220 lines
34 KiB
XML
220 lines
34 KiB
XML
<document id="61263FC68A9E625389DAD3F1B2182063" ID-CLB-Dataset="45454" ID-DOI="10.11646/zootaxa.3815.4.6" ID-GBIF-Dataset="8c9e9a24-cc17-4abc-a27b-08a81ad9b982" ID-ISSN="1175-5326" ID-Zenodo-Dep="230822" ID-ZooBank="13A3E4D1-A4D4-4C6B-AB43-5268594EA221" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460438438669" checkinUser="plazi" docAuthor="Husana, Daniel Edison M., Yamamuro, Masumi & Ng, Peter K. L." docDate="2014" docId="03DC8797FF86FFB8FF1880A026BB2BEF" docLanguage="en" docName="zt03815p574.pdf" docOrigin="Zootaxa 3815 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sundathelphusa vienae Husana, Yamamuro & Ng, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="572" masterDocId="FFE5FFEFFF82FFB0FF8F860322142B17" masterDocTitle="Two new species of freshwater crabs of the genus Sundathelphusa Bott, 1969 (Decapoda: Brachyura: Gecarcinucidae) from caves in Luzon, Philippines" masterLastPageNumber="574" masterPageNumber="565" pageNumber="569" updateTime="1698332556256" updateUser="ExternalLinkService">
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<mods:titleInfo id="6E58AD6224ED5035FFAF8166237926A7">
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<mods:title id="E0F2CF0072EF3BDA4ACDC991BB13E454">Two new species of freshwater crabs of the genus Sundathelphusa Bott, 1969 (Decapoda: Brachyura: Gecarcinucidae) from caves in Luzon, Philippines</mods:title>
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<mods:name id="0AC946A685ADC228807F06BE27BD91E4" type="personal">
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<mods:roleTerm id="FE5E39DE1945D7CC52B7C56D65D8196E">Author</mods:roleTerm>
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<mods:namePart id="080A8536A164AF45973DB4EB4AF7B6F6">Husana, Daniel Edison M.</mods:namePart>
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</mods:name>
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<mods:name id="1F1330D36DB286B389194A7CB72EB8B6" type="personal">
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<mods:roleTerm id="14E472AEB9F306ABE88A697934AD40CE">Author</mods:roleTerm>
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<mods:namePart id="6774FDC9437C7F43CE049CDE969A5B5F">Yamamuro, Masumi</mods:namePart>
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</mods:name>
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<mods:name id="2E0F71935FED19670F653B75F1939792" type="personal">
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<mods:role id="B4FFA4D11653FBD9571C71EB4AFAA031">
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<mods:roleTerm id="00A33D03EDC8CE866B750AFA38EA5578">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="554283AB91635FB44A4FEB8A744CA70A">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="D66C920563B3A50DCB71760D62B8B2FF">text</mods:typeOfResource>
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<mods:title id="F0E46E7D97435A5BAE92A9D325E3CD6F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6D18E9AB2C947DA008D406730BD8D558">
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<mods:date id="B50E5C9168322EDC45DFFBEFE32B26CA">2014</mods:date>
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<mods:number id="4E8873D781295D832B7E53D73942F6C2">3815</mods:number>
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<mods:number id="BE01F673C93DD9B00D592DF482DFFF71">4</mods:number>
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<mods:identifier id="DF385967433C53DF462DFD4B575C8233" type="DOI">10.11646/zootaxa.3815.4.6</mods:identifier>
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<treatment id="03DC8797FF86FFB8FF1880A026BB2BEF" ID-DOI="http://doi.org/10.5281/zenodo.6132784" ID-GBIF-Taxon="119507212" ID-Zenodo-Dep="6132784" LSID="urn:lsid:plazi:treatment:03DC8797FF86FFB8FF1880A026BB2BEF" httpUri="http://treatment.plazi.org/id/03DC8797FF86FFB8FF1880A026BB2BEF" lastPageId="8" lastPageNumber="572" pageId="4" pageNumber="569">
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<subSubSection id="C36F650AFF86FFB4FF1880A023762DF7" pageId="4" pageNumber="569" type="nomenclature">
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<paragraph id="8BCA3681FF86FFB4FF1880A020112DAA" blockId="4.[151,517,1699,1760]" box="[151,517,1699,1725]" pageId="4" pageNumber="569">
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<heading id="D08281EDFF86FFB4FF1880A020112DAA" bold="true" box="[151,517,1699,1725]" fontSize="10" level="2" pageId="4" pageNumber="569" reason="2">
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<emphasis id="B901EA93FF86FFB4FF1880A020112DAA" bold="true" box="[151,517,1699,1725]" pageId="4" pageNumber="569">
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<taxonomicName id="4C754D02FF86FFB4FF1880A023B12DAB" ID-CoL="53GCD" box="[151,421,1699,1724]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="vienae" status="sp. nov.">
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<emphasis id="B901EA93FF86FFB4FF1880A023B12DAB" bold="true" box="[151,421,1699,1724]" italics="true" pageId="4" pageNumber="569">Sundathelphusa vienae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23257E8FF86FFB4FE2380A720112DAA" box="[428,517,1700,1725]" pageId="4" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCA3681FF86FFB4FF1880C423762DF7" blockId="4.[151,517,1699,1760]" box="[151,354,1735,1760]" pageId="4" pageNumber="569">
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(
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<figureCitation id="134E2A04FF86FFB4FF1080C4231F2DF7" box="[159,267,1735,1760]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="5.[151,250,1869,1891]" captionStartId-1="6.[151,250,1325,1347]" captionStartId-2="7.[151,250,1644,1666]" captionTargetBox-0="[313,1273,733,1847]" captionTargetBox-1="[160,1424,199,1307]" captionTargetBox-2="[313,1273,193,1621]" captionTargetId-0="figure@5.[313,1273,733,1848]" captionTargetId-1="figure@6.[151,1436,193,1317]" captionTargetId-2="figure@7.[313,1273,193,1623]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 4. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." captionText-1="FIGURE 5. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen [telson and somite 6 damaged]; B, left second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 2.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." captionText-2="FIGURE 6. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, buccal field and thoracic sternum; B, abdomen. Scale bars, A, B = 5.0 mm." httpUri-0="https://zenodo.org/record/230826/files/figure.png" httpUri-1="https://zenodo.org/record/230827/files/figure.png" httpUri-2="https://zenodo.org/record/230828/files/figure.png" pageId="4" pageNumber="569">Figs. 4–6</figureCitation>
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,
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<figureCitation id="134E2A04FF86FFB4FE9980CB23302DF7" box="[278,292,1736,1760]" captionStart="FIGURE 7. A, B" captionStartId="8.[151,250,1219,1241]" captionTargetBox="[151,1436,303,1196]" captionTargetId="figure@8.[151,1436,303,1196]" captionTargetPageId="8" captionText="FIGURE 7. A, B, Sundathelphusa danae sp. nov., female (20.4 × 16.8 mm) (NMCR 39093), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines; C, D, Sundathelphusa vienae sp. nov., paratype female (19.8 × 16.5 mm) (NSMT-Cr 22317), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, C, female abdomens; B, D, thoracic sternums showing vulvae." httpUri="https://zenodo.org/record/230829/files/figure.png" pageId="4" pageNumber="569">7</figureCitation>
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C, D)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F650AFF86FFB4FF18810C202E2C67" pageId="4" pageNumber="569" type="materials_examined">
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<paragraph id="8BCA3681FF86FFB4FF18810C202E2C67" blockId="4.[151,1436,1807,2013]" pageId="4" pageNumber="569">
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<emphasis id="B901EA93FF86FFB4FF18810C23972C3F" bold="true" box="[151,387,1807,1832]" pageId="4" pageNumber="569">Material examined.</emphasis>
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<typeStatus id="54CE8823FF86FFB4FE02810C23EE2C3F" box="[397,506,1807,1832]" pageId="4" pageNumber="569" type="holotype">Holotype</typeStatus>
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: male (21.3 ×
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<quantity id="4C8D9B64FF86FFB4FD2A811321182C3F" box="[677,780,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="4" pageNumber="569" unit="mm" value="17.9">17.9 mm</quantity>
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) (
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<collectionCode id="ED64AE44FF86FFB4FCA98113216E2C3F" box="[806,890,1808,1832]" pageId="4" pageNumber="569">NMCR</collectionCode>
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39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon,
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<collectingCountry id="F3627611FF86FFB4FED2813723CA2C5A" box="[349,478,1844,1869]" name="Philippines" pageId="4" pageNumber="569">Philippines</collectingCountry>
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, 14˚
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<geoCoordinate id="EE415046FF86FFB4FD86813720652C5A" box="[521,625,1844,1869]" direction="north" orientation="latitude" pageId="4" pageNumber="569" precision="55" value="1.0">01.000N</geoCoordinate>
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, 121˚
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<geoCoordinate id="EE415046FF86FFB4FD25813721042C5A" box="[682,784,1844,1869]" direction="east" orientation="longitude" pageId="4" pageNumber="569" precision="55" value="47.775">47.775E</geoCoordinate>
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, coll. D.E.M. Husana,
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<date id="FFCB1041FF86FFB4FB82813726D62C5B" box="[1037,1218,1844,1869]" pageId="4" pageNumber="569" value="2009-12">December 2009</date>
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.
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<typeStatus id="54CE8823FF86FFB4FB44813627262C5A" box="[1227,1330,1845,1869]" pageId="4" pageNumber="569" type="paratype">Paratype</typeStatus>
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:
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<specimenCount id="9D73FD08FF86FFB4FAB3813727882C5A" box="[1340,1436,1844,1869]" pageId="4" pageNumber="569" type="female">1 female</specimenCount>
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(19.8 ×
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<quantity id="4C8D9B64FF86FFB4FF60815423402C67" box="[239,340,1879,1904]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="4" pageNumber="569" unit="mm" value="16.5">16.5 mm</quantity>
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) (NSMT-Cr 22317).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F650AFF86FFB7FF48817F275A2C5A" lastPageId="7" lastPageNumber="572" pageId="4" pageNumber="569" type="description">
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<paragraph id="8BCA3681FF86FFB5FF48817F2653290F" blockId="4.[151,1436,1807,2013]" lastBlockId="5.[151,1436,151,680]" lastPageId="5" lastPageNumber="570" pageId="4" pageNumber="569">
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<emphasis id="B901EA93FF86FFB4FF48817F23452C82" bold="true" box="[199,337,1916,1941]" pageId="4" pageNumber="569">Description</emphasis>
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. Carapace distinctively quadrate (
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<figureCitation id="134E2A04FF86FFB4FD55817F21342C83" box="[730,800,1916,1941]" captionStart="FIGURE 4" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[313,1273,733,1847]" captionTargetId="figure@5.[313,1273,733,1848]" captionTargetPageId="5" captionText="FIGURE 4. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230826/files/figure.png" pageId="4" pageNumber="569">Fig. 4</figureCitation>
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A), widest at anterior quarter, broader than long, dorsal surface convex longitudinally, dorso-ventrally inflated, regions distinct; branchial regions covered with distinct oblique striae; cervical groove prominent; H-shaped gastric groove distinct. Epigastric cristae distinct, edges sharp, separated by distinct median furrow; postorbital cristae, sharp; epigastric, postorbital cristae, not confluent; epibranchial teeth, postorbital cristae not confluent. Frontal margin (
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<figureCitation id="134E2A04FF87FFB5FC3186BF26072BC3" box="[958,1043,188,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[313,1273,733,1847]" captionTargetId="figure@5.[313,1273,733,1848]" captionTargetPageId="5" captionText="FIGURE 4. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230826/files/figure.png" pageId="5" pageNumber="570">Figs. 4</figureCitation>
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B, 5C) protruded, gently sinuous, deflexed. External orbital tooth triangular, produced anteriorly; outer margin convex, longer than inner margin; epibranchial tooth distinct, small, well separated from external orbital tooth by notch; anterolateral margin convex, armed with small granules, clearly demarcated from posterolateral margin; posterolateral margin almost straight to gently concave, with oblique striae converging gradually towards posterior margin of carapace. Frontal median triangle (
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<figureCitation id="134E2A04FF87FFB5FF73876C235F2A9F" box="[252,331,367,392]" captionStart="FIGURE 4" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[313,1273,733,1847]" captionTargetId="figure@5.[313,1273,733,1848]" captionTargetPageId="5" captionText="FIGURE 4. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230826/files/figure.png" pageId="5" pageNumber="570">Figs. 4</figureCitation>
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B, 5C) distinct but not complete, with protruded dorsal margin; orbit well demarcated; supraorbital margin smooth; infraorbital margin beaded with distinct granules; outer edge reaching, fused with anterolateral margin; suborbital, subbranchial regions covered with few scattered oblique striae of various sizes; pterygostomial region with streak of oblique ridges on upper part. Posterior margin of epistome (
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<figureCitation id="134E2A04FF87FFB5FBD787DF26BF2AE3" box="[1112,1195,476,501]" captionStart="FIGURE 4" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[313,1273,733,1847]" captionTargetId="figure@5.[313,1273,733,1848]" captionTargetPageId="5" captionText="FIGURE 4. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230826/files/figure.png" pageId="5" pageNumber="570">Figs. 4</figureCitation>
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B, 5D) with 3 lobes, median lobe, triangular; lateral lobes wider, distal margin more protruded anteriorly.
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</paragraph>
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<paragraph id="8BCA3681FF87FFB5FF48842620D629B0" blockId="5.[151,1436,151,680]" pageId="5" pageNumber="570">
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Eyes well developed (
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<figureCitation id="134E2A04FF87FFB5FE3084272017292B" box="[447,515,548,573]" captionStart="FIGURE 4" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[313,1273,733,1847]" captionTargetId="figure@5.[313,1273,733,1848]" captionTargetPageId="5" captionText="FIGURE 4. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230826/files/figure.png" pageId="5" pageNumber="570">Fig. 4</figureCitation>
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B), occupying almost entire orbit. Third maxilliped ischium (
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<figureCitation id="134E2A04FF87FFB5FB3E842726E7292A" box="[1201,1267,548,573]" captionStart="FIGURE 6" captionStartId="7.[151,250,1644,1666]" captionTargetBox="[313,1273,193,1621]" captionTargetId="figure@7.[313,1273,193,1623]" captionTargetPageId="7" captionText="FIGURE 6. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, buccal field and thoracic sternum; B, abdomen. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230828/files/figure.png" pageId="5" pageNumber="570">Fig. 6</figureCitation>
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A) rectangular, much longer than broad, bearing distinct oblique submedian sulcus closer to mesial margin; merus quadrate, distalexternal angle convex, upswept; tip of exopod not reaching midpoint of outer margin of merus, with flagellum reaching beyond edge of mesial margin of merus.
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</paragraph>
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<caption id="DF0A6609FF87FFB5FF18814E27602C95" httpUri="https://zenodo.org/record/230826/files/figure.png" pageId="5" pageNumber="570" targetBox="[313,1273,733,1847]" targetPageId="5">
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<paragraph id="8BCA3681FF87FFB5FF18814E27602C95" blockId="5.[151,1436,1869,1922]" pageId="5" pageNumber="570">
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<emphasis id="B901EA93FF87FFB5FF18814E23072C75" bold="true" box="[151,275,1869,1891]" pageId="5" pageNumber="570">FIGURE 4.</emphasis>
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<taxonomicName id="4C754D02FF87FFB5FE95814E20102C74" box="[282,516,1869,1891]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="570" phylum="Arthropoda" rank="species" species="vienae" status="sp. nov.">
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<emphasis id="B901EA93FF87FFB5FE95814E20102C74" box="[282,516,1869,1891]" italics="true" pageId="5" pageNumber="570">Sundathelphusa vienae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23257E8FF87FFB5FD84814D20482C74" box="[523,604,1870,1891]" pageId="5" pageNumber="570" rank="species">
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<emphasis id="B901EA93FF87FFB5FD84814D20482C74" bold="true" box="[523,604,1870,1891]" pageId="5" pageNumber="570">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm.
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</paragraph>
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</caption>
|
||
<caption id="DF0A6609FF84FFB6FF18832E21092EAA" httpUri="https://zenodo.org/record/230827/files/figure.png" pageId="6" pageNumber="571" targetBox="[160,1424,199,1307]" targetPageId="6">
|
||
<paragraph id="8BCA3681FF84FFB6FF18832E21092EAA" blockId="6.[151,1436,1325,1469]" pageId="6" pageNumber="571">
|
||
<emphasis id="B901EA93FF84FFB6FF18832E23072E55" bold="true" box="[151,275,1325,1347]" pageId="6" pageNumber="571">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C754D02FF84FFB6FE95832E20102E54" box="[282,516,1325,1347]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="vienae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF84FFB6FE95832E20102E54" box="[282,516,1325,1347]" italics="true" pageId="6" pageNumber="571">Sundathelphusa vienae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257E8FF84FFB6FD84832E20482E55" box="[523,604,1325,1346]" pageId="6" pageNumber="571" rank="species">
|
||
<emphasis id="B901EA93FF84FFB6FD84832E20482E55" bold="true" box="[523,604,1325,1346]" pageId="6" pageNumber="571">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen [telson and somite 6 damaged]; B, left second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G1, ventral view; F, left G1, dorsal view; G, left G2; H, terminal segment of left G1, ventral view; I, terminal segment of left G1, dorsal view; J, flagellum of left G2. Scale bars, A, B = 2.0 mm; C, D = 2.0 mm; E‒G = 1.0 mm; H‒J = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA3681FF84FFB6FF4883E627022D51" blockId="6.[151,1436,1509,2038]" pageId="6" pageNumber="571">Chelipeds not noticeably elongated, subequal, stronger in males; margins of merus serrated, dorsal margin with small subdistal tooth; carpus armed with strong distal sharply pointed inner tooth, dorso-ventrally flattened; palm with smooth outer surface; fingers robust, cutting edges armed with medium-sized to large sharp teeth.</paragraph>
|
||
<paragraph id="8BCA3681FF84FFB6FF48805223782DC1" blockId="6.[151,1436,1509,2038]" pageId="6" pageNumber="571">
|
||
Ambulatory legs (
|
||
<figureCitation id="134E2A04FF84FFB6FE1C805223F62D7E" box="[403,482,1617,1642]" captionStart="FIGURE 4" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[313,1273,733,1847]" captionTargetId="figure@5.[313,1273,733,1848]" captionTargetPageId="5" captionText="FIGURE 4. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230826/files/figure.png" pageId="6" pageNumber="571">Figs. 4</figureCitation>
|
||
A, 5B) long, slender, second leg longest, anterior margins of meri distinctly serrated, without subdistal tooth or spine, posterior margins smooth on all legs; second, third pair shorter than carapace length; carpi short, with longitudinal submedian ridge on dorsal surface, widened distally; propodi, dactyli subequal in length.
|
||
</paragraph>
|
||
<paragraph id="8BCA3681FF84FFB6FF4880E226612CE1" blockId="6.[151,1436,1509,2038]" pageId="6" pageNumber="571">
|
||
Male abdominal cavity reaching level of proximal quarter of coxae of chelipeds. Male abdomen (
|
||
<figureCitation id="134E2A04FF84FFB6FA8B80E227442DED" box="[1284,1360,1761,1786]" captionStart="FIGURE 5" captionStartId="6.[151,250,1325,1347]" captionTargetBox="[160,1424,199,1307]" captionTargetId="figure@6.[151,1436,193,1317]" captionTargetPageId="6" captionText="FIGURE 5. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen [telson and somite 6 damaged]; B, left second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 2.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." httpUri="https://zenodo.org/record/230827/files/figure.png" pageId="6" pageNumber="571">Figs. 5</figureCitation>
|
||
A, 6B) narrow, T-shaped; somite 1 short, somite 6 longer than broad; proximal, distal margins of somites 3–5 sinuous; lateral margins of somite 2 convex; lateral margins of somite 3 convex; lateral margins of somite 4 straight, narrows gradually to distal end; somites 3–5 narrow gradually; lateral margins of somite 5 concave, proximally wide distally narrow; lateral margins of somite 6 straight; telson subtriangular, broader than long, lateral margin slightly concave medially, rounded distally. G1 (
|
||
<figureCitation id="134E2A04FF84FFB6FD428196210A2CB9" box="[717,798,1941,1966]" captionStart="FIGURE 5" captionStartId="6.[151,250,1325,1347]" captionTargetBox="[160,1424,199,1307]" captionTargetId="figure@6.[151,1436,193,1317]" captionTargetPageId="6" captionText="FIGURE 5. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen [telson and somite 6 damaged]; B, left second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 2.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." httpUri="https://zenodo.org/record/230827/files/figure.png" pageId="6" pageNumber="571">Figs. 5</figureCitation>
|
||
E, F, H, I) slender, slightly bent outwards; subterminal segment gradually tapering towards terminal segment, segments separated by distinct suture; terminal segment straight, tapering, cylindrical. G2 (
|
||
<figureCitation id="134E2A04FF84FFB6FD9581DE20722CE1" box="[538,614,2013,2038]" captionStart="FIGURE 5" captionStartId="6.[151,250,1325,1347]" captionTargetBox="[160,1424,199,1307]" captionTargetId="figure@6.[151,1436,193,1317]" captionTargetPageId="6" captionText="FIGURE 5. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen [telson and somite 6 damaged]; B, left second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 2.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." httpUri="https://zenodo.org/record/230827/files/figure.png" pageId="6" pageNumber="571">Figs. 5</figureCitation>
|
||
G, J) subequal in length to G1, flagellum short.
|
||
</paragraph>
|
||
<caption id="DF0A6609FF85FFB7FF18806F27232DB6" httpUri="https://zenodo.org/record/230828/files/figure.png" pageId="7" pageNumber="572" targetBox="[313,1273,193,1621]" targetPageId="7">
|
||
<paragraph id="8BCA3681FF85FFB7FF18806F27232DB6" blockId="7.[151,1436,1644,1697]" pageId="7" pageNumber="572">
|
||
<emphasis id="B901EA93FF85FFB7FF18806F23072D96" bold="true" box="[151,275,1644,1666]" pageId="7" pageNumber="572">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C754D02FF85FFB7FE95806F20102D95" box="[282,516,1644,1666]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="vienae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF85FFB7FE95806F20102D95" box="[282,516,1644,1666]" italics="true" pageId="7" pageNumber="572">Sundathelphusa vienae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257E8FF85FFB7FD84806E20482D95" box="[523,604,1645,1666]" pageId="7" pageNumber="572" rank="species">
|
||
<emphasis id="B901EA93FF85FFB7FD84806E20482D95" bold="true" box="[523,604,1645,1666]" pageId="7" pageNumber="572">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines: A, buccal field and thoracic sternum; B, abdomen. Scale bars, A, B = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA3681FF85FFB7FF4880CB219B2C3E" blockId="7.[151,1436,1736,2013]" pageId="7" pageNumber="572">
|
||
<emphasis id="B901EA93FF85FFB7FF4880CB23312DF6" bold="true" box="[199,293,1736,1761]" pageId="7" pageNumber="572">Female.</emphasis>
|
||
Female chelipeds subequal, neither inflated; abdomen ovate, covering entire thoracic sternum, all somites and telson freely articulating; telson broadly triangular with convex lateral margins (
|
||
<figureCitation id="134E2A04FF85FFB7FB1280EF26F42C12" box="[1181,1248,1772,1797]" captionStart="FIGURE 7. A, B" captionStartId="8.[151,250,1219,1241]" captionTargetBox="[151,1436,303,1196]" captionTargetId="figure@8.[151,1436,303,1196]" captionTargetPageId="8" captionText="FIGURE 7. A, B, Sundathelphusa danae sp. nov., female (20.4 × 16.8 mm) (NMCR 39093), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines; C, D, Sundathelphusa vienae sp. nov., paratype female (19.8 × 16.5 mm) (NSMT-Cr 22317), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, C, female abdomens; B, D, thoracic sternums showing vulvae." httpUri="https://zenodo.org/record/230829/files/figure.png" pageId="7" pageNumber="572">Fig. 7</figureCitation>
|
||
C). Vulvae large, subovate, without operculum; on median part of somite 6 (
|
||
<figureCitation id="134E2A04FF85FFB7FCA78113217F2C3E" box="[808,875,1808,1833]" captionStart="FIGURE 7. A, B" captionStartId="8.[151,250,1219,1241]" captionTargetBox="[151,1436,303,1196]" captionTargetId="figure@8.[151,1436,303,1196]" captionTargetPageId="8" captionText="FIGURE 7. A, B, Sundathelphusa danae sp. nov., female (20.4 × 16.8 mm) (NMCR 39093), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines; C, D, Sundathelphusa vienae sp. nov., paratype female (19.8 × 16.5 mm) (NSMT-Cr 22317), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, C, female abdomens; B, D, thoracic sternums showing vulvae." httpUri="https://zenodo.org/record/230829/files/figure.png" pageId="7" pageNumber="572">Fig. 7</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="8BCA3681FF85FFB7FF488137275A2C5A" blockId="7.[151,1436,1736,2013]" box="[199,1358,1844,1869]" pageId="7" pageNumber="572">
|
||
<emphasis id="B901EA93FF85FFB7FF488137235C2C5A" bold="true" box="[199,328,1844,1869]" pageId="7" pageNumber="572">Coloration</emphasis>
|
||
. Dorsal surface of carapace is brownish to orange in life, chelipeds are yellowish to orange.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F650AFF85FFB7FF48815B23272C82" pageId="7" pageNumber="572" type="etymology">
|
||
<paragraph id="8BCA3681FF85FFB7FF48815B23272C82" blockId="7.[151,1436,1736,2013]" pageId="7" pageNumber="572">
|
||
<emphasis id="B901EA93FF85FFB7FF48815B23522C66" bold="true" box="[199,326,1880,1905]" pageId="7" pageNumber="572">Etymology</emphasis>
|
||
. Named after a daughter of the first author, Vien Danille Husana, who accompanied him in some of his fieldwork.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F650AFF85FFB8FF4881A326BB2BEF" lastPageId="8" lastPageNumber="573" pageId="7" pageNumber="572" type="discussion">
|
||
<paragraph id="8BCA3681FF85FFB8FF4881A326BB2BEF" blockId="7.[151,1436,1736,2013]" lastBlockId="8.[151,1436,151,248]" lastPageId="8" lastPageNumber="573" pageId="7" pageNumber="572">
|
||
<emphasis id="B901EA93FF85FFB7FF4881A323202CAE" bold="true" box="[199,308,1952,1977]" pageId="7" pageNumber="572">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D02FF85FFB7FECD81A320512CAF" box="[322,581,1952,1977]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="vienae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF85FFB7FECD81A320512CAF" box="[322,581,1952,1977]" italics="true" pageId="7" pageNumber="572">Sundathelphusa vienae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257E8FF85FFB7FDC381A320B32CAE" box="[588,679,1952,1977]" pageId="7" pageNumber="572" rank="species">
|
||
<emphasis id="B901EA93FF85FFB7FDC381A320B32CAE" bold="true" box="[588,679,1952,1977]" pageId="7" pageNumber="572">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is unusual among
|
||
<taxonomicName id="4C754D02FF85FFB7FCF581A326392CAE" box="[890,1069,1952,1977]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="572" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EA93FF85FFB7FCF581A326392CAE" box="[890,1069,1952,1977]" italics="true" pageId="7" pageNumber="572">Sundathelphusa</emphasis>
|
||
</taxonomicName>
|
||
species in that it has a relatively high carapace like
|
||
<taxonomicName id="4C754D02FF85FFB7FEFB81C623DF2CCB" box="[372,459,1989,2012]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="picta">
|
||
<emphasis id="B901EA93FF85FFB7FEFB81C623DF2CCB" box="[372,459,1989,2012]" italics="true" pageId="7" pageNumber="572">S. picta</emphasis>
|
||
</taxonomicName>
|
||
(von
|
||
<bibRefCitation id="EFE44B70FF85FFB7FD9C81C620AD2CCA" author="Martens" box="[531,697,1988,2013]" pageId="7" pageNumber="572" refString="Martens, E. von (1868) Ueber einige neue crustaceen. Monatsberichte k. - p. Akademie Wissenschaften Berlin, 1868, 608 - 615." type="journal article" year="1868">Martens, 1868</bibRefCitation>
|
||
) and its allies. The new species, however, has a more quadrate carapace and proportionately longer ambulatory legs in contrast to the more ovate inflated carapace and shorter legs of the latter (cf.
|
||
<bibRefCitation id="EFE44B70FF8AFFB8FEF286BE23E52BC3" author="Bott" box="[381,497,188,212]" pageId="8" pageNumber="573" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott 1970</bibRefCitation>
|
||
: pl. 12 figs. 36–38). The G1 is also more slender than in
|
||
<taxonomicName id="4C754D02FF8AFFB8FBE886BE26AE2BC3" box="[1127,1210,189,212]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="picta">
|
||
<emphasis id="B901EA93FF8AFFB8FBE886BE26AE2BC3" box="[1127,1210,189,212]" italics="true" pageId="8" pageNumber="573">S. picta</emphasis>
|
||
</taxonomicName>
|
||
and almost straight, with its tapering pointed terminal segment (
|
||
<figureCitation id="134E2A04FF8AFFB8FD0F86DC20D92BEF" box="[640,717,223,248]" captionStart="FIGURE 5" captionStartId="6.[151,250,1325,1347]" captionTargetBox="[160,1424,199,1307]" captionTargetId="figure@6.[151,1436,193,1317]" captionTargetPageId="6" captionText="FIGURE 5. Sundathelphusa vienae sp. nov., holotype male (21.3 × 17.9 mm) (NMCR 39077), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen [telson and somite 6 damaged]; B, left second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 2.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." httpUri="https://zenodo.org/record/230827/files/figure.png" pageId="8" pageNumber="573">Figs. 5</figureCitation>
|
||
E, F, H, I; versus
|
||
<bibRefCitation id="EFE44B70FF8AFFB8FC1E86E326102BEF" author="Bott" box="[913,1028,224,248]" pageId="8" pageNumber="573" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott 1970</bibRefCitation>
|
||
: pl. 28 fig. 50).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |