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<document id="9B11315D0D03FC09778E015FBE98DDC9" ID-CLB-Dataset="26919" ID-DOI="10.11646/zootaxa.4598.1.1" ID-GBIF-Dataset="475fd53e-c5c6-4a99-bb2e-cc2382f0171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668063" ID-ZooBank="6BA5F086-1014-46B5-B6D5-77FE0CD07AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127411994" checkinUser="plazi" docAuthor="Seldon, David S. &amp; Buckley, Thomas R." docDate="2019" docId="03BBD0260278B77190FBB0B0E9C4FBD8" docLanguage="en" docName="zootaxa.4598.1.1.pdf.imf" docOrigin="Zootaxa 4598 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mecodema tuhoe Seldon &amp; Buckley 2019, sp. n." docType="treatment" docUuid="D89D2690-32FB-4BC5-AD76-0D0330AB99E2" docUuidSource="ZooBank" docVersion="11" lastPageNumber="128" masterDocId="FF82A85E0205B70E906CB267ED0BFFA4" masterDocTitle="The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand" masterLastPageNumber="148" masterPageNumber="1" pageNumber="126" updateTime="1698726038790" updateUser="ExternalLinkService">
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<mods:title id="002D8D7D2C40708F95EE3E41DE7FDB09">The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand</mods:title>
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<mods:namePart id="E10903E007FEED4173140E0B4E7D5D3B">Seldon, David S.</mods:namePart>
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<mods:namePart id="82EF04F9FE3135DAE38BBF9B0BF1B241">Buckley, Thomas R.</mods:namePart>
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<mods:date id="2B42E973BBF7C9081A89D2099118E180">2019</mods:date>
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<treatment id="03BBD0260278B77190FBB0B0E9C4FBD8" ID-DOI="http://doi.org/10.5281/zenodo.5678450" ID-GBIF-Taxon="157062701" ID-Zenodo-Dep="5678450" LSID="urn:lsid:zoobank.org:act:D89D2690-32FB-4BC5-AD76-0D0330AB99E2" httpUri="http://treatment.plazi.org/id/03BBD0260278B77190FBB0B0E9C4FBD8" lastPageId="127" lastPageNumber="128" pageId="125" pageNumber="126">
<subSubSection id="C30832BB0278B77390FBB0B0ECBBFD56" box="[151,432,727,754]" pageId="125" pageNumber="126" type="nomenclature">
<paragraph id="8BAD61300278B77390FBB0B0ECBBFD56" blockId="125.[151,432,727,787]" box="[151,432,727,754]" pageId="125" pageNumber="126">
<heading id="D0E5D65C0278B77390FBB0B0ECBBFD56" bold="true" box="[151,432,727,754]" fontSize="11" level="1" pageId="125" pageNumber="126" reason="1">
<emphasis id="B966BD220278B77390FBB0B0ECBBFD56" bold="true" box="[151,432,727,754]" pageId="125" pageNumber="126">
<taxonomicName id="4C121AB30278B77390FBB0B0EC6DFD55" ID-CoL="8LXZF" authority="Seldon &amp; Buckley, 2019" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[151,358,727,753]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="tuhoe" status="sp. nov.">
<emphasis id="B966BD220278B77390FBB0B0EC6DFD55" bold="true" box="[151,358,727,753]" italics="true" pageId="125" pageNumber="126">Mecodema tuhoe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25500590278B7739102B0BFECBBFD56" box="[366,432,728,754]" pageId="125" pageNumber="126" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30832BB0278B77390FBB09DEC01FCB7" box="[151,266,762,787]" pageId="125" pageNumber="126" type="description">
<paragraph id="8BAD61300278B77390FBB09DEC01FCB7" blockId="125.[151,432,727,787]" box="[151,266,762,787]" pageId="125" pageNumber="126">
<figureCitation id="13297DB50278B77390FBB09DEC0DFCB7" box="[151,262,762,787]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="125" pageNumber="126">Figure 61</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0278B77390FBB125E80FFCDB" pageId="125" pageNumber="126" type="diagnosis">
<paragraph id="8BAD61300278B77390FBB125E80FFCDB" blockId="125.[151,1437,834,2014]" pageId="125" pageNumber="126">
<emphasis id="B966BD220278B77390FBB125EC18FCFF" bold="true" box="[151,275,834,859]" pageId="125" pageNumber="126">Diagnosis:</emphasis>
Distinguishable from other North Island
<taxonomicName id="4C121AB30278B773928EB124EE56FCFF" authorityName="Blanchard" authorityYear="1853" box="[738,861,835,859]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B966BD220278B773928EB124EE56FCFF" box="[738,861,835,859]" italics="true" pageId="125" pageNumber="126">Mecodema</emphasis>
</taxonomicName>
species by:
<emphasis id="B966BD220278B773938AB125EEFFFCFF" bold="true" box="[998,1012,834,859]" pageId="125" pageNumber="126">1</emphasis>
, elytral intervals 16 slightly convex, intervals 79 strongly convex;
<emphasis id="B966BD220278B773919EB101EF0BFCDB" bold="true" box="[498,512,870,895]" pageId="125" pageNumber="126">2</emphasis>
, distinctive shape of the apical portion of the penis lobe (
<figureCitation id="13297DB50278B77394EFB101E9D8FCDB" box="[1155,1235,870,895]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="125" pageNumber="126">Fig. 61</figureCitation>
PL).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0278B77190ABB1EDEF53FD69" lastPageId="127" lastPageNumber="128" pageId="125" pageNumber="126" type="description">
<paragraph id="8BAD61300278B77390ABB1EDEE5DFC63" blockId="125.[151,1437,834,2014]" pageId="125" pageNumber="126">
<emphasis id="B966BD220278B77390ABB1EDEC50FC07" bold="true" box="[199,347,906,931]" pageId="125" pageNumber="126">Description:</emphasis>
Length
<quantity id="4CEACCD50278B77391D7B1EDEF69FC06" box="[443,610,906,931]" metricMagnitude="-2" metricUnit="m" metricValue="2.8649999999999998" metricValueMax="3.18" metricValueMin="2.55" pageId="125" pageNumber="126" unit="mm" value="28.65" valueMax="31.8" valueMin="25.5">25.531.8 mm</quantity>
, pronotal width
<quantity id="4CEACCD50278B7739372B1EDEEA3FC06" box="[798,936,906,931]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="8.8" metricValueMin="6.9" pageId="125" pageNumber="126" unit="mm" value="7.85" valueMax="8.8" valueMin="6.9">6.98.8 mm</quantity>
, elytral width
<quantity id="4CEACCD50278B7739422B1ECE9D1FC06" box="[1102,1242,907,931]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.7" metricValueMin="7.8" pageId="125" pageNumber="126" unit="mm" value="8.75" valueMax="9.7" valueMin="7.8">7.89.7 mm</quantity>
. Colour of entire body matte to glossy black, coxae may be dark reddish-brown.
</paragraph>
<paragraph id="8BAD61300278B77390ABB1B4EE88FAF6" blockId="125.[151,1437,834,2014]" pageId="125" pageNumber="126">
<emphasis id="B966BD220278B77390ABB1B4EC06FC4F" box="[199,269,979,1003]" italics="true" pageId="125" pageNumber="126">Head:</emphasis>
Broad and flat (
<figureCitation id="13297DB50278B77391BDB1B5EF36FC4F" box="[465,573,978,1003]" captionStart="FIGURE 14" captionStartId="20.[151,250,936,958]" captionTargetBox="[171,1412,204,911]" captionTargetId="figure@20.[160,1412,193,921]" captionTargetPageId="20" captionText="FIGURE 14. Diagrammatic cross sections that represent the width and convexity of head or deflection of pronotum (frontal view): (A) broad and flat; (B) broad and convex; (C) narrow and flat; (D) narrow and convex." httpUri="https://zenodo.org/record/2668091/files/figure.png" pageId="125" pageNumber="126">Fig. 14A</figureCitation>
). Vertex with very fine wrinkles forming an isodiametric pattern laterally; vertexal groove defined by punctures and small grooves, broadened laterally (
<figureCitation id="13297DB50278B773946DB191E95FFBAB" box="[1025,1108,1014,1039]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="125" pageNumber="126">Fig. 61</figureCitation>
); small supraorbital puncture bearing 45 setae; 3 well-defined supraorbital grooves; frons with a large and shallow depression each side of midline, otherwise smooth; frontoclypeal suture and tentorial pits (
<figureCitation id="13297DB50278B77393E7B659EEC4FBF3" box="[907,975,1086,1111]" captionStart="FIGURE 9" captionStartId="15.[151,250,1706,1728]" captionTargetBox="[157,1433,759,1683]" captionTargetId="figure@15.[151,1436,757,1683]" captionTargetPageId="15" captionText="FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove." httpUri="https://zenodo.org/record/2668081/files/figure.png" pageId="125" pageNumber="126">Fig. 9</figureCitation>
) indistinct; anterior area of clypeus with narrow grooves laterally, broad scoop-like depression medially, 1 setose puncture each side and 1 medial setose puncture, all bearing 2 setae (medial puncture may be absent) (
<figureCitation id="13297DB50278B77393C9B6E1EEF4FB3B" box="[933,1023,1158,1183]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="125" pageNumber="126">Fig. 61</figureCitation>
). Labrum rectangular, apical edge moderately emarginated with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes rounded (
<figureCitation id="13297DB50278B77390F3B6A9EC0DFB43" box="[159,262,1230,1255]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="125" pageNumber="126">Fig. 12F</figureCitation>
), median process broad and short, moderately angled upward (30°), moderately indentate (
<figureCitation id="13297DB50278B7739549B6A9E885FB43" box="[1317,1422,1230,1255]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="125" pageNumber="126">Fig. 12B</figureCitation>
); mentum setae absent. Submentum sclerite (
<figureCitation id="13297DB50278B77392FAB695EFD5FAAF" box="[662,734,1266,1291]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="125" pageNumber="126">Fig. 8</figureCitation>
) constriction narrow with 6 evenly-spaced setae (large gap medially). Stipes with 2 basal setae. Gula pits small and deep, suture indistinct, gula flat and smooth. Gena with rugose wrinkles forming an isodiametric pattern across entire area.
</paragraph>
<paragraph id="8BAD61300278B77390ABB739EDDDF9EB" blockId="125.[151,1437,834,2014]" pageId="125" pageNumber="126">
<emphasis id="B966BD220278B77390ABB739EC49FAD2" box="[199,322,1374,1398]" italics="true" pageId="125" pageNumber="126">Prothorax:</emphasis>
Prothoracic carina narrow the entire length, crenulations absent with 1014 setae each side (
<figureCitation id="13297DB50278B7739503B739EDBEFA3F" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="125" pageNumber="126">Fig. 61</figureCitation>
), extended to anterior angle; posterior lateral sinuation evidently carinate, slightly angled outward; pronotum broad and laterally deflected, overall shape cordate; midline well-defined, anterior medial and posterior medial diamond-shaped impressions present, disc with fine wrinkles present (may only be laterally) (
<figureCitation id="13297DB50278B77394BDB7ADE82CFA47" box="[1233,1319,1482,1507]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="125" pageNumber="126">Fig. 61</figureCitation>
); pronotal foveae broad and shallow; anterior edge inwardly curved, posterior edge straight. Prosternum flat and smooth; proepisternum without microsculpture. Procoxal setae absent; protibia distally expanded and shovel-like (
<figureCitation id="13297DB50278B7739503B475EDC2F9EB" captionStart="FIGURE 10" captionStartId="16.[151,250,1442,1464]" captionTargetBox="[152,1428,191,1421]" captionTargetId="figure@16.[151,1435,183,1427]" captionTargetPageId="16" captionText="FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used." httpUri="https://zenodo.org/record/2668083/files/figure.png" pageId="125" pageNumber="126">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="8BAD61300278B77390ABB43CE880F88E" blockId="125.[151,1437,834,2014]" pageId="125" pageNumber="126">
<emphasis id="B966BD220278B77390ABB43CEC1DF9D7" box="[199,278,1627,1651]" italics="true" pageId="125" pageNumber="126">Elytra:</emphasis>
Broad and deflected laterally, posteriorly steep to apex; humeral angle evenly convex (
<figureCitation id="13297DB50278B7739498B43DE846F9D7" box="[1268,1357,1626,1651]" captionStart="FIGURE 6" captionStartId="10.[151,250,519,541]" captionTargetBox="[156,1432,193,498]" captionTargetId="figure@10.[151,1436,193,498]" captionTargetPageId="10" captionText="FIGURE 6. Detail of the three different forms of the elytral humeral angle: (A) = anteriorly convergent (M. parataiko), (B) = evenly convex (M. oconnori), (C) = subangulate (M. atuanui)." httpUri="https://zenodo.org/record/2668075/files/figure.png" pageId="125" pageNumber="126">Fig. 6B</figureCitation>
), basal margin slightly curved and gently sloped to base, interval 1 extended to scutellum; lateral carina narrow the entire length, extended beyond humerus; humerus with 34 setose punctures (
<figureCitation id="13297DB50278B773938EB4C5E931F91F" box="[994,1082,1698,1723]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="125" pageNumber="126">Fig. 61</figureCitation>
); suture poorly defined; striae impressed with regularly spaced asetose punctures, striae 7 and 8 with slightly larger asetose punctures; intervals 16 weakly convex, intervals 79 strongly convex; interval microsculpture present as fine transverse lines; 7
<superScript id="7C67CC780278B773953DB48DE856F95C" attach="left" box="[1361,1373,1770,1784]" fontSize="6" pageId="125" pageNumber="126">th</superScript>
strial setal pattern with 34 setose punctures in anterior ½, 34 setose punctures in posterior ½, setose punctures small.
</paragraph>
<paragraph id="8BAD61300278B77090ABB552EC65FF71" blockId="125.[151,1437,834,2014]" lastBlockId="126.[151,1436,151,213]" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126">
<emphasis id="B966BD220278B77390ABB552EC88F8EA" box="[199,387,1845,1870]" italics="true" pageId="125" pageNumber="126">Ventral surface:</emphasis>
Mesepisternum (
<figureCitation id="13297DB50278B773923BB552EFA8F8EA" box="[599,675,1845,1870]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="125" pageNumber="126">Fig. 2</figureCitation>
) with numerous obsolescent punctures; metepisternum without microsculpture (may have wrinkles); setose punctures present on mesocoxae (1) and metacoxae (2). All abdominal ventrites finely lineate laterally, ventrites 1 and 2 with numerous obsolescent punctures; ventrites 35 with 1 setose puncture each side of midline; ventrite 6 setae present:
<specimenCount id="9D14AAB90278B7739363B5C5EE2FF81D" box="[783,804,1954,1977]" pageId="125" pageNumber="126" type="male"></specimenCount>
with 1 setose puncture each side at junction of straight apical edge and curve anterad,
<specimenCount id="9D14AAB90278B773919DB5A1EF0DF879" box="[497,518,1990,2013]" pageId="125" pageNumber="126" type="female"></specimenCount>
with 2 setose punctures each side, apical edge rounded; ventrites foveate laterally (shallow). Anterior metaventrite process (
<figureCitation id="13297DB5027BB7709205B2F0EFA7FF14" box="[617,684,151,176]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="126" pageNumber="127">Fig. 2</figureCitation>
) a rounded triangle with the carina well-defined, broad anterad and narrowed posterad.
</paragraph>
<caption id="DF6D31B8027BB77090FBB5EAE808F845" httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="126" pageNumber="127" startId="126.[151,250,1933,1955]" targetBox="[272,1317,269,1906]" targetPageId="126">
<paragraph id="8BAD6130027BB77090FBB5EAE808F845" blockId="126.[151,1436,1933,2017]" pageId="126" pageNumber="127">
<emphasis id="B966BD22027BB77090FBB5EAEC2BF807" bold="true" box="[151,288,1933,1955]" pageId="126" pageNumber="127">FIGURE 61.</emphasis>
<taxonomicName id="4C121AB3027BB770914AB5EAECDFF807" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[294,468,1933,1955]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="tuhoe">
<emphasis id="B966BD22027BB770914AB5EAECDFF807" box="[294,468,1933,1955]" italics="true" pageId="126" pageNumber="127">Mecodema tuhoe</emphasis>
</taxonomicName>
plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe.
</paragraph>
</caption>
<paragraph id="8BAD6130027AB77190ABB2FFEE62FDC4" blockId="127.[151,1437,151,1149]" pageId="127" pageNumber="128">
<emphasis id="B966BD22027AB77190ABB2FFEC73FF14" box="[199,376,152,176]" italics="true" pageId="127" pageNumber="128">Male genitalia:</emphasis>
Apical portion of penis lobe asymmetrically rounded with a distinct deflection to right of vertical axis (VV) (
<figureCitation id="13297DB5027AB7719113B2DBECE7FF71" box="[383,492,188,213]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="127" pageNumber="128">Fig. 15A</figureCitation>
); shaft curved down to form bluntly rounded point of the ventral process, apex symmetrically curved upward to form a continuous curving dorsal process (flattened discus-shaped), that terminates with sharp curve to shaft (
<figureCitation id="13297DB5027AB771925AB363EF8DFEB9" box="[566,646,260,285]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="127" pageNumber="128">Fig. 61</figureCitation>
PL); ventral edge of penis lobe moderately curved to right (VV); penis lobe shaft narrow the entire length; overall length of penis lobe slightly curved ventrally (LV) (
<figureCitation id="13297DB5027AB771954DB340E884FEE4" box="[1313,1423,295,320]" captionStart="FIGURE 16" captionStartId="21.[151,250,897,919]" captionTargetBox="[151,1436,181,869]" captionTargetId="figure@21.[151,1436,181,876]" captionTargetPageId="21" captionText="FIGURE 16. Relative strength of the ventral curve over the length of the penis lobe (lateral view): (A) slightly curved (M. oblongum); (B) moderately curved (M. ponaiti); (C) distinctly curved (M. manaia). Scale bar = 1 mm." httpUri="https://zenodo.org/record/2668095/files/figure.png" pageId="127" pageNumber="128">Fig. 16A</figureCitation>
). Structures of the endophallus (
<figureCitation id="13297DB5027AB771919AB32BEF36FEC0" box="[502,573,332,357]" captionStart="FIGURE 4" captionStartId="7.[151,250,973,995]" captionTargetBox="[354,1232,193,952]" captionTargetId="figure@7.[354,1232,193,952]" captionTargetPageId="7" captionText="FIGURE 4. An Environmental Scanning Electron Microscope (ESEM) photograph of the four main structures of the apical plate of the endophallus within the penis lobe. This is an example from Mecodema haunoho showing the central spicule (c.s), left (l.s.f) and the right setose flange (r.s.f), and microtrichial field (m.f). These structures are variable among species but within a species, they are very consistent, even over distant geographical ranges. The microtrichial field can be covered in flattened scales as in photo above, or with elongated setae-like scales." httpUri="https://zenodo.org/record/2668071/files/figure.png" pageId="127" pageNumber="128">Fig. 4</figureCitation>
): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule greatly expanded without projection; setal (scales) coverage of the apical plate extensive (76100%); size of left setose flange small; size of right setose flange absent. Left paramere basal lobe articulated at right angle to penis lobe, small and rectangular with a slight dorsal hump forming a moderate (45°) slope to arm; arm very short and broad (
<figureCitation id="13297DB5027AB77191A1B3BBEF2BFE51" box="[461,544,476,501]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="127" pageNumber="128">Fig. 61</figureCitation>
LP); terminal lobe short and broad, apical tuft of a few short setae and short, sparsely distributed setae at arm-basal lobe junction (
<figureCitation id="13297DB5027AB771937FB398EE62FDBC" box="[787,873,511,536]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="127" pageNumber="128">Fig. 61</figureCitation>
LP); ventral edge curved. Right paramere long, broadly rectangular (
<figureCitation id="13297DB5027AB77191E5B043ECD6FD99" box="[393,477,548,573]" captionStart="FIGURE 61" captionStartId="126.[151,250,1933,1955]" captionTargetBox="[272,1317,269,1906]" captionTargetId="figure@126.[257,1330,265,1912]" captionTargetPageId="126" captionText="FIGURE 61. Mecodema tuhoe plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668189/files/figure.png" pageId="127" pageNumber="128">Fig. 61</figureCitation>
RP), gradually narrowed from ½ length to terminus with a double row of long to medium-lengthed setae extended along apical ½ of ventral edge.
</paragraph>
<paragraph id="8BAD6130027AB77190ABB00BEF53FD69" blockId="127.[151,1437,151,1149]" pageId="127" pageNumber="128">
<emphasis id="B966BD22027AB77190ABB00BEC98FD20" box="[199,403,620,644]" italics="true" pageId="127" pageNumber="128">Female genitalia:</emphasis>
Basal gonocoxite 1 long and broad, ventral surface with many rugose grooves, internal dorso-lateral carina with 3 setae. Gonocoxite 2 paddle-like and ovate in shape with apical edge strongly curved upward. Ramus (
<figureCitation id="13297DB5027AB771913AB0D3EC93FD69" box="[342,408,692,717]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="127" pageNumber="128">Fig. 5</figureCitation>
) long and broad.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB027AB77190ABB0B0ECE0FC93" pageId="127" pageNumber="128" type="discussion">
<paragraph id="8BAD6130027AB77190ABB0B0ECE0FC93" blockId="127.[151,1437,151,1149]" pageId="127" pageNumber="128">
<emphasis id="B966BD22027AB77190ABB0B0EC59FD54" bold="true" box="[199,338,727,752]" pageId="127" pageNumber="128">Comments:</emphasis>
This species appears very much like
<taxonomicName id="4C121AB3027AB771929CB0BEEE61FD54" authorityName="Castelnau" authorityYear="1867" box="[752,874,728,752]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B966BD22027AB771929CB0BEEE61FD54" box="[752,874,728,752]" italics="true" pageId="127" pageNumber="128">M. simplex</emphasis>
</taxonomicName>
, and the dorsal external morphology confirms this conflation. However, the aedeagus of
<taxonomicName id="4C121AB3027AB7719223B09AEFBDFCB0" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[591,694,764,788]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="tuhoe">
<emphasis id="B966BD22027AB7719223B09AEFBDFCB0" box="[591,694,764,788]" italics="true" pageId="127" pageNumber="128">M. tuhoe</emphasis>
</taxonomicName>
, including the parameres are completely different in shape than those structures in
<taxonomicName id="4C121AB3027AB7719106B146ECEFFC9C" authorityName="Castelnau" authorityYear="1867" box="[362,484,800,824]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B966BD22027AB7719106B146ECEFFC9C" box="[362,484,800,824]" italics="true" pageId="127" pageNumber="128">M. simplex</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB027AB77190ABB123E975FCF9" box="[199,1150,836,861]" pageId="127" pageNumber="128" type="distribution">
<paragraph id="8BAD6130027AB77190ABB123E975FCF9" blockId="127.[151,1437,151,1149]" box="[199,1150,836,861]" pageId="127" pageNumber="128">
<emphasis id="B966BD22027AB77190ABB123EC69FCF9" bold="true" box="[199,354,836,861]" pageId="127" pageNumber="128">Distribution:</emphasis>
<collectingCountry id="F30521A0027AB7719105B122EF09FCF9" box="[361,514,836,861]" name="New Zealand" pageId="127" pageNumber="128">New Zealand</collectingCountry>
, North Island, Bay of Plenty, Ruatahuna, Kakanui River.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB027AB77190ABB100EFF2FBFC" pageId="127" pageNumber="128" type="materials_examined">
<paragraph id="8BAD6130027AB77190ABB100E861FC6C" blockId="127.[151,1437,151,1149]" pageId="127" pageNumber="128">
<materialsCitation id="3B7A6B6D027AB77190ABB100EDB8FC6C" ID-GBIF-Occurrence="2237851872" collectingDate="1992" collectingDateMax="1993-02-01" collectingDateMin="1992" collectionCode="LUNZ, GB, EAST, CAPE, INSECT, SURVEY" collectorName="J. S. Dugdale &amp; R. M. Emberson" country="New Zealand" elevation="200" location="Kakanui" pageId="127" pageNumber="128" specimenCode="NZMS260" specimenCount="1" typeStatus="holotype">
<emphasis id="B966BD22027AB77190ABB100EC36FC24" bold="true" box="[199,317,871,896]" pageId="127" pageNumber="128">
<typeStatus id="54A9DF92027AB77190ABB100EC33FC24" box="[199,312,871,896]" pageId="127" pageNumber="128" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED03F9F5027AB7719128B10FEC84FCDB" box="[324,399,872,895]" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="127" pageNumber="128">LUNZ</collectionCode>
male labelled.
<collectingCountry id="F30521A0027AB7719254B10FEE0EFC24" box="[568,773,872,896]" name="New Zealand" pageId="127" pageNumber="128">NEW ZEALAND</collectingCountry>
,
<collectionCode id="ED03F9F5027AB771937CB10FEE32FC24" box="[784,825,872,896]" country="Sweden" name="University of Gothenburg" pageId="127" pageNumber="128" type="Herbarium">GB</collectionCode>
,
<location id="8ECD37EB027AB7719328B100EEA8FC24" LSID="urn:lsid:plazi:treatment:03BBD0260278B77190FBB0B0E9C4FBD8:8ECD37EB027AB7719328B100EEA8FC24" box="[836,931,871,896]" country="New Zealand" name="Kakanui" pageId="127" pageNumber="128">Kakanui</location>
<quantity id="4CEACCD5027AB77193C6B10FEEE2FC24" box="[938,1001,872,896]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="127" pageNumber="128" unit="m" value="200.0">
<elevation id="003F8603027AB77193C6B10FEEE2FC24" box="[938,1001,872,896]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="127" pageNumber="128" unit="m" value="200.0">200m</elevation>
</quantity>
<specimenCode id="DBB4C94B027AB771939CB100E960FC24" box="[1008,1131,871,896]" collectionCode="NZMS" pageId="127" pageNumber="128">NZMS260</specimenCode>
Z14-886818,
<date id="FFAC47F0027AB771956CB10FE862FC24" box="[1280,1385,872,896]" pageId="127" pageNumber="128" value="1993-02-01">
<collectingDate id="EFE8BE18027AB771956CB10FE862FC24" box="[1280,1385,872,896]" pageId="127" pageNumber="128" value="1993-02-01">1.II.1993</collectingDate>
</date>
,
<collectorName id="26E704E6027AB7719518B10FEDF2FC01" pageId="127" pageNumber="128">J.S. Dugdale</collectorName>
,
<collectorName id="26E704E6027AB7719169B1EAECB3FC01" box="[261,440,908,933]" pageId="127" pageNumber="128">R.M. Emberson</collectorName>
/ under log in puriri (
<taxonomicName id="4C121AB3027AB77192C3B1EAEE3EFC00" box="[687,821,908,932]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="127" pageNumber="128" phylum="Tracheophyta" rank="species" species="lucens">
<emphasis id="B966BD22027AB77192C3B1EAEE3EFC00" box="[687,821,908,932]" italics="true" pageId="127" pageNumber="128">Vitex lucens</emphasis>
</taxonomicName>
)-nikau forest /
<collectionCode id="ED03F9F5027AB771938AB1EBE855FC00" box="[998,1374,908,932]" pageId="127" pageNumber="128">EAST CAPE INSECT SURVEY</collectionCode>
<date id="FFAC47F0027AB7719532B1EBEDB8FC6C" pageId="127" pageNumber="128" value="1992" valueMax="1993">
<collectingDate id="EFE8BE18027AB7719532B1EBEDB8FC6C" pageId="127" pageNumber="128" value="1992" valueMax="1993">1992- 93</collectingDate>
</date>
</materialsCitation>
/
<materialsCitation id="3B7A6B6D027AB77190A5B1D7E86DFC6C" ID-GBIF-Occurrence="2237851851" box="[201,1382,943,968]" collectionCode="LUNZ" collectorName="DS Seldon &amp; TR Buckley" country="New Zealand" pageId="127" pageNumber="128" specimenCode="LUNZ00002675" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A9DF92027AB77190A5B1D7EC56FC6C" box="[201,349,944,968]" pageId="127" pageNumber="128" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C121AB3027AB771910FB1C8EF2EFC6C" authority="Seldon &amp; Buckley, 2019" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[355,549,943,968]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="tuhoe" status="sp. nov.">Mecodema tuhoe</taxonomicName>
<emphasis id="B966BD22027AB7719240B1C8EF60FC6C" bold="true" box="[556,619,943,968]" pageId="127" pageNumber="128">
<taxonomicNameLabel id="A2550059027AB7719240B1C8EF60FC6C" box="[556,619,943,968]" pageId="127" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
design.
<collectorName id="26E704E6027AB77192A5B1D7EE4EFC6C" box="[713,837,943,968]" pageId="127" pageNumber="128">DS Seldon</collectorName>
,
<collectorName id="26E704E6027AB771933CB1D7EEDCFC6C" box="[848,983,943,968]" pageId="127" pageNumber="128">TR Buckley</collectorName>
2018 [red label] /
<specimenCode id="DBB4C94B027AB77194C4B1C8E86DFC6C" box="[1192,1382,943,968]" collectionCode="LUNZ" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="127" pageNumber="128">LUNZ00002675</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD6130027AB77190ABB1B3EFF2FBFC" blockId="127.[151,1437,151,1149]" pageId="127" pageNumber="128">
<materialsCitation id="3B7A6B6D027AB77190ABB1B3EF81FC48" ID-GBIF-Occurrence="2237851859" box="[199,650,980,1005]" collectionCode="GB" country="New Zealand" location="NEW ZEALAND" pageId="127" pageNumber="128" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B966BD22027AB77190ABB1B3EC43FC49" bold="true" box="[199,328,980,1005]" pageId="127" pageNumber="128">
<typeStatus id="54A9DF92027AB77190ABB1B3EC48FC49" box="[199,323,980,1005]" pageId="127" pageNumber="128" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D14AAB9027AB771913DB1B3EC7FFC48" box="[337,372,980,1004]" pageId="127" pageNumber="128" type="female">1♀</specimenCount>
,
<collectingCountry id="F30521A0027AB77191E9B1B2EF5EFC48" box="[389,597,980,1004]" name="New Zealand" pageId="127" pageNumber="128">NEW ZEALAND</collectingCountry>
,
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(
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/ J.S.
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(
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.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB027AB77190ABB603E9C4FBD8" box="[199,1231,1124,1149]" pageId="127" pageNumber="128" type="etymology">
<paragraph id="8BAD6130027AB77190ABB603E9C4FBD8" blockId="127.[151,1437,151,1149]" box="[199,1231,1124,1149]" pageId="127" pageNumber="128">
<emphasis id="B966BD22027AB77190ABB603EC5AFBD9" bold="true" box="[199,337,1124,1149]" pageId="127" pageNumber="128">Etymology:</emphasis>
<taxonomicName id="4C121AB3027AB7719134B603EF13FBD8" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[344,536,1124,1148]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="tuhoe">
<emphasis id="B966BD22027AB7719134B603EF13FBD8" box="[344,536,1124,1148]" italics="true" pageId="127" pageNumber="128">Mecodema tuhoe</emphasis>
</taxonomicName>
is named after Ngati Tūhoe whose rohe includes Te Urewera.
</paragraph>
</subSubSection>
</treatment>
</document>