the Tianmu Shan range in the northwest of Zhejiang. The specimens were sifted from leaf litter in broad-leaved and primary mixed forests at altitudes of 300-1500 m.</paragraph>
</subSubSection>
</treatment>
</document>