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<document id="38F1A0C16697E5D59B061138A7B495C2" ID-ISSN="0003-0090" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630348657882" checkinUser="felipe" docAuthor="Ríos-Tamayo, D. &amp; Goloboff, P. A." docDate="2018" docId="C22D980AFFB7F324730CFD47D32EFCDB" docLanguage="en" docName="B419.pdf" docOrigin="Bulletin of the American Museum of Natural History 2018 (419)" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Actinopus taragui Ríos-Tamayo &amp; Goloboff 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="73" masterDocId="3E14E072FFF2F36C702CFFD7D734FFB4" masterDocTitle="Taxonomic Revision And Morphology Of The Trapdoor Spider Genus Actinopus (Mygalomorphae: Actinopodidae) In Argentina" masterLastPageNumber="84" masterPageNumber="1" pageNumber="70" updateTime="1699240534849" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A2277F1C64F9E1F809A14EEBEA094ADE">Taxonomic Revision And Morphology Of The Trapdoor Spider Genus Actinopus (Mygalomorphae: Actinopodidae) In Argentina</mods:title>
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<mods:namePart id="DADD354137B8D7F0FD195F61D5225C7B">Ríos-Tamayo, D.</mods:namePart>
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<mods:namePart id="9D4A56A094F6793F32F59052B47A1452">Goloboff, P. A.</mods:namePart>
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<mods:number id="39AFDA364E3A79F19979C00AD5B7B921">2018-03-20</mods:number>
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<subSubSection id="029E7A97FFB7F329730CFD47D37DFD1C" box="[800,1097,656,680]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="4A3B291CFFB7F329730CFD47D37DFD1C" blockId="69.[800,1097,656,680]" box="[800,1097,656,680]" pageId="69" pageNumber="70">
<heading id="11739E70FFB7F329730CFD47D37DFD1C" box="[800,1097,656,680]" centered="true" fontSize="9" level="2" pageId="69" pageNumber="70" reason="2">
<taxonomicName id="8D84529FFFB7F329730CFD47D4DDFD1C" ID-CoL="64PM3" authority="Ríos-Tamayo &amp; Goloboff, 2018" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[800,1001,656,680]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="taragui" status="sp. nov.">
<emphasis id="78F0F50EFFB7F329730CFD47D4DDFD1C" bold="true" box="[800,1001,656,680]" italics="true" pageId="69" pageNumber="70">Actinopus taragui</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="63C34875FFB7F32973DAFD46D37DFD1C" box="[1014,1097,657,680]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="029E7A97FFB7F3297308FD15D370FD6E" box="[804,1092,706,730]" pageId="69" pageNumber="70" type="description">
<paragraph id="4A3B291CFFB7F3297308FD15D370FD6E" blockId="69.[804,1092,706,730]" box="[804,1092,706,730]" pageId="69" pageNumber="70">
<heading id="11739E70FFB7F3297308FD15D370FD6E" box="[804,1092,706,730]" centered="true" fontSize="9" level="2" pageId="69" pageNumber="70" reason="2">
<figureCitation id="D2BF3599FFB7F3297308FD15D4FFFD6E" box="[804,971,706,730]" captionStart="FIGURE 36" captionStartId="70.[109,195,1113,1134]" captionTargetBox="[209,1111,225,1080]" captionTargetId="figure-271@70.[209,1111,225,1085]" captionTargetPageId="70" captionText="FIGURE 36. Actinopus taragui, sp. nov., male holotype (MACN-Ar 36624). A, carapace. B, tibia II, retrolateral. C, palpal tibia. DF, copulatory bulb, three different views. G, sternum. Scale bars = 1 mm." pageId="69" pageNumber="70">Figures 36AG</figureCitation>
,
<figureCitation id="D2BF3599FFB7F32973FBFD15D32BFD6E" box="[983,1055,706,730]" captionStart="FIGURE 37" captionStartId="71.[108,194,816,837]" captionTargetBox="[247,1074,224,781]" captionTargetId="figure-444@71.[247,1074,224,789]" captionTargetPageId="71" captionText="FIGURE 37. Actinopus taragui, sp. nov., female paratype (MACN-Ar 36625). A, carapace. B, tibia II, retrolateral. C, tibia I, prolateral. DE, spermathecae. D, Parque Provincial “Islas Malvinas,” Misiones. E, Corrientes, Corrientes. F, sternum. Scale bars = 1 mm." pageId="69" pageNumber="70">37AF</figureCitation>
,
<figureCitation id="D2BF3599FFB7F3297406FD15D370FD6E" box="[1066,1092,706,730]" captionStart="FIGURE 44" captionStartId="78.[109,195,1450,1471]" captionTargetBox="[225,1096,836,1416]" captionTargetId="figure-104@78.[220,1103,832,1420]" captionTargetPageId="78" captionText="FIGURE 44. Known distribution of the species with type II morphology (part) in Argentina. Actinopus argenteus sp. nov., Actinopus ariasi sp. nov., Actinopus coylei sp. nov., Actinopus gerschiapelliarum sp. nov., Actinopus indiamuerta sp. nov., Actinopus insignis (Holmberg, 1881), Actinopus longipalpis Koch, 1842, Actinopus ramirezi sp. nov., Actinopus taragui sp. nov. Additional records (in Uruguay) for A. gerschiapelliarum and A. longipalpis are given in the list of specimens examined." pageId="69" pageNumber="70">44</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="029E7A97FFB7F32972E0FD23D497FC60" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="4A3B291CFFB7F32972E0FD23D497FC60" blockId="69.[683,1213,755,1747]" pageId="69" pageNumber="70">
<collectionCode id="2C95B1D9FFB7F32972E0FD23D4B0FCB8" box="[716,900,756,780]" pageId="69" pageNumber="70">TYPE MATERIAL</collectionCode>
:
<typeStatus id="953F97BEFFB7F32973BEFD23D33EFCB8" box="[914,1034,756,780]" pageId="69" pageNumber="70" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="5C82E295FFB7F329743BFD24D31AFCB8" box="[1047,1070,755,780]" pageId="69" pageNumber="70" type="male"></specimenCount>
(MACN-Ar 36624).
<emphasis id="78F0F50EFFB7F3297328FCC2D44DFC99" bold="true" box="[772,889,789,813]" italics="true" pageId="69" pageNumber="70">Corrientes</emphasis>
: Capital [
<geoCoordinate id="2FB04FDBFFB7F32973CDFCC2D30AFC99" box="[993,1086,788,813]" degrees="27" direction="south" minutes="28" orientation="latitude" pageId="69" pageNumber="70" precision="925" value="-27.466667">27°28 S</geoCoordinate>
<geoCoordinate id="2FB04FDBFFB7F3297469FCC2D385FC99" box="[1093,1201,788,813]" degrees="58" direction="west" minutes="49" orientation="longitude" pageId="69" pageNumber="70" precision="925" value="-58.816666">58°49 W</geoCoordinate>
],
<date id="3E3A0FDCFFB7F3297281FCE0D408FCFB" box="[685,828,823,847]" pageId="69" pageNumber="70" value="2007-05-13">13 May. 2007</date>
, G. Rubio,
<specimenCount id="5C82E295FFB7F329739DFCE0D4E0FCFB" box="[945,980,822,847]" pageId="69" pageNumber="70" type="male">1♂</specimenCount>
.
<typeStatus id="953F97BEFFB7F32973CCFCE0D379FCFB" box="[992,1101,823,847]" pageId="69" pageNumber="70" type="paratype">PARATYPE</typeStatus>
:
<emphasis id="78F0F50EFFB7F3297475FCE0D389FCFA" box="[1113,1213,823,846]" italics="true" pageId="69" pageNumber="70">Misiones.</emphasis>
<specimenCount id="5C82E295FFB7F3297280FC8FD5E4FCC4" box="[684,720,855,880]" pageId="69" pageNumber="70" type="female">3♀</specimenCount>
, (together with
<specimenCount id="5C82E295FFB7F32973AAFC8FD327FCC4" box="[902,1043,856,880]" pageId="69" pageNumber="70" type="immature">2 immatures</specimenCount>
, not
<typeStatus id="953F97BEFFB7F3297461FC8FD38CFCC4" box="[1101,1208,856,880]" pageId="69" pageNumber="70" type="paratype">paratypes</typeStatus>
, MACN-Ar 36625), Parque Provincial “
<collectingCountry id="3293698CFFB7F32974A5FCAED424FC07" name="Falkland Islands" pageId="69" pageNumber="70">Islas Malvinas</collectingCountry>
,” Arroyo Uruzú [
<geoCoordinate id="2FB04FDBFFB7F32973FAFC4DD303FC07" box="[982,1079,922,947]" degrees="25" direction="south" minutes="58" orientation="latitude" pageId="69" pageNumber="70" precision="925" value="-25.966667">25°58 S</geoCoordinate>
<geoCoordinate id="2FB04FDBFFB7F329746DFC4DD385FC06" box="[1089,1201,922,947]" degrees="54" direction="west" minutes="11" orientation="longitude" pageId="69" pageNumber="70" precision="925" value="-54.183334">54°11 W</geoCoordinate>
],
<date id="3E3A0FDCFFB7F3297280FC6BD422FC60" box="[684,790,956,980]" pageId="69" pageNumber="70" value="1988-02">Feb. 1988</date>
, P. Goloboff.
</paragraph>
</subSubSection>
<subSubSection id="029E7A97FFB7F32972E0FC0AD33DFB8C" pageId="69" pageNumber="70" type="etymology">
<paragraph id="4A3B291CFFB7F32972E0FC0AD33DFB8C" blockId="69.[683,1213,755,1747]" pageId="69" pageNumber="70">
ETYMOLOGY: The specific name is a noun in apposition to the
<typeStatus id="953F97BEFFB7F3297354FC28D493FBA3" box="[888,935,1023,1047]" pageId="69" pageNumber="70">type</typeStatus>
locality; in the Guarani language, it means “
<collectingRegion id="8840E7FEFFB7F32973A4FBF7D4C8FB8C" box="[904,1020,1056,1080]" country="Argentina" name="Corrientes" pageId="69" pageNumber="70">Corrientes</collectingRegion>
.”
</paragraph>
</subSubSection>
<subSubSection id="029E7A97FFB7F32972E0FB96D32AF99F" pageId="69" pageNumber="70" type="diagnosis">
<paragraph id="4A3B291CFFB7F32972E0FB96D32AF99F" blockId="69.[683,1213,755,1747]" pageId="69" pageNumber="70">
DIAGNOSIS: Males of
<taxonomicName id="8D84529FFFB7F3297380FB95D327FBED" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[940,1043,1090,1113]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="taragui">
<emphasis id="78F0F50EFFB7F3297380FB95D327FBED" box="[940,1043,1090,1113]" italics="true" pageId="69" pageNumber="70">A. taragui</emphasis>
</taxonomicName>
present a copulatory bulb (fig. 36DF) with Ak poorly developed, extending to the apex, and MRk reduced. They differ from
<taxonomicName id="8D84529FFFB7F3297329FB71D46FFB09" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[773,859,1190,1213]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="ariasi">
<emphasis id="78F0F50EFFB7F3297329FB71D46FFB09" box="[773,859,1190,1213]" italics="true" pageId="69" pageNumber="70">A. ariasi</emphasis>
</taxonomicName>
by having more retrolateral spines on tibia II, but fewer than present in
<taxonomicName id="8D84529FFFB7F329746CFB10D381FB6A" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[1088,1205,1223,1246]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="ramirezi">
<emphasis id="78F0F50EFFB7F329746CFB10D381FB6A" box="[1088,1205,1223,1246]" italics="true" pageId="69" pageNumber="70">A. ramirezi</emphasis>
</taxonomicName>
. Males differ from both by the lighter color in apical leg segments (patellae to tarsi) and by the presence of rastellum. Females can be distinguished by the rectangular shape of the spermathecae (fig. 37D, E), with base wide and a lobe on the outer part. Differ from
<taxonomicName id="8D84529FFFB7F32972CBFA58D450FA13" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[743,868,1423,1447]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="szumikae">
<emphasis id="78F0F50EFFB7F32972CBFA58D450FA13" box="[743,868,1423,1447]" italics="true" pageId="69" pageNumber="70">A. szumikae</emphasis>
</taxonomicName>
and
<taxonomicName id="8D84529FFFB7F32973B5FA58D4C5FA13" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[921,1009,1423,1447]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="coylei">
<emphasis id="78F0F50EFFB7F32973B5FA58D4C5FA13" box="[921,1009,1423,1447]" italics="true" pageId="69" pageNumber="70">A. coylei</emphasis>
</taxonomicName>
by the shape of the fovea; from
<taxonomicName id="8D84529FFFB7F3297307FA67D49DFA7C" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[811,937,1456,1480]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="szumikae">
<emphasis id="78F0F50EFFB7F3297307FA67D49DFA7C" box="[811,937,1456,1480]" italics="true" pageId="69" pageNumber="70">A. szumikae</emphasis>
</taxonomicName>
also by the lower number of thorns apical to the apical crown on tibia III, and darker coloration; from
<taxonomicName id="8D84529FFFB7F3297381FA24D332F9BF" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[941,1030,1523,1547]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="coylei">
<emphasis id="78F0F50EFFB7F3297381FA24D332F9BF" box="[941,1030,1523,1547]" italics="true" pageId="69" pageNumber="70">A. coylei</emphasis>
</taxonomicName>
by the sigilla not reaching the center of the sternum.
</paragraph>
</subSubSection>
<subSubSection id="029E7A97FFB7F32472E0F9E2D53BFA56" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70" type="description">
<paragraph id="4A3B291CFFB7F32A72E0F9E2D3BAF9D6" blockId="69.[683,1213,755,1747]" lastBlockId="70.[683,1213,1213,1733]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
DESCRIPTION:
<typeStatus id="953F97BEFFB7F3297356F9E2D4D4F9F9" box="[890,992,1589,1613]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
male (MACN-Ar 36624, fig. 36AG):
<emphasis id="78F0F50EFFB7F32973A1F980D327F9DB" box="[909,1043,1623,1647]" italics="true" pageId="69" pageNumber="70">Total length:</emphasis>
7.44.
<emphasis id="78F0F50EFFB7F3297476F980D388F9DA" box="[1114,1212,1623,1646]" italics="true" pageId="69" pageNumber="70">Carapace</emphasis>
(fig. 36A): length 3.30, width 3.06.
<emphasis id="78F0F50EFFB7F3297435F9AFD389F924" box="[1049,1213,1656,1680]" italics="true" pageId="69" pageNumber="70">Cephalic region:</emphasis>
length 1.80, width 2.16; clypeus small with few small bristles, 9 bristles between PME and 4 behind each PME-PLE, 6 on each side of the OQ.
<emphasis id="78F0F50EFFB4F32A7216FB69D548FB61" box="[570,636,1214,1237]" italics="true" pageId="70" pageNumber="71">Fovea:</emphasis>
length 0.54, width 1.08.
<emphasis id="78F0F50EFFB4F32A7142FB08D7F8FAA3" italics="true" pageId="70" pageNumber="71">Eyes, diameters and interdistances:</emphasis>
AME 0.16 (small), ALE 0.28, PME 0.12, PLE 0.16, OQ length 0.64, anterior width 1.60, 1.40 posterior width; AME-ALE 0.32, PME-PLE 0.08, AME-PME 0.32, ALE-PLE 0.20, AME-AME 0.20.
<emphasis id="78F0F50EFFB4F32A7042FA54D7E8FA2F" box="[110,220,1411,1435]" italics="true" pageId="70" pageNumber="71">Chelicerae:</emphasis>
Length 2.76, width 1.38; cheliceral apex with 9 recumbent and elongated bristles that cover the base of the fang. Rastellum with 34 marginal cusps; cheliceral furrow with 6 large promarginal teeth and 6 retromarginal (small ones between 3d-4th and 4th-5th) and 9 denticles, near the promarginal teeth.
<emphasis id="78F0F50EFFB4F32A7135F99DD645F9D6" box="[281,369,1610,1634]" italics="true" pageId="70" pageNumber="71">Labium:</emphasis>
length 0.68, width 0.76.
<emphasis id="78F0F50EFFB4F32A7040F9BCD7FBF937" box="[108,207,1643,1667]" italics="true" pageId="70" pageNumber="71">Maxillae:</emphasis>
anterior length 0.88, posterior length 1.44, width 1.04.
<emphasis id="78F0F50EFFB4F32A7136F95BD646F917" box="[282,370,1676,1699]" italics="true" pageId="70" pageNumber="71">Sternum</emphasis>
(fig. 36G): Center slightly depressed; length 2.40, maximum width 2.08, mini- mum width 0.76; postlabial sigillum shallow, posterior sigilla well defined and shallow.
<emphasis id="78F0F50EFFB4F32A747DFB09D388FB42" box="[1105,1212,1246,1270]" italics="true" pageId="70" pageNumber="71">Abdomen:</emphasis>
Length 2.82, small, covered with thin bristles, thicker and more numerous on the dorsum. PMS: length 0.32; PLS: length of basal:medial:apical segments 0.44:0.16:0.08.
<emphasis id="78F0F50EFFB4F32A73BAFAB4D4D0FACE" box="[918,996,1379,1402]" italics="true" pageId="70" pageNumber="71">Spigots:</emphasis>
1 large, 6 small on PMS; PLS, numbers of large(small) spigots on basal:medial:apical article 2(10): (the other articles deformed, could not be observed).
<emphasis id="78F0F50EFFB4F32A7409FA11D424FA4B" italics="true" pageId="70" pageNumber="71">Lengths of legs and palp:</emphasis>
I: 3.42, 1.44, 2.10, 2.70, 1.80, 11.46. II: 3.18, 1.38, 2.10, 2.70, 1.80, 11.16. III: 2.58, 1.50, 1.56, 2.82, 1.92, 10.38. IV: 3.30, 1.50, 2.70, 3.00, 2.10, 12.60. Palp: 3.60, 1.80, 3.12, , 1.02, 9.54.
</paragraph>
<caption id="1EFB7994FFB4F32A7041FB8ED326FB38" pageId="70" pageNumber="71" startId="70.[109,195,1113,1134]" targetBox="[209,1111,225,1080]" targetPageId="70">
<paragraph id="4A3B291CFFB4F32A7041FB8ED326FB38" blockId="70.[108,1213,1111,1164]" pageId="70" pageNumber="71">
FIGURE 36.
<taxonomicName id="8D84529FFFB4F32A70C0FB80D6AFFBDB" authority="Ríos-Tamayo &amp; Goloboff, 2018" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[236,411,1111,1135]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="taragui" status="sp. nov.">
<emphasis id="78F0F50EFFB4F32A70C0FB80D6AFFBDB" box="[236,411,1111,1135]" italics="true" pageId="70" pageNumber="71">Actinopus taragui</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="63C34875FFB4F32A718AFB80D6C7FBDA" box="[422,499,1111,1134]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (MACN-Ar 36624).
<emphasis id="78F0F50EFFB4F32A7371FB80D45BFBDB" bold="true" box="[861,879,1111,1135]" pageId="70" pageNumber="71">A</emphasis>
, carapace.
<emphasis id="78F0F50EFFB4F32A73F1FB80D4D9FBDB" bold="true" box="[989,1005,1111,1135]" pageId="70" pageNumber="71">B</emphasis>
, tibia II, retrolateral.
<emphasis id="78F0F50EFFB4F32A7040FBA3D749FB38" bold="true" box="[108,125,1140,1164]" pageId="70" pageNumber="71">C</emphasis>
, palpal tibia.
<emphasis id="78F0F50EFFB4F32A7125FBA3D603FB38" bold="true" box="[265,311,1140,1164]" pageId="70" pageNumber="71">DF</emphasis>
, copulatory bulb, three different views.
<emphasis id="78F0F50EFFB4F32A72E2FBA3D5EBFB38" bold="true" box="[718,735,1140,1164]" pageId="70" pageNumber="71">G</emphasis>
, sternum. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="4A3B291CFFB4F32A72E1F9BCD389F971" blockId="70.[683,1213,1213,1733]" pageId="70" pageNumber="71">
<emphasis id="78F0F50EFFB4F32A72E1F9BCD47CF937" box="[717,840,1643,1667]" italics="true" pageId="70" pageNumber="71">Chaetotaxy:</emphasis>
All femora, 0. Patellae: I-II, 0; III, 22 P SUP-D ANT, +8 on the margin, 1-1-1 D (3:4 B), 6 R A +4 on the margin; IV, 44 P SUP-D ANT +4
</paragraph>
<caption id="1EFB7994FFB5F32B7040FCE7D517FC34" pageId="71" pageNumber="72" startId="71.[108,194,816,837]" targetBox="[247,1074,224,781]" targetPageId="71">
<paragraph id="4A3B291CFFB5F32B7040FCE7D517FC34" blockId="71.[107,1209,813,896]" pageId="71" pageNumber="72">
FIGURE 37.
<taxonomicName id="8D84529FFFB5F32B70C0FCF9D6A8FCF1" authority="Ríos-Tamayo &amp; Goloboff, 2018" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[236,412,814,837]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="taragui" status="sp. nov.">
<emphasis id="78F0F50EFFB5F32B70C0FCF9D6A8FCF1" box="[236,412,814,837]" italics="true" pageId="71" pageNumber="72">Actinopus taragui</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="63C34875FFB5F32B7185FCF9D6C3FCF1" box="[425,503,814,837]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype (MACN-Ar 36625).
<emphasis id="78F0F50EFFB5F32B7354FCFAD4BDFCF1" bold="true" box="[888,905,813,837]" pageId="71" pageNumber="72">A</emphasis>
, carapace.
<emphasis id="78F0F50EFFB5F32B73D5FCF9D33DFCF2" bold="true" box="[1017,1033,814,838]" pageId="71" pageNumber="72">B</emphasis>
, tibia II, retrolateral.
<emphasis id="78F0F50EFFB5F32B70B0FC9CD799FCD7" bold="true" box="[156,173,843,867]" pageId="71" pageNumber="72">C</emphasis>
, tibia I, prolateral.
<emphasis id="78F0F50EFFB5F32B7143FC9CD6A9FCD7" bold="true" box="[367,413,843,867]" pageId="71" pageNumber="72">DE</emphasis>
, spermathecae.
<emphasis id="78F0F50EFFB5F32B726DFC9CD560FCD7" bold="true" box="[577,596,843,867]" pageId="71" pageNumber="72">D</emphasis>
, Parque Provincial “Islas Malvinas,” Misiones.
<emphasis id="78F0F50EFFB5F32B7401FC9CD30FFCD7" bold="true" box="[1069,1083,843,867]" pageId="71" pageNumber="72">E</emphasis>
, Corrientes, Corrientes.
<emphasis id="78F0F50EFFB5F32B70CFFCBFD7C5FC34" bold="true" box="[227,241,872,896]" pageId="71" pageNumber="72">F</emphasis>
, sternum. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="4A3B291CFFB5F32B7040FC40D548F97C" blockId="71.[107,638,919,1737]" pageId="71" pageNumber="72">
near the edge. Tibiae: I, 2 V ANT A, 1-1-1-1-2 V POST; II, 1-1-2-3-3-4 R, 1-3-2-4 V; III, 1 P, 1 D B, with an apical crown of 15 thorns, 1-1-1 R, 1-1-1 V POST, 2 V ANT A; IV, 2-2 v, 2 A POST, 1-1-2 V ANT. Metatarsi: I, 1-2-1-1-2-V POST, 2-0-2-1-1 V ANT; II, 1-1 V ANT, 1-1 V, 1-2-1-2-1-2-4 V POST- R INF; III, 1-1-2-2-4 on row D POST-R, 1-2-1 V POST, 1-1-0-1-1-1 V ANT, 1-1-1 D ANT; IV, 1-2- 2-1-1 V ANT-P INF, 1-1-2 V POST. Tarsi: I, 1-1- 1-1 P INF, 1-1-2-1-1-3-3 R INF, 1-1 V; II, 1-1-2-1-1 P INF, 1-1-3-1-4-2-3 R INF, 1-1 V; III, 1-1-1-1-1-1- 2-2 P INF, 2-2-3-2-2 R INF (3:4 A); IV, 2-1-2-4-1-5 P INF, 2-1-2-1 R INF. Total number of retrolateral spines on tibiae I and II (fig. 36B): 0 and 14 respectively.
<emphasis id="78F0F50EFFB5F32B709DFAB1D7D1FACA" box="[177,229,1382,1406]" italics="true" pageId="71" pageNumber="72">Palp:</emphasis>
tibia long and slender (width/length: 0.31±0.01) (fig. 36C). Copulatory bulb (fig. 36DF) thin, BTA and ATA developed, embolus with a series of denticles at the base.
<emphasis id="78F0F50EFFB5F32B7188FA1ED503FA55" box="[420,567,1481,1505]" italics="true" pageId="71" pageNumber="72">Pseudoscopula</emphasis>
: Tarsi: I sparse (1:2 A); II sparse (3:4 A); III-IV, dense and uniform. Metatarsi: IV sparse and apical.
<emphasis id="78F0F50EFFB5F32B7040F9FBD7C9F9F0" box="[108,253,1580,1604]" italics="true" pageId="71" pageNumber="72">Trichobothria:</emphasis>
Tibiae: I 3-2; II 4-3; III 3-5; IV 6-6. Metatarsi: I-II 10; III 7; IV 6. Tarsi: I-II 7; III 9; IV 6.
<emphasis id="78F0F50EFFB5F32B70AAF9B9D7F2F932" box="[134,198,1646,1670]" italics="true" pageId="71" pageNumber="72">Color:</emphasis>
Carapace dark reddish brown. Abdomen dark brown, with small bristles, booklungs with dark markings. Sternum, labium, and maxillae yel-
</paragraph>
<paragraph id="4A3B291CFFB5F32B7280FC40D36DFC64" blockId="71.[684,1215,919,1737]" pageId="71" pageNumber="72">lowish. Legs like carapace; tarsi-metatarsi-tibiae and patellae lighter. Spinnerets yellowish.</paragraph>
<paragraph id="4A3B291CFFB5F32472E0FC0DD6DCFD5E" blockId="71.[684,1215,919,1737]" lastBlockId="72.[107,639,226,1738]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<typeStatus id="953F97BEFFB5F32B72E0FC0DD41DFC46" box="[716,809,986,1010]" pageId="71" pageNumber="72" type="paratype">Paratype</typeStatus>
female (MACN-Ar 36625, figs. 37A F):
<emphasis id="78F0F50EFFB5F32B72FCFC2DD466FBA6" box="[720,850,1018,1042]" italics="true" pageId="71" pageNumber="72">Total length:</emphasis>
24.42.
<emphasis id="78F0F50EFFB5F32B73B2FC2CD334FBA6" box="[926,1024,1019,1042]" italics="true" pageId="71" pageNumber="72">Carapace</emphasis>
(fig. 37A): length 8.16, width 7.56, with 50 bristles on each posterior margin.
<emphasis id="78F0F50EFFB5F32B7324FBEBD487FBE0" box="[776,947,1084,1108]" italics="true" pageId="71" pageNumber="72">Cephalic region:</emphasis>
length 5.28, width 6.48; clypeus with 90 bristles and one central large; chilum with 70 bristles, 68 bristles between AME- ALE, 42 between PME; abundant bristles behind each PME-PLE dyad, and on each side of the OQ.
<emphasis id="78F0F50EFFB5F32B7280FB35D5DBFB4D" box="[684,751,1250,1273]" italics="true" pageId="71" pageNumber="72">Fovea:</emphasis>
length 1.44, width 3.6.
<emphasis id="78F0F50EFFB5F32B73C2FB35D470FAAF" italics="true" pageId="71" pageNumber="72">Eyes, diameters and interdistances:</emphasis>
AME 0.20, ALE 0.48, PME 0.28, PLE 0.40, AME-ALE 1.00, PME-PLE 0.12, AME- PME 0.56, ALE-PLE 0.68, AME-AME 0.48, OQ length 1.28, anterior width 3.68, posterior width 3.56.
<emphasis id="78F0F50EFFB5F32B72C4FA50D46EFA2B" box="[744,858,1415,1439]" italics="true" pageId="71" pageNumber="72">Chelicerae:</emphasis>
Length 4.20; width 2.70; cheliceral apex with 19 recumbent and elongated bristles covering the base of the fang; dorsal anterior region with shallow ridges. Rastellum with 12 marginal, blunt cusps, and 10 small dorsal ones. Fang groove with 7 promarginal teeth and 6 retromarginal (a small one between 3d-4th and 4th- 5th; and 3 internal teeth next to the last two) and 12 denticles in the furrow, near the promargin.
<emphasis id="78F0F50EFFB5F32B7280F967D430F97C" box="[684,772,1712,1736]" italics="true" pageId="71" pageNumber="72">Labium:</emphasis>
length 1.86, width 1.92, with 24 anterior cuspules.
<emphasis id="78F0F50EFFBAF32470F8FF35D602FF4E" box="[212,310,226,250]" italics="true" pageId="72" pageNumber="73">Maxillae:</emphasis>
anterior length 2.34, posterior length 3.00, width 2.34, with +120 cuspules 1:3 V ANT, expanding to the anterior face.
<emphasis id="78F0F50EFFBAF32471DFFEF3D578FE8F" box="[499,588,292,315]" italics="true" pageId="72" pageNumber="73">Sternum</emphasis>
(fig. 37F): Center slightly depressed; length 5.58, maximum width 4.62, minimum width 1.92. Postlabial sigillum deep, posterior sigilla shallow but well defined.
<emphasis id="78F0F50EFFBAF32470E4FE7FD600FE74" box="[200,308,424,448]" italics="true" pageId="72" pageNumber="73">Abdomen:</emphasis>
Length 11.16, globose covered with small bristles. PMS: length 0.88; PLS: length of basal:medial:apical segments 1.00:0.72:0.40.
<emphasis id="78F0F50EFFBAF3247040FDDBD78FFD97" box="[108,187,524,547]" italics="true" pageId="72" pageNumber="73">Spigots:</emphasis>
4 large, 50 small on PMS; PLS, numbers of large(small) spigots on basal:medial:apical article 6(38):9(52):3(+70).
<emphasis id="78F0F50EFFBAF3247148FD9AD553FDD1" box="[356,615,589,613]" italics="true" pageId="72" pageNumber="73">Lengths of legs and palp:</emphasis>
I: 4.74, 3.18, 2.34, 2.70, 1.20, 14.16. II: 4.74, 3.18, 2.28, 2.94, 1.20, 14.34. III: 4.32, 3.60, 1.80, 3.18, 1.20, 14.10. IV: 6.00, 3.36, 3.36, 3.60, 1.38, 17.70. Palp: 4.38, 2.88, 3.18, , 3.00, 13.44.
</paragraph>
<paragraph id="4A3B291CFFBAF32470A1FD24D53BFA56" blockId="72.[107,639,226,1738]" pageId="72" pageNumber="73">
<emphasis id="78F0F50EFFBAF32470A1FD24D63CFCBF" box="[141,264,755,779]" italics="true" pageId="72" pageNumber="73">Chaetotaxy:</emphasis>
All femora: 0. Patellae: I-II, 0; III, 12 P A, +10 on the margin, 24 R (1:2 A) +16 on the margin; IV, 40 P SUP-D ANT +5 on the edge; palp, 1-1 V POST, 5 P (serial). Tibiae: I, 1-1-1-1 P (1:3 A), 24 R INF, 1-1 V POST; II, 120 R (3:4 A), 1 V POST between 1-1 V POST; III, 0/1 D BAS, 70 R (3:4 A), with apical crown of 19 thorns and 3 apical (at the apex, apical to the crown); IV, 0; palp, 13 P, 24 R INF. Metatarsi: I, 25 P INF, 25 R INF, 1-1-1-2 V POST; II, 27 P INF, 22 R INF, 1-1-1-2 V POST; III, 55 on row D POST-R (more abundant R A), 21 D ANT, 2 V A; IV, 16 P, 1 D ANT A. Tarsi: I, 9 P INF, 12 R INF, 2 V A; II, 15 P INF, 10 R INF, 2 VA; III, 25 R, 98 V-P; IV, 49 V-P; palp, 18 P INF, 29 R INF, 2 V A. Total number of retrolateral spines on tibiae I and II: 24 and 120 (fig. 37B) respectively.
<emphasis id="78F0F50EFFBAF3247040FAD3D7C8FAA8" box="[108,252,1284,1308]" italics="true" pageId="72" pageNumber="73">Trichobothria:</emphasis>
Tibiae: I-II 6-7; III 6-9; IV 7-7. Metatarsi: I 12; II 16; III-IV 12. Tarsi: I 10; II-III 12; IV 8.
<emphasis id="78F0F50EFFBAF32470A8FA91D7F0FAEA" box="[132,196,1350,1374]" italics="true" pageId="72" pageNumber="73">Color:</emphasis>
carapace dark reddish brown, dark chelicerae with light dorsal bands and dark dorsal spots. Abdomen brown, booklungs with dark markings. Sternum yellowish, labium and maxillae darker. Legs like carapace. Spinnerets yellowish.
</paragraph>
</subSubSection>
<subSubSection id="029E7A97FFBAF32470A0FA3CD4ACFDF1" pageId="72" pageNumber="73" type="distribution">
<paragraph id="4A3B291CFFBAF32470A0FA3CD66CF990" blockId="72.[107,639,226,1738]" pageId="72" pageNumber="73">
DISTRIBUTION: Provinces of
<collectingRegion id="8840E7FEFFBAF32471D7FA3CD543F9B7" box="[507,631,1515,1539]" country="Argentina" name="Corrientes" pageId="72" pageNumber="73">Corrientes</collectingRegion>
,
<taxonomicName id="8D84529FFFBAF3247040F9DBD787F990" authorityName="Tullgren" authorityYear="1905" box="[108,179,1548,1572]" class="Arachnida" family="Nemesiidae" genus="Chaco" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">Chaco</taxonomicName>
, and
<collectingRegion id="8840E7FEFFBAF32470C3F9DBD660F990" box="[239,340,1548,1572]" country="Paraguay" name="Misiones" pageId="72" pageNumber="73">Misiones</collectingRegion>
.
</paragraph>
<paragraph id="4A3B291CFFBAF32470A0F9FAD4ACFDF1" blockId="72.[107,639,226,1738]" lastBlockId="72.[684,1212,225,879]" pageId="72" pageNumber="73">
OTHER MATERIAL EXAMINED:
<emphasis id="78F0F50EFFBAF32471D0F9FAD541F9F1" bold="true" box="[508,629,1581,1605]" italics="true" pageId="72" pageNumber="73">
<collectingRegion id="8840E7FEFFBAF32471D0F9FAD541F9F1" box="[508,629,1581,1605]" country="Argentina" name="Corrientes" pageId="72" pageNumber="73">Corrientes</collectingRegion>
</emphasis>
: Capital [
<geoCoordinate id="2FB04FDBFFBAF32470E7F998D619F9D3" box="[203,301,1613,1639]" degrees="27" direction="south" minutes="28" orientation="latitude" pageId="72" pageNumber="73" precision="925" value="-27.466667">27°28 S</geoCoordinate>
<geoCoordinate id="2FB04FDBFFBAF324711AF999D693F9D2" box="[310,423,1613,1638]" degrees="58" direction="west" minutes="49" orientation="longitude" pageId="72" pageNumber="73" precision="925" value="-58.816666">58°49 W</geoCoordinate>
],
<date id="3E3A0FDCFFBAF3247190F998D57BF9D2" box="[444,591,1614,1639]" pageId="72" pageNumber="73" value="2006-04-13">13 Apr. 2006</date>
, M. Chatelanaz,
<specimenCount id="5C82E295FFBAF32470DCF9A7D620F933" box="[240,276,1646,1672]" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
(CARTROUNNE 7285).
<date id="3E3A0FDCFFBAF3247204F9A7D790F91D" pageId="72" pageNumber="73" value="2007-05-13">13 May. 2007</date>
, G. Rubio,
<specimenCount id="5C82E295FFBAF324711FF946D66CF91C" box="[307,344,1679,1705]" pageId="72" pageNumber="73" type="male">1♂</specimenCount>
(CARTROUNNE 8168). Paraje Yahapé, Departamento Berón de Astrada [
<geoCoordinate id="2FB04FDBFFBAF3247298FF34D424FF4F" box="[692,784,225,251]" degrees="27" direction="south" minutes="22" orientation="latitude" pageId="72" pageNumber="73" precision="925" value="-27.366667">27°22 S</geoCoordinate>
<geoCoordinate id="2FB04FDBFFBAF324733AFF35D4B7FF4E" box="[790,899,225,250]" degrees="57" direction="west" minutes="39" orientation="longitude" pageId="72" pageNumber="73" precision="925" value="-57.65">57°39 W</geoCoordinate>
],
<date id="3E3A0FDCFFBAF32473B9FF35D323FF4F" box="[917,1047,226,251]" pageId="72" pageNumber="73" value="2008-03-06">6 Mar. 2008</date>
, G. Ávalos,
<specimenCount id="5C82E295FFBAF32474B4FF34D388FF4E" box="[1176,1212,225,251]" pageId="72" pageNumber="73" type="female">2♀</specimenCount>
(CARTROUNNE 8170). Cerca de Galarza (grassland), no collector,
<date id="3E3A0FDCFFBAF32473EFFEF2D365FE88" box="[963,1105,292,317]" pageId="72" pageNumber="73" value="2005-04-28">28 Apr. 2005</date>
,
<specimenCount id="5C82E295FFBAF3247472FEF2D5E3FEE9" pageId="72" pageNumber="73" type="immature">1 immature</specimenCount>
(CARTROUNNE 8169). No date and collector,
<specimenCount id="5C82E295FFBAF32472F3FEB0D430FECB" box="[735,772,358,383]" pageId="72" pageNumber="73" type="male">1♂</specimenCount>
(CARTROUNNE 8171).
<emphasis id="78F0F50EFFBAF324741BFEB1D3B1FECA" box="[1079,1157,358,382]" italics="true" pageId="72" pageNumber="73">
<taxonomicName id="8D84529FFFBAF324741BFEB1D34BFECA" authorityName="Tullgren" authorityYear="1905" box="[1079,1151,358,382]" class="Arachnida" family="Nemesiidae" genus="Chaco" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="78F0F50EFFBAF324741BFEB1D34BFECA" bold="true" box="[1079,1151,358,382]" italics="true" pageId="72" pageNumber="73">Chaco</emphasis>
</taxonomicName>
:
</emphasis>
San Bernardo [
<geoCoordinate id="2FB04FDBFFBAF324730DFE5FD44BFE14" box="[801,895,391,416]" degrees="27" direction="south" minutes="28" orientation="latitude" pageId="72" pageNumber="73" precision="925" value="-27.466667">27°28 S</geoCoordinate>
<geoCoordinate id="2FB04FDBFFBAF32473AAFE5FD4C7FE2B" box="[902,1011,391,416]" degrees="58" direction="west" minutes="55" orientation="longitude" pageId="72" pageNumber="73" precision="925" value="-58.916668">58°55 W</geoCoordinate>
],
<date id="3E3A0FDCFFBAF324742AFE5FD3A3FE14" box="[1030,1175,392,416]" pageId="72" pageNumber="73" value="1991-03-16">16 Mar. 1991</date>
, O. Di Iorio,
<specimenCount id="5C82E295FFBAF3247332FE7ED475FE75" box="[798,833,424,449]" pageId="72" pageNumber="73" type="male">1♂</specimenCount>
(MACN-Ar 30737). No specific locality,
<date id="3E3A0FDCFFBAF3247327FE1DD4A7FE56" box="[779,915,458,482]" pageId="72" pageNumber="73" value="2006-04-03">3 Apr. 2006</date>
, V. Fernandez,
<specimenCount id="5C82E295FFBAF3247469FE1DD35EFE56" box="[1093,1130,457,482]" pageId="72" pageNumber="73" type="male">1♂</specimenCount>
(CAR- TROUNNE 8167).
<emphasis id="78F0F50EFFBAF32473AAFE3CD4DDFDB7" bold="true" box="[902,1001,491,515]" italics="true" pageId="72" pageNumber="73">
<collectingRegion id="8840E7FEFFBAF32473AAFE3CD4DDFDB7" box="[902,1001,491,515]" country="Paraguay" name="Misiones" pageId="72" pageNumber="73">Misiones</collectingRegion>
</emphasis>
: San Ignacio, near banks of
<collectingRegion id="8840E7FEFFBAF3247333FDDBD45AFD90" box="[799,878,524,548]" country="Brazil" name="Parana" pageId="72" pageNumber="73">Paraná</collectingRegion>
,
<date id="3E3A0FDCFFBAF32473ACFDDAD4C4FD90" box="[896,1008,524,548]" pageId="72" pageNumber="73" value="2012-01">Jan. 2012</date>
, P. Goloboff,
<specimenCount id="5C82E295FFBAF32474B4FDDBD388FD90" box="[1176,1212,523,548]" pageId="72" pageNumber="73" type="male">1♂</specimenCount>
(FML-ARAN 02231).
</paragraph>
</subSubSection>
<subSubSection id="029E7A97FFBAF32472E1FD99D32EFCDB" pageId="72" pageNumber="73" type="discussion">
<paragraph id="4A3B291CFFBAF32472E1FD99D32EFCDB" blockId="72.[684,1212,225,879]" pageId="72" pageNumber="73">
COMMENTS: The specimens from Yahapé differ in the color of the booklung markings and the shape of the fovea. The specimen of San Ignacio, differs by its larger size and the legs with just metatarsi and tarsi lighter than the rest of the segments. In the absence of additional specimens, it seems best to include them in
<taxonomicName id="8D84529FFFBAF3247460FCC2D382FC98" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2018" box="[1100,1206,789,812]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="taragui">
<emphasis id="78F0F50EFFBAF3247460FCC2D382FC98" box="[1100,1206,789,812]" italics="true" pageId="72" pageNumber="73">A. taragui</emphasis>
</taxonomicName>
, because of their similarity in other characters and their geographical proximity.
</paragraph>
</subSubSection>
</treatment>
</document>