1333 lines
290 KiB
XML
1333 lines
290 KiB
XML
<document ID-DOI="10.35929/RSZ.0083" ID-ISSN="0035-418" ID-Zenodo-Dep="7761487" checkinTime="1679514280647" checkinUser="valdenar" docAuthor="Schwendinger, Peter J., Huber, Siegfried, Lehmann-Graber, Christina, Ono, Hirotsugu, Aung, Mu Mu & Hongpadharakiree, Komsan" docDate="2022" docId="03E18D717244036FB106F9F85699FCAB" docLanguage="en" docName="RevSuisseZool.129.2.375-424.pdf" docOrigin="Revue suisse de Zoologie 129 (2)" docSource="http://dx.doi.org/10.35929/rsz.0083" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Liphistius birmanicus Thorell 1897" docType="treatment" docVersion="3" lastPageNumber="380" masterDocId="FFD8F50972460369B239FFEC5305FF92" masterDocTitle="A taxonomic revision of the Liphistius birmanicus-group (Araneae: Liphistiidae) with the description of five new species" masterLastPageNumber="424" masterPageNumber="375" pageNumber="376" updateTime="1679571001943" updateUser="valdenar">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
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<mods:titleInfo>
|
||
<mods:title>A taxonomic revision of the Liphistius birmanicus-group (Araneae: Liphistiidae) with the description of five new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
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<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Schwendinger, Peter J.</mods:namePart>
|
||
<mods:affiliation>Muséum d’histoire naturelle de Genève, C. P. 6404, CH- 1211 Genève 6, Switzerland</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Huber, Siegfried</mods:namePart>
|
||
<mods:affiliation>Ottenbohlstrasse 12, D- 88690 Mühlhofen, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Lehmann-Graber, Christina</mods:namePart>
|
||
<mods:affiliation>Muséum d’histoire naturelle de Genève, C. P. 6404, CH- 1211 Genève 6, Switzerland</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Ono, Hirotsugu</mods:namePart>
|
||
<mods:affiliation>Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Aung, Mu Mu</mods:namePart>
|
||
<mods:affiliation>Forest Research Institute, Forest Department, Ministry of Natural Resources and Environmental Conservation, Yezin, Nay Pyi Taw, Myanmar</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Hongpadharakiree, Komsan</mods:namePart>
|
||
<mods:affiliation>Sirinart Rajini Mangrove Ecosystem Learning Center, Pranburi, Prachuab Khiri Khan Province, Thailand</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Revue suisse de Zoologie</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2022</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2022-10-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>129</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>375</mods:start>
|
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<mods:end>424</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.35929/rsz.0083</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.35929/RSZ.0083</mods:identifier>
|
||
<mods:identifier type="ISSN">0035-418</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7761487</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7761493" ID-Zenodo-Dep="7761493" LSID="urn:lsid:plazi:treatment:03E18D717244036FB106F9F85699FCAB" httpUri="http://treatment.plazi.org/id/03E18D717244036FB106F9F85699FCAB" lastPageId="6" lastPageNumber="380" pageId="2" pageNumber="376">
|
||
<subSubSection box="[831,1084,1555,1579]" pageId="2" pageNumber="376" type="nomenclature">
|
||
<paragraph blockId="2.[831,1437,1555,1772]" box="[831,1084,1555,1579]" pageId="2" pageNumber="376">
|
||
<emphasis bold="true" box="[831,1084,1555,1579]" pageId="2" pageNumber="376">
|
||
The
|
||
<taxonomicName ID-CoL="72J64" authorityName="Thorell" authorityYear="1897" box="[883,1007,1555,1579]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis bold="true" box="[883,1007,1555,1579]" italics="true" pageId="2" pageNumber="376">birmanicus</emphasis>
|
||
</taxonomicName>
|
||
-group
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="378" pageId="2" pageNumber="376" type="description">
|
||
<paragraph blockId="2.[831,1437,1555,1772]" pageId="2" pageNumber="376">
|
||
When this species group was defined by Schwendinger in 1998 it contained only three species (
|
||
<emphasis italics="true" pageId="2" pageNumber="376">
|
||
<taxonomicName authorityName="Thorell" authorityYear="1897" box="[1278,1433,1620,1643]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="birmanicus">L. birmanicus</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Platnick & Sedgwick" authorityYear="1984" box="[831,931,1652,1675]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="lordae">L. lordae</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authorityName="Schwendinger" authorityYear="1998" box="[988,1066,1652,1675]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="lahu">
|
||
<emphasis box="[988,1066,1652,1675]" italics="true" pageId="2" pageNumber="376">L. lahu</emphasis>
|
||
</taxonomicName>
|
||
). As four additional species were added later and five are newly described here, a refined diagnosis and description of this group is necessary and given in the following.
|
||
</paragraph>
|
||
<paragraph blockId="2.[831,1437,1796,2077]" lastBlockId="3.[151,756,258,993]" lastPageId="3" lastPageNumber="377" pageId="2" pageNumber="376">
|
||
<emphasis bold="true" box="[831,951,1796,1819]" pageId="2" pageNumber="376">Diagnosis:</emphasis>
|
||
Essentially distinguished by the following four characters: (1) Contrategular area below (proximal of) tegulum completely unpigmented (e.g.
|
||
<figureCitation box="[1297,1379,1861,1884]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="2" pageNumber="376">Figs 8F</figureCitation>
|
||
, H-I, 13F-G, 21K, M, 23L-M). In males of all other species groups this area is more or less extensively pigmented. (2) A more or less distinct (least so in
|
||
<taxonomicName box="[1270,1396,1957,1980]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[1270,1396,1957,1980]" italics="true" pageId="2" pageNumber="376">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="2" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<figureCitation box="[895,993,1989,2012]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="2" pageNumber="376">Fig. 23F</figureCitation>
|
||
) proximal ledge on retrodorsal side of contrategulum (e.g.
|
||
<figureCitation box="[1066,1137,2021,2044]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="2" pageNumber="376">Fig. 4</figureCitation>
|
||
A-C, G). A similar ledge is present only in
|
||
<taxonomicName authority="Schiodte, 1849" authorityName="Schiodte" authorityYear="1849" box="[1063,1381,2054,2077]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="desultor">
|
||
<emphasis box="[1063,1190,2054,2077]" italics="true" pageId="2" pageNumber="376">L. desultor</emphasis>
|
||
<bibRefCitation author="Schiodte J. C." box="[1206,1381,2054,2077]" pageId="2" pageNumber="376" pagination="617 - 624" refId="ref38042" refString="Schiodte J. C. 1849. Om en afvigende Slaegt af Spindlernes Orden. Naturhistorisk Tidsskrift 2: 617 - 624." type="journal article" year="1849">Schiödte, 1849</bibRefCitation>
|
||
</taxonomicName>
|
||
, but there it is situated medially, not proximally, and its most prominent part is on the proventral side, not on the retroventral side (
|
||
<bibRefCitation author="Schwendinger P. J. & Syuhadah N. & Lehmann-Graber C. & Price L. & Huber S. & Hashim R. & Bhassu S. & Monod L." box="[399,705,323,346]" pageId="3" pageNumber="377" pagination="321 - 353" refId="ref38357" refString="Schwendinger P. J., Syuhadah N., Lehmann-Graber C., Price L., Huber S., Hashim R., Bhassu S., Monod L. 2019. A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in Peninsular Malaysia; part 2. Revue suisse de Zoologie 126 (2): 321 - 353." type="journal article" year="2019">
|
||
Schwendinger
|
||
<emphasis box="[567,620,323,346]" italics="true" pageId="3" pageNumber="377">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
: fig. 5F-G, I-K). (3) Embolus proper narrowly divided, with a longer sclerotised part and a distinctly shorter membranous part lying close to each other (e.g.
|
||
<figureCitation captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4</figureCitation>
|
||
C-D, G). This is also found in the
|
||
<taxonomicName authorityName="Platnick & Sedgwick" authorityYear="1984" box="[585,685,452,475]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="bristowei">
|
||
<emphasis box="[585,685,452,475]" italics="true" pageId="3" pageNumber="377">bristowei</emphasis>
|
||
</taxonomicName>
|
||
- (e.g.
|
||
<bibRefCitation author="Schwendinger P. J." box="[151,388,485,508]" pageId="3" pageNumber="377" pagination="331 - 351" refId="ref38065" refString="Schwendinger P. J. 1990. On the spider genus Liphistius (Araneae: Mesothelae) in Thailand and Burma. Zoologica Scripta 19 (3): 331 - 351." type="journal article" year="1990">Schwendinger, 1990</bibRefCitation>
|
||
: figs 57-59),
|
||
<taxonomicName authorityName="Abraham" authorityYear="1923" box="[563,680,485,508]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis box="[563,680,485,508]" italics="true" pageId="3" pageNumber="377">malayanus</emphasis>
|
||
</taxonomicName>
|
||
- (e.g.
|
||
<bibRefCitation author="Schwendinger P. J." box="[151,390,517,540]" pageId="3" pageNumber="377" pagination="391 - 445" refId="ref38223" refString="Schwendinger P. J. 2017. A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1. Revue suisse de Zoologie 124 (2): 391 - 445." type="journal article" year="2017">Schwendinger, 2017</bibRefCitation>
|
||
: fig. 3B, D, F, H),
|
||
<emphasis box="[665,740,517,540]" italics="true" pageId="3" pageNumber="377">tioman</emphasis>
|
||
-,
|
||
<emphasis box="[151,220,549,572]" italics="true" pageId="3" pageNumber="377">linang</emphasis>
|
||
- and
|
||
<taxonomicName authorityName="Abraham" authorityYear="1923" box="[300,402,549,572]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="batuensis">
|
||
<emphasis box="[300,402,549,572]" italics="true" pageId="3" pageNumber="377">batuensis</emphasis>
|
||
</taxonomicName>
|
||
-group, whereas in the
|
||
<emphasis box="[688,747,549,572]" italics="true" pageId="3" pageNumber="377">trang</emphasis>
|
||
- group the embolus is widely divided, with both parts clearly apart from each other (
|
||
<bibRefCitation author="Schwendinger P. J." box="[518,753,614,637]" pageId="3" pageNumber="377" pagination="331 - 351" refId="ref38065" refString="Schwendinger P. J. 1990. On the spider genus Liphistius (Araneae: Mesothelae) in Thailand and Burma. Zoologica Scripta 19 (3): 331 - 351." type="journal article" year="1990">Schwendinger, 1990</bibRefCitation>
|
||
: figs 60-62;
|
||
<bibRefCitation author="Schwendinger P. J." box="[279,505,646,669]" pageId="3" pageNumber="377" pagination="391 - 445" refId="ref38223" refString="Schwendinger P. J. 2017. A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1. Revue suisse de Zoologie 124 (2): 391 - 445." type="journal article" year="2017">Schwendinger, 2017</bibRefCitation>
|
||
: fig. 3A, C, E, G). (4) Poreplate with a pair of more or less developed (least so in
|
||
<taxonomicName authorityName="Yu, Zhang & Zhang" authorityYear="2021" box="[182,294,711,734]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[182,294,711,734]" italics="true" pageId="3" pageNumber="377">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[353,469,711,734]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[353,469,711,734]" italics="true" pageId="3" pageNumber="377">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[479,568,711,734]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[581,753,711,734]" pageId="3" pageNumber="377" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[621,671,711,734]" italics="true" pageId="3" pageNumber="377">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 4 and
|
||
<figureCitation box="[280,364,744,767]" captionStart="Fig" captionStartId="40.[151,186,1942,1963]" captionTargetBox="[155,1436,262,1898]" captionTargetId="figure-107@40.[151,1439,257,1908]" captionTargetPageId="40" captionText="Fig. 22. Liphistius cupreus sp. nov., vulval plates of ten females (all taken from exuviae), dorsal view (E with uterus externus attached, H, K, N), ventral view (A-D, I-J, L-M, O), ventral and slightly anterior view (P), posterior view (G), and isolated uterus externus (F). (A) Moult of 10.II.2019. (B) Moult of 12.V.2020. (C) Moult of 5.VII.2019. (D-F) Moult of 22.XI.2020. (G) Moult of VI.2019. (H-I) Moult of 13.II.2021. (J) Allotype; moult of 19.I.2019. (K-L) Moult of 14.II.2021. (M) Moult of 1.V.2021. (N-P) Moult of 23.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761549" httpUri="https://zenodo.org/record/7761549/files/figure.png" pageId="3" pageNumber="377">Fig. 22</figureCitation>
|
||
) anterior lobes (e.g.
|
||
<figureCitation box="[615,687,744,767]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="3" pageNumber="377">Fig. 5</figureCitation>
|
||
). The females of several species in the
|
||
<taxonomicName authorityName="Abraham" authorityYear="1923" box="[524,640,776,799]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis box="[524,640,776,799]" italics="true" pageId="3" pageNumber="377">malayanus</emphasis>
|
||
</taxonomicName>
|
||
-,
|
||
<emphasis box="[666,740,776,799]" italics="true" pageId="3" pageNumber="377">tioman</emphasis>
|
||
-,
|
||
<emphasis box="[151,220,808,831]" italics="true" pageId="3" pageNumber="377">linang</emphasis>
|
||
- and
|
||
<taxonomicName authorityName="Abraham" authorityYear="1923" box="[305,407,808,831]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="batuensis">
|
||
<emphasis box="[305,407,808,831]" italics="true" pageId="3" pageNumber="377">batuensis</emphasis>
|
||
</taxonomicName>
|
||
-group also have pronounced anterior lobes on their poreplates, but these co-occur with either a very large receptacular cluster, a very large CDO or a reduced sclerotisation of the posterior part of the poreplate and/or of the posterior stalk (see various illustrations in
|
||
<bibRefCitation author="Schwendinger P. J." box="[311,534,970,993]" pageId="3" pageNumber="377" pagination="391 - 445" refId="ref38223" refString="Schwendinger P. J. 2017. A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1. Revue suisse de Zoologie 124 (2): 391 - 445." type="journal article" year="2017">Schwendinger, 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,757,1018,2077]" lastBlockId="4.[151,756,258,953]" lastPageId="4" lastPageNumber="378" pageId="3" pageNumber="377">
|
||
<emphasis bold="true" box="[151,441,1018,1041]" pageId="3" pageNumber="377">Additions to description:</emphasis>
|
||
Medium-sized to very large species (CL of males 4.92-12.67, CW 4.56-11.47). All males and the females of most species with a brown to almost black body colouration, often with annulated legs (
|
||
<figureCitation box="[330,404,1147,1170]" captionStart="Fig" captionStartId="8.[151,186,1932,1953]" captionTargetBox="[155,1431,783,1902]" captionTargetId="figure-159@8.[151,1436,777,1910]" captionTargetPageId="8" captionText="Fig. 2. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius tung sp. nov., male holotype.(B) Liphistius pyinoolwin, male (matured 1.X.2014) from the Anisakan Falls; note exuviae of ectoparasitic mites (Ljunghia sp.) on the opisthosoma. (C) Liphistius platnicki sp. nov., female paratype (moulted 10.III.2021) from near Loi Kan Village. (D, F) Liphistius cupreus sp. nov., male paratype from near Kong Nyaung Village. (E) Same species, different male paratype. (G) Same species, female paratype (moulted 13.II.2021). (H-I) Same species, different female paratype (moulted 21.V.2021). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761491" httpUri="https://zenodo.org/record/7761491/files/figure.png" pageId="3" pageNumber="377">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation box="[421,433,1147,1170]" captionStart="Fig" captionStartId="9.[151,186,1970,1991]" captionTargetBox="[160,1426,651,1915]" captionTargetId="figure-160@9.[151,1436,641,1924]" captionTargetPageId="9" captionText="Fig. 3. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius ferox sp. nov., male paratype (matured 1.IX.2014) from Pathi Village. (B) Same species, female paratype from Thandaung Gyi. (C) Liphistius pinlaung, female from near Tong Htiwaw Temple. (D) Liphistius birmanicus from Yado, male (matured 20.IX.2017). (E) Same species, large female n°1. (F, H) Same species, large female n°2. (G) Same species, small female. (I) Same species, large female n°3. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761495" httpUri="https://zenodo.org/record/7761495/files/figure.png" pageId="3" pageNumber="377">3</figureCitation>
|
||
A-D); only in
|
||
<taxonomicName authorityName="Thorell" authorityYear="1897" box="[602,756,1147,1170]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis box="[602,756,1147,1170]" italics="true" pageId="3" pageNumber="377">L. birmanicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu & Li" authorityYear="2019" box="[200,305,1179,1202]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="hpruso">
|
||
<emphasis box="[200,305,1179,1202]" italics="true" pageId="3" pageNumber="377">L. hpruso</emphasis>
|
||
</taxonomicName>
|
||
females with yellow or orange-coloured areas on carapace and proximal portions of legs and palps (
|
||
<figureCitation box="[234,308,1244,1267]" captionStart="Fig" captionStartId="9.[151,186,1970,1991]" captionTargetBox="[160,1426,651,1915]" captionTargetId="figure-160@9.[151,1436,641,1924]" captionTargetPageId="9" captionText="Fig. 3. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius ferox sp. nov., male paratype (matured 1.IX.2014) from Pathi Village. (B) Same species, female paratype from Thandaung Gyi. (C) Liphistius pinlaung, female from near Tong Htiwaw Temple. (D) Liphistius birmanicus from Yado, male (matured 20.IX.2017). (E) Same species, large female n°1. (F, H) Same species, large female n°2. (G) Same species, small female. (I) Same species, large female n°3. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761495" httpUri="https://zenodo.org/record/7761495/files/figure.png" pageId="3" pageNumber="377">Fig. 3</figureCitation>
|
||
E-I). Males with medium-sized tibial apophysis, triangular in ventral view, usually not much set back from distal margin of palpal tibia (e.g.
|
||
<figureCitation captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="3" pageNumber="377">Fig. 6K, N</figureCitation>
|
||
; except in
|
||
<taxonomicName box="[355,436,1341,1364]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="tung" status="sp. nov.">
|
||
<emphasis box="[355,436,1341,1364]" italics="true" pageId="3" pageNumber="377">L. tung</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[448,538,1341,1364]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<figureCitation box="[554,641,1341,1364]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="3" pageNumber="377">Fig. 8B</figureCitation>
|
||
), carrying four long tapering apical megaspines (e.g.
|
||
<figureCitation box="[655,744,1374,1397]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4F</figureCitation>
|
||
). Paracymbium rather short, moderately deep, with indistinctly conical, almost plane distal side (e.g. 4H-J), or with distinctly conical distal side (only in
|
||
<taxonomicName box="[151,264,1503,1526]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[151,264,1503,1526]" italics="true" pageId="3" pageNumber="377">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[272,359,1503,1526]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName box="[371,493,1503,1526]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[371,493,1503,1526]" italics="true" pageId="3" pageNumber="377">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[502,586,1503,1526]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authorityName="Yu, Zhang & Zhang" authorityYear="2021" box="[640,749,1503,1526]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[640,749,1503,1526]" italics="true" pageId="3" pageNumber="377">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation box="[151,258,1536,1559]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="3" pageNumber="377">Figs 21A</figureCitation>
|
||
,
|
||
<figureCitation box="[273,322,1536,1559]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="3" pageNumber="377">23A</figureCitation>
|
||
,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[337,513,1536,1559]" pageId="3" pageNumber="377" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[378,430,1536,1559]" italics="true" pageId="3" pageNumber="377">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3D-F), and with (e.g.
|
||
<figureCitation box="[215,305,1568,1591]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="3" pageNumber="377">Fig. 6L</figureCitation>
|
||
) or without (e.g. 10A) a retrolateralproximal heel; the latter widely (e.g.
|
||
<figureCitation box="[575,646,1600,1623]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Figs 4</figureCitation>
|
||
H-I, 15G) or narrowly rounded (e.g.
|
||
<figureCitation box="[450,539,1633,1656]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="3" pageNumber="377">Figs 6L</figureCitation>
|
||
,
|
||
<figureCitation box="[554,599,1633,1656]" captionStart="Fig" captionStartId="35.[151,186,1288,1309]" captionTargetBox="[156,1431,261,1232]" captionTargetId="figure-280@35.[151,1436,257,1238]" captionTargetPageId="35" captionText="Fig. 19. Liphistius pyinoolwin, palps of two males: matured 16.X.2014 (A-E), matured 24.IX.2014 (F). (A) Left palpal organ, distal view (dorsal side up). (B) Left embolus complex and dorsal part of contrategulum, distal and slightly proventral view. (C) Left palpal organ and distal margin of tarsus, prolateral view. (D) Left palpal organ, ventral and slightly proximal view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Distal part of left tibia and proximal part of tarsus, ventral view. (F) Distal margin of right tibia and proximal part of tarsus, ventral view. Scale lines 1.0 mm (A-D; E-F). Abbreviation: l - lobate prolateral protrusion on base of embolus complex." figureDoi="http://doi.org/10.5281/zenodo.7761541" httpUri="https://zenodo.org/record/7761541/files/figure.png" pageId="3" pageNumber="377">19F</figureCitation>
|
||
), in
|
||
<taxonomicName authorityName="Platnick & Sedgwick" authorityYear="1984" box="[654,756,1633,1656]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="lordae">
|
||
<emphasis box="[654,756,1633,1656]" italics="true" pageId="3" pageNumber="377">L. lordae</emphasis>
|
||
</taxonomicName>
|
||
heel protruding proximally (
|
||
<figureCitation box="[484,593,1665,1688]" captionStart="Fig" captionStartId="32.[151,186,1942,1963]" captionTargetBox="[159,1427,1125,1893]" captionTargetId="figure-353@32.[151,1436,1117,1903]" captionTargetPageId="32" captionText="Fig. 17. Liphistius lordae, left palps of two males: male n°1 (A-B, D-G), male n°2 (C). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Contrategulum, distal view. (D) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Palpal organ and distal margin of tarsus, prolateral view. (F) Tibial apophysis, retrolateral and slightly proximal view. (G) Proximal part of tarsus, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761535" httpUri="https://zenodo.org/record/7761535/files/figure.png" pageId="3" pageNumber="377">Fig. 17G</figureCitation>
|
||
), in all other cases retrolaterally (e.g.
|
||
<figureCitation box="[424,503,1697,1720]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="3" pageNumber="377">Fig. 6J</figureCitation>
|
||
, L-N); cumulus not or only indistinctly elevated (e.g.
|
||
<figureCitation box="[495,561,1730,1753]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4</figureCitation>
|
||
H-J), or distinctly elevated (only in
|
||
<taxonomicName box="[353,470,1762,1785]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[353,470,1762,1785]" italics="true" pageId="3" pageNumber="377">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[481,572,1762,1785]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName box="[587,714,1762,1785]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[587,714,1762,1785]" italics="true" pageId="3" pageNumber="377">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authorityName="Yu, Zhang & Zhang" authorityYear="2021" box="[253,364,1795,1818]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[253,364,1795,1818]" italics="true" pageId="3" pageNumber="377">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation box="[380,483,1795,1818]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="3" pageNumber="377">Figs 21B</figureCitation>
|
||
,
|
||
<figureCitation box="[496,523,1795,1818]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="3" pageNumber="377">23</figureCitation>
|
||
A-B,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[584,752,1795,1818]" pageId="3" pageNumber="377" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[623,673,1795,1818]" italics="true" pageId="3" pageNumber="377">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3D-F), carrying several long, strong bristles. Ventral side of palpal tarsus with a distinct subdistal suture (
|
||
<figureCitation box="[159,262,1892,1915]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="3" pageNumber="377">Figs 21A</figureCitation>
|
||
,
|
||
<figureCitation box="[275,324,1892,1915]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="3" pageNumber="377">23A</figureCitation>
|
||
), with a subdistal trench or without any depression there; distal margin of palpal tarsus widely but shallowly invaginated (e.g.
|
||
<figureCitation box="[511,602,1956,1979]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4G</figureCitation>
|
||
). Subtegulum with (e.g.
|
||
<figureCitation box="[269,407,1989,2012]" captionStart="Fig" captionStartId="19.[151,186,1745,1766]" captionTargetBox="[156,1430,263,1692]" captionTargetId="figure-155@19.[151,1436,257,1698]" captionTargetPageId="19" captionText="Fig. 10. Liphistius ferox sp. nov., left palps of four males; holotype (A-B, E-K), male from Naw Bubaw, matured IV.2019 (C, M), male from Thandaung Gyi, matured 7.X.2014 (D), male from Thandaung Gyi, matured 18.IX.2016 (L). (A) Tarsus and palpal organ, ventral view. (B) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (C-D) Embolus complex, ventral view. (F) Embolus proper, prodorsal view. (G) Palpal organ, retroventral view. (H) Palpal organ, distal view (dorsal side up). (I) Dorsal part of contrategulum, distal and slightly prolateral view. (J) Distal part of tarsus, ventral view. (K-M) Same, retrolateral and slightly proximal view. Scale lines 1.0 mm (A; B, F-G; C-E; H-I; J; K-M). Abbreviations: tp - triangular distal protrusion of longitudinal rib on sclerotised part of embolus proper; so - angular retrolateral side of sperm duct opening." figureDoi="http://doi.org/10.5281/zenodo.7761515" httpUri="https://zenodo.org/record/7761515/files/figure.png" pageId="3" pageNumber="377">Fig. 10A, G</figureCitation>
|
||
) or without (e.g.
|
||
<figureCitation captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="3" pageNumber="377">Fig. 21A, K, M</figureCitation>
|
||
) a small apophysis, never with a large one. Tegulum well-developed and wide; its distal margin not elevated in most species (e.g.
|
||
<figureCitation box="[1084,1152,258,281]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="3" pageNumber="377">Fig. 8</figureCitation>
|
||
F-G), only in
|
||
<taxonomicName authorityName="Yu, Zhang & Zhang" authorityYear="2021" box="[1321,1437,258,281]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[1321,1437,258,281]" italics="true" pageId="3" pageNumber="377">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[879,1003,290,313]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[879,1003,290,313]" italics="true" pageId="3" pageNumber="377">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1012,1097,290,313]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
distinctly elevated into a short wide keel (
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[950,1115,322,345]" pageId="3" pageNumber="377" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[990,1038,322,345]" italics="true" pageId="3" pageNumber="377">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3A-B, D-E and
|
||
<figureCitation box="[1342,1419,322,345]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="3" pageNumber="377">Fig. 23</figureCitation>
|
||
I- M); proximal margin of tegulum with a wide, finely to coarsely dentate edge, in most species more or less bent and overhanging the unpigmented membranous contrategular area below it (e.g.
|
||
<figureCitation box="[1180,1259,450,473]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="3" pageNumber="377">Fig. 6F</figureCitation>
|
||
) or adpressed to it (e.g.
|
||
<figureCitation box="[914,1014,482,505]" captionStart="Fig" captionStartId="19.[151,186,1745,1766]" captionTargetBox="[156,1430,263,1692]" captionTargetId="figure-155@19.[151,1436,257,1698]" captionTargetPageId="19" captionText="Fig. 10. Liphistius ferox sp. nov., left palps of four males; holotype (A-B, E-K), male from Naw Bubaw, matured IV.2019 (C, M), male from Thandaung Gyi, matured 7.X.2014 (D), male from Thandaung Gyi, matured 18.IX.2016 (L). (A) Tarsus and palpal organ, ventral view. (B) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (C-D) Embolus complex, ventral view. (F) Embolus proper, prodorsal view. (G) Palpal organ, retroventral view. (H) Palpal organ, distal view (dorsal side up). (I) Dorsal part of contrategulum, distal and slightly prolateral view. (J) Distal part of tarsus, ventral view. (K-M) Same, retrolateral and slightly proximal view. Scale lines 1.0 mm (A; B, F-G; C-E; H-I; J; K-M). Abbreviations: tp - triangular distal protrusion of longitudinal rib on sclerotised part of embolus proper; so - angular retrolateral side of sperm duct opening." figureDoi="http://doi.org/10.5281/zenodo.7761515" httpUri="https://zenodo.org/record/7761515/files/figure.png" pageId="3" pageNumber="377">Fig. 10B</figureCitation>
|
||
), only in
|
||
<taxonomicName box="[1125,1251,482,505]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[1125,1251,482,505]" italics="true" pageId="3" pageNumber="377">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1260,1347,482,505]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
straight and strongly salient (
|
||
<figureCitation box="[1089,1206,514,537]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="3" pageNumber="377">Fig. 23M</figureCitation>
|
||
). Pigmented bridge between tegulum and contrategulum on retrodorsal side of palpal organ well developed. Contrategulum carrying a medium-sized, more or less conical proventral process with a rounded (e.g.
|
||
<figureCitation box="[1281,1348,643,666]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4</figureCitation>
|
||
A-C) or widely truncate apex (
|
||
<figureCitation box="[1078,1214,675,698]" captionStart="Fig" captionStartId="32.[151,186,1942,1963]" captionTargetBox="[159,1427,1125,1893]" captionTargetId="figure-353@32.[151,1436,1117,1903]" captionTargetPageId="32" captionText="Fig. 17. Liphistius lordae, left palps of two males: male n°1 (A-B, D-G), male n°2 (C). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Contrategulum, distal view. (D) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Palpal organ and distal margin of tarsus, prolateral view. (F) Tibial apophysis, retrolateral and slightly proximal view. (G) Proximal part of tarsus, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761535" httpUri="https://zenodo.org/record/7761535/files/figure.png" pageId="3" pageNumber="377">Fig. 17A, C</figureCitation>
|
||
), in most species at a roughly 45-90° angle to the axis of the palpal organ (e.g.
|
||
<figureCitation box="[887,974,739,762]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4G</figureCitation>
|
||
), in
|
||
<taxonomicName authorityName="Thorell" authorityYear="1897" box="[1024,1174,739,762]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis box="[1024,1174,739,762]" italics="true" pageId="3" pageNumber="377">L. birmanicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu & Li" authorityYear="2019" box="[1230,1355,739,762]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="pinlaung">
|
||
<emphasis box="[1230,1355,739,762]" italics="true" pageId="3" pageNumber="377">L. pinlaung</emphasis>
|
||
</taxonomicName>
|
||
distaddirected, almost parallel to axis of palpal organ (
|
||
<figureCitation captionStart="Fig" captionStartId="26.[151,186,1411,1432]" captionTargetBox="[157,1431,263,1314]" captionTargetId="figure-229@26.[151,1436,257,1320]" captionTargetPageId="26" captionText="Fig. 13. Liphistius birmanicus, left palps of two males from the type locality; male matured 27.I.2016 (A-B), male matured 15.XI.2014 (C-H). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum, distal and slightly prolateral view. (C) Distal part of tarsus, prodorsal view; knob-shaped process on distodorsal corner indicated by short arrow. (D) Palpal organ and distal margin of tarsus, prolateral view. (E) Same, ventral view; bent proximal edge of tegulum indicated by long arrow. (F) Distal part of palpal organ, retroventral and slightly proximal view. (G) Palpal organ, retrolateral view. (H) Proximal part of tarsus, ventral view. Scale line 1.0 mm.Abbreviation: cp - proventral contrategular process." figureDoi="http://doi.org/10.5281/zenodo.7761527" httpUri="https://zenodo.org/record/7761527/files/figure.png" pageId="3" pageNumber="377">Figs 13D, F</figureCitation>
|
||
,
|
||
<figureCitation box="[925,1008,803,826]" captionStart="Fig" captionStartId="30.[151,186,1936,1957]" captionTargetBox="[155,1431,946,1889]" captionTargetId="figure-412@30.[151,1436,941,1894]" captionTargetPageId="30" captionText="Fig. 15. Liphistius pinlaung, left palps of two males: matured 23.XI.2014 (A-B), matured 4.VII.2015 (C-G). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Palpal organ and distal margin of tarsus, prolateral view. (D) Palpal organ, ventral view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of palpal organ, retroventral view. (F) Tegulum, retrolateral view. (G) Proximal part of tarsus, ventral view.All to same scale; scale line 1.0 mm. Abbreviation: cp - proventral process of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761531" httpUri="https://zenodo.org/record/7761531/files/figure.png" pageId="3" pageNumber="377">15C, E</figureCitation>
|
||
); distal edge of contrategulum narrow (e.g.
|
||
<figureCitation box="[891,990,835,858]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="3" pageNumber="377">Fig. 21E</figureCitation>
|
||
) to wide (e.g.
|
||
<figureCitation box="[1167,1271,835,858]" captionStart="Fig" captionStartId="35.[151,186,1288,1309]" captionTargetBox="[156,1431,261,1232]" captionTargetId="figure-280@35.[151,1436,257,1238]" captionTargetPageId="35" captionText="Fig. 19. Liphistius pyinoolwin, palps of two males: matured 16.X.2014 (A-E), matured 24.IX.2014 (F). (A) Left palpal organ, distal view (dorsal side up). (B) Left embolus complex and dorsal part of contrategulum, distal and slightly proventral view. (C) Left palpal organ and distal margin of tarsus, prolateral view. (D) Left palpal organ, ventral and slightly proximal view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Distal part of left tibia and proximal part of tarsus, ventral view. (F) Distal margin of right tibia and proximal part of tarsus, ventral view. Scale lines 1.0 mm (A-D; E-F). Abbreviation: l - lobate prolateral protrusion on base of embolus complex." figureDoi="http://doi.org/10.5281/zenodo.7761541" httpUri="https://zenodo.org/record/7761541/files/figure.png" pageId="3" pageNumber="377">Fig. 19A</figureCitation>
|
||
), its prolateral part with a more (e.g.
|
||
<figureCitation box="[1096,1173,867,890]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="3" pageNumber="377">Fig. 6I</figureCitation>
|
||
) or less (e.g.
|
||
<figureCitation box="[1340,1428,867,890]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4G</figureCitation>
|
||
) elevated keel, its prodorsal part evenly arched (e.g. 4A-C), or with an angular (in
|
||
<taxonomicName box="[1169,1283,931,954]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[1169,1283,931,954]" italics="true" pageId="3" pageNumber="377">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1293,1381,931,954]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<figureCitation captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="3" pageNumber="377">Fig. 21A</figureCitation>
|
||
, E-F, H-I) or arched protrusion (in
|
||
<taxonomicName authority=", Yu et al., 2021" authorityName="Yu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[1278,1397,963,986]" italics="true" pageId="3" pageNumber="377">L. nabang,</emphasis>
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." pageId="3" pageNumber="377" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[831,881,995,1018]" italics="true" pageId="3" pageNumber="377">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 3B); retrodorsal side of contrategulum with a pronounced (e.g.
|
||
<figureCitation box="[1106,1171,1027,1050]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4</figureCitation>
|
||
A-C, G) or rather small (only in
|
||
<taxonomicName box="[931,1047,1060,1083]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[931,1047,1060,1083]" italics="true" pageId="3" pageNumber="377">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1058,1148,1060,1083]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName box="[1163,1289,1060,1083]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[1163,1289,1060,1083]" italics="true" pageId="3" pageNumber="377">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1300,1387,1060,1083]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName box="[831,948,1092,1115]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[831,948,1092,1115]" italics="true" pageId="3" pageNumber="377">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation box="[967,1052,1092,1115]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="3" pageNumber="377">Figs 21</figureCitation>
|
||
E-F, H-I, N, 23F, H-I,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." pageId="3" pageNumber="377" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[1369,1423,1092,1115]" italics="true" pageId="3" pageNumber="377">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3A-B) proximal ledge; contrategular area proximal of tegulum completely unpigmented (e.g.
|
||
<figureCitation captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="3" pageNumber="377">Fig. 8</figureCitation>
|
||
F-I). Para-embolic plate short in most species, with (e.g.
|
||
<figureCitation box="[892,960,1220,1243]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="3" pageNumber="377">Fig. 6</figureCitation>
|
||
F-H) or without (e.g.
|
||
<figureCitation box="[1215,1284,1220,1243]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="3" pageNumber="377">Fig. 4</figureCitation>
|
||
C-D) a slight indentation between it and keel-shaped retroventral edge of embolus complex; in
|
||
<taxonomicName box="[1110,1191,1284,1307]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="tung" status="sp. nov.">
|
||
<emphasis box="[1110,1191,1284,1307]" italics="true" pageId="3" pageNumber="377">L. tung</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1201,1289,1284,1307]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName box="[1349,1436,1284,1307]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
|
||
<emphasis box="[1349,1436,1284,1307]" italics="true" pageId="3" pageNumber="377">L. ferox</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[831,923,1316,1339]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
para-embolic plate long and tounge-shaped (
|
||
<figureCitation box="[839,930,1348,1371]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="3" pageNumber="377">Figs 8A</figureCitation>
|
||
, I-H, 10A-D). Retrolateral margin of embolus complex unpigmented and overhanging distal margin of tegulum (e.g.
|
||
<figureCitation box="[985,1049,1412,1435]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="3" pageNumber="377">Fig. 8</figureCitation>
|
||
F-G); prolateral margin of embolus complex with a lobate protrusion in
|
||
<taxonomicName authorityName="Xu, Yu, Aung, Yu, Liu, Lwin, Sang & Li" authorityYear="2021" box="[1228,1377,1444,1467]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="pyinoolwin">
|
||
<emphasis box="[1228,1377,1444,1467]" italics="true" pageId="3" pageNumber="377">L. pyinoolwin</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Fig" captionStartId="35.[151,186,1288,1309]" captionTargetBox="[156,1431,261,1232]" captionTargetId="figure-280@35.[151,1436,257,1238]" captionTargetPageId="35" captionText="Fig. 19. Liphistius pyinoolwin, palps of two males: matured 16.X.2014 (A-E), matured 24.IX.2014 (F). (A) Left palpal organ, distal view (dorsal side up). (B) Left embolus complex and dorsal part of contrategulum, distal and slightly proventral view. (C) Left palpal organ and distal margin of tarsus, prolateral view. (D) Left palpal organ, ventral and slightly proximal view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Distal part of left tibia and proximal part of tarsus, ventral view. (F) Distal margin of right tibia and proximal part of tarsus, ventral view. Scale lines 1.0 mm (A-D; E-F). Abbreviation: l - lobate prolateral protrusion on base of embolus complex." figureDoi="http://doi.org/10.5281/zenodo.7761541" httpUri="https://zenodo.org/record/7761541/files/figure.png" pageId="3" pageNumber="377">Fig. 19</figureCitation>
|
||
A-B), unmodified in all other species of the group. Embolus proper narrowly divided; its sclerotised part strengthened by 2-6 ribs reaching apex; only in
|
||
<taxonomicName box="[1353,1436,1541,1564]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
|
||
<emphasis box="[1353,1436,1541,1564]" italics="true" pageId="3" pageNumber="377">L. ferox</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[831,918,1573,1596]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with 2 ribs, one of them carrying a triangular distal protrusion (
|
||
<figureCitation box="[1022,1098,1605,1628]" captionStart="Fig" captionStartId="19.[151,186,1745,1766]" captionTargetBox="[156,1430,263,1692]" captionTargetId="figure-155@19.[151,1436,257,1698]" captionTargetPageId="19" captionText="Fig. 10. Liphistius ferox sp. nov., left palps of four males; holotype (A-B, E-K), male from Naw Bubaw, matured IV.2019 (C, M), male from Thandaung Gyi, matured 7.X.2014 (D), male from Thandaung Gyi, matured 18.IX.2016 (L). (A) Tarsus and palpal organ, ventral view. (B) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (C-D) Embolus complex, ventral view. (F) Embolus proper, prodorsal view. (G) Palpal organ, retroventral view. (H) Palpal organ, distal view (dorsal side up). (I) Dorsal part of contrategulum, distal and slightly prolateral view. (J) Distal part of tarsus, ventral view. (K-M) Same, retrolateral and slightly proximal view. Scale lines 1.0 mm (A; B, F-G; C-E; H-I; J; K-M). Abbreviations: tp - triangular distal protrusion of longitudinal rib on sclerotised part of embolus proper; so - angular retrolateral side of sperm duct opening." figureDoi="http://doi.org/10.5281/zenodo.7761515" httpUri="https://zenodo.org/record/7761515/files/figure.png" pageId="3" pageNumber="377">Fig. 10</figureCitation>
|
||
E-H); in distal view retrolateral side of sclerotised embolus part rounded and concave in most species, only in
|
||
<taxonomicName box="[1105,1191,1669,1692]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
|
||
<emphasis box="[1105,1191,1669,1692]" italics="true" pageId="3" pageNumber="377">L. ferox</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1200,1286,1669,1692]" pageId="3" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
angular (
|
||
<figureCitation captionStart="Fig" captionStartId="19.[151,186,1745,1766]" captionTargetBox="[156,1430,263,1692]" captionTargetId="figure-155@19.[151,1436,257,1698]" captionTargetPageId="19" captionText="Fig. 10. Liphistius ferox sp. nov., left palps of four males; holotype (A-B, E-K), male from Naw Bubaw, matured IV.2019 (C, M), male from Thandaung Gyi, matured 7.X.2014 (D), male from Thandaung Gyi, matured 18.IX.2016 (L). (A) Tarsus and palpal organ, ventral view. (B) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (C-D) Embolus complex, ventral view. (F) Embolus proper, prodorsal view. (G) Palpal organ, retroventral view. (H) Palpal organ, distal view (dorsal side up). (I) Dorsal part of contrategulum, distal and slightly prolateral view. (J) Distal part of tarsus, ventral view. (K-M) Same, retrolateral and slightly proximal view. Scale lines 1.0 mm (A; B, F-G; C-E; H-I; J; K-M). Abbreviations: tp - triangular distal protrusion of longitudinal rib on sclerotised part of embolus proper; so - angular retrolateral side of sperm duct opening." figureDoi="http://doi.org/10.5281/zenodo.7761515" httpUri="https://zenodo.org/record/7761515/files/figure.png" pageId="3" pageNumber="377">Fig. 10H</figureCitation>
|
||
); membranous part of embolus proper distinctly shorter than sclerotised part, at its base a more or less strongly pigmented area with longitudinal or inclined wrinkles (e.g.
|
||
<figureCitation box="[989,1068,1797,1820]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="3" pageNumber="377">Figs 6I</figureCitation>
|
||
,
|
||
<figureCitation box="[1082,1131,1797,1820]" captionStart="Fig" captionStartId="26.[151,186,1411,1432]" captionTargetBox="[157,1431,263,1314]" captionTargetId="figure-229@26.[151,1436,257,1320]" captionTargetPageId="26" captionText="Fig. 13. Liphistius birmanicus, left palps of two males from the type locality; male matured 27.I.2016 (A-B), male matured 15.XI.2014 (C-H). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum, distal and slightly prolateral view. (C) Distal part of tarsus, prodorsal view; knob-shaped process on distodorsal corner indicated by short arrow. (D) Palpal organ and distal margin of tarsus, prolateral view. (E) Same, ventral view; bent proximal edge of tegulum indicated by long arrow. (F) Distal part of palpal organ, retroventral and slightly proximal view. (G) Palpal organ, retrolateral view. (H) Proximal part of tarsus, ventral view. Scale line 1.0 mm.Abbreviation: cp - proventral contrategular process." figureDoi="http://doi.org/10.5281/zenodo.7761527" httpUri="https://zenodo.org/record/7761527/files/figure.png" pageId="3" pageNumber="377">13D</figureCitation>
|
||
). Females with lateral folds of vulval plate well developed, glabrous or carrying few to several hairs; no hairs in genital atrium, and only exceptionally a single hair on poreplate or on posterior stalk (
|
||
<figureCitation box="[897,990,1925,1948]" captionStart="Fig" captionStartId="31.[151,186,1817,1838]" captionTargetBox="[155,1436,267,1763]" captionTargetId="figure-78@31.[151,1440,255,1768]" captionTargetPageId="31" captionText="Fig. 16. Liphistius pinlaung (A-J) and Liphistius sp. (specimen from Kalaw; K-L), vulval plates of seven females (all taken from exuviae), dorsal view (A, D, G, I, K) and ventral view (B-C, E-F, H, J, L). (A-B) Moult of 10.II.2019. (C) Moult of 7.XII.2018. (D-E) Moult of 10.IV.2021. (F) Moult of 21.X.2018. (G-H) Moult of 16.V.2020. (I-J) Moult of 8.III.2020. (K-L) Moult of 15.VIII.2017. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761533" httpUri="https://zenodo.org/record/7761533/files/figure.png" pageId="3" pageNumber="377">Figs 16I</figureCitation>
|
||
,
|
||
<figureCitation box="[1002,1048,1925,1948]" captionStart="Fig" captionStartId="33.[151,186,1998,2019]" captionTargetBox="[154,1432,751,1943]" captionTargetId="figure-212@33.[151,1436,748,1947]" captionTargetPageId="33" captionText="Fig. 18. Liphistius lordae, vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H); all taken from exuviae. (A-B) Female n°1. (C-D) Female n°2. (E-F) Female n°3. (G-H) Female n°4; note pathologically enlarged vesicle on left side of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761539" httpUri="https://zenodo.org/record/7761539/files/figure.png" pageId="3" pageNumber="377">18E</figureCitation>
|
||
). Poreplates mostly wider than long (with only very few exceptions, e.g.
|
||
<figureCitation box="[1259,1364,1957,1980]" captionStart="Fig" captionStartId="40.[151,186,1942,1963]" captionTargetBox="[155,1436,262,1898]" captionTargetId="figure-107@40.[151,1439,257,1908]" captionTargetPageId="40" captionText="Fig. 22. Liphistius cupreus sp. nov., vulval plates of ten females (all taken from exuviae), dorsal view (E with uterus externus attached, H, K, N), ventral view (A-D, I-J, L-M, O), ventral and slightly anterior view (P), posterior view (G), and isolated uterus externus (F). (A) Moult of 10.II.2019. (B) Moult of 12.V.2020. (C) Moult of 5.VII.2019. (D-F) Moult of 22.XI.2020. (G) Moult of VI.2019. (H-I) Moult of 13.II.2021. (J) Allotype; moult of 19.I.2019. (K-L) Moult of 14.II.2021. (M) Moult of 1.V.2021. (N-P) Moult of 23.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761549" httpUri="https://zenodo.org/record/7761549/files/figure.png" pageId="3" pageNumber="377">Fig. 22N</figureCitation>
|
||
), with a more or less strongly invaginated anterior margin (slightly arched only in one documented female of
|
||
<taxonomicName authority=", Yu et al. 2021" authorityName="Yu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis italics="true" pageId="3" pageNumber="377">L. nabang</emphasis>
|
||
,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[930,1099,2054,2077]" pageId="3" pageNumber="377" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[971,1023,2054,2077]" italics="true" pageId="3" pageNumber="377">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 4E-F) carrying a pair of anterior lobes ranging from large and wide (in
|
||
<taxonomicName authorityName="Platnick & Sedgwick" authorityYear="1984" box="[652,750,258,281]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="lordae">
|
||
<emphasis box="[652,750,258,281]" italics="true" pageId="4" pageNumber="378">L. lordae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation box="[151,229,290,313]" captionStart="Fig" captionStartId="33.[151,186,1998,2019]" captionTargetBox="[154,1432,751,1943]" captionTargetId="figure-212@33.[151,1436,748,1947]" captionTargetPageId="33" captionText="Fig. 18. Liphistius lordae, vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H); all taken from exuviae. (A-B) Female n°1. (C-D) Female n°2. (E-F) Female n°3. (G-H) Female n°4; note pathologically enlarged vesicle on left side of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761539" httpUri="https://zenodo.org/record/7761539/files/figure.png" pageId="4" pageNumber="378">Fig. 18</figureCitation>
|
||
) to small and reduced to 1-2 clusters of vesicles (in
|
||
<taxonomicName box="[192,285,322,345]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="cupre">
|
||
<emphasis box="[192,285,322,345]" italics="true" pageId="4" pageNumber="378">L. cupre</emphasis>
|
||
</taxonomicName>
|
||
us sp. nov.,
|
||
<figureCitation box="[425,505,322,345]" captionStart="Fig" captionStartId="40.[151,186,1942,1963]" captionTargetBox="[155,1436,262,1898]" captionTargetId="figure-107@40.[151,1439,257,1908]" captionTargetPageId="40" captionText="Fig. 22. Liphistius cupreus sp. nov., vulval plates of ten females (all taken from exuviae), dorsal view (E with uterus externus attached, H, K, N), ventral view (A-D, I-J, L-M, O), ventral and slightly anterior view (P), posterior view (G), and isolated uterus externus (F). (A) Moult of 10.II.2019. (B) Moult of 12.V.2020. (C) Moult of 5.VII.2019. (D-F) Moult of 22.XI.2020. (G) Moult of VI.2019. (H-I) Moult of 13.II.2021. (J) Allotype; moult of 19.I.2019. (K-L) Moult of 14.II.2021. (M) Moult of 1.V.2021. (N-P) Moult of 23.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761549" httpUri="https://zenodo.org/record/7761549/files/figure.png" pageId="4" pageNumber="378">Fig. 22</figureCitation>
|
||
and
|
||
<taxonomicName authority=", Yu et al., 2021" authorityName="Yu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[566,678,322,345]" italics="true" pageId="4" pageNumber="378">L. nabang</emphasis>
|
||
,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." pageId="4" pageNumber="378" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis italics="true" pageId="4" pageNumber="378">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 4); lateral margins of poreplate ventrally with (e.g.
|
||
<figureCitation box="[262,327,386,409]" captionStart="Fig" captionStartId="13.[149,184,1407,1428]" captionTargetBox="[154,1434,257,1369]" captionTargetId="figure-257@13.[149,1437,254,1374]" captionTargetPageId="13" captionText="Fig. 7. Liphistius metopiae sp. nov., vulval plates of five female paratypes (all taken from exuviae); dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Allotype; moult of 23.IV.1998. (D-E) Moult of 16.IV.1999. (E) Moult of 24.II.2000. (G-H) Moult of 28.VI.2001. (I) Moult of 4.III.1999. Scale lines 1.0 mm. Abbreviations: al - anterior lobe of poreplate; alp - anterolateral process of poreplate." figureDoi="http://doi.org/10.5281/zenodo.7761503" httpUri="https://zenodo.org/record/7761503/files/figure.png" pageId="4" pageNumber="378">Fig. 7</figureCitation>
|
||
) or without small, knob-shaped anterolateral processes (e.g.
|
||
<figureCitation box="[399,466,418,441]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="4" pageNumber="378">Fig. 5</figureCitation>
|
||
); CDO mostly small (e.g.
|
||
<figureCitation box="[151,239,450,473]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="4" pageNumber="378">Fig. 5A</figureCitation>
|
||
), slightly larger only in
|
||
<taxonomicName box="[517,597,450,473]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="tung" status="sp. nov.">
|
||
<emphasis box="[517,597,450,473]" italics="true" pageId="4" pageNumber="378">L. tung</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[607,695,450,473]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="4" pageNumber="378">Fig. 9A, C, E</figureCitation>
|
||
), distinctly larger only in juveniles (
|
||
<figureCitation box="[655,744,482,505]" captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="4" pageNumber="378">Fig. 9G</figureCitation>
|
||
), mostly rounded (e.g.
|
||
<figureCitation box="[417,559,514,537]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="4" pageNumber="378">Fig. 5A, D</figureCitation>
|
||
), rarely slit-like (
|
||
<figureCitation box="[159,260,546,569]" captionStart="Fig" captionStartId="26.[151,186,1819,1840]" captionTargetId="figure-9@27.[151,1436,257,2027]" captionTargetPageId="27" captionText="Fig. 14. Liphistius birmanicus, vulval plates (all taken from exuviae) of ten females from the type locality (A-M) and of one female from the Shan State (N-O); dorsal view [A, D (with uterus externus attached), G, J, N], ventral view (B-C, E-F, H-I, K-M, O). (A-B) Moult of 6.II.2021. (C) Moult of 27.XII.2020. (D) Moult of 20.XI.2021. (E) Moult of 19.II.2021 (same specimen as D). (F) Moult of 21.II.2021. (G-H) Moult of 25.XII.2020. (I) Moult of 14.V.2021. (J-K) Moult of 20.III.2021. (L) Moult of 26.X.2017. (M) Moult of 24.VIII.2017. (N-O) Moult of 2.IV.2018. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761529" httpUri="https://zenodo.org/record/7761529/files/figure.png" pageId="4" pageNumber="378">Fig. 14D</figureCitation>
|
||
); receptacular cluster always racemose (never digitiform as in some
|
||
<emphasis box="[396,454,578,601]" italics="true" pageId="4" pageNumber="378">trang</emphasis>
|
||
-group species), moderately large, only in a few females slightly surpassing anterior margin of poreplate (e.g.
|
||
<figureCitation box="[471,548,642,665]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="4" pageNumber="378">Figs 5</figureCitation>
|
||
A-B, 18D, G-H). Poreplate and posterior stalk always fully sclerotised and connected to each other. The latter ranging from very short (e.g.
|
||
<figureCitation box="[321,418,738,761]" captionStart="Fig" captionStartId="40.[151,186,1942,1963]" captionTargetBox="[155,1436,262,1898]" captionTargetId="figure-107@40.[151,1439,257,1908]" captionTargetPageId="40" captionText="Fig. 22. Liphistius cupreus sp. nov., vulval plates of ten females (all taken from exuviae), dorsal view (E with uterus externus attached, H, K, N), ventral view (A-D, I-J, L-M, O), ventral and slightly anterior view (P), posterior view (G), and isolated uterus externus (F). (A) Moult of 10.II.2019. (B) Moult of 12.V.2020. (C) Moult of 5.VII.2019. (D-F) Moult of 22.XI.2020. (G) Moult of VI.2019. (H-I) Moult of 13.II.2021. (J) Allotype; moult of 19.I.2019. (K-L) Moult of 14.II.2021. (M) Moult of 1.V.2021. (N-P) Moult of 23.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761549" httpUri="https://zenodo.org/record/7761549/files/figure.png" pageId="4" pageNumber="378">Fig. 22K</figureCitation>
|
||
) to quite long (e.g.
|
||
<figureCitation box="[630,710,738,761]" captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="4" pageNumber="378">Fig. 9C</figureCitation>
|
||
and
|
||
<bibRefCitation author="Aung K. P. P. & Xu X. & Lwin W. W. & Sang M. Z. & Yu L. & Liu H. & Liu F. & Li D." box="[151,347,770,793]" pageId="4" pageNumber="378" pagination="29 - 39" refId="ref37364" refString="Aung K. P. P., Xu X., Lwin W. W., Sang M. Z., Yu L., Liu H., Liu F., Li D. 2019. Two new species of the primitively segmented spider genus Liphistius Schiodte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 29 - 39." type="journal article" year="2019">
|
||
Aung
|
||
<emphasis box="[219,268,770,793]" italics="true" pageId="4" pageNumber="378">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
: fig. 3B-E), and from much narrower than poreplate (e.g.
|
||
<figureCitation box="[384,468,802,825]" captionStart="Fig" captionStartId="36.[151,186,1326,1347]" captionTargetBox="[155,1436,271,1286]" captionTargetId="figure-385@36.[151,1439,257,1291]" captionTargetPageId="36" captionText="Fig. 20. Liphistius pyinoolwin, vulval plates of three females (all taken from exuviae), dorsal view (A, D, F), ventral view (B, E, G) and ventral and slightly anterior view (C). (A-C) Moult of 1.IX.2017. (D-E) Moult of 15.X.2017. (F-G) Moult of 22.I.2016. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761545" httpUri="https://zenodo.org/record/7761545/files/figure.png" pageId="4" pageNumber="378">Fig. 20</figureCitation>
|
||
) to distinctly wider than poreplate (e.g.
|
||
<figureCitation box="[315,411,834,857]" captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="4" pageNumber="378">Fig. 24D</figureCitation>
|
||
and
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[465,630,834,857]" pageId="4" pageNumber="378" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[503,552,834,857]" italics="true" pageId="4" pageNumber="378">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 4C-F), from widely eliptically (e.g.
|
||
<figureCitation box="[476,564,866,889]" captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="4" pageNumber="378">Fig. 9A</figureCitation>
|
||
) to quadrangular (e.g.
|
||
<figureCitation box="[211,310,898,921]" captionStart="Fig" captionStartId="26.[151,186,1819,1840]" captionTargetId="figure-9@27.[151,1436,257,2027]" captionTargetPageId="27" captionText="Fig. 14. Liphistius birmanicus, vulval plates (all taken from exuviae) of ten females from the type locality (A-M) and of one female from the Shan State (N-O); dorsal view [A, D (with uterus externus attached), G, J, N], ventral view (B-C, E-F, H-I, K-M, O). (A-B) Moult of 6.II.2021. (C) Moult of 27.XII.2020. (D) Moult of 20.XI.2021. (E) Moult of 19.II.2021 (same specimen as D). (F) Moult of 21.II.2021. (G-H) Moult of 25.XII.2020. (I) Moult of 14.V.2021. (J-K) Moult of 20.III.2021. (L) Moult of 26.X.2017. (M) Moult of 24.VIII.2017. (N-O) Moult of 2.IV.2018. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761529" httpUri="https://zenodo.org/record/7761529/files/figure.png" pageId="4" pageNumber="378">Fig. 14B</figureCitation>
|
||
) and axe-blade-shaped (e.g.
|
||
<figureCitation box="[640,743,898,921]" captionStart="Fig" captionStartId="31.[151,186,1817,1838]" captionTargetBox="[155,1436,267,1763]" captionTargetId="figure-78@31.[151,1440,255,1768]" captionTargetPageId="31" captionText="Fig. 16. Liphistius pinlaung (A-J) and Liphistius sp. (specimen from Kalaw; K-L), vulval plates of seven females (all taken from exuviae), dorsal view (A, D, G, I, K) and ventral view (B-C, E-F, H, J, L). (A-B) Moult of 10.II.2019. (C) Moult of 7.XII.2018. (D-E) Moult of 10.IV.2021. (F) Moult of 21.X.2018. (G-H) Moult of 16.V.2020. (I-J) Moult of 8.III.2020. (K-L) Moult of 15.VIII.2017. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761533" httpUri="https://zenodo.org/record/7761533/files/figure.png" pageId="4" pageNumber="378">Fig. 16A</figureCitation>
|
||
), with a short and indistinct (e.g.
|
||
<figureCitation box="[496,578,930,953]" captionStart="Fig" captionStartId="40.[151,186,1942,1963]" captionTargetBox="[155,1436,262,1898]" captionTargetId="figure-107@40.[151,1439,257,1908]" captionTargetPageId="40" captionText="Fig. 22. Liphistius cupreus sp. nov., vulval plates of ten females (all taken from exuviae), dorsal view (E with uterus externus attached, H, K, N), ventral view (A-D, I-J, L-M, O), ventral and slightly anterior view (P), posterior view (G), and isolated uterus externus (F). (A) Moult of 10.II.2019. (B) Moult of 12.V.2020. (C) Moult of 5.VII.2019. (D-F) Moult of 22.XI.2020. (G) Moult of VI.2019. (H-I) Moult of 13.II.2021. (J) Allotype; moult of 19.I.2019. (K-L) Moult of 14.II.2021. (M) Moult of 1.V.2021. (N-P) Moult of 23.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761549" httpUri="https://zenodo.org/record/7761549/files/figure.png" pageId="4" pageNumber="378">Figs 22</figureCitation>
|
||
,
|
||
<figureCitation box="[590,618,930,953]" captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="4" pageNumber="378">24</figureCitation>
|
||
) to long and
|
||
</paragraph>
|
||
<paragraph blockId="4.[831,1436,258,313]" pageId="4" pageNumber="378">
|
||
pronounced (e.g.
|
||
<figureCitation box="[1021,1098,258,281]" captionStart="Fig" captionStartId="36.[151,186,1326,1347]" captionTargetBox="[155,1436,271,1286]" captionTargetId="figure-385@36.[151,1439,257,1291]" captionTargetPageId="36" captionText="Fig. 20. Liphistius pyinoolwin, vulval plates of three females (all taken from exuviae), dorsal view (A, D, F), ventral view (B, E, G) and ventral and slightly anterior view (C). (A-C) Moult of 1.IX.2017. (D-E) Moult of 15.X.2017. (F-G) Moult of 22.I.2016. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761545" httpUri="https://zenodo.org/record/7761545/files/figure.png" pageId="4" pageNumber="378">Fig. 20</figureCitation>
|
||
F-G) constriction in its anterior part.
|
||
</paragraph>
|
||
<paragraph blockId="4.[831,1437,337,648]" pageId="4" pageNumber="378">
|
||
<emphasis bold="true" box="[831,1171,337,360]" pageId="4" pageNumber="378">Species included (n = 12):</emphasis>
|
||
<taxonomicName authority="Thorell, 1897" authorityName="Thorell" authorityYear="1897" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis box="[1189,1436,337,360]" italics="true" pageId="4" pageNumber="378">Liphistius birmanicus</emphasis>
|
||
<bibRefCitation author="Thorell T." box="[831,998,369,392]" pageId="4" pageNumber="378" pagination="161 - 267" refId="ref38425" refString="Thorell T. 1897. Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXIII. Secondo saggio sui Ragni Birmani. I. Parallelodontes - Tubitelariae. Annali del Museo Civico di Storia Naturale di Genova (serie 2) 17: 161 - 267." type="journal article" year="1897">Thorell, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
(Myanmar),
|
||
<taxonomicName box="[1178,1311,369,392]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[1178,1311,369,392]" italics="true" pageId="4" pageNumber="378">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1335,1437,369,392]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar),
|
||
<taxonomicName box="[975,1064,401,424]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
|
||
<emphasis box="[975,1064,401,424]" italics="true" pageId="4" pageNumber="378">L. ferox</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1079,1169,401,424]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar),
|
||
<taxonomicName authority="Aung et al., 2019" authorityName="Aung" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="hpruso">
|
||
<emphasis box="[1326,1436,401,424]" italics="true" pageId="4" pageNumber="378">L. hpruso</emphasis>
|
||
<bibRefCitation author="Aung K. P. P. & Xu X. & Lwin W. W. & Sang M. Z. & Yu L. & Liu H. & Liu F. & Li D." box="[831,1033,433,456]" pageId="4" pageNumber="378" pagination="29 - 39" refId="ref37364" refString="Aung K. P. P., Xu X., Lwin W. W., Sang M. Z., Yu L., Liu H., Liu F., Li D. 2019. Two new species of the primitively segmented spider genus Liphistius Schiodte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 29 - 39." type="journal article" year="2019">
|
||
Aung
|
||
<emphasis box="[903,961,433,456]" italics="true" pageId="4" pageNumber="378">et al.</emphasis>
|
||
, 2019
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(Myanmar),
|
||
<taxonomicName authority="Schwendinger, 1998" authorityName="Schwendinger" authorityYear="1998" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="lahu">
|
||
<emphasis box="[1185,1266,433,456]" italics="true" pageId="4" pageNumber="378">L. lahu</emphasis>
|
||
<bibRefCitation author="Schwendinger P. J." pageId="4" pageNumber="378" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
(Thailand),
|
||
<taxonomicName authority="Platnick & Sedgwick, 1984" authorityName="Platnick & Sedgwick" authorityYear="1984" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="lordae">
|
||
<emphasis box="[1045,1156,465,488]" italics="true" pageId="4" pageNumber="378">L. lordae</emphasis>
|
||
<bibRefCitation author="Platnick N. I. & Sedgwick W. C." pageId="4" pageNumber="378" pagination="1 - 31" refId="ref37980" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." type="journal article" year="1984">Platnick & Sedgwick, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
(Myanmar),
|
||
<taxonomicName box="[1050,1187,497,520]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="metopiae" status="sp. nov.">
|
||
<emphasis box="[1050,1187,497,520]" italics="true" pageId="4" pageNumber="378">L. metopiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1205,1299,497,520]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Thailand),
|
||
<taxonomicName authority="Yu et al., 2021" authorityName="Yu" authorityYear="2021" box="[831,1111,529,552]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[831,942,529,552]" italics="true" pageId="4" pageNumber="378">L. nabang</emphasis>
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[949,1111,529,552]" pageId="4" pageNumber="378" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[988,1036,529,552]" italics="true" pageId="4" pageNumber="378">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(China),
|
||
<taxonomicName authority="Aung et al., 2019" authorityName="Aung" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="pinlaung">
|
||
<emphasis box="[1217,1342,529,552]" italics="true" pageId="4" pageNumber="378">L. pinlaung</emphasis>
|
||
<bibRefCitation author="Aung K. P. P. & Xu X. & Lwin W. W. & Sang M. Z. & Yu L. & Liu H. & Liu F. & Li D." pageId="4" pageNumber="378" pagination="29 - 39" refId="ref37364" refString="Aung K. P. P., Xu X., Lwin W. W., Sang M. Z., Yu L., Liu H., Liu F., Li D. 2019. Two new species of the primitively segmented spider genus Liphistius Schiodte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 29 - 39." type="journal article" year="2019">
|
||
Aung
|
||
<emphasis italics="true" pageId="4" pageNumber="378">et al.</emphasis>
|
||
, 2019
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(Myanmar),
|
||
<taxonomicName box="[1077,1202,561,584]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[1077,1202,561,584]" italics="true" pageId="4" pageNumber="378">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1212,1299,561,584]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar),
|
||
<taxonomicName authority="Xu et al., 2021" authorityName="Xu" authorityYear="2021" box="[831,1166,593,616]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="pyinoolwin">
|
||
<emphasis box="[831,983,593,616]" italics="true" pageId="4" pageNumber="378">L. pyinoolwin</emphasis>
|
||
<bibRefCitation author="Xu X. & Yu L. & Aung K. P. P. & Yu L. & Liu F. & Lwin W. W. & Sang M. Z. & Li D." box="[994,1166,593,616]" pageId="4" pageNumber="378" pagination="41 - 58" refId="ref38558" refString="Xu X., Yu L., Aung K. P. P., Yu L., Liu F., Lwin W. W., Sang M. Z., Li D. 2021. A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae). ZooKeys 1031: 41 - 58." type="journal article" year="2021">
|
||
Xu
|
||
<emphasis box="[1037,1101,593,616]" italics="true" pageId="4" pageNumber="378">et al.,</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(Myanmar),
|
||
<taxonomicName box="[1315,1395,593,616]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="tung" status="sp. nov.">
|
||
<emphasis box="[1315,1395,593,616]" italics="true" pageId="4" pageNumber="378">L. tung</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="378" type="distribution">
|
||
<paragraph blockId="4.[831,1437,672,887]" pageId="4" pageNumber="378">
|
||
<emphasis bold="true" box="[831,983,672,695]" pageId="4" pageNumber="378">Distribution:</emphasis>
|
||
All species known so far occur in the mountainous area that lies roughly between the Ayeyarwady (= Irrawaddy) River in the west, the Mekong River in the east and the Thai-Burmese border in the south (
|
||
<figureCitation box="[994,1065,800,823]" captionStart="Fig" captionStartId="7.[151,186,1431,1452]" captionTargetBox="[151,1436,257,1395]" captionTargetId="figure-270@7.[151,1436,257,1395]" captionTargetPageId="7" captionText="Fig. 1. Localities of species in the Liphistius birmanicus-group in northern Thailand, eastern Myanmar and southern China. Blue lines indicate major rivers, black lines national borders, grey lines major roads." figureDoi="http://doi.org/10.5281/zenodo.7761489" httpUri="https://zenodo.org/record/7761489/files/figure.png" pageId="4" pageNumber="378">Fig. 1</figureCitation>
|
||
). At least two of these species (
|
||
<taxonomicName authorityName="Schwendinger" authorityYear="1998" box="[840,921,832,855]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="lahu">
|
||
<emphasis box="[840,921,832,855]" italics="true" pageId="4" pageNumber="378">L. lahu</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[981,1068,832,855]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
|
||
<emphasis box="[981,1068,832,855]" italics="true" pageId="4" pageNumber="378">L. ferox</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1079,1168,832,855]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) occur together with or very close to species of the
|
||
<taxonomicName authorityName="Platnick & Sedgwick" authorityYear="1984" box="[1126,1225,864,887]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="bristowei">
|
||
<emphasis box="[1126,1225,864,887]" italics="true" pageId="4" pageNumber="378">bristowei</emphasis>
|
||
</taxonomicName>
|
||
-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="380" pageId="4" pageNumber="378" type="key">
|
||
<paragraph blockId="4.[151,362,1038,1061]" box="[151,362,1038,1061]" pageId="4" pageNumber="378">
|
||
<emphasis bold="true" box="[151,362,1038,1061]" pageId="4" pageNumber="378">Key to the species:</emphasis>
|
||
</paragraph>
|
||
<key lastPageId="6" lastPageNumber="380" pageId="4" pageNumber="378">
|
||
<keyStep pageId="4" pageNumber="378">
|
||
<paragraph blockId="4.[151,1437,1102,2053]" box="[151,1436,1102,1125]" pageId="4" pageNumber="378">
|
||
<keyLead box="[151,1436,1102,1125]" pageId="4" pageNumber="378">
|
||
1 Males (unknown for
|
||
<taxonomicName box="[466,568,1102,1125]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="hpruso">
|
||
<emphasis box="[466,568,1102,1125]" italics="true" pageId="4" pageNumber="378">L. hpruso</emphasis>
|
||
</taxonomicName>
|
||
) ............................................................................................................................ 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1102,2053]" box="[151,1437,1134,1157]" pageId="4" pageNumber="378">
|
||
<keyLead box="[151,1437,1134,1157]" pageId="4" pageNumber="378">- Females................................................................................................................................................................ 12</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="378">
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
2(
|
||
<emphasis box="[173,186,1166,1189]" italics="true" pageId="4" pageNumber="378">1</emphasis>
|
||
) Proventral process of contrategulum strongly distad-directed, almost parallel to axis of palpal organ (
|
||
<figureCitation captionStart="Fig" captionStartId="26.[151,186,1411,1432]" captionTargetBox="[157,1431,263,1314]" captionTargetId="figure-229@26.[151,1436,257,1320]" captionTargetPageId="26" captionText="Fig. 13. Liphistius birmanicus, left palps of two males from the type locality; male matured 27.I.2016 (A-B), male matured 15.XI.2014 (C-H). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum, distal and slightly prolateral view. (C) Distal part of tarsus, prodorsal view; knob-shaped process on distodorsal corner indicated by short arrow. (D) Palpal organ and distal margin of tarsus, prolateral view. (E) Same, ventral view; bent proximal edge of tegulum indicated by long arrow. (F) Distal part of palpal organ, retroventral and slightly proximal view. (G) Palpal organ, retrolateral view. (H) Proximal part of tarsus, ventral view. Scale line 1.0 mm.Abbreviation: cp - proventral contrategular process." figureDoi="http://doi.org/10.5281/zenodo.7761527" httpUri="https://zenodo.org/record/7761527/files/figure.png" pageId="4" pageNumber="378">Figs 13D, F</figureCitation>
|
||
,
|
||
<figureCitation box="[268,313,1198,1221]" captionStart="Fig" captionStartId="30.[151,186,1936,1957]" captionTargetBox="[155,1431,946,1889]" captionTargetId="figure-412@30.[151,1436,941,1894]" captionTargetPageId="30" captionText="Fig. 15. Liphistius pinlaung, left palps of two males: matured 23.XI.2014 (A-B), matured 4.VII.2015 (C-G). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Palpal organ and distal margin of tarsus, prolateral view. (D) Palpal organ, ventral view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of palpal organ, retroventral view. (F) Tegulum, retrolateral view. (G) Proximal part of tarsus, ventral view.All to same scale; scale line 1.0 mm. Abbreviation: cp - proventral process of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761531" httpUri="https://zenodo.org/record/7761531/files/figure.png" pageId="4" pageNumber="378">15C</figureCitation>
|
||
) .................................................................................................................................................................. 3
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
- Proventral process of contrategulum inclined from axis of palpal organ at an angle of roughly 45-90° (
|
||
<figureCitation box="[1342,1432,1230,1253]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="4" pageNumber="378">Figs 4G</figureCitation>
|
||
,
|
||
<figureCitation box="[242,266,1262,1285]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="4" pageNumber="378">6I</figureCitation>
|
||
,
|
||
<figureCitation box="[277,301,1262,1285]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="4" pageNumber="378">8I</figureCitation>
|
||
,
|
||
<figureCitation box="[313,355,1262,1285]" captionStart="Fig" captionStartId="19.[151,186,1745,1766]" captionTargetBox="[156,1430,263,1692]" captionTargetId="figure-155@19.[151,1436,257,1698]" captionTargetPageId="19" captionText="Fig. 10. Liphistius ferox sp. nov., left palps of four males; holotype (A-B, E-K), male from Naw Bubaw, matured IV.2019 (C, M), male from Thandaung Gyi, matured 7.X.2014 (D), male from Thandaung Gyi, matured 18.IX.2016 (L). (A) Tarsus and palpal organ, ventral view. (B) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (C-D) Embolus complex, ventral view. (F) Embolus proper, prodorsal view. (G) Palpal organ, retroventral view. (H) Palpal organ, distal view (dorsal side up). (I) Dorsal part of contrategulum, distal and slightly prolateral view. (J) Distal part of tarsus, ventral view. (K-M) Same, retrolateral and slightly proximal view. Scale lines 1.0 mm (A; B, F-G; C-E; H-I; J; K-M). Abbreviations: tp - triangular distal protrusion of longitudinal rib on sclerotised part of embolus proper; so - angular retrolateral side of sperm duct opening." figureDoi="http://doi.org/10.5281/zenodo.7761515" httpUri="https://zenodo.org/record/7761515/files/figure.png" pageId="4" pageNumber="378">10F</figureCitation>
|
||
,
|
||
<figureCitation box="[366,411,1262,1285]" captionStart="Fig" captionStartId="32.[151,186,1942,1963]" captionTargetBox="[159,1427,1125,1893]" captionTargetId="figure-353@32.[151,1436,1117,1903]" captionTargetPageId="32" captionText="Fig. 17. Liphistius lordae, left palps of two males: male n°1 (A-B, D-G), male n°2 (C). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Contrategulum, distal view. (D) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Palpal organ and distal margin of tarsus, prolateral view. (F) Tibial apophysis, retrolateral and slightly proximal view. (G) Proximal part of tarsus, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761535" httpUri="https://zenodo.org/record/7761535/files/figure.png" pageId="4" pageNumber="378">17E</figureCitation>
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||
,
|
||
<figureCitation box="[422,468,1262,1285]" captionStart="Fig" captionStartId="35.[151,186,1288,1309]" captionTargetBox="[156,1431,261,1232]" captionTargetId="figure-280@35.[151,1436,257,1238]" captionTargetPageId="35" captionText="Fig. 19. Liphistius pyinoolwin, palps of two males: matured 16.X.2014 (A-E), matured 24.IX.2014 (F). (A) Left palpal organ, distal view (dorsal side up). (B) Left embolus complex and dorsal part of contrategulum, distal and slightly proventral view. (C) Left palpal organ and distal margin of tarsus, prolateral view. (D) Left palpal organ, ventral and slightly proximal view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Distal part of left tibia and proximal part of tarsus, ventral view. (F) Distal margin of right tibia and proximal part of tarsus, ventral view. Scale lines 1.0 mm (A-D; E-F). Abbreviation: l - lobate prolateral protrusion on base of embolus complex." figureDoi="http://doi.org/10.5281/zenodo.7761541" httpUri="https://zenodo.org/record/7761541/files/figure.png" pageId="4" pageNumber="378">19C</figureCitation>
|
||
,
|
||
<figureCitation box="[480,528,1262,1285]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="4" pageNumber="378">21N</figureCitation>
|
||
,
|
||
<figureCitation box="[539,586,1262,1285]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="4" pageNumber="378">23H</figureCitation>
|
||
) ......................................................................................................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="378">
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
3(
|
||
<emphasis box="[173,186,1294,1317]" italics="true" pageId="4" pageNumber="378">2</emphasis>
|
||
) Large spiders (CL and CW of males 7.79-9.30 and 7.29-8.48, respectively); paracymbium without retrolateralproximal heel (
|
||
<figureCitation box="[404,500,1326,1349]" captionStart="Fig" captionStartId="26.[151,186,1411,1432]" captionTargetBox="[157,1431,263,1314]" captionTargetId="figure-229@26.[151,1436,257,1320]" captionTargetPageId="26" captionText="Fig. 13. Liphistius birmanicus, left palps of two males from the type locality; male matured 27.I.2016 (A-B), male matured 15.XI.2014 (C-H). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum, distal and slightly prolateral view. (C) Distal part of tarsus, prodorsal view; knob-shaped process on distodorsal corner indicated by short arrow. (D) Palpal organ and distal margin of tarsus, prolateral view. (E) Same, ventral view; bent proximal edge of tegulum indicated by long arrow. (F) Distal part of palpal organ, retroventral and slightly proximal view. (G) Palpal organ, retrolateral view. (H) Proximal part of tarsus, ventral view. Scale line 1.0 mm.Abbreviation: cp - proventral contrategular process." figureDoi="http://doi.org/10.5281/zenodo.7761527" httpUri="https://zenodo.org/record/7761527/files/figure.png" pageId="4" pageNumber="378">Fig. 13H</figureCitation>
|
||
) ...............................................................................................
|
||
<taxonomicName authorityName="Thorell" authorityYear="1897" box="[1160,1308,1326,1349]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis box="[1160,1308,1326,1349]" italics="true" pageId="4" pageNumber="378">L. birmanicus</emphasis>
|
||
</taxonomicName>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
- Medium-sized spiders (CL and CW of males 4.92-6.27 and 4.56-5.40, respectively); paracymbium with retrolateral-proximal heel (
|
||
<figureCitation box="[460,557,1390,1413]" captionStart="Fig" captionStartId="30.[151,186,1936,1957]" captionTargetBox="[155,1431,946,1889]" captionTargetId="figure-412@30.[151,1436,941,1894]" captionTargetPageId="30" captionText="Fig. 15. Liphistius pinlaung, left palps of two males: matured 23.XI.2014 (A-B), matured 4.VII.2015 (C-G). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Palpal organ and distal margin of tarsus, prolateral view. (D) Palpal organ, ventral view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of palpal organ, retroventral view. (F) Tegulum, retrolateral view. (G) Proximal part of tarsus, ventral view.All to same scale; scale line 1.0 mm. Abbreviation: cp - proventral process of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761531" httpUri="https://zenodo.org/record/7761531/files/figure.png" pageId="4" pageNumber="378">Fig. 15G</figureCitation>
|
||
) ...........................................................................................
|
||
<taxonomicName box="[1185,1308,1390,1413]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="pinlaung">
|
||
<emphasis box="[1185,1308,1390,1413]" italics="true" pageId="4" pageNumber="378">L. pinlaung</emphasis>
|
||
</taxonomicName>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="378">
|
||
<paragraph blockId="4.[151,1437,1102,2053]" box="[151,1436,1422,1445]" pageId="4" pageNumber="378">
|
||
<keyLead box="[151,1436,1422,1445]" pageId="4" pageNumber="378">
|
||
4(
|
||
<emphasis box="[173,186,1422,1445]" italics="true" pageId="4" pageNumber="378">2</emphasis>
|
||
) Para-embolic plate much longer than retroventral edge of embolus complex (
|
||
<figureCitation box="[1062,1151,1422,1445]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="4" pageNumber="378">Figs 8A</figureCitation>
|
||
, H-I, 10A-D ................. 5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
- Para-embolic plate as long as or only slightly longer than retroventral edge of embolus complex (
|
||
<figureCitation box="[1301,1391,1454,1477]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="4" pageNumber="378">Figs 4D</figureCitation>
|
||
,
|
||
<figureCitation box="[1404,1433,1454,1477]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="4" pageNumber="378">6F</figureCitation>
|
||
,
|
||
<figureCitation box="[242,287,1486,1509]" captionStart="Fig" captionStartId="26.[151,186,1411,1432]" captionTargetBox="[157,1431,263,1314]" captionTargetId="figure-229@26.[151,1436,257,1320]" captionTargetPageId="26" captionText="Fig. 13. Liphistius birmanicus, left palps of two males from the type locality; male matured 27.I.2016 (A-B), male matured 15.XI.2014 (C-H). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum, distal and slightly prolateral view. (C) Distal part of tarsus, prodorsal view; knob-shaped process on distodorsal corner indicated by short arrow. (D) Palpal organ and distal margin of tarsus, prolateral view. (E) Same, ventral view; bent proximal edge of tegulum indicated by long arrow. (F) Distal part of palpal organ, retroventral and slightly proximal view. (G) Palpal organ, retrolateral view. (H) Proximal part of tarsus, ventral view. Scale line 1.0 mm.Abbreviation: cp - proventral contrategular process." figureDoi="http://doi.org/10.5281/zenodo.7761527" httpUri="https://zenodo.org/record/7761527/files/figure.png" pageId="4" pageNumber="378">13E</figureCitation>
|
||
,
|
||
<figureCitation box="[298,347,1486,1509]" captionStart="Fig" captionStartId="30.[151,186,1936,1957]" captionTargetBox="[155,1431,946,1889]" captionTargetId="figure-412@30.[151,1436,941,1894]" captionTargetPageId="30" captionText="Fig. 15. Liphistius pinlaung, left palps of two males: matured 23.XI.2014 (A-B), matured 4.VII.2015 (C-G). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Palpal organ and distal margin of tarsus, prolateral view. (D) Palpal organ, ventral view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of palpal organ, retroventral view. (F) Tegulum, retrolateral view. (G) Proximal part of tarsus, ventral view.All to same scale; scale line 1.0 mm. Abbreviation: cp - proventral process of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761531" httpUri="https://zenodo.org/record/7761531/files/figure.png" pageId="4" pageNumber="378">15D</figureCitation>
|
||
,
|
||
<figureCitation box="[357,405,1486,1509]" captionStart="Fig" captionStartId="32.[151,186,1942,1963]" captionTargetBox="[159,1427,1125,1893]" captionTargetId="figure-353@32.[151,1436,1117,1903]" captionTargetPageId="32" captionText="Fig. 17. Liphistius lordae, left palps of two males: male n°1 (A-B, D-G), male n°2 (C). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Contrategulum, distal view. (D) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Palpal organ and distal margin of tarsus, prolateral view. (F) Tibial apophysis, retrolateral and slightly proximal view. (G) Proximal part of tarsus, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761535" httpUri="https://zenodo.org/record/7761535/files/figure.png" pageId="4" pageNumber="378">17D</figureCitation>
|
||
,
|
||
<figureCitation box="[417,465,1486,1509]" captionStart="Fig" captionStartId="35.[151,186,1288,1309]" captionTargetBox="[156,1431,261,1232]" captionTargetId="figure-280@35.[151,1436,257,1238]" captionTargetPageId="35" captionText="Fig. 19. Liphistius pyinoolwin, palps of two males: matured 16.X.2014 (A-E), matured 24.IX.2014 (F). (A) Left palpal organ, distal view (dorsal side up). (B) Left embolus complex and dorsal part of contrategulum, distal and slightly proventral view. (C) Left palpal organ and distal margin of tarsus, prolateral view. (D) Left palpal organ, ventral and slightly proximal view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Distal part of left tibia and proximal part of tarsus, ventral view. (F) Distal margin of right tibia and proximal part of tarsus, ventral view. Scale lines 1.0 mm (A-D; E-F). Abbreviation: l - lobate prolateral protrusion on base of embolus complex." figureDoi="http://doi.org/10.5281/zenodo.7761541" httpUri="https://zenodo.org/record/7761541/files/figure.png" pageId="4" pageNumber="378">19D</figureCitation>
|
||
,
|
||
<figureCitation box="[476,524,1486,1509]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="4" pageNumber="378">21A</figureCitation>
|
||
,
|
||
<figureCitation box="[535,582,1486,1509]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="4" pageNumber="378">23A</figureCitation>
|
||
) .......................................................................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="378">
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
5(
|
||
<emphasis box="[173,186,1518,1541]" italics="true" pageId="4" pageNumber="378">4</emphasis>
|
||
) Medium-sized spiders (CL and CW of males 6.30-6.70 and 5.70-6.26, respectively); sclerotised part of embolus proper strengthened by 3 longitudinal ribs reaching the apex, none of them with an angular distal protrusion (
|
||
<figureCitation captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="4" pageNumber="378">Fig. 8A</figureCitation>
|
||
, C-D, F-I); proximal edge of tegulum salient, distinctly protruding from surface of palpal organ (
|
||
<figureCitation box="[1318,1380,1582,1605]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="4" pageNumber="378">Fig. 8</figureCitation>
|
||
H-J); tibial apophysis distinctly set back from anterior margin of tibia (
|
||
<figureCitation box="[937,1018,1614,1637]" captionStart="Fig" captionStartId="16.[151,186,1775,1796]" captionTargetBox="[154,1433,261,1719]" captionTargetId="figure-111@16.[151,1436,257,1722]" captionTargetPageId="16" captionText="Fig. 8. Liphistius tung sp. nov., left palps of male holotype (A-C, G, I-J) and of male paratype (D-F, H). (A) Tarsus and palpal organ, ventral view. (B) Distal part of tibia and proximal part of tarsus, ventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Distal edge of contrategulum, distal and slightly prodorsal view. (F) Palpal organ, retrolateral view. (G) Same, retroventral view. (H-I) Distal part of palpal organ, ventral and slightly proximal view. (J) Distal part of tarsus and palpal organ, prolateral view. Arrows indicating bent and salient proximal edge of tegulum. Scale lines 1.0 mm (A-B; C-J)." figureDoi="http://doi.org/10.5281/zenodo.7761509" httpUri="https://zenodo.org/record/7761509/files/figure.png" pageId="4" pageNumber="378">Fig. 8B</figureCitation>
|
||
) ................
|
||
<taxonomicName box="[1153,1228,1614,1637]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="tung" status="sp. nov.">
|
||
<emphasis box="[1153,1228,1614,1637]" italics="true" pageId="4" pageNumber="378">L. tung</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1235,1318,1614,1637]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Thailand)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
- Very large spiders (CL and CW of males 11.47-12.67 and 10.45-11.47, respectively); sclerotised part of embolus proper strengthened by only 2 longitudinal ribs reaching the apex, one of them carrying an angular distal protrusion (
|
||
<figureCitation box="[302,380,1710,1733]" captionStart="Fig" captionStartId="19.[151,186,1745,1766]" captionTargetBox="[156,1430,263,1692]" captionTargetId="figure-155@19.[151,1436,257,1698]" captionTargetPageId="19" captionText="Fig. 10. Liphistius ferox sp. nov., left palps of four males; holotype (A-B, E-K), male from Naw Bubaw, matured IV.2019 (C, M), male from Thandaung Gyi, matured 7.X.2014 (D), male from Thandaung Gyi, matured 18.IX.2016 (L). (A) Tarsus and palpal organ, ventral view. (B) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (C-D) Embolus complex, ventral view. (F) Embolus proper, prodorsal view. (G) Palpal organ, retroventral view. (H) Palpal organ, distal view (dorsal side up). (I) Dorsal part of contrategulum, distal and slightly prolateral view. (J) Distal part of tarsus, ventral view. (K-M) Same, retrolateral and slightly proximal view. Scale lines 1.0 mm (A; B, F-G; C-E; H-I; J; K-M). Abbreviations: tp - triangular distal protrusion of longitudinal rib on sclerotised part of embolus proper; so - angular retrolateral side of sperm duct opening." figureDoi="http://doi.org/10.5281/zenodo.7761515" httpUri="https://zenodo.org/record/7761515/files/figure.png" pageId="4" pageNumber="378">Fig. 10</figureCitation>
|
||
E-H); proximal edge of tegulum adpressed to surface of palpal organ (
|
||
<figureCitation box="[1146,1245,1710,1733]" captionStart="Fig" captionStartId="19.[151,186,1745,1766]" captionTargetBox="[156,1430,263,1692]" captionTargetId="figure-155@19.[151,1436,257,1698]" captionTargetPageId="19" captionText="Fig. 10. Liphistius ferox sp. nov., left palps of four males; holotype (A-B, E-K), male from Naw Bubaw, matured IV.2019 (C, M), male from Thandaung Gyi, matured 7.X.2014 (D), male from Thandaung Gyi, matured 18.IX.2016 (L). (A) Tarsus and palpal organ, ventral view. (B) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (C-D) Embolus complex, ventral view. (F) Embolus proper, prodorsal view. (G) Palpal organ, retroventral view. (H) Palpal organ, distal view (dorsal side up). (I) Dorsal part of contrategulum, distal and slightly prolateral view. (J) Distal part of tarsus, ventral view. (K-M) Same, retrolateral and slightly proximal view. Scale lines 1.0 mm (A; B, F-G; C-E; H-I; J; K-M). Abbreviations: tp - triangular distal protrusion of longitudinal rib on sclerotised part of embolus proper; so - angular retrolateral side of sperm duct opening." figureDoi="http://doi.org/10.5281/zenodo.7761515" httpUri="https://zenodo.org/record/7761515/files/figure.png" pageId="4" pageNumber="378">Fig. 10B</figureCitation>
|
||
); tibial apophysis only slightly set back from anterior margin of tibia (
|
||
<figureCitation box="[797,884,1742,1765]" captionStart="Fig" captionStartId="19.[151,186,1745,1766]" captionTargetBox="[156,1430,263,1692]" captionTargetId="figure-155@19.[151,1436,257,1698]" captionTargetPageId="19" captionText="Fig. 10. Liphistius ferox sp. nov., left palps of four males; holotype (A-B, E-K), male from Naw Bubaw, matured IV.2019 (C, M), male from Thandaung Gyi, matured 7.X.2014 (D), male from Thandaung Gyi, matured 18.IX.2016 (L). (A) Tarsus and palpal organ, ventral view. (B) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (C-D) Embolus complex, ventral view. (F) Embolus proper, prodorsal view. (G) Palpal organ, retroventral view. (H) Palpal organ, distal view (dorsal side up). (I) Dorsal part of contrategulum, distal and slightly prolateral view. (J) Distal part of tarsus, ventral view. (K-M) Same, retrolateral and slightly proximal view. Scale lines 1.0 mm (A; B, F-G; C-E; H-I; J; K-M). Abbreviations: tp - triangular distal protrusion of longitudinal rib on sclerotised part of embolus proper; so - angular retrolateral side of sperm duct opening." figureDoi="http://doi.org/10.5281/zenodo.7761515" httpUri="https://zenodo.org/record/7761515/files/figure.png" pageId="4" pageNumber="378">Fig. 10J</figureCitation>
|
||
) .................................
|
||
<taxonomicName box="[1138,1220,1742,1765]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
|
||
<emphasis box="[1138,1220,1742,1765]" italics="true" pageId="4" pageNumber="378">L. ferox</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1226,1309,1742,1765]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="378">
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
6(
|
||
<emphasis box="[173,186,1774,1797]" italics="true" pageId="4" pageNumber="378">4</emphasis>
|
||
) Cumulus distinctly elevated (
|
||
<figureCitation box="[560,663,1774,1797]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="4" pageNumber="378">Figs 21B</figureCitation>
|
||
,
|
||
<figureCitation box="[676,723,1774,1797]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="4" pageNumber="378">23B</figureCitation>
|
||
,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[735,902,1774,1797]" pageId="4" pageNumber="378" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[774,823,1774,1797]" italics="true" pageId="4" pageNumber="378">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3D-F); paracymbium with distinctly conical distal side (21A, 23A,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[483,643,1806,1829]" pageId="4" pageNumber="378" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[519,566,1806,1829]" italics="true" pageId="4" pageNumber="378">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3D-F); proximal ledge on retrodorsal side of contrategulum indistinct or small (
|
||
<figureCitation box="[343,422,1838,1861]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="4" pageNumber="378">Figs 21</figureCitation>
|
||
E-F, H-I, L-N, 23F, H-I) ............................................................................................................ 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
- Cumulus not or only slightly elevated (
|
||
<figureCitation box="[657,724,1870,1893]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="4" pageNumber="378">Figs 4</figureCitation>
|
||
H-J, 6J, L-N, 17G, 19E-F); paracymbium with indistinctly conical, almost plane distal side (
|
||
<figureCitation box="[509,576,1902,1925]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="4" pageNumber="378">Figs 4</figureCitation>
|
||
H-J, 6J, L-N, 17G, 19E-F); proximal ledge on retrodorsal side of contrategulum pronounced (
|
||
<figureCitation box="[382,449,1934,1957]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="4" pageNumber="378">Figs 4</figureCitation>
|
||
A-C, 6A-D, 17A, C, 19A) ..................................................................................................... 9
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="5" lastPageNumber="379" pageId="4" pageNumber="378">
|
||
<paragraph blockId="4.[151,1437,1102,2053]" pageId="4" pageNumber="378">
|
||
<keyLead pageId="4" pageNumber="378">
|
||
7(
|
||
<emphasis box="[173,186,1966,1989]" italics="true" pageId="4" pageNumber="378">6</emphasis>
|
||
) Distal edge of contrategulum evenly rounded, without a prodorsal protrusion in distal view (
|
||
<figureCitation box="[1237,1315,1966,1989]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="4" pageNumber="378">Fig. 23</figureCitation>
|
||
E-F); distal margin of tegulum distinctly elevated, developed as a short and wide keel (
|
||
<figureCitation box="[1042,1119,1998,2021]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="4" pageNumber="378">Fig. 23</figureCitation>
|
||
I-M); paracymbium without a retrolateral-proximal heel (
|
||
<figureCitation box="[530,626,2030,2053]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="4" pageNumber="378">Fig. 23A</figureCitation>
|
||
) ...................................................................
|
||
<taxonomicName box="[1098,1219,2030,2053]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[1098,1219,2030,2053]" italics="true" pageId="4" pageNumber="378">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1226,1309,2030,2053]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Distal edge of contrategulum with an angular or arched prodorsal protrusion in distal view (
|
||
<figureCitation box="[1207,1303,258,281]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="5" pageNumber="379">Fig. 21A</figureCitation>
|
||
, E-F, H-I, L,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[242,403,290,313]" pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[279,326,290,313]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3B); distal margin of tegulum distinctly, indistinctly or not elevated (
|
||
<figureCitation box="[1193,1329,290,313]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="5" pageNumber="379">
|
||
Fig. 21
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
K, M, Yu
|
||
<emphasis box="[1376,1423,290,313]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
</figureCitation>
|
||
: fig. 3A-B, D-E); paracymbium with or without a retrolateral-proximal heel (
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[1131,1296,322,345]" pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[1171,1218,322,345]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3E,
|
||
<figureCitation captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="5" pageNumber="379">Fig. 21A</figureCitation>
|
||
) ...................................................................................................................................................................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
8(
|
||
<emphasis box="[173,186,386,409]" italics="true" pageId="5" pageNumber="379">7</emphasis>
|
||
) Distal edge of contrategulum with an angular prodorsal protrusion (
|
||
<figureCitation box="[972,1071,386,409]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="5" pageNumber="379">Fig. 21A</figureCitation>
|
||
, E-F, H-I, L) and with a narrowly rounded dorsal apex (
|
||
<figureCitation box="[475,551,418,441]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="5" pageNumber="379">Fig. 21</figureCitation>
|
||
E-G); distal edge of tegulum not or only indistinctly elevated (
|
||
<figureCitation box="[1224,1361,418,441]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="5" pageNumber="379">Fig. 21K, M</figureCitation>
|
||
); paracymbium without a retrolateral-proximal heel (
|
||
<figureCitation box="[744,841,450,473]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="5" pageNumber="379">Fig. 21A</figureCitation>
|
||
) ....................................
|
||
<taxonomicName box="[1108,1219,450,473]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[1108,1219,450,473]" italics="true" pageId="5" pageNumber="379">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1226,1309,450,473]" pageId="5" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Distal edge of contrategulum with an arched prodorsal protrusion and with a pointed dorsal apex (
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[1273,1432,482,505]" pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[1311,1357,482,505]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3B); distal edge of tegulum distinctly elevated, developed as a short, wide keel (
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[1151,1316,514,537]" pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[1191,1239,514,537]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3A-B, D-E); paracymbium with a retrolateral-proximal heel (
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[822,984,546,569]" pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[860,907,546,569]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 3E) ......................
|
||
<taxonomicName box="[1240,1348,546,569]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[1240,1348,546,569]" italics="true" pageId="5" pageNumber="379">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
(China)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
9(
|
||
<emphasis box="[173,186,578,601]" italics="true" pageId="5" pageNumber="379">6</emphasis>
|
||
) Proventral process of contrategulum very wide, with widely truncate apex in distal view (
|
||
<figureCitation box="[1209,1341,578,601]" captionStart="Fig" captionStartId="32.[151,186,1942,1963]" captionTargetBox="[159,1427,1125,1893]" captionTargetId="figure-353@32.[151,1436,1117,1903]" captionTargetPageId="32" captionText="Fig. 17. Liphistius lordae, left palps of two males: male n°1 (A-B, D-G), male n°2 (C). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Contrategulum, distal view. (D) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Palpal organ and distal margin of tarsus, prolateral view. (F) Tibial apophysis, retrolateral and slightly proximal view. (G) Proximal part of tarsus, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761535" httpUri="https://zenodo.org/record/7761535/files/figure.png" pageId="5" pageNumber="379">Fig. 17A, C</figureCitation>
|
||
); heel of paracymbium protruding proximally (
|
||
<figureCitation box="[646,743,610,633]" captionStart="Fig" captionStartId="32.[151,186,1942,1963]" captionTargetBox="[159,1427,1125,1893]" captionTargetId="figure-353@32.[151,1436,1117,1903]" captionTargetPageId="32" captionText="Fig. 17. Liphistius lordae, left palps of two males: male n°1 (A-B, D-G), male n°2 (C). (A) Palpal organ, distal view (dorsal side up). (B) Dorsal part of contrategulum and embolus proper, distal and slightly prolateral view. (C) Contrategulum, distal view. (D) Distal part of palpal organ, ventral view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Palpal organ and distal margin of tarsus, prolateral view. (F) Tibial apophysis, retrolateral and slightly proximal view. (G) Proximal part of tarsus, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761535" httpUri="https://zenodo.org/record/7761535/files/figure.png" pageId="5" pageNumber="379">Fig. 17G</figureCitation>
|
||
) ...................................................................
|
||
<taxonomicName authorityName="Platnick & Sedgwick" authorityYear="1984" box="[1211,1308,610,633]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="lordae">
|
||
<emphasis box="[1211,1308,610,633]" italics="true" pageId="5" pageNumber="379">L. lordae</emphasis>
|
||
</taxonomicName>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Proventral process of contrategulum conical, with narrowly rounded or narrowly and obliquely truncate apex in distal view (
|
||
<figureCitation box="[372,439,674,697]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="5" pageNumber="379">Figs 4</figureCitation>
|
||
A-C, 6A-D, 19A); heel of paracymbium protruding retrolaterally (
|
||
<figureCitation box="[1146,1213,674,697]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="5" pageNumber="379">Figs 4</figureCitation>
|
||
H-J, 6J, L-N, 19E-F) ............................................................................................................................................................................. 10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
10(
|
||
<emphasis box="[187,200,738,761]" italics="true" pageId="5" pageNumber="379">9</emphasis>
|
||
) Retrolateral-proximal heel of paracymbium carrying 2-3 enlarged spicules (
|
||
<figureCitation box="[1057,1134,738,761]" captionStart="Fig" captionStartId="35.[151,186,1288,1309]" captionTargetBox="[156,1431,261,1232]" captionTargetId="figure-280@35.[151,1436,257,1238]" captionTargetPageId="35" captionText="Fig. 19. Liphistius pyinoolwin, palps of two males: matured 16.X.2014 (A-E), matured 24.IX.2014 (F). (A) Left palpal organ, distal view (dorsal side up). (B) Left embolus complex and dorsal part of contrategulum, distal and slightly proventral view. (C) Left palpal organ and distal margin of tarsus, prolateral view. (D) Left palpal organ, ventral and slightly proximal view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Distal part of left tibia and proximal part of tarsus, ventral view. (F) Distal margin of right tibia and proximal part of tarsus, ventral view. Scale lines 1.0 mm (A-D; E-F). Abbreviation: l - lobate prolateral protrusion on base of embolus complex." figureDoi="http://doi.org/10.5281/zenodo.7761541" httpUri="https://zenodo.org/record/7761541/files/figure.png" pageId="5" pageNumber="379">Fig. 19</figureCitation>
|
||
E-F); base of embolus complex with a lobate prolateral protrusion in distal view (
|
||
<figureCitation box="[822,897,770,793]" captionStart="Fig" captionStartId="35.[151,186,1288,1309]" captionTargetBox="[156,1431,261,1232]" captionTargetId="figure-280@35.[151,1436,257,1238]" captionTargetPageId="35" captionText="Fig. 19. Liphistius pyinoolwin, palps of two males: matured 16.X.2014 (A-E), matured 24.IX.2014 (F). (A) Left palpal organ, distal view (dorsal side up). (B) Left embolus complex and dorsal part of contrategulum, distal and slightly proventral view. (C) Left palpal organ and distal margin of tarsus, prolateral view. (D) Left palpal organ, ventral and slightly proximal view; bent and adpressed proximal edge of tegulum indicated by arrow. (E) Distal part of left tibia and proximal part of tarsus, ventral view. (F) Distal margin of right tibia and proximal part of tarsus, ventral view. Scale lines 1.0 mm (A-D; E-F). Abbreviation: l - lobate prolateral protrusion on base of embolus complex." figureDoi="http://doi.org/10.5281/zenodo.7761541" httpUri="https://zenodo.org/record/7761541/files/figure.png" pageId="5" pageNumber="379">Fig. 19</figureCitation>
|
||
A-B) ............................
|
||
<taxonomicName box="[1161,1309,770,793]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="pyinoolwin">
|
||
<emphasis box="[1161,1309,770,793]" italics="true" pageId="5" pageNumber="379">L. pyinoolwin</emphasis>
|
||
</taxonomicName>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Retrolateral-proximal heel of paracymbium without enlarged spicules but normal spicules or stiff bristles instead (
|
||
<figureCitation box="[250,313,834,857]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="5" pageNumber="379">Fig. 4</figureCitation>
|
||
H-J, 6J, L-N); base of embolus complex without prolateral protrusion in distal view (
|
||
<figureCitation box="[1219,1303,834,857]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="5" pageNumber="379">Fig. 4C</figureCitation>
|
||
,
|
||
<figureCitation box="[1314,1348,834,857]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="5" pageNumber="379">6A</figureCitation>
|
||
) ..... 11
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
11(
|
||
<emphasis box="[186,213,866,889]" italics="true" pageId="5" pageNumber="379">10</emphasis>
|
||
) Depth/length ratio of tibial apophysis of palp ~ 1.4 (
|
||
<figureCitation box="[799,881,866,889]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="5" pageNumber="379">Fig. 4F</figureCitation>
|
||
); retrolateral-proximal heel of paracymbium widely rounded (
|
||
<figureCitation box="[344,407,898,921]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="5" pageNumber="379">Fig. 4</figureCitation>
|
||
H-J); prolateral part of distal edge of contrategulum indistinctly elevated (
|
||
<figureCitation box="[1193,1308,898,921]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="5" pageNumber="379">Fig. 4E, G</figureCitation>
|
||
); no invagination between para-embolic plate and retroventral edge of embolus complex (
|
||
<figureCitation box="[1082,1145,930,953]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="5" pageNumber="379">Fig. 4</figureCitation>
|
||
C-D) ...
|
||
<taxonomicName authorityName="Schwendinger" authorityYear="1998" box="[1241,1317,930,953]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="lahu">
|
||
<emphasis box="[1241,1317,930,953]" italics="true" pageId="5" pageNumber="379">L. lahu</emphasis>
|
||
</taxonomicName>
|
||
(Thailand)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Depth/length ratio of tibial apophysis of palp ~ 2.2 (
|
||
<figureCitation box="[790,875,962,985]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="5" pageNumber="379">Fig. 6O</figureCitation>
|
||
); retrolateral-proximal heel of paracymbium narrowly rounded (
|
||
<figureCitation box="[369,444,994,1017]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="5" pageNumber="379">Fig. 6J</figureCitation>
|
||
, L-N); prolateral part of distal edge of contrategulum distinctly elevated (
|
||
<figureCitation box="[1215,1277,994,1017]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="5" pageNumber="379">Fig. 6</figureCitation>
|
||
H-I); a shallow invagination between para-embolic plate and retroventral edge of embolus complex (
|
||
<figureCitation box="[1149,1211,1026,1049]" captionStart="Fig" captionStartId="13.[151,186,1821,1842]" captionTargetId="figure-7@12.[151,1436,257,2016]" captionTargetPageId="12" captionText="Fig. 6. Liphistius metopiae sp. nov., palps of male holotype (A, D-K, M, O) and of male paratype (B-C, L, N). (A) Left palpal organ, distal view (dorsal side up). (B) Distal edge of contrategulum and proventral process of left palp, distal view. (C-D) Same of right palp, distal view. (E) Left palpal organ, retrolateral view. (F) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (G) Same, ventral and slightly proximal view. (H) Entire palpal organ and distal margin of tibia, ventral and more proximal view than in G. (I) Same, prolateral view. (J) Left tarsus and palpal organ, ventral view. (K) Distal part of left tibia, ventral view. (L) Left tarsus and subtegulum, ventral view. (M) Right paracymbium, ventral view.(N) Right paracymbium and distal part of tibia, ventral view. (O) Distal part of tibia, retrolateral and slightly proximal view. Scale line 1.0 mm (A-I; J-O).Abbreviations: de - prolateral part of distal contrategular edge; h - retrolateral-proximal heel of paracymbium." figureDoi="http://doi.org/10.5281/zenodo.7761507" httpUri="https://zenodo.org/record/7761507/files/figure.png" pageId="5" pageNumber="379">Fig. 6</figureCitation>
|
||
F-H) ........................ ...............................................................................................................................
|
||
<taxonomicName box="[1102,1229,1058,1081]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="metopiae" status="sp. nov.">
|
||
<emphasis box="[1102,1229,1058,1081]" italics="true" pageId="5" pageNumber="379">L. metopiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1235,1318,1058,1081]" pageId="5" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Thailand)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
12(
|
||
<emphasis box="[187,200,1090,1113]" italics="true" pageId="5" pageNumber="379">1</emphasis>
|
||
) Portions of carapace, legs and palps of females and of last immature instars of males orange-coloured (
|
||
<figureCitation box="[1347,1410,1090,1113]" captionStart="Fig" captionStartId="9.[151,186,1970,1991]" captionTargetBox="[160,1426,651,1915]" captionTargetId="figure-160@9.[151,1436,641,1924]" captionTargetPageId="9" captionText="Fig. 3. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius ferox sp. nov., male paratype (matured 1.IX.2014) from Pathi Village. (B) Same species, female paratype from Thandaung Gyi. (C) Liphistius pinlaung, female from near Tong Htiwaw Temple. (D) Liphistius birmanicus from Yado, male (matured 20.IX.2017). (E) Same species, large female n°1. (F, H) Same species, large female n°2. (G) Same species, small female. (I) Same species, large female n°3. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761495" httpUri="https://zenodo.org/record/7761495/files/figure.png" pageId="5" pageNumber="379">Fig. 3</figureCitation>
|
||
EI) ......................................................................................................................................................................... 13
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" box="[151,1437,1154,1177]" pageId="5" pageNumber="379">
|
||
<keyLead box="[151,1437,1154,1177]" pageId="5" pageNumber="379">
|
||
- Body and legs uniformly dark, or body brown and legs more or less distinctly annulated (
|
||
<figureCitation box="[1194,1263,1154,1177]" captionStart="Fig" captionStartId="8.[151,186,1932,1953]" captionTargetBox="[155,1431,783,1902]" captionTargetId="figure-159@8.[151,1436,777,1910]" captionTargetPageId="8" captionText="Fig. 2. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius tung sp. nov., male holotype.(B) Liphistius pyinoolwin, male (matured 1.X.2014) from the Anisakan Falls; note exuviae of ectoparasitic mites (Ljunghia sp.) on the opisthosoma. (C) Liphistius platnicki sp. nov., female paratype (moulted 10.III.2021) from near Loi Kan Village. (D, F) Liphistius cupreus sp. nov., male paratype from near Kong Nyaung Village. (E) Same species, different male paratype. (G) Same species, female paratype (moulted 13.II.2021). (H-I) Same species, different female paratype (moulted 21.V.2021). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761491" httpUri="https://zenodo.org/record/7761491/files/figure.png" pageId="5" pageNumber="379">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation box="[1275,1286,1154,1177]" captionStart="Fig" captionStartId="9.[151,186,1970,1991]" captionTargetBox="[160,1426,651,1915]" captionTargetId="figure-160@9.[151,1436,641,1924]" captionTargetPageId="9" captionText="Fig. 3. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius ferox sp. nov., male paratype (matured 1.IX.2014) from Pathi Village. (B) Same species, female paratype from Thandaung Gyi. (C) Liphistius pinlaung, female from near Tong Htiwaw Temple. (D) Liphistius birmanicus from Yado, male (matured 20.IX.2017). (E) Same species, large female n°1. (F, H) Same species, large female n°2. (G) Same species, small female. (I) Same species, large female n°3. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761495" httpUri="https://zenodo.org/record/7761495/files/figure.png" pageId="5" pageNumber="379">3</figureCitation>
|
||
A-C) ....... 14
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
13(
|
||
<emphasis box="[187,214,1186,1209]" italics="true" pageId="5" pageNumber="379">12</emphasis>
|
||
) Large spiders (CL and CW of females up to 14.73 and 13.27, respectively); posterior stalk of vulval plate usually wider than long or rarely as wide as long, its posterior margin usually straight, rarely widely rounded (
|
||
<figureCitation box="[1333,1410,1218,1241]" captionStart="Fig" captionStartId="26.[151,186,1819,1840]" captionTargetId="figure-9@27.[151,1436,257,2027]" captionTargetPageId="27" captionText="Fig. 14. Liphistius birmanicus, vulval plates (all taken from exuviae) of ten females from the type locality (A-M) and of one female from the Shan State (N-O); dorsal view [A, D (with uterus externus attached), G, J, N], ventral view (B-C, E-F, H-I, K-M, O). (A-B) Moult of 6.II.2021. (C) Moult of 27.XII.2020. (D) Moult of 20.XI.2021. (E) Moult of 19.II.2021 (same specimen as D). (F) Moult of 21.II.2021. (G-H) Moult of 25.XII.2020. (I) Moult of 14.V.2021. (J-K) Moult of 20.III.2021. (L) Moult of 26.X.2017. (M) Moult of 24.VIII.2017. (N-O) Moult of 2.IV.2018. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761529" httpUri="https://zenodo.org/record/7761529/files/figure.png" pageId="5" pageNumber="379">Fig. 14</figureCitation>
|
||
) ........................................................................................................................................
|
||
<taxonomicName authorityName="Thorell" authorityYear="1897" box="[1160,1308,1250,1273]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis box="[1160,1308,1250,1273]" italics="true" pageId="5" pageNumber="379">L. birmanicus</emphasis>
|
||
</taxonomicName>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Medium-sized spiders [CL and CW of female holotype (fully grown?) 7.02 and 6.16, respectively]; posterior stalk of vulval plate clearly longer than wide, its posterior margin narrowly rounded or angular (
|
||
<bibRefCitation author="Aung K. P. P. & Xu X. & Lwin W. W. & Sang M. Z. & Yu L. & Liu H. & Liu F. & Li D." pageId="5" pageNumber="379" pagination="29 - 39" refId="ref37364" refString="Aung K. P. P., Xu X., Lwin W. W., Sang M. Z., Yu L., Liu H., Liu F., Li D. 2019. Two new species of the primitively segmented spider genus Liphistius Schiodte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 29 - 39." type="journal article" year="2019">
|
||
Aung
|
||
<emphasis box="[1374,1423,1314,1337]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
: fig. 3B-E) ....................................................................................................................
|
||
<taxonomicName box="[1206,1308,1346,1369]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="hpruso">
|
||
<emphasis box="[1206,1308,1346,1369]" italics="true" pageId="5" pageNumber="379">L. hpruso</emphasis>
|
||
</taxonomicName>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
14(
|
||
<emphasis box="[187,214,1378,1401]" italics="true" pageId="5" pageNumber="379">12</emphasis>
|
||
) Very large spiders (CL and CW in females up to 18.32 and 16.61, respectively); anterior lobes of poreplate folded ventrad (
|
||
<figureCitation box="[336,454,1410,1433]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="21.[153,188,1845,1866]" captionStartId-1="21.[151,186,1651,1672]" captionTargetBox-1="[156,1436,901,1604]" captionTargetId-0="figure-7@20.[151,1439,257,2021]" captionTargetId-1="figure-411@21.[151,1439,879,1625]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="Fig. 11. Liphistius ferox, vulvae of seven females; dorsal view (A, C, E, G, I, K, N, P), ventral view (B, D, F, H, J, L-M, O). (A-B) Thandaung Gyi, moulted 21.VII.2015; with uterus externus attached (in B hidden on other side). (C-D) Thandaung Gyi, moulted 8.1.2015. (E-F) Thandaung Gyi, moulted 15.VII.2015; with uterus externus attached (in F only pocket of one side visible). (G-H) Thandaung Gyi, moulted 16.VII.2015; vulval plate. (I-J) Same female; isolated uterus externus. (K-L) Naw Bubaw, moulted 26.II.2015. (M) Allotype; Thandaung Lay, moulted 10.VII.2015, vulval plate. (N) Same female; isolated uterus externus. (O) Thandaung Gyi, moulted 20.IX.2014; vulval plate. (P) Same female; isolated uterus externus. Arrows indicating paired lateral pockets of still attached membranous uterus externus. Scale lines 1.0 mm." captionText-1="Fig. 12. Liphistius ferox, vulvae of three females; dorsal view (A, D, F), ventral view (B-C, E). (A-B) Thandaung Gyi, moulted 26.X.2014; vulval plate. (C) Thandaung Gyi, moulted 11.1.2015; vulval plate. (D) Same female; isolated uterus externus. (E) Thandaung Gyi, moulted 6.XII.2014; vulval plate. (F) Same female; isolated uterus externus. Scale lines 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7761521" figureDoi-1="http://doi.org/10.5281/zenodo.7761519" httpUri-0="https://zenodo.org/record/7761521/files/figure.png" httpUri-1="https://zenodo.org/record/7761519/files/figure.png" pageId="5" pageNumber="379">Figs 11-12</figureCitation>
|
||
); body and legs uniformly very dark, without annulations (
|
||
<figureCitation box="[1083,1145,1410,1433]" captionStart="Fig" captionStartId="9.[151,186,1970,1991]" captionTargetBox="[160,1426,651,1915]" captionTargetId="figure-160@9.[151,1436,641,1924]" captionTargetPageId="9" captionText="Fig. 3. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius ferox sp. nov., male paratype (matured 1.IX.2014) from Pathi Village. (B) Same species, female paratype from Thandaung Gyi. (C) Liphistius pinlaung, female from near Tong Htiwaw Temple. (D) Liphistius birmanicus from Yado, male (matured 20.IX.2017). (E) Same species, large female n°1. (F, H) Same species, large female n°2. (G) Same species, small female. (I) Same species, large female n°3. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761495" httpUri="https://zenodo.org/record/7761495/files/figure.png" pageId="5" pageNumber="379">Fig. 3</figureCitation>
|
||
A-B) ................................. .....................................................................................................................................
|
||
<taxonomicName box="[1138,1220,1442,1465]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
|
||
<emphasis box="[1138,1220,1442,1465]" italics="true" pageId="5" pageNumber="379">L. ferox</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1226,1309,1442,1465]" pageId="5" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Medium-sized spiders [CL and CW in females up to 8.50 and 7.90, respectively (for
|
||
<taxonomicName authorityName="Platnick & Sedgwick" authorityYear="1984" box="[1135,1232,1474,1497]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="lordae">
|
||
<emphasis box="[1135,1232,1474,1497]" italics="true" pageId="5" pageNumber="379">L. lordae</emphasis>
|
||
</taxonomicName>
|
||
)]; anterior lobes of poreplate not folded ventrad (
|
||
<figureCitation box="[556,625,1506,1529]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="5" pageNumber="379">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation box="[636,651,1506,1529]" captionStart="Fig" captionStartId="13.[149,184,1407,1428]" captionTargetBox="[154,1434,257,1369]" captionTargetId="figure-257@13.[149,1437,254,1374]" captionTargetPageId="13" captionText="Fig. 7. Liphistius metopiae sp. nov., vulval plates of five female paratypes (all taken from exuviae); dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Allotype; moult of 23.IV.1998. (D-E) Moult of 16.IV.1999. (E) Moult of 24.II.2000. (G-H) Moult of 28.VI.2001. (I) Moult of 4.III.1999. Scale lines 1.0 mm. Abbreviations: al - anterior lobe of poreplate; alp - anterolateral process of poreplate." figureDoi="http://doi.org/10.5281/zenodo.7761503" httpUri="https://zenodo.org/record/7761503/files/figure.png" pageId="5" pageNumber="379">7</figureCitation>
|
||
,
|
||
<figureCitation box="[663,678,1506,1529]" captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="5" pageNumber="379">9</figureCitation>
|
||
,
|
||
<figureCitation box="[689,715,1506,1529]" captionStart="Fig" captionStartId="31.[151,186,1817,1838]" captionTargetBox="[155,1436,267,1763]" captionTargetId="figure-78@31.[151,1440,255,1768]" captionTargetPageId="31" captionText="Fig. 16. Liphistius pinlaung (A-J) and Liphistius sp. (specimen from Kalaw; K-L), vulval plates of seven females (all taken from exuviae), dorsal view (A, D, G, I, K) and ventral view (B-C, E-F, H, J, L). (A-B) Moult of 10.II.2019. (C) Moult of 7.XII.2018. (D-E) Moult of 10.IV.2021. (F) Moult of 21.X.2018. (G-H) Moult of 16.V.2020. (I-J) Moult of 8.III.2020. (K-L) Moult of 15.VIII.2017. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761533" httpUri="https://zenodo.org/record/7761533/files/figure.png" pageId="5" pageNumber="379">16</figureCitation>
|
||
A-J, 18, 20, 22, 24,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[925,1084,1506,1529]" pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[961,1008,1506,1529]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 4); body brown to dark, legs annulated or not .................................................................................................................................................. 15
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
15(
|
||
<emphasis box="[187,214,1570,1593]" italics="true" pageId="5" pageNumber="379">14</emphasis>
|
||
) Posterior stalk as wide as or wider than poreplate; anterior lobes of poreplate reduced to 1-2 clusters of vesicles on each side (
|
||
<figureCitation box="[388,471,1602,1625]" captionStart="Fig" captionStartId="40.[151,186,1942,1963]" captionTargetBox="[155,1436,262,1898]" captionTargetId="figure-107@40.[151,1439,257,1908]" captionTargetPageId="40" captionText="Fig. 22. Liphistius cupreus sp. nov., vulval plates of ten females (all taken from exuviae), dorsal view (E with uterus externus attached, H, K, N), ventral view (A-D, I-J, L-M, O), ventral and slightly anterior view (P), posterior view (G), and isolated uterus externus (F). (A) Moult of 10.II.2019. (B) Moult of 12.V.2020. (C) Moult of 5.VII.2019. (D-F) Moult of 22.XI.2020. (G) Moult of VI.2019. (H-I) Moult of 13.II.2021. (J) Allotype; moult of 19.I.2019. (K-L) Moult of 14.II.2021. (M) Moult of 1.V.2021. (N-P) Moult of 23.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761549" httpUri="https://zenodo.org/record/7761549/files/figure.png" pageId="5" pageNumber="379">Figs 22</figureCitation>
|
||
,
|
||
<figureCitation box="[482,511,1602,1625]" captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="5" pageNumber="379">24</figureCitation>
|
||
,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[521,681,1602,1625]" pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[557,604,1602,1625]" italics="true" pageId="5" pageNumber="379">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: fig. 4) ............................................................................................... 16
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Posterior stalk narrower than poreplate; anterior lobes of poreplate fully developed (
|
||
<figureCitation box="[1139,1208,1634,1657]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="5" pageNumber="379">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation box="[1219,1234,1634,1657]" captionStart="Fig" captionStartId="13.[149,184,1407,1428]" captionTargetBox="[154,1434,257,1369]" captionTargetId="figure-257@13.[149,1437,254,1374]" captionTargetPageId="13" captionText="Fig. 7. Liphistius metopiae sp. nov., vulval plates of five female paratypes (all taken from exuviae); dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Allotype; moult of 23.IV.1998. (D-E) Moult of 16.IV.1999. (E) Moult of 24.II.2000. (G-H) Moult of 28.VI.2001. (I) Moult of 4.III.1999. Scale lines 1.0 mm. Abbreviations: al - anterior lobe of poreplate; alp - anterolateral process of poreplate." figureDoi="http://doi.org/10.5281/zenodo.7761503" httpUri="https://zenodo.org/record/7761503/files/figure.png" pageId="5" pageNumber="379">7</figureCitation>
|
||
,
|
||
<figureCitation box="[1246,1261,1634,1657]" captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="5" pageNumber="379">9</figureCitation>
|
||
,
|
||
<figureCitation box="[1273,1299,1634,1657]" captionStart="Fig" captionStartId="31.[151,186,1817,1838]" captionTargetBox="[155,1436,267,1763]" captionTargetId="figure-78@31.[151,1440,255,1768]" captionTargetPageId="31" captionText="Fig. 16. Liphistius pinlaung (A-J) and Liphistius sp. (specimen from Kalaw; K-L), vulval plates of seven females (all taken from exuviae), dorsal view (A, D, G, I, K) and ventral view (B-C, E-F, H, J, L). (A-B) Moult of 10.II.2019. (C) Moult of 7.XII.2018. (D-E) Moult of 10.IV.2021. (F) Moult of 21.X.2018. (G-H) Moult of 16.V.2020. (I-J) Moult of 8.III.2020. (K-L) Moult of 15.VIII.2017. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761533" httpUri="https://zenodo.org/record/7761533/files/figure.png" pageId="5" pageNumber="379">16</figureCitation>
|
||
A-J, 18, 20) ............................................................................................................................................................................. 18
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
16(
|
||
<emphasis box="[187,214,1698,1721]" italics="true" pageId="5" pageNumber="379">15</emphasis>
|
||
) Ventral side of vulval plate with a widely separated pair of “bulging margins” laterally between poreplate and posterior stalk; posterior part of genital atrium strongly curved ventrad (
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[1009,1170,1730,1753]" pageId="5" pageNumber="379" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[1047,1101,1730,1753]" italics="true" pageId="5" pageNumber="379">et al.</emphasis>
|
||
, 2021
|
||
</bibRefCitation>
|
||
: fig. 4) ........................... ....................................................................................................................................................
|
||
<taxonomicName box="[1240,1348,1762,1785]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[1240,1348,1762,1785]" italics="true" pageId="5" pageNumber="379">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
(China)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" box="[151,1437,1794,1817]" pageId="5" pageNumber="379">
|
||
<keyLead box="[151,1437,1794,1817]" pageId="5" pageNumber="379">- “Bulging margins” absent; posterior part of genital atrium not strongly curved ventrad .................................. 17</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
17(
|
||
<emphasis box="[187,214,1826,1849]" italics="true" pageId="5" pageNumber="379">16</emphasis>
|
||
) Anterior margin of poreplate with 1-2 rather small clusters of vesicles on each side; lateral margins of poreplate without anterolateral processes; posterior stalk much shorter than poreplate (
|
||
<figureCitation box="[1048,1127,1858,1881]" captionStart="Fig" captionStartId="40.[151,186,1942,1963]" captionTargetBox="[155,1436,262,1898]" captionTargetId="figure-107@40.[151,1439,257,1908]" captionTargetPageId="40" captionText="Fig. 22. Liphistius cupreus sp. nov., vulval plates of ten females (all taken from exuviae), dorsal view (E with uterus externus attached, H, K, N), ventral view (A-D, I-J, L-M, O), ventral and slightly anterior view (P), posterior view (G), and isolated uterus externus (F). (A) Moult of 10.II.2019. (B) Moult of 12.V.2020. (C) Moult of 5.VII.2019. (D-F) Moult of 22.XI.2020. (G) Moult of VI.2019. (H-I) Moult of 13.II.2021. (J) Allotype; moult of 19.I.2019. (K-L) Moult of 14.II.2021. (M) Moult of 1.V.2021. (N-P) Moult of 23.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761549" httpUri="https://zenodo.org/record/7761549/files/figure.png" pageId="5" pageNumber="379">Fig. 22</figureCitation>
|
||
); prefoveal setae on carapace absent .....................................................................................................................
|
||
<taxonomicName box="[1108,1219,1890,1913]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[1108,1219,1890,1913]" italics="true" pageId="5" pageNumber="379">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1226,1309,1890,1913]" pageId="5" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" pageId="5" pageNumber="379">
|
||
<keyLead pageId="5" pageNumber="379">
|
||
- Anterior margin of poreplate with only one rather large cluster of vesicles on each side; lateral margin of poreplate with a small knob-shaped anterolateral process on each side; posterior stalk almost as long as poreplate (
|
||
<figureCitation captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="5" pageNumber="379">Fig. 24</figureCitation>
|
||
); prefoveal setae present ..................................................................................
|
||
<taxonomicName box="[1098,1219,1986,2009]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[1098,1219,1986,2009]" italics="true" pageId="5" pageNumber="379">L. platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1226,1309,1986,2009]" pageId="5" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="379">
|
||
<paragraph blockId="5.[151,1437,258,2073]" box="[151,1437,2018,2041]" pageId="5" pageNumber="379">
|
||
<keyLead box="[151,1437,2018,2041]" pageId="5" pageNumber="379">
|
||
18(
|
||
<emphasis box="[187,214,2018,2041]" italics="true" pageId="5" pageNumber="379">15</emphasis>
|
||
) No knob-shaped anterolateral processes on ventral side of poreplate (
|
||
<figureCitation box="[979,1048,2018,2041]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="5" pageNumber="379">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation box="[1059,1074,2018,2041]" captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="5" pageNumber="379">9</figureCitation>
|
||
,
|
||
<figureCitation box="[1086,1113,2018,2041]" captionStart="Fig" captionStartId="33.[151,186,1998,2019]" captionTargetBox="[154,1432,751,1943]" captionTargetId="figure-212@33.[151,1436,748,1947]" captionTargetPageId="33" captionText="Fig. 18. Liphistius lordae, vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H); all taken from exuviae. (A-B) Female n°1. (C-D) Female n°2. (E-F) Female n°3. (G-H) Female n°4; note pathologically enlarged vesicle on left side of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761539" httpUri="https://zenodo.org/record/7761539/files/figure.png" pageId="5" pageNumber="379">18</figureCitation>
|
||
) ........................................ 19
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,258,2073]" box="[151,1437,2050,2073]" pageId="5" pageNumber="379">
|
||
<keyLead box="[151,1437,2050,2073]" pageId="5" pageNumber="379">
|
||
- Each lateral side of poreplate ventrally with a knob-shaped anterolateral process (
|
||
<figureCitation box="[1111,1180,2050,2073]" captionStart="Fig" captionStartId="13.[149,184,1407,1428]" captionTargetBox="[154,1434,257,1369]" captionTargetId="figure-257@13.[149,1437,254,1374]" captionTargetPageId="13" captionText="Fig. 7. Liphistius metopiae sp. nov., vulval plates of five female paratypes (all taken from exuviae); dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Allotype; moult of 23.IV.1998. (D-E) Moult of 16.IV.1999. (E) Moult of 24.II.2000. (G-H) Moult of 28.VI.2001. (I) Moult of 4.III.1999. Scale lines 1.0 mm. Abbreviations: al - anterior lobe of poreplate; alp - anterolateral process of poreplate." figureDoi="http://doi.org/10.5281/zenodo.7761503" httpUri="https://zenodo.org/record/7761503/files/figure.png" pageId="5" pageNumber="379">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation box="[1191,1217,2050,2073]" captionStart="Fig" captionStartId="31.[151,186,1817,1838]" captionTargetBox="[155,1436,267,1763]" captionTargetId="figure-78@31.[151,1440,255,1768]" captionTargetPageId="31" captionText="Fig. 16. Liphistius pinlaung (A-J) and Liphistius sp. (specimen from Kalaw; K-L), vulval plates of seven females (all taken from exuviae), dorsal view (A, D, G, I, K) and ventral view (B-C, E-F, H, J, L). (A-B) Moult of 10.II.2019. (C) Moult of 7.XII.2018. (D-E) Moult of 10.IV.2021. (F) Moult of 21.X.2018. (G-H) Moult of 16.V.2020. (I-J) Moult of 8.III.2020. (K-L) Moult of 15.VIII.2017. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761533" httpUri="https://zenodo.org/record/7761533/files/figure.png" pageId="5" pageNumber="379">16</figureCitation>
|
||
A-J, 20) ............ 21
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="380">
|
||
<paragraph blockId="6.[151,1437,258,825]" pageId="6" pageNumber="380">
|
||
<keyLead pageId="6" pageNumber="380">
|
||
19(
|
||
<emphasis box="[187,214,258,281]" italics="true" pageId="6" pageNumber="380">18</emphasis>
|
||
) Anterior lobes of poreplate very wide and continuous with lateral poreplate margins, no step between them step (
|
||
<figureCitation box="[250,329,290,313]" captionStart="Fig" captionStartId="33.[151,186,1998,2019]" captionTargetBox="[154,1432,751,1943]" captionTargetId="figure-212@33.[151,1436,748,1947]" captionTargetPageId="33" captionText="Fig. 18. Liphistius lordae, vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H); all taken from exuviae. (A-B) Female n°1. (C-D) Female n°2. (E-F) Female n°3. (G-H) Female n°4; note pathologically enlarged vesicle on left side of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761539" httpUri="https://zenodo.org/record/7761539/files/figure.png" pageId="6" pageNumber="380">Fig. 18</figureCitation>
|
||
); no annulations on legs and palps
|
||
<emphasis box="[705,1308,290,313]" italics="true" pageId="6" pageNumber="380">
|
||
...........................................................................
|
||
<taxonomicName authorityName="Platnick & Sedgwick" authorityYear="1984" box="[1211,1308,290,313]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="lordae">L. lordae</taxonomicName>
|
||
</emphasis>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,258,825]" pageId="6" pageNumber="380">
|
||
<keyLead pageId="6" pageNumber="380">
|
||
- Anterior lobes of poreplate mostly rather narrow, with a more or less distinct step between them and lateral poreplate margins (
|
||
<figureCitation box="[400,469,354,377]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="6" pageNumber="380">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation box="[481,496,354,377]" captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="6" pageNumber="380">9</figureCitation>
|
||
); legs and palps with or without annulations ............................................................... 20
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="380">
|
||
<paragraph blockId="6.[151,1437,258,825]" pageId="6" pageNumber="380">
|
||
<keyLead pageId="6" pageNumber="380">
|
||
20(
|
||
<emphasis box="[187,214,386,409]" italics="true" pageId="6" pageNumber="380">19</emphasis>
|
||
) Step between anterior lobes and lateral margins of poreplate pronounced; length of posterior stalk clearly more than half of poreplate length (
|
||
<figureCitation box="[559,623,418,441]" captionStart="Fig" captionStartId="17.[151,186,1966,1987]" captionTargetBox="[160,1422,958,1921]" captionTargetId="figure-390@17.[151,1428,949,1929]" captionTargetPageId="17" captionText="Fig. 9. Liphistius tung sp. nov., vulval plates of four females; dorsal view (A, C, E, G) and ventral view (B, D, F, H). (A-B) Allotype (taken from specimen). (C-D) Large paratype (taken from specimen). (E-F) Medium-sized paratype (from exuvia 18.IV.2013). (G-H) Small paratype (from exuvia 28.III.2012). Arrow indicating pronounced step between anterior lobes and lateral margin of poreplate. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761513" httpUri="https://zenodo.org/record/7761513/files/figure.png" pageId="6" pageNumber="380">Fig. 9</figureCitation>
|
||
); legs and palps not annulated ..........................
|
||
<taxonomicName box="[1144,1219,418,441]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="tung" status="sp. nov.">
|
||
<emphasis box="[1144,1219,418,441]" italics="true" pageId="6" pageNumber="380">L. tung</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1226,1309,418,441]" pageId="6" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,258,825]" pageId="6" pageNumber="380">
|
||
<keyLead pageId="6" pageNumber="380">
|
||
- Step between anterior lobes and lateral margins of poreplate rather indistinct; length of posterior stalk less than or little more than half of poreplate length (
|
||
<figureCitation box="[705,769,482,505]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="6" pageNumber="380">Fig. 5</figureCitation>
|
||
); legs and palps annulated .........................
|
||
<taxonomicName authorityName="Schwendinger" authorityYear="1998" box="[1241,1317,482,505]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="lahu">
|
||
<emphasis box="[1241,1317,482,505]" italics="true" pageId="6" pageNumber="380">L. lahu</emphasis>
|
||
</taxonomicName>
|
||
(Thailand)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="380">
|
||
<paragraph blockId="6.[151,1437,258,825]" pageId="6" pageNumber="380">
|
||
<keyLead pageId="6" pageNumber="380">
|
||
21(
|
||
<emphasis box="[187,214,514,537]" italics="true" pageId="6" pageNumber="380">18</emphasis>
|
||
) Posterior stalk of vulval plate clearly longer than wide or only slightly wider than long, its anterior portion with a long constriction (
|
||
<figureCitation box="[455,535,546,569]" captionStart="Fig" captionStartId="36.[151,186,1326,1347]" captionTargetBox="[155,1436,271,1286]" captionTargetId="figure-385@36.[151,1439,257,1291]" captionTargetPageId="36" captionText="Fig. 20. Liphistius pyinoolwin, vulval plates of three females (all taken from exuviae), dorsal view (A, D, F), ventral view (B, E, G) and ventral and slightly anterior view (C). (A-C) Moult of 1.IX.2017. (D-E) Moult of 15.X.2017. (F-G) Moult of 22.I.2016. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761545" httpUri="https://zenodo.org/record/7761545/files/figure.png" pageId="6" pageNumber="380">Fig. 20</figureCitation>
|
||
;
|
||
<bibRefCitation author="Xu X. & Yu L. & Aung K. P. P. & Yu L. & Liu F. & Lwin W. W. & Sang M. Z. & Li D." box="[546,708,546,569]" pageId="6" pageNumber="380" pagination="41 - 58" refId="ref38558" refString="Xu X., Yu L., Aung K. P. P., Yu L., Liu F., Lwin W. W., Sang M. Z., Li D. 2021. A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae). ZooKeys 1031: 41 - 58." type="journal article" year="2021">
|
||
Xu
|
||
<emphasis box="[585,632,546,569]" italics="true" pageId="6" pageNumber="380">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
: figs 5-7) ..................................................
|
||
<taxonomicName box="[1161,1309,546,569]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="pyinoolwin">
|
||
<emphasis box="[1161,1309,546,569]" italics="true" pageId="6" pageNumber="380">L. pyinoolwin</emphasis>
|
||
</taxonomicName>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,258,825]" pageId="6" pageNumber="380">
|
||
<keyLead pageId="6" pageNumber="380">
|
||
- Posterior stalk of vulval plate clearly wider than long, its anterior portion with a short constriction (
|
||
<figureCitation box="[1285,1353,578,601]" captionStart="Fig" captionStartId="13.[149,184,1407,1428]" captionTargetBox="[154,1434,257,1369]" captionTargetId="figure-257@13.[149,1437,254,1374]" captionTargetPageId="13" captionText="Fig. 7. Liphistius metopiae sp. nov., vulval plates of five female paratypes (all taken from exuviae); dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Allotype; moult of 23.IV.1998. (D-E) Moult of 16.IV.1999. (E) Moult of 24.II.2000. (G-H) Moult of 28.VI.2001. (I) Moult of 4.III.1999. Scale lines 1.0 mm. Abbreviations: al - anterior lobe of poreplate; alp - anterolateral process of poreplate." figureDoi="http://doi.org/10.5281/zenodo.7761503" httpUri="https://zenodo.org/record/7761503/files/figure.png" pageId="6" pageNumber="380">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation box="[1364,1390,578,601]" captionStart="Fig" captionStartId="31.[151,186,1817,1838]" captionTargetBox="[155,1436,267,1763]" captionTargetId="figure-78@31.[151,1440,255,1768]" captionTargetPageId="31" captionText="Fig. 16. Liphistius pinlaung (A-J) and Liphistius sp. (specimen from Kalaw; K-L), vulval plates of seven females (all taken from exuviae), dorsal view (A, D, G, I, K) and ventral view (B-C, E-F, H, J, L). (A-B) Moult of 10.II.2019. (C) Moult of 7.XII.2018. (D-E) Moult of 10.IV.2021. (F) Moult of 21.X.2018. (G-H) Moult of 16.V.2020. (I-J) Moult of 8.III.2020. (K-L) Moult of 15.VIII.2017. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761533" httpUri="https://zenodo.org/record/7761533/files/figure.png" pageId="6" pageNumber="380">16</figureCitation>
|
||
A-J;
|
||
<bibRefCitation author="Aung K. P. P. & Xu X. & Lwin W. W. & Sang M. Z. & Yu L. & Liu H. & Liu F. & Li D." box="[242,431,610,633]" pageId="6" pageNumber="380" pagination="29 - 39" refId="ref37364" refString="Aung K. P. P., Xu X., Lwin W. W., Sang M. Z., Yu L., Liu H., Liu F., Li D. 2019. Two new species of the primitively segmented spider genus Liphistius Schiodte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 29 - 39." type="journal article" year="2019">
|
||
Aung
|
||
<emphasis box="[308,355,610,633]" italics="true" pageId="6" pageNumber="380">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
: fig. 5) .................................................................................................................................... 22
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="380">
|
||
<paragraph blockId="6.[151,1437,258,825]" pageId="6" pageNumber="380">
|
||
<keyLead pageId="6" pageNumber="380">
|
||
22(
|
||
<emphasis box="[187,214,642,665]" italics="true" pageId="6" pageNumber="380">21</emphasis>
|
||
) Anterior margin of poreplate mostly with a deep and narrowly U-shaped invagination between anterior lobes (indistinct in
|
||
<figureCitation box="[384,467,674,697]" captionStart="Fig" captionStartId="13.[149,184,1407,1428]" captionTargetBox="[154,1434,257,1369]" captionTargetId="figure-257@13.[149,1437,254,1374]" captionTargetPageId="13" captionText="Fig. 7. Liphistius metopiae sp. nov., vulval plates of five female paratypes (all taken from exuviae); dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Allotype; moult of 23.IV.1998. (D-E) Moult of 16.IV.1999. (E) Moult of 24.II.2000. (G-H) Moult of 28.VI.2001. (I) Moult of 4.III.1999. Scale lines 1.0 mm. Abbreviations: al - anterior lobe of poreplate; alp - anterolateral process of poreplate." figureDoi="http://doi.org/10.5281/zenodo.7761503" httpUri="https://zenodo.org/record/7761503/files/figure.png" pageId="6" pageNumber="380">Fig. 7E</figureCitation>
|
||
); posterior stalk of vulval plate small in comparison with poreplate (
|
||
<figureCitation box="[1198,1263,674,697]" captionStart="Fig" captionStartId="13.[149,184,1407,1428]" captionTargetBox="[154,1434,257,1369]" captionTargetId="figure-257@13.[149,1437,254,1374]" captionTargetPageId="13" captionText="Fig. 7. Liphistius metopiae sp. nov., vulval plates of five female paratypes (all taken from exuviae); dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Allotype; moult of 23.IV.1998. (D-E) Moult of 16.IV.1999. (E) Moult of 24.II.2000. (G-H) Moult of 28.VI.2001. (I) Moult of 4.III.1999. Scale lines 1.0 mm. Abbreviations: al - anterior lobe of poreplate; alp - anterolateral process of poreplate." figureDoi="http://doi.org/10.5281/zenodo.7761503" httpUri="https://zenodo.org/record/7761503/files/figure.png" pageId="6" pageNumber="380">Fig. 7</figureCitation>
|
||
); legs and palps annulated ..............................................................................................................
|
||
<taxonomicName box="[1102,1229,706,729]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="metopiae" status="sp. nov.">
|
||
<emphasis box="[1102,1229,706,729]" italics="true" pageId="6" pageNumber="380">L. metopiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1235,1318,706,729]" pageId="6" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Thailand)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,258,825]" pageId="6" pageNumber="380">
|
||
<keyLead pageId="6" pageNumber="380">
|
||
- Anterior margin of poreplate with a less deep, more widely U-shaped or widely V-shaped invagination between anterior lobes; posterior stalk of vulval plate distinctly larger in comparison with poreplate (
|
||
<figureCitation box="[1224,1300,770,793]" captionStart="Fig" captionStartId="31.[151,186,1817,1838]" captionTargetBox="[155,1436,267,1763]" captionTargetId="figure-78@31.[151,1440,255,1768]" captionTargetPageId="31" captionText="Fig. 16. Liphistius pinlaung (A-J) and Liphistius sp. (specimen from Kalaw; K-L), vulval plates of seven females (all taken from exuviae), dorsal view (A, D, G, I, K) and ventral view (B-C, E-F, H, J, L). (A-B) Moult of 10.II.2019. (C) Moult of 7.XII.2018. (D-E) Moult of 10.IV.2021. (F) Moult of 21.X.2018. (G-H) Moult of 16.V.2020. (I-J) Moult of 8.III.2020. (K-L) Moult of 15.VIII.2017. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761533" httpUri="https://zenodo.org/record/7761533/files/figure.png" pageId="6" pageNumber="380">Fig. 16</figureCitation>
|
||
A-J,
|
||
<bibRefCitation author="Aung K. P. P. & Xu X. & Lwin W. W. & Sang M. Z. & Yu L. & Liu H. & Liu F. & Li D." pageId="6" pageNumber="380" pagination="29 - 39" refId="ref37364" refString="Aung K. P. P., Xu X., Lwin W. W., Sang M. Z., Yu L., Liu H., Liu F., Li D. 2019. Two new species of the primitively segmented spider genus Liphistius Schiodte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 29 - 39." type="journal article" year="2019">
|
||
Aung
|
||
<emphasis italics="true" pageId="6" pageNumber="380">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
: fig. 5); legs and palps not annulated ..................................................................
|
||
<taxonomicName box="[1185,1308,802,825]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="pinlaung">
|
||
<emphasis box="[1185,1308,802,825]" italics="true" pageId="6" pageNumber="380">L. pinlaung</emphasis>
|
||
</taxonomicName>
|
||
(Myanmar)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |