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<document ID-DOI="10.1670/20-094" ID-Zenodo-Dep="7877128" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1682707425169" checkinUser="diego" docAuthor="Ching, Ann Hui, Tay, Tricia, Brown, Bryan, Mohareb, Amir M, Sethi, Aisha &amp; Annamalai, Aniyizhai" docDate="2023" docId="039D49574E2EB05BA9A4FA3ACA4021CA" docLanguage="en" docName="JourHerpetol.55.2.105-111.pdf" docOrigin="Journal of Herpetology 7 (7)" docSource="http://dx.doi.org/10.1016/j.jmh.2023.100183" docStyle="DocumentStyle:55072EE3B1C90D1F2334E9D3FE399900.1:JourHerpetol.2019-.journal_article.1cover" docStyleId="55072EE3B1C90D1F2334E9D3FE399900" docStyleName="JourHerpetol.2019-.journal_article.1cover" docStyleVersion="1" docTitle="Hemidactylus gramineus Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai 2023, sp. nov." docType="treatment" docVersion="5" lastPageNumber="111" masterDocId="FFA4312F4E2CB05EAA55FF8CCF13261D" masterDocTitle="Dermatologic conditions of adult refugees following resettlement in the United States, 2015 to 2018." masterLastPageNumber="111" masterPageNumber="105" pageNumber="105" updateTime="1683049242774" updateUser="juliana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Dermatologic conditions of adult refugees following resettlement in the United States, 2015 to 2018.</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ching, Ann Hui</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tay, Tricia</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brown, Bryan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mohareb, Amir M</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sethi, Aisha</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Annamalai, Aniyizhai</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Journal of Herpetology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-01-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>7</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>105</mods:start>
<mods:end>111</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1016/j.jmh.2023.100183</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1670/20-094</mods:identifier>
<mods:identifier type="Zenodo-Dep">7877128</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:039D49574E2EB05BA9A4FA3ACA4021CA" httpUri="http://treatment.plazi.org/id/039D49574E2EB05BA9A4FA3ACA4021CA" lastPageId="5" pageId="2">
<subSubSection box="[1009,1342,1462,1484]" pageId="2" type="nomenclature">
<paragraph blockId="2.[1009,1342,1462,1484]" box="[1009,1342,1462,1484]" pageId="2">
<heading box="[1009,1342,1462,1484]" centered="true" fontSize="9" level="2" pageId="2" reason="2">
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[1009,1251,1462,1484]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[1009,1251,1462,1484]" italics="true" pageId="2">Hemidactylus gramineus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1259,1342,1463,1484]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" type="description">
<paragraph blockId="2.[896,1454,1492,1572]" box="[1068,1281,1492,1514]" pageId="2">
<figureCitation box="[1068,1166,1492,1514]" captionStart="FIG" captionStartId="2.[146,158,1019,1038]" captionTargetBox="[124,790,159,992]" captionTargetId="figure-819@2.[124,790,159,992]" captionTargetPageId="2" captionText="FIG. 1. Holotype (UTEP 22208) and paratype (UTEP 22209) of Hemidactylus gramineus sp. nov. Photos by Luis M. P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.7877130" httpUri="https://zenodo.org/record/7877130/files/figure.png" pageId="2">Figures 1</figureCitation>
,
<figureCitation box="[1180,1194,1492,1513]" captionStart="FIG" captionStartId="3.[146,174,1169,1188]" captionTargetBox="[124,790,159,1142]" captionTargetId="figure-612@3.[124,790,159,1142]" captionTargetPageId="3" captionText="FIG. 2. Photos of live Hemidactylus gramineus sp. nov. (Top: holotype UTEP 22208; bottom: paratype UTEP 22209). Photos by Eli Greenbaum." figureDoi="http://doi.org/10.5281/zenodo.7886504" httpUri="https://zenodo.org/record/7886504/files/figure.png" pageId="2">2</figureCitation>
;
<tableCitation box="[1206,1281,1492,1514]" captionStart="TABLE" captionStartId="3.[864,876,160,179]" captionTargetBox="[124,790,159,1142]" captionTargetId="figure-612@3.[124,790,159,1142]" captionTargetPageId="3" captionText="TABLE 1. Measurements and scale counts of the holotype and paratype of Hemidactylus gramineus sp. nov. Abbreviations are the same as those described in the Materials and Methods." pageId="2">Table 1</tableCitation>
</paragraph>
<paragraph blockId="2.[896,1454,1492,1572]" pageId="2">
<uri pageId="2">
urn:lsid:zoobank.org:act:
<uuid pageId="2">8C27856D-9BAD-4880-930F-2503794734C3</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="2" type="reference_group">
<paragraph blockId="2.[840,1362,1585,1634]" box="[840,1362,1585,1607]" pageId="2">
<treatmentCitationGroup box="[840,1362,1585,1607]" pageId="2">
<treatmentCitation author="CERIACO, L. M. P., I. &amp; AGARWAL, M. P. &amp; MARQUES, AND &amp; A. M. BAUER" box="[840,1362,1585,1607]" page=")" pageId="2" year="2020">
<taxonomicName authority=": Ceriaco et al. (2020 a: 14)" authorityName="Ceriaco" authorityPageNumber="14" authorityYear="2020" box="[840,1362,1585,1607]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="nzingae">
<emphasis box="[840,973,1585,1606]" italics="true" pageId="2">Hemidactylus</emphasis>
cf.
<emphasis box="[1013,1088,1586,1607]" italics="true" pageId="2">nzingae</emphasis>
:
<bibRefCitation author="CERIACO, L. M. P., I. &amp; AGARWAL, M. P. &amp; MARQUES, AND &amp; A. M. BAUER" box="[1102,1362,1585,1607]" pageId="2" pagination="1 - 71" refId="ref7198" refString="CERIACO, L. M. P., I. AGARWAL, M. P. MARQUES, AND, A. M. BAUER. 2020 a. A review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two new species. Zootaxa 4746: 1 - 71." type="journal article" year="2020">Ceríaco et al. (2020a: 14)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="2.[840,1362,1585,1634]" box="[840,1285,1612,1634]" pageId="2">
<treatmentCitationGroup box="[840,1285,1612,1634]" pageId="2">
<taxonomicName box="[840,1013,1612,1633]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="undetermined">
<emphasis box="[840,973,1612,1633]" italics="true" pageId="2">Hemidactylus</emphasis>
sp.
</taxonomicName>
:
<treatmentCitation author="CERIACO, L. M. P., I. &amp; AGARWAL, M. P. &amp; MARQUES, AND &amp; A. M. BAUER" box="[1025,1285,1612,1634]" page="19" pageId="2" year="2020">
<bibRefCitation author="CERIACO, L. M. P., I. &amp; AGARWAL, M. P. &amp; MARQUES, AND &amp; A. M. BAUER" box="[1025,1285,1612,1634]" pageId="2" pagination="1 - 71" refId="ref7198" refString="CERIACO, L. M. P., I. AGARWAL, M. P. MARQUES, AND, A. M. BAUER. 2020 a. A review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two new species. Zootaxa 4746: 1 - 71." type="journal article" year="2020">Ceríaco et al. (2020a: 19)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="2" type="materials_examined">
<paragraph blockId="2.[840,1510,1662,2007]" pageId="2">
<materialsCitation collectingDate="2013-06-07" collectionCode="UTEP" collectorName="Chifundera Kusamba &amp; Wandege M. Muninga" country="Democratic Republic of the Congo" county="Kinshasa Province" elevation="548" location="Lumene River" municipality="Bombo-Lumene Reserve" pageId="2" specimenCode="UTEP 22208" specimenCount="1" typeStatus="holotype">
<emphasis box="[864,949,1662,1683]" italics="true" pageId="2">
<typeStatus box="[864,949,1662,1683]" pageId="2">Holotype</typeStatus>
</emphasis>
.—
<specimenCode box="[977,1110,1663,1684]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="2" type="Museum">UTEP 22208</specimenCode>
(field number ELI 1645;
<figureCitation box="[1394,1466,1662,1684]" captionStart="FIG" captionStartId="2.[146,158,1019,1038]" captionTargetBox="[124,790,159,992]" captionTargetId="figure-819@2.[124,790,159,992]" captionTargetPageId="2" captionText="FIG. 1. Holotype (UTEP 22208) and paratype (UTEP 22209) of Hemidactylus gramineus sp. nov. Photos by Luis M. P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.7877130" httpUri="https://zenodo.org/record/7877130/files/figure.png" pageId="2">Figs. 1</figureCitation>
,
<figureCitation box="[1484,1497,1663,1685]" captionStart="FIG" captionStartId="3.[146,174,1169,1188]" captionTargetBox="[124,790,159,1142]" captionTargetId="figure-612@3.[124,790,159,1142]" captionTargetPageId="3" captionText="FIG. 2. Photos of live Hemidactylus gramineus sp. nov. (Top: holotype UTEP 22208; bottom: paratype UTEP 22209). Photos by Eli Greenbaum." figureDoi="http://doi.org/10.5281/zenodo.7886504" httpUri="https://zenodo.org/record/7886504/files/figure.png" pageId="2">2</figureCitation>
), adult male collected near bank of
<location LSID="urn:lsid:plazi:treatment:039D49574E2EB05BA9A4FA3ACA4021CA:8EEBAE9A4E2EB05CAEF1F910CA2420AF" box="[1188,1335,1692,1714]" country="Democratic Republic of the Congo" county="Kinshasa Province" municipality="Bombo-Lumene Reserve" name="Lumene River" pageId="2">Lumene River</location>
,
<collectingMunicipality pageId="2">Bombo-Lumene Reserve</collectingMunicipality>
(-4.42027
<emphasis bold="true" box="[1031,1039,1721,1743]" italics="true" pageId="2">°</emphasis>
, 16.04687
<emphasis bold="true" box="[1140,1148,1721,1743]" italics="true" pageId="2">°</emphasis>
,
<quantity box="[1162,1229,1721,1743]" metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" pageId="2" unit="m" value="548.0">
<elevation box="[1162,1229,1721,1743]" metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" pageId="2" unit="m" value="548.0">548 m</elevation>
</quantity>
),
<collectingCounty box="[1248,1441,1721,1743]" pageId="2">Kinshasa Province</collectingCounty>
,
<collectingCountry box="[1453,1506,1721,1742]" name="Democratic Republic of the Congo" pageId="2">DRC</collectingCountry>
, collected by
<collectorName box="[969,1191,1750,1772]" pageId="2">Chifundera Kusamba</collectorName>
and
<collectorName box="[1244,1476,1750,1772]" pageId="2">Wandege M. Muninga</collectorName>
on
<date box="[840,963,1780,1801]" pageId="2" value="2013-06-07">
<collectingDate box="[840,963,1780,1801]" pageId="2" value="2013-06-07">7 June 2013</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="2.[840,1510,1662,2007]" pageId="2">
<materialsCitation collectingDate="2013-06-07" collectionCode="UTEP" collectorName="Chifundera Kusamba; Wandege M. Muninga; Mwenebatu M. Aristote &amp; Eli Greenbaum" country="Democratic Republic of the Congo" elevation="555" location="Bombo-Lumene Reserve" municipality="Kinshasa Province" pageId="2" specimenCode="UTEP 22209" specimenCount="1" typeStatus="paratype">
<emphasis box="[864,947,1810,1831]" italics="true" pageId="2">
<typeStatus box="[864,947,1810,1831]" pageId="2">Paratype</typeStatus>
</emphasis>
.—
<specimenCode box="[975,1108,1809,1830]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="2" type="Museum">UTEP 22209</specimenCode>
(field number ELI 1674), adult male collected in
<location LSID="urn:lsid:plazi:treatment:039D49574E2EB05BA9A4FA3ACA4021CA:8EEBAE9A4E2EB05CA99FF8A2CBD92159" box="[970,1226,1838,1860]" country="Democratic Republic of the Congo" municipality="Kinshasa Province" name="Bombo-Lumene Reserve" pageId="2">Bombo-Lumene Reserve</location>
(-4.41843
<emphasis bold="true" box="[1338,1346,1838,1860]" italics="true" pageId="2">°</emphasis>
, 16.04420
<emphasis bold="true" box="[1450,1458,1838,1860]" italics="true" pageId="2">°</emphasis>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="2" unit="m" value="555.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="2" unit="m" value="555.0">555 m</elevation>
</quantity>
),
<collectingMunicipality box="[890,1091,1867,1889]" pageId="2">Kinshasa Province</collectingMunicipality>
,
<collectingCountry box="[1110,1506,1867,1889]" name="Democratic Republic of the Congo" pageId="2">Democratic Republic of the Congo</collectingCountry>
, collected by
<collectorName box="[991,1229,1897,1919]" pageId="2">Chifundera Kusamba</collectorName>
,
<collectorName box="[1249,1505,1897,1919]" pageId="2">Wandege M. Muninga</collectorName>
,
<collectorName box="[840,1091,1926,1948]" pageId="2">Mwenebatu M. Aristote</collectorName>
, and
<collectorName box="[1150,1307,1926,1948]" pageId="2">Eli Greenbaum</collectorName>
on
<date box="[1349,1472,1926,1948]" pageId="2" value="2013-06-07">
<collectingDate box="[1349,1472,1926,1948]" pageId="2" value="2013-06-07">7 June 2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" pageId="2" type="diagnosis">
<paragraph blockId="2.[840,1510,1662,2007]" lastBlockId="3.[123,793,1257,2007]" lastPageId="3" pageId="2">
<emphasis box="[864,958,1956,1977]" italics="true" pageId="2">Diagnosis</emphasis>
.—A small-sized
<taxonomicName authorityName="Goldfuss" authorityYear="1820" box="[1145,1275,1955,1976]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="genus">
<emphasis box="[1145,1275,1955,1976]" italics="true" pageId="2">Hemidactylus</emphasis>
</taxonomicName>
, maximum
<collectionCode box="[1412,1456,1956,1977]" pageId="2">SVL</collectionCode>
<quantity metricMagnitude="-2" metricUnit="m" metricValue="3.95" pageId="2" unit="mm" value="39.5">39.5 mm</quantity>
, with a short, blunt snout (
<figureCitation box="[1221,1288,1985,2007]" captionStart="FIG" captionStartId="2.[146,158,1019,1038]" captionTargetBox="[124,790,159,992]" captionTargetId="figure-819@2.[124,790,159,992]" captionTargetPageId="2" captionText="FIG. 1. Holotype (UTEP 22208) and paratype (UTEP 22209) of Hemidactylus gramineus sp. nov. Photos by Luis M. P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.7877130" httpUri="https://zenodo.org/record/7877130/files/figure.png" pageId="2">Fig. 1</figureCitation>
). Dorsal pholidosis heterogeneous, with 1214 irregularly arranged longitudinal rows of subtrihedral, striated, strongly keeled tubercles at midbody. Two well-developed pairs of postmentals, the inner pair longer than the outer pair, about half the size of the mental, and in contact behind the mental. Ventrolateral folds distinct, 23 25 scale rows across midventer. Five divided scansors beneath first digit of both manus and pes, seven beneath fourth digit of manus, seven beneath the fourth digit of pes. Males with a continuous series of eight precloacal pores. Body dorsum grayish brown, bordered by irregular, squiggly, dark brown markings that are interrupted by pairs of light gray spots, bordered by a cream/tan band that passes from the tip of the snout through the eye to the upper portion of the flanks and onto the tail, with numerous whitish tubercles on the flanks.
</paragraph>
</subSubSection>
<caption pageId="3" startId="3.[864,915,160,179]" targetBox="[840,1510,266,809]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph blockId="3.[840,1510,159,232]" pageId="3">
TABLE 1. Measurements and scale counts of the holotype and paratype of
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[967,1189,186,205]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[967,1189,186,205]" italics="true" pageId="3">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1202,1281,186,205]" pageId="3">
<taxonomicNameLabel box="[1202,1281,186,205]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Abbreviations are the same as those described in the Materials and Methods.
</paragraph>
</caption>
<paragraph pageId="3">
<table box="[840,1510,266,809]" gridcols="3" gridrows="22" pageId="3">
<tr box="[840,1510,266,285]" gridrow="0" pageId="3">
<th box="[840,1090,266,285]" gridcol="0" gridrow="0" pageId="3">Type status</th>
<th box="[1171,1300,266,285]" gridcol="1" gridrow="0" pageId="3">Holotype (mm)</th>
<th box="[1380,1510,266,285]" gridcol="2" gridrow="0" pageId="3">Paratype (mm)</th>
</tr>
<tr box="[840,1510,308,330]" gridrow="1" pageId="3">
<th box="[840,1090,308,330]" gridcol="0" gridrow="1" pageId="3">Catalog no.</th>
<td box="[1171,1300,308,330]" gridcol="1" gridrow="1" pageId="3">UTEP 22208</td>
<td box="[1380,1510,308,330]" gridcol="2" gridrow="1" pageId="3">UTEP 22209</td>
</tr>
<tr box="[840,1510,332,354]" gridrow="2" pageId="3">
<th box="[840,1090,332,354]" gridcol="0" gridrow="2" pageId="3">Sex</th>
<td box="[1171,1300,332,354]" gridcol="1" gridrow="2" pageId="3">Male</td>
<td box="[1380,1510,332,354]" gridcol="2" gridrow="2" pageId="3">Male</td>
</tr>
<tr box="[840,1510,356,378]" gridrow="3" pageId="3">
<th box="[840,1090,356,378]" gridcol="0" gridrow="3" pageId="3">SVL</th>
<td box="[1171,1300,356,378]" gridcol="1" gridrow="3" pageId="3">33.8</td>
<td box="[1380,1510,356,378]" gridcol="2" gridrow="3" pageId="3">39.5</td>
</tr>
<tr box="[840,1510,380,402]" gridrow="4" pageId="3">
<th box="[840,1090,380,402]" gridcol="0" gridrow="4" pageId="3">TL</th>
<td box="[1171,1300,380,402]" gridcol="1" gridrow="4" pageId="3">33.7</td>
<td box="[1380,1510,380,402]" gridcol="2" gridrow="4" pageId="3">35.1</td>
</tr>
<tr box="[840,1510,404,426]" gridrow="5" pageId="3">
<th box="[840,1090,404,426]" gridcol="0" gridrow="5" pageId="3">TRL</th>
<td box="[1171,1300,404,426]" gridcol="1" gridrow="5" pageId="3">14.2</td>
<td box="[1380,1510,404,426]" gridcol="2" gridrow="5" pageId="3">17.5</td>
</tr>
<tr box="[840,1510,428,450]" gridrow="6" pageId="3">
<th box="[840,1090,428,450]" gridcol="0" gridrow="6" pageId="3">BW</th>
<td box="[1171,1300,428,450]" gridcol="1" gridrow="6" pageId="3">5.1</td>
<td box="[1380,1510,428,450]" gridcol="2" gridrow="6" pageId="3">4.9</td>
</tr>
<tr box="[840,1510,452,474]" gridrow="7" pageId="3">
<th box="[840,1090,452,474]" gridcol="0" gridrow="7" pageId="3">FL</th>
<td box="[1171,1300,452,474]" gridcol="1" gridrow="7" pageId="3">4.9</td>
<td box="[1380,1510,452,474]" gridcol="2" gridrow="7" pageId="3">4.5</td>
</tr>
<tr box="[840,1510,476,498]" gridrow="8" pageId="3">
<th box="[840,1090,476,498]" gridcol="0" gridrow="8" pageId="3">CL</th>
<td box="[1171,1300,476,498]" gridcol="1" gridrow="8" pageId="3">5.4</td>
<td box="[1380,1510,476,498]" gridcol="2" gridrow="8" pageId="3">5.7</td>
</tr>
<tr box="[840,1510,500,521]" gridrow="9" pageId="3">
<th box="[840,1090,500,521]" gridcol="0" gridrow="9" pageId="3">HL</th>
<td box="[1171,1300,500,521]" gridcol="1" gridrow="9" pageId="3">10.0</td>
<td box="[1380,1510,500,521]" gridcol="2" gridrow="9" pageId="3">11.0</td>
</tr>
<tr box="[840,1510,523,545]" gridrow="10" pageId="3">
<th box="[840,1090,523,545]" gridcol="0" gridrow="10" pageId="3">HW</th>
<td box="[1171,1300,523,545]" gridcol="1" gridrow="10" pageId="3">6.0</td>
<td box="[1380,1510,523,545]" gridcol="2" gridrow="10" pageId="3">6.7</td>
</tr>
<tr box="[840,1510,547,569]" gridrow="11" pageId="3">
<th box="[840,1090,547,569]" gridcol="0" gridrow="11" pageId="3">HH</th>
<td box="[1171,1300,547,569]" gridcol="1" gridrow="11" pageId="3">3.8</td>
<td box="[1380,1510,547,569]" gridcol="2" gridrow="11" pageId="3">4.5</td>
</tr>
<tr box="[840,1510,572,593]" gridrow="12" pageId="3">
<th box="[840,1090,572,593]" gridcol="0" gridrow="12" pageId="3">ED</th>
<td box="[1171,1300,572,593]" gridcol="1" gridrow="12" pageId="3">1.8</td>
<td box="[1380,1510,572,593]" gridcol="2" gridrow="12" pageId="3">2.4</td>
</tr>
<tr box="[840,1510,595,617]" gridrow="13" pageId="3">
<th box="[840,1090,595,617]" gridcol="0" gridrow="13" pageId="3">NE</th>
<td box="[1171,1300,595,617]" gridcol="1" gridrow="13" pageId="3">2.7</td>
<td box="[1380,1510,595,617]" gridcol="2" gridrow="13" pageId="3">3.2</td>
</tr>
<tr box="[840,1510,619,640]" gridrow="14" pageId="3">
<th box="[840,1090,619,640]" gridcol="0" gridrow="14" pageId="3">SE</th>
<td box="[1171,1300,619,640]" gridcol="1" gridrow="14" pageId="3">4.3</td>
<td box="[1380,1510,619,640]" gridcol="2" gridrow="14" pageId="3">4.2</td>
</tr>
<tr box="[840,1510,643,665]" gridrow="15" pageId="3">
<th box="[840,1090,643,665]" gridcol="0" gridrow="15" pageId="3">EE</th>
<td box="[1171,1300,643,665]" gridcol="1" gridrow="15" pageId="3">2.6</td>
<td box="[1380,1510,643,665]" gridcol="2" gridrow="15" pageId="3">2.9</td>
</tr>
<tr box="[840,1510,667,689]" gridrow="16" pageId="3">
<th box="[840,1090,667,689]" gridcol="0" gridrow="16" pageId="3">Ear height</th>
<td box="[1171,1300,667,689]" gridcol="1" gridrow="16" pageId="3">0.7</td>
<td box="[1380,1510,667,689]" gridcol="2" gridrow="16" pageId="3">0.7</td>
</tr>
<tr box="[840,1510,691,713]" gridrow="17" pageId="3">
<th box="[840,1090,691,713]" gridcol="0" gridrow="17" pageId="3">Pores (right/left)</th>
<td box="[1171,1300,691,713]" gridcol="1" gridrow="17" pageId="3">8</td>
<td box="[1380,1510,691,713]" gridcol="2" gridrow="17" pageId="3">8</td>
</tr>
<tr box="[840,1510,715,737]" gridrow="18" pageId="3">
<th box="[840,1090,715,737]" gridcol="0" gridrow="18" pageId="3">Scales across belly</th>
<td box="[1171,1300,715,737]" gridcol="1" gridrow="18" pageId="3">25</td>
<td box="[1380,1510,715,737]" gridcol="2" gridrow="18" pageId="3">23</td>
</tr>
<tr box="[840,1510,739,761]" gridrow="19" pageId="3">
<th box="[840,1090,739,761]" gridcol="0" gridrow="19" pageId="3">Dorsal tubercle rows</th>
<td box="[1171,1300,739,761]" gridcol="1" gridrow="19" pageId="3">14</td>
<td box="[1380,1510,739,761]" gridcol="2" gridrow="19" pageId="3">12</td>
</tr>
<tr box="[840,1510,763,785]" gridrow="20" pageId="3">
<th box="[840,1090,763,785]" gridcol="0" gridrow="20" pageId="3">Supralabials (right/left)</th>
<td box="[1171,1300,763,785]" gridcol="1" gridrow="20" pageId="3">10/10</td>
<td box="[1380,1510,763,785]" gridcol="2" gridrow="20" pageId="3">10/9</td>
</tr>
<tr box="[840,1510,787,809]" gridrow="21" pageId="3">
<th box="[840,1090,787,809]" gridcol="0" gridrow="21" pageId="3">Infralabials (right/left)</th>
<td box="[1171,1300,787,809]" gridcol="1" gridrow="21" pageId="3">9/9</td>
<td box="[1380,1510,787,809]" gridcol="2" gridrow="21" pageId="3">8/8</td>
</tr>
</table>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7886504" ID-Zenodo-Dep="7886504" httpUri="https://zenodo.org/record/7886504/files/figure.png" pageId="3" startId="3.[146,174,1169,1188]" targetBox="[124,790,159,1142]" targetPageId="3" targetType="figure">
<paragraph blockId="3.[123,794,1169,1212]" pageId="3">
FIG. 2. Photos of live
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[358,572,1169,1188]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[358,572,1169,1188]" italics="true" pageId="3">Hemidactylus gramineus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[578,651,1169,1188]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
(Top: holotype UTEP 22208; bottom: paratype UTEP 22209). Photos by Eli Greenbaum.
</paragraph>
</caption>
<subSubSection lastPageId="4" pageId="3" type="description">
<paragraph blockId="3.[123,793,1257,2007]" lastBlockId="4.[123,793,921,2007]" lastPageId="4" pageId="3">
<emphasis box="[146,370,1682,1703]" italics="true" pageId="3">
Description of
<typeStatus box="[285,370,1682,1703]" pageId="3">Holotype</typeStatus>
</emphasis>
.—The
<typeStatus box="[445,536,1682,1704]" pageId="3">holotype</typeStatus>
is an adult male in good condition (
<figureCitation box="[237,296,1712,1734]" captionStart="FIG" captionStartId="2.[146,158,1019,1038]" captionTargetBox="[124,790,159,992]" captionTargetId="figure-819@2.[124,790,159,992]" captionTargetPageId="2" captionText="FIG. 1. Holotype (UTEP 22208) and paratype (UTEP 22209) of Hemidactylus gramineus sp. nov. Photos by Luis M. P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.7877130" httpUri="https://zenodo.org/record/7877130/files/figure.png" pageId="3">Fig. 1</figureCitation>
). The body shape is somewhat dorsoventrally flattened. Measurements and meristic characters of the
<typeStatus box="[701,792,1742,1764]" pageId="3">holotype</typeStatus>
are presented in
<tableCitation box="[302,379,1773,1795]" captionStart="TABLE" captionStartId="3.[864,876,160,179]" captionTargetBox="[124,790,159,1142]" captionTargetId="figure-612@3.[124,790,159,1142]" captionTargetPageId="3" captionText="TABLE 1. Measurements and scale counts of the holotype and paratype of Hemidactylus gramineus sp. nov. Abbreviations are the same as those described in the Materials and Methods." pageId="3">Table 1</tableCitation>
. Head large (HL/
<collectionCode box="[569,618,1773,1794]" pageId="3">SVL</collectionCode>
<quantity box="[626,721,1773,1794]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" unit="mm" value="0.3">0.30 mm</quantity>
), wide (HW/HL
<quantity box="[230,324,1803,1825]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" unit="mm" value="0.6">0.60 mm</quantity>
), not depressed (HH/HL
<quantity box="[603,697,1803,1824]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="3" unit="mm" value="0.38">0.38 mm</quantity>
), distinct from neck. Canthus rostralis moderately distinct. Snout short (SE/HL
<quantity box="[215,311,1864,1885]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="3" unit="mm" value="0.43">0.43 mm</quantity>
), longer than eye diameter; scales on snout, canthus rostralis, forehead and between eyes heterogeneous, juxtaposed, and weakly pointed; scales on snout, canthus rostralis and forehead twice the size of those on the occipital and interorbital regions. Eye small (ED/HL
<quantity box="[607,704,1985,2006]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="3" unit="mm" value="0.18">0.18 mm</quantity>
), supraciliaries large, pointed, with those at the anterior end of orbit slightly larger. Ear opening oval (greatest diameter
<quantity box="[1369,1449,902,923]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" unit="mm" value="0.7">0.7 mm</quantity>
); eyeto-ear distance greater than diameter of eye. Rostral wider than deep; three internasals, with the two outer ones enlarged and separated from each other by the smaller one in the middle; one supranasal on each side that is smaller than internasal, one pair of still smaller postnasals; rostral in contact with nostril, supralabial I, supranasal and internasal; nostrils small (&gt;
<quantity box="[1320,1385,1078,1099]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" unit="mm" value="1.0">1 mm</quantity>
), oval, each surrounded by supranasal, rostral, supralabial I and postnasal; one or two rows of scales separate orbit from supralabials. Mental triangular; two postmentals, smaller than the mental and in extensive contact with each other behind mental; outer pair about one third the size of the inner pair, separated from each other by inner pair and four gular scales. Inner postmental bordered by mental, infralabial I, outer postmental and two gular scales; outer postmental bordered by infralabials I and II, inner postmental, three to four small gular scales of which the outer one is enlarged and continue as a single row of enlarged scales below infralabials. Infralabials bordered by a double row of enlarged scales. Supralabials and infralabials to angle of jaw 10 (right and left) and 9 (right and left), respectively. Body relatively stout, moderately elongate (
<collectionCode box="[1144,1192,1487,1508]" pageId="3">TRL</collectionCode>
/
<collectionCode box="[1200,1247,1487,1508]" pageId="3">SVL</collectionCode>
<quantity box="[1259,1356,1487,1509]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="3" unit="mm" value="0.42">0.42 mm</quantity>
), ventrolateral folds without denticulate scales. Dorsal pholidosis heterogeneous, comprising conical, granular, striated scales intermixed with enlarged, fairly regularly arranged, longitudinal rows of 14 subtrihedral, strongly keeled, striated tubercles at midbody, extending from occipital region to tail, each enlarged tubercle roughly six to eight times longer than adjacent granules, surrounded by rosette of 1012 small granules, 2 or 3 granules between two adjacent enlarged tubercles; enlarged tubercles on nape and shoulder small and pointed, those on occipital region heterogeneous. Ventral scales larger than dorsal, smooth, imbricate; midbody scale rows across belly 25 (between ventrolateral folds); gular region with still smaller, subimbricate scales, those on lateral aspect of neck granular, anterior gular scales slightly larger than the rest. Eight precloacal pores. Scales on the palm and sole smooth, imbricate, rounded; scales on dorsal aspect of upper arm larger than granules on dorsum, subimbricate and striated, dorsal aspect of forearm with smaller, striated, conical and granular scales, intermixed with a few enlarged conical tubercles; those on dorsal part of thigh and shank conical, granular, striated, intermixed with enlarged, striated, subtrihedral tubercles, which are numerous on shank compared with anterior aspect of thigh; posterior aspect of thigh lacks enlarged tubercles.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7877132" ID-Zenodo-Dep="7877132" httpUri="https://zenodo.org/record/7877132/files/figure.png" pageId="4" startId="4.[864,876,424,443]" targetBox="[842,1508,159,397]" targetPageId="4" targetType="figure">
<paragraph blockId="4.[840,1510,424,491]" pageId="4">
<smallCapsWord baselines="438,438" box="[864,893,424,443]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Fig" pageId="4">FIG</smallCapsWord>
. 4. Photo of live
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[1109,1343,424,443]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[1109,1343,424,443]" italics="true" pageId="4">Hemidactylus gramineus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1358,1445,424,443]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
, from Mayongongo village, Pool department, Republic of the Congo. Photo by Sylvestre Boudzoumou.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7877134" ID-Zenodo-Dep="7877134" httpUri="https://zenodo.org/record/7877134/files/figure.png" pageId="4" startId="4.[146,158,762,781]" targetBox="[124,791,159,734]" targetPageId="4" targetType="figure">
<paragraph blockId="4.[123,794,761,876]" pageId="4">
<smallCapsWord baselines="776,775" box="[146,174,762,781]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Fig" pageId="4">FIG</smallCapsWord>
. 3. Distribution map for
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[423,638,761,780]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[423,638,761,780]" italics="true" pageId="4">Hemidactylus gramineus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[644,717,762,781]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
in DRC and Republic of the Congo. Blue star represents the type and paratype localities. White circle represents the photographic record of the new species from Republic of Congo. Black triangles represent the sister species
<taxonomicName box="[194,385,857,876]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nzingae">
<emphasis box="[194,385,857,876]" italics="true" pageId="4">Hemidactylus nzingae</emphasis>
</taxonomicName>
in Angola (data from
<bibRefCitation author="CERIACO, L. M. P., I. &amp; AGARWAL, M. P. &amp; MARQUES, AND &amp; A. M. BAUER" box="[594,784,857,876]" pageId="4" pagination="1 - 71" refId="ref7198" refString="CERIACO, L. M. P., I. AGARWAL, M. P. MARQUES, AND, A. M. BAUER. 2020 a. A review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two new species. Zootaxa 4746: 1 - 71." type="journal article" year="2020">Ceríaco et al., 2020a</bibRefCitation>
).
</paragraph>
</caption>
<paragraph blockId="4.[123,793,921,2007]" pageId="4">
Forelimbs and hind limbs relatively short, stout; forearm short (FL/
<collectionCode box="[230,278,1134,1155]" pageId="4">SVL</collectionCode>
<quantity box="[286,380,1134,1155]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="4" unit="mm" value="0.14">0.14 mm</quantity>
); tibia short (CL/
<collectionCode box="[563,612,1134,1155]" pageId="4">SVL</collectionCode>
<quantity box="[620,714,1134,1156]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="4" unit="mm" value="0.16">0.16 mm</quantity>
); digits moderately short, strongly clawed; all digits of manus and digits IIV of pes indistinctly webbed; terminal phalanx of all digits curved, arising angularly from distal portion of expanded lamellar pad, half or more than half as long as associated toepad; scansors beneath each toe in straight transverse series, divided except for distal and two basal scansors on digit I and one or two distal and basal scansors in all other digits that are single; scansors from proximal-most part of the toe at least twice diameter of palmar scales to distal-most single scansor: 5-6-6-7-7 (right manus), 5-8-7-7-6 (right pes). Relative length of digits:
<collectionCode box="[768,792,1438,1459]" pageId="4">III</collectionCode>
= IV&gt;
<collectionCode box="[202,219,1468,1489]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" type="Museum">V</collectionCode>
&gt; II&gt; I (right manus); IV&gt;
<collectionCode box="[518,542,1468,1489]" pageId="4">III</collectionCode>
&gt;
<collectionCode box="[572,589,1468,1489]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" type="Museum">V</collectionCode>
&gt; II&gt; I (right pes). The tail bears spinose tubercles dorsally, and subimbricate subcaudal scales, about one fourth of the width of the tail.
</paragraph>
<paragraph blockId="4.[123,793,921,2007]" pageId="4">
<emphasis box="[147,486,1560,1581]" italics="true" pageId="4">Coloration (in Life and PreserƲatiƲe)</emphasis>
.—Vertebral region of dorsum grayish brown, bordered by irregular, squiggly, dark brown markings that are interrupted by pairs of light gray spots; the dark brown markings transition to chevrons on the tail. Crown of head with scattered, poorly defined brown markings that resemble irregular chevrons. A cream/tan band passes from the tip of the snout through the eye to the upper portion of the flanks and onto the tail, with numerous whitish tubercles on the flanks. The lower portion of the flanks has a grayish brown band that is bordered by numerous light gray or whitish tubercles. The lowest margin of the flanks is light brown with irregular brown markings, and transitions to the venter. Forelimbs and hind limbs grayish brown with irregular dark brown crossbars. Venter cream; palms and soles cream. In preservative the specimens look more brownish with the pattern not so clearly delimited.
</paragraph>
<paragraph blockId="4.[840,1511,535,2007]" pageId="4">
<emphasis box="[864,952,536,557]" italics="true" pageId="4">Variation</emphasis>
.—Variation in scalation and body measurements of the
<typeStatus box="[883,975,566,587]" pageId="4">paratype</typeStatus>
of
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[1016,1258,565,587]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[1016,1258,565,587]" italics="true" pageId="4">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1268,1352,565,586]" pageId="4">
<taxonomicNameLabel box="[1268,1352,565,586]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is reported in
<tableCitation box="[840,916,595,617]" captionStart="TABLE" captionStartId="3.[864,876,160,179]" captionTargetBox="[124,790,159,1142]" captionTargetId="figure-612@3.[124,790,159,1142]" captionTargetPageId="3" captionText="TABLE 1. Measurements and scale counts of the holotype and paratype of Hemidactylus gramineus sp. nov. Abbreviations are the same as those described in the Materials and Methods." pageId="4">Table 1</tableCitation>
. The
<typeStatus box="[977,1069,595,616]" pageId="4">paratype</typeStatus>
agrees almost entirely with the
<typeStatus box="[1413,1506,595,617]" pageId="4">holotype</typeStatus>
, although its coloration is less contrasting than that of the
<typeStatus box="[840,934,654,676]" pageId="4">holotype</typeStatus>
. Coloration of a tentatively assigned third specimen (based on a photo of a live specimen) from
<location LSID="urn:lsid:plazi:treatment:039D49574E2EB05BA9A4FA3ACA4021CA:8EEBAE9A4E28B05AAF5AFD20CAF524DC" box="[1295,1510,683,705]" country="Republic of the Congo" name="Mayongongo village" pageId="4">Mayongongo village</location>
[-4.19, 14.96,
<elevation box="[986,1103,712,735]" metricMagnitude="2" metricUnit="m" metricValue="3.6799999999999997" pageId="4" unit="m" value="368.0">
<quantity box="[986,1050,713,735]" metricMagnitude="2" metricUnit="m" metricValue="3.6799999999999997" pageId="4" unit="m" value="368.0">368 m</quantity>
a.s.l.
</elevation>
], Pool Department,
<collectingCountry box="[1314,1510,713,735]" name="Republic of the Congo" pageId="4">Republic of Congo</collectingCountry>
(
<figureCitation box="[847,907,742,765]" captionStart="FIG" captionStartId="4.[864,876,424,443]" captionTargetBox="[842,1508,159,397]" captionTargetId="figure-909@4.[842,1508,159,397]" captionTargetPageId="4" captionText="FIG. 4. Photo of live Hemidactylus gramineus sp. nov., from Mayongongo village, Pool department, Republic of the Congo. Photo by Sylvestre Boudzoumou." figureDoi="http://doi.org/10.5281/zenodo.7877132" httpUri="https://zenodo.org/record/7877132/files/figure.png" pageId="4">Fig. 4</figureCitation>
), is nearly identical to that of the
<typeStatus box="[1263,1354,742,764]" pageId="4">holotype</typeStatus>
(
<figureCitation box="[1369,1429,742,765]" captionStart="FIG" captionStartId="3.[146,174,1169,1188]" captionTargetBox="[124,790,159,1142]" captionTargetId="figure-612@3.[124,790,159,1142]" captionTargetPageId="3" captionText="FIG. 2. Photos of live Hemidactylus gramineus sp. nov. (Top: holotype UTEP 22208; bottom: paratype UTEP 22209). Photos by Eli Greenbaum." figureDoi="http://doi.org/10.5281/zenodo.7886504" httpUri="https://zenodo.org/record/7886504/files/figure.png" pageId="4">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" pageId="4" type="discussion">
<paragraph blockId="4.[840,1511,535,2007]" pageId="4">
<emphasis box="[864,1441,772,794]" italics="true" pageId="4">Comparison aeith West and Central African Congeners</emphasis>
.—The newly described species can be morphologically distinguished from all of its regional congeners, including those occurring in the
<collectingCountry box="[840,891,861,882]" name="Democratic Republic of the Congo" pageId="4">DRC</collectingCountry>
and in the areas neighboring the western
<collectingCountry box="[1367,1421,861,882]" name="Democratic Republic of the Congo" pageId="4">DRC</collectingCountry>
:
<collectingCountry box="[1435,1506,861,883]" name="Gabon" pageId="4">Gabon</collectingCountry>
,
<collectingCountry box="[840,1081,890,912]" name="Republic of the Congo" pageId="4">Republic of the Congo</collectingCountry>
, and
<collectingCountry box="[1141,1219,890,912]" name="Angola" pageId="4">Angola</collectingCountry>
.
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[1232,1470,890,912]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[1232,1470,890,912]" italics="true" pageId="4">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="4">
<taxonomicNameLabel pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
may be readily distinguished from
<taxonomicName authorityName="Schmidt" authorityYear="1919" box="[1321,1453,921,942]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="ituriensis">
<emphasis box="[1321,1453,921,942]" italics="true" pageId="4">H. ituriensis</emphasis>
</taxonomicName>
and
<taxonomicName authority="Wagner, Leache and Fujita, 2014" authorityName="Wagner, Leache and Fujita" authorityYear="2014" box="[840,1410,950,972]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="coalescens">
<emphasis box="[840,1072,950,971]" italics="true" pageId="4">Hemidactylus coalescens</emphasis>
<bibRefCitation author="WAGNER, P. &amp; A. D. LEACHE &amp; M. K. FUJITA" box="[1077,1410,950,972]" pageId="4" pagination="1 - 14" refId="ref8782" refString="WAGNER, P., A. D. LEACHE, AND M. K. FUJITA. 2014. Description of four new West African forest geckos of the Hemidactylus fasciatus Gray, 1842 complex, revealed by coalescent species delimitation. Bonn Zoological Bulletin 63: 1 - 14." type="journal article" year="2014">Wagner, Leaché and Fujita, 2014</bibRefCitation>
</taxonomicName>
, by being much smaller (maximum [max]
<collectionCode box="[1203,1247,979,1000]" pageId="4">SVL</collectionCode>
39.5 vs.
<quantity box="[1358,1437,979,1001]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="4" unit="mm" value="78.0">78 mm</quantity>
in
<taxonomicName authorityName="Wagner, Leache and Fujita" authorityYear="2014" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="coalescens">
<emphasis italics="true" pageId="4">H. coalescens</emphasis>
</taxonomicName>
and
<quantity box="[998,1076,1009,1030]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="4" unit="mm" value="89.0">89 mm</quantity>
in
<taxonomicName authorityName="Schmidt" authorityYear="1919" box="[1120,1244,1009,1030]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="ituriensis">
<emphasis box="[1120,1244,1009,1030]" italics="true" pageId="4">H. ituriensis</emphasis>
</taxonomicName>
) and in not having the typical dorsal crossbands that characterize these two species (and the
<taxonomicName authorityName="Gray" authorityYear="1842" box="[880,993,1068,1089]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="fasciatus">
<emphasis box="[880,993,1068,1089]" italics="true" pageId="4">H. fasciatus</emphasis>
</taxonomicName>
group as a whole). It may be distinguished from
<taxonomicName authority="Hallowell, 1854" authorityName="Hallowell" authorityYear="1854" box="[840,1139,1097,1119]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="angulatus">
<emphasis box="[840,967,1097,1119]" italics="true" pageId="4">H. angulatus</emphasis>
Hallowell, 1854
</taxonomicName>
by having only 8 precloacal pores (vs. 2046 uninterrupted precloacal-femoral pores in males), by having a lower number of tubercle rows on the dorsum (1214 vs. 1425), and by having small subcaudal scales (vs. transversely enlarged).
</paragraph>
<paragraph blockId="4.[840,1511,535,2007]" pageId="4">
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[864,1106,1245,1267]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[864,1106,1245,1267]" italics="true" pageId="4">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1113,1196,1246,1267]" pageId="4">
<taxonomicNameLabel box="[1113,1196,1246,1267]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is readily distinguished from
<taxonomicName authority="Bauer and Pauwels, 2002" authorityName="Bauer and Pauwels" authorityYear="2002" box="[840,1299,1275,1297]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="kamdemtohami">
<emphasis box="[840,1017,1275,1297]" italics="true" pageId="4">H. kamdemtohami</emphasis>
Bauer and Pauwels, 2002
</taxonomicName>
, and
<taxonomicName authority="(Gray, 1845)" baseAuthorityName="Gray" baseAuthorityYear="1845" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="richardsonii">
<emphasis box="[1361,1510,1275,1297]" italics="true" pageId="4">H. richardsonii</emphasis>
(
<bibRefCitation author="GRAY, J. E." box="[847,962,1304,1326]" pageId="4" refId="ref7360" refString="GRAY, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. British Museum, UK." type="book" year="1845">Gray, 1845</bibRefCitation>
)
</taxonomicName>
by the lack of basal digital webbing and from
<taxonomicName authority="(Tornier, 1902)" baseAuthorityName="Tornier" baseAuthorityYear="1902" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="matschiei">
<emphasis italics="true" pageId="4">H. matschiei</emphasis>
(
<bibRefCitation author="TORNIER, G." box="[952,1098,1334,1356]" pageId="4" pagination="663 - 667" refId="ref8595" refString="TORNIER, G. 1902. Die Crocodile, Schildkroten und Eidechsen in Kamerun. Zoologische Jahrbucher 15: 663 - 667, pl. 35." type="journal article" year="1902">Tornier, 1902</bibRefCitation>
)
</taxonomicName>
by having spiny tubercles on the dorsum and small subcaudal scales. It may be distinguished from
<taxonomicName authorityName="Tornier" authorityYear="1902" box="[901,1064,1393,1415]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="steindachneri">
<emphasis box="[901,1064,1393,1415]" italics="true" pageId="4">H. steindachneri</emphasis>
</taxonomicName>
by lacking a longitudinal row of keeled tubercles on the ventrolateral border of the flanks, and from
<taxonomicName authorityName="Boulenger" authorityYear="1901" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="hecqui">
<emphasis italics="true" pageId="4">H. hecqui</emphasis>
</taxonomicName>
by not having the nostril in contact with the first supralabial, by having a relatively larger head depth (vs. flat head in
<taxonomicName authorityName="Boulenger" authorityYear="1901" box="[940,1039,1511,1533]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="hecqui">
<emphasis box="[940,1039,1511,1533]" italics="true" pageId="4">H. hecqui</emphasis>
</taxonomicName>
), lower number of tubercle rows on the dorsum (1214 vs. 16), by having fewer rows of ventral scales (2325 vs. 26), and a higher number of precloacal-femoral pores in males (8 vs. 6).
</paragraph>
<paragraph blockId="4.[840,1511,535,2007]" pageId="4">
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[864,1105,1630,1652]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[864,1105,1630,1652]" italics="true" pageId="4">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1113,1195,1630,1651]" pageId="4">
<taxonomicNameLabel box="[1113,1195,1630,1651]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName authorityName="Bocage" authorityYear="1873" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="longicephalus">
<emphasis italics="true" pageId="4">H. longicephalus</emphasis>
</taxonomicName>
by having a lower number of tubercle rows on the dorsum (1214 vs. 1718), fewer granular scales between the dorsal tubercles (13 vs. 36), and fewer rows of ventral scales (2325 vs. 3033). It can be distinguished from
<emphasis box="[1348,1441,1749,1770]" italics="true" pageId="4">H. paiƲae</emphasis>
by its smaller size (max SVL 39.5 vs.
<quantity box="[1186,1282,1778,1800]" metricMagnitude="-2" metricUnit="m" metricValue="6.84" pageId="4" unit="mm" value="68.4">68.4 mm</quantity>
), a lower number of granular scales between the dorsal tubercles (13 vs. 49), and a lower number of rows of ventral scales (2325 vs. 2834).
</paragraph>
<paragraph blockId="4.[840,1511,535,2007]" lastBlockId="5.[123,793,1273,2007]" lastPageId="5" pageId="4">
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[864,1104,1867,1888]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[864,1104,1867,1888]" italics="true" pageId="4">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1110,1191,1867,1888]" pageId="4">
<taxonomicNameLabel box="[1110,1191,1867,1888]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
may be distinguished from
<taxonomicName authorityName="Bocage" authorityYear="1893" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="benguellensis">
<emphasis italics="true" pageId="4">H. benguellensis</emphasis>
</taxonomicName>
by a much lower number of precloacal femoral pores in males (8 vs. 2333) and fewer granular scales between the dorsal tubercles (13 vs. 49). The new species can be distinguished from
<taxonomicName authorityName="Bocage" authorityYear="1893" box="[1049,1151,1985,2007]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="bayonii">
<emphasis box="[1049,1151,1985,2007]" italics="true" pageId="4">H. bayonii</emphasis>
</taxonomicName>
and
<emphasis box="[1206,1310,1985,2007]" italics="true" pageId="4">H. Ʋernayi</emphasis>
by fewer granular scales between the dorsal tubercles (13 vs. 34), and fewer rows of ventral scales (2325 vs.
<quantity box="[415,503,1303,1325]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" metricValueMax="8.128" metricValueMin="7.112" pageId="5" unit="in" value="30.0" valueMax="32.0" valueMin="28.0">2832 in</quantity>
<taxonomicName authorityName="Bocage" authorityYear="1893" box="[512,615,1303,1324]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="bayonii">
<emphasis box="[512,615,1303,1324]" italics="true" pageId="5">H. bayonii</emphasis>
</taxonomicName>
and
<quantity box="[671,759,1303,1325]" metricMagnitude="-1" metricUnit="m" metricValue="7.493" metricValueMax="7.874" metricValueMin="7.112" pageId="5" unit="in" value="29.5" valueMax="31.0" valueMin="28.0">2831 in</quantity>
<emphasis italics="true" pageId="5">H. Ʋernayi</emphasis>
).
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[219,464,1332,1354]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[219,464,1332,1354]" italics="true" pageId="5">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[474,559,1332,1353]" pageId="5">
<taxonomicNameLabel box="[474,559,1332,1353]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is easily distinguishable from
<taxonomicName authorityName="Moreau de Jonnes" authorityYear="1818" box="[232,344,1362,1384]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="mabouia">
<emphasis box="[232,344,1362,1384]" italics="true" pageId="5">H. mabouia</emphasis>
</taxonomicName>
by its smaller size (max SVL 39.5 vs.
<quantity metricMagnitude="-2" metricUnit="m" metricValue="6.74" pageId="5" unit="mm" value="67.4">67.4 mm</quantity>
), lower number of precloacal femoral pores in males (8 vs. 2839), fewer granular scales between the dorsal tubercles (13 vs. 510), and lower number of rows of ventral scales (2325 vs. 3437).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7877136" ID-Zenodo-Dep="7877136" httpUri="https://zenodo.org/record/7877136/files/figure.png" pageId="5" startId="5.[146,158,1162,1181]" targetBox="[124,790,159,1134]" targetPageId="5" targetType="figure">
<paragraph blockId="5.[123,792,1161,1228]" pageId="5">
<smallCapsWord baselines="1176,1175" box="[146,176,1162,1181]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Fig" pageId="5">FIG</smallCapsWord>
. 5. Grassy habitats of Bombo-Lumene Reserve, Kinshasa Province, southwestern Democratic Republic of the Congo. Photos by Eli Greenbaum.
</paragraph>
</caption>
<paragraph blockId="5.[123,793,1273,2007]" lastBlockId="5.[840,1511,159,2007]" pageId="5">
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[146,388,1510,1532]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[146,388,1510,1532]" italics="true" pageId="5">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[395,478,1510,1531]" pageId="5">
<taxonomicNameLabel box="[395,478,1510,1531]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName authorityName="Peters" authorityYear="1870" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="muriceus">
<emphasis italics="true" pageId="5">H. muriceus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[220,353,1540,1562]" pageId="5" sensu="stricto">sensu stricto</taxonomicNameLabel>
by a higher number of tubercle rows on the dorsum (1214 vs. 712) and fewer granular scales between the dorsal tubercles (13 vs. 510). The new species can be distinguished from
<taxonomicName authorityName="Henle &amp; Bohme" authorityYear="2003" box="[331,519,1629,1651]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="pseudomuriceus">
<emphasis box="[331,519,1629,1651]" italics="true" pageId="5">H. pseudomuriceus</emphasis>
</taxonomicName>
by having fewer precloacal femoral pores in males (8 vs. 1417) and fewer granular scales between the dorsal tubercles (13 vs. 510). The newly described species can be distinguished from
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1875" box="[613,721,1718,1740]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="echinus">
<emphasis box="[613,721,1718,1740]" italics="true" pageId="5">H. echinus</emphasis>
</taxonomicName>
by its lack of two rows of scattered spiny tubercles on the ventral side of the tail (a character that differentiates
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1875" box="[542,645,1777,1799]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="echinus">
<emphasis box="[542,645,1777,1799]" italics="true" pageId="5">H. echinus</emphasis>
</taxonomicName>
from all other known
<taxonomicName authorityName="Goldfuss" authorityYear="1820" box="[205,338,1807,1828]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="genus">
<emphasis box="[205,338,1807,1828]" italics="true" pageId="5">Hemidactylus</emphasis>
</taxonomicName>
), and by being smaller (max SVL 39.5 vs. 68.0 mm).
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[250,499,1837,1858]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[250,499,1837,1858]" italics="true" pageId="5">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[515,605,1837,1858]" pageId="5">
<taxonomicNameLabel box="[515,605,1837,1858]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName authorityName="Boulenger" authorityYear="1901" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="ansorgii">
<emphasis italics="true" pageId="5">H. ansorgii</emphasis>
</taxonomicName>
by a lower number of precloacal femoral pores (8 vs. 10 11) and a stockier body (
<taxonomicName authorityName="Boulenger" authorityYear="1901" box="[405,517,1896,1917]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="ansorgii">
<emphasis box="[405,517,1896,1917]" italics="true" pageId="5">H. ansorgii</emphasis>
</taxonomicName>
has a slender body and narrow head with a short interorbital distance).
<taxonomicName authorityName="Boulenger" authorityYear="1901" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="ansorgii">
<emphasis italics="true" pageId="5">Hemidactylus ansorgii</emphasis>
</taxonomicName>
also has a row of pointed tubercles separating the venter and the flanks, which are completely devoid of any tubercles (see figures in
<bibRefCitation author="TORNIER, G." box="[1020,1165,159,181]" pageId="5" pagination="663 - 667" refId="ref8595" refString="TORNIER, G. 1902. Die Crocodile, Schildkroten und Eidechsen in Kamerun. Zoologische Jahrbucher 15: 663 - 667, pl. 35." type="journal article" year="1902">Tornier, 1902</bibRefCitation>
; Perret, 1975), whereas in
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis italics="true" pageId="5">H. gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[949,1031,189,210]" pageId="5">
<taxonomicNameLabel box="[949,1031,189,210]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
this ventrolateral row of tubercles is lacking, and its tubercles extend across the flanks and dorsum.
</paragraph>
<paragraph blockId="5.[840,1511,159,2007]" pageId="5">
<taxonomicName authority="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai, 2023" authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[864,1105,248,270]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="gramineus" status="sp. nov.">
<emphasis box="[864,1105,248,270]" italics="true" pageId="5">Hemidactylus gramineus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1113,1195,248,269]" pageId="5">
<taxonomicNameLabel box="[1113,1195,248,269]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName authority="Ceriaco, Agarwal, Marques, and Bauer" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nzingae">
<emphasis italics="true" pageId="5">H. nzingae</emphasis>
</taxonomicName>
by having a lower number of tubercle rows on the dorsum (1214 vs. 1621) and by having irregular, undulate, dark brown markings that are interrupted by pairs of dorsal light gray spots (vs. well-marked series of dark-brown continuous W-shaped transverse markings from occiput to sacrum in
<taxonomicName authority="Ceriaco, Agarwal, Marques, and Bauer" authorityYear="2020" box="[960,1071,425,446]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nzingae">
<emphasis box="[960,1071,425,446]" italics="true" pageId="5">H. nzingae</emphasis>
</taxonomicName>
); the new species can be distinguished from
<taxonomicName authorityName="Ceríaco, Agarwal, Marques &amp; Bauer" authorityYear="2020" box="[896,1055,454,476]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="hannasabinae">
<emphasis box="[896,1055,454,476]" italics="true" pageId="5">H. hannasabinae</emphasis>
</taxonomicName>
by having a lower number of tubercle rows on the dorsum (1214 vs. 1618), by having strongly keeled, striated tubercles at midbody (vs. smoothly keeled in
<taxonomicName authorityName="Ceríaco, Agarwal, Marques &amp; Bauer" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="hannasabinae">
<emphasis italics="true" pageId="5">H. hannasabinae</emphasis>
</taxonomicName>
), and by being smaller (max SVL 39.5 vs.
<quantity box="[1406,1499,542,564]" metricMagnitude="-2" metricUnit="m" metricValue="4.74" pageId="5" unit="mm" value="47.4">47.4 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" type="distribution">
<paragraph blockId="5.[840,1511,159,2007]" pageId="5">
<emphasis box="[864,981,572,593]" italics="true" pageId="5">Distribution</emphasis>
.—The new species is confirmed from Bombo-Lumene Reserve in Kinshasa Province, southwestern
<collectingCountry box="[1404,1455,601,622]" name="Democratic Republic of the Congo" pageId="5">DRC</collectingCountry>
(
<figureCitation captionStart="FIG" captionStartId="4.[146,158,762,781]" captionTargetBox="[124,791,159,734]" captionTargetId="figure-851@4.[124,791,159,734]" captionTargetPageId="4" captionText="FIG. 3. Distribution map for Hemidactylus gramineus sp. nov. in DRC and Republic of the Congo. Blue star represents the type and paratype localities. White circle represents the photographic record of the new species from Republic of Congo. Black triangles represent the sister species Hemidactylus nzingae in Angola (data from Ceríaco et al., 2020a)." figureDoi="http://doi.org/10.5281/zenodo.7877134" httpUri="https://zenodo.org/record/7877134/files/figure.png" pageId="5">Fig. 3</figureCitation>
). A photo of a third specimen from Mayongongo village [-4.19, 14.96,
<quantity box="[905,967,660,682]" metricMagnitude="2" metricUnit="m" metricValue="3.6799999999999997" pageId="5" unit="m" value="368.0">368 m</quantity>
a.s.l.], Pool Department,
<collectingCountry box="[1227,1458,660,682]" name="Republic of the Congo" pageId="5">Republic of the Congo</collectingCountry>
(
<figureCitation captionStart="FIG" captionStartId="4.[864,876,424,443]" captionTargetBox="[842,1508,159,397]" captionTargetId="figure-909@4.[842,1508,159,397]" captionTargetPageId="4" captionText="FIG. 4. Photo of live Hemidactylus gramineus sp. nov., from Mayongongo village, Pool department, Republic of the Congo. Photo by Sylvestre Boudzoumou." figureDoi="http://doi.org/10.5281/zenodo.7877132" httpUri="https://zenodo.org/record/7877132/files/figure.png" pageId="5">Fig. 4</figureCitation>
) presents a remarkable morphological resemblance with the newly described species. The specimen is deposited in the laboratory of the Institut National des Sciences Exactes et Naturelles of the
<collectingCountry box="[1015,1206,778,800]" name="Republic of the Congo" pageId="5">Republic of Congo</collectingCountry>
in Brazzaville. We tentatively assign this specimen to the new species, but it was not possible to include it in our morphological or molecular datasets, and further research on this specimen and additional material are needed to confirm the presence of the species in the
<collectingCountry box="[1272,1507,895,917]" name="Republic of the Congo" pageId="5">Republic of the Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" type="biology_ecology">
<paragraph blockId="5.[840,1511,159,2007]" pageId="5">
<emphasis box="[864,1198,925,947]" italics="true" pageId="5">Habitat and Natural History Notes</emphasis>
.—The
<typeStatus box="[1273,1364,925,947]" pageId="5">holotype</typeStatus>
was collected in the early evening while on the ground near the bank of the Lumene River. The
<typeStatus box="[1049,1141,984,1005]" pageId="5">paratype</typeStatus>
was collected in the early evening while it was crawling through grass in sandy soil near a small pond.
</paragraph>
<paragraph blockId="5.[840,1511,159,2007]" pageId="5">
Bombo-Lumene Reserve is located ~
<quantity box="[1285,1418,1072,1094]" metricMagnitude="5" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="5" unit="km" value="140.0" valueMax="150.0" valueMin="130.0">130150 km</quantity>
east of
<collectingRegion box="[840,937,1102,1124]" country="Republic of the Congo" name="Kinshasa" pageId="5">Kinshasa</collectingRegion>
, in a hilly area known locally as the Bateke
<collectingRegion box="[1431,1509,1102,1124]" country="Benin" name="Plateau" pageId="5">Plateau</collectingRegion>
(name derived from the local Bateke tribal people), at an elevation of ~
<quantity box="[996,1111,1161,1183]" metricMagnitude="2" metricUnit="m" metricValue="6.8" metricValueMax="9.6" metricValueMin="4.0" pageId="5" unit="m" value="680.0" valueMax="960.0" valueMin="400.0">400960 m</quantity>
a.s.l. (
<bibRefCitation author="KAYUMBA, M. &amp; C. LUBINI &amp; T. E. KIDIKWADI &amp; J. - P. HABARI" box="[1185,1421,1160,1183]" pageId="5" pagination="716 - 727" refId="ref7704" refString="KAYUMBA, M., C. LUBINI, T. E. KIDIKWADI, AND J. - P. HABARI. 2015. Floristic study of the mature formation of Bombo-Lumene Domain and Reserve (Kinshasa / RD Congo). International Journal of Innovation and Applied Studies, 11: 716 - 727." type="journal article" year="2015">Kayumba et al., 2015</bibRefCitation>
). It was originally created as a Game Reserve in 1968, but was elevated in 1976 to Natural Reserve International Union for Conservation of Nature (IUCN) category VI because of its ecological importance (
<bibRefCitation author="KAYUMBA, M. &amp; C. LUBINI &amp; T. E. KIDIKWADI &amp; J. - P. HABARI" box="[976,1210,1278,1300]" pageId="5" pagination="716 - 727" refId="ref7704" refString="KAYUMBA, M., C. LUBINI, T. E. KIDIKWADI, AND J. - P. HABARI. 2015. Floristic study of the mature formation of Bombo-Lumene Domain and Reserve (Kinshasa / RD Congo). International Journal of Innovation and Applied Studies, 11: 716 - 727." type="journal article" year="2015">Kayumba et al., 2015</bibRefCitation>
). The rainy season lasts for 8 mo from mid-September to mid-May, whereas the 4-mo dry season occurs from mid-May to mid-September. The rainfalls reach an annual range from
<quantity box="[1173,1405,1366,1388]" metricMagnitude="0" metricUnit="m" metricValue="1.61485" metricValueMax="1.6297" metricValueMin="1.6" pageId="5" unit="mm" value="1614.85" valueMax="1629.7" valueMin="1600.0">1,600 to 1,629.7 mm</quantity>
, and the average annual temperature is 24.4
<emphasis bold="true" box="[1222,1230,1396,1418]" italics="true" pageId="5">°</emphasis>
C (
<bibRefCitation author="KAYUMBA, M. &amp; C. LUBINI &amp; T. E. KIDIKWADI &amp; J. - P. HABARI" box="[1264,1500,1396,1418]" pageId="5" pagination="716 - 727" refId="ref7704" refString="KAYUMBA, M., C. LUBINI, T. E. KIDIKWADI, AND J. - P. HABARI. 2015. Floristic study of the mature formation of Bombo-Lumene Domain and Reserve (Kinshasa / RD Congo). International Journal of Innovation and Applied Studies, 11: 716 - 727." type="journal article" year="2015">Kayumba et al., 2015</bibRefCitation>
). Bombo-Lumene Reserve is a forestsavanna mosaic (i.e., western Congolian forestsavanna mosaic sensu
<bibRefCitation author="BURGESS, N. &amp; J. D. HALES &amp; E. UNDERWOOD AND &amp; E. DINERSTEIN" pageId="5" refId="ref7159" refString="BURGESS, N., J. D. HALES, E. UNDERWOOD AND, E. DINERSTEIN. 2004. Terrestrial Ecoregions of Africa and Madagascar: A Conservation Assessment. World Wildlife Fund, USA." type="book" year="2004">Burgess et al. [2004]</bibRefCitation>
), with nearly all forest occurring as gallery forest adjacent to the Bombo or Lumene Rivers (Greenbaum, pers. obs.).
</paragraph>
<paragraph blockId="5.[840,1511,159,2007]" pageId="5">
We encountered the new species during the dry season that typically lasts for 4 mo (
<bibRefCitation author="MILAU, F. &amp; C. KIFUKIETO &amp; C. KACHAKA &amp; J. ALONI &amp; F. FRANCIS" box="[1109,1302,1573,1595]" pageId="5" pagination="37 - 44" refId="ref8022" refString="MILAU, F., C. KIFUKIETO, C. KACHAKA, J. ALONI, AND F. FRANCIS. 2016. Contribution a l'etude de la fauna associee a la decomposition du bois (Isoptera et Haplotaxida) a Bombo-Lumene au plateau des Batekes (RDC). Entomologie Faunistique 69: 37 - 44." type="journal article" year="2016">Milau et al., 2016</bibRefCitation>
), and we observed large brush fires during our (CK and EG) brief visit to the reserve. Other lizards encountered at the reserve included
<taxonomicName authorityName="Moreau de Jonnes" authorityYear="1818" box="[840,1060,1661,1682]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="mabouia">
<emphasis box="[840,1060,1661,1682]" italics="true" pageId="5">Hemidactylus mabouia</emphasis>
</taxonomicName>
(in or near human habitations),
<taxonomicName authority="Gray, 1845" authorityName="Gray" authorityYear="1845" class="Reptilia" family="Scincidae" genus="Trachylepis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="maculilabris">
<emphasis italics="true" pageId="5">Trachylepis maculilabris</emphasis>
<bibRefCitation author="GRAY, J. E." box="[966,1080,1690,1712]" pageId="5" refId="ref7360" refString="GRAY, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. British Museum, UK." type="book" year="1845">Gray, 1845</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="Hallowell, 1857" authorityName="Hallowell" authorityYear="1857" class="Reptilia" family="Gerrhosauridae" genus="Gerrhosaurus" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nigrolineatus">
<emphasis box="[1093,1224,1690,1711]" italics="true" pageId="5">Gerrhosaurus</emphasis>
cf.
<emphasis box="[1265,1392,1690,1711]" italics="true" pageId="5">nigrolineatus</emphasis>
Hallowell, 1857
</taxonomicName>
; and
<taxonomicName authority="Bocage, 1866" authorityName="Bocage" authorityYear="1866" box="[953,1279,1720,1742]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="cabindae">
<emphasis box="[953,1131,1720,1741]" italics="true" pageId="5">Panaspis cabindae</emphasis>
Bocage, 1866
</taxonomicName>
(
<bibRefCitation author="MEDINA, M. F. &amp; A. M. BAUER &amp; W. R. BRANCH &amp; A. SCHMITZ &amp; W. CONRADIE &amp; Z. T. NAGY &amp; T. J. HIBBITTS &amp; R. ERNST &amp; D. M. PORTIK &amp; S. V. NIELSEN &amp; C. KUSAMBA &amp; T. J. COLSTON &amp; M. BEHANGANA &amp; M. - O. RODEL &amp; E. GREENBAUM" box="[1295,1506,1720,1742]" pageId="5" pagination="409 - 423" refId="ref7910" refString="MEDINA, M. F., A. M. BAUER, W. R. BRANCH, A. SCHMITZ, W. CONRADIE, Z. T. NAGY, T. J. HIBBITTS, R. ERNST, D. M. PORTIK, S. V. NIELSEN, C. KUSAMBA, T. J. COLSTON, M. BEHANGANA, M. - O. RODEL, AND E. GREENBAUM. 2016. Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution 100: 409 - 423." type="journal article" year="2016">Medina et al., 2016</bibRefCitation>
;
<bibRefCitation author="ALLEN, K. E. &amp; W. P. TAPONDJOU &amp; E. GREENBAUM &amp; L. J. WELTON &amp; A. M. BAUER" box="[840,1025,1749,1771]" pageId="5" pagination="231 - 241" refId="ref6739" refString="ALLEN, K. E., W. P. TAPONDJOU, E. GREENBAUM, L. J. WELTON, AND A. M. BAUER. 2019. High levels of hidden phylogenetic structure and diversity within Central and West African Trachylepis skinks. Salamandra 55: 231 - 241." type="journal article" year="2019">Allen et al., 2019</bibRefCitation>
). The specimen from
<collectingCountry box="[1258,1459,1749,1771]" name="Republic of the Congo" pageId="5">Republic of Congo</collectingCountry>
was collected in savannah near a village (K. Jackson, pers. com.
<date pageId="5" value="2009-01-14">14 January 2009</date>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" type="etymology">
<paragraph blockId="5.[840,1511,159,2007]" pageId="5">
<emphasis box="[864,964,1838,1859]" italics="true" pageId="5">Etymology</emphasis>
.—The specific epithet
<taxonomicName authorityName="Ching &amp; Tay &amp; Brown &amp; Mohareb &amp; Sethi &amp; Annamalai" authorityYear="2023" box="[1240,1340,1838,1859]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="gramineus">
<emphasis box="[1240,1340,1838,1859]" italics="true" pageId="5">gramineus</emphasis>
</taxonomicName>
refers to the grassy
<typeStatus box="[916,962,1868,1889]" pageId="5">type</typeStatus>
of habitat from which the species was collected in the
<collectingCountry box="[881,932,1897,1918]" name="Democratic Republic of the Congo" pageId="5">DRC</collectingCountry>
and the
<collectingCountry box="[1029,1267,1897,1919]" name="Republic of the Congo" pageId="5">Republic of the Congo</collectingCountry>
(
<figureCitation box="[1283,1342,1896,1919]" captionStart="FIG" captionStartId="5.[146,158,1162,1181]" captionTargetBox="[124,790,159,1134]" captionTargetId="figure-866@5.[124,790,159,1134]" captionTargetPageId="5" captionText="FIG. 5. Grassy habitats of Bombo-Lumene Reserve, Kinshasa Province, southwestern Democratic Republic of the Congo. Photos by Eli Greenbaum." figureDoi="http://doi.org/10.5281/zenodo.7877136" httpUri="https://zenodo.org/record/7877136/files/figure.png" pageId="5">Fig. 5</figureCitation>
) and is applied here as an adjective in the genitive singular. We propose the French common name of Gecko de lherbe and the English common name of grass-dwelling tropical gecko.
</paragraph>
</subSubSection>
</treatment>
</document>