239 lines
40 KiB
XML
239 lines
40 KiB
XML
<document id="6D5E0522BD90F9783DEB108702281137" ID-CLB-Dataset="26919" ID-DOI="10.11646/zootaxa.4598.1.1" ID-GBIF-Dataset="475fd53e-c5c6-4a99-bb2e-cc2382f0171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668063" ID-ZooBank="6BA5F086-1014-46B5-B6D5-77FE0CD07AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127411994" checkinUser="plazi" docAuthor="Seldon, David S. & Buckley, Thomas R." docDate="2019" docId="03BBD0260245B74C90FBB2F0EF6BFE2C" docLanguage="en" docName="zootaxa.4598.1.1.pdf.imf" docOrigin="Zootaxa 4598 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mecodema kokoromatua Seldon, Leschen & Liebherr 2012" docType="treatment" docVersion="10" lastPageNumber="67" masterDocId="FF82A85E0205B70E906CB267ED0BFFA4" masterDocTitle="The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand" masterLastPageNumber="148" masterPageNumber="1" pageNumber="65" updateTime="1698726038790" updateUser="ExternalLinkService">
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<mods:title id="1CCCEAD36D7EA80278E7DEABCC88514E">The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand</mods:title>
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<mods:namePart id="A9824597F885CD83EEEAB6F92EBD7C2E">Seldon, David S.</mods:namePart>
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<mods:namePart id="3738427FF64F73D1F8C11B72B54DDCB0">Buckley, Thomas R.</mods:namePart>
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<mods:typeOfResource id="C6BDDF54E99DC5DC806D02469E6888D6">text</mods:typeOfResource>
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<mods:title id="A58D459125FCF620355EF64BC437DF9E">Zootaxa</mods:title>
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<mods:part id="5899C5E3DC51589FE3F055361E8DBA9A">
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<mods:date id="28863DF0D6526115DE0E1534EFB04A51">2019</mods:date>
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<mods:number id="4B70250CC8FF541F54B8BA78767411C9">2019-05-06</mods:number>
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<mods:number id="A6895D280B52E7F5FA91B77B02FAD256">4598</mods:number>
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<mods:identifier id="98A6695649CCEA58879B73EDC5F0438E" type="DOI">10.11646/zootaxa.4598.1.1</mods:identifier>
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<treatment id="03BBD0260245B74C90FBB2F0EF6BFE2C" ID-GBIF-Taxon="157062681" LSID="urn:lsid:plazi:treatment:03BBD0260245B74C90FBB2F0EF6BFE2C" httpUri="http://treatment.plazi.org/id/03BBD0260245B74C90FBB2F0EF6BFE2C" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65">
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<subSubSection id="C30832BB0245B74E90FBB2F0EE72FF16" box="[151,889,151,178]" pageId="64" pageNumber="65" type="nomenclature">
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<paragraph id="8BAD61300245B74E90FBB2F0EE72FF16" blockId="64.[151,889,151,211]" box="[151,889,151,178]" pageId="64" pageNumber="65">
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<heading id="D0E5D65C0245B74E90FBB2F0EE72FF16" bold="true" box="[151,889,151,178]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
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<emphasis id="B966BD220245B74E90FBB2F0EE72FF16" bold="true" box="[151,889,151,178]" pageId="64" pageNumber="65">
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<taxonomicName id="4C121AB30245B74E90FBB2F0EE7FFF16" ID-CoL="3YKT7" authority="Seldon, Leschen & Liebherr, 2012" authorityName="Seldon, Leschen & Liebherr" authorityYear="2012" box="[151,884,151,178]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="kokoromatua">
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<emphasis id="B966BD220245B74E90FBB2F0ECC8FF15" bold="true" box="[151,451,151,177]" italics="true" pageId="64" pageNumber="65">Mecodema kokoromatua</emphasis>
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<bibRefCitation id="EF831CC10245B74E91A6B2FFEE7FFF16" author="Seldon, D. S. & Leschen, R. A. B. & Liebherr, J. K." box="[458,884,152,178]" pageId="64" pageNumber="65" pagination="39 - 50" refId="ref93318" refString="Seldon, D. S., Leschen, R. A. B. & Liebherr, J. K. (2012) A new species of Mecodema (Carabidae: Broscini) from Northland, New Zealand, with notes on a newly observed structure within the female genitalia. New Zealand Entomologist, 35 (1), 39 - 50. https: // doi. org / 10.1080 / 00779962.2012.649707" type="journal article" year="2012">Seldon, Leschen & Liebherr, 2012</bibRefCitation>
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</taxonomicName>
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.
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30832BB0245B74E90FBB2DDEC01FF77" box="[151,266,186,211]" pageId="64" pageNumber="65" type="description">
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<paragraph id="8BAD61300245B74E90FBB2DDEC01FF77" blockId="64.[151,889,151,211]" box="[151,266,186,211]" pageId="64" pageNumber="65">
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||
<figureCitation id="13297DB50245B74E90FBB2DDEC0DFF77" box="[151,262,186,211]" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Figure 35</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30832BB0245B74E90FBB365EFC7FE23" pageId="64" pageNumber="65" type="diagnosis">
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<paragraph id="8BAD61300245B74E90FBB365EFC7FE23" blockId="64.[151,1437,258,1979]" pageId="64" pageNumber="65">
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<emphasis id="B966BD220245B74E90FBB365EC18FEBF" bold="true" box="[151,275,258,283]" pageId="64" pageNumber="65">Diagnosis:</emphasis>
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Differs from other North Island
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<taxonomicName id="4C121AB30245B74E92F8B364EE04FEBF" authorityName="Blanchard" authorityYear="1853" box="[660,783,259,283]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis id="B966BD220245B74E92F8B364EE04FEBF" box="[660,783,259,283]" italics="true" pageId="64" pageNumber="65">Mecodema</emphasis>
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</taxonomicName>
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species due to having:
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<emphasis id="B966BD220245B74E9448B365E939FEBF" bold="true" box="[1060,1074,258,283]" pageId="64" pageNumber="65">1</emphasis>
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, less than or equal to six setae along the pronotal carina (
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<taxonomicName id="4C121AB30245B74E91D0B341EF22FE9A" authorityName="Broun" authorityYear="1915" box="[444,553,294,318]" class="Insecta" family="Carabidae" genus="Metaglymma" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="curvidens">
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<emphasis id="B966BD220245B74E91D0B341EF22FE9A" box="[444,553,294,318]" italics="true" pageId="64" pageNumber="65">curvidens</emphasis>
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</taxonomicName>
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species group);
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<emphasis id="B966BD220245B74E92B3B341EFE6FE9B" bold="true" box="[735,749,294,319]" pageId="64" pageNumber="65">2</emphasis>
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, mesocoxa and metacoxa with setose punctures absent;
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<emphasis id="B966BD220245B74E9532B341E867FE9B" bold="true" box="[1374,1388,294,319]" pageId="64" pageNumber="65">3</emphasis>
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, the shape of the left paramere basal lobe is a small and very narrowly rectangular with a short slope to arm, terminal lobe with tuft of medium length setae (
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<figureCitation id="13297DB50245B74E9226B309EF90FE23" box="[586,667,366,391]" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Fig. 35</figureCitation>
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PL).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30832BB0245B74E90ABB3F4EEBEF81F" pageId="64" pageNumber="65" type="description">
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<paragraph id="8BAD61300245B74E90ABB3F4EFD9FE74" blockId="64.[151,1437,258,1979]" pageId="64" pageNumber="65">
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<emphasis id="B966BD220245B74E90ABB3F4EC50FE08" bold="true" box="[199,347,403,428]" pageId="64" pageNumber="65">Description:</emphasis>
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Length
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<quantity id="4CEACCD50245B74E91DCB3F4EF2CFE0F" box="[432,551,403,427]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="64" pageNumber="65" unit="mm" value="23.5" valueMax="25.0" valueMin="22.0">22–25 mm</quantity>
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, pronotal width
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<quantity id="4CEACCD50245B74E92BFB3F4EE4DFE0F" box="[723,838,403,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="64" pageNumber="65" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
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, elytral width
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<quantity id="4CEACCD50245B74E938AB3F4E951FE0F" box="[998,1114,403,428]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="64" pageNumber="65" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">5–5.5 mm</quantity>
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. Colour of entire body matte to glossy black, coxae may be dark reddish-brown.
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</paragraph>
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<paragraph id="8BAD61300245B74E90ABB3BBE995FCB7" blockId="64.[151,1437,258,1979]" pageId="64" pageNumber="65">
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<emphasis id="B966BD220245B74E90ABB3BBEC06FE50" box="[199,269,476,500]" italics="true" pageId="64" pageNumber="65">Head:</emphasis>
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Narrow and flat (
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<figureCitation id="13297DB50245B74E9185B3BCEF5EFE50" box="[489,597,475,500]" captionStart="FIGURE 14" captionStartId="20.[151,250,936,958]" captionTargetBox="[171,1412,204,911]" captionTargetId="figure@20.[160,1412,193,921]" captionTargetPageId="20" captionText="FIGURE 14. Diagrammatic cross sections that represent the width and convexity of head or deflection of pronotum (frontal view): (A) broad and flat; (B) broad and convex; (C) narrow and flat; (D) narrow and convex." httpUri="https://zenodo.org/record/2668091/files/figure.png" pageId="64" pageNumber="65">Fig. 14C</figureCitation>
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). Vertex smooth; vertexal groove narrow and well-defined, posteriorly curved medially; large supraorbital puncture (
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<figureCitation id="13297DB50245B74E92E6B398EFC0FDBC" box="[650,715,511,536]" captionStart="FIGURE 9" captionStartId="15.[151,250,1706,1728]" captionTargetBox="[157,1433,759,1683]" captionTargetId="figure@15.[151,1436,757,1683]" captionTargetPageId="15" captionText="FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove." httpUri="https://zenodo.org/record/2668081/files/figure.png" pageId="64" pageNumber="65">Fig. 9</figureCitation>
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) bearing 2–3 setae; 2 defined supraorbital grooves; frons with a small and shallow depression either side of midline; frontoclypeal suture straight, very narrow and well defined between small tentorial pits; anterior clypeal microsculpture indistinct, 1 large setose puncture each side bearing 2 setae. Labrum (
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<figureCitation id="13297DB50245B74E9122B00CEC9EFD20" box="[334,405,619,644]" captionStart="FIGURE 9" captionStartId="15.[151,250,1706,1728]" captionTargetBox="[157,1433,759,1683]" captionTargetId="figure@15.[151,1436,757,1683]" captionTargetPageId="15" captionText="FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove." httpUri="https://zenodo.org/record/2668081/files/figure.png" pageId="64" pageNumber="65">Fig. 9</figureCitation>
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) rectangular, anterior edge outwardly curved with 2 proximate central setae, 2 each side evenly spaced. Mentum lobes squared (
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<figureCitation id="13297DB50245B74E9238B0E8EFB5FD0C" box="[596,702,655,680]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations A–D are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations E–G are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="64" pageNumber="65">Fig. 12G</figureCitation>
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), median process long and narrow, slightly angled upward (15°), indentation notched (
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<figureCitation id="13297DB50245B74E911AB0D4ECEBFD68" box="[374,480,691,716]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations A–D are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations E–G are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="64" pageNumber="65">Fig. 12D</figureCitation>
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); mentum setae variably present or absent. Submentum sclerite constriction narrow bearing 6–8 evenly spaced setae. Stipes with 2 basal setae. Gula pits broad and deep, gula suture well-defined, gula flat and smooth. Gena with fine wrinkles forming an isodiametric pattern across entire area.
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</paragraph>
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<paragraph id="8BAD61300245B74E90ABB178ECCDFBB4" blockId="64.[151,1437,258,1979]" pageId="64" pageNumber="65">
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<emphasis id="B966BD220245B74E90ABB178EC49FC93" box="[199,322,799,823]" italics="true" pageId="64" pageNumber="65">Prothorax:</emphasis>
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Prothoracic carina very narrow the entire length, crenulations absent with 4–6 setae each side (
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<figureCitation id="13297DB50245B74E9503B178EDBDFCF8" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Fig. 35</figureCitation>
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), extended to anterior angle; posterior lateral sinuation evidently carinate, angled slightly inward; pronotum narrow, medially flattened but deflected laterally, overall shape squared (
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<figureCitation id="13297DB50245B74E938DB100E933FC24" box="[993,1080,871,896]" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Fig. 35</figureCitation>
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); midline well-defined without medial impressions, disc with fine, transverse lines from midline to laterally, pronotal foveae narrow and shallow; anterior edge slightly curved to straight, posterior edge curved medially. Prosternum weakly convex with very fine transverse lines present; proepisternum dorso-ventrally wrinkled. Procoxal setae absent; protibia distally expanded and shovel-like (
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<figureCitation id="13297DB50245B74E913EB190ECB2FBB4" box="[338,441,1015,1040]" captionStart="FIGURE 10" captionStartId="16.[151,250,1442,1464]" captionTargetBox="[152,1428,191,1421]" captionTargetId="figure@16.[151,1435,183,1427]" captionTargetPageId="16" captionText="FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used." httpUri="https://zenodo.org/record/2668083/files/figure.png" pageId="64" pageNumber="65">Fig. 10B</figureCitation>
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).
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</paragraph>
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<paragraph id="8BAD61300245B74E90ABB67BEC44FAAB" blockId="64.[151,1437,258,1979]" pageId="64" pageNumber="65">
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<emphasis id="B966BD220245B74E90ABB67BEC1DFB90" box="[199,278,1052,1076]" italics="true" pageId="64" pageNumber="65">Elytra:</emphasis>
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||
Narrow and flat to moderately deflected laterally; humeral angle evenly convex (
|
||
<figureCitation id="13297DB50245B74E94CDB67CE9F3FB90" box="[1185,1272,1051,1076]" captionStart="FIGURE 6" captionStartId="10.[151,250,519,541]" captionTargetBox="[156,1432,193,498]" captionTargetId="figure@10.[151,1436,193,498]" captionTargetPageId="10" captionText="FIGURE 6. Detail of the three different forms of the elytral humeral angle: (A) = anteriorly convergent (M. parataiko), (B) = evenly convex (M. oconnori), (C) = subangulate (M. atuanui)." httpUri="https://zenodo.org/record/2668075/files/figure.png" pageId="64" pageNumber="65">Fig. 6B</figureCitation>
|
||
); basal margin slightly curved and bevelled to base, interval 1 extended to scutellum; lateral carina narrow the entire length, extended beyond humeral angle; humerus with 3 proximate setose punctures along angle, 1 double-spaced posterad; suture impressed but no more so than striae; striae with regularly spaced asetose punctures that increased in size laterally; intervals 1–6 flat or weakly convex, intervals 7–9 weakly convex; interval microsculpture absent; 7
|
||
<superScript id="7C67CC780245B74E90C9B6A9EDBAFB78" attach="left" box="[165,177,1230,1244]" fontSize="6" pageId="64" pageNumber="65">th</superScript>
|
||
strial setal pattern with 1 setose puncture in anterior ½, 3–4 setose punctures in posterior ½ (
|
||
<figureCitation id="13297DB50245B74E9487B6B5E84AFB4F" box="[1259,1345,1234,1259]" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Fig. 35</figureCitation>
|
||
), setose punctures small.
|
||
</paragraph>
|
||
<paragraph id="8BAD61300245B74E90ABB77DE900FA43" blockId="64.[151,1437,258,1979]" pageId="64" pageNumber="65">
|
||
<emphasis id="B966BD220245B74E90ABB77DEC75FA96" box="[199,382,1306,1330]" italics="true" pageId="64" pageNumber="65">Ventral surface:</emphasis>
|
||
Mesepisternum (
|
||
<figureCitation id="13297DB50245B74E922BB77DEF85FA97" box="[583,654,1306,1331]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1–Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="64" pageNumber="65">Fig. 2</figureCitation>
|
||
) wrinkled; metepisternum without microsculpture; setose punctures absent on mesocoxae and metacoxae. Abdominal ventrites 1 and 2 without microsculpture and punctures; ventrites 3 and 4 setose punctures absent, ventrite 5 with 1 setose puncture each side of midline; ventrite 6 (
|
||
<figureCitation id="13297DB50245B74E9561B705E859FADF" box="[1293,1362,1378,1403]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1–Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="64" pageNumber="65">Fig. 2</figureCitation>
|
||
) setae present:
|
||
<specimenCount id="9D14AAB90245B74E909BB7E0EC07FA3A" box="[247,268,1415,1438]" pageId="64" pageNumber="65" type="male">♂</specimenCount>
|
||
with 3 setae each side, apical edge obtusely rounded,
|
||
<specimenCount id="9D14AAB90245B74E9302B7E0EE88FA3A" box="[878,899,1415,1438]" pageId="64" pageNumber="65" type="female">♀</specimenCount>
|
||
with 2 setae each side of midline, apical edge a flattened curve; ventrites 3–5 foveate laterally (very shallow, almost indistinct). Anterior metaventrite process a bluntly rounded triangle with a well-defined and broad carina the entire length.
|
||
</paragraph>
|
||
<paragraph id="8BAD61300245B74E90ABB795EE23F8EB" blockId="64.[151,1437,258,1979]" pageId="64" pageNumber="65">
|
||
<emphasis id="B966BD220245B74E90ABB795EC7EF9AE" box="[199,373,1522,1546]" italics="true" pageId="64" pageNumber="65">Male genitalia:</emphasis>
|
||
Apical portion of penis lobe symmetrically rounded (
|
||
<figureCitation id="13297DB50245B74E93B2B795E924F9AF" box="[990,1071,1522,1547]" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Fig. 35</figureCitation>
|
||
PL) with a distinct deflection to right of vertical axis (VV) (
|
||
<figureCitation id="13297DB50245B74E91A4B471EF3BF98B" box="[456,560,1558,1583]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="64" pageNumber="65">Fig. 15A</figureCitation>
|
||
); apex with dorsal curve slightly pushed forward of vertical axis (
|
||
<figureCitation id="13297DB50245B74E9560B471E844F98B" box="[1292,1359,1558,1583]" captionStart="FIGURE 3" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[155,1433,1029,1723]" captionTargetId="figure@6.[151,1436,1023,1723]" captionTargetPageId="6" captionText="FIGURE 3. Detail of the apical portion of aedeagus (male genitalia) with parameres removed showing taxonomic structures, vertical and horizontal planes that are referred to in the species descriptions and redescriptions: aedeagus shaft width (ASW); vertical central line (ƑCL) [dashed line, not a structure]; horizontal central line (HCL) [dashed line, not a structure]." httpUri="https://zenodo.org/record/2668069/files/figure.png" pageId="64" pageNumber="65">Fig. 3</figureCitation>
|
||
); shaft of penis lobe straight (VV), greatly broadened toward base; overall length of penis lobe slightly curved ventrally (LV) (
|
||
<figureCitation id="13297DB50245B74E90B3B439EC41F9D3" box="[223,330,1630,1655]" captionStart="FIGURE 16" captionStartId="21.[151,250,897,919]" captionTargetBox="[151,1436,181,869]" captionTargetId="figure@21.[151,1436,181,876]" captionTargetPageId="21" captionText="FIGURE 16. Relative strength of the ventral curve over the length of the penis lobe (lateral view): (A) slightly curved (M. oblongum); (B) moderately curved (M. ponaiti); (C) distinctly curved (M. manaia). Scale bar = 1 mm." httpUri="https://zenodo.org/record/2668095/files/figure.png" pageId="64" pageNumber="65">Fig. 16A</figureCitation>
|
||
). Structures of the endophallus (
|
||
<figureCitation id="13297DB50245B74E92A8B439EE06F9D3" box="[708,781,1630,1655]" captionStart="FIGURE 4" captionStartId="7.[151,250,973,995]" captionTargetBox="[354,1232,193,952]" captionTargetId="figure@7.[354,1232,193,952]" captionTargetPageId="7" captionText="FIGURE 4. An Environmental Scanning Electron Microscope (ESEM) photograph of the four main structures of the apical plate of the endophallus within the penis lobe. This is an example from Mecodema haunoho showing the central spicule (c.s), left (l.s.f) and the right setose flange (r.s.f), and microtrichial field (m.f). These structures are variable among species but within a species, they are very consistent, even over distant geographical ranges. The microtrichial field can be covered in flattened scales as in photo above, or with elongated setae-like scales." httpUri="https://zenodo.org/record/2668071/files/figure.png" pageId="64" pageNumber="65">Fig. 4</figureCitation>
|
||
): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule narrow and short; setal (scales) coverage of the apical plate extensive (76–100%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe small and very narrowly rectangular with a short slope to arm (
|
||
<figureCitation id="13297DB50245B74E9297B4ADEE45F947" box="[763,846,1738,1763]" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Fig. 35</figureCitation>
|
||
LP); arm long (nearly equal to basal lobe); terminal lobe indistinct from arm, apical tuft of medium-lengthed setae, with very short, sparse setae extended along apical ½ of ventral edge (
|
||
<figureCitation id="13297DB50245B74E9104B575ECB2F88F" box="[360,441,1810,1835]" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Fig. 35</figureCitation>
|
||
LP). Right paramere similar in size and shape to left with a double-row of medium length setae extended along apical ¾ of ventral edge (
|
||
<figureCitation id="13297DB50245B74E92C8B551EFFDF8EB" box="[676,758,1846,1871]" captionStart="FIGURE 35" captionStartId="65.[151,250,1847,1869]" captionTargetBox="[259,1325,208,1812]" captionTargetId="figure@65.[251,1336,193,1826]" captionTargetPageId="65" captionText="FIGURE 35. Mecodema kokoromatua plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="64" pageNumber="65">Fig. 35</figureCitation>
|
||
RP).
|
||
</paragraph>
|
||
<paragraph id="8BAD61300245B74E90ABB53DEEBEF81F" blockId="64.[151,1437,258,1979]" pageId="64" pageNumber="65">
|
||
<emphasis id="B966BD220245B74E90ABB53DEC85F8D6" box="[199,398,1882,1906]" italics="true" pageId="64" pageNumber="65">Female genitalia:</emphasis>
|
||
Basal gonocoxite 1 narrow at base, width doubled apically, internal dorso-lateral carina with 2 apical setae, ventral surface entirely covered with grooves. Gonocoxite 2 (
|
||
<figureCitation id="13297DB50245B74E9460B519E95AF833" box="[1036,1105,1918,1943]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="64" pageNumber="65">Fig. 5</figureCitation>
|
||
) paddle-like, ventral surface convex with sensillar pits. Ramus long and narrow, broadened apically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6D31B80244B74F90FBB550E808F82E" httpUri="https://zenodo.org/record/2668133/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1847,1869]" targetBox="[259,1325,208,1812]" targetPageId="65">
|
||
<paragraph id="8BAD61300244B74F90FBB550E808F82E" blockId="65.[151,1437,1847,1930]" pageId="65" pageNumber="66">
|
||
<emphasis id="B966BD220244B74F90FBB550EC2AF8E9" bold="true" box="[151,289,1847,1869]" pageId="65" pageNumber="66">FIGURE 35.</emphasis>
|
||
<taxonomicName id="4C121AB30244B74F9144B550EF2FF8E9" authorityName="Seldon, Leschen & Liebherr" authorityYear="2012" box="[296,548,1847,1869]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="kokoromatua">
|
||
<emphasis id="B966BD220244B74F9144B550EF2FF8E9" box="[296,548,1847,1869]" italics="true" pageId="65" pageNumber="66">Mecodema kokoromatua</emphasis>
|
||
</taxonomicName>
|
||
plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30832BB0247B74C90ABB2F0EF6BFE2C" pageId="66" pageNumber="67" type="discussion">
|
||
<paragraph id="8BAD61300247B74C90ABB2F0EF6BFE2C" blockId="66.[151,1436,151,392]" pageId="66" pageNumber="67">
|
||
<emphasis id="B966BD220247B74C90ABB2F0EC5AFF14" bold="true" box="[199,337,151,176]" pageId="66" pageNumber="67">Comments:</emphasis>
|
||
<taxonomicName id="4C121AB30247B74C9131B2FFEF73FF14" authorityName="Seldon, Leschen & Liebherr" authorityYear="2012" box="[349,632,152,176]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="kokoromatua">
|
||
<emphasis id="B966BD220247B74C9131B2FFEF73FF14" box="[349,632,152,176]" italics="true" pageId="66" pageNumber="67">Mecodema kokoromatua</emphasis>
|
||
</taxonomicName>
|
||
is restricted to coastal sand dune forest at the base of the Ahipara
|
||
<materialsCitation id="3B7A6B6D0247B74C90FBB2DAE897FEB8" ID-GBIF-Occurrence="2237851776" collectionCode="NZAC" country="New Zealand" county="North Head" location="New Zealand" municipality="North Head" pageId="66" pageNumber="67" specimenCount="1" stateProvince="Northland" typeStatus="holotype">
|
||
Escarpment.
|
||
<emphasis id="B966BD220247B74C90ABB2B8EC69FF5C" bold="true" box="[199,354,223,248]" pageId="66" pageNumber="67">Distribution:</emphasis>
|
||
<location id="8ECD37EB0247B74C9105B287EF09FF5C" LSID="urn:lsid:plazi:treatment:03BBD0260245B74C90FBB2F0EF6BFE2C:8ECD37EB0247B74C9105B287EF09FF5C" box="[361,514,223,248]" country="New Zealand" county="North Head" municipality="North Head" name="New Zealand" pageId="66" pageNumber="67" stateProvince="Northland">New Zealand</location>
|
||
,
|
||
<location id="8ECD37EB0247B74C9262B2B8EF96FF5C" LSID="urn:lsid:plazi:treatment:03BBD0260245B74C90FBB2F0EF6BFE2C:8ECD37EB0247B74C9262B2B8EF96FF5C" box="[526,669,223,248]" country="New Zealand" county="North Head" municipality="North Head" name="North Island" pageId="66" pageNumber="67" stateProvince="Northland">North Island</location>
|
||
,
|
||
<collectingRegion id="49D6AFD20247B74C92C4B2B8EE16FF5C" box="[680,797,223,248]" country="New Zealand" name="Northland" pageId="66" pageNumber="67">Northland</collectingRegion>
|
||
,
|
||
<location id="8ECD37EB0247B74C9344B2B8EE98FF5C" LSID="urn:lsid:plazi:treatment:03BBD0260245B74C90FBB2F0EF6BFE2C:8ECD37EB0247B74C9344B2B8EE98FF5C" box="[808,915,223,248]" country="New Zealand" county="North Head" municipality="North Head" name="Herekino" pageId="66" pageNumber="67" stateProvince="Northland">Herekino</location>
|
||
,
|
||
<collectingMunicipality id="6BC9FB4A0247B74C93F2B2B8E92FFF5C" box="[926,1060,223,248]" pageId="66" pageNumber="67">North Head</collectingMunicipality>
|
||
, Waiatua Stream.
|
||
<emphasis id="B966BD220247B74C90ABB363EC36FEB9" bold="true" box="[199,317,260,285]" pageId="66" pageNumber="67">
|
||
<typeStatus id="54A9DF920247B74C90ABB363EC33FEB9" box="[199,312,260,285]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectionCode id="ED03F9F50247B74C912AB363EC99FEB8" box="[326,402,260,284]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="66" pageNumber="67" type="Other institutional type">NZAC</collectionCode>
|
||
male labelled.
|
||
<collectingCountry id="F30521A00247B74C9229B362EE1EFEB8" box="[581,789,260,284]" name="New Zealand" pageId="66" pageNumber="67">NEW ZEALAND</collectingCountry>
|
||
, ND, Herekino,
|
||
<collectingCounty id="62CC19BC0247B74C93B8B363E957FEB9" box="[980,1116,260,285]" pageId="66" pageNumber="67">North Head</collectingCounty>
|
||
, Waiatua Stream, 8–23 Dec
|
||
</materialsCitation>
|
||
2010. D.S. Seldon,
|
||
<geoCoordinate id="EE2607F70247B74C911FB340EF15FEE4" box="[371,542,295,320]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="1" value="-35.269737">35°16’11.052S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2607F70247B74C9245B340EFE9FEE4" box="[553,738,295,320]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="1" value="173.1598">173°09’35.314E</geoCoordinate>
|
||
/
|
||
<typeStatus id="54A9DF920247B74C9295B34FEE86FEE4" box="[761,909,296,320]" pageId="66" pageNumber="67" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="4C121AB30247B74C93F8B340E9A7FEE4" authority="Seldon & Buckley, 2019" authorityName="Seldon & Buckley" authorityYear="2019" box="[916,1196,295,320]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="kokoromatua" status="sp. nov.">Mecodema kokoromatua</taxonomicName>
|
||
<taxonomicNameLabel id="A25500590247B74C94DFB34FE9E3FE9B" box="[1203,1256,296,319]" pageId="66" pageNumber="67" rank="species">sp.n.</taxonomicNameLabel>
|
||
design.
|
||
<bibRefCitation id="EF831CC10247B74C952BB340E897FEE4" author="Seldon, D. S. & Leschen, R. A. B." box="[1351,1436,295,320]" pageId="66" pageNumber="67" pagination="1 - 45" refId="ref93265" refString="Seldon, D. S. & Leschen, R. A. B. (2011) Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini). Zootaxa, 2829, 1 - 45. https: // doi. org / 10.11646 / zootaxa. 2829.1.1" type="journal article" year="2011">Seldon,</bibRefCitation>
|
||
Leschen & Liebherr 2011 [red label].
|
||
<emphasis id="B966BD220247B74C90ABB308EC43FE2C" bold="true" box="[199,328,367,392]" pageId="66" pageNumber="67">
|
||
<typeStatus id="54A9DF920247B74C90ABB308EC48FE2C" box="[199,323,367,392]" pageId="66" pageNumber="67" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
(see
|
||
<bibRefCitation id="EF831CC10247B74C91EFB308EF5FFE2C" author="Seldon, D. S. & Leschen, R. A. B. & Liebherr, J. K." box="[387,596,367,392]" pageId="66" pageNumber="67" pagination="39 - 50" refId="ref93318" refString="Seldon, D. S., Leschen, R. A. B. & Liebherr, J. K. (2012) A new species of Mecodema (Carabidae: Broscini) from Northland, New Zealand, with notes on a newly observed structure within the female genitalia. New Zealand Entomologist, 35 (1), 39 - 50. https: // doi. org / 10.1080 / 00779962.2012.649707" type="journal article" year="2012">
|
||
Seldon
|
||
<emphasis id="B966BD220247B74C91B4B316EF1BFE2C" box="[472,528,368,392]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |