644 lines
63 KiB
XML
644 lines
63 KiB
XML
<document ID-DOI="10.1590/S1679-62252010000200004" ID-ISSN="1982-0224" ID-Zenodo-Dep="5421513" approvalRequired="12" approvalRequired_for_textStreams="12" checkinTime="1630379354900" checkinUser="carolina" docAuthor="Bertaco, Vinicius A., Carvalho, Fernando R. & Jerep, Fernando C." docDate="2010" docId="D67F87BEFFCAFFF7FCE1FCEBFDBBFBDE" docLanguage="en" docName="NeotropIchthyol.2010.25.pdf" docOrigin="Neotropical Ichthyology 8 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252010000200004&lng=en&tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Astyanax goyanensis Bertaco & Carvalho & Jerep 2010, new combination" docType="treatment" docVersion="2" lastPageId="4" lastPageNumber="269" masterDocId="2A46FFC6FFCBFFF3FF89FFDFFF88FFD7" masterDocTitle="Astyanax goyanensis (Miranda-Ribeiro, 1944), new combination and Astyanax courensis, new species (Ostariophysi: Characiformes): two Characidae from the upper rio Tocantins basin, Central Brazil" masterLastPageNumber="275" masterPageNumber="265" pageId="1" pageNumber="266" updateTime="1630687045039" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Astyanax goyanensis (Miranda-Ribeiro, 1944), new combination and Astyanax courensis, new species (Ostariophysi: Characiformes): two Characidae from the upper rio Tocantins basin, Central Brazil</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bertaco, Vinicius A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Carvalho, Fernando R.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jerep, Fernando C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>Neotropical Ichthyology</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>8</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>265</mods:start>
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<mods:end>275</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252010000200004&lng=en&tlng=en</mods:url>
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||
</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1590/S1679-62252010000200004</mods:identifier>
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<mods:identifier type="ISSN">1982-0224</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5421513</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:D67F87BEFFCAFFF7FCE1FCEBFDBBFBDE" httpUri="http://treatment.plazi.org/id/D67F87BEFFCAFFF7FCE1FCEBFDBBFBDE" lastPageId="4" lastPageNumber="269" pageId="1" pageNumber="266">
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||
<subSubSection pageId="1" pageNumber="266" type="nomenclature">
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||
<paragraph blockId="1.[872,1388,820,903]" pageId="1" pageNumber="266">
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<heading centered="true" fontSize="10" level="2" pageId="1" pageNumber="266" reason="2">
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<emphasis bold="true" pageId="1" pageNumber="266">
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<taxonomicName authority="(Miranda-Ribeiro, 1944)" authorityName="Bertaco & Carvalho & Jerep" authorityYear="2010" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1944" box="[872,1383,820,839]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="goyanensis" status="new combination">
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<emphasis bold="true" box="[872,1100,820,839]" italics="true" pageId="1" pageNumber="266">Astyanax goyanensis</emphasis>
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(Miranda-Ribeiro, 1944)
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1039,1221,852,871]" pageId="1" pageNumber="266" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection box="[1081,1179,884,903]" pageId="1" pageNumber="266" type="description">
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||
<paragraph blockId="1.[872,1388,820,903]" box="[1081,1179,884,903]" pageId="1" pageNumber="266">
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<figureCitation box="[1081,1179,884,903]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[136,175,1153,1172]" captionStartId-1="3.[129,168,763,782]" captionTargetBox-0="[136,1459,174,1140]" captionTargetBox-1="[129,772,174,745]" captionTargetId-0="figure-512@2.[136,1459,174,1140]" captionTargetId-1="figure-872@3.[129,772,174,745]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Astyanax goyanensis. (a) MNRJ 4129, holotype, female, 89.2 mm SL; rio dos Couros, Chapada dos Veadeiros, Goiás State, Brazil (anal-fin incomplete); (b) UFRGS 11291, topotype, male, 56.7 mm SL; small stream tributary of rio dos Couros at Parque Portal da Chapada, tributary of rio Tocantinzinho, upper rio Tocantins basin, Alto Paraíso de Goiás, Goiás State, Brazil." captionText-1="Fig. 2. Astyanax goyanensis, UFRGS 11291, 59.2 mm SL. Scanning electronic microscopy images of the upper and lower jaws, left side, lateral view. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5421515" figureDoi-1="http://doi.org/10.5281/zenodo.5421517" httpUri-0="https://zenodo.org/record/5421515/files/figure.png" httpUri-1="https://zenodo.org/record/5421517/files/figure.png" pageId="1" pageNumber="266">
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<emphasis bold="true" box="[1081,1179,884,903]" pageId="1" pageNumber="266">Figs. 1-2</emphasis>
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</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="1" pageNumber="266" type="reference_group">
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<paragraph blockId="1.[809,1452,948,1127]" pageId="1" pageNumber="266">
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<treatmentCitationGroup pageId="1" pageNumber="266">
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<taxonomicName authority="Miranda-Ribeiro, 1944: 1 - 2" authorityName="Miranda-Ribeiro" authorityPageNumber="1 - 2" authorityYear="1944" box="[809,1356,948,967]" class="Actinopterygii" family="Characidae" genus="Astyanacinus" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="goyanensis">
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<emphasis box="[809,1070,948,967]" italics="true" pageId="1" pageNumber="266">Astyanacinus goyanensis</emphasis>
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Miranda-Ribeiro, 1944:1-2
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</taxonomicName>
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(original description, type-locality: “Rio do Couro, Veadeiros, estado de Goiaz”; picture of
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<typeStatus box="[1151,1250,1012,1031]" pageId="1" pageNumber="266">holotype</typeStatus>
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). -
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<treatmentCitation author="Fowler, H" box="[1274,1452,1012,1031]" page="34" pageId="1" pageNumber="266" year="1948">
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<bibRefCitation author="Fowler, H" box="[1274,1452,1012,1031]" pageId="1" pageNumber="266" pagination="1 - 204" refId="ref9730" refString="Fowler, H. W. 1948. Os peixes de agua doce do Brasil. Arquivos de Zoologia do Estado de Sao Paulo (1 ª entrega), 6: 1 - 204." type="journal article" year="1948">Fowler, 1948:34</bibRefCitation>
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||
</treatmentCitation>
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(distribution). -
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<treatmentCitation author="Lima, F & Malabarba, P & Buckup, J & Pezzi da Silva, R & Vari, A & Harold, R & Benine, O & Oyakawa, C & Pavanelli, N & Menezes, C & Lucena, M & Malabarba, Z & Lucena, R & Reis, F & Langeani, L & Casatti, V & Bertaco, C" box="[1001,1221,1044,1063]" page="108" pageId="1" pageNumber="266" year="2003">
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<bibRefCitation author="Lima, F & Malabarba, P & Buckup, J & Pezzi da Silva, R & Vari, A & Harold, R & Benine, O & Oyakawa, C & Pavanelli, N & Menezes, C & Lucena, M & Malabarba, Z & Lucena, R & Reis, F & Langeani, L & Casatti, V & Bertaco, C" box="[1001,1221,1044,1063]" pageId="1" pageNumber="266" pagination="106 - 169" refId="ref10029" refString="Lima, F. C. T., L. R. Malabarba, P. A. Buckup, J. F. Pezzi da Silva, R. P. Vari, A. Harold, R. Benine, O. T. Oyakawa, C. S. Pavanelli, N. A. Menezes, C. A. S. Lucena, M. C. S. L. Malabarba, Z. M. S. Lucena, R. E. Reis, F. Langeani, L. Casatti, V. A. Bertaco, C. Moreira & P. H. F. Lucinda. 2003. Genera incertae sedis in Characidae. Pp. 106 - 169. In: R. E. Reis, S. O. Kullander & C. J. Ferraris (Eds.). Check List of the Freshwater Fishes of South and Central America. Porto Alegre, Edipucrs, 729 p." type="book chapter" year="2003">
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Lima
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<emphasis box="[1062,1106,1044,1063]" italics="true" pageId="1" pageNumber="266">et al</emphasis>
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., 2003:108
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</bibRefCitation>
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||
</treatmentCitation>
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(maximum length and distribution). -
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<treatmentCitation author="Lima, F & Buckup, N & Menezes, C & Lucena, M & Malabarba, Z & Lucena, M" box="[1007,1243,1076,1095]" page="44" pageId="1" pageNumber="266" year="2007">
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<bibRefCitation author="Lima, F & Buckup, N & Menezes, C & Lucena, M & Malabarba, Z & Lucena, M" box="[1007,1243,1076,1095]" pageId="1" pageNumber="266" pagination="44 - 62" refId="ref10206" refString="Lima, F. C. T., P. A. Buckup, N. A. Menezes, C. A. S. Lucena, M. C. S. L. Malabarba, Z. M. S. Lucena, M. Toledo-Piza & A. Zanata. 2007. Familia Characidae: Generos incertae sedis. Pp. 44 - 62. In: P. A. Buckup, N. A. Menezes & M. S. Ghazzi (Eds.). Catalogo das especies de peixes de agua doce do Brasil. Rio de Janeiro, Museu Nacional, 195 p." type="book chapter" year="2007">
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Lima
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||
<emphasis box="[1077,1129,1076,1095]" italics="true" pageId="1" pageNumber="266">et al</emphasis>
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., 2007:44
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</bibRefCitation>
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||
</treatmentCitation>
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(type-locality and distribution).
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="1" pageNumber="266" type="materials_examined">
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||
<paragraph blockId="1.[809,1452,1181,1428]" pageId="1" pageNumber="266">
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||
<emphasis bold="true" box="[809,1010,1181,1198]" pageId="1" pageNumber="266">Material examined.</emphasis>
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||
<materialsCitation collectionCode="MNRJ" collectorName="dos Couros & Chapada dos Veadeiros" country="Brazil" location="Goias State" pageId="1" pageNumber="266" specimenCode="MNRJ 4129" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
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||
<specimenCode box="[1017,1138,1181,1198]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="1" pageNumber="266">MNRJ 4129</specimenCode>
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,
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||
<typeStatus box="[1149,1235,1181,1198]" pageId="1" pageNumber="266">holotype</typeStatus>
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||
, female,
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||
<quantity box="[1324,1410,1181,1198]" metricMagnitude="-2" metricUnit="m" metricValue="8.22" pageId="1" pageNumber="266" unit="mm" value="82.2">82.2 mm</quantity>
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SL, rio
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<collectorName box="[840,948,1209,1226]" pageId="1" pageNumber="266">dos Couros</collectorName>
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,
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<collectorName box="[956,1178,1209,1226]" pageId="1" pageNumber="266">Chapada dos Veadeiros</collectorName>
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||
,
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||
<collectingRegion box="[1186,1295,1209,1226]" country="Brazil" name="Goias" pageId="1" pageNumber="266">Goiás State</collectingRegion>
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,
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<collectingCountry box="[1303,1364,1209,1226]" name="Brazil" pageId="1" pageNumber="266">Brazil</collectingCountry>
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, 1929
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</materialsCitation>
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,
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<materialsCitation collectionCode="R" collectorName="Pfrimmer" country="Brazil" pageId="1" pageNumber="266" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
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<collectionCode box="[1430,1449,1209,1226]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="266">R</collectionCode>
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||
.
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||
<collectorName box="[809,901,1238,1255]" pageId="1" pageNumber="266">Pfrimmer</collectorName>
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</materialsCitation>
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||
.
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||
<materialsCitation collectionCode="MNRJ" country="Brazil" pageId="1" pageNumber="266" specimenCode="MNRJ 2843" specimenCount="2" stateProvince="Goias" typeStatus="paratype">
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||
<specimenCode box="[908,1024,1238,1255]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="1" pageNumber="266">MNRJ 2843</specimenCode>
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,
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||
<typeStatus box="[1032,1124,1238,1255]" pageId="1" pageNumber="266">paratypes</typeStatus>
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, 5 of 7,
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<quantity box="[1194,1325,1238,1255]" metricMagnitude="-2" metricUnit="m" metricValue="7.209999999999999" metricValueMax="8.48" metricValueMin="5.9399999999999995" pageId="1" pageNumber="266" unit="mm" value="72.1" valueMax="84.8" valueMin="59.4">59.4-84.8 mm</quantity>
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SL, collected with the
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<typeStatus box="[897,984,1267,1284]" pageId="1" pageNumber="266">holotype</typeStatus>
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</materialsCitation>
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.
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<materialsCitation accessionNumber="MCP 44402" box="[995,1448,1267,1284]" collectionCode="MCP" country="Brazil" pageId="1" pageNumber="266" specimenCount="1" stateProvince="Goias" typeStatus="paratype">
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||
<accessionNumber box="[995,1115,1267,1284]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP44402" pageId="1" pageNumber="266">
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||
<collectionCode box="[995,1046,1267,1284]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="1" pageNumber="266">MCP</collectionCode>
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||
44402
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||
</accessionNumber>
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, topotypes, 20,
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<quantity box="[1272,1409,1267,1284]" metricMagnitude="-2" metricUnit="m" metricValue="4.54" metricValueMax="6.39" metricValueMin="2.69" pageId="1" pageNumber="266" unit="mm" value="45.4" valueMax="63.9" valueMin="26.9">26.9-63.9 mm</quantity>
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||
SL
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||
</materialsCitation>
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||
,
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||
<materialsCitation collectingDate="2009-09-10" collectionCode="UFRGS" collectorName="dos Couros" country="Brazil" county="Alto Paraiso" elevation="1135" latitude="-14.170279" location="Parque Portal da Chapada" longLatPrecision="21" longitude="-47.609722" municipality="rio Tocantinzinho" pageId="1" pageNumber="266" specimenCode="UFRGS 11291, 80, 5" specimenCount="1" stateProvince="Tocantins" typeStatus="paratype">
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||
<specimenCode box="[809,961,1296,1313]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">UFRGS 11291</specimenCode>
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||
,
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||
<specimenCode box="[974,1001,1296,1313]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">80</specimenCode>
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||
,
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||
<specimenCode box="[1014,1026,1296,1313]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">5</specimenCode>
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||
c&s, topotypes,
|
||
<quantity box="[1203,1345,1296,1313]" metricMagnitude="-2" metricUnit="m" metricValue="4.92" metricValueMax="7.99" metricValueMin="1.85" pageId="1" pageNumber="266" unit="mm" value="49.2" valueMax="79.9" valueMin="18.5">18.5-79.9 mm</quantity>
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||
SL, small tributary of rio
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||
<collectorName box="[955,1063,1325,1342]" pageId="1" pageNumber="266">dos Couros</collectorName>
|
||
at
|
||
<location LSID="urn:lsid:plazi:treatment:D67F87BEFFCAFFF7FCE1FCEBFDBBFBDE:5B096073FFCAFFF2FBCDFAF2FAB6FAE9" box="[1092,1342,1325,1342]" country="Brazil" county="Alto Paraiso" latitude="-14.170279" longLatPrecision="21" longitude="-47.609722" municipality="rio Tocantinzinho" name="Parque Portal da Chapada" pageId="1" pageNumber="266" stateProvince="Tocantins">Parque Portal da Chapada</location>
|
||
, affluent of
|
||
<collectingMunicipality box="[809,973,1353,1370]" pageId="1" pageNumber="266">rio Tocantinzinho</collectingMunicipality>
|
||
, upper rio
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||
<collectingRegion box="[1067,1157,1353,1370]" country="Brazil" name="Tocantins" pageId="1" pageNumber="266">Tocantins</collectingRegion>
|
||
basin,
|
||
<quantity box="[1220,1288,1353,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.135" pageId="1" pageNumber="266" unit="m" value="1135.0">
|
||
<elevation box="[1220,1288,1353,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.135" pageId="1" pageNumber="266" unit="m" value="1135.0">1135 m</elevation>
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||
</quantity>
|
||
a.s.l,
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||
<collectingCounty box="[1337,1452,1353,1370]" pageId="1" pageNumber="266">Alto Paraíso</collectingCounty>
|
||
de
|
||
<collectingRegion box="[836,894,1382,1399]" country="Brazil" name="Goias" pageId="1" pageNumber="266">Goiás</collectingRegion>
|
||
,
|
||
<collectingRegion box="[902,1011,1382,1399]" country="Brazil" name="Goias" pageId="1" pageNumber="266">Goiás State</collectingRegion>
|
||
,
|
||
<collectingCountry box="[1019,1079,1382,1399]" name="Brazil" pageId="1" pageNumber="266">Brazil</collectingCountry>
|
||
,
|
||
<geoCoordinate box="[1087,1198,1382,1399]" degrees="14" direction="south" minutes="10" orientation="latitude" pageId="1" pageNumber="266" precision="15" seconds="13" value="-14.170279">14º10’13”S</geoCoordinate>
|
||
<geoCoordinate box="[1202,1322,1382,1399]" degrees="47" direction="west" minutes="36" orientation="longitude" pageId="1" pageNumber="266" precision="15" seconds="35" value="-47.609722">47º36’35”W</geoCoordinate>
|
||
,
|
||
<date box="[1330,1448,1382,1399]" pageId="1" pageNumber="266" value="2009-09-10">
|
||
<collectingDate box="[1330,1448,1382,1399]" pageId="1" pageNumber="266" value="2009-09-10">10 Sep 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation box="[809,961,1411,1428]" collectionCode="V" collectorName="Bertaco" country="Brazil" pageId="1" pageNumber="266" specimenCount="1" stateProvince="Tocantins" typeStatus="paratype">
|
||
<collectionCode box="[809,826,1411,1428]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="266" type="Museum">V</collectionCode>
|
||
. A.
|
||
<collectorName box="[862,939,1411,1428]" pageId="1" pageNumber="266">Bertaco</collectorName>
|
||
, F
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation box="[970,1210,1411,1428]" collectionCode="R" collectorName="Carvalho & G. Frainer" country="Brazil" pageId="1" pageNumber="266" specimenCount="1" stateProvince="Tocantins" typeStatus="paratype">
|
||
<collectionCode box="[970,989,1411,1428]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="266">R</collectionCode>
|
||
.
|
||
<collectorName box="[997,1085,1411,1428]" pageId="1" pageNumber="266">Carvalho</collectorName>
|
||
&
|
||
<collectorName box="[1114,1210,1411,1428]" pageId="1" pageNumber="266">G. Frainer</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="1.[809,1453,1472,1863]" pageId="1" pageNumber="266">
|
||
<emphasis bold="true" box="[809,1015,1472,1489]" pageId="1" pageNumber="266">Additional material.</emphasis>
|
||
<materialsCitation accessionNumber="MCP 23937" collectingDate="1996-08-04" collectingDateMax="2008-05-25" collectingDateMin="1996-08-04" collectionCode="MCP, R, T" collectorName="dos Couros & da Guabiroba & Koch, P & K. M. Grosser. & Rita Maria & F. C. Jerep & P. Carvalho. & Alto Paraiso & Parque Nacional da Chapada dos Veadeiros & das Cobras & P. Carvalho" country="Brazil" county="Goias State" latitude="-14.160833" location="Alto Paraiso" longLatPrecision="21" longitude="-47.631943" municipality="Capao da Guabiroba" pageId="1" pageNumber="266" specimenCode="6, UFRGS 9905, 19, UFRGS 9935, 31, 2, UFRGS 9960, 12" specimenCount="1" stateProvince="Tocantins">
|
||
<collectingCounty box="[1020,1130,1472,1489]" pageId="1" pageNumber="266">Goiás State</collectingCounty>
|
||
,
|
||
<collectingCountry box="[1139,1200,1472,1489]" name="Brazil" pageId="1" pageNumber="266">Brazil</collectingCountry>
|
||
, upper rio
|
||
<collectingRegion box="[1298,1391,1472,1489]" country="Brazil" name="Tocantins" pageId="1" pageNumber="266">Tocantins</collectingRegion>
|
||
basin, rio
|
||
<collectorName box="[844,955,1501,1518]" pageId="1" pageNumber="266">dos Couros</collectorName>
|
||
basin.
|
||
<accessionNumber box="[1029,1151,1501,1518]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP23937" pageId="1" pageNumber="266">
|
||
<collectionCode box="[1029,1080,1501,1518]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="1" pageNumber="266">MCP</collectionCode>
|
||
23937
|
||
</accessionNumber>
|
||
,
|
||
<specimenCode box="[1162,1176,1501,1518]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="1" pageNumber="266">6</specimenCode>
|
||
,
|
||
<quantity box="[1188,1328,1501,1518]" metricMagnitude="-2" metricUnit="m" metricValue="4.18" metricValueMax="6.15" metricValueMin="2.21" pageId="1" pageNumber="266" unit="mm" value="41.8" valueMax="61.5" valueMin="22.1">22.1-61.5 mm</quantity>
|
||
SL, córrego
|
||
<collectingMunicipality box="[809,1012,1529,1546]" pageId="1" pageNumber="266">
|
||
Capão
|
||
<collectorName box="[879,1012,1529,1546]" pageId="1" pageNumber="266">da Guabiroba</collectorName>
|
||
</collectingMunicipality>
|
||
tributary of rio
|
||
<collectorName box="[1174,1288,1529,1546]" pageId="1" pageNumber="266">dos Couros</collectorName>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:D67F87BEFFCAFFF7FCE1FCEBFDBBFBDE:5B096073FFCAFFF2FA9BFA26FA05F9DD" box="[1298,1421,1529,1546]" country="Brazil" county="Goias State" latitude="-14.160833" longLatPrecision="21" longitude="-47.631943" municipality="Capao da Guabiroba" name="Alto Paraiso" pageId="1" pageNumber="266" stateProvince="Tocantins">Alto Paraíso</location>
|
||
de
|
||
<collectingRegion box="[809,868,1558,1575]" country="Brazil" name="Goias" pageId="1" pageNumber="266">Goiás</collectingRegion>
|
||
,
|
||
<geoCoordinate box="[876,989,1558,1575]" degrees="14" direction="south" minutes="07" orientation="latitude" pageId="1" pageNumber="266" precision="15" seconds="43" value="-14.128612">14º07’43”S</geoCoordinate>
|
||
<geoCoordinate box="[993,1115,1558,1575]" degrees="47" direction="west" minutes="38" orientation="longitude" pageId="1" pageNumber="266" precision="15" seconds="27" value="-47.640835">47º38’27”W</geoCoordinate>
|
||
,
|
||
<date box="[1124,1236,1558,1575]" pageId="1" pageNumber="266" value="1996-08-04">
|
||
<collectingDate box="[1124,1236,1558,1575]" pageId="1" pageNumber="266" value="1996-08-04">4 Aug 1996</collectingDate>
|
||
</date>
|
||
, W.
|
||
<collectionCode box="[1275,1294,1558,1575]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="266">R</collectionCode>
|
||
.
|
||
<collectorName box="[1302,1379,1558,1575]" pageId="1" pageNumber="266">Koch, P</collectorName>
|
||
. Milani &
|
||
<collectorName box="[832,974,1587,1604]" pageId="1" pageNumber="266">K. M. Grosser.</collectorName>
|
||
<specimenCode box="[977,1111,1587,1604]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">UFRGS 9905</specimenCode>
|
||
,
|
||
<specimenCode box="[1119,1145,1587,1604]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">19</specimenCode>
|
||
,
|
||
<quantity box="[1154,1289,1587,1604]" metricMagnitude="-2" metricUnit="m" metricValue="4.17" metricValueMax="5.56" metricValueMin="2.78" pageId="1" pageNumber="266" unit="mm" value="41.7" valueMax="55.6" valueMin="27.8">27.8-55.6 mm</quantity>
|
||
SL, córrego
|
||
<collectorName pageId="1" pageNumber="266">Rita Maria</collectorName>
|
||
<locationDeviation box="[870,1175,1616,1633]" firstLocation="Cavalcante" pageId="1" pageNumber="266" secondLocation="Minacu">between Cavalcante and Minaçú</locationDeviation>
|
||
, rio
|
||
<collectingRegion box="[1213,1277,1616,1633]" country="Brazil" name="Parana" pageId="1" pageNumber="266">Paranã</collectingRegion>
|
||
basin,
|
||
<location LSID="urn:lsid:plazi:treatment:D67F87BEFFCAFFF7FCE1FCEBFDBBFBDE:5B096073FFCAFFF2FAB4F98FFA20F9B6" box="[1341,1448,1616,1633]" country="Brazil" county="Goias State" latitude="-14.160833" longLatPrecision="21" longitude="-47.631943" municipality="Capao da Guabiroba" name="Cavalcante" pageId="1" pageNumber="266" stateProvince="Tocantins">Cavalcante</location>
|
||
,
|
||
<geoCoordinate box="[809,920,1645,1662]" degrees="13" direction="south" minutes="45" orientation="latitude" pageId="1" pageNumber="266" precision="15" seconds="16" value="-13.754444">13º45’16”S</geoCoordinate>
|
||
<geoCoordinate box="[924,1044,1645,1662]" degrees="47" direction="west" minutes="27" orientation="longitude" pageId="1" pageNumber="266" precision="15" seconds="18" value="-47.455">47º27’18”W</geoCoordinate>
|
||
,
|
||
<date box="[1053,1179,1645,1662]" pageId="1" pageNumber="266" value="2008-05-25">
|
||
<collectingDate box="[1053,1179,1645,1662]" pageId="1" pageNumber="266" value="2008-05-25">25 May 2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1188,1286,1645,1662]" pageId="1" pageNumber="266">F. C. Jerep</collectorName>
|
||
&
|
||
<collectionCode box="[1314,1330,1645,1662]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="266">T</collectionCode>
|
||
.
|
||
<collectorName box="[1337,1453,1645,1662]" pageId="1" pageNumber="266">P. Carvalho.</collectorName>
|
||
<specimenCode box="[809,945,1673,1690]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">UFRGS 9935</specimenCode>
|
||
,
|
||
<specimenCode box="[955,981,1673,1690]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">31</specimenCode>
|
||
,
|
||
<specimenCode box="[992,1004,1673,1690]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">2</specimenCode>
|
||
c&s,
|
||
<quantity box="[1062,1200,1673,1690]" metricMagnitude="-2" metricUnit="m" metricValue="3.3650000000000007" metricValueMax="5.0200000000000005" metricValueMin="1.7100000000000002" pageId="1" pageNumber="266" unit="mm" value="33.65" valueMax="50.2" valueMin="17.1">17.1-50.2 mm</quantity>
|
||
SL, córrego on road GO- 239 between
|
||
<collectorName box="[932,1049,1702,1719]" pageId="1" pageNumber="266">Alto Paraíso</collectorName>
|
||
de
|
||
<collectingRegion box="[1080,1136,1702,1719]" country="Brazil" name="Goias" pageId="1" pageNumber="266">Goiás</collectingRegion>
|
||
and
|
||
<collectorName pageId="1" pageNumber="266">
|
||
<location LSID="urn:lsid:plazi:treatment:D67F87BEFFCAFFF7FCE1FCEBFDBBFBDE:5B096073FFCAFFF2FB15F979FA24F960" box="[1180,1452,1702,1719]" country="Brazil" county="Goias State" latitude="-14.160833" longLatPrecision="21" longitude="-47.631943" municipality="Capao da Guabiroba" name="Parque Nacional da Chapada" pageId="1" pageNumber="266" stateProvince="Tocantins">Parque Nacional da Chapada</location>
|
||
dos Veadeiros
|
||
</collectorName>
|
||
,
|
||
<collectorName box="[954,1075,1731,1748]" pageId="1" pageNumber="266">Alto Paraíso</collectorName>
|
||
de
|
||
<collectingRegion box="[1108,1167,1731,1748]" country="Brazil" name="Goias" pageId="1" pageNumber="266">Goiás</collectingRegion>
|
||
,
|
||
<geoCoordinate box="[1177,1290,1731,1748]" degrees="14" direction="south" minutes="08" orientation="latitude" pageId="1" pageNumber="266" precision="15" seconds="24" value="-14.14">14º08’24”S</geoCoordinate>
|
||
<geoCoordinate box="[1295,1418,1731,1748]" degrees="47" direction="west" minutes="39" orientation="longitude" pageId="1" pageNumber="266" precision="15" seconds="36" value="-47.66">47º39’36”W</geoCoordinate>
|
||
,
|
||
<date pageId="1" pageNumber="266" value="2008-05-25">
|
||
<collectingDate pageId="1" pageNumber="266" value="2008-05-25">25 May 2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[923,1028,1760,1777]" pageId="1" pageNumber="266">F. C. Jerep</collectorName>
|
||
&
|
||
<collectionCode box="[1062,1078,1760,1777]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="266">T</collectionCode>
|
||
.
|
||
<collectorName box="[1088,1209,1760,1777]" pageId="1" pageNumber="266">P. Carvalho.</collectorName>
|
||
<specimenCode box="[1215,1352,1760,1777]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">UFRGS 9960</specimenCode>
|
||
,
|
||
<specimenCode box="[1363,1389,1760,1777]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="266">12</specimenCode>
|
||
,
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.26" metricValueMax="8.24" metricValueMin="2.28" pageId="1" pageNumber="266" unit="mm" value="52.6" valueMax="82.4" valueMin="22.8">22.8- 82.4 mm</quantity>
|
||
SL, rio
|
||
<collectorName box="[983,1093,1789,1806]" pageId="1" pageNumber="266">das Cobras</collectorName>
|
||
between
|
||
<collectorName box="[1189,1312,1789,1806]" pageId="1" pageNumber="266">Alto Paraíso</collectorName>
|
||
de
|
||
<collectingRegion box="[1351,1408,1789,1806]" country="Brazil" name="Goias" pageId="1" pageNumber="266">Goiás</collectingRegion>
|
||
and
|
||
<collectorName box="[809,1210,1817,1834]" pageId="1" pageNumber="266">
|
||
<location LSID="urn:lsid:plazi:treatment:D67F87BEFFCAFFF7FCE1FCEBFDBBFBDE:5B096073FFCAFFF2FCA0F8C6FBBBF8FD" box="[809,1075,1817,1834]" country="Brazil" county="Goias State" latitude="-14.160833" longLatPrecision="21" longitude="-47.631943" municipality="Capao da Guabiroba" name="Parque Nacional da Chapada" pageId="1" pageNumber="266" stateProvince="Tocantins">Parque Nacional da Chapada</location>
|
||
dos Veadeiros
|
||
</collectorName>
|
||
,
|
||
<geoCoordinate box="[1217,1326,1817,1834]" degrees="14" direction="south" minutes="09" orientation="latitude" pageId="1" pageNumber="266" precision="15" seconds="39" value="-14.160833">14º09’39”S</geoCoordinate>
|
||
<geoCoordinate box="[1330,1447,1817,1834]" degrees="47" direction="west" minutes="37" orientation="longitude" pageId="1" pageNumber="266" precision="15" seconds="55" value="-47.631943">47º37’55”W</geoCoordinate>
|
||
,
|
||
<date box="[809,943,1846,1863]" pageId="1" pageNumber="266" value="2008-05-25">
|
||
<collectingDate box="[809,943,1846,1863]" pageId="1" pageNumber="266" value="2008-05-25">25 May 2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[953,1058,1846,1863]" pageId="1" pageNumber="266">F. C. Jerep</collectorName>
|
||
&
|
||
<collectionCode box="[1091,1107,1846,1863]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="266">T</collectionCode>
|
||
.
|
||
<collectorName box="[1117,1234,1846,1863]" pageId="1" pageNumber="266">P. Carvalho</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266" type="diagnosis">
|
||
<paragraph blockId="1.[809,1452,1909,1962]" lastBlockId="2.[816,1459,1301,1608]" lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266">
|
||
<emphasis bold="true" box="[809,941,1909,1928]" pageId="1" pageNumber="266">Diagnosis.</emphasis>
|
||
<taxonomicName authorityName="Bertaco & Carvalho & Jerep" authorityYear="2010" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1944" box="[956,1216,1909,1928]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="goyanensis">
|
||
<emphasis box="[956,1216,1909,1928]" italics="true" pageId="1" pageNumber="266">Astyanax goyanensis</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName baseAuthorityName="Jenyns" baseAuthorityYear="1842" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="scabripinnis">
|
||
<emphasis box="[1427,1443,1909,1928]" italics="true" pageId="1" pageNumber="266">A</emphasis>
|
||
.
|
||
<emphasis box="[809,943,1943,1962]" italics="true" pageId="1" pageNumber="266">scabripinnis</emphasis>
|
||
</taxonomicName>
|
||
species complex and is distinguished from all species of this complex by the total number of pelvic-fin rays (7
|
||
<emphasis box="[162,183,1333,1352]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 8), except from
|
||
<taxonomicName authority="Bertaco & Malabarba" authorityName="Bertaco & Malabarba" authorityYear="2001" box="[350,775,1333,1352]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="brachypterygium">
|
||
<emphasis box="[350,366,1333,1352]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[376,551,1333,1352]" italics="true" pageId="2" pageNumber="267">brachypterygium</emphasis>
|
||
Bertaco & Malabarba
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Almiron, Azpelicueta & Casciotta" authorityName="Almiron, Azpelicueta & Casciotta" authorityYear="2002" box="[136,585,1365,1384]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="ita">
|
||
<emphasis box="[136,152,1365,1384]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[169,197,1365,1384]" italics="true" pageId="2" pageNumber="267">ita</emphasis>
|
||
Almirón, Azpelicueta & Casciotta
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="(Hensel)" baseAuthorityName="Hensel" baseAuthorityYear="1870" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis box="[648,664,1365,1384]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[681,778,1365,1384]" italics="true" pageId="2" pageNumber="267">obscurus</emphasis>
|
||
(Hensel)
|
||
</taxonomicName>
|
||
, Bertaco & Lucena (2010). It differs from
|
||
<taxonomicName authorityName="Bertaco & Malabarba" authorityYear="2001" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="brachypterygium">
|
||
<emphasis box="[755,771,1397,1416]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[136,318,1429,1448]" italics="true" pageId="2" pageNumber="267">brachypterygium</emphasis>
|
||
</taxonomicName>
|
||
by caudal peduncle length (14.1-18.5% SL, mean = 16.1
|
||
<emphasis box="[279,301,1461,1480]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 11.9-14.4, mean = 13.0), by the number of caudal-peduncle scales (14
|
||
<emphasis box="[420,442,1493,1512]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 16), and by the number of cusps of the premaxillary inner series and the larger dentary teeth (4-5
|
||
<emphasis box="[187,209,1557,1576]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 5-7); from
|
||
<taxonomicName authorityName="Almiron, Azpelicueta & Casciotta" authorityYear="2002" box="[339,396,1557,1576]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="ita">
|
||
<emphasis box="[339,355,1557,1576]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[368,396,1557,1576]" italics="true" pageId="2" pageNumber="267">ita</emphasis>
|
||
</taxonomicName>
|
||
by the number of branched anal-fin rays (13-16
|
||
<emphasis box="[262,284,1589,1608]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 20-24), number of humeral spots (2
|
||
<emphasis box="[672,694,1589,1608]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 1), and by the orbital diameter (20.2-33.2% HL, mean = 27.4
|
||
<emphasis box="[690,712,1621,1640]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 38.6- 42.0, mean = 40.2); and from
|
||
<taxonomicName baseAuthorityName="Hensel" baseAuthorityYear="1870" box="[459,586,1653,1672]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis box="[459,475,1653,1672]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[489,586,1653,1672]" italics="true" pageId="2" pageNumber="267">obscurus</emphasis>
|
||
</taxonomicName>
|
||
by the number of maxillary teeth (2-5
|
||
<emphasis box="[344,366,1685,1704]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 1), number of lateral line scales (35-37, mean = 36
|
||
<emphasis box="[245,266,1717,1736]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 37-39, mean = 38), and by the number of branched anal-fin rays (13-16, mean = 14.7
|
||
<emphasis box="[512,534,1749,1768]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 16-21, mean = 18.3). Furthermore, the lower number of the branched anal-fin rays in
|
||
<taxonomicName authorityName="Bertaco & Carvalho & Jerep" authorityYear="2010" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1944" box="[168,324,1813,1832]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="goyanensis">
|
||
<emphasis box="[168,184,1813,1832]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[202,324,1813,1832]" italics="true" pageId="2" pageNumber="267">goyanensis</emphasis>
|
||
</taxonomicName>
|
||
distinguish it from its congeners of the species complex, except from
|
||
<taxonomicName authorityName="Bertaco & Malabarba" authorityYear="2001" box="[502,734,1845,1864]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="brachypterygium">
|
||
<emphasis box="[502,518,1845,1864]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[538,734,1845,1864]" italics="true" pageId="2" pageNumber="267">brachypterygium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Bertaco & Malabarba" authorityName="Bertaco & Malabarba" authorityYear="2001" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="cremnobates">
|
||
<emphasis box="[755,771,1845,1864]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[136,274,1877,1896]" italics="true" pageId="2" pageNumber="267">cremnobates</emphasis>
|
||
Bertaco & Malabarba
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Cope)" baseAuthorityName="Cope" baseAuthorityYear="1894" box="[538,742,1877,1896]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="laticeps">
|
||
<emphasis box="[538,554,1877,1896]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[571,655,1877,1896]" italics="true" pageId="2" pageNumber="267">laticeps</emphasis>
|
||
(Cope)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Eigenmann" authorityName="Eigenmann" authorityYear="1908" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis box="[756,772,1877,1896]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[136,271,1909,1928]" italics="true" pageId="2" pageNumber="267">intermedius</emphasis>
|
||
Eigenmann
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Steindachner)" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[430,739,1909,1928]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="jenynsii">
|
||
<emphasis box="[430,446,1909,1928]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[466,555,1909,1928]" italics="true" pageId="2" pageNumber="267">jenynsii</emphasis>
|
||
(Steindachner)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Alcaraz, Pavanelli & Bertaco" authorityName="Alcaraz, Pavanelli & Bertaco" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="jordanensis">
|
||
<emphasis box="[755,771,1909,1928]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[136,261,1941,1960]" italics="true" pageId="2" pageNumber="267">jordanensis</emphasis>
|
||
Alcaraz, Pavanelli & Bertaco
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Bertaco & Lucena" authorityName="Bertaco & Lucena" authorityYear="2006" box="[596,1010,1301,1960]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="microschemos">
|
||
<emphasis box="[596,612,1941,1960]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[625,778,1941,1960]" italics="true" pageId="2" pageNumber="267">microschemos</emphasis>
|
||
Bertaco & Lucena
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Haluch & Abilhoa" authorityName="Haluch & Abilhoa" box="[1019,1297,1301,1320]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="totae">
|
||
<emphasis box="[1019,1035,1301,1320]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[1046,1099,1301,1320]" italics="true" pageId="2" pageNumber="267">totae</emphasis>
|
||
Haluch & Abilhoa
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="Abilhoa & Duboc. It" authorityName="Abilhoa & Duboc. It" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="varzeae">
|
||
<emphasis box="[1349,1365,1301,1320]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[1377,1459,1301,1320]" italics="true" pageId="2" pageNumber="267">varzeae</emphasis>
|
||
Abilhoa & Duboc. It
|
||
</taxonomicName>
|
||
differs from most these species by the number of humeral spot (2
|
||
<emphasis box="[1099,1121,1365,1384]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 1), except
|
||
<taxonomicName authorityName="Bertaco & Malabarba" authorityYear="2001" box="[1242,1452,1365,1384]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="brachypterygium">
|
||
<emphasis box="[1242,1258,1365,1384]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[1270,1452,1365,1384]" italics="true" pageId="2" pageNumber="267">brachypterygium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Bertaco & Malabarba" authorityYear="2001" box="[816,984,1397,1416]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="cremnobates">
|
||
<emphasis box="[816,832,1397,1416]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[846,984,1397,1416]" italics="true" pageId="2" pageNumber="267">cremnobates</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName box="[1044,1157,1397,1416]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="varzeae">
|
||
<emphasis box="[1044,1060,1397,1416]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[1073,1157,1397,1416]" italics="true" pageId="2" pageNumber="267">varzeae</emphasis>
|
||
</taxonomicName>
|
||
. From these three species it differs by the number of cups in the premaxillary and dentary teeth (5-7
|
||
<emphasis box="[926,948,1461,1480]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 3-5), and by the caudal-peduncle length (14.1- 18.5% SL, mean = 16.1
|
||
<emphasis box="[1065,1087,1493,1512]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 11.9-14.4, mean = 13.0, 11.0-14.9, mean = 13.1, 10.5-13.9, mean = 12.4, respectively), and additionally from
|
||
<taxonomicName box="[998,1105,1557,1576]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="varzeae">
|
||
<emphasis box="[998,1014,1557,1576]" italics="true" pageId="2" pageNumber="267">A</emphasis>
|
||
.
|
||
<emphasis box="[1025,1105,1557,1576]" italics="true" pageId="2" pageNumber="267">varzeae</emphasis>
|
||
</taxonomicName>
|
||
by the number of lateral line scales (35-37, mean = 36
|
||
<emphasis box="[1006,1027,1589,1608]" italics="true" pageId="2" pageNumber="267">vs</emphasis>
|
||
. 37-42, mean = 39).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5421515" ID-Zenodo-Dep="5421515" httpUri="https://zenodo.org/record/5421515/files/figure.png" pageId="2" pageNumber="267" startId="2.[136,175,1153,1172]" targetBox="[136,1459,174,1140]" targetPageId="2">
|
||
<paragraph blockId="2.[136,1457,1153,1236]" pageId="2" pageNumber="267">
|
||
<emphasis bold="true" box="[136,205,1153,1172]" pageId="2" pageNumber="267">Fig. 1.</emphasis>
|
||
<taxonomicName authorityName="Bertaco & Carvalho & Jerep" authorityYear="2010" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1944" box="[211,433,1153,1172]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="goyanensis">
|
||
<emphasis box="[211,433,1153,1172]" italics="true" pageId="2" pageNumber="267">Astyanax goyanensis</emphasis>
|
||
</taxonomicName>
|
||
. (
|
||
<emphasis bold="true" box="[455,468,1153,1172]" pageId="2" pageNumber="267">a</emphasis>
|
||
) MNRJ 4129, holotype, female, 89.2 mm SL; rio dos Couros, Chapada dos Veadeiros, Goiás State, Brazil (anal-fin incomplete); (
|
||
<emphasis bold="true" box="[513,528,1185,1204]" pageId="2" pageNumber="267">b</emphasis>
|
||
) UFRGS 11291, topotype, male, 56.7 mm SL; small stream tributary of rio dos Couros at Parque Portal da Chapada, tributary of rio Tocantinzinho, upper rio Tocantins basin, Alto Paraíso de Goiás, Goiás State, Brazil.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="4" lastPageNumber="269" pageId="2" pageNumber="267" type="description">
|
||
<paragraph blockId="2.[816,1460,1653,1960]" lastBlockId="3.[129,772,885,1960]" lastPageId="3" lastPageNumber="268" pageId="2" pageNumber="267">
|
||
<emphasis bold="true" box="[816,951,1653,1672]" pageId="2" pageNumber="267">Description.</emphasis>
|
||
Morphometric data summarized in
|
||
<tableCitation box="[1316,1392,1653,1672]" captionStart="Table 1" captionStartId="7.[129,192,182,201]" captionText="Table 1. Morphometric data of holotype (H), paratypes, and non-types of Astyanax goyanensis, and holotype and paratypes of Astyanax courensis, new species, from the rio dos Couros basin, upper rio Tocantins basin, Alto Paraíso de Goiás, Goiás State, Brazil. The range of paratypes and non-types of A. goyanensis not includes the holotype, but of A. courensis includes." pageId="2" pageNumber="267">Table 1</tableCitation>
|
||
. Body compressed and elongate; greatest body depth usually immediately anterior to dorsal-fin origin. Dorsal profile of head between vertical through posterior nostril and tip of supraoccipital spine straight or slightly convex. Profile of body convex from tip of supraocciptal spine to base of last dorsalfin ray, and slightly concave from that point to adipose-fin origin. Ventral profile of head convex. Ventral profile of body slightly convex from pectoral-fin insertion to anal-fin origin. Body profile along anal-fin base posterodorsally slanted. Caudal peduncle elongate and nearly straight to slightly concave along both dorsal and ventral margins.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5421517" ID-Zenodo-Dep="5421517" httpUri="https://zenodo.org/record/5421517/files/figure.png" pageId="3" pageNumber="268" startId="3.[129,168,763,782]" targetBox="[129,772,174,745]" targetPageId="3">
|
||
<paragraph blockId="3.[129,772,763,846]" pageId="3" pageNumber="268">
|
||
<emphasis bold="true" box="[129,201,763,782]" pageId="3" pageNumber="268">Fig. 2.</emphasis>
|
||
<taxonomicName authority=", UFRGS" authorityName="UFRGS" box="[209,537,763,782]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="goyanensis">
|
||
<emphasis box="[209,436,763,782]" italics="true" pageId="3" pageNumber="268">Astyanax goyanensis</emphasis>
|
||
, UFRGS
|
||
</taxonomicName>
|
||
11291, 59.2 mm SL. Scanning electronic microscopy images of the upper and lower jaws, left side, lateral view. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="3.[129,772,885,1960]" pageId="3" pageNumber="268">Snout rounded from margin of upper lip to vertical through anterior nostrils. Head relatively small (one third of SL). Mouth terminal, lower jaw projecting slightly more than upper jaw. Maxilla extending posteriorly to vertical through anterior half of orbit, slightly curved, and aligned at angle of approximately 45 degrees relative to longitudinal body axis. Maxilla slightly widened anteroposteriorly.</paragraph>
|
||
<paragraph blockId="3.[129,772,885,1960]" pageId="3" pageNumber="268">
|
||
Premaxilla with two tooth rows: outer row with three to five tri- to pentacuspid teeth, central cusp longest. Five teeth in inner premaxillary row; teeth gradually decreasing in length from first towards fourth teeth with last tooth distinctly smaller: teeth with five to seven cusps with central cusp twice as long and broad as other cusps. Maxilla with two to five (five only in the
|
||
<typeStatus box="[202,296,1365,1384]" pageId="3" pageNumber="268">holotype</typeStatus>
|
||
and
|
||
<specimenCount box="[355,498,1365,1384]" pageId="3" pageNumber="268" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
) teeth with three to five cusps; central cusp longest. Three to four anteriormost dentary teeth larger, with five or seven cusps, followed by one medium-sized tooth with three to five cusps, and five to seven teeth with one to three cusps. Central cusp in all teeth two to three times as long and broad as other cusps. All cusp tips slightly curved posteriorly towards oral cavity (
|
||
<figureCitation box="[682,748,1557,1576]" captionStart="Fig" captionStartId="3.[129,168,763,782]" captionTargetBox="[129,772,174,745]" captionTargetId="figure-872@3.[129,772,174,745]" captionTargetPageId="3" captionText="Fig. 2. Astyanax goyanensis, UFRGS 11291, 59.2 mm SL. Scanning electronic microscopy images of the upper and lower jaws, left side, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5421517" httpUri="https://zenodo.org/record/5421517/files/figure.png" pageId="3" pageNumber="268">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="3.[129,772,885,1960]" lastBlockId="3.[809,1452,182,969]" pageId="3" pageNumber="268">
|
||
Dorsal-fin rays ii,9 (
|
||
<typeStatus box="[391,487,1589,1608]" pageId="3" pageNumber="268">holotype</typeStatus>
|
||
with ii,8;
|
||
<specimenCount box="[605,771,1589,1608]" pageId="3" pageNumber="268" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
|
||
with i,9, seven topotypes with i,9, and ii,
|
||
<quantity box="[616,662,1621,1640]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="3" pageNumber="268" unit="in" value="8.0">8 in</quantity>
|
||
two c&s specimens, n = 32); first unbranched ray approximately half length of second ray. Distal margin of dorsal fin nearly straight or slightly convex. Dorsal-fin origin anterior at middle of SL. Adipose-fin origin approximately at vertical through insertion of base of last anal-fin ray. Anal-fin rays ii-v, 13-16 (
|
||
<typeStatus box="[676,771,1781,1800]" pageId="3" pageNumber="268">holotype</typeStatus>
|
||
with anal incomplete, 13, mean = 14.7, n = 32). First unbranched ray apparent only in c&s specimens. Anal-fin profile nearly straight in males, and smoothly concave in females. Anal-fin origin posterior to vertical through base of last dorsal-fin ray. Anal-fin rays of males bearing one pair of small bony hooks along posterolateral border of each segment of lepidotrichia, usually along last unbranched ray to eighth anterior branched rays. Hooks more numerous along second through fifth branched ray, and usually located along posteriormost branch and distal 1/2 to 2/3 of each ray.
|
||
</paragraph>
|
||
<paragraph blockId="3.[809,1452,182,969]" pageId="3" pageNumber="268">Pectoral-fin rays i,10-13 (mean = 12.0, n = 32). Tip of pectoral fin reaching pelvic-fin insertion in males and not reaching that insertion in females. Pelvic-fin rays i,6 (n = 32). Pelvic-fin origin slightly anterior to vertical through dorsal-fin origin. Tip of pelvic fin reaching anal-fin origin in males, falling short of that point in females. Caudal fin forked and unscaled, lobes similar in size, with 19 principal rays. Dorsal procurrent rays 11-14, and ventral procurrent rays 10-12 (n = 7).</paragraph>
|
||
<paragraph blockId="3.[809,1452,182,969]" pageId="3" pageNumber="268">Lateral line complete with 35-37 scales (mean = 36.0, n = 32). Scale rows between dorsal-fin origin and lateral line 6-7 (mean = 6.1, n = 32); 4-5 scale rows between lateral line and pelvic-fin origin (mean = 4.5, n = 32). Predorsal scales 11-14 (mean = 12.5, n = 32) usually arranged in regular series. Scale rows around caudal peduncle 14 (n = 32). Axillary scale on pelvic-fin origin extends posteriorly covering 2-3 scales. Scale sheath along anal-fin base 5-12 scales, in single series, covering base of anteriormost rays (n = 30).</paragraph>
|
||
<paragraph blockId="3.[809,1452,182,969]" pageId="3" pageNumber="268">Precaudal vertebrae 17-18; caudal vertebrae 17-18; total vertebrae 35 (n = 7). Supraneurals 5-7 (n = 7). Upper branch of gill-rakers 7-8, lower branch 11-12 (n = 7).</paragraph>
|
||
<paragraph blockId="3.[809,1452,1014,1609]" pageId="3" pageNumber="268">
|
||
<emphasis bold="true" box="[809,994,1014,1033]" pageId="3" pageNumber="268">Color in alcohol.</emphasis>
|
||
General body color yellowish-brown. Dark chromatophores scattered on lateral portion of head, more densely concentrated on snout and anterior border of eye. Lateral portion of body with scales bordered with dark pigment, forming slightly reticulate pattern.Body with black, pigmented, midlateral stripe extending from humeral region to base of middle caudal fin; faint dark pigmentation present on middle caudal-fin rays. Midlateral body stripe expanded dorsally and ventrally to caudalfin base, forming small caudal spot.Two humeral spots: anterior spot conspicuous, vertically elongate, with superior portion wider, located over second to fourth vertical series of scales, extending over 2 to 3 horizontal series of scales above lateral line; and inferior portion of spot narrow, and extending over 1 to 2 horizontal series of scales below lateral line; posterior humeral spot diffuse, not ventrally surpassing lateral-line, extending over 2 to 3 horizontal series and 2 to 3 vertical series of scales. Region between spots pale. Fins with dispersed dark chromatophores (
|
||
<figureCitation box="[817,890,1558,1577]" captionStart="Fig" captionStartId="2.[136,175,1153,1172]" captionTargetBox="[136,1459,174,1140]" captionTargetId="figure-512@2.[136,1459,174,1140]" captionTargetPageId="2" captionText="Fig. 1. Astyanax goyanensis. (a) MNRJ 4129, holotype, female, 89.2 mm SL; rio dos Couros, Chapada dos Veadeiros, Goiás State, Brazil (anal-fin incomplete); (b) UFRGS 11291, topotype, male, 56.7 mm SL; small stream tributary of rio dos Couros at Parque Portal da Chapada, tributary of rio Tocantinzinho, upper rio Tocantins basin, Alto Paraíso de Goiás, Goiás State, Brazil." figureDoi="http://doi.org/10.5281/zenodo.5421515" httpUri="https://zenodo.org/record/5421515/files/figure.png" pageId="3" pageNumber="268">Fig. 1b</figureCitation>
|
||
). Specimens retained orange and yellowish pigment on all fins after several days fixation in formalin.
|
||
</paragraph>
|
||
<paragraph blockId="3.[809,1452,1654,1801]" pageId="3" pageNumber="268">
|
||
<emphasis bold="true" box="[809,940,1654,1673]" pageId="3" pageNumber="268">Color in life.</emphasis>
|
||
The specimens immediately after capture presented overall golden body coloration. Fins yellowish-orange to red. Humeral spots faint black, and midlateral stripe and caudalpeduncle spot less conspicuous than after preservation. Caudalfin middle rays black (
|
||
<figureCitation box="[1029,1102,1782,1801]" captionStart="Fig" captionStartId="2.[136,175,1153,1172]" captionTargetBox="[136,1459,174,1140]" captionTargetId="figure-512@2.[136,1459,174,1140]" captionTargetPageId="2" captionText="Fig. 1. Astyanax goyanensis. (a) MNRJ 4129, holotype, female, 89.2 mm SL; rio dos Couros, Chapada dos Veadeiros, Goiás State, Brazil (anal-fin incomplete); (b) UFRGS 11291, topotype, male, 56.7 mm SL; small stream tributary of rio dos Couros at Parque Portal da Chapada, tributary of rio Tocantinzinho, upper rio Tocantins basin, Alto Paraíso de Goiás, Goiás State, Brazil." figureDoi="http://doi.org/10.5281/zenodo.5421515" httpUri="https://zenodo.org/record/5421515/files/figure.png" pageId="3" pageNumber="268">Fig. 1b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="3.[809,1453,1846,1961]" pageId="3" pageNumber="268">
|
||
<emphasis bold="true" box="[809,1030,1846,1865]" pageId="3" pageNumber="268">Sexual dimorphism.</emphasis>
|
||
Males of
|
||
<taxonomicName box="[1132,1348,1846,1865]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="goyanensis">
|
||
<emphasis box="[1132,1348,1846,1865]" italics="true" pageId="3" pageNumber="268">Astyanax goyanensis</emphasis>
|
||
</taxonomicName>
|
||
are easily recognized by the presence of small bony hooks on the anal-fin rays (see Description).Also, males and females differ by anal-fin shape, which is nearly straight in males and concave in females.
|
||
</paragraph>
|
||
<paragraph blockId="4.[136,779,182,233]" pageId="4" pageNumber="269">Gill glands (Burns & Weitzman, 1996) were not found macroscopically on first gill arch in either males or females.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="269" type="distribution">
|
||
<paragraph blockId="4.[136,779,278,393]" pageId="4" pageNumber="269">
|
||
<emphasis bold="true" box="[136,282,278,297]" pageId="4" pageNumber="269">Distribution.</emphasis>
|
||
<taxonomicName box="[287,508,278,297]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="269" phylum="Chordata" rank="species" species="goyanensis">
|
||
<emphasis box="[287,508,278,297]" italics="true" pageId="4" pageNumber="269">Astyanax goyanensis</emphasis>
|
||
</taxonomicName>
|
||
is known from tributaries of the rio dos Couros, affluent of rio Tocantinzinho, and rio
|
||
<collectingRegion box="[136,214,342,361]" country="Brazil" name="Parana" pageId="4" pageNumber="269">Paranã</collectingRegion>
|
||
basin, upper rio
|
||
<collectingRegion box="[425,536,342,361]" country="Brazil" name="Tocantins" pageId="4" pageNumber="269">Tocantins</collectingRegion>
|
||
basin, Chapada dos Veadeiros,
|
||
<collectingRegion box="[250,313,374,393]" country="Brazil" name="Goias" pageId="4" pageNumber="269">Goiás</collectingRegion>
|
||
,
|
||
<collectingCountry box="[321,385,374,393]" name="Brazil" pageId="4" pageNumber="269">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation box="[397,459,374,393]" captionStart="Fig" captionStartId="4.[136,175,1877,1896]" captionTargetBox="[185,1398,1135,1847]" captionTargetId="figure-560@4.[166,1411,1129,1863]" captionTargetPageId="4" captionText="Fig. 3. Distribution of Astyanax goyanensis (circles) and A. courensis, new species (square), on upper rio Tocantins, Alto Paraíso de Goiás, Goiás State, Brazil. A, rio Tocantinzinho; B, rio Tocantins; C, rio Paranã; DF, Distrito Federal. Symbols can represent more than one lot or locality. Base map prepared by Marilyn Weitzman." figureDoi="http://doi.org/10.5281/zenodo.5421519" httpUri="https://zenodo.org/record/5421519/files/figure.png" pageId="4" pageNumber="269">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[136,779,438,1033]" pageId="4" pageNumber="269">
|
||
<emphasis bold="true" box="[136,313,438,457]" pageId="4" pageNumber="269">Ecological notes.</emphasis>
|
||
The collection locality is around
|
||
<quantity box="[638,714,438,457]" metricMagnitude="3" metricUnit="m" metricValue="1.135" pageId="4" pageNumber="269" unit="m" value="1135.0">1135 m</quantity>
|
||
above sea level. The rio dos Couros in that point has moderate riparian vegetation and it is typically between 10 and
|
||
<quantity box="[588,639,502,521]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="269" unit="m" value="15.0">15 m</quantity>
|
||
wide, and
|
||
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="269" unit="m" value="1.5">1.5 m</quantity>
|
||
deep. Although presenting some rapids, the water is clear. The substrate is constituted by stones, rocks, and some stretches with mud. A natural
|
||
<quantity box="[475,533,598,617]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="269" unit="m" value="10.0">10 m</quantity>
|
||
high waterfall named Cachoeira São Bento is located downriver.All topotypes were collected on the stretch downstream from the waterfall in a small stream draining directly into the river (about
|
||
<quantity box="[658,723,694,713]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="269" unit="m" value="100.0">100 m</quantity>
|
||
from the channel river). The stream is about
|
||
<quantity box="[524,581,726,745]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="269" unit="m" value="0.8">0.8 m</quantity>
|
||
deep and
|
||
<quantity box="[680,718,726,745]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="269" unit="m" value="6.0">6 m</quantity>
|
||
wide, has transparent water with moderate vegetation covering the channel (trees and shrubs), and bottom with small stones, submerged logs and roots (
|
||
<figureCitation box="[415,492,822,841]" captionStart="Fig" captionStartId="5.[129,168,835,854]" captionTargetBox="[129,1452,174,817]" captionTargetId="figure-404@5.[129,1452,174,817]" captionTargetPageId="5" captionText="Fig. 4. (a) Rio dos Couros downstream from the Cachoeira São Bento, type-locality of Astyanax courensis, new species; (b) small stream tributary of rio dos Couros at Parque Portal da Chapada, locality of collection of Astyanax goyanensis." figureDoi="http://doi.org/10.5281/zenodo.5421521" httpUri="https://zenodo.org/record/5421521/files/figure.png" pageId="4" pageNumber="269">Fig. 4b</figureCitation>
|
||
). No other fish species was collected with
|
||
<taxonomicName box="[288,432,854,873]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="269" phylum="Chordata" rank="species" species="goyanensis">
|
||
<emphasis box="[288,304,854,873]" italics="true" pageId="4" pageNumber="269">A</emphasis>
|
||
.
|
||
<emphasis box="[315,432,854,873]" italics="true" pageId="4" pageNumber="269">goyanensis</emphasis>
|
||
</taxonomicName>
|
||
in the stream. The rio dos Couros has several waterfalls, some with about
|
||
<quantity box="[621,695,886,905]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="269" unit="m" value="100.0">100 m</quantity>
|
||
height (Cachoeira de
|
||
<collectingCountry box="[286,410,918,937]" name="Saint Vincent and the Grenadines" pageId="4" pageNumber="269">São Vicente</collectingCountry>
|
||
and Catarata dos Couros). Stomach contents of
|
||
<specimenCount box="[289,474,950,969]" pageId="4" pageNumber="269" type="generic">five specimens</specimenCount>
|
||
contained fragments of allochthonous insect, fish scales, and vegetal organic matter (leaf parts). The stomachs were not full.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |