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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.205.76790" ID-Pensoft-Pub="1314-2003-205-99" ID-Pensoft-UUID="4024A478048757B197E5CF8734331A9E" ModsDocID="1314-2003-205-99" checkinTime="1661230211668" checkinUser="pensoft" docAuthor="O'Donnell, Shawn A., Ringelberg, Jens J. &amp; Lewis, Gwilym P." docDate="2022" docId="8DCC3817207E5061AE4EFBCDFE7EBB54" docLanguage="en" docName="PhytoKeys 205: 99-145" docOrigin="PhytoKeys 205" docPubDate="2022-08-22" docSource="http://dx.doi.org/10.3897/phytokeys.205.76790" docTitle="Entada rheedei Spreng., Syst. Veg. 2: 325. 1825." docType="treatment" docVersion="1" id="4024A478048757B197E5CF8734331A9E" lastPageNumber="99" masterDocId="4024A478048757B197E5CF8734331A9E" masterDocTitle="Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1661230211668" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>O'Donnell, Shawn A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0731-7425</mods:nameIdentifier>
<mods:affiliation>Department of Geography and Environmental Sciences, Northumbria University, Newcastle upon Tyne, NE 1 8 ST, UK</mods:affiliation>
<mods:nameIdentifier type="email">shawn.odonnell@cantab.net</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ringelberg, Jens J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0567-5210</mods:nameIdentifier>
<mods:affiliation>Department of Systematic and Evolutionary Botany, University of Zurich, 8008 Zurich, Switzerland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lewis, Gwilym P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2599-4577</mods:nameIdentifier>
<mods:affiliation>Accelerated Taxonomy Department, Royal Botanic Gardens, Kew, Richmond, TW 9 3 AE, UK</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>205</mods:number>
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<mods:extent unit="page">
<mods:start>99</mods:start>
<mods:end>145</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.205.76790</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.205.76790</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-205-99</mods:identifier>
<mods:identifier type="Pensoft-UUID">4024A478048757B197E5CF8734331A9E</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:8DCC3817207E5061AE4EFBCDFE7EBB54" httpUri="http://treatment.plazi.org/id/8DCC3817207E5061AE4EFBCDFE7EBB54" lastPageNumber="99" pageId="0" pageNumber="99">
<subSubSection pageId="0" pageNumber="99" type="nomenclature">
<paragraph pageId="0" pageNumber="99">
<taxonomicName LSID="8DCC3817-207E-5061-AE4E-FBCDFE7EBB54" authority="Spreng., Syst. Veg. 2: 325. 1825." authorityName="Spreng., Syst. Veg. 2: 325. 1825." class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada rheedei" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="rheedei">Entada rheedei Spreng., Syst. Veg. 2: 325. 1825.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="reference_group">
<paragraph pageId="0" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada pursaetha" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="pursaetha">= Entada pursaetha</taxonomicName>
DC.,
<normalizedToken originalValue="Mém">Mem</normalizedToken>
.
<normalizedToken originalValue="Légum">Legum</normalizedToken>
.: 421. 1826.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada monostachya" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="monostachya">= Entada monostachya</taxonomicName>
DC.,
<normalizedToken originalValue="Mém">Mem</normalizedToken>
.
<normalizedToken originalValue="Légum">Legum</normalizedToken>
.: 422. 1826.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada gogo" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="gogo">= Entada gogo</taxonomicName>
I.M. Johnst., Sargentia 8: 137. 1949.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="materials_examined">
<paragraph pageId="0" pageNumber="99">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<collectingCountry name="India">INDIA</collectingCountry>
. Malabar coast, illustration in
<emphasis italics="true" pageId="0" pageNumber="99">Rheede</emphasis>
Hort. Malab. 9: 151, tab. 77. 1689.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="description">
<paragraph pageId="0" pageNumber="99">Description.</paragraph>
<paragraph pageId="0" pageNumber="99">
Liana to 75 m long, stems to 30 cm diameter at base (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genus-wide variation in morphological characters. A-D habit A large woody liana, E. rheedei (photo: B Wursten, Hyde et al. (2021)) B small tree, E. abyssinica (photo: G Baumann, Dressler et al. (2014 a)) C shrub, E. burkei (photo: M Schmidt, Dressler et al. (2014 a, b )) D geoxylic suffrutex, E. dolichorrhachis (photo: M Bingham, Bingham et al. (2021)) E, F leaves E bipinnate leaf with few pinnae, few large leaflets and ending in a bifurcating tendril, E. rheedei (photo: AP Balan, Balan and Predeep (2021)) F bipinnate leaf with many pinnae, many small leaflets and no tendril, E. rangei (photo: A Dreyer, Dressler et al. (2014 a)) G-I inflorescences G terminal panicle of up-turned spikes, E. polyphylla (photo: R Vasquez Martinez, CC BY-NC-SA 3.0, MBG (2021)) H axillary fascicles of spiciform racemes, E. goetzei (photo: G Baumann, Dressler et al. (2014 a)) I short spiciform raceme of dark red flowers, E. wahlbergii (photo: R Mangelsdorff, Dressler et al. (2014 a)) J-M fruits J immature, weakly falcate, segmented craspedium up to 2 m long, E. rheedei (photo: photographer unknown, Centre for Australian National Biodiversity Research (CANBR), 2000) K immature, segmented, laxly spiralled craspedium up to 120 cm long, E. gigas (photo: R Aguilar CC BY-NC-SA 2.0, Aguilar (2021)) L ripe segmented craspedia breaking up into one-seeded segments with exfoliating epicarp, E. africana (photo: B Eichhorn, Dressler et al. (2014 a)) M ripe unsegmented craspedium, the entire valve breaking away from the persistent replum, E. burkei (photo: M Kriek CC BY-SA 4.0, Ueda (2021) observation 85675968) N, O seeds N one-seeded endocarp segment and small ovoid, flattened seed with elliptic pleurogram, E. africana (photo: B Eichhorn, Dressler et al. (2014 a)) O ripe one-seeded fruit segments with large circular, laterally compressed seeds lacking a pleurogram, E. gigas (photo: J Stevens, Dressler et al. (2014 a)). Scale bars: 1 cm (N, O)." figureDoi="10.3897/phytokeys.205.76790.figure2" httpUri="https://binary.pensoft.net/fig/732697" pageId="0" pageNumber="99">2A</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Entada rheedei habit, vegetative and reproductive structures. A liana with twisted woody stems, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 28499551) B adaxial view of mature leaf, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 11095941) C axillary spikes of open, sessile flowers, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 11095941) D immature pods, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 28499551) E mature pods of collection Schleiben 769 a, Tanzania (photographer unknown)." figureDoi="10.3897/phytokeys.205.76790.figure21" httpUri="https://binary.pensoft.net/fig/732716" pageId="0" pageNumber="99">21A</figureCitation>
), with elevated nectaries at nodes.
<emphasis bold="true" pageId="0" pageNumber="99">Leaves</emphasis>
: rachis 6.4-12.9 cm long, terminating in a bifurcating tendril; pinnae 1-2 pairs per leaf, 5-14 cm long, with 3-5 pairs of opposite leaflets, sometimes terminating in a glandular mucro (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Putative extrafloral nectaries on herbarium specimens of five of the six native Madagascan species of Entada A E. leptostachya, Du Puy et al. M 235 B E. louvelii, Reserves Naturelles RN 1447 C E. pervillei, Service Forestier 10525 _ SF D E. pervillei, Service Forestier 11481 _ SF E E. rheedei, Gautier LG 3153 F E. tuberosa, Jongkind et al. 3264. Scale bars: 2 mm (A-F). Photos: S O'Donnell." figureDoi="10.3897/phytokeys.205.76790.figure4" httpUri="https://binary.pensoft.net/fig/732699" pageId="0" pageNumber="99">4E</figureCitation>
); leaflets 1.8-6.6
<normalizedToken originalValue="×">x</normalizedToken>
1.2-2.9 cm, chartaceous, elliptic to oblanceolate, asymmetric, apex obtuse to acuminate, retuse, base rounded to attenuate, mid-rib above pubescent, lamina glabrous, except below near the base (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genus-wide variation in morphological characters. A-D habit A large woody liana, E. rheedei (photo: B Wursten, Hyde et al. (2021)) B small tree, E. abyssinica (photo: G Baumann, Dressler et al. (2014 a)) C shrub, E. burkei (photo: M Schmidt, Dressler et al. (2014 a, b )) D geoxylic suffrutex, E. dolichorrhachis (photo: M Bingham, Bingham et al. (2021)) E, F leaves E bipinnate leaf with few pinnae, few large leaflets and ending in a bifurcating tendril, E. rheedei (photo: AP Balan, Balan and Predeep (2021)) F bipinnate leaf with many pinnae, many small leaflets and no tendril, E. rangei (photo: A Dreyer, Dressler et al. (2014 a)) G-I inflorescences G terminal panicle of up-turned spikes, E. polyphylla (photo: R Vasquez Martinez, CC BY-NC-SA 3.0, MBG (2021)) H axillary fascicles of spiciform racemes, E. goetzei (photo: G Baumann, Dressler et al. (2014 a)) I short spiciform raceme of dark red flowers, E. wahlbergii (photo: R Mangelsdorff, Dressler et al. (2014 a)) J-M fruits J immature, weakly falcate, segmented craspedium up to 2 m long, E. rheedei (photo: photographer unknown, Centre for Australian National Biodiversity Research (CANBR), 2000) K immature, segmented, laxly spiralled craspedium up to 120 cm long, E. gigas (photo: R Aguilar CC BY-NC-SA 2.0, Aguilar (2021)) L ripe segmented craspedia breaking up into one-seeded segments with exfoliating epicarp, E. africana (photo: B Eichhorn, Dressler et al. (2014 a)) M ripe unsegmented craspedium, the entire valve breaking away from the persistent replum, E. burkei (photo: M Kriek CC BY-SA 4.0, Ueda (2021) observation 85675968) N, O seeds N one-seeded endocarp segment and small ovoid, flattened seed with elliptic pleurogram, E. africana (photo: B Eichhorn, Dressler et al. (2014 a)) O ripe one-seeded fruit segments with large circular, laterally compressed seeds lacking a pleurogram, E. gigas (photo: J Stevens, Dressler et al. (2014 a)). Scale bars: 1 cm (N, O)." figureDoi="10.3897/phytokeys.205.76790.figure2" httpUri="https://binary.pensoft.net/fig/732697" pageId="0" pageNumber="99">2E</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Entada rheedei habit, vegetative and reproductive structures. A liana with twisted woody stems, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 28499551) B adaxial view of mature leaf, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 11095941) C axillary spikes of open, sessile flowers, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 11095941) D immature pods, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 28499551) E mature pods of collection Schleiben 769 a, Tanzania (photographer unknown)." figureDoi="10.3897/phytokeys.205.76790.figure21" httpUri="https://binary.pensoft.net/fig/732716" pageId="0" pageNumber="99">21B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Inflorescence</emphasis>
: a spike, 8-25 cm long, axillary, solitary, or sometimes several spikes from a short shoot, peduncle 1-8.5 cm long, peduncle and rachis puberulous to villose (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Entada rheedei habit, vegetative and reproductive structures. A liana with twisted woody stems, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 28499551) B adaxial view of mature leaf, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 11095941) C axillary spikes of open, sessile flowers, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 11095941) D immature pods, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 28499551) E mature pods of collection Schleiben 769 a, Tanzania (photographer unknown)." figureDoi="10.3897/phytokeys.205.76790.figure21" httpUri="https://binary.pensoft.net/fig/732716" pageId="0" pageNumber="99">21C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Flowers</emphasis>
: cream or greenish, sessile to sub-sessile, staminate or bisexual, with an unpleasant odour; calyx cupular, 0.75-1.2 mm long, shallowly toothed; petals pale green to white, 2.5-3.5 mm long; stamen filaments 2-6.5 mm long, white turning yellow; stigma shallowly cupular.
<emphasis bold="true" pageId="0" pageNumber="99">Fruit</emphasis>
: a gigantic, torulose craspedium, 50-200
<normalizedToken originalValue="×">x</normalizedToken>
7-15 cm, straight to slightly curved, with transverse septa between seeds dividing the fruit into one-seeded segments which, upon ripening, fall from the persistent replum; segments 6.5-7.5 cm long; epicarp and endocarp woody (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genus-wide variation in morphological characters. A-D habit A large woody liana, E. rheedei (photo: B Wursten, Hyde et al. (2021)) B small tree, E. abyssinica (photo: G Baumann, Dressler et al. (2014 a)) C shrub, E. burkei (photo: M Schmidt, Dressler et al. (2014 a, b )) D geoxylic suffrutex, E. dolichorrhachis (photo: M Bingham, Bingham et al. (2021)) E, F leaves E bipinnate leaf with few pinnae, few large leaflets and ending in a bifurcating tendril, E. rheedei (photo: AP Balan, Balan and Predeep (2021)) F bipinnate leaf with many pinnae, many small leaflets and no tendril, E. rangei (photo: A Dreyer, Dressler et al. (2014 a)) G-I inflorescences G terminal panicle of up-turned spikes, E. polyphylla (photo: R Vasquez Martinez, CC BY-NC-SA 3.0, MBG (2021)) H axillary fascicles of spiciform racemes, E. goetzei (photo: G Baumann, Dressler et al. (2014 a)) I short spiciform raceme of dark red flowers, E. wahlbergii (photo: R Mangelsdorff, Dressler et al. (2014 a)) J-M fruits J immature, weakly falcate, segmented craspedium up to 2 m long, E. rheedei (photo: photographer unknown, Centre for Australian National Biodiversity Research (CANBR), 2000) K immature, segmented, laxly spiralled craspedium up to 120 cm long, E. gigas (photo: R Aguilar CC BY-NC-SA 2.0, Aguilar (2021)) L ripe segmented craspedia breaking up into one-seeded segments with exfoliating epicarp, E. africana (photo: B Eichhorn, Dressler et al. (2014 a)) M ripe unsegmented craspedium, the entire valve breaking away from the persistent replum, E. burkei (photo: M Kriek CC BY-SA 4.0, Ueda (2021) observation 85675968) N, O seeds N one-seeded endocarp segment and small ovoid, flattened seed with elliptic pleurogram, E. africana (photo: B Eichhorn, Dressler et al. (2014 a)) O ripe one-seeded fruit segments with large circular, laterally compressed seeds lacking a pleurogram, E. gigas (photo: J Stevens, Dressler et al. (2014 a)). Scale bars: 1 cm (N, O)." figureDoi="10.3897/phytokeys.205.76790.figure2" httpUri="https://binary.pensoft.net/fig/732697" pageId="0" pageNumber="99">2J</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Entada rheedei habit, vegetative and reproductive structures. A liana with twisted woody stems, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 28499551) B adaxial view of mature leaf, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 11095941) C axillary spikes of open, sessile flowers, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 11095941) D immature pods, South Africa (photo: R Taylor CC BY-NC 4.0, Ueda (2021) observation 28499551) E mature pods of collection Schleiben 769 a, Tanzania (photographer unknown)." figureDoi="10.3897/phytokeys.205.76790.figure21" httpUri="https://binary.pensoft.net/fig/732716" pageId="0" pageNumber="99">21D, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Seeds</emphasis>
: subcircular, laterally compressed, 5
<normalizedToken originalValue="×">x</normalizedToken>
3.5-5 cm, hard, brown, pleurogram lacking.
</paragraph>
<caption doi="10.3897/phytokeys.205.76790.figure21" httpUri="https://binary.pensoft.net/fig/732716" pageId="0" pageNumber="99" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Figure 21.</emphasis>
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada rheedei" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="rheedei">
<emphasis italics="true" pageId="0" pageNumber="99">Entada rheedei</emphasis>
</taxonomicName>
habit, vegetative and reproductive structures.
<emphasis bold="true" pageId="0" pageNumber="99">A</emphasis>
liana with twisted woody stems, South Africa (photo: R Taylor
<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC 4.0</emphasis>
,
<bibRefCitation author="Ueda, K" journalOrPublisher="Acta Phytotaxonomica et Geobotanica" pageId="0" pageNumber="99" publicationUrl="iNaturalist.org" refId="B59" refString="Ueda, K, 2021. iNaturalist Research-grade Observations. iNaturalist.org" title="iNaturalist Research-grade Observations." url="iNaturalist.org" year="2021">Ueda (2021)</bibRefCitation>
observation 28499551)
<emphasis bold="true" pageId="0" pageNumber="99">B</emphasis>
adaxial view of mature leaf, South Africa (photo: R Taylor
<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC 4.0</emphasis>
,
<bibRefCitation author="Ueda, K" journalOrPublisher="Acta Phytotaxonomica et Geobotanica" pageId="0" pageNumber="99" publicationUrl="iNaturalist.org" refId="B59" refString="Ueda, K, 2021. iNaturalist Research-grade Observations. iNaturalist.org" title="iNaturalist Research-grade Observations." url="iNaturalist.org" year="2021">Ueda (2021)</bibRefCitation>
observation 11095941)
<emphasis bold="true" pageId="0" pageNumber="99">C</emphasis>
axillary spikes of open, sessile flowers, South Africa (photo: R Taylor
<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC 4.0</emphasis>
,
<bibRefCitation author="Ueda, K" journalOrPublisher="Acta Phytotaxonomica et Geobotanica" pageId="0" pageNumber="99" publicationUrl="iNaturalist.org" refId="B59" refString="Ueda, K, 2021. iNaturalist Research-grade Observations. iNaturalist.org" title="iNaturalist Research-grade Observations." url="iNaturalist.org" year="2021">Ueda (2021)</bibRefCitation>
observation 11095941)
<emphasis bold="true" pageId="0" pageNumber="99">D</emphasis>
immature pods, South Africa (photo: R Taylor
<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC 4.0</emphasis>
,
<bibRefCitation author="Ueda, K" journalOrPublisher="Acta Phytotaxonomica et Geobotanica" pageId="0" pageNumber="99" publicationUrl="iNaturalist.org" refId="B59" refString="Ueda, K, 2021. iNaturalist Research-grade Observations. iNaturalist.org" title="iNaturalist Research-grade Observations." url="iNaturalist.org" year="2021">Ueda (2021)</bibRefCitation>
observation 28499551)
<emphasis bold="true" pageId="0" pageNumber="99">E</emphasis>
mature pods of collection Schleiben 769a, Tanzania (photographer unknown).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>