156 lines
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156 lines
16 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.976.53859" ID-GBIF-Dataset="3c8d8e79-b176-41e6-87c9-fb6f4b5ff9a6" ID-PMC="PMC7591553" ID-Pensoft-Pub="1313-2970-976-1" ID-Pensoft-UUID="1C1677B3CFE65ECEADF5CA56DACD0B9C" ID-PubMed="33173395" ID-ZooBank="B755A1D5D42D4CA589BE10C11EAB4580" ModsDocID="1313-2970-976-1" checkinTime="1603239875264" checkinUser="pensoft" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Tongkerd, Piyoros & Panha, Somsak" docDate="2020" docId="C9B5B1BBDB3F5CD18FBF0DE40EAAF7E5" docLanguage="en" docName="ZooKeys 976: 1-100" docOrigin="ZooKeys 976" docSource="http://dx.doi.org/10.3897/zookeys.976.53859" docTitle="Sarika subcornea" docType="treatment" docVersion="5" id="1C1677B3CFE65ECEADF5CA56DACD0B9C" lastPageNumber="1" masterDocId="1C1677B3CFE65ECEADF5CA56DACD0B9C" masterDocTitle="Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae)" masterLastPageNumber="100" masterPageNumber="1" pageNumber="1" updateTime="1668169619115" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pholyotha, Arthit</mods:namePart>
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<mods:affiliation>Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6677-1164</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>976</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>100</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.976.53859</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.976.53859</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-976-1</mods:identifier>
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<mods:identifier type="ZooBank">B755A1D5D42D4CA589BE10C11EAB4580</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1C1677B3CFE65ECEADF5CA56DACD0B9C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168504527" LSID="urn:lsid:plazi:treatment:C9B5B1BBDB3F5CD18FBF0DE40EAAF7E5" httpUri="http://treatment.plazi.org/id/C9B5B1BBDB3F5CD18FBF0DE40EAAF7E5" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="C9B5B1BB-DB3F-5CD1-8FBF-0DE40EAAF7E5" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika subcornea" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subcornea">Sarika subcornea (Pfeiffer, 1861)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix subcornea" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subcornea">Helix subcornea</taxonomicName>
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Pfeiffer, 1861: 20. Type locality:
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<normalizedToken originalValue="“Siam”">"Siam"</normalizedToken>
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[Thailand];
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceeding of the Zoological Society of London" pageId="0" pageNumber="1" refId="B64" refString="Pfeiffer, L, 1868. Monographia Heliceorum Viventium, Supplementum Tertium. Volume 5. Lipsiae: F.A. Brockhaus, 565 pp." title="Monographia Heliceorum Viventium, Supplementum Tertium. Volume 5. Lipsiae: F. A. Brockhaus, 565 pp." year="1868">Pfeiffer 1868</bibRefCitation>
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: 102, 103.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Orobia) subcornea" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subcornea" subGenus="Orobia">Nanina (Orobia) subcornea</taxonomicName>
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:
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<bibRefCitation author="Martens von, E" journalOrPublisher="Unpublished Master's Thesis, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok" pageId="0" pageNumber="1" refId="B45" refString="Martens von, E, 1867. Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." title="Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." year="1867">Martens 1867</bibRefCitation>
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: 73.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Macrochlamys) subcornea" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subcornea" subGenus="Macrochlamys">Nanina (Macrochlamys) subcornea</taxonomicName>
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: Tryon 1886: 92;
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<bibRefCitation author="Fischer, H" journalOrPublisher="In: Mission Pavie, Etudes Diverses" pageId="0" pageNumber="1" pagination="390 - 450" refId="B19" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo- Chine orientale cites jusqu' a ce jour. In: Mission Pavie, Etudes Diverses 3: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo- Chine orientale cites jusqu' a ce jour." volume="3" year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 395.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Ariophanta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariophanta (Macrochlamys) subcornea" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subcornea" subGenus="Macrochlamys">Ariophanta (Macrochlamys) subcornea</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.14809" author="Fischer, P" journalOrPublisher="Imprimerie Dejussieu Pere et Fils, Autun" pageId="0" pageNumber="1" refId="B18" refString="Fischer, P, 1891. Catalogue et distribution geographique des Mollusques terrestres, fluviatiles & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Imprimerie Dejussieu Pere et Fils, Autun, DOI: https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution geographique des Mollusques terrestres, fluviatiles & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." url="https://doi.org/10.5962/bhl.title.14809" year="1891">Fischer 1891</bibRefCitation>
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: 21.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina subcornea" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subcornea">Nanina subcornea</taxonomicName>
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:
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<bibRefCitation author="Fischer, H" journalOrPublisher="In: Mission Pavie, Etudes Diverses" pageId="0" pageNumber="1" pagination="390 - 450" refId="B19" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo- Chine orientale cites jusqu' a ce jour. In: Mission Pavie, Etudes Diverses 3: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo- Chine orientale cites jusqu' a ce jour." volume="3" year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 395;
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<bibRefCitation author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie aus dem Haus der Natur-Cismar" pageId="0" pageNumber="1" pagination="35 - 70" refId="B31" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie aus dem Haus der Natur-Cismar 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
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: 45.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika subcornea" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subcornea">Sarika subcornea</taxonomicName>
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:
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<bibRefCitation author="Maassen, WJM" journalOrPublisher="De Kreukel, Amsterdam" pageId="0" pageNumber="1" refId="B43" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia. "A Handlist". De Kreukel, Amsterdam" title="A preliminary checklist of the non-marine molluscs of West-Malaysia. " A Handlist "." year="2001">Maassen 2001</bibRefCitation>
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: 114.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">The type specimen could not be located in the NHM collections.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shell depressed, medium size (shell width up to 13.6 mm, shell height up to 6.0 mm) and thin. Shell surface smooth and shining; shell colour whitish horny. Whorls
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<normalizedToken originalValue="7½">71/2</normalizedToken>
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, increasing regularly; body whorl well rounded. Spire elevated; suture impressed. Aperture crescent-shaped and obliquely opened. Peristome simple and slightly thickened. Columellar margin simple. Umbilicus narrowly opened.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The original description is not illustrated, type specimen could not be traced, and no new specimens have been reported so far. Hanley and Theobald (1876: 59, pl. 149, Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Schematic drawing of the mantle terminology of Sarika showing five mantle lobes (two shell lobes and three dorsal lobes)." figureDoi="10.3897/zookeys.976.53859.figure2" httpUri="https://binary.pensoft.net/fig/464384" pageId="0" pageNumber="1">2</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. An illustrated synopsis of the shell and penis. A shell shape B whorl periphery C inner sculpture of the penis D appendage inside penis (no penial verge, pseudo-verge, and penial verge). Yellow arrowheads indicate the invagination of the penial wall." figureDoi="10.3897/zookeys.976.53859.figure3" httpUri="https://binary.pensoft.net/fig/464385" pageId="0" pageNumber="1">3</figureCitation>
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) illustrated this species based on a specimen from "Phie Than" [Payathonzu, Myanmar]. But,
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<bibRefCitation author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B8" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London" title="The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." year="1908">Blanford and Godwin-Austen (1908)</bibRefCitation>
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stated that this specimen was instead a juvenile shell of
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<taxonomicName lsidName="S. resplendens" pageId="0" pageNumber="1" rank="species" species="resplendens">
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<emphasis italics="true" pageId="0" pageNumber="1">S. resplendens</emphasis>
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</taxonomicName>
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. Recently,
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<bibRefCitation author="Maassen, WJM" journalOrPublisher="De Kreukel, Amsterdam" pageId="0" pageNumber="1" refId="B43" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia. "A Handlist". De Kreukel, Amsterdam" title="A preliminary checklist of the non-marine molluscs of West-Malaysia. " A Handlist "." year="2001">Maassen (2001)</bibRefCitation>
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also reported
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<taxonomicName lsidName="S. subcornea" pageId="0" pageNumber="1" rank="species" species="subcornea">
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<emphasis italics="true" pageId="0" pageNumber="1">S. subcornea</emphasis>
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</taxonomicName>
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as a new record from Kedah, Malaysia, but unfortunately without illustration. Therefore, the taxonomic status of this species is still unclear, and we retained this species within the genus
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
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</taxonomicName>
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as per
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<bibRefCitation author="Maassen, WJM" journalOrPublisher="De Kreukel, Amsterdam" pageId="0" pageNumber="1" refId="B43" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia. "A Handlist". De Kreukel, Amsterdam" title="A preliminary checklist of the non-marine molluscs of West-Malaysia. " A Handlist "." year="2001">Maassen (2001)</bibRefCitation>
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.
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<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Theobald and Hanley's Conchologia Indica, Taylor & Francis, London. Volume 1" pageId="0" pageNumber="1" refId="B24" refString="Godwin-Austen, HH, 1883. [1888 [1882-1888]] Land and Freshwater Mollusca of India, Including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other Islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica, Taylor & Francis, London. Volume 1" title="[1888 [1882 - 1888 ]] Land and Freshwater Mollusca of India, Including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other Islands of the Indian Ocean. Supplementary to Messrs." year="1883">Godwin-Austen (1883)</bibRefCitation>
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noted that
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<taxonomicName lsidName="S. subcornea" pageId="0" pageNumber="1" rank="species" species="subcornea">
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<emphasis italics="true" pageId="0" pageNumber="1">S. subcornea</emphasis>
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</taxonomicName>
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is distinguished from other species by the relatively more numerous and more closely wound whorls.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |