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<document id="10D9BFD628266EAC44E26EBDBA3B344F" ID-CLB-Dataset="26919" ID-DOI="10.11646/zootaxa.4598.1.1" ID-GBIF-Dataset="475fd53e-c5c6-4a99-bb2e-cc2382f0171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668063" ID-ZooBank="6BA5F086-1014-46B5-B6D5-77FE0CD07AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127411994" checkinUser="plazi" docAuthor="Seldon, David S. &amp; Buckley, Thomas R." docDate="2019" docId="03BBD0260276B77B90FBB759EC26F833" docLanguage="en" docName="zootaxa.4598.1.1.pdf.imf" docOrigin="Zootaxa 4598 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mecodema teparawhau Seldon &amp; Buckley 2019, sp. n." docType="treatment" docUuid="F914C688-C16F-4B23-93A5-093B3468B722" docUuidSource="ZooBank" docVersion="11" lastPageNumber="118" masterDocId="FF82A85E0205B70E906CB267ED0BFFA4" masterDocTitle="The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand" masterLastPageNumber="148" masterPageNumber="1" pageNumber="116" updateTime="1698726038790" updateUser="ExternalLinkService">
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<mods:title id="410DB37EB08D34774C3178D03DE88C54">The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand</mods:title>
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<mods:namePart id="78ADE82C97FCD35974C18E048039F7EE">Seldon, David S.</mods:namePart>
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<mods:namePart id="81544F5CE9CD835A1930DF3C7C4237EA">Buckley, Thomas R.</mods:namePart>
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<mods:title id="702B5FAE52BED44EE34F96043E109C32">Zootaxa</mods:title>
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<mods:date id="50E65EEA8691DD184D76EB0E0D5E8B36">2019</mods:date>
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<mods:number id="0214416CD7B44A9F9EB96CF625DE08B3">2019-05-06</mods:number>
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<mods:classification id="709CCA670BD15A80641CA397D8F0C794">journal article</mods:classification>
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<mods:identifier id="CBFEE0555DAE44D5B05228FD4C53DFA7" type="DOI">10.11646/zootaxa.4598.1.1</mods:identifier>
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<treatment id="03BBD0260276B77B90FBB759EC26F833" ID-DOI="http://doi.org/10.5281/zenodo.5678442" ID-GBIF-Taxon="157062702" ID-Zenodo-Dep="5678442" LSID="urn:lsid:zoobank.org:act:F914C688-C16F-4B23-93A5-093B3468B722" httpUri="http://treatment.plazi.org/id/03BBD0260276B77B90FBB759EC26F833" lastPageId="117" lastPageNumber="118" pageId="115" pageNumber="116">
<subSubSection id="C30832BB0276B77D90FBB759ECF0FAFC" box="[151,507,1342,1368]" pageId="115" pageNumber="116" type="nomenclature">
<paragraph id="8BAD61300276B77D90FBB759ECF0FAFC" blockId="115.[151,507,1342,1402]" box="[151,507,1342,1368]" pageId="115" pageNumber="116">
<heading id="D0E5D65C0276B77D90FBB759ECF0FAFC" bold="true" box="[151,507,1342,1368]" fontSize="11" level="1" pageId="115" pageNumber="116" reason="1">
<emphasis id="B966BD220276B77D90FBB759ECF0FAFC" bold="true" box="[151,507,1342,1368]" pageId="115" pageNumber="116">
<taxonomicName id="4C121AB30276B77D90FBB759ECBAFAFC" ID-CoL="8LXZB" authority="Seldon &amp; Buckley, 2019" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[151,433,1342,1368]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="teparawhau" status="sp. nov.">
<emphasis id="B966BD220276B77D90FBB759ECBAFAFC" bold="true" box="[151,433,1342,1368]" italics="true" pageId="115" pageNumber="116">Mecodema teparawhau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25500590276B77D91D5B759ECF0FAFC" box="[441,507,1342,1368]" pageId="115" pageNumber="116" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30832BB0276B77D90FBB706EC01FADE" box="[151,266,1377,1402]" pageId="115" pageNumber="116" type="description">
<paragraph id="8BAD61300276B77D90FBB706EC01FADE" blockId="115.[151,507,1342,1402]" box="[151,266,1377,1402]" pageId="115" pageNumber="116">
<figureCitation id="13297DB50276B77D90FBB706EC0DFADE" box="[151,262,1377,1402]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="115" pageNumber="116">Figure 57</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0276B77D90FBB7CEECB7F9AE" pageId="115" pageNumber="116" type="diagnosis">
<paragraph id="8BAD61300276B77D90FBB7CEECB7F9AE" blockId="115.[151,1436,1449,2014]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90FBB7CEEC18FA66" bold="true" box="[151,275,1449,1474]" pageId="115" pageNumber="116">Diagnosis:</emphasis>
Differing from other North Island
<taxonomicName id="4C121AB30276B77D92CFB7CEEE15FA65" authorityName="Blanchard" authorityYear="1853" box="[675,798,1449,1473]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B966BD220276B77D92CFB7CEEE15FA65" box="[675,798,1449,1473]" italics="true" pageId="115" pageNumber="116">Mecodema</emphasis>
</taxonomicName>
species by having:
<emphasis id="B966BD220276B77D946CB7CEE905FA66" bold="true" box="[1024,1038,1449,1474]" pageId="115" pageNumber="116">1</emphasis>
, an indistinct vertexal groove with obsolescent punctures laterally;
<emphasis id="B966BD220276B77D9268B7AAEF19FA42" bold="true" box="[516,530,1485,1510]" pageId="115" pageNumber="116">2</emphasis>
, 810 setae along prothoracic carina;
<emphasis id="B966BD220276B77D93AAB7AAEEDFFA42" bold="true" box="[966,980,1485,1510]" pageId="115" pageNumber="116">3</emphasis>
, setal distribution along ventral edge of left paramere (
<figureCitation id="13297DB50276B77D9157B796EC87F9AE" box="[315,396,1521,1546]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="115" pageNumber="116">Fig. 57</figureCitation>
LP).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0276B77B90ABB472EDD5FADF" lastPageId="117" lastPageNumber="118" pageId="115" pageNumber="116" type="description">
<paragraph id="8BAD61300276B77D90ABB472E9E3F9F6" blockId="115.[151,1436,1449,2014]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB472EC50F98A" bold="true" box="[199,347,1557,1582]" pageId="115" pageNumber="116">Description:</emphasis>
Length
<quantity id="4CEACCD50276B77D91A8B472EF4EF989" box="[452,581,1557,1582]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="115" pageNumber="116" unit="mm" value="29.0" valueMax="32.0" valueMin="26.0">2632 mm</quantity>
, pronotal width
<quantity id="4CEACCD50276B77D9362B472EE96F989" box="[782,925,1557,1582]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.5" metricValueMin="7.5" pageId="115" pageNumber="116" unit="mm" value="8.5" valueMax="9.5" valueMin="7.5">7.59.5 mm</quantity>
, elytral
<quantity id="4CEACCD50276B77D9468B472E9A9F989" box="[1028,1186,1557,1582]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="10.6" metricValueMin="8.6" pageId="115" pageNumber="116" unit="mm" value="9.6" valueMax="10.6" valueMin="8.6">8.610.6 mm</quantity>
. Colour of head and pronotum glossy black, elytra and ventrally matte black, coxae dark reddish-brown and legs black.
</paragraph>
<paragraph id="8BAD61300276B77D90ABB43AEC46F87A" blockId="115.[151,1436,1449,2014]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB43AEC06F9D1" box="[199,269,1629,1653]" italics="true" pageId="115" pageNumber="116">Head:</emphasis>
Broad and flat (
<figureCitation id="13297DB50276B77D91ADB43AEF22F9D2" box="[449,553,1629,1654]" captionStart="FIGURE 14" captionStartId="20.[151,250,936,958]" captionTargetBox="[171,1412,204,911]" captionTargetId="figure@20.[160,1412,193,921]" captionTargetPageId="20" captionText="FIGURE 14. Diagrammatic cross sections that represent the width and convexity of head or deflection of pronotum (frontal view): (A) broad and flat; (B) broad and convex; (C) narrow and flat; (D) narrow and convex." httpUri="https://zenodo.org/record/2668091/files/figure.png" pageId="115" pageNumber="116">Fig. 14A</figureCitation>
). Vertex smooth; vertexal groove shallow but defined by obsolescent punctures laterally; large supraorbital puncture bearing 5 setae (
<figureCitation id="13297DB50276B77D9294B4E6EE45F93E" box="[760,846,1665,1690]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="115" pageNumber="116">Fig. 57</figureCitation>
); 1 indistinct supraorbital groove mesad eyes; frons smooth, except shallow grooves in large depression each side proximate pits; frontoclypeal suture very narrow medially to indistinct in grooves around very small tentorial pits; anterior area of clypeus with well-defined grooves; clypeus with 1 setose puncture each side bearing 12 setae (
<figureCitation id="13297DB50276B77D93C3B48AE908F8A2" box="[943,1027,1773,1798]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="115" pageNumber="116">Fig. 57</figureCitation>
), some specimens may have medial setose puncture bearing 1 seta. Labrum rectangular, anterior edge rounded laterally, slightly emarginated medially with with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes rounded (
<figureCitation id="13297DB50276B77D94A2B552E83FF8EA" box="[1230,1332,1845,1870]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="115" pageNumber="116">Fig. 12F</figureCitation>
), median process broad and short, slightly angled upward (15°), moderately indentate (
<figureCitation id="13297DB50276B77D9462B53EE97CF8D6" box="[1038,1143,1881,1906]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="115" pageNumber="116">Fig. 12B</figureCitation>
); setae variably absent or present. Submentum sclerite constriction narrow with 6 evenly spaced setae. Stipes with 2 basal setae. Gula pits small, suture defined, gula (
<figureCitation id="13297DB50276B77D91B7B5C6EF2AF81E" box="[475,545,1953,1978]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="115" pageNumber="116">Fig. 8</figureCitation>
) flat and smooth. Gena with fine wrinkles medially forming an isodiametric pattern laterally.
</paragraph>
<caption id="DF6D31B80271B77A90FBB520E808F83E" httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="116" pageNumber="117" startId="116.[151,250,1863,1885]" targetBox="[278,1307,203,1832]" targetPageId="116">
<paragraph id="8BAD61300271B77A90FBB520E808F83E" blockId="116.[151,1436,1863,1946]" pageId="116" pageNumber="117">
<emphasis id="B966BD220271B77A90FBB520EC14F8F9" bold="true" box="[151,287,1863,1885]" pageId="116" pageNumber="117">FIGURE 57.</emphasis>
<taxonomicName id="4C121AB30271B77A9149B520EF18F8F9" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[293,531,1863,1885]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="teparawhau">
<emphasis id="B966BD220271B77A9149B520EF18F8F9" box="[293,531,1863,1885]" italics="true" pageId="116" pageNumber="117">Mecodema teparawhau</emphasis>
</taxonomicName>
plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe.
</paragraph>
</caption>
<paragraph id="8BAD61300270B77B90ABB2FFEFE4FE2C" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<emphasis id="B966BD220270B77B90ABB2FFEC49FF14" box="[199,322,152,176]" italics="true" pageId="117" pageNumber="118">Prothorax:</emphasis>
Prothoracic carina narrow the entire length (slightly broadened at shoulder), slightly crenulated with 811 setae each side (
<figureCitation id="13297DB50270B77B91B9B2DBEF27FF71" box="[469,556,188,213]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="117" pageNumber="118">Fig. 57</figureCitation>
), extended to anterior angle; posterior lateral sinuation indistinctly carinate, parallel; pronotum broad and slightly deflected laterally, overall shape broadly cordate; midline poorly impressed, anterior medial impression triangular, posterior medial impression diamond-shaped present, disc with transverse wrinkles; pronotal foveae (
<figureCitation id="13297DB50270B77B91A3B340EF29FEE4" box="[463,546,295,320]" captionStart="FIGURE 11" captionStartId="17.[151,250,949,971]" captionTargetBox="[221,1358,181,923]" captionTargetId="figure@17.[221,1359,179,929]" captionTargetPageId="17" captionText="FIGURE 11. Detail of pronotum showing the taxonomic structures used in the species descriptions. PPC = posterior pronotum constriction; PS = pronotal setae; PAA = pronotum anterior angle; AE = anterior edge; PC = pronotal carina; ML = midline; PF= pronotal foveae; PPS = pronotum posterior sinuation; PE = posterior edge." httpUri="https://zenodo.org/record/2668085/files/figure.png" pageId="117" pageNumber="118">Fig. 11</figureCitation>
) shallow and narrow; anterior edge inwardly curved, posterior edge slightly emarginated medially. Prosternum concave and smooth; proepisternum with microsculpture. Procoxal setae absent; protibia distally expanded and shovel-like (
<figureCitation id="13297DB50270B77B9210B308EFE9FE2C" box="[636,738,367,392]" captionStart="FIGURE 10" captionStartId="16.[151,250,1442,1464]" captionTargetBox="[152,1428,191,1421]" captionTargetId="figure@16.[151,1435,183,1427]" captionTargetPageId="16" captionText="FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used." httpUri="https://zenodo.org/record/2668083/files/figure.png" pageId="117" pageNumber="118">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="8BAD61300270B77B90ABB3F3EF01FDC7" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<emphasis id="B966BD220270B77B90ABB3F3EC1DFE08" box="[199,278,404,428]" italics="true" pageId="117" pageNumber="118">Elytra:</emphasis>
Broad and deflected laterally; humeral angle evenly convex (
<figureCitation id="13297DB50270B77B93A4B3F3E914FE09" box="[968,1055,404,429]" captionStart="FIGURE 6" captionStartId="10.[151,250,519,541]" captionTargetBox="[156,1432,193,498]" captionTargetId="figure@10.[151,1436,193,498]" captionTargetPageId="10" captionText="FIGURE 6. Detail of the three different forms of the elytral humeral angle: (A) = anteriorly convergent (M. parataiko), (B) = evenly convex (M. oconnori), (C) = subangulate (M. atuanui)." httpUri="https://zenodo.org/record/2668075/files/figure.png" pageId="117" pageNumber="118">Fig. 6B</figureCitation>
); basal margin moderately curved and gently sloped to base, interval 1 extended to base; lateral carina narrow the entire length, extended to humeral angle; humerus with 3 setose punctures; suture well-defined; striae with small, regularly spaced, asetose punctures, slightly increased in size laterally; intervals 18 slightly convex, interval 9 moderately convex; interval microsculpture absent; 7
<superScript id="7C67CC780270B77B91C5B044ECBEFD95" attach="left" box="[425,437,547,561]" fontSize="6" pageId="117" pageNumber="118">th</superScript>
strial setal pattern with 3 setose punctures in anterior ½, 5 setose punctures in posterior ½ (
<figureCitation id="13297DB50270B77B90F3B02DEDF8FDC7" box="[159,243,586,611]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="117" pageNumber="118">Fig. 57</figureCitation>
), setose punctures small.
</paragraph>
<paragraph id="8BAD61300270B77B90ABB008E931FC9F" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<emphasis id="B966BD220270B77B90ABB008EC8AFD23" box="[199,385,623,647]" italics="true" pageId="117" pageNumber="118">Ventral surface:</emphasis>
Mesepisternum and metepisternum without microsculpture; setose punctures present on mesocoxa (2) and metacoxa (2). All abdominal ventrites (
<figureCitation id="13297DB50270B77B935AB0F5EE76FD0F" box="[822,893,658,683]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="117" pageNumber="118">Fig. 2</figureCitation>
) finely lineate laterally; ventrites 1 and 2 with scattered obsolescent punctures, ventrites 35 with 1 setose puncture each side of midline; ventrite 6 setae present:
<specimenCount id="9D14AAB90270B77B90FBB0BCEDA7FD56" box="[151,172,731,754]" pageId="117" pageNumber="118" type="male"></specimenCount>
with 12 setose punctures each side distantly situated at junction of straight apical edge and curve anterad;
<specimenCount id="9D14AAB90270B77B95EBB0BCE897FD56" box="[1415,1436,731,754]" pageId="117" pageNumber="118" type="female"></specimenCount>
with 23 setose punctures each side, medially proximate, along rounded apical edge; lateral foveae absent. Anterior metaventrite process (
<figureCitation id="13297DB50270B77B91E1B145ECDBFC9F" box="[397,464,802,827]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="117" pageNumber="118">Fig. 2</figureCitation>
) a short triangle with a narrow carina the entire length.
</paragraph>
<paragraph id="8BAD61300270B77B90ABB120EE6BFAAB" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<emphasis id="B966BD220270B77B90ABB120EC72FCFB" box="[199,377,839,863]" italics="true" pageId="117" pageNumber="118">Male genitalia:</emphasis>
Apical portion of penis lobe asymmetrically hooked with a distinct deflection to right of vertical axis (VV) (
<figureCitation id="13297DB50270B77B9102B10DECDDFC27" box="[366,470,874,899]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="117" pageNumber="118">Fig. 15A</figureCitation>
); ventral process a very small, bluntly rounded point (
<figureCitation id="13297DB50270B77B9441B10DE975FC27" box="[1069,1150,874,899]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="117" pageNumber="118">Fig. 57</figureCitation>
PL), apex asymmetrically curved upward to form a long and narrow dorsal process with a slightly recurved hook that is perpendicular to shaft, short, shape return curve to broadened apical part of shaft (
<figureCitation id="13297DB50270B77B93E7B1D5EED4FC6F" box="[907,991,946,971]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="117" pageNumber="118">Fig. 57</figureCitation>
PL); penis lobe shaft narrow the entire length, very gradually broadened to base; ventral edge of penis lobe with slight curve to right (VV); overall length of penis lobe moderate curved ventrally (LV) (
<figureCitation id="13297DB50270B77B92CEB19DEE02FBB7" box="[674,777,1018,1043]" captionStart="FIGURE 16" captionStartId="21.[151,250,897,919]" captionTargetBox="[151,1436,181,869]" captionTargetId="figure@21.[151,1436,181,876]" captionTargetPageId="21" captionText="FIGURE 16. Relative strength of the ventral curve over the length of the penis lobe (lateral view): (A) slightly curved (M. oblongum); (B) moderately curved (M. ponaiti); (C) distinctly curved (M. manaia). Scale bar = 1 mm." httpUri="https://zenodo.org/record/2668095/files/figure.png" pageId="117" pageNumber="118">Fig. 16B</figureCitation>
). Structures of the endophallus (
<figureCitation id="13297DB50270B77B9419B19DE9B0FBB7" box="[1141,1211,1018,1043]" captionStart="FIGURE 4" captionStartId="7.[151,250,973,995]" captionTargetBox="[354,1232,193,952]" captionTargetId="figure@7.[354,1232,193,952]" captionTargetPageId="7" captionText="FIGURE 4. An Environmental Scanning Electron Microscope (ESEM) photograph of the four main structures of the apical plate of the endophallus within the penis lobe. This is an example from Mecodema haunoho showing the central spicule (c.s), left (l.s.f) and the right setose flange (r.s.f), and microtrichial field (m.f). These structures are variable among species but within a species, they are very consistent, even over distant geographical ranges. The microtrichial field can be covered in flattened scales as in photo above, or with elongated setae-like scales." httpUri="https://zenodo.org/record/2668071/files/figure.png" pageId="117" pageNumber="118">Fig. 4</figureCitation>
): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule greatly expanded with projection; setal (scales) coverage of the apical plate moderate (2675%); size of left setose flange small; size of right setose flange very large. Left paramere articulated at right angle to penis lobe, rectangular with slight dorsal hump forming a steep (60°) slope to arm (
<figureCitation id="13297DB50270B77B924AB6EDEF73FB07" box="[550,632,1162,1187]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="117" pageNumber="118">Fig. 57</figureCitation>
LP); arm short and narrow; terminal lobe short and broad with an apical tuft of a few short setae; medium-lengthed setae in a comb at arm-basal lobe junction with shorter setae extended a little way along curved ventral edge (
<figureCitation id="13297DB50270B77B922FB6B5EF9DFB4F" box="[579,662,1234,1259]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="117" pageNumber="118">Fig. 57</figureCitation>
LP). Right paramere long, narrowly triangular with a double row of long setae extended along apical ¾ of ventral edge (
<figureCitation id="13297DB50270B77B92B1B691EE25FAAB" box="[733,814,1270,1295]" captionStart="FIGURE 57" captionStartId="116.[151,250,1863,1885]" captionTargetBox="[278,1307,203,1832]" captionTargetId="figure@116.[260,1328,193,1842]" captionTargetPageId="116" captionText="FIGURE 57. Mecodema teparawhau plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668181/files/figure.png" pageId="117" pageNumber="118">Fig. 57</figureCitation>
RP).
</paragraph>
<paragraph id="8BAD61300270B77B90ABB77CEDD5FADF" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<emphasis id="B966BD220270B77B90ABB77CEC99FA97" box="[199,402,1307,1331]" italics="true" pageId="117" pageNumber="118">Female genitalia:</emphasis>
Basal gonocoxite 1 short and broad, ventral surface smooth, internal dorso-lateral carina with 2 setae. Gonocoxite 2 a short rounded triangle with a slight upward curve apically. Ramus (
<figureCitation id="13297DB50270B77B9488B759E823FAF3" box="[1252,1320,1342,1367]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="117" pageNumber="118">Fig. 5</figureCitation>
) long and broad.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0270B77B90ABB7E1EC0FFA43" pageId="117" pageNumber="118" type="discussion">
<paragraph id="8BAD61300270B77B90ABB7E1EC0FFA43" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<emphasis id="B966BD220270B77B90ABB7E1EC59FA3B" bold="true" box="[199,338,1414,1439]" pageId="117" pageNumber="118">Comments:</emphasis>
The Whangarei area has a number of
<taxonomicName id="4C121AB30270B77B9364B7E0EE88FA3B" authorityName="Blanchard" authorityYear="1853" box="[776,899,1415,1439]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B966BD220270B77B9364B7E0EE88FA3B" box="[776,899,1415,1439]" italics="true" pageId="117" pageNumber="118">Mecodema</emphasis>
</taxonomicName>
species that inhabit the forested hills, and with all species it is difficult to determine their natural distributions due to the isolation induced by anthropogenic activities.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0270B77B90ABB795E825F9AF" box="[199,1326,1522,1547]" pageId="117" pageNumber="118" type="distribution">
<paragraph id="8BAD61300270B77B90ABB795E825F9AF" blockId="117.[151,1437,151,1943]" box="[199,1326,1522,1547]" pageId="117" pageNumber="118">
<emphasis id="B966BD220270B77B90ABB795EC69F9AF" bold="true" box="[199,354,1522,1547]" pageId="117" pageNumber="118">Distribution:</emphasis>
<collectingCountry id="F30521A00270B77B9105B794EF09F9AF" box="[361,514,1522,1547]" name="New Zealand" pageId="117" pageNumber="118">New Zealand</collectingCountry>
, North Island, Northland, Whangarei area, Pukenui Forest, Mt Parahaki.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0270B77B90ABB471E898F8EB" pageId="117" pageNumber="118" type="materials_examined">
<paragraph id="8BAD61300270B77B90ABB471EF22F9D3" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<materialsCitation id="3B7A6B6D0270B77B90ABB471ECC8F9F7" ID-GBIF-Occurrence="2237851807" collectingDate="2000-05-09" collectionCode="AMNZ" collectorName="E. Conaghan" country="New Zealand" location="Pukenui Forest" pageId="117" pageNumber="118" specimenCode="NZMS 260" specimenCount="1" typeStatus="holotype">
<emphasis id="B966BD220270B77B90ABB471EC36F98B" bold="true" box="[199,317,1558,1583]" pageId="117" pageNumber="118">
<typeStatus id="54A9DF920270B77B90ABB471EC33F98B" box="[199,312,1558,1583]" pageId="117" pageNumber="118" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED03F9F50270B77B9128B470EC9DF98B" box="[324,406,1559,1583]" country="New Zealand" httpUri="http://grbio.org/cool/76et-m1kt" name="Auckland Institute and Museum" pageId="117" pageNumber="118" type="Museum">AMNZ</collectionCode>
male labelled.
<collectingCountry id="F30521A00270B77B9228B47FEE18F98B" box="[580,787,1559,1583]" name="New Zealand" pageId="117" pageNumber="118">NEW ZEALAND</collectingCountry>
, ND,
<location id="8ECD37EB0270B77B933AB471E909F98B" LSID="urn:lsid:plazi:treatment:03BBD0260276B77B90FBB759EC26F833:8ECD37EB0270B77B933AB471E909F98B" box="[854,1026,1558,1583]" country="New Zealand" name="Pukenui Forest" pageId="117" pageNumber="118">Pukenui Forest</location>
,
<date id="FFAC47F00270B77B9462B470E991F98B" box="[1038,1178,1559,1584]" pageId="117" pageNumber="118" value="2000-05-09">
<collectingDate id="EFE8BE180270B77B9462B470E991F98B" box="[1038,1178,1559,1584]" pageId="117" pageNumber="118" value="2000-05-09">9 May 2000</collectingDate>
</date>
,
<collectorName id="26E704E60270B77B94C9B47FE83DF98B" box="[1189,1334,1558,1583]" pageId="117" pageNumber="118">E. Conaghan</collectorName>
/
<specimenCode id="DBB4C94B0270B77B9522B470EDCAF9F7" collectionCode="NZMS" pageId="117" pageNumber="118">NZMS 260</specimenCode>
Q07 245 0 96, on track
</materialsCitation>
/
<materialsCitation id="3B7A6B6D0270B77B91B4B45CEF2EF9D3" ID-GBIF-Occurrence="2237851829" collectionCode="AMNZ" collectorName="DS Seldon &amp; TR Buckley" country="New Zealand" pageId="117" pageNumber="118" specimenCode="AMNZ 40702" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A9DF920270B77B91B4B45CEF67F9F7" box="[472,620,1595,1619]" pageId="117" pageNumber="118" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C121AB30270B77B921FB45DEE7CF9F7" authority="Seldon &amp; Buckley, 2019" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[627,887,1594,1619]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="teparawhau" status="sp. nov.">Mecodema teparawhau</taxonomicName>
<emphasis id="B966BD220270B77B9312B45DEEB6F9F7" bold="true" box="[894,957,1594,1619]" pageId="117" pageNumber="118">
<taxonomicNameLabel id="A25500590270B77B9312B45DEEB6F9F7" box="[894,957,1594,1619]" pageId="117" pageNumber="118" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
design.
<collectorName id="26E704E60270B77B9477B45CE99DF9F7" box="[1051,1174,1594,1619]" pageId="117" pageNumber="118">DS Seldon</collectorName>
,
<collectorName id="26E704E60270B77B94CDB45CE822F9F7" box="[1185,1321,1594,1619]" pageId="117" pageNumber="118">TR Buckley</collectorName>
2018 [red label] /
<specimenCode id="DBB4C94B0270B77B9080B438EC87F9D3" box="[236,396,1631,1655]" collectionCode="AMNZ" country="New Zealand" httpUri="http://grbio.org/cool/76et-m1kt" name="Auckland Institute and Museum" pageId="117" pageNumber="118" type="Museum">AMNZ 40702</specimenCode>
[green label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD61300270B77B90ABB4E5E898F8EB" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<materialsCitation id="3B7A6B6D0270B77B90ABB4E5E95DF93F" ID-GBIF-Occurrence="2237851830" box="[199,1110,1666,1691]" collectingDate="1999-03-03" collectionCode="AM" country="New Zealand" location="Hirupaki Pa" pageId="117" pageNumber="118" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B966BD220270B77B90ABB4E5EC43F93F" bold="true" box="[199,328,1666,1691]" pageId="117" pageNumber="118">
<typeStatus id="54A9DF920270B77B90ABB4E5EC48F93F" box="[199,323,1666,1691]" pageId="117" pageNumber="118" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D14AAB90270B77B9122B4E4EC79F93F" box="[334,370,1667,1691]" pageId="117" pageNumber="118" type="male">1♂</specimenCount>
,
<specimenCount id="9D14AAB90270B77B9113B4E4ECA9F93F" box="[383,418,1667,1691]" pageId="117" pageNumber="118" type="female">1♀</specimenCount>
,
<collectingCountry id="F30521A00270B77B91C3B4E4EF76F93F" box="[431,637,1667,1691]" name="New Zealand" pageId="117" pageNumber="118">NEW ZEALAND</collectingCountry>
, ND,
<location id="8ECD37EB0270B77B92D2B4E5EE4FF93E" LSID="urn:lsid:plazi:treatment:03BBD0260276B77B90FBB759EC26F833:8ECD37EB0270B77B92D2B4E5EE4FF93E" box="[702,836,1666,1691]" country="New Zealand" name="Hirupaki Pa" pageId="117" pageNumber="118">Hirupaki Pa</location>
site,
<date id="FFAC47F00270B77B9313B4E4E915F93F" box="[895,1054,1666,1691]" pageId="117" pageNumber="118" value="1999-03-03">
<collectingDate id="EFE8BE180270B77B9313B4E4E915F93F" box="[895,1054,1666,1691]" pageId="117" pageNumber="118" value="1999-03-03">3 March 1999</collectingDate>
</date>
,
<collectionCode id="ED03F9F50270B77B9445B4E4E95DF93F" box="[1065,1110,1667,1691]" country="Australia" name="Australian Museum" pageId="117" pageNumber="118" type="Museum">AM</collectionCode>
</materialsCitation>
<materialsCitation id="3B7A6B6D0270B77B9430B4E5E83EF93F" ID-GBIF-Occurrence="2237851876" box="[1116,1333,1666,1691]" collectionCode="NZMS" collectorName="Booth" country="New Zealand" pageId="117" pageNumber="118" specimenCode="NZMS 260" specimenCount="1" typeStatus="paratype">
<collectorName id="26E704E60270B77B9430B4E5E9ABF93F" box="[1116,1184,1666,1691]" pageId="117" pageNumber="118">Booth</collectorName>
/
<specimenCode id="DBB4C94B0270B77B94DAB4E4E83EF93F" box="[1206,1333,1666,1691]" collectionCode="NZMS" pageId="117" pageNumber="118">NZMS 260</specimenCode>
</materialsCitation>
<materialsCitation id="3B7A6B6D0270B77B9557B4E5EF76F91B" ID-GBIF-Occurrence="2237851934" collectionCode="AMNZ" country="New Zealand" pageId="117" pageNumber="118" specimenCode="AMNZ 40700, 40701" specimenCount="1" typeStatus="paratype">
Q06 269 124,
<collectingMethod id="525319270270B77B90BDB4C1EC62F91B" box="[209,361,1702,1727]" pageId="117" pageNumber="118">in pitfall trap</collectingMethod>
/
<specimenCode id="DBB4C94B0270B77B91E8B4C0EF22F91B" box="[388,553,1703,1727]" collectionCode="AMNZ" country="New Zealand" httpUri="http://grbio.org/cool/76et-m1kt" name="Auckland Institute and Museum" pageId="117" pageNumber="118" type="Museum">AMNZ 40700</specimenCode>
,
<specimenCode id="DBB4C94B0270B77B925BB4C0EF76F91B" box="[567,637,1703,1727]" collectionCode="AMNZ" country="New Zealand" httpUri="http://grbio.org/cool/76et-m1kt" name="Auckland Institute and Museum" pageId="117" pageNumber="118" type="Museum">40701</specimenCode>
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[green labels];
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(
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/ 0 0 22, E.
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1925 /
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/ 2593 /
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<collectorName id="26E704E60270B77B9251B575EF7FF88F" box="[573,628,1810,1835]" pageId="117" pageNumber="118">Coll.</collectorName>
<taxonomicName id="4C121AB30270B77B9211B574EE15F88F" authorityName="Castelnau" authorityYear="1867" box="[637,798,1811,1835]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="128" phylum="Arthropoda" rank="species" species="simplex">M. SIMPLEX</taxonomicName>
[hw] / 2007. 163. 10866; 1 0867; 10868; 10869; 10870 (
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:10:32 COLL.
<taxonomicName id="4C121AB30270B77B934BB55FEECEF8EB" authorityName="Castelnau" authorityYear="1867" box="[807,965,1847,1871]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="128" phylum="Arthropoda" rank="species" species="simplex">M. SIMPLEX</taxonomicName>
[hw] / 2 007.163.1086610870 (CMNZ).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0270B77B90ABB53DEC26F833" pageId="117" pageNumber="118" type="etymology">
<paragraph id="8BAD61300270B77B90ABB53DEC26F833" blockId="117.[151,1437,151,1943]" pageId="117" pageNumber="118">
<emphasis id="B966BD220270B77B90ABB53DEC5AF8D7" bold="true" box="[199,337,1882,1907]" pageId="117" pageNumber="118">Etymology:</emphasis>
This species is named to honour Te Parawhau, an inland Whangarei hapu with close connections to Ngati Wai.
</paragraph>
</subSubSection>
</treatment>
</document>