treatments-xml/data/CE/0F/8E/CE0F8E0D4F2E8D5DFC1EF91CFA154768.xml
2024-06-21 12:52:15 +02:00

2069 lines
286 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="3C849156F6FFAF6C77A6FA9B3042FFD7" ID-DOI="10.1093/jmammal/gyaa104" ID-ISSN="0022-2372" ID-Zenodo-Dep="10632393" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1707335569122" checkinUser="felipe" docAuthor="Moreno Cárdenas, Pablo A, Tinoco, Nicolás, Albuja, Luis &amp; Patterson, Bruce D" docDate="2021" docId="CE0F8E0D4F2E8D5DFC1EF91CFA154768" docLanguage="en" docName="JMammal.102.1.123-138.pdf" docOrigin="Journal of Mammalogy (J. Mammal.) 102 (1)" docSource="http://dx.doi.org/10.1093/jmammal/gyaa104" docStyle="DocumentStyle:8488A0FD6434D222CC79ECAD85E66EFE.4:JMammal.2015-.journal_article" docStyleId="8488A0FD6434D222CC79ECAD85E66EFE" docStyleName="JMammal.2015-.journal_article" docStyleVersion="4" docTitle="Rhagomys septentrionalis Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson 2021, sp. nov." docType="treatment" docVersion="1" lastPageNumber="135" masterDocId="3236F6754F2D8D51FFDAFFA3FFE94228" masterDocTitle="A new species of Rhagomys (Rodentia, Sigmodontinae) from southeastern Ecuador" masterLastPageNumber="138" masterPageNumber="123" pageNumber="126" updateTime="1708032465684" updateUser="karina">
<mods:mods id="024E3B2847722B27C735427A7B8DB70A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A9E9FCDDC8BC68F4F50C928D7B3425C1">
<mods:title id="0CB5986DFC4843D2895FB67FB3831194">A new species of Rhagomys (Rodentia, Sigmodontinae) from southeastern Ecuador</mods:title>
</mods:titleInfo>
<mods:name id="AFA3A9EEE16D0BB154FC7B4454FAF408" type="personal">
<mods:role id="AA6EE0506262D968DF3E1B9F9031E9AF">
<mods:roleTerm id="E037046EA8043140AF97F120C20BDD47">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7EF2EE6927122F1C2B1B5E0B930B9F52">Moreno Cárdenas, Pablo A</mods:namePart>
</mods:name>
<mods:name id="5328DC7A2BB71C669ACA2C04FAC7541B" type="personal">
<mods:role id="DDDB199BA2975D4E13808A8376BE4336">
<mods:roleTerm id="42B085B1E0AC17FF64BE6F5BAA089CD6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="149D5C3335A1D5DFE7CB5ADA09B73A96">Tinoco, Nicolás</mods:namePart>
</mods:name>
<mods:name id="BFB8895F2690A3F604C868562FD2BA8D" type="personal">
<mods:role id="CEFDF9774935E51587C22325ED4494C6">
<mods:roleTerm id="950F90AFDA0D1D5A6BDFCF56392DF476">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF54E5AA51F37C1E9727B4A8AA632F95">Albuja, Luis</mods:namePart>
</mods:name>
<mods:name id="B2F26BC47224CE821D9381923E21B09A" type="personal">
<mods:role id="738E758077C9D21606550B6CCD2B0794">
<mods:roleTerm id="BBCC5ACC6A6D944A3A9087EBB6D8A0DD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BFF31A0BC2FEA8B538E4DB98F7E1511C">Patterson, Bruce D</mods:namePart>
</mods:name>
<mods:typeOfResource id="8B3EFE03FB39855D659ABFF718DD7F57">text</mods:typeOfResource>
<mods:relatedItem id="E8AF2E1DEFAB417F5344536E1EB1373D" type="host">
<mods:titleInfo id="D07191AE609C48982C181C3EED89D705">
<mods:title id="93241A1848945094786DA7075D015D6F">Journal of Mammalogy</mods:title>
</mods:titleInfo>
<mods:part id="149E5B0A31530780DB17B0B0928F95CB">
<mods:date id="47F853352A627A0F00843ABC15748121">2021</mods:date>
<mods:detail id="C6EC6B2A869829CFE9DE004875A764CA" type="series">
<mods:title id="CEF0FB43104626F8F190814D4D8BD81E">J. Mammal.</mods:title>
</mods:detail>
<mods:detail id="26821CCEF5D3470D62FA40E7DC68040C" type="pubDate">
<mods:number id="6D8A25D24B01BECF4329E2AD75EC08BA">2021-03-17</mods:number>
</mods:detail>
<mods:detail id="23E4F22C9D30FB41F4DA8A62893C2681" type="volume">
<mods:number id="415607D9C34692A5B633408F104AD8E2">102</mods:number>
</mods:detail>
<mods:detail id="9297B7ADC41C8BBE92E225C6CE7E8B54" type="issue">
<mods:number id="2F2296CD688F06CC4997EDA3E3F2F0F7">1</mods:number>
</mods:detail>
<mods:extent id="440310A1D0A288F6D00B71CDE0BA5193" unit="page">
<mods:start id="1FD75C2B2D47C81F95032C84B2044F7A">123</mods:start>
<mods:end id="E49DD2DF92B2DA6A5C0611877911C323">138</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="842A8278B16419D392B18B5C9A414A98">
<mods:url id="B451D6632642E1AC4BE25A3D18BF1417">http://dx.doi.org/10.1093/jmammal/gyaa104</mods:url>
</mods:location>
<mods:classification id="580A60B7FC5F88E8A12AB140A1873C7D">journal article</mods:classification>
<mods:identifier id="73361DDAED5FA1761732357C7ECBD34F" type="DOI">10.1093/jmammal/gyaa104</mods:identifier>
<mods:identifier id="1ECE094BE78F9227C61272370AF43785" type="ISSN">0022-2372</mods:identifier>
<mods:identifier id="1B817D23631815D272BDEC6320D2096E" type="Zenodo-Dep">10632393</mods:identifier>
</mods:mods>
<treatment id="CE0F8E0D4F2E8D5DFC1EF91CFA154768" LSID="urn:lsid:plazi:treatment:CE0F8E0D4F2E8D5DFC1EF91CFA154768" httpUri="http://treatment.plazi.org/id/CE0F8E0D4F2E8D5DFC1EF91CFA154768" lastPageId="12" lastPageNumber="135" pageId="3" pageNumber="126">
<subSubSection id="0EBC6C904F2E8D52FC1EF91CFAD644FE" box="[964,1343,1727,1751]" pageId="3" pageNumber="126" type="nomenclature">
<paragraph id="46193F1B4F2E8D52FC1EF91CFAD644FE" blockId="3.[964,1343,1727,1751]" box="[964,1343,1727,1751]" pageId="3" pageNumber="126">
<emphasis id="74D2E3094F2E8D52FC1EF91CFB0E44FE" box="[964,1255,1727,1751]" italics="true" pageId="3" pageNumber="126">
<taxonomicName id="81A644984F2E8D52FC1EF91CFB0944FF" authority="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson, 2021" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[964,1248,1727,1751]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="126" phylum="Chordata" rank="species" species="septentrionalis" status="sp. nov.">
<emphasis id="74D2E3094F2E8D52FC1EF91CFB0944FF" bold="true" box="[964,1248,1727,1751]" italics="true" pageId="3" pageNumber="126">Rhagomys septentrionalis</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="6FE15E724F2E8D52FB37F91CFAD644FE" box="[1261,1343,1727,1750]" pageId="3" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0EBC6C904F2E8D52FBCFF94CFB07452E" box="[1045,1262,1775,1798]" pageId="3" pageNumber="126" type="vernacular_names">
<paragraph id="46193F1B4F2E8D52FBCFF94CFB07452E" blockId="3.[1045,1262,1775,1798]" box="[1045,1262,1775,1798]" pageId="3" pageNumber="126">
<heading id="1D5188774F2E8D52FBCFF94CFB07452E" box="[1045,1262,1775,1798]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="126" reason="8">
Northern
<taxonomicName id="81A644984F2E8D52FBA6F94CFB07452E" ID-CoL="78BH" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1148,1262,1775,1798]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="126" phylum="Chordata" rank="genus">Rhagomys</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0EBC6C904F2E8D52FBFAF8BCFB0A451E" box="[1056,1251,1823,1846]" pageId="3" pageNumber="126" type="description">
<paragraph id="46193F1B4F2E8D52FBFAF8BCFB0A451E" blockId="3.[1056,1251,1823,1846]" box="[1056,1251,1823,1846]" pageId="3" pageNumber="126">
<figureCitation id="DE9D239E4F2E8D52FBFAF8BCFB6C451E" box="[1056,1157,1823,1846]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="7.[101,136,1365,1386]" captionStartId-1="8.[144,179,1875,1896]" captionStartId-2="9.[101,136,1893,1914]" captionStartId-3="9.[819,854,1923,1944]" captionTargetBox-0="[114,1475,200,1352]" captionTargetBox-1="[168,789,1240,1861]" captionTargetBox-2="[126,747,1216,1880]" captionTargetBox-3="[843,1464,1082,1910]" captionTargetId-0="figure-6@7.[114,1475,200,1352]" captionTargetId-1="figure-889@8.[168,789,1240,1861]" captionTargetId-2="figure-6@9.[126,747,1216,1880]" captionTargetId-3="figure-54@9.[843,1464,1082,1910]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="9" captionText-0="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." captionText-1="Fig. 4.—Basicranium of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype): PPP, posterolateral palatal pits; PPF, parapterygoid fossa; TC, transverse canal; MLF, middle lacerate foramen; BOC, basioccipital. Scale bar = 5 mm." captionText-2="Fig. 5.—Lateral view of the mandibles of Rhagomys: (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype); (B) R. rufescens (MZUSP AB401; from Pavan and Leite 2011); and (C) R. longilingua (MUSM 20009). Scale bar = 10 mm." captionText-3="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10632401" figureDoi-1="http://doi.org/10.5281/zenodo.10632403" figureDoi-2="http://doi.org/10.5281/zenodo.10632405" figureDoi-3="http://doi.org/10.5281/zenodo.10632407" httpUri-0="https://zenodo.org/record/10632401/files/figure.png" httpUri-1="https://zenodo.org/record/10632403/files/figure.png" httpUri-2="https://zenodo.org/record/10632405/files/figure.png" httpUri-3="https://zenodo.org/record/10632407/files/figure.png" pageId="3" pageNumber="126">Figs. 36</figureCitation>
;
<tableCitation id="0B240AA04F2E8D52FB4EF8BCFB0A451E" box="[1172,1251,1823,1846]" captionStart="Table 1" captionStartId="4.[126,182,506,527]" captionTargetPageId="4" captionText="Table 1.—External, vibrissal, and craniodental measurements (mm), and age class, for specimens of Rhagomys. Acronyms are: Natural History Museum, London (BMNH); Colección Boliviana de Fauna, La Paz (CBF); Field Museum of Natural History, Chicago (FMNH); Laboratory of Vertebrates, Universidade Federal de Ouro Preto (LZV-UFOP) with the accession number R; Instituto de Ciencias Biológicas, Escuela Politécnica Nacional, Quito (MEPN); Museu Nacional, Universidade Federal do Rio de Janeiro (MN); Museu de Zoologia, Universidade de Sao Paulo (MZUSP). The tabulated measurements were previously published, except for the MEPN holotype. Sex: Female (F), Male (M), unknown (?)." pageId="3" pageNumber="126">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="0EBC6C904F2E8D57FC97F8E4FED743E8" lastPageId="6" lastPageNumber="129" pageId="3" pageNumber="126" type="materials_examined">
<paragraph id="46193F1B4F2E8D57FC97F8E4FD594308" blockId="3.[819,1489,1863,1982]" lastBlockId="6.[144,816,201,1984]" lastPageId="6" lastPageNumber="129" pageId="3" pageNumber="126">
<emphasis id="74D2E3094F2E8D52FC97F8E4FC5F4576" box="[845,950,1863,1886]" italics="true" pageId="3" pageNumber="126">
<typeStatus id="991D81B94F2E8D52FC97F8E4FC5B4576" box="[845,946,1863,1886]" pageId="3" pageNumber="126" type="holotype">Holotype</typeStatus>
.
</emphasis>
—An adult specimen, with all molars fully erupted at occlusal level, age class 2, sex unknown, prepared as dry skin, cranium and skeleton in good condition and deposited in the Mammalogy collection of the Departamento de Ciencias
<materialsCitation id="F6CE35464F2B8D57FF4AFF6AFD454308" collectingDate="2008-11-06" collectionCode="MEPN" collectorName="Biologicas de la Escuela Politecnica Nacional &amp; Ramiro Barriga" country="Ecuador" location="Biologicas de la Escuela Politecnica Nacional" pageId="6" pageNumber="129" specimenCode="MEPN 10898" specimenCount="1" typeStatus="holotype">
<collectorName id="EB535ACD4F2B8D57FF4AFF6AFD7042C8" box="[144,665,201,224]" pageId="6" pageNumber="129">
<location id="437969C04F2B8D57FF4AFF6AFD7042C8" LSID="urn:lsid:plazi:treatment:CE0F8E0D4F2E8D5DFC1EF91CFA154768:437969C04F2B8D57FF4AFF6AFD7042C8" box="[144,665,201,224]" country="Ecuador" name="Biologicas de la Escuela Politecnica Nacional" pageId="6" pageNumber="129">Biológicas de la Escuela Politécnica Nacional</location>
</collectorName>
del
<collectingCountry id="3EB17F8B4F2B8D57FD0EFF6AFCC442C8" box="[724,813,201,224]" name="Ecuador" pageId="6" pageNumber="129">Ecuador</collectingCountry>
(
<specimenCode id="160097604F2B8D57FF4DFF4AFED94328" box="[151,304,233,256]" collectionCode="MEPN" pageId="6" pageNumber="129">MEPN 10898</specimenCode>
). The
<typeStatus id="991D81B94F2B8D57FEB5FF4AFE254328" box="[367,460,233,256]" pageId="6" pageNumber="129" type="holotype">holotype</typeStatus>
was collected by
<collectorName id="EB535ACD4F2B8D57FD52FF4AFCC44328" box="[648,813,233,256]" pageId="6" pageNumber="129">Ramiro Barriga</collectorName>
on
<date id="321819DB4F2B8D57FF6BFEAAFE984308" box="[177,369,265,288]" pageId="6" pageNumber="129" value="2008-11-06">
<collectingDate id="225CE0334F2B8D57FF6BFEAAFE984308" box="[177,369,265,288]" pageId="6" pageNumber="129" value="2008-11-06">6 November 2008</collectingDate>
</date>
and prepared by Luis Albuja
</materialsCitation>
.
</paragraph>
<caption id="12D96F934F298D55FFA4FE59F9DF404F" pageId="4" pageNumber="127" startId="4.[126,182,506,527]" targetBox="[101,1900,642,1609]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="46193F1B4F298D55FFA4FE59F9DF404F" blockId="4.[102,1899,506,615]" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FFA4FE59FF0E4027" bold="true" box="[126,231,506,527]" pageId="4" pageNumber="127">Table 1.—</emphasis>
External, vibrissal, and craniodental measurements (mm), and age class, for specimens of
<taxonomicName id="81A644984F298D55FB8BFE59FB5C4027" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1105,1205,506,527]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="127" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F298D55FB8BFE59FB5C4027" box="[1105,1205,506,527]" italics="true" pageId="4" pageNumber="127">Rhagomys</emphasis>
</taxonomicName>
. Acronyms are: Natural History Museum, London (BMNH); Colección Boliviana de Fauna, La Paz (CBF); Field Museum of Natural History, Chicago (FMNH); Laboratory of Vertebrates, Universidade Federal de Ouro Preto (LZV-UFOP) with the accession number R; Instituto de Ciencias Biológicas, Escuela Politécnica Nacional, Quito (MEPN); Museu Nacional, Universidade Federal do Rio de Janeiro (MN); Museu de Zoologia, Universidade de Sao Paulo (MZUSP). The tabulated measurements were previously published, except for the MEPN holotype. Sex: Female (F), Male (M), unknown (?).
</paragraph>
</caption>
<paragraph id="46193F1B4F298D55FE78FD20F8AE4461" pageId="4" pageNumber="127">
<table id="34A6CDBB4F2972AEFFBFFD21F8854461" box="[101,1900,642,1609]" gridcols="16" gridrows="38" pageId="4" pageNumber="127">
<tr id="F8963D594F2972AEFFBFFD21F88540BE" box="[101,1900,642,662]" gridrow="0" pageId="4" pageNumber="127" rowspan-0="1" rowspan-1="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="BB4754254F2972AEFE78FD21FD8140BE" box="[418,616,642,662]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="4" pageNumber="127">
<taxonomicName id="81A644984F298D55FE78FD20FD8B40BE" authorityName="Luna and Patterson" authorityYear="2003" box="[418,610,643,662]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="127" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F298D55FE78FD20FD8B40BE" box="[418,610,643,662]" italics="true" pageId="4" pageNumber="127">Rhagomys longilingua</emphasis>
</taxonomicName>
a
</th>
<th id="BB4754254F2972AEFB1BFD21FA9F40BE" box="[1217,1398,642,662]" colspan="2" colspanRight="1" gridcol="10" gridrow="0" pageId="4" pageNumber="127">
<taxonomicName id="81A644984F298D55FB1BFD20FA9940BE" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1217,1392,643,662]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="127" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F298D55FB1BFD20FA9940BE" box="[1217,1392,643,662]" italics="true" pageId="4" pageNumber="127">Rhagomys rufescens</emphasis>
</taxonomicName>
b
</th>
<th id="BB4754254F2972AEF92FFD21F88540BE" box="[1781,1900,642,662]" gridcol="15" gridrow="0" pageId="4" pageNumber="127">
<taxonomicName id="81A644984F298D55F92FFD20F8A740BE" authority="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson, 2021" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1781,1870,643,662]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="127" phylum="Chordata" rank="genus" status="sp. nov.">
<emphasis id="74D2E3094F298D55F92FFD20F8A740BE" box="[1781,1870,643,662]" italics="true" pageId="4" pageNumber="127">Rhagomys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FE15E724F298D55F889FD20F88540BE" box="[1875,1900,643,662]" pageId="4" pageNumber="127" rank="species">sp.</taxonomicNameLabel>
</th>
</tr>
<tr id="F8963D594F2972AEFFBFFD3DF8854099" box="[101,1900,670,689]" gridrow="1" pageId="4" pageNumber="127" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td id="BB4754254F2972AEF92FFD3DF8854099" box="[1781,1900,670,689]" gridcol="15" gridrow="1" pageId="4" pageNumber="127">
<taxonomicNameLabel id="6FE15E724F298D55F8C4FD3DF8AA4099" box="[1822,1859,670,689]" pageId="4" pageNumber="127" rank="species">nov.</taxonomicNameLabel>
</td>
</tr>
<tr id="F8963D594F2972AEFFBFFD61F88540FD" box="[101,1900,706,725]" gridrow="2" pageId="4" pageNumber="127" rowspan-0="1">
<td id="BB4754254F2972AEFF3AFD61FEF940FD" box="[224,272,706,725]" gridcol="1" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FF3CFD61FEE740FD" box="[230,270,706,725]" italics="true" pageId="4" pageNumber="127">CBF</emphasis>
</td>
<td id="BB4754254F2972AEFEE2FD61FE6C40FD" box="[312,389,706,725]" gridcol="2" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FE9CFD61FE6B40FD" box="[326,386,706,725]" italics="true" pageId="4" pageNumber="127">FMNH</emphasis>
</td>
<td id="BB4754254F2972AEFE78FD61FE1840FD" box="[418,497,706,725]" gridcol="3" gridrow="2" pageId="4" pageNumber="127">
<taxonomicName id="81A644984F298D55FE62FD61FE1840FD" box="[440,497,706,725]" class="Liliopsida" family="Musaceae" genus="Musa" kingdom="Plantae" order="Zingiberales" pageId="4" pageNumber="127" phylum="Tracheophyta" rank="genus">
<emphasis id="74D2E3094F298D55FE62FD61FE1840FD" box="[440,497,706,725]" italics="true" pageId="4" pageNumber="127">MUSA</emphasis>
</taxonomicName>
</td>
<td id="BB4754254F2972AEFDFBFD61FD8140FD" box="[545,616,706,725]" gridcol="4" gridrow="2" pageId="4" pageNumber="127">
<taxonomicName id="81A644984F298D55FDF9FD61FDB540FD" box="[547,604,706,725]" class="Liliopsida" family="Musaceae" genus="Musa" kingdom="Plantae" order="Zingiberales" pageId="4" pageNumber="127" phylum="Tracheophyta" rank="genus">
<emphasis id="74D2E3094F298D55FDF9FD61FDB540FD" box="[547,604,706,725]" italics="true" pageId="4" pageNumber="127">MUSA</emphasis>
</taxonomicName>
</td>
<td id="BB4754254F2972AEFD55FD61FD2140FD" box="[655,712,706,725]" gridcol="5" gridrow="2" pageId="4" pageNumber="127">
<taxonomicName id="81A644984F298D55FD55FD61FD2140FD" box="[655,712,706,725]" class="Liliopsida" family="Musaceae" genus="Musa" kingdom="Plantae" order="Zingiberales" pageId="4" pageNumber="127" phylum="Tracheophyta" rank="genus">
<emphasis id="74D2E3094F298D55FD55FD61FD2140FD" box="[655,712,706,725]" italics="true" pageId="4" pageNumber="127">MUSA</emphasis>
</taxonomicName>
</td>
<td id="BB4754254F2972AEFD24FD61FCD540FD" box="[766,828,706,725]" gridcol="6" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FD24FD61FCD540FD" box="[766,828,706,725]" italics="true" pageId="4" pageNumber="127">MUSM</emphasis>
</td>
<td id="BB4754254F2972AEFCBBFD61FC4540FD" box="[865,940,706,725]" gridcol="7" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FCBFFD61FC4840FD" box="[869,929,706,725]" italics="true" pageId="4" pageNumber="127">BMNH</emphasis>
</td>
<td id="BB4754254F2972AEFC03FD61FBFE40FD" box="[985,1047,706,725]" gridcol="8" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FC3EFD61FBED40FD" box="[996,1028,706,725]" italics="true" pageId="4" pageNumber="127">MN</emphasis>
</td>
<td id="BB4754254F2972AEFB9BFD61FB6F40FD" box="[1089,1158,706,725]" gridcol="9" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FB9BFD61FB6F40FD" box="[1089,1158,706,725]" italics="true" pageId="4" pageNumber="127">MZUSP</emphasis>
</td>
<td id="BB4754254F2972AEFB1BFD61FB1140FD" box="[1217,1272,706,725]" gridcol="10" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FB11FD61FB1140FD" box="[1227,1272,706,725]" italics="true" pageId="4" pageNumber="127">163R</emphasis>
</td>
<td id="BB4754254F2972AEFAF6FD61FA9F40FD" box="[1324,1398,706,725]" gridcol="11" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FAEBFD61FA8A40FD" box="[1329,1379,706,725]" italics="true" pageId="4" pageNumber="127">213 R</emphasis>
</td>
<td id="BB4754254F2972AEFA4DFD61FA2140FD" box="[1431,1480,706,725]" gridcol="12" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FA41FD61FA2140FD" box="[1435,1480,706,725]" italics="true" pageId="4" pageNumber="127">214R</emphasis>
</td>
<td id="BB4754254F2972AEFA28FD61F9C540FD" box="[1522,1580,706,725]" gridcol="13" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FA25FD61F9C540FD" box="[1535,1580,706,725]" italics="true" pageId="4" pageNumber="127">215R</emphasis>
</td>
<td id="BB4754254F2972AEF9BBFD61F93540FD" box="[1633,1756,706,725]" gridcol="14" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55F9BBFD61F93540FD" box="[1633,1756,706,725]" italics="true" pageId="4" pageNumber="127">CMUFLA 905</emphasis>
</td>
<td id="BB4754254F2972AEF92FFD61F88540FD" box="[1781,1900,706,725]" gridcol="15" gridrow="2" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55F92CFD61F88340FD" box="[1782,1898,706,725]" italics="true" pageId="4" pageNumber="127">MEPN 10898</emphasis>
</td>
</tr>
<tr id="F8963D594F2972AEFFBFFD7EF88540D1" box="[101,1900,733,761]" gridrow="3" pageId="4" pageNumber="127" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1">
<td id="BB4754254F2972AEFF3AFD7EFEF940D1" box="[224,272,733,761]" gridcol="1" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FF3EFD45FEE640D1" box="[228,271,742,761]" italics="true" pageId="4" pageNumber="127">7620</emphasis>
</td>
<td id="BB4754254F2972AEFEE2FD7EFE6C40D1" box="[312,389,733,761]" gridcol="2" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FE9FFD45FE6C40D1" box="[325,389,742,761]" italics="true" pageId="4" pageNumber="127">170687</emphasis>
</td>
<td id="BB4754254F2972AEFE78FD7EFE1840D1" box="[418,497,733,761]" gridcol="3" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FE63FD7EFE0740D8" box="[441,494,733,752]" italics="true" pageId="4" pageNumber="127">13453</emphasis>
</td>
<td id="BB4754254F2972AEFDFBFD7EFD8140D1" box="[545,616,733,761]" gridcol="4" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FDFFFD7EFDB340D8" box="[549,602,733,752]" italics="true" pageId="4" pageNumber="127">13856</emphasis>
</td>
<td id="BB4754254F2972AEFD55FD7EFD2140D1" box="[655,712,733,761]" gridcol="5" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FD4BFD7EFD2F40D8" box="[657,710,733,752]" italics="true" pageId="4" pageNumber="127">15895</emphasis>
</td>
<td id="BB4754254F2972AEFD24FD7EFCD540D1" box="[766,828,733,761]" gridcol="6" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FCD8FD45FCDE40D1" box="[770,823,742,761]" italics="true" pageId="4" pageNumber="127">20009</emphasis>
</td>
<td id="BB4754254F2972AEFCBBFD7EFC4540D1" box="[865,940,733,761]" gridcol="7" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FCBBFD45FC4F40D1" box="[865,934,742,761]" italics="true" pageId="4" pageNumber="127">86.2.8.5</emphasis>
</td>
<td id="BB4754254F2972AEFC03FD7EFBFE40D1" box="[985,1047,733,761]" gridcol="8" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FC03FD45FBE740D1" box="[985,1038,742,761]" italics="true" pageId="4" pageNumber="127">66056</emphasis>
</td>
<td id="BB4754254F2972AEFB9BFD7EFB6F40D1" box="[1089,1158,733,761]" gridcol="9" gridrow="3" pageId="4" pageNumber="127">
<emphasis id="74D2E3094F298D55FB93FD45FB9740D1" box="[1097,1150,742,761]" italics="true" pageId="4" pageNumber="127">31952</emphasis>
</td>
</tr>
<tr id="F8963D594F2972AEFFBFFCA9F8854135" box="[101,1900,778,797]" gridrow="4" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFCA9FF7C4135" box="[101,149,778,797]" gridcol="0" gridrow="4" pageId="4" pageNumber="127">Sex</th>
<td id="BB4754254F2972AEFF3AFCA9FEF94135" box="[224,272,778,797]" gridcol="1" gridrow="4" pageId="4" pageNumber="127">F</td>
<td id="BB4754254F2972AEFEE2FCA9FE6C4135" box="[312,389,778,797]" gridcol="2" gridrow="4" pageId="4" pageNumber="127">F</td>
<td id="BB4754254F2972AEFE78FCA9FE184135" box="[418,497,778,797]" gridcol="3" gridrow="4" pageId="4" pageNumber="127">M</td>
<td id="BB4754254F2972AEFDFBFCA9FD814135" box="[545,616,778,797]" gridcol="4" gridrow="4" pageId="4" pageNumber="127">M</td>
<td id="BB4754254F2972AEFD55FCA9FD214135" box="[655,712,778,797]" gridcol="5" gridrow="4" pageId="4" pageNumber="127">M</td>
<td id="BB4754254F2972AEFD24FCA9FCD54135" box="[766,828,778,797]" gridcol="6" gridrow="4" pageId="4" pageNumber="127">M</td>
<td id="BB4754254F2972AEFCBBFCA9FC454135" box="[865,940,778,797]" gridcol="7" gridrow="4" pageId="4" pageNumber="127">F</td>
<td id="BB4754254F2972AEFC03FCA9FBFE4135" box="[985,1047,778,797]" gridcol="8" gridrow="4" pageId="4" pageNumber="127">F</td>
<td id="BB4754254F2972AEFB9BFCA9FB6F4135" box="[1089,1158,778,797]" gridcol="9" gridrow="4" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFCA9FB114135" box="[1217,1272,778,797]" gridcol="10" gridrow="4" pageId="4" pageNumber="127">F</td>
<td id="BB4754254F2972AEFAF6FCA9FA9F4135" box="[1324,1398,778,797]" gridcol="11" gridrow="4" pageId="4" pageNumber="127">M</td>
<td id="BB4754254F2972AEFA4DFCA9FA214135" box="[1431,1480,778,797]" gridcol="12" gridrow="4" pageId="4" pageNumber="127">F</td>
<td id="BB4754254F2972AEFA28FCA9F9C54135" box="[1522,1580,778,797]" gridcol="13" gridrow="4" pageId="4" pageNumber="127">M</td>
<td id="BB4754254F2972AEF9BBFCA9F9354135" box="[1633,1756,778,797]" gridcol="14" gridrow="4" pageId="4" pageNumber="127">M</td>
<td id="BB4754254F2972AEF92FFCA9F8854135" box="[1781,1900,778,797]" gridcol="15" gridrow="4" pageId="4" pageNumber="127">?</td>
</tr>
<tr id="F8963D594F2972AEFFBFFC81F885411D" box="[101,1900,802,821]" gridrow="5" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFC81FF7C411D" box="[101,149,802,821]" gridcol="0" gridrow="5" pageId="4" pageNumber="127">Age</th>
<td id="BB4754254F2972AEFF3AFC81FEF9411D" box="[224,272,802,821]" gridcol="1" gridrow="5" pageId="4" pageNumber="127">1</td>
<td id="BB4754254F2972AEFEE2FC81FE6C411D" box="[312,389,802,821]" gridcol="2" gridrow="5" pageId="4" pageNumber="127">2</td>
<td id="BB4754254F2972AEFE78FC81FE18411D" box="[418,497,802,821]" gridcol="3" gridrow="5" pageId="4" pageNumber="127">4</td>
<td id="BB4754254F2972AEFDFBFC81FD81411D" box="[545,616,802,821]" gridcol="4" gridrow="5" pageId="4" pageNumber="127">1</td>
<td id="BB4754254F2972AEFD55FC81FD21411D" box="[655,712,802,821]" gridcol="5" gridrow="5" pageId="4" pageNumber="127">4</td>
<td id="BB4754254F2972AEFD24FC81FCD5411D" box="[766,828,802,821]" gridcol="6" gridrow="5" pageId="4" pageNumber="127">2</td>
<td id="BB4754254F2972AEFCBBFC81FC45411D" box="[865,940,802,821]" gridcol="7" gridrow="5" pageId="4" pageNumber="127">3</td>
<td id="BB4754254F2972AEFC03FC81FBFE411D" box="[985,1047,802,821]" gridcol="8" gridrow="5" pageId="4" pageNumber="127">2</td>
<td id="BB4754254F2972AEFB9BFC81FB6F411D" box="[1089,1158,802,821]" gridcol="9" gridrow="5" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFC81FB11411D" box="[1217,1272,802,821]" gridcol="10" gridrow="5" pageId="4" pageNumber="127">1</td>
<td id="BB4754254F2972AEFAF6FC81FA9F411D" box="[1324,1398,802,821]" gridcol="11" gridrow="5" pageId="4" pageNumber="127">2</td>
<td id="BB4754254F2972AEFA4DFC81FA21411D" box="[1431,1480,802,821]" gridcol="12" gridrow="5" pageId="4" pageNumber="127">2</td>
<td id="BB4754254F2972AEFA28FC81F9C5411D" box="[1522,1580,802,821]" gridcol="13" gridrow="5" pageId="4" pageNumber="127">4</td>
<td id="BB4754254F2972AEF9BBFC81F935411D" box="[1633,1756,802,821]" gridcol="14" gridrow="5" pageId="4" pageNumber="127">2</td>
<td id="BB4754254F2972AEF92FFC81F885411D" box="[1781,1900,802,821]" gridcol="15" gridrow="5" pageId="4" pageNumber="127">2</td>
</tr>
<tr id="F8963D594F2972AEFFBFFC98F8854166" box="[101,1900,827,846]" gridrow="6" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFC98FF7C4166" box="[101,149,827,846]" gridcol="0" gridrow="6" pageId="4" pageNumber="127">HBL</th>
<td id="BB4754254F2972AEFF3AFC98FEF94166" box="[224,272,827,846]" gridcol="1" gridrow="6" pageId="4" pageNumber="127">80</td>
<td id="BB4754254F2972AEFEE2FC98FE6C4166" box="[312,389,827,846]" gridcol="2" gridrow="6" pageId="4" pageNumber="127">103</td>
<td id="BB4754254F2972AEFE78FC98FE184166" box="[418,497,827,846]" gridcol="3" gridrow="6" pageId="4" pageNumber="127">89</td>
<td id="BB4754254F2972AEFDFBFC98FD814166" box="[545,616,827,846]" gridcol="4" gridrow="6" pageId="4" pageNumber="127">52</td>
<td id="BB4754254F2972AEFD55FC98FD214166" box="[655,712,827,846]" gridcol="5" gridrow="6" pageId="4" pageNumber="127">92</td>
<td id="BB4754254F2972AEFD24FC98FCD54166" box="[766,828,827,846]" gridcol="6" gridrow="6" pageId="4" pageNumber="127">90</td>
<td id="BB4754254F2972AEFCBBFC98FC454166" box="[865,940,827,846]" gridcol="7" gridrow="6" pageId="4" pageNumber="127">94</td>
<td id="BB4754254F2972AEFC03FC98FBFE4166" box="[985,1047,827,846]" gridcol="8" gridrow="6" pageId="4" pageNumber="127">75</td>
<td id="BB4754254F2972AEFB9BFC98FB6F4166" box="[1089,1158,827,846]" gridcol="9" gridrow="6" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFC98FB114166" box="[1217,1272,827,846]" gridcol="10" gridrow="6" pageId="4" pageNumber="127">85</td>
<td id="BB4754254F2972AEFAF6FC98FA9F4166" box="[1324,1398,827,846]" gridcol="11" gridrow="6" pageId="4" pageNumber="127">83</td>
<td id="BB4754254F2972AEFA4DFC98FA214166" box="[1431,1480,827,846]" gridcol="12" gridrow="6" pageId="4" pageNumber="127">82</td>
<td id="BB4754254F2972AEFA28FC98F9C54166" box="[1522,1580,827,846]" gridcol="13" gridrow="6" pageId="4" pageNumber="127">98</td>
<td id="BB4754254F2972AEF9BBFC98F9354166" box="[1633,1756,827,846]" gridcol="14" gridrow="6" pageId="4" pageNumber="127">65</td>
<td id="BB4754254F2972AEF92FFC98F8854166" box="[1781,1900,827,846]" gridcol="15" gridrow="6" pageId="4" pageNumber="127">90</td>
</tr>
<tr id="F8963D594F2972AEFFBFFCF7F885414F" box="[101,1900,852,871]" gridrow="7" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFCF7FF7C414F" box="[101,149,852,871]" gridcol="0" gridrow="7" pageId="4" pageNumber="127">TL</th>
<td id="BB4754254F2972AEFF3AFCF7FEF9414F" box="[224,272,852,871]" gridcol="1" gridrow="7" pageId="4" pageNumber="127">75</td>
<td id="BB4754254F2972AEFEE2FCF7FE6C414F" box="[312,389,852,871]" gridcol="2" gridrow="7" pageId="4" pageNumber="127">104</td>
<td id="BB4754254F2972AEFE78FCF7FE18414F" box="[418,497,852,871]" gridcol="3" gridrow="7" pageId="4" pageNumber="127">101</td>
<td id="BB4754254F2972AEFDFBFCF7FD81414F" box="[545,616,852,871]" gridcol="4" gridrow="7" pageId="4" pageNumber="127">87</td>
<td id="BB4754254F2972AEFD55FCF7FD21414F" box="[655,712,852,871]" gridcol="5" gridrow="7" pageId="4" pageNumber="127">97</td>
<td id="BB4754254F2972AEFD24FCF7FCD5414F" box="[766,828,852,871]" gridcol="6" gridrow="7" pageId="4" pageNumber="127">93</td>
<td id="BB4754254F2972AEFCBBFCF7FC45414F" box="[865,940,852,871]" gridcol="7" gridrow="7" pageId="4" pageNumber="127">93</td>
<td id="BB4754254F2972AEFC03FCF7FBFE414F" box="[985,1047,852,871]" gridcol="8" gridrow="7" pageId="4" pageNumber="127">94</td>
<td id="BB4754254F2972AEFB9BFCF7FB6F414F" box="[1089,1158,852,871]" gridcol="9" gridrow="7" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFCF7FB11414F" box="[1217,1272,852,871]" gridcol="10" gridrow="7" pageId="4" pageNumber="127">85</td>
<td id="BB4754254F2972AEFAF6FCF7FA9F414F" box="[1324,1398,852,871]" gridcol="11" gridrow="7" pageId="4" pageNumber="127">86</td>
<td id="BB4754254F2972AEFA4DFCF7FA21414F" box="[1431,1480,852,871]" gridcol="12" gridrow="7" pageId="4" pageNumber="127">88</td>
<td id="BB4754254F2972AEFA28FCF7F9C5414F" box="[1522,1580,852,871]" gridcol="13" gridrow="7" pageId="4" pageNumber="127">107</td>
<td id="BB4754254F2972AEF9BBFCF7F935414F" box="[1633,1756,852,871]" gridcol="14" gridrow="7" pageId="4" pageNumber="127">105</td>
<td id="BB4754254F2972AEF92FFCF7F885414F" box="[1781,1900,852,871]" gridcol="15" gridrow="7" pageId="4" pageNumber="127">100</td>
</tr>
<tr id="F8963D594F2972AEFFBFFCCEF88541A8" box="[101,1900,877,896]" gridrow="8" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFCCEFF7C41A8" box="[101,149,877,896]" gridcol="0" gridrow="8" pageId="4" pageNumber="127">LHF</th>
<td id="BB4754254F2972AEFF3AFCCEFEF941A8" box="[224,272,877,896]" gridcol="1" gridrow="8" pageId="4" pageNumber="127">13</td>
<td id="BB4754254F2972AEFEE2FCCEFE6C41A8" box="[312,389,877,896]" gridcol="2" gridrow="8" pageId="4" pageNumber="127">20</td>
<td id="BB4754254F2972AEFE78FCCEFE1841A8" box="[418,497,877,896]" gridcol="3" gridrow="8" pageId="4" pageNumber="127">18.1</td>
<td id="BB4754254F2972AEFDFBFCCEFD8141A8" box="[545,616,877,896]" gridcol="4" gridrow="8" pageId="4" pageNumber="127">16.1</td>
<td id="BB4754254F2972AEFD55FCCEFD2141A8" box="[655,712,877,896]" gridcol="5" gridrow="8" pageId="4" pageNumber="127">17.6</td>
<td id="BB4754254F2972AEFD24FCCEFCD541A8" box="[766,828,877,896]" gridcol="6" gridrow="8" pageId="4" pageNumber="127">17</td>
<td id="BB4754254F2972AEFCBBFCCEFC4541A8" box="[865,940,877,896]" gridcol="7" gridrow="8" pageId="4" pageNumber="127">20</td>
<td id="BB4754254F2972AEFC03FCCEFBFE41A8" box="[985,1047,877,896]" gridcol="8" gridrow="8" pageId="4" pageNumber="127">19</td>
<td id="BB4754254F2972AEFB9BFCCEFB6F41A8" box="[1089,1158,877,896]" gridcol="9" gridrow="8" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFCCEFB1141A8" box="[1217,1272,877,896]" gridcol="10" gridrow="8" pageId="4" pageNumber="127">18</td>
<td id="BB4754254F2972AEFAF6FCCEFA9F41A8" box="[1324,1398,877,896]" gridcol="11" gridrow="8" pageId="4" pageNumber="127">20</td>
<td id="BB4754254F2972AEFA4DFCCEFA2141A8" box="[1431,1480,877,896]" gridcol="12" gridrow="8" pageId="4" pageNumber="127">19</td>
<td id="BB4754254F2972AEFA28FCCEF9C541A8" box="[1522,1580,877,896]" gridcol="13" gridrow="8" pageId="4" pageNumber="127">22</td>
<td id="BB4754254F2972AEF9BBFCCEF93541A8" box="[1633,1756,877,896]" gridcol="14" gridrow="8" pageId="4" pageNumber="127">17</td>
<td id="BB4754254F2972AEF92FFCCEF88541A8" box="[1781,1900,877,896]" gridcol="15" gridrow="8" pageId="4" pageNumber="127">17</td>
</tr>
<tr id="F8963D594F2972AEFFBFFC25F88541B1" box="[101,1900,902,921]" gridrow="9" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFC25FF7C41B1" box="[101,149,902,921]" gridcol="0" gridrow="9" pageId="4" pageNumber="127">EL</th>
<td id="BB4754254F2972AEFF3AFC25FEF941B1" box="[224,272,902,921]" gridcol="1" gridrow="9" pageId="4" pageNumber="127">9</td>
<td id="BB4754254F2972AEFEE2FC25FE6C41B1" box="[312,389,902,921]" gridcol="2" gridrow="9" pageId="4" pageNumber="127">14</td>
<td id="BB4754254F2972AEFE78FC25FE1841B1" box="[418,497,902,921]" gridcol="3" gridrow="9" pageId="4" pageNumber="127">12.1</td>
<td id="BB4754254F2972AEFDFBFC25FD8141B1" box="[545,616,902,921]" gridcol="4" gridrow="9" pageId="4" pageNumber="127">10</td>
<td id="BB4754254F2972AEFD55FC25FD2141B1" box="[655,712,902,921]" gridcol="5" gridrow="9" pageId="4" pageNumber="127">13.1</td>
<td id="BB4754254F2972AEFD24FC25FCD541B1" box="[766,828,902,921]" gridcol="6" gridrow="9" pageId="4" pageNumber="127">13.6</td>
<td id="BB4754254F2972AEFCBBFC25FC4541B1" box="[865,940,902,921]" gridcol="7" gridrow="9" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03FC25FBFE41B1" box="[985,1047,902,921]" gridcol="8" gridrow="9" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFC25FB6F41B1" box="[1089,1158,902,921]" gridcol="9" gridrow="9" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFC25FB1141B1" box="[1217,1272,902,921]" gridcol="10" gridrow="9" pageId="4" pageNumber="127">10</td>
<td id="BB4754254F2972AEFAF6FC25FA9F41B1" box="[1324,1398,902,921]" gridcol="11" gridrow="9" pageId="4" pageNumber="127">13</td>
<td id="BB4754254F2972AEFA4DFC25FA2141B1" box="[1431,1480,902,921]" gridcol="12" gridrow="9" pageId="4" pageNumber="127">13</td>
<td id="BB4754254F2972AEFA28FC25F9C541B1" box="[1522,1580,902,921]" gridcol="13" gridrow="9" pageId="4" pageNumber="127">15</td>
<td id="BB4754254F2972AEF9BBFC25F93541B1" box="[1633,1756,902,921]" gridcol="14" gridrow="9" pageId="4" pageNumber="127">12</td>
<td id="BB4754254F2972AEF92FFC25F88541B1" box="[1781,1900,902,921]" gridcol="15" gridrow="9" pageId="4" pageNumber="127">11</td>
</tr>
<tr id="F8963D594F2972AEFFBFFC3DF8854199" box="[101,1900,926,945]" gridrow="10" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFC3DFF7C4199" box="[101,149,926,945]" gridcol="0" gridrow="10" pageId="4" pageNumber="127">DFL</th>
<td id="BB4754254F2972AEFF3AFC3DFEF94199" box="[224,272,926,945]" gridcol="1" gridrow="10" pageId="4" pageNumber="127">7</td>
<td id="BB4754254F2972AEFEE2FC3DFE6C4199" box="[312,389,926,945]" gridcol="2" gridrow="10" pageId="4" pageNumber="127">8</td>
<td id="BB4754254F2972AEFE78FC3DFE184199" box="[418,497,926,945]" gridcol="3" gridrow="10" pageId="4" pageNumber="127">9.2</td>
<td id="BB4754254F2972AEFDFBFC3DFD814199" box="[545,616,926,945]" gridcol="4" gridrow="10" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FC3DFD214199" box="[655,712,926,945]" gridcol="5" gridrow="10" pageId="4" pageNumber="127">9</td>
<td id="BB4754254F2972AEFD24FC3DFCD54199" box="[766,828,926,945]" gridcol="6" gridrow="10" pageId="4" pageNumber="127">8</td>
<td id="BB4754254F2972AEFCBBFC3DFC454199" box="[865,940,926,945]" gridcol="7" gridrow="10" pageId="4" pageNumber="127">7</td>
<td id="BB4754254F2972AEFC03FC3DFBFE4199" box="[985,1047,926,945]" gridcol="8" gridrow="10" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFC3DFB6F4199" box="[1089,1158,926,945]" gridcol="9" gridrow="10" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFC3DFB114199" box="[1217,1272,926,945]" gridcol="10" gridrow="10" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FC3DFA9F4199" box="[1324,1398,926,945]" gridcol="11" gridrow="10" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFC3DFA214199" box="[1431,1480,926,945]" gridcol="12" gridrow="10" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FC3DF9C54199" box="[1522,1580,926,945]" gridcol="13" gridrow="10" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFC3DF9354199" box="[1633,1756,926,945]" gridcol="14" gridrow="10" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFC3DF8854199" box="[1781,1900,926,945]" gridcol="15" gridrow="10" pageId="4" pageNumber="127">9</td>
</tr>
<tr id="F8963D594F2972AEFFBFFC14F88541E2" box="[101,1900,951,970]" gridrow="11" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFC14FF7C41E2" box="[101,149,951,970]" gridcol="0" gridrow="11" pageId="4" pageNumber="127">LMV</th>
<td id="BB4754254F2972AEFF3AFC14FEF941E2" box="[224,272,951,970]" gridcol="1" gridrow="11" pageId="4" pageNumber="127">29</td>
<td id="BB4754254F2972AEFEE2FC14FE6C41E2" box="[312,389,951,970]" gridcol="2" gridrow="11" pageId="4" pageNumber="127">37</td>
<td id="BB4754254F2972AEFE78FC14FE1841E2" box="[418,497,951,970]" gridcol="3" gridrow="11" pageId="4" pageNumber="127">36</td>
<td id="BB4754254F2972AEFDFBFC14FD8141E2" box="[545,616,951,970]" gridcol="4" gridrow="11" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FC14FD2141E2" box="[655,712,951,970]" gridcol="5" gridrow="11" pageId="4" pageNumber="127">33.5</td>
<td id="BB4754254F2972AEFD24FC14FCD541E2" box="[766,828,951,970]" gridcol="6" gridrow="11" pageId="4" pageNumber="127">35</td>
<td id="BB4754254F2972AEFCBBFC14FC4541E2" box="[865,940,951,970]" gridcol="7" gridrow="11" pageId="4" pageNumber="127">34</td>
<td id="BB4754254F2972AEFC03FC14FBFE41E2" box="[985,1047,951,970]" gridcol="8" gridrow="11" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFC14FB6F41E2" box="[1089,1158,951,970]" gridcol="9" gridrow="11" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFC14FB1141E2" box="[1217,1272,951,970]" gridcol="10" gridrow="11" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FC14FA9F41E2" box="[1324,1398,951,970]" gridcol="11" gridrow="11" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFC14FA2141E2" box="[1431,1480,951,970]" gridcol="12" gridrow="11" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FC14F9C541E2" box="[1522,1580,951,970]" gridcol="13" gridrow="11" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFC14F93541E2" box="[1633,1756,951,970]" gridcol="14" gridrow="11" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFC14F88541E2" box="[1781,1900,951,970]" gridcol="15" gridrow="11" pageId="4" pageNumber="127">36</td>
</tr>
<tr id="F8963D594F2972AEFFBFFC6CF88541CA" box="[101,1900,975,994]" gridrow="12" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFC6CFF7C41CA" box="[101,149,975,994]" gridcol="0" gridrow="12" pageId="4" pageNumber="127">LSV</th>
<td id="BB4754254F2972AEFF3AFC6CFEF941CA" box="[224,272,975,994]" gridcol="1" gridrow="12" pageId="4" pageNumber="127">19</td>
<td id="BB4754254F2972AEFEE2FC6CFE6C41CA" box="[312,389,975,994]" gridcol="2" gridrow="12" pageId="4" pageNumber="127">26</td>
<td id="BB4754254F2972AEFE78FC6CFE1841CA" box="[418,497,975,994]" gridcol="3" gridrow="12" pageId="4" pageNumber="127">24</td>
<td id="BB4754254F2972AEFDFBFC6CFD8141CA" box="[545,616,975,994]" gridcol="4" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FC6CFD2141CA" box="[655,712,975,994]" gridcol="5" gridrow="12" pageId="4" pageNumber="127">17.5</td>
<td id="BB4754254F2972AEFD24FC6CFCD541CA" box="[766,828,975,994]" gridcol="6" gridrow="12" pageId="4" pageNumber="127">25</td>
<td id="BB4754254F2972AEFCBBFC6CFC4541CA" box="[865,940,975,994]" gridcol="7" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03FC6CFBFE41CA" box="[985,1047,975,994]" gridcol="8" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFC6CFB6F41CA" box="[1089,1158,975,994]" gridcol="9" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFC6CFB1141CA" box="[1217,1272,975,994]" gridcol="10" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FC6CFA9F41CA" box="[1324,1398,975,994]" gridcol="11" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFC6CFA2141CA" box="[1431,1480,975,994]" gridcol="12" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FC6CF9C541CA" box="[1522,1580,975,994]" gridcol="13" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFC6CF93541CA" box="[1633,1756,975,994]" gridcol="14" gridrow="12" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFC6CF88541CA" box="[1781,1900,975,994]" gridcol="15" gridrow="12" pageId="4" pageNumber="127">?</td>
</tr>
<tr id="F8963D594F2972AEFFBFFC4BF88541D3" box="[101,1900,1000,1019]" gridrow="13" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFC4BFF7C41D3" box="[101,149,1000,1019]" gridcol="0" gridrow="13" pageId="4" pageNumber="127">LGV</th>
<td id="BB4754254F2972AEFF3AFC4BFEF941D3" box="[224,272,1000,1019]" gridcol="1" gridrow="13" pageId="4" pageNumber="127">14</td>
<td id="BB4754254F2972AEFEE2FC4BFE6C41D3" box="[312,389,1000,1019]" gridcol="2" gridrow="13" pageId="4" pageNumber="127">21.5</td>
<td id="BB4754254F2972AEFE78FC4BFE1841D3" box="[418,497,1000,1019]" gridcol="3" gridrow="13" pageId="4" pageNumber="127">20</td>
<td id="BB4754254F2972AEFDFBFC4BFD8141D3" box="[545,616,1000,1019]" gridcol="4" gridrow="13" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FC4BFD2141D3" box="[655,712,1000,1019]" gridcol="5" gridrow="13" pageId="4" pageNumber="127">17.3</td>
<td id="BB4754254F2972AEFD24FC4BFCD541D3" box="[766,828,1000,1019]" gridcol="6" gridrow="13" pageId="4" pageNumber="127">20</td>
<td id="BB4754254F2972AEFCBBFC4BFC4541D3" box="[865,940,1000,1019]" gridcol="7" gridrow="13" pageId="4" pageNumber="127">17</td>
<td id="BB4754254F2972AEFC03FC4BFBFE41D3" box="[985,1047,1000,1019]" gridcol="8" gridrow="13" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFC4BFB6F41D3" box="[1089,1158,1000,1019]" gridcol="9" gridrow="13" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFC4BFB1141D3" box="[1217,1272,1000,1019]" gridcol="10" gridrow="13" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FC4BFA9F41D3" box="[1324,1398,1000,1019]" gridcol="11" gridrow="13" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFC4BFA2141D3" box="[1431,1480,1000,1019]" gridcol="12" gridrow="13" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FC4BF9C541D3" box="[1522,1580,1000,1019]" gridcol="13" gridrow="13" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFC4BF93541D3" box="[1633,1756,1000,1019]" gridcol="14" gridrow="13" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFC4BF88541D3" box="[1781,1900,1000,1019]" gridcol="15" gridrow="13" pageId="4" pageNumber="127">18</td>
</tr>
<tr id="F8963D594F2972AEFFBFFBA3F885463B" box="[101,1900,1024,1043]" gridrow="14" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFBA3FF7C463B" box="[101,149,1024,1043]" gridcol="0" gridrow="14" pageId="4" pageNumber="127">CL</th>
<td id="BB4754254F2972AEFF3AFBA3FEF9463B" box="[224,272,1024,1043]" gridcol="1" gridrow="14" pageId="4" pageNumber="127">25.34</td>
<td id="BB4754254F2972AEFEE2FBA3FE6C463B" box="[312,389,1024,1043]" gridcol="2" gridrow="14" pageId="4" pageNumber="127">28.33</td>
<td id="BB4754254F2972AEFE78FBA3FE18463B" box="[418,497,1024,1043]" gridcol="3" gridrow="14" pageId="4" pageNumber="127">28.58</td>
<td id="BB4754254F2972AEFDFBFBA3FD81463B" box="[545,616,1024,1043]" gridcol="4" gridrow="14" pageId="4" pageNumber="127">22.31</td>
<td id="BB4754254F2972AEFD55FBA3FD21463B" box="[655,712,1024,1043]" gridcol="5" gridrow="14" pageId="4" pageNumber="127">27.72</td>
<td id="BB4754254F2972AEFD24FBA3FCD5463B" box="[766,828,1024,1043]" gridcol="6" gridrow="14" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFCBBFBA3FC45463B" box="[865,940,1024,1043]" gridcol="7" gridrow="14" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03FBA3FBFE463B" box="[985,1047,1024,1043]" gridcol="8" gridrow="14" pageId="4" pageNumber="127">25.26</td>
<td id="BB4754254F2972AEFB9BFBA3FB6F463B" box="[1089,1158,1024,1043]" gridcol="9" gridrow="14" pageId="4" pageNumber="127">26.24</td>
<td id="BB4754254F2972AEFB1BFBA3FB11463B" box="[1217,1272,1024,1043]" gridcol="10" gridrow="14" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FBA3FA9F463B" box="[1324,1398,1024,1043]" gridcol="11" gridrow="14" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFBA3FA21463B" box="[1431,1480,1024,1043]" gridcol="12" gridrow="14" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FBA3F9C5463B" box="[1522,1580,1024,1043]" gridcol="13" gridrow="14" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFBA3F935463B" box="[1633,1756,1024,1043]" gridcol="14" gridrow="14" pageId="4" pageNumber="127">26.41</td>
<td id="BB4754254F2972AEF92FFBA3F885463B" box="[1781,1900,1024,1043]" gridcol="15" gridrow="14" pageId="4" pageNumber="127">26.44</td>
</tr>
<tr id="F8963D594F2972AEFFBFFBBAF8854604" box="[101,1900,1049,1068]" gridrow="15" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFBBAFF7C4604" box="[101,149,1049,1068]" gridcol="0" gridrow="15" pageId="4" pageNumber="127">CIL</th>
<td id="BB4754254F2972AEFF3AFBBAFEF94604" box="[224,272,1049,1068]" gridcol="1" gridrow="15" pageId="4" pageNumber="127">22.99</td>
<td id="BB4754254F2972AEFEE2FBBAFE6C4604" box="[312,389,1049,1068]" gridcol="2" gridrow="15" pageId="4" pageNumber="127">26.01</td>
<td id="BB4754254F2972AEFE78FBBAFE184604" box="[418,497,1049,1068]" gridcol="3" gridrow="15" pageId="4" pageNumber="127">25.95</td>
<td id="BB4754254F2972AEFDFBFBBAFD814604" box="[545,616,1049,1068]" gridcol="4" gridrow="15" pageId="4" pageNumber="127">19.97</td>
<td id="BB4754254F2972AEFD55FBBAFD214604" box="[655,712,1049,1068]" gridcol="5" gridrow="15" pageId="4" pageNumber="127">25.66</td>
<td id="BB4754254F2972AEFD24FBBAFCD54604" box="[766,828,1049,1068]" gridcol="6" gridrow="15" pageId="4" pageNumber="127">25.02</td>
<td id="BB4754254F2972AEFCBBFBBAFC454604" box="[865,940,1049,1068]" gridcol="7" gridrow="15" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03FBBAFBFE4604" box="[985,1047,1049,1068]" gridcol="8" gridrow="15" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFBBAFB6F4604" box="[1089,1158,1049,1068]" gridcol="9" gridrow="15" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFBBAFB114604" box="[1217,1272,1049,1068]" gridcol="10" gridrow="15" pageId="4" pageNumber="127">24.25</td>
<td id="BB4754254F2972AEFAF6FBBAFA9F4604" box="[1324,1398,1049,1068]" gridcol="11" gridrow="15" pageId="4" pageNumber="127">24.85</td>
<td id="BB4754254F2972AEFA4DFBBAFA214604" box="[1431,1480,1049,1068]" gridcol="12" gridrow="15" pageId="4" pageNumber="127">24.89</td>
<td id="BB4754254F2972AEFA28FBBAF9C54604" box="[1522,1580,1049,1068]" gridcol="13" gridrow="15" pageId="4" pageNumber="127">27.89</td>
<td id="BB4754254F2972AEF9BBFBBAF9354604" box="[1633,1756,1049,1068]" gridcol="14" gridrow="15" pageId="4" pageNumber="127">22.2</td>
<td id="BB4754254F2972AEF92FFBBAF8854604" box="[1781,1900,1049,1068]" gridcol="15" gridrow="15" pageId="4" pageNumber="127">23.58</td>
</tr>
<tr id="F8963D594F2972AEFFBFFB91F885466D" box="[101,1900,1074,1093]" gridrow="16" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFB91FF7C466D" box="[101,149,1074,1093]" gridcol="0" gridrow="16" pageId="4" pageNumber="127">CML</th>
<td id="BB4754254F2972AEFF3AFB91FEF9466D" box="[224,272,1074,1093]" gridcol="1" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFEE2FB91FE6C466D" box="[312,389,1074,1093]" gridcol="2" gridrow="16" pageId="4" pageNumber="127">16.68</td>
<td id="BB4754254F2972AEFE78FB91FE18466D" box="[418,497,1074,1093]" gridcol="3" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFDFBFB91FD81466D" box="[545,616,1074,1093]" gridcol="4" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FB91FD21466D" box="[655,712,1074,1093]" gridcol="5" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD24FB91FCD5466D" box="[766,828,1074,1093]" gridcol="6" gridrow="16" pageId="4" pageNumber="127">15.91</td>
<td id="BB4754254F2972AEFCBBFB91FC45466D" box="[865,940,1074,1093]" gridcol="7" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03FB91FBFE466D" box="[985,1047,1074,1093]" gridcol="8" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFB91FB6F466D" box="[1089,1158,1074,1093]" gridcol="9" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFB91FB11466D" box="[1217,1272,1074,1093]" gridcol="10" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FB91FA9F466D" box="[1324,1398,1074,1093]" gridcol="11" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFB91FA21466D" box="[1431,1480,1074,1093]" gridcol="12" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FB91F9C5466D" box="[1522,1580,1074,1093]" gridcol="13" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFB91F935466D" box="[1633,1756,1074,1093]" gridcol="14" gridrow="16" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFB91F885466D" box="[1781,1900,1074,1093]" gridcol="15" gridrow="16" pageId="4" pageNumber="127">15.5</td>
</tr>
<tr id="F8963D594F2972AEFFBFFBE9F8854675" box="[101,1900,1098,1117]" gridrow="17" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFBE9FF7C4675" box="[101,149,1098,1117]" gridcol="0" gridrow="17" pageId="4" pageNumber="127">RL</th>
<td id="BB4754254F2972AEFF3AFBE9FEF94675" box="[224,272,1098,1117]" gridcol="1" gridrow="17" pageId="4" pageNumber="127">6.92</td>
<td id="BB4754254F2972AEFEE2FBE9FE6C4675" box="[312,389,1098,1117]" gridcol="2" gridrow="17" pageId="4" pageNumber="127">7.7</td>
<td id="BB4754254F2972AEFE78FBE9FE184675" box="[418,497,1098,1117]" gridcol="3" gridrow="17" pageId="4" pageNumber="127">7.66</td>
<td id="BB4754254F2972AEFDFBFBE9FD814675" box="[545,616,1098,1117]" gridcol="4" gridrow="17" pageId="4" pageNumber="127">5.24</td>
<td id="BB4754254F2972AEFD55FBE9FD214675" box="[655,712,1098,1117]" gridcol="5" gridrow="17" pageId="4" pageNumber="127">6.94</td>
<td id="BB4754254F2972AEFD24FBE9FCD54675" box="[766,828,1098,1117]" gridcol="6" gridrow="17" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFCBBFBE9FC454675" box="[865,940,1098,1117]" gridcol="7" gridrow="17" pageId="4" pageNumber="127">8.61</td>
<td id="BB4754254F2972AEFC03FBE9FBFE4675" box="[985,1047,1098,1117]" gridcol="8" gridrow="17" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFBE9FB6F4675" box="[1089,1158,1098,1117]" gridcol="9" gridrow="17" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFBE9FB114675" box="[1217,1272,1098,1117]" gridcol="10" gridrow="17" pageId="4" pageNumber="127">8.35</td>
<td id="BB4754254F2972AEFAF6FBE9FA9F4675" box="[1324,1398,1098,1117]" gridcol="11" gridrow="17" pageId="4" pageNumber="127">8.54</td>
<td id="BB4754254F2972AEFA4DFBE9FA214675" box="[1431,1480,1098,1117]" gridcol="12" gridrow="17" pageId="4" pageNumber="127">8.38</td>
<td id="BB4754254F2972AEFA28FBE9F9C54675" box="[1522,1580,1098,1117]" gridcol="13" gridrow="17" pageId="4" pageNumber="127">10.14</td>
<td id="BB4754254F2972AEF9BBFBE9F9354675" box="[1633,1756,1098,1117]" gridcol="14" gridrow="17" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFBE9F8854675" box="[1781,1900,1098,1117]" gridcol="15" gridrow="17" pageId="4" pageNumber="127">7.4</td>
</tr>
<tr id="F8963D594F2972AEFFBFFBC0F885465E" box="[101,1900,1123,1142]" gridrow="18" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFBC0FF7C465E" box="[101,149,1123,1142]" gridcol="0" gridrow="18" pageId="4" pageNumber="127">RW</th>
<td id="BB4754254F2972AEFF3AFBC0FEF9465E" box="[224,272,1123,1142]" gridcol="1" gridrow="18" pageId="4" pageNumber="127">4.44</td>
<td id="BB4754254F2972AEFEE2FBC0FE6C465E" box="[312,389,1123,1142]" gridcol="2" gridrow="18" pageId="4" pageNumber="127">5.06</td>
<td id="BB4754254F2972AEFE78FBC0FE18465E" box="[418,497,1123,1142]" gridcol="3" gridrow="18" pageId="4" pageNumber="127">5.03</td>
<td id="BB4754254F2972AEFDFBFBC0FD81465E" box="[545,616,1123,1142]" gridcol="4" gridrow="18" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FBC0FD21465E" box="[655,712,1123,1142]" gridcol="5" gridrow="18" pageId="4" pageNumber="127">5.14</td>
<td id="BB4754254F2972AEFD24FBC0FCD5465E" box="[766,828,1123,1142]" gridcol="6" gridrow="18" pageId="4" pageNumber="127">4.99</td>
<td id="BB4754254F2972AEFCBBFBC0FC45465E" box="[865,940,1123,1142]" gridcol="7" gridrow="18" pageId="4" pageNumber="127">4.48</td>
<td id="BB4754254F2972AEFC03FBC0FBFE465E" box="[985,1047,1123,1142]" gridcol="8" gridrow="18" pageId="4" pageNumber="127">4.15</td>
<td id="BB4754254F2972AEFB9BFBC0FB6F465E" box="[1089,1158,1123,1142]" gridcol="9" gridrow="18" pageId="4" pageNumber="127">4.36</td>
<td id="BB4754254F2972AEFB1BFBC0FB11465E" box="[1217,1272,1123,1142]" gridcol="10" gridrow="18" pageId="4" pageNumber="127">3.65</td>
<td id="BB4754254F2972AEFAF6FBC0FA9F465E" box="[1324,1398,1123,1142]" gridcol="11" gridrow="18" pageId="4" pageNumber="127">4.45</td>
<td id="BB4754254F2972AEFA4DFBC0FA21465E" box="[1431,1480,1123,1142]" gridcol="12" gridrow="18" pageId="4" pageNumber="127">3.68</td>
<td id="BB4754254F2972AEFA28FBC0F9C5465E" box="[1522,1580,1123,1142]" gridcol="13" gridrow="18" pageId="4" pageNumber="127">4.11</td>
<td id="BB4754254F2972AEF9BBFBC0F935465E" box="[1633,1756,1123,1142]" gridcol="14" gridrow="18" pageId="4" pageNumber="127">3.91</td>
<td id="BB4754254F2972AEF92FFBC0F885465E" box="[1781,1900,1123,1142]" gridcol="15" gridrow="18" pageId="4" pageNumber="127">4.55</td>
</tr>
<tr id="F8963D594F2972AEFFBFFBD8F88546A6" box="[101,1900,1147,1166]" gridrow="19" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFBD8FF7C46A6" box="[101,149,1147,1166]" gridcol="0" gridrow="19" pageId="4" pageNumber="127">LOF</th>
<td id="BB4754254F2972AEFF3AFBD8FEF946A6" box="[224,272,1147,1166]" gridcol="1" gridrow="19" pageId="4" pageNumber="127">9.69</td>
<td id="BB4754254F2972AEFEE2FBD8FE6C46A6" box="[312,389,1147,1166]" gridcol="2" gridrow="19" pageId="4" pageNumber="127">10.72</td>
<td id="BB4754254F2972AEFE78FBD8FE1846A6" box="[418,497,1147,1166]" gridcol="3" gridrow="19" pageId="4" pageNumber="127">10.5</td>
<td id="BB4754254F2972AEFDFBFBD8FD8146A6" box="[545,616,1147,1166]" gridcol="4" gridrow="19" pageId="4" pageNumber="127">8.29</td>
<td id="BB4754254F2972AEFD55FBD8FD2146A6" box="[655,712,1147,1166]" gridcol="5" gridrow="19" pageId="4" pageNumber="127">10.52</td>
<td id="BB4754254F2972AEFD24FBD8FCD546A6" box="[766,828,1147,1166]" gridcol="6" gridrow="19" pageId="4" pageNumber="127">10.5</td>
<td id="BB4754254F2972AEFCBBFBD8FC4546A6" box="[865,940,1147,1166]" gridcol="7" gridrow="19" pageId="4" pageNumber="127">10.65</td>
<td id="BB4754254F2972AEFC03FBD8FBFE46A6" box="[985,1047,1147,1166]" gridcol="8" gridrow="19" pageId="4" pageNumber="127">9.72</td>
<td id="BB4754254F2972AEFB9BFBD8FB6F46A6" box="[1089,1158,1147,1166]" gridcol="9" gridrow="19" pageId="4" pageNumber="127">9.86</td>
<td id="BB4754254F2972AEFB1BFBD8FB1146A6" box="[1217,1272,1147,1166]" gridcol="10" gridrow="19" pageId="4" pageNumber="127">9.34</td>
<td id="BB4754254F2972AEFAF6FBD8FA9F46A6" box="[1324,1398,1147,1166]" gridcol="11" gridrow="19" pageId="4" pageNumber="127">9.89</td>
<td id="BB4754254F2972AEFA4DFBD8FA2146A6" box="[1431,1480,1147,1166]" gridcol="12" gridrow="19" pageId="4" pageNumber="127">9.92</td>
<td id="BB4754254F2972AEFA28FBD8F9C546A6" box="[1522,1580,1147,1166]" gridcol="13" gridrow="19" pageId="4" pageNumber="127">10.49</td>
<td id="BB4754254F2972AEF9BBFBD8F93546A6" box="[1633,1756,1147,1166]" gridcol="14" gridrow="19" pageId="4" pageNumber="127">10.12</td>
<td id="BB4754254F2972AEF92FFBD8F88546A6" box="[1781,1900,1147,1166]" gridcol="15" gridrow="19" pageId="4" pageNumber="127">9.35</td>
</tr>
<tr id="F8963D594F2972AEFFBFFB37F885468F" box="[101,1900,1172,1191]" gridrow="20" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFB37FF7C468F" box="[101,149,1172,1191]" gridcol="0" gridrow="20" pageId="4" pageNumber="127">NL</th>
<td id="BB4754254F2972AEFF3AFB37FEF9468F" box="[224,272,1172,1191]" gridcol="1" gridrow="20" pageId="4" pageNumber="127">7.8</td>
<td id="BB4754254F2972AEFEE2FB37FE6C468F" box="[312,389,1172,1191]" gridcol="2" gridrow="20" pageId="4" pageNumber="127">8.55</td>
<td id="BB4754254F2972AEFE78FB37FE18468F" box="[418,497,1172,1191]" gridcol="3" gridrow="20" pageId="4" pageNumber="127">8.51</td>
<td id="BB4754254F2972AEFDFBFB37FD81468F" box="[545,616,1172,1191]" gridcol="4" gridrow="20" pageId="4" pageNumber="127">5.91</td>
<td id="BB4754254F2972AEFD55FB37FD21468F" box="[655,712,1172,1191]" gridcol="5" gridrow="20" pageId="4" pageNumber="127">8.84</td>
<td id="BB4754254F2972AEFD24FB37FCD5468F" box="[766,828,1172,1191]" gridcol="6" gridrow="20" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFCBBFB37FC45468F" box="[865,940,1172,1191]" gridcol="7" gridrow="20" pageId="4" pageNumber="127">8.69</td>
<td id="BB4754254F2972AEFC03FB37FBFE468F" box="[985,1047,1172,1191]" gridcol="8" gridrow="20" pageId="4" pageNumber="127">7.72</td>
<td id="BB4754254F2972AEFB9BFB37FB6F468F" box="[1089,1158,1172,1191]" gridcol="9" gridrow="20" pageId="4" pageNumber="127">8.3</td>
<td id="BB4754254F2972AEFB1BFB37FB11468F" box="[1217,1272,1172,1191]" gridcol="10" gridrow="20" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FB37FA9F468F" box="[1324,1398,1172,1191]" gridcol="11" gridrow="20" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFB37FA21468F" box="[1431,1480,1172,1191]" gridcol="12" gridrow="20" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FB37F9C5468F" box="[1522,1580,1172,1191]" gridcol="13" gridrow="20" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFB37F935468F" box="[1633,1756,1172,1191]" gridcol="14" gridrow="20" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFB37F885468F" box="[1781,1900,1172,1191]" gridcol="15" gridrow="20" pageId="4" pageNumber="127">8.26</td>
</tr>
<tr id="F8963D594F2972AEFFBFFB0FF8854697" box="[101,1900,1196,1215]" gridrow="21" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFB0FFF7C4697" box="[101,149,1196,1215]" gridcol="0" gridrow="21" pageId="4" pageNumber="127">NW</th>
<td id="BB4754254F2972AEFF3AFB0FFEF94697" box="[224,272,1196,1215]" gridcol="1" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFEE2FB0FFE6C4697" box="[312,389,1196,1215]" gridcol="2" gridrow="21" pageId="4" pageNumber="127">2.95</td>
<td id="BB4754254F2972AEFE78FB0FFE184697" box="[418,497,1196,1215]" gridcol="3" gridrow="21" pageId="4" pageNumber="127">2.91</td>
<td id="BB4754254F2972AEFDFBFB0FFD814697" box="[545,616,1196,1215]" gridcol="4" gridrow="21" pageId="4" pageNumber="127">2.46</td>
<td id="BB4754254F2972AEFD55FB0FFD214697" box="[655,712,1196,1215]" gridcol="5" gridrow="21" pageId="4" pageNumber="127">2.73</td>
<td id="BB4754254F2972AEFD24FB0FFCD54697" box="[766,828,1196,1215]" gridcol="6" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFCBBFB0FFC454697" box="[865,940,1196,1215]" gridcol="7" gridrow="21" pageId="4" pageNumber="127">3.09</td>
<td id="BB4754254F2972AEFC03FB0FFBFE4697" box="[985,1047,1196,1215]" gridcol="8" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFB0FFB6F4697" box="[1089,1158,1196,1215]" gridcol="9" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFB0FFB114697" box="[1217,1272,1196,1215]" gridcol="10" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FB0FFA9F4697" box="[1324,1398,1196,1215]" gridcol="11" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFB0FFA214697" box="[1431,1480,1196,1215]" gridcol="12" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FB0FF9C54697" box="[1522,1580,1196,1215]" gridcol="13" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFB0FF9354697" box="[1633,1756,1196,1215]" gridcol="14" gridrow="21" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFB0FF8854697" box="[1781,1900,1196,1215]" gridcol="15" gridrow="21" pageId="4" pageNumber="127">?</td>
</tr>
<tr id="F8963D594F2972AEFFBFFB66F88546F0" box="[101,1900,1221,1240]" gridrow="22" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFB66FF7C46F0" box="[101,149,1221,1240]" gridcol="0" gridrow="22" pageId="4" pageNumber="127">LIB</th>
<td id="BB4754254F2972AEFF3AFB66FEF946F0" box="[224,272,1221,1240]" gridcol="1" gridrow="22" pageId="4" pageNumber="127">5.3</td>
<td id="BB4754254F2972AEFEE2FB66FE6C46F0" box="[312,389,1221,1240]" gridcol="2" gridrow="22" pageId="4" pageNumber="127">5.78</td>
<td id="BB4754254F2972AEFE78FB66FE1846F0" box="[418,497,1221,1240]" gridcol="3" gridrow="22" pageId="4" pageNumber="127">5.3</td>
<td id="BB4754254F2972AEFDFBFB66FD8146F0" box="[545,616,1221,1240]" gridcol="4" gridrow="22" pageId="4" pageNumber="127">4.98</td>
<td id="BB4754254F2972AEFD55FB66FD2146F0" box="[655,712,1221,1240]" gridcol="5" gridrow="22" pageId="4" pageNumber="127">5.45</td>
<td id="BB4754254F2972AEFD24FB66FCD546F0" box="[766,828,1221,1240]" gridcol="6" gridrow="22" pageId="4" pageNumber="127">5.5</td>
<td id="BB4754254F2972AEFCBBFB66FC4546F0" box="[865,940,1221,1240]" gridcol="7" gridrow="22" pageId="4" pageNumber="127">5.48</td>
<td id="BB4754254F2972AEFC03FB66FBFE46F0" box="[985,1047,1221,1240]" gridcol="8" gridrow="22" pageId="4" pageNumber="127">4.98</td>
<td id="BB4754254F2972AEFB9BFB66FB6F46F0" box="[1089,1158,1221,1240]" gridcol="9" gridrow="22" pageId="4" pageNumber="127">4.98</td>
<td id="BB4754254F2972AEFB1BFB66FB1146F0" box="[1217,1272,1221,1240]" gridcol="10" gridrow="22" pageId="4" pageNumber="127">4.86</td>
<td id="BB4754254F2972AEFAF6FB66FA9F46F0" box="[1324,1398,1221,1240]" gridcol="11" gridrow="22" pageId="4" pageNumber="127">4.97</td>
<td id="BB4754254F2972AEFA4DFB66FA2146F0" box="[1431,1480,1221,1240]" gridcol="12" gridrow="22" pageId="4" pageNumber="127">5</td>
<td id="BB4754254F2972AEFA28FB66F9C546F0" box="[1522,1580,1221,1240]" gridcol="13" gridrow="22" pageId="4" pageNumber="127">5.06</td>
<td id="BB4754254F2972AEF9BBFB66F93546F0" box="[1633,1756,1221,1240]" gridcol="14" gridrow="22" pageId="4" pageNumber="127">4.92</td>
<td id="BB4754254F2972AEF92FFB66F88546F0" box="[1781,1900,1221,1240]" gridcol="15" gridrow="22" pageId="4" pageNumber="127">5.21</td>
</tr>
<tr id="F8963D594F2972AEFFBFFB7DF88546D9" box="[101,1900,1246,1265]" gridrow="23" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFB7DFF7C46D9" box="[101,149,1246,1265]" gridcol="0" gridrow="23" pageId="4" pageNumber="127">DL</th>
<td id="BB4754254F2972AEFF3AFB7DFEF946D9" box="[224,272,1246,1265]" gridcol="1" gridrow="23" pageId="4" pageNumber="127">6.14</td>
<td id="BB4754254F2972AEFEE2FB7DFE6C46D9" box="[312,389,1246,1265]" gridcol="2" gridrow="23" pageId="4" pageNumber="127">7.33</td>
<td id="BB4754254F2972AEFE78FB7DFE1846D9" box="[418,497,1246,1265]" gridcol="3" gridrow="23" pageId="4" pageNumber="127">7.5</td>
<td id="BB4754254F2972AEFDFBFB7DFD8146D9" box="[545,616,1246,1265]" gridcol="4" gridrow="23" pageId="4" pageNumber="127">5.66</td>
<td id="BB4754254F2972AEFD55FB7DFD2146D9" box="[655,712,1246,1265]" gridcol="5" gridrow="23" pageId="4" pageNumber="127">7.7</td>
<td id="BB4754254F2972AEFD24FB7DFCD546D9" box="[766,828,1246,1265]" gridcol="6" gridrow="23" pageId="4" pageNumber="127">7.24</td>
<td id="BB4754254F2972AEFCBBFB7DFC4546D9" box="[865,940,1246,1265]" gridcol="7" gridrow="23" pageId="4" pageNumber="127">7.98</td>
<td id="BB4754254F2972AEFC03FB7DFBFE46D9" box="[985,1047,1246,1265]" gridcol="8" gridrow="23" pageId="4" pageNumber="127">6.04</td>
<td id="BB4754254F2972AEFB9BFB7DFB6F46D9" box="[1089,1158,1246,1265]" gridcol="9" gridrow="23" pageId="4" pageNumber="127">7.02</td>
<td id="BB4754254F2972AEFB1BFB7DFB1146D9" box="[1217,1272,1246,1265]" gridcol="10" gridrow="23" pageId="4" pageNumber="127">6.18</td>
<td id="BB4754254F2972AEFAF6FB7DFA9F46D9" box="[1324,1398,1246,1265]" gridcol="11" gridrow="23" pageId="4" pageNumber="127">6.49</td>
<td id="BB4754254F2972AEFA4DFB7DFA2146D9" box="[1431,1480,1246,1265]" gridcol="12" gridrow="23" pageId="4" pageNumber="127">6.5</td>
<td id="BB4754254F2972AEFA28FB7DF9C546D9" box="[1522,1580,1246,1265]" gridcol="13" gridrow="23" pageId="4" pageNumber="127">7.79</td>
<td id="BB4754254F2972AEF9BBFB7DF93546D9" box="[1633,1756,1246,1265]" gridcol="14" gridrow="23" pageId="4" pageNumber="127">8.21</td>
<td id="BB4754254F2972AEF92FFB7DF88546D9" box="[1781,1900,1246,1265]" gridcol="15" gridrow="23" pageId="4" pageNumber="127">6.52</td>
</tr>
<tr id="F8963D594F2972AEFFBFFB55F8854721" box="[101,1900,1270,1289]" gridrow="24" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFB55FF7C4721" box="[101,149,1270,1289]" gridcol="0" gridrow="24" pageId="4" pageNumber="127">LBP</th>
<td id="BB4754254F2972AEFF3AFB55FEF94721" box="[224,272,1270,1289]" gridcol="1" gridrow="24" pageId="4" pageNumber="127">6.09</td>
<td id="BB4754254F2972AEFEE2FB55FE6C4721" box="[312,389,1270,1289]" gridcol="2" gridrow="24" pageId="4" pageNumber="127">6.86</td>
<td id="BB4754254F2972AEFE78FB55FE184721" box="[418,497,1270,1289]" gridcol="3" gridrow="24" pageId="4" pageNumber="127">6.56</td>
<td id="BB4754254F2972AEFDFBFB55FD814721" box="[545,616,1270,1289]" gridcol="4" gridrow="24" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FB55FD214721" box="[655,712,1270,1289]" gridcol="5" gridrow="24" pageId="4" pageNumber="127">6.05</td>
<td id="BB4754254F2972AEFD24FB55FCD54721" box="[766,828,1270,1289]" gridcol="6" gridrow="24" pageId="4" pageNumber="127">6.65</td>
<td id="BB4754254F2972AEFCBBFB55FC454721" box="[865,940,1270,1289]" gridcol="7" gridrow="24" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03FB55FBFE4721" box="[985,1047,1270,1289]" gridcol="8" gridrow="24" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFB55FB6F4721" box="[1089,1158,1270,1289]" gridcol="9" gridrow="24" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFB55FB114721" box="[1217,1272,1270,1289]" gridcol="10" gridrow="24" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FB55FA9F4721" box="[1324,1398,1270,1289]" gridcol="11" gridrow="24" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFB55FA214721" box="[1431,1480,1270,1289]" gridcol="12" gridrow="24" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FB55F9C54721" box="[1522,1580,1270,1289]" gridcol="13" gridrow="24" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFB55F9354721" box="[1633,1756,1270,1289]" gridcol="14" gridrow="24" pageId="4" pageNumber="127">11.59</td>
<td id="BB4754254F2972AEF92FFB55F8854721" box="[1781,1900,1270,1289]" gridcol="15" gridrow="24" pageId="4" pageNumber="127">5.57</td>
</tr>
<tr id="F8963D594F2972AEFFBFFAACF885470A" box="[101,1900,1295,1314]" gridrow="25" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFAACFF7C470A" box="[101,149,1295,1314]" gridcol="0" gridrow="25" pageId="4" pageNumber="127">BPB</th>
<td id="BB4754254F2972AEFF3AFAACFEF9470A" box="[224,272,1295,1314]" gridcol="1" gridrow="25" pageId="4" pageNumber="127">4.82</td>
<td id="BB4754254F2972AEFEE2FAACFE6C470A" box="[312,389,1295,1314]" gridcol="2" gridrow="25" pageId="4" pageNumber="127">5.22</td>
<td id="BB4754254F2972AEFE78FAACFE18470A" box="[418,497,1295,1314]" gridcol="3" gridrow="25" pageId="4" pageNumber="127">4.99</td>
<td id="BB4754254F2972AEFDFBFAACFD81470A" box="[545,616,1295,1314]" gridcol="4" gridrow="25" pageId="4" pageNumber="127">4.59</td>
<td id="BB4754254F2972AEFD55FAACFD21470A" box="[655,712,1295,1314]" gridcol="5" gridrow="25" pageId="4" pageNumber="127">5.09</td>
<td id="BB4754254F2972AEFD24FAACFCD5470A" box="[766,828,1295,1314]" gridcol="6" gridrow="25" pageId="4" pageNumber="127">4.97</td>
<td id="BB4754254F2972AEFCBBFAACFC45470A" box="[865,940,1295,1314]" gridcol="7" gridrow="25" pageId="4" pageNumber="127">5.29</td>
<td id="BB4754254F2972AEFC03FAACFBFE470A" box="[985,1047,1295,1314]" gridcol="8" gridrow="25" pageId="4" pageNumber="127">4.99</td>
<td id="BB4754254F2972AEFB9BFAACFB6F470A" box="[1089,1158,1295,1314]" gridcol="9" gridrow="25" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFAACFB11470A" box="[1217,1272,1295,1314]" gridcol="10" gridrow="25" pageId="4" pageNumber="127">4.81</td>
<td id="BB4754254F2972AEFAF6FAACFA9F470A" box="[1324,1398,1295,1314]" gridcol="11" gridrow="25" pageId="4" pageNumber="127">4.91</td>
<td id="BB4754254F2972AEFA4DFAACFA21470A" box="[1431,1480,1295,1314]" gridcol="12" gridrow="25" pageId="4" pageNumber="127">4.85</td>
<td id="BB4754254F2972AEFA28FAACF9C5470A" box="[1522,1580,1295,1314]" gridcol="13" gridrow="25" pageId="4" pageNumber="127">5.09</td>
<td id="BB4754254F2972AEF9BBFAACF935470A" box="[1633,1756,1295,1314]" gridcol="14" gridrow="25" pageId="4" pageNumber="127">2.61</td>
<td id="BB4754254F2972AEF92FFAACF885470A" box="[1781,1900,1295,1314]" gridcol="15" gridrow="25" pageId="4" pageNumber="127">4.72</td>
</tr>
<tr id="F8963D594F2972AEFFBFFA84F8854712" box="[101,1900,1319,1338]" gridrow="26" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFA84FF7C4712" box="[101,149,1319,1338]" gridcol="0" gridrow="26" pageId="4" pageNumber="127">PPL</th>
<td id="BB4754254F2972AEFF3AFA84FEF94712" box="[224,272,1319,1338]" gridcol="1" gridrow="26" pageId="4" pageNumber="127">8.46</td>
<td id="BB4754254F2972AEFEE2FA84FE6C4712" box="[312,389,1319,1338]" gridcol="2" gridrow="26" pageId="4" pageNumber="127">9.11</td>
<td id="BB4754254F2972AEFE78FA84FE184712" box="[418,497,1319,1338]" gridcol="3" gridrow="26" pageId="4" pageNumber="127">9.26</td>
<td id="BB4754254F2972AEFDFBFA84FD814712" box="[545,616,1319,1338]" gridcol="4" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FA84FD214712" box="[655,712,1319,1338]" gridcol="5" gridrow="26" pageId="4" pageNumber="127">9.24</td>
<td id="BB4754254F2972AEFD24FA84FCD54712" box="[766,828,1319,1338]" gridcol="6" gridrow="26" pageId="4" pageNumber="127">8.47</td>
<td id="BB4754254F2972AEFCBBFA84FC454712" box="[865,940,1319,1338]" gridcol="7" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03FA84FBFE4712" box="[985,1047,1319,1338]" gridcol="8" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFA84FB6F4712" box="[1089,1158,1319,1338]" gridcol="9" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFA84FB114712" box="[1217,1272,1319,1338]" gridcol="10" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6FA84FA9F4712" box="[1324,1398,1319,1338]" gridcol="11" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DFA84FA214712" box="[1431,1480,1319,1338]" gridcol="12" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28FA84F9C54712" box="[1522,1580,1319,1338]" gridcol="13" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBFA84F9354712" box="[1633,1756,1319,1338]" gridcol="14" gridrow="26" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFA84F8854712" box="[1781,1900,1319,1338]" gridcol="15" gridrow="26" pageId="4" pageNumber="127">8.13</td>
</tr>
<tr id="F8963D594F2972AEFFBFFAE3F885477B" box="[101,1900,1344,1363]" gridrow="27" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFAE3FF7C477B" box="[101,149,1344,1363]" gridcol="0" gridrow="27" pageId="4" pageNumber="127">IFL</th>
<td id="BB4754254F2972AEFF3AFAE3FEF9477B" box="[224,272,1344,1363]" gridcol="1" gridrow="27" pageId="4" pageNumber="127">3.25</td>
<td id="BB4754254F2972AEFEE2FAE3FE6C477B" box="[312,389,1344,1363]" gridcol="2" gridrow="27" pageId="4" pageNumber="127">3.98</td>
<td id="BB4754254F2972AEFE78FAE3FE18477B" box="[418,497,1344,1363]" gridcol="3" gridrow="27" pageId="4" pageNumber="127">3.98</td>
<td id="BB4754254F2972AEFDFBFAE3FD81477B" box="[545,616,1344,1363]" gridcol="4" gridrow="27" pageId="4" pageNumber="127">3.42</td>
<td id="BB4754254F2972AEFD55FAE3FD21477B" box="[655,712,1344,1363]" gridcol="5" gridrow="27" pageId="4" pageNumber="127">4.45</td>
<td id="BB4754254F2972AEFD24FAE3FCD5477B" box="[766,828,1344,1363]" gridcol="6" gridrow="27" pageId="4" pageNumber="127">3.5</td>
<td id="BB4754254F2972AEFCBBFAE3FC45477B" box="[865,940,1344,1363]" gridcol="7" gridrow="27" pageId="4" pageNumber="127">4.62</td>
<td id="BB4754254F2972AEFC03FAE3FBFE477B" box="[985,1047,1344,1363]" gridcol="8" gridrow="27" pageId="4" pageNumber="127">3.78</td>
<td id="BB4754254F2972AEFB9BFAE3FB6F477B" box="[1089,1158,1344,1363]" gridcol="9" gridrow="27" pageId="4" pageNumber="127">3.87</td>
<td id="BB4754254F2972AEFB1BFAE3FB11477B" box="[1217,1272,1344,1363]" gridcol="10" gridrow="27" pageId="4" pageNumber="127">3.6</td>
<td id="BB4754254F2972AEFAF6FAE3FA9F477B" box="[1324,1398,1344,1363]" gridcol="11" gridrow="27" pageId="4" pageNumber="127">3.92</td>
<td id="BB4754254F2972AEFA4DFAE3FA21477B" box="[1431,1480,1344,1363]" gridcol="12" gridrow="27" pageId="4" pageNumber="127">3.29</td>
<td id="BB4754254F2972AEFA28FAE3F9C5477B" box="[1522,1580,1344,1363]" gridcol="13" gridrow="27" pageId="4" pageNumber="127">4.2</td>
<td id="BB4754254F2972AEF9BBFAE3F935477B" box="[1633,1756,1344,1363]" gridcol="14" gridrow="27" pageId="4" pageNumber="127">3.81</td>
<td id="BB4754254F2972AEF92FFAE3F885477B" box="[1781,1900,1344,1363]" gridcol="15" gridrow="27" pageId="4" pageNumber="127">4.2</td>
</tr>
<tr id="F8963D594F2972AEFFBFFAFBF8854743" box="[101,1900,1368,1387]" gridrow="28" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFAFBFF7C4743" box="[101,149,1368,1387]" gridcol="0" gridrow="28" pageId="4" pageNumber="127">IFB</th>
<td id="BB4754254F2972AEFF3AFAFBFEF94743" box="[224,272,1368,1387]" gridcol="1" gridrow="28" pageId="4" pageNumber="127">1.48</td>
<td id="BB4754254F2972AEFEE2FAFBFE6C4743" box="[312,389,1368,1387]" gridcol="2" gridrow="28" pageId="4" pageNumber="127">1.44</td>
<td id="BB4754254F2972AEFE78FAFBFE184743" box="[418,497,1368,1387]" gridcol="3" gridrow="28" pageId="4" pageNumber="127">1.49</td>
<td id="BB4754254F2972AEFDFBFAFBFD814743" box="[545,616,1368,1387]" gridcol="4" gridrow="28" pageId="4" pageNumber="127">1.35</td>
<td id="BB4754254F2972AEFD55FAFBFD214743" box="[655,712,1368,1387]" gridcol="5" gridrow="28" pageId="4" pageNumber="127">1.57</td>
<td id="BB4754254F2972AEFD24FAFBFCD54743" box="[766,828,1368,1387]" gridcol="6" gridrow="28" pageId="4" pageNumber="127">1.28</td>
<td id="BB4754254F2972AEFCBBFAFBFC454743" box="[865,940,1368,1387]" gridcol="7" gridrow="28" pageId="4" pageNumber="127">1.58</td>
<td id="BB4754254F2972AEFC03FAFBFBFE4743" box="[985,1047,1368,1387]" gridcol="8" gridrow="28" pageId="4" pageNumber="127">1.38</td>
<td id="BB4754254F2972AEFB9BFAFBFB6F4743" box="[1089,1158,1368,1387]" gridcol="9" gridrow="28" pageId="4" pageNumber="127">1.34</td>
<td id="BB4754254F2972AEFB1BFAFBFB114743" box="[1217,1272,1368,1387]" gridcol="10" gridrow="28" pageId="4" pageNumber="127">1.55</td>
<td id="BB4754254F2972AEFAF6FAFBFA9F4743" box="[1324,1398,1368,1387]" gridcol="11" gridrow="28" pageId="4" pageNumber="127">1.5</td>
<td id="BB4754254F2972AEFA4DFAFBFA214743" box="[1431,1480,1368,1387]" gridcol="12" gridrow="28" pageId="4" pageNumber="127">1.42</td>
<td id="BB4754254F2972AEFA28FAFBF9C54743" box="[1522,1580,1368,1387]" gridcol="13" gridrow="28" pageId="4" pageNumber="127">1.46</td>
<td id="BB4754254F2972AEF9BBFAFBF9354743" box="[1633,1756,1368,1387]" gridcol="14" gridrow="28" pageId="4" pageNumber="127">1.37</td>
<td id="BB4754254F2972AEF92FFAFBF8854743" box="[1781,1900,1368,1387]" gridcol="15" gridrow="28" pageId="4" pageNumber="127">1.24</td>
</tr>
<tr id="F8963D594F2972AEFFBFFAD2F88547AC" box="[101,1900,1393,1412]" gridrow="29" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFAD2FF7C47AC" box="[101,149,1393,1412]" gridcol="0" gridrow="29" pageId="4" pageNumber="127">MTL</th>
<td id="BB4754254F2972AEFF3AFAD2FEF947AC" box="[224,272,1393,1412]" gridcol="1" gridrow="29" pageId="4" pageNumber="127">4.23</td>
<td id="BB4754254F2972AEFEE2FAD2FE6C47AC" box="[312,389,1393,1412]" gridcol="2" gridrow="29" pageId="4" pageNumber="127">4.52</td>
<td id="BB4754254F2972AEFE78FAD2FE1847AC" box="[418,497,1393,1412]" gridcol="3" gridrow="29" pageId="4" pageNumber="127">4.26</td>
<td id="BB4754254F2972AEFDFBFAD2FD8147AC" box="[545,616,1393,1412]" gridcol="4" gridrow="29" pageId="4" pageNumber="127">3.95</td>
<td id="BB4754254F2972AEFD55FAD2FD2147AC" box="[655,712,1393,1412]" gridcol="5" gridrow="29" pageId="4" pageNumber="127">4.08</td>
<td id="BB4754254F2972AEFD24FAD2FCD547AC" box="[766,828,1393,1412]" gridcol="6" gridrow="29" pageId="4" pageNumber="127">4.29</td>
<td id="BB4754254F2972AEFCBBFAD2FC4547AC" box="[865,940,1393,1412]" gridcol="7" gridrow="29" pageId="4" pageNumber="127">4.73</td>
<td id="BB4754254F2972AEFC03FAD2FBFE47AC" box="[985,1047,1393,1412]" gridcol="8" gridrow="29" pageId="4" pageNumber="127">4.59</td>
<td id="BB4754254F2972AEFB9BFAD2FB6F47AC" box="[1089,1158,1393,1412]" gridcol="9" gridrow="29" pageId="4" pageNumber="127">4.39</td>
<td id="BB4754254F2972AEFB1BFAD2FB1147AC" box="[1217,1272,1393,1412]" gridcol="10" gridrow="29" pageId="4" pageNumber="127">4.45</td>
<td id="BB4754254F2972AEFAF6FAD2FA9F47AC" box="[1324,1398,1393,1412]" gridcol="11" gridrow="29" pageId="4" pageNumber="127">4.46</td>
<td id="BB4754254F2972AEFA4DFAD2FA2147AC" box="[1431,1480,1393,1412]" gridcol="12" gridrow="29" pageId="4" pageNumber="127">4.43</td>
<td id="BB4754254F2972AEFA28FAD2F9C547AC" box="[1522,1580,1393,1412]" gridcol="13" gridrow="29" pageId="4" pageNumber="127">4.82</td>
<td id="BB4754254F2972AEF9BBFAD2F93547AC" box="[1633,1756,1393,1412]" gridcol="14" gridrow="29" pageId="4" pageNumber="127">4,24</td>
<td id="BB4754254F2972AEF92FFAD2F88547AC" box="[1781,1900,1393,1412]" gridcol="15" gridrow="29" pageId="4" pageNumber="127">4.45</td>
</tr>
<tr id="F8963D594F2972AEFFBFFA29F88547B5" box="[101,1900,1418,1437]" gridrow="30" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFA29FF7C47B5" box="[101,149,1418,1437]" gridcol="0" gridrow="30" pageId="4" pageNumber="127">PBB</th>
<td id="BB4754254F2972AEFF3AFA29FEF947B5" box="[224,272,1418,1437]" gridcol="1" gridrow="30" pageId="4" pageNumber="127">2.21</td>
<td id="BB4754254F2972AEFEE2FA29FE6C47B5" box="[312,389,1418,1437]" gridcol="2" gridrow="30" pageId="4" pageNumber="127">2.52</td>
<td id="BB4754254F2972AEFE78FA29FE1847B5" box="[418,497,1418,1437]" gridcol="3" gridrow="30" pageId="4" pageNumber="127">2.53</td>
<td id="BB4754254F2972AEFDFBFA29FD8147B5" box="[545,616,1418,1437]" gridcol="4" gridrow="30" pageId="4" pageNumber="127">2</td>
<td id="BB4754254F2972AEFD55FA29FD2147B5" box="[655,712,1418,1437]" gridcol="5" gridrow="30" pageId="4" pageNumber="127">2.55</td>
<td id="BB4754254F2972AEFD24FA29FCD547B5" box="[766,828,1418,1437]" gridcol="6" gridrow="30" pageId="4" pageNumber="127">2.36</td>
<td id="BB4754254F2972AEFCBBFA29FC4547B5" box="[865,940,1418,1437]" gridcol="7" gridrow="30" pageId="4" pageNumber="127">2.71</td>
<td id="BB4754254F2972AEFC03FA29FBFE47B5" box="[985,1047,1418,1437]" gridcol="8" gridrow="30" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFA29FB6F47B5" box="[1089,1158,1418,1437]" gridcol="9" gridrow="30" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFA29FB1147B5" box="[1217,1272,1418,1437]" gridcol="10" gridrow="30" pageId="4" pageNumber="127">5.52</td>
<td id="BB4754254F2972AEFAF6FA29FA9F47B5" box="[1324,1398,1418,1437]" gridcol="11" gridrow="30" pageId="4" pageNumber="127">5.85</td>
<td id="BB4754254F2972AEFA4DFA29FA2147B5" box="[1431,1480,1418,1437]" gridcol="12" gridrow="30" pageId="4" pageNumber="127">5.75</td>
<td id="BB4754254F2972AEFA28FA29F9C547B5" box="[1522,1580,1418,1437]" gridcol="13" gridrow="30" pageId="4" pageNumber="127">6.62</td>
<td id="BB4754254F2972AEF9BBFA29F93547B5" box="[1633,1756,1418,1437]" gridcol="14" gridrow="30" pageId="4" pageNumber="127">6,24</td>
<td id="BB4754254F2972AEF92FFA29F88547B5" box="[1781,1900,1418,1437]" gridcol="15" gridrow="30" pageId="4" pageNumber="127">2.32</td>
</tr>
<tr id="F8963D594F2972AEFFBFFA01F885479D" box="[101,1900,1442,1461]" gridrow="31" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFA01FF7C479D" box="[101,149,1442,1461]" gridcol="0" gridrow="31" pageId="4" pageNumber="127">M1B</th>
<td id="BB4754254F2972AEFF3AFA01FEF9479D" box="[224,272,1442,1461]" gridcol="1" gridrow="31" pageId="4" pageNumber="127">1.21</td>
<td id="BB4754254F2972AEFEE2FA01FE6C479D" box="[312,389,1442,1461]" gridcol="2" gridrow="31" pageId="4" pageNumber="127">1.3</td>
<td id="BB4754254F2972AEFE78FA01FE18479D" box="[418,497,1442,1461]" gridcol="3" gridrow="31" pageId="4" pageNumber="127">1.15</td>
<td id="BB4754254F2972AEFDFBFA01FD81479D" box="[545,616,1442,1461]" gridcol="4" gridrow="31" pageId="4" pageNumber="127">1.1</td>
<td id="BB4754254F2972AEFD55FA01FD21479D" box="[655,712,1442,1461]" gridcol="5" gridrow="31" pageId="4" pageNumber="127">1.22</td>
<td id="BB4754254F2972AEFD24FA01FCD5479D" box="[766,828,1442,1461]" gridcol="6" gridrow="31" pageId="4" pageNumber="127">1.26</td>
<td id="BB4754254F2972AEFCBBFA01FC45479D" box="[865,940,1442,1461]" gridcol="7" gridrow="31" pageId="4" pageNumber="127">1.26</td>
<td id="BB4754254F2972AEFC03FA01FBFE479D" box="[985,1047,1442,1461]" gridcol="8" gridrow="31" pageId="4" pageNumber="127">1.31</td>
<td id="BB4754254F2972AEFB9BFA01FB6F479D" box="[1089,1158,1442,1461]" gridcol="9" gridrow="31" pageId="4" pageNumber="127">1.23</td>
<td id="BB4754254F2972AEFB1BFA01FB11479D" box="[1217,1272,1442,1461]" gridcol="10" gridrow="31" pageId="4" pageNumber="127">1.69</td>
<td id="BB4754254F2972AEFAF6FA01FA9F479D" box="[1324,1398,1442,1461]" gridcol="11" gridrow="31" pageId="4" pageNumber="127">1.3</td>
<td id="BB4754254F2972AEFA4DFA01FA21479D" box="[1431,1480,1442,1461]" gridcol="12" gridrow="31" pageId="4" pageNumber="127">1.39</td>
<td id="BB4754254F2972AEFA28FA01F9C5479D" box="[1522,1580,1442,1461]" gridcol="13" gridrow="31" pageId="4" pageNumber="127">1.32</td>
<td id="BB4754254F2972AEF9BBFA01F935479D" box="[1633,1756,1442,1461]" gridcol="14" gridrow="31" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FFA01F885479D" box="[1781,1900,1442,1461]" gridcol="15" gridrow="31" pageId="4" pageNumber="127">1.26</td>
</tr>
<tr id="F8963D594F2972AEFFBFFA18F88547E6" box="[101,1900,1467,1486]" gridrow="32" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFA18FF7C47E6" box="[101,149,1467,1486]" gridcol="0" gridrow="32" pageId="4" pageNumber="127">ZB</th>
<td id="BB4754254F2972AEFF3AFA18FEF947E6" box="[224,272,1467,1486]" gridcol="1" gridrow="32" pageId="4" pageNumber="127">14.71</td>
<td id="BB4754254F2972AEFEE2FA18FE6C47E6" box="[312,389,1467,1486]" gridcol="2" gridrow="32" pageId="4" pageNumber="127">15.88</td>
<td id="BB4754254F2972AEFE78FA18FE1847E6" box="[418,497,1467,1486]" gridcol="3" gridrow="32" pageId="4" pageNumber="127">15.9</td>
<td id="BB4754254F2972AEFDFBFA18FD8147E6" box="[545,616,1467,1486]" gridcol="4" gridrow="32" pageId="4" pageNumber="127">12.59</td>
<td id="BB4754254F2972AEFD55FA18FD2147E6" box="[655,712,1467,1486]" gridcol="5" gridrow="32" pageId="4" pageNumber="127">15.9</td>
<td id="BB4754254F2972AEFD24FA18FCD547E6" box="[766,828,1467,1486]" gridcol="6" gridrow="32" pageId="4" pageNumber="127">15.86</td>
<td id="BB4754254F2972AEFCBBFA18FC4547E6" box="[865,940,1467,1486]" gridcol="7" gridrow="32" pageId="4" pageNumber="127">16.75</td>
<td id="BB4754254F2972AEFC03FA18FBFE47E6" box="[985,1047,1467,1486]" gridcol="8" gridrow="32" pageId="4" pageNumber="127">13.71</td>
<td id="BB4754254F2972AEFB9BFA18FB6F47E6" box="[1089,1158,1467,1486]" gridcol="9" gridrow="32" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFA18FB1147E6" box="[1217,1272,1467,1486]" gridcol="10" gridrow="32" pageId="4" pageNumber="127">12.96</td>
<td id="BB4754254F2972AEFAF6FA18FA9F47E6" box="[1324,1398,1467,1486]" gridcol="11" gridrow="32" pageId="4" pageNumber="127">14.08</td>
<td id="BB4754254F2972AEFA4DFA18FA2147E6" box="[1431,1480,1467,1486]" gridcol="12" gridrow="32" pageId="4" pageNumber="127">13.81</td>
<td id="BB4754254F2972AEFA28FA18F9C547E6" box="[1522,1580,1467,1486]" gridcol="13" gridrow="32" pageId="4" pageNumber="127">15.49</td>
<td id="BB4754254F2972AEF9BBFA18F93547E6" box="[1633,1756,1467,1486]" gridcol="14" gridrow="32" pageId="4" pageNumber="127">14.62</td>
<td id="BB4754254F2972AEF92FFA18F88547E6" box="[1781,1900,1467,1486]" gridcol="15" gridrow="32" pageId="4" pageNumber="127">14.34</td>
</tr>
<tr id="F8963D594F2972AEFFBFFA70F88547CE" box="[101,1900,1491,1510]" gridrow="33" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFA70FF7C47CE" box="[101,149,1491,1510]" gridcol="0" gridrow="33" pageId="4" pageNumber="127">BB</th>
<td id="BB4754254F2972AEFF3AFA70FEF947CE" box="[224,272,1491,1510]" gridcol="1" gridrow="33" pageId="4" pageNumber="127">12.01</td>
<td id="BB4754254F2972AEFEE2FA70FE6C47CE" box="[312,389,1491,1510]" gridcol="2" gridrow="33" pageId="4" pageNumber="127">13.89</td>
<td id="BB4754254F2972AEFE78FA70FE1847CE" box="[418,497,1491,1510]" gridcol="3" gridrow="33" pageId="4" pageNumber="127">13.2</td>
<td id="BB4754254F2972AEFDFBFA70FD8147CE" box="[545,616,1491,1510]" gridcol="4" gridrow="33" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFD55FA70FD2147CE" box="[655,712,1491,1510]" gridcol="5" gridrow="33" pageId="4" pageNumber="127">11.24</td>
<td id="BB4754254F2972AEFD24FA70FCD547CE" box="[766,828,1491,1510]" gridcol="6" gridrow="33" pageId="4" pageNumber="127">13.37</td>
<td id="BB4754254F2972AEFCBBFA70FC4547CE" box="[865,940,1491,1510]" gridcol="7" gridrow="33" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03FA70FBFE47CE" box="[985,1047,1491,1510]" gridcol="8" gridrow="33" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BFA70FB6F47CE" box="[1089,1158,1491,1510]" gridcol="9" gridrow="33" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BFA70FB1147CE" box="[1217,1272,1491,1510]" gridcol="10" gridrow="33" pageId="4" pageNumber="127">12.83</td>
<td id="BB4754254F2972AEFAF6FA70FA9F47CE" box="[1324,1398,1491,1510]" gridcol="11" gridrow="33" pageId="4" pageNumber="127">13.1</td>
<td id="BB4754254F2972AEFA4DFA70FA2147CE" box="[1431,1480,1491,1510]" gridcol="12" gridrow="33" pageId="4" pageNumber="127">13.22</td>
<td id="BB4754254F2972AEFA28FA70F9C547CE" box="[1522,1580,1491,1510]" gridcol="13" gridrow="33" pageId="4" pageNumber="127">13.33</td>
<td id="BB4754254F2972AEF9BBFA70F93547CE" box="[1633,1756,1491,1510]" gridcol="14" gridrow="33" pageId="4" pageNumber="127">12.28</td>
<td id="BB4754254F2972AEF92FFA70F88547CE" box="[1781,1900,1491,1510]" gridcol="15" gridrow="33" pageId="4" pageNumber="127">13.08</td>
</tr>
<tr id="F8963D594F2972AEFFBFFA4FF88547D7" box="[101,1900,1516,1535]" gridrow="34" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFFA4FFF7C47D7" box="[101,149,1516,1535]" gridcol="0" gridrow="34" pageId="4" pageNumber="127">ZPB</th>
<td id="BB4754254F2972AEFF3AFA4FFEF947D7" box="[224,272,1516,1535]" gridcol="1" gridrow="34" pageId="4" pageNumber="127">2.75</td>
<td id="BB4754254F2972AEFEE2FA4FFE6C47D7" box="[312,389,1516,1535]" gridcol="2" gridrow="34" pageId="4" pageNumber="127">3.3</td>
<td id="BB4754254F2972AEFE78FA4FFE1847D7" box="[418,497,1516,1535]" gridcol="3" gridrow="34" pageId="4" pageNumber="127">3.03</td>
<td id="BB4754254F2972AEFDFBFA4FFD8147D7" box="[545,616,1516,1535]" gridcol="4" gridrow="34" pageId="4" pageNumber="127">2.18</td>
<td id="BB4754254F2972AEFD55FA4FFD2147D7" box="[655,712,1516,1535]" gridcol="5" gridrow="34" pageId="4" pageNumber="127">2.92</td>
<td id="BB4754254F2972AEFD24FA4FFCD547D7" box="[766,828,1516,1535]" gridcol="6" gridrow="34" pageId="4" pageNumber="127">2.96</td>
<td id="BB4754254F2972AEFCBBFA4FFC4547D7" box="[865,940,1516,1535]" gridcol="7" gridrow="34" pageId="4" pageNumber="127">3.65</td>
<td id="BB4754254F2972AEFC03FA4FFBFE47D7" box="[985,1047,1516,1535]" gridcol="8" gridrow="34" pageId="4" pageNumber="127">2.91</td>
<td id="BB4754254F2972AEFB9BFA4FFB6F47D7" box="[1089,1158,1516,1535]" gridcol="9" gridrow="34" pageId="4" pageNumber="127">3.08</td>
<td id="BB4754254F2972AEFB1BFA4FFB1147D7" box="[1217,1272,1516,1535]" gridcol="10" gridrow="34" pageId="4" pageNumber="127">3.31</td>
<td id="BB4754254F2972AEFAF6FA4FFA9F47D7" box="[1324,1398,1516,1535]" gridcol="11" gridrow="34" pageId="4" pageNumber="127">3.13</td>
<td id="BB4754254F2972AEFA4DFA4FFA2147D7" box="[1431,1480,1516,1535]" gridcol="12" gridrow="34" pageId="4" pageNumber="127">3.5</td>
<td id="BB4754254F2972AEFA28FA4FF9C547D7" box="[1522,1580,1516,1535]" gridcol="13" gridrow="34" pageId="4" pageNumber="127">3.58</td>
<td id="BB4754254F2972AEF9BBFA4FF93547D7" box="[1633,1756,1516,1535]" gridcol="14" gridrow="34" pageId="4" pageNumber="127">3,51</td>
<td id="BB4754254F2972AEF92FFA4FF88547D7" box="[1781,1900,1516,1535]" gridcol="15" gridrow="34" pageId="4" pageNumber="127">2.86</td>
</tr>
<tr id="F8963D594F2972AEFFBFF9A7F885443F" box="[101,1900,1540,1559]" gridrow="35" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFF9A7FF7C443F" box="[101,149,1540,1559]" gridcol="0" gridrow="35" pageId="4" pageNumber="127">ID</th>
<td id="BB4754254F2972AEFF3AF9A7FEF9443F" box="[224,272,1540,1559]" gridcol="1" gridrow="35" pageId="4" pageNumber="127">1.47</td>
<td id="BB4754254F2972AEFEE2F9A7FE6C443F" box="[312,389,1540,1559]" gridcol="2" gridrow="35" pageId="4" pageNumber="127">1.91</td>
<td id="BB4754254F2972AEFE78F9A7FE18443F" box="[418,497,1540,1559]" gridcol="3" gridrow="35" pageId="4" pageNumber="127">1.87</td>
<td id="BB4754254F2972AEFDFBF9A7FD81443F" box="[545,616,1540,1559]" gridcol="4" gridrow="35" pageId="4" pageNumber="127">1.25</td>
<td id="BB4754254F2972AEFD55F9A7FD21443F" box="[655,712,1540,1559]" gridcol="5" gridrow="35" pageId="4" pageNumber="127">1.78</td>
<td id="BB4754254F2972AEFD24F9A7FCD5443F" box="[766,828,1540,1559]" gridcol="6" gridrow="35" pageId="4" pageNumber="127">1.74</td>
<td id="BB4754254F2972AEFCBBF9A7FC45443F" box="[865,940,1540,1559]" gridcol="7" gridrow="35" pageId="4" pageNumber="127">1.8</td>
<td id="BB4754254F2972AEFC03F9A7FBFE443F" box="[985,1047,1540,1559]" gridcol="8" gridrow="35" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BF9A7FB6F443F" box="[1089,1158,1540,1559]" gridcol="9" gridrow="35" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BF9A7FB11443F" box="[1217,1272,1540,1559]" gridcol="10" gridrow="35" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6F9A7FA9F443F" box="[1324,1398,1540,1559]" gridcol="11" gridrow="35" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DF9A7FA21443F" box="[1431,1480,1540,1559]" gridcol="12" gridrow="35" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28F9A7F9C5443F" box="[1522,1580,1540,1559]" gridcol="13" gridrow="35" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBF9A7F935443F" box="[1633,1756,1540,1559]" gridcol="14" gridrow="35" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FF9A7F885443F" box="[1781,1900,1540,1559]" gridcol="15" gridrow="35" pageId="4" pageNumber="127">1.53</td>
</tr>
<tr id="F8963D594F2972AEFFBFF9BEF8854418" box="[101,1900,1565,1584]" gridrow="36" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFF9BEFF7C4418" box="[101,149,1565,1584]" gridcol="0" gridrow="36" pageId="4" pageNumber="127">BH</th>
<td id="BB4754254F2972AEFF3AF9BEFEF94418" box="[224,272,1565,1584]" gridcol="1" gridrow="36" pageId="4" pageNumber="127">8.52</td>
<td id="BB4754254F2972AEFEE2F9BEFE6C4418" box="[312,389,1565,1584]" gridcol="2" gridrow="36" pageId="4" pageNumber="127">9.31</td>
<td id="BB4754254F2972AEFE78F9BEFE184418" box="[418,497,1565,1584]" gridcol="3" gridrow="36" pageId="4" pageNumber="127">8.96</td>
<td id="BB4754254F2972AEFDFBF9BEFD814418" box="[545,616,1565,1584]" gridcol="4" gridrow="36" pageId="4" pageNumber="127">8.24</td>
<td id="BB4754254F2972AEFD55F9BEFD214418" box="[655,712,1565,1584]" gridcol="5" gridrow="36" pageId="4" pageNumber="127">9.02</td>
<td id="BB4754254F2972AEFD24F9BEFCD54418" box="[766,828,1565,1584]" gridcol="6" gridrow="36" pageId="4" pageNumber="127">9</td>
<td id="BB4754254F2972AEFCBBF9BEFC454418" box="[865,940,1565,1584]" gridcol="7" gridrow="36" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03F9BEFBFE4418" box="[985,1047,1565,1584]" gridcol="8" gridrow="36" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BF9BEFB6F4418" box="[1089,1158,1565,1584]" gridcol="9" gridrow="36" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BF9BEFB114418" box="[1217,1272,1565,1584]" gridcol="10" gridrow="36" pageId="4" pageNumber="127">8.69</td>
<td id="BB4754254F2972AEFAF6F9BEFA9F4418" box="[1324,1398,1565,1584]" gridcol="11" gridrow="36" pageId="4" pageNumber="127">9.02</td>
<td id="BB4754254F2972AEFA4DF9BEFA214418" box="[1431,1480,1565,1584]" gridcol="12" gridrow="36" pageId="4" pageNumber="127">8.72</td>
<td id="BB4754254F2972AEFA28F9BEF9C54418" box="[1522,1580,1565,1584]" gridcol="13" gridrow="36" pageId="4" pageNumber="127">9.22</td>
<td id="BB4754254F2972AEF9BBF9BEF9354418" box="[1633,1756,1565,1584]" gridcol="14" gridrow="36" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FF9BEF8854418" box="[1781,1900,1565,1584]" gridcol="15" gridrow="36" pageId="4" pageNumber="127">9.05</td>
</tr>
<tr id="F8963D594F2972AEFFBFF995F8854461" box="[101,1900,1590,1609]" gridrow="37" pageId="4" pageNumber="127">
<th id="BB4754254F2972AEFFBFF995FF7C4461" box="[101,149,1590,1609]" gridcol="0" gridrow="37" pageId="4" pageNumber="127">MFB</th>
<td id="BB4754254F2972AEFF3AF995FEF94461" box="[224,272,1590,1609]" gridcol="1" gridrow="37" pageId="4" pageNumber="127">1.34</td>
<td id="BB4754254F2972AEFEE2F995FE6C4461" box="[312,389,1590,1609]" gridcol="2" gridrow="37" pageId="4" pageNumber="127">1.72</td>
<td id="BB4754254F2972AEFE78F995FE184461" box="[418,497,1590,1609]" gridcol="3" gridrow="37" pageId="4" pageNumber="127">1.46</td>
<td id="BB4754254F2972AEFDFBF995FD814461" box="[545,616,1590,1609]" gridcol="4" gridrow="37" pageId="4" pageNumber="127">1.17</td>
<td id="BB4754254F2972AEFD55F995FD214461" box="[655,712,1590,1609]" gridcol="5" gridrow="37" pageId="4" pageNumber="127">1.36</td>
<td id="BB4754254F2972AEFD24F995FCD54461" box="[766,828,1590,1609]" gridcol="6" gridrow="37" pageId="4" pageNumber="127">1.4</td>
<td id="BB4754254F2972AEFCBBF995FC454461" box="[865,940,1590,1609]" gridcol="7" gridrow="37" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFC03F995FBFE4461" box="[985,1047,1590,1609]" gridcol="8" gridrow="37" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB9BF995FB6F4461" box="[1089,1158,1590,1609]" gridcol="9" gridrow="37" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFB1BF995FB114461" box="[1217,1272,1590,1609]" gridcol="10" gridrow="37" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFAF6F995FA9F4461" box="[1324,1398,1590,1609]" gridcol="11" gridrow="37" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA4DF995FA214461" box="[1431,1480,1590,1609]" gridcol="12" gridrow="37" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEFA28F995F9C54461" box="[1522,1580,1590,1609]" gridcol="13" gridrow="37" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF9BBF995F9354461" box="[1633,1756,1590,1609]" gridcol="14" gridrow="37" pageId="4" pageNumber="127">?</td>
<td id="BB4754254F2972AEF92FF995F8854461" box="[1781,1900,1590,1609]" gridcol="15" gridrow="37" pageId="4" pageNumber="127">1.52</td>
</tr>
</table>
</paragraph>
<tableNote id="BB403E954F298D55FFBCF9CEFB8544B4" pageId="4" pageNumber="127" targetBox="[101,1900,642,1609]" targetPageId="4">
<paragraph id="46193F1B4F298D55FFBCF9CEFCB844A9" blockId="4.[102,1132,1645,1692]" box="[102,849,1645,1665]" pageId="4" pageNumber="127">
<superScript id="B1D392534F298D55FFBCF9CEFF4244A9" attach="right" box="[102,171,1645,1665]" fontSize="5" pageId="4" pageNumber="127">aSource</superScript>
: Luna and Patterson (2003),
<bibRefCitation id="223742EA4F298D55FE44F9CDFD8644A9" author="VILLALPANDO, G. &amp; J. VARGAS &amp; J. SALAZAR-BRAVO" box="[414,623,1646,1665]" pageId="4" pageNumber="127" pagination="143 - 149" refId="ref14774" refString="VILLALPANDO, G., J. VARGAS, AND J. SALAZAR-BRAVO. 2006. First record of Rhagomys (Mammalia: Sigmodontinae) in Bolivia. Mastozoologia Neotropical 13: 143 - 149." type="journal article" year="2006">Villalpando et al. (2006)</bibRefCitation>
, and
<bibRefCitation id="223742EA4F298D55FD44F9CDFCA544A9" author="MEDINA, C. E. &amp; D. R. DIAZ &amp; K. PINO &amp; A. PARI &amp; H. ZEBALLOS" box="[670,844,1646,1665]" pageId="4" pageNumber="127" pagination="2136" refId="ref13384" refString="MEDINA, C. E., D. R. DIAZ, K. PINO, A. PARI, AND H. ZEBALLOS. 2017. New locality records of Rhagomys longilingua Luna &amp; Patterson, 2003 (Rodentia: Cricetidae) in Peru. Check List 13: 2136." type="journal article" year="2017">Medina et al. (2017)</bibRefCitation>
.
</paragraph>
<paragraph id="46193F1B4F298D55FFBCF924FB8544B4" blockId="4.[102,1132,1645,1692]" box="[102,1132,1671,1692]" pageId="4" pageNumber="127">
<superScript id="B1D392534F298D55FFBCF924FF4244B4" attach="right" box="[102,171,1671,1692]" fontSize="5" pageId="4" pageNumber="127">bSource</superScript>
: Luna and Patterson (2003),
<bibRefCitation id="223742EA4F298D55FE7AF92AFD8744B4" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[416,622,1673,1692]" pageId="4" pageNumber="127" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. (2004)</bibRefCitation>
,
<bibRefCitation id="223742EA4F298D55FDA3F92AFCA944B4" author="PASSAMANI, M. &amp; R. A. SERPA CERBONCINI &amp; J. E. DE OLIVEIRA" box="[633,832,1673,1692]" pageId="4" pageNumber="127" pagination="827 - 831" refId="ref13707" refString="PASSAMANI, M., R. A. SERPA CERBONCINI, AND J. E. DE OLIVEIRA. 2011. Distribution extension of Phaenomys ferrugineus (Thomas, 1894), and new data on Abrawayaomys ruschii Cunha and Cruz, 1979 and Rhagomys rufescens (Thomas, 1886), three rare species of rodents (Rodentia: Cricetidae) in Minas Gerais, Brazil. Check List 7: 827 - 831" type="journal article" year="2011">Passamani et al. (2011)</bibRefCitation>
, and Carvalho and Silvério (2012).
</paragraph>
</tableNote>
<caption id="12D96F934F288D54FFBFF8B1FABA456C" ID-DOI="http://doi.org/10.5281/zenodo.10632397" ID-Zenodo-Dep="10632397" httpUri="https://zenodo.org/record/10632397/files/figure.png" pageId="5" pageNumber="128" startId="5.[101,136,1810,1831]" targetBox="[116,1473,202,1794]" targetPageId="5" targetType="figure">
<paragraph id="46193F1B4F288D54FFBFF8B1FABA456C" blockId="5.[101,1488,1810,1860]" pageId="5" pageNumber="128">
<emphasis id="74D2E3094F288D54FFBFF8B1FF4F450F" bold="true" box="[101,166,1810,1831]" pageId="5" pageNumber="128">Fig. 2.</emphasis>
—Phylogenetic tree based on the concatenated alignment of
<emphasis id="74D2E3094F288D54FD2EF8B1FCC8450F" box="[756,801,1810,1831]" italics="true" pageId="5" pageNumber="128">Cytb</emphasis>
,
<emphasis id="74D2E3094F288D54FCF4F8B1FCB2450F" box="[814,859,1810,1831]" italics="true" pageId="5" pageNumber="128">CO1</emphasis>
, and
<emphasis id="74D2E3094F288D54FC48F8B1FC2F450F" box="[914,966,1810,1831]" italics="true" pageId="5" pageNumber="128">IBRP</emphasis>
genes; Maximum Likelihood and Bayesian Inference support values (BS / PP) are presented at tribal and generic nodes. The dashed rectangle marks the position of the genus
<taxonomicName id="81A644984F288D54FB33F88CFAA4456C" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1257,1357,1839,1860]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="128" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F288D54FB33F88CFAA4456C" box="[1257,1357,1839,1860]" italics="true" pageId="5" pageNumber="128">Rhagomys</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="46193F1B4F2B8D57FF71FE8AFED743E8" blockId="6.[144,816,201,1984]" pageId="6" pageNumber="129">
<emphasis id="74D2E3094F2B8D57FF71FE8AFEA64368" box="[171,335,297,320]" italics="true" pageId="6" pageNumber="129">Type locality.</emphasis>
<materialsCitation id="F6CE35464F2B8D57FEB0FE8AFED043E8" country="Ecuador" county="Refugio de Vida Silvestre" elevation="1442" latitude="-3.803611" location="Cordillera del Condor" longLatPrecision="21" longitude="-78.50278" municipality="Rio Blanco" pageId="6" pageNumber="129" specimenCount="1" stateProvince="Zamora-Chinchipe">
<collectingCountry id="3EB17F8B4F2B8D57FEB0FE8AFE2D4368" box="[362,452,297,320]" name="Ecuador" pageId="6" pageNumber="129">Ecuador</collectingCountry>
,
<collectingRegion id="8462F1F94F2B8D57FE3EFE8AFCC34368" box="[484,810,297,320]" country="Ecuador" name="Zamora-Chinchipe" pageId="6" pageNumber="129">Zamora-Chinchipe province</collectingRegion>
,
<collectingCounty id="AF7847974F2B8D57FF4AFEEAFE4F4348" box="[144,422,329,352]" pageId="6" pageNumber="129">Refugio de Vida Silvestre</collectingCounty>
“El Zarza,” on the shore of the
<collectingMunicipality id="A67DA5614F2B8D57FCDDFEEAFF3743A8" pageId="6" pageNumber="129">Río Blanco</collectingMunicipality>
,
<locationDeviation id="57AB26394F2B8D57FF32FECAFE2B43A8" box="[232,450,361,384]" location="San Antonio" pageId="6" pageNumber="129">
<quantity id="815E92FE4F2B8D57FF32FECAFEF743A8" box="[232,286,361,384]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="129" unit="km" value="2.0">2 km</quantity>
N San Antonio
</locationDeviation>
(town),
<geoCoordinate id="239259DC4F2B8D57FDC6FECAFD7143A8" box="[540,664,359,385]" degrees="03" direction="south" minutes="48" orientation="latitude" pageId="6" pageNumber="129" precision="15" seconds="13" value="-3.803611">
03°48
<emphasis id="74D2E3094F2B8D57FD86FEC4FD8A43A8" box="[604,611,359,384]" italics="true" pageId="6" pageNumber="129"></emphasis>
13
<emphasis id="74D2E3094F2B8D57FDA7FECBFD6143A9" box="[637,648,360,385]" italics="true" pageId="6" pageNumber="129">ʺ</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="239259DC4F2B8D57FD7EFECAFCC543A8" box="[676,812,359,385]" degrees="78" direction="west" minutes="30" orientation="longitude" pageId="6" pageNumber="129" precision="15" seconds="10" value="-78.50278">
78°30
<emphasis id="74D2E3094F2B8D57FD3EFEC4FD0243A8" box="[740,747,359,384]" italics="true" pageId="6" pageNumber="129"></emphasis>
10
<emphasis id="74D2E3094F2B8D57FCDFFECBFCF943A9" box="[773,784,360,385]" italics="true" pageId="6" pageNumber="129">ʺ</emphasis>
W
</geoCoordinate>
,
<quantity id="815E92FE4F2B8D57FF4AFE2AFF0E4388" box="[144,231,393,416]" metricMagnitude="3" metricUnit="m" metricValue="1.4419999999999997" pageId="6" pageNumber="129" unit="m" value="1442.0">
<elevation id="CD8BD8284F2B8D57FF4AFE2AFF0E4388" box="[144,231,393,416]" metricMagnitude="3" metricUnit="m" metricValue="1.4419999999999997" pageId="6" pageNumber="129" unit="m" value="1442.0">1,442 m</elevation>
</quantity>
elevation, in the western foothills of the
<location id="437969C04F2B8D57FD40FE2AFF0943E8" LSID="urn:lsid:plazi:treatment:CE0F8E0D4F2E8D5DFC1EF91CFA154768:437969C04F2B8D57FD40FE2AFF0943E8" country="Ecuador" county="Refugio de Vida Silvestre" latitude="-3.803611" longLatPrecision="21" longitude="-78.50278" municipality="Rio Blanco" name="Cordillera del Condor" pageId="6" pageNumber="129" stateProvince="Zamora-Chinchipe">Cordillera del Cóndor</location>
(
<figureCitation id="DE9D239E4F2B8D57FF2AFE0AFEC443E8" box="[240,301,425,448]" captionStart="Fig" captionStartId="1.[101,136,1893,1914]" captionTargetBox="[116,1473,975,1878]" captionTargetId="figure-6@1.[114,1475,973,1880]" captionTargetPageId="1" captionText="Fig. 1.—Known geographic distribution of the genus Rhagomys. Solid triangles represent records of R. rufescens inAtlantic Forest, Brazil; the inverted triangle marks Rhagomys sp., from Brazilian Amazonia; squares denote records of R. longilingua in Andean Peru and Bolivia, and a record in Brazilian Amazonia. The star marks the location of Rhagomys septentrionalis sp. nov., near the Cordillera del Cóndor of Ecuador. Numbers refer to collecting localities in the Appendix." figureDoi="http://doi.org/10.5281/zenodo.10632395" httpUri="https://zenodo.org/record/10632395/files/figure.png" pageId="6" pageNumber="129">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0EBC6C904F2B8D57FF71FE6AFD9E4008" pageId="6" pageNumber="129" type="etymology">
<paragraph id="46193F1B4F2B8D57FF71FE6AFD9E4008" blockId="6.[144,816,201,1984]" pageId="6" pageNumber="129">
<emphasis id="74D2E3094F2B8D57FF71FE6AFECA43C8" box="[171,291,457,480]" italics="true" pageId="6" pageNumber="129">Etymology.</emphasis>
—The name
<taxonomicName id="81A644984F2B8D57FE75FE6AFDB343C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[431,602,457,480]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F2B8D57FE61FE6AFDB343C8" box="[443,602,457,480]" italics="true" pageId="6" pageNumber="129">septentrionalis</emphasis>
</taxonomicName>
” comes from Latin meaning “of the north,” referring to the distribution of this species at the northern edge of the generic range.
</paragraph>
</subSubSection>
<subSubSection id="0EBC6C904F2B8D57FF71FD8AFDC54728" pageId="6" pageNumber="129" type="diagnosis">
<paragraph id="46193F1B4F2B8D57FF71FD8AFD9C4088" blockId="6.[144,816,201,1984]" pageId="6" pageNumber="129">
<emphasis id="74D2E3094F2B8D57FF71FD8AFEF74068" box="[171,286,553,576]" italics="true" pageId="6" pageNumber="129">Diagnosis.</emphasis>
—Orange-colored mouse with spiny hairs on the back and rump to the base of the tail, without a black ring of skin around the eyes, and without superciliary vibrissae. Hair color on the tail is the same above and below.
</paragraph>
<paragraph id="46193F1B4F2B8D57FF71FD0AFDC54728" blockId="6.[144,816,201,1984]" pageId="6" pageNumber="129">
The interorbital region is narrow and hourglass-shaped, with slightly marked supraorbital cristae. The postorbital process is short and inconspicuous (
<figureCitation id="DE9D239E4F2B8D57FE7FFD4AFE0C4128" box="[421,485,745,768]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="6" pageNumber="129">Fig. 3</figureCitation>
). The ectotympanics are large and round. The lower extreme of the squamosal hamular process reaches halfway up the mastoid. A conspicuous transverse canal is located at the sides of the cranium at the base of the pterygoid. The incisive foramina are long and narrow. The M1 presents a procingulum interrupted by a prominent anterolabial conule that occupies virtually the entire leading margin of M1, and the anterolingual conule is almost inconspicuous, marked by a minute anteromedian flexus. The anteroloph of the M1 is directly connected to the posterior section of the anterolabial conule, without an intermediate connection (anterolophule). The anterior mure of M1 and M2 are directly connected to the anterior margin of the protocone and surpass the posterior margin of the protocone to shape the median mure, while the paracones of M1 and M2 do not connect with the median mure, joining the paraflexus with the mesoflexus. The m3 does not have an ectostylid on its labial margin.
</paragraph>
</subSubSection>
<subSubSection id="0EBC6C904F2B8D5DFF71FAAAFBD44628" lastPageId="12" lastPageNumber="135" pageId="6" pageNumber="129" type="description">
<paragraph id="46193F1B4F2B8D57FF71FAAAFCC74408" blockId="6.[144,816,201,1984]" pageId="6" pageNumber="129">
<emphasis id="74D2E3094F2B8D57FF71FAAAFED94708" box="[171,304,1289,1312]" italics="true" pageId="6" pageNumber="129">Description.</emphasis>
—The description is based solely on the
<typeStatus id="991D81B94F2B8D57FD2FFAAAFF574768" pageId="6" pageNumber="129" type="holotype">holotype</typeStatus>
(
<collectionCode id="20B7A7DE4F2B8D57FF17FA8AFEF04768" box="[205,281,1321,1344]" pageId="6" pageNumber="129">MEPN</collectionCode>
10898). This specimen closely resembles the descriptions of other
<taxonomicName id="81A644984F2B8D57FE85FAEAFE274748" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[351,462,1353,1376]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="129" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F2B8D57FE85FAEAFE274748" box="[351,462,1353,1376]" italics="true" pageId="6" pageNumber="129">Rhagomys</emphasis>
</taxonomicName>
(
<bibRefCitation id="223742EA4F2B8D57FE05FAEAFD9B4748" author="THOMAS, O." box="[479,626,1353,1376]" pageId="6" pageNumber="129" pagination="250 - 251" refId="ref14652" refString="THOMAS, O. 1886. Description on a new Brazilian species of Hesperomys. Annals &amp; Magazine of Natural History, series 5, 17: 250 - 251." type="journal article" year="1886">Thomas 1886</bibRefCitation>
,
<bibRefCitation id="223742EA4F2B8D57FD58FAEAFD5E4748" author="THOMAS, O." box="[642,695,1353,1376]" pageId="6" pageNumber="129" pagination="192 - 198" refId="ref14683" refString="THOMAS, O. 1917. On the arrangement of the South American rats allied to Oryzomys and Rhipidomys. Annals &amp; Magazine of Natural History, series 8, 20: 192 - 198." type="journal article" year="1917">1917</bibRefCitation>
; Luna and Patterson 2003;
<bibRefCitation id="223742EA4F2B8D57FEE4FACAFDC647A8" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[318,559,1385,1408]" pageId="6" pageNumber="129" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
) and is a mouse of medium size and weight (total length
<quantity id="815E92FE4F2B8D57FDD6FA2AFD8C4788" box="[524,613,1417,1440]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="129" unit="mm" value="190.0">190 mm</quantity>
and weight
<quantity id="815E92FE4F2B8D57FD34FA2AFCC84788" box="[750,801,1417,1440]" metricMagnitude="-2" metricUnit="kg" metricValue="2.3" pageId="6" pageNumber="129" unit="g" value="23.0">23 g</quantity>
). The
<typeStatus id="991D81B94F2B8D57FF1FFA0AFECB47E8" box="[197,290,1449,1472]" pageId="6" pageNumber="129" type="holotype">holotype</typeStatus>
has orange spiny pelage, small ears, flattened dental structure, without folds, with simple, conical cusps like those of the rodent genus
<taxonomicName id="81A644984F2B8D57FE47FA4AFDA84428" authorityName="Cunha &amp; Cruz" authorityYear="1979" box="[413,577,1513,1536]" class="Mammalia" family="Cricetidae" genus="Abrawayaomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="129" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F2B8D57FE47FA4AFDA84428" box="[413,577,1513,1536]" italics="true" pageId="6" pageNumber="129">Abrawayaomys</emphasis>
</taxonomicName>
, with a round nail embedded in the hallux of the foot, with large callous plantar pads.
</paragraph>
<paragraph id="46193F1B4F2B8D57FF71F98AFC1043E8" blockId="6.[144,816,201,1984]" lastBlockId="6.[861,1531,201,1984]" pageId="6" pageNumber="129">
<emphasis id="74D2E3094F2B8D57FF71F98AFE634468" box="[171,394,1577,1600]" italics="true" pageId="6" pageNumber="129">External characters.</emphasis>
—The body is covered by short hairs DFL =
<quantity id="815E92FE4F2B8D57FF3AF9EAFEF74448" box="[224,286,1609,1632]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="129" unit="mm" value="9.0">9 mm</quantity>
(
<tableCitation id="0B240AA04F2B8D57FEF4F9EAFE944448" box="[302,381,1609,1632]" captionStart="Table 1" captionStartId="4.[126,182,506,527]" captionTargetPageId="4" captionText="Table 1.—External, vibrissal, and craniodental measurements (mm), and age class, for specimens of Rhagomys. Acronyms are: Natural History Museum, London (BMNH); Colección Boliviana de Fauna, La Paz (CBF); Field Museum of Natural History, Chicago (FMNH); Laboratory of Vertebrates, Universidade Federal de Ouro Preto (LZV-UFOP) with the accession number R; Instituto de Ciencias Biológicas, Escuela Politécnica Nacional, Quito (MEPN); Museu Nacional, Universidade Federal do Rio de Janeiro (MN); Museu de Zoologia, Universidade de Sao Paulo (MZUSP). The tabulated measurements were previously published, except for the MEPN holotype. Sex: Female (F), Male (M), unknown (?)." pageId="6" pageNumber="129">Table 1</tableCitation>
), a bit longer on the nape and shoulders (
<quantity id="815E92FE4F2B8D57FF42F9CAFF0C44A8" box="[152,229,1641,1664]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="129" unit="mm" value="10.0">10 mm</quantity>
) and completely covering the skin. The texture is spiny on the dorsum and belly, over which latter the pelage is shorter (
<quantity id="815E92FE4F2B8D57FF43F90AFF3344E8" box="[153,218,1705,1728]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="129" unit="mm" value="5.0">5 mm</quantity>
). The color on the dorsum, the forehead, back, and sides of body, is Raw Umber (color 23). The muzzle, cheeks, rostrum, and post-auricular patches are paler and Cinnamon (color 39). The lower jaw, the insides of the arms and manus are Clay (color 123 B); a paler color extends towards the belly and continues inside the hind legs and towards the backs of the feet. There are
<specimenCount id="50A0F4924F2B8D57FED8F8CAFE8545A8" box="[258,364,1897,1920]" count="2" pageId="6" pageNumber="129" type="generic" typeStatus="types">two types</specimenCount>
of hair on the body: on the dorsum, the hair is flattened and coarse, like spines, with a transparent keel in the middle, thick edges, as in
<taxonomicName id="81A644984F2B8D57FE35F80AFDB445E8" authorityName="Thomas" authorityYear="1900" box="[495,605,1961,1984]" class="Mammalia" family="Cricetidae" genus="Neacomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="129" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F2B8D57FE35F80AFDB445E8" box="[495,605,1961,1984]" italics="true" pageId="6" pageNumber="129">Neacomys</emphasis>
</taxonomicName>
, but with a smaller hair diameter than the spiny hair of
<taxonomicName id="81A644984F2B8D57FB32FF6AFABF42C8" authorityName="Thomas" authorityYear="1900" box="[1256,1366,201,224]" class="Mammalia" family="Cricetidae" genus="Neacomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="129" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F2B8D57FB32FF6AFABF42C8" box="[1256,1366,201,224]" italics="true" pageId="6" pageNumber="129">Neacomys</emphasis>
</taxonomicName>
. The spiny nature of hair is less pronounced on the shoulders, neck, and head. This condition is probably a function of the specimens age. The color along the edges of the spiny hair of
<taxonomicName id="81A644984F2B8D57FAE5FE8AFA124368" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1343,1531,297,320]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F2B8D57FAE5FE8AFA124368" box="[1343,1531,297,320]" italics="true" pageId="6" pageNumber="129">R. septentrionalis</emphasis>
</taxonomicName>
is Neutral Dark Grey (color 83) along the middle and Blackish Gray neutral (color 82) at the apex, where the transparent keel disappears. The color tone of the spiny hairs on the nape and head is darker.
</paragraph>
<paragraph id="46193F1B4F2B8D57FCA2FE6AFAC74068" blockId="6.[861,1531,201,1984]" pageId="6" pageNumber="129">The second class of hair is thin and curly, with the basal portion Dark Grey neutral color and the apical third of the hair colored in Spectrum Orange (color 17) that resembles the general tone over the dorsum and sides of the body.</paragraph>
<paragraph id="46193F1B4F2B8D57FCA2FDEAFB0540E8" blockId="6.[861,1531,201,1984]" pageId="6" pageNumber="129">The belly with spiny hair, Cream (color 54) at the base and Clay apically from the belly extending to the anus and insides of the legs. The thin hair on the belly is Clay throughout its length and denser than the spiny hair.</paragraph>
<paragraph id="46193F1B4F2B8D57FCA2FD6AFC524168" blockId="6.[861,1531,201,1984]" pageId="6" pageNumber="129">The ears are small, covered inside by inconspicuous silverplated hair, fringed by tiny orangish hairs; externally, the pinnae are blackish, clothed in short brown hairs, becoming whitish at the base.</paragraph>
<paragraph id="46193F1B4F2B8D57FCA2FCEAFAC24628" blockId="6.[861,1531,201,1984]" pageId="6" pageNumber="129">
The eye margins do not have a blackish ring of skin. The longer mystacial vibrissae (Weksler and Percequillo 2011) extend
<quantity id="815E92FE4F2B8D57FC4EFC2AFC224188" box="[916,971,905,928]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="129" unit="cm" value="1.0">1 cm</quantity>
beyond the ear. No superciliary vibrissa is present, but an elongated genal vibrissa exceeds the margins of the ear, there are two interramal vibrissae, and one very small submental vibrissa at the edge of the lower lip.
</paragraph>
<paragraph id="46193F1B4F2B8D57FCA2FBAAFB4A47E8" blockId="6.[861,1531,201,1984]" pageId="6" pageNumber="129">
The tail is longer than the head-and-body, covered by hairs similar in color to the rump for about
<quantity id="815E92FE4F2B8D57FB28FB8AFAD54668" box="[1266,1340,1065,1088]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="129" unit="mm" value="10.0">10 mm</quantity>
from its base and with a little tuft,
<quantity id="815E92FE4F2B8D57FBC1FBEAFBB34648" box="[1051,1114,1097,1120]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="129" unit="mm" value="4.0">4 mm</quantity>
long, at its tip. The scales on the tail form rings around it. The color on the upper part of the tail is Vandyke Brown (color 221) and it is covered by short spiny hairs
<quantity id="815E92FE4F2B8D57FC43FB0AFC0546E8" box="[921,1004,1193,1216]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="129" unit="mm" value="1.2">1.2 mm</quantity>
long of the same color that exceed by 2.5 times the length of the scales in the tail, and are projected in triads from the edge of each scale. The hair in the middle is a little longer than the ones at either side of each scale. Below, the color is paler and for about a third of its length, the tone is Draft (color 27), covered by shorter hairs than those dorsally, fading into Pale Pinkish Buff (color 121D) along the length and tip of each hair. Along the apical third of the tail, the ventral section is as dark as the upper section.
</paragraph>
<paragraph id="46193F1B4F2B8D57FCA2FA6AFABB45E8" blockId="6.[861,1531,201,1984]" pageId="6" pageNumber="129">
The characters of the manus are described from the dry skin. The digits match the description of Luna and Patterson (2003). They are long with round fingertips, with a deep depression in the front end of the apical pads. More proximally, the digits have parallel and transverse grooves, almost invisible as seen on the dry skin. There also are three interdigital pads, one hypothenar, and one thenar (Luna and Patterson 2003), all of them well-defined. The D3 and D4 are almost the same length, the D4 slightly longer. There is a vestigial thumb with a minute nail. The nails of the manus are very small (
<quantity id="815E92FE4F2B8D57FAD1F94AFAA14528" box="[1291,1352,1769,1792]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="129" unit="mm" value="1.0">1 mm</quantity>
length), slightly surpassing the fingertips; the ungual claws are embedded in the fingertips, so they can scarcely be seen in ventral view, where one appreciates that they are concave and sharp at the tips. All the hairs at the base of the claws are transparent and greatly surpass the tips of the claws. Along their dorsal surface, the skin of the hands is covered by short transparent hairs.
</paragraph>
<caption id="12D96F934F2A8D56FFBFFAF6FA97478D" ID-DOI="http://doi.org/10.5281/zenodo.10632401" ID-Zenodo-Dep="10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="7" pageNumber="130" startId="7.[101,136,1365,1386]" targetBox="[114,1475,200,1352]" targetPageId="7" targetType="figure">
<paragraph id="46193F1B4F2A8D56FFBFFAF6FA97478D" blockId="7.[101,1488,1365,1445]" pageId="7" pageNumber="130">
<emphasis id="74D2E3094F2A8D56FFBFFAF6FF4F4742" bold="true" box="[101,166,1365,1386]" pageId="7" pageNumber="130">Fig. 3.</emphasis>
—Skulls of
<taxonomicName id="81A644984F2A8D56FEC7FAF6FE684742" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[285,385,1365,1386]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="130" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F2A8D56FEC7FAF6FE684742" box="[285,385,1365,1386]" italics="true" pageId="7" pageNumber="130">Rhagomys</emphasis>
</taxonomicName>
to the same scale. (
<emphasis id="74D2E3094F2A8D56FD9EFAF6FDBC4742" bold="true" box="[580,597,1365,1386]" pageId="7" pageNumber="130">A</emphasis>
)
<taxonomicName id="81A644984F2A8D56FDBFFAF6FCB64742" authority="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson, 2021" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[613,863,1365,1386]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="130" phylum="Chordata" rank="species" species="septentrionalis" status="sp. nov.">
<emphasis id="74D2E3094F2A8D56FDBFFAF6FCB64742" box="[613,863,1365,1386]" italics="true" pageId="7" pageNumber="130">Rhagomys septentrionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FE15E724F2A8D56FCBCFAF6FC5B4742" box="[870,946,1365,1386]" pageId="7" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
(MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (
<emphasis id="74D2E3094F2A8D56FD69FAD1FD2A47AF" bold="true" box="[691,707,1394,1415]" pageId="7" pageNumber="130">B</emphasis>
)
<taxonomicName id="81A644984F2A8D56FD0BFAD1FC7D47AF" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[721,916,1394,1415]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="130" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F2A8D56FD0BFAD1FC7D47AF" box="[721,916,1394,1415]" italics="true" pageId="7" pageNumber="130">Rhagomys rufescens</emphasis>
</taxonomicName>
(MZUSP 65545), in dorsal and ventral view; (
<emphasis id="74D2E3094F2A8D56FA94FAD1FAB647AF" bold="true" box="[1358,1375,1394,1415]" pageId="7" pageNumber="130">C</emphasis>
)
<taxonomicName id="81A644984F2A8D56FAB6FAD1FF3D478D" authorityName="Luna and Patterson" authorityYear="2003" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="130" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F2A8D56FAB6FAD1FF3D478D" italics="true" pageId="7" pageNumber="130">Rhagomys longilingua</emphasis>
</taxonomicName>
(MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm.
</paragraph>
</caption>
<paragraph id="46193F1B4F2A8D56FF5AFA73FAFA442F" blockId="7.[101,771,1488,1991]" lastBlockId="7.[819,1488,1488,1991]" pageId="7" pageNumber="130">
The pes is blackish dorsally with five digits entirely covered by short orange hairs (Clay colored). As on the manus, pedal digits show transverse ventral grooves and rounded fingertips with apical indentations. The D2, D3, and D4, are the longest toes; the D5 exceeds by more than half the second phalanx of D4. The hallux is medium-sized, with a length of
<quantity id="815E92FE4F2A8D56FD7AF9D3FCEB44AF" box="[672,770,1648,1671]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="7" pageNumber="130" unit="mm" value="3.25">3.25 mm</quantity>
from the union of the first phalanx with the metatarsus, surpassing the lower edge of the second interdigital pad to reach the middle of the first phalanx of D2. The hallux shows a flattened nail on the fingertip that does not appear as a claw. The pes has six plantar pads: four are interdigital, one hypothenar, and one thenar. The hypothenar is smaller than the thenar pad and is located beside the fourth interdigital pad. The claws on the pedal digits are longer than those of the manus, with an approximate length of
<quantity id="815E92FE4F2A8D56FEE6F833FE65458F" box="[316,396,1936,1959]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="130" unit="mm" value="1.9">1.9 mm</quantity>
(except on the hallux) and width of
<quantity id="815E92FE4F2A8D56FFBFF813FF2D45EF" box="[101,196,1968,1991]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="130" unit="mm" value="0.4">0.40 mm</quantity>
(on D3); on all of these, the claws surpass the tips in ventral view. The hair at the base of the claws is transparent, as on the manus, and slightly surpasses the tips.
</paragraph>
<paragraph id="46193F1B4F2A8D56FC97F9B3FA45446F" blockId="7.[819,1488,1488,1991]" pageId="7" pageNumber="130">The soles of the pes are completely naked. However, hair along the margins completely covers the calcaneum region.</paragraph>
<paragraph id="46193F1B4F2A8D56FC97F9F3FA2445EF" blockId="7.[819,1488,1488,1991]" pageId="7" pageNumber="130">
<emphasis id="74D2E3094F2A8D56FC97F9F3FC60444F" box="[845,905,1616,1639]" italics="true" pageId="7" pageNumber="130">Skull.</emphasis>
—The skull is broad and robust. The rostrum is long, narrow, and ends in a tip. The interorbital region is narrow, with a broad, rounded braincase. The zygomatic arches are parallelsided, with a conspicuous jugal bone. The anterior margin of the upper incisors does not reach the anterior margin of the nasals, nor do their tips exceed the anterior edge of the premaxillary. The nasolacrimal capsule joins the premaxillary with an opening to the posterior proximal to the anterior margin of the zygomatic plate. The nasals are elongate, wider anteriorly but narrowed posteriorly to form a “
<collectionCode id="20B7A7DE4F2A8D56FC2CF8D3FBE245AF" box="[1014,1035,1904,1927]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="130" type="Museum">V</collectionCode>
” shape at their frontal suture. The posterior projection of the premaxillary exceeds by
<quantity id="815E92FE4F2A8D56FB39F833FAF7458F" box="[1251,1310,1936,1959]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="130" unit="mm" value="1.0">1 mm</quantity>
the hindmost extent of the nasal bones. The zygomatic notch is shallow (
<figureCitation id="DE9D239E4F2A8D56FA5BF813FA5445EF" box="[1409,1469,1968,1991]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="7" pageNumber="130">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="46193F1B4F258D59FF71FF6AFE0243C8" blockId="8.[144,814,201,1184]" pageId="8" pageNumber="131">
The zygomatic arches converge anteriorly. The zygomatic plate is moderately wide, low, extending
<quantity id="815E92FE4F258D59FDBAFF4AFD2B4328" box="[608,706,233,256]" metricMagnitude="-3" metricUnit="m" metricValue="5.31" pageId="8" pageNumber="131" unit="mm" value="5.31">5.31 mm</quantity>
from the maxillary root of the zygomatic arch to the lower part of the plate. The anterior border of the plate is slightly tilted posteriorly and is totally posterior to the nasolacrimal capsule. The antorbital bridge of the zygomatic maxillary root stands at almost the same height as the tip of the nasals in lateral view. The first maxillary molar (M1) lies at a level just behind the posterior edge of the zygomatic plate.
</paragraph>
<paragraph id="46193F1B4F258D59FF71FE4AFDF54128" blockId="8.[144,814,201,1184]" pageId="8" pageNumber="131">
The interorbital region is wide posteriorly (on the frontals) and convergent anteriorly (towards the lacrimals), appearing hourglass-shaped (
<figureCitation id="DE9D239E4F258D59FE80FD8AFE724068" box="[346,411,553,576]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="8" pageNumber="131">Fig. 3</figureCitation>
). The specimen has faint but noticeable supraorbital cristae. The interparietal is wide and does not contact the squamosal bone. The length of the supraoccipitalparietal suture is longer than the occipito-squamosal suture. The anterior section of the squamosal bone presents a faint postorbital process, visible from a dorsal view of the cranium as a small protrusion inside the orbit.
</paragraph>
<paragraph id="46193F1B4F258D59FF71FCAAFC574368" blockId="8.[144,814,201,1184]" lastBlockId="8.[861,1531,201,1984]" pageId="8" pageNumber="131">
The alisphenoid strut is broad and partially hides the opening of the alisphenoid canal from lateral view. The foramen ovale accessorius is completely visible (
<figureCitation id="DE9D239E4F258D59FDD0FCEAFDA24148" box="[522,587,841,864]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="8" pageNumber="131">Fig. 3</figureCitation>
). The pattern of the carotid circulation resembles pattern 3 (
<bibRefCitation id="223742EA4F258D59FD8AFCCAFD2A41A8" author="VOSS, R. S." box="[592,707,873,896]" pageId="8" pageNumber="131" pagination="259 - 493" refId="ref14810" refString="VOSS, R. S. 1988. Systematics and ecology of ichthyomyine rodents (Muroidea): patterns of morphological evolution in a small adaptive radiation. Bulletin of the American Museum of Natural History 188: 259 - 493." type="journal article" year="1988">Voss 1988</bibRefCitation>
): lacking squamosal-alisphenoidal grooves or a sphenofrontal foramen; in the ear region, without a stapedial foramen, the periotic is hidden by the ectotympanic. The auditory bullae have well-developed carotid canals. Under the basisphenoid, a conspicuous transverse canal is situated at the level of the hamular pterygoid processes (
<figureCitation id="DE9D239E4F258D59FEA2FB8AFE504668" box="[376,441,1065,1088]" captionStart="Fig" captionStartId="8.[144,179,1875,1896]" captionTargetBox="[168,789,1240,1861]" captionTargetId="figure-889@8.[168,789,1240,1861]" captionTargetPageId="8" captionText="Fig. 4.—Basicranium of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype): PPP, posterolateral palatal pits; PPF, parapterygoid fossa; TC, transverse canal; MLF, middle lacerate foramen; BOC, basioccipital. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10632403" httpUri="https://zenodo.org/record/10632403/files/figure.png" pageId="8" pageNumber="131">Fig. 4</figureCitation>
). The posterior region of alisphenoid shows a marked posterior opening of the alisphenoid canal in oval shape, with a small middle lacerate foramen with a laterally widened shape (
<figureCitation id="DE9D239E4F258D59FE61FB2AFE144688" box="[443,509,1161,1184]" captionStart="Fig" captionStartId="8.[144,179,1875,1896]" captionTargetBox="[168,789,1240,1861]" captionTargetId="figure-889@8.[168,789,1240,1861]" captionTargetPageId="8" captionText="Fig. 4.—Basicranium of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype): PPP, posterolateral palatal pits; PPF, parapterygoid fossa; TC, transverse canal; MLF, middle lacerate foramen; BOC, basioccipital. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10632403" httpUri="https://zenodo.org/record/10632403/files/figure.png" pageId="8" pageNumber="131">Fig. 4</figureCitation>
). There is a groove for the infraorbital branch of stapedial artery in the posterior part of the pterygoid plate; this groove is long, running from the posterior opening of the alisphenoid canal into the middle lacerate foramen.
</paragraph>
<paragraph id="46193F1B4F258D59FCA2FEEAFA714008" blockId="8.[861,1531,201,1984]" pageId="8" pageNumber="131">The foramen ovale is similar in size to the posterior opening of the alisphenoidal canal and is visible in ventral view, because the edge of the pterygoid plate located between the posterior opening of the alisphenoidal canal and the foramen ovale is flattened in the cranium of this genus and does not project outside it, as happens in other cricetids in which the foramen ovale is hidden ventrally by the edge of the pterygoid plate.</paragraph>
<paragraph id="46193F1B4F258D59FCA2FD8AFAAD4608" blockId="8.[861,1531,201,1984]" pageId="8" pageNumber="131">
The parapterygoid fossa is shallow and wide (
<figureCitation id="DE9D239E4F258D59FAA8FD8AFA5B4068" box="[1394,1458,553,576]" captionStart="Fig" captionStartId="8.[144,179,1875,1896]" captionTargetBox="[168,789,1240,1861]" captionTargetId="figure-889@8.[168,789,1240,1861]" captionTargetPageId="8" captionText="Fig. 4.—Basicranium of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype): PPP, posterolateral palatal pits; PPF, parapterygoid fossa; TC, transverse canal; MLF, middle lacerate foramen; BOC, basioccipital. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10632403" httpUri="https://zenodo.org/record/10632403/files/figure.png" pageId="8" pageNumber="131">Fig. 4</figureCitation>
), with a prominent pit. These pits are round, with thin and irregular edges. The tegmen tympani or petrosal overlaps slightly the posterior suspensory process of the squamosal bone. The squamosal hamular process is wide and widens posteriorly to contact the mastoid and the ectotympanic. The mastoid fenestra lies between the mastoid periotic suture and the exoccipital and is less conspicuous and elongated, located at the left side of the cranium. Another fenestra is located between the mastoid and the exoccipital over the paraoccipital process. The ectotympanic ends near the pterygoid hamulars in a short, wide eustachian tube. The carotid canal is small and situated near the end of the eustachian tube. The basioccipital surface is almost flat, and there is a small but noticeable median crista that extends from the anteroventral edge of the magnum foramen towards the anterior edge of the basioccipital.
</paragraph>
<paragraph id="46193F1B4F258D59FCA2FB8AFACC4728" blockId="8.[861,1531,201,1984]" pageId="8" pageNumber="131">The subsquamosal fenestra and the postglenoid foramen are well-developed. These foramina are shaped like parentheses, so they narrow and converge at the anterior and posterior margins. The premaxillary and maxillary suture ventrally bisects the incisive foramen. The septum of the incisive foramen occupies almost half of its total length. The masseteric tubercle is a small projection in the maxillary, anterior to M1.</paragraph>
<paragraph id="46193F1B4F258D59FCA2FAAAFA424548" blockId="8.[861,1531,201,1984]" pageId="8" pageNumber="131">
The palate is large, with the posterior margin that extends beyond the M3 roughly the length of that tooth (
<quantity id="815E92FE4F258D59FA86FA8AFA294768" box="[1372,1472,1321,1344]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="8" pageNumber="131" unit="mm" value="0.52">0.52 mm</quantity>
). The posterior margin of the palate ends at a wide mesopterygoid fossa without a medial process or spine (
<figureCitation id="DE9D239E4F258D59FACFFACAFABD47A8" box="[1301,1364,1385,1408]" captionStart="Fig" captionStartId="8.[144,179,1875,1896]" captionTargetBox="[168,789,1240,1861]" captionTargetId="figure-889@8.[168,789,1240,1861]" captionTargetPageId="8" captionText="Fig. 4.—Basicranium of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype): PPP, posterolateral palatal pits; PPF, parapterygoid fossa; TC, transverse canal; MLF, middle lacerate foramen; BOC, basioccipital. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10632403" httpUri="https://zenodo.org/record/10632403/files/figure.png" pageId="8" pageNumber="131">Fig. 4</figureCitation>
). The suture of the anterior part of the palatine with the maxillary lies at the level of the M1 hypocone, and its posterior sutures with the pterygoids lies at half the height of the parapterygoid fossa. The posterolateral palatal pits are very conspicuous in a defined fossa, containing two foramina at the right side and one at the left of the palatine (
<figureCitation id="DE9D239E4F258D59FBB7F98AFB424468" box="[1133,1195,1577,1600]" captionStart="Fig" captionStartId="8.[144,179,1875,1896]" captionTargetBox="[168,789,1240,1861]" captionTargetId="figure-889@8.[168,789,1240,1861]" captionTargetPageId="8" captionText="Fig. 4.—Basicranium of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype): PPP, posterolateral palatal pits; PPF, parapterygoid fossa; TC, transverse canal; MLF, middle lacerate foramen; BOC, basioccipital. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10632403" httpUri="https://zenodo.org/record/10632403/files/figure.png" pageId="8" pageNumber="131">Fig. 4</figureCitation>
). The posterolateral palatal pits surpass the posterior edge of the palatine and extend for a sixth part of its length into the mesopterygoid foramen. The dental maxillary rows are sub-parallel because they converge towards the anterior. The anterior margin of the mesopterygoid foramen presents square angles. In the roof of the mesopterygoid foramen, two small sphenopalatine vacuities are visible in the leading edges of the basisphenoids. The pterygoid hamular processes were slightly damaged during preparation, but they appear to diverge slightly towards the eustachian tubes.
</paragraph>
<caption id="12D96F934F258D59FF4AF8F0FE2F45E8" ID-DOI="http://doi.org/10.5281/zenodo.10632403" ID-Zenodo-Dep="10632403" httpUri="https://zenodo.org/record/10632403/files/figure.png" pageId="8" pageNumber="131" startId="8.[144,179,1875,1896]" targetBox="[168,789,1240,1861]" targetPageId="8" targetType="figure">
<paragraph id="46193F1B4F258D59FF4AF8F0FE2F45E8" blockId="8.[144,814,1875,1984]" pageId="8" pageNumber="131">
<emphasis id="74D2E3094F258D59FF4AF8F0FF3B4540" bold="true" box="[144,210,1875,1896]" pageId="8" pageNumber="131">Fig. 4.</emphasis>
—Basicranium of
<taxonomicName id="81A644984F258D59FE53F8F0FD6C4540" authority="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson, 2021" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[393,645,1875,1896]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="131" phylum="Chordata" rank="species" species="septentrionalis" status="sp. nov.">
<emphasis id="74D2E3094F258D59FE53F8F0FD6C4540" box="[393,645,1875,1896]" italics="true" pageId="8" pageNumber="131">Rhagomys septentrionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FE15E724F258D59FD57F8F0FD324540" box="[653,731,1875,1896]" pageId="8" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
(MEPN 10898, holotype): PPP, posterolateral palatal pits; PPF, parapterygoid fossa; TC, transverse canal; MLF, middle lacerate foramen; BOC, basioccipital. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="46193F1B4F258D58FCA2F8CAFF344068" blockId="8.[861,1531,201,1984]" lastBlockId="9.[101,772,201,1152]" lastPageId="9" lastPageNumber="132" pageId="8" pageNumber="131">
<emphasis id="74D2E3094F258D59FCA2F8CAFC0C45A8" box="[888,997,1897,1920]" italics="true" pageId="8" pageNumber="131">Mandible.</emphasis>
—The angular process is short and sharp (
<figureCitation id="DE9D239E4F258D59FA76F8CAFA0245A8" box="[1452,1515,1897,1920]" captionStart="Fig" captionStartId="9.[101,136,1893,1914]" captionTargetBox="[126,747,1216,1880]" captionTargetId="figure-6@9.[126,747,1216,1880]" captionTargetPageId="9" captionText="Fig. 5.—Lateral view of the mandibles of Rhagomys: (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype); (B) R. rufescens (MZUSP AB401; from Pavan and Leite 2011); and (C) R. longilingua (MUSM 20009). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632405" httpUri="https://zenodo.org/record/10632405/files/figure.png" pageId="8" pageNumber="131">Fig. 5</figureCitation>
). The coronoid process is short, with an anterior border straight up, that does not reach the level of articular condyle. The articular condyle is enlarged, narrow, oriented behind and with the same thickness as the neck of the condyloid. The angular notch of the mandible is shallow. The capsular process of the lower incisor is very conspicuous and visible in dorsal view. In lateral view, it is located at the midpoint of the mandibular notch. There is a conspicuous lower masseteric crest, whose anterior extent surpasses the anterior cingulum of m1. There is a wide and deep retromolar fossa, whose anterior margin reaches the entoconid of the m2, and its posterior margin is at the level of the posterior part of the articular process. There also is a well-developed digastric apophysis at the mandibular symphysis.
</paragraph>
<paragraph id="46193F1B4F248D58FF5AFDEAFC304328" blockId="9.[101,772,201,1152]" lastBlockId="9.[819,1488,201,1024]" pageId="9" pageNumber="132">
<emphasis id="74D2E3094F248D58FF5AFDEAFF054048" box="[128,236,585,608]" italics="true" pageId="9" pageNumber="132">Dentition.</emphasis>
—The incisors are orthodont and narrow, with a fine, straight fissure in the dentine that is visible in posterior view as a black line on the incisors. The molars are pentalophodont, containing a paracone, protocone, metacone, hypocone, and mesoloph (
<figureCitation id="DE9D239E4F248D58FF3BFD6AFECB40C8" box="[225,290,713,736]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="9" pageNumber="132">Fig. 6</figureCitation>
). The molars are low crowned, with cusps similar to tubercles (bunodont) that are very noticeable. These cusps are narrow and oriented posteriorly on the upper molars; the flexi and flexids are wide and well-defined. The paracone and metacone of the M1 and M2 are not at the same level with the protocone and hypocone but rather are staggered behind them. The same occurs with the conids of the lower molars, where the protoconid and hypoconid are situated posterior to the metaconid and entoconid, respectively. M1 is longer than M2, M2 is longer than wide, and M3 is rounded and half the length of M2. The paraflexus and mesoflexus completely encircle the paracone, because posterior to the paracone, the paraflexus connects with the mesoflexus, but there is no connection between the paracone and the median mure (
<figureCitation id="DE9D239E4F248D58FD68FBCAFD1A46A8" box="[690,755,1129,1152]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="9" pageNumber="132">Fig. 6</figureCitation>
). The metacone is almost completely encircled by the metaflexus on M1 and M2.
</paragraph>
<paragraph id="46193F1B4F248D5BFC97FEAAFEF643A8" blockId="9.[819,1488,201,1024]" lastBlockId="10.[144,814,201,2016]" lastPageId="10" lastPageNumber="133" pageId="9" pageNumber="132">
The anteromedial flexus is present but short and shallow, located between a well-developed anterolabial conule and a minute, inconspicuous anterolingual conule (
<figureCitation id="DE9D239E4F248D58FAEAFEEAFA9C4348" box="[1328,1397,329,352]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="9" pageNumber="132">Fig. 6</figureCitation>
). There is no internal projection of the enamel on the anterolabial conule of M1 designated by Luna and Patterson (2003) as the anteromedial crista. The anteroloph adheres to the posterior part of the anterolabial conule. A small crista extends from the posterior part of the anterolingual conule to connect it with the anterior mure. On the M1 and M2, the anterior mure connects directly with the anterior margin of the protocone; on the protocones posterior margin, it shapes the median mure. This mure extends medially towards the mesoloph on both anterior molars. The anterolabial conule does not give any sign of an anterolophule or of a connection with the anteroloph, because the anterolabial conule and anteroloph are fused and reach the labial edge of M1, where there is a conspicuous parastyle. The paracones of M1 and M2 do not have an additional connection with the median mure and are isolated from the other molar cones (
<figureCitation id="DE9D239E4F248D58FCA5FCEAFC574148" box="[895,958,841,864]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="9" pageNumber="132">Fig. 6</figureCitation>
). The anteroloph of M2 is fused labially with the anterior edge of the paracone that extends postero-lingually to form a shallow protoflexus. A posteroflexus is lacking on M1 and M2, but the posteroloph joins with the posterior part of the metacone. A long, conspicuous mesoloph is fused with a small mesostyle on M1 and M2. M3 is rounded and has an anterior mure that extends from the posterior edge of the anteroloph towards the anterior margin of the protocone and turns obliquely as a median mure from the posterior margin of the protocone to the posterior portion of the molar, where there is a structure that may be the hypocone. There also is a paraflexus anterior to the paracone.
</paragraph>
<caption id="12D96F934F248D58FFBFF8C6FE2045FA" ID-DOI="http://doi.org/10.5281/zenodo.10632405" ID-Zenodo-Dep="10632405" httpUri="https://zenodo.org/record/10632405/files/figure.png" pageId="9" pageNumber="132" startId="9.[101,136,1893,1914]" targetBox="[126,747,1216,1880]" targetPageId="9" targetType="figure">
<paragraph id="46193F1B4F248D58FFBFF8C6FE2045FA" blockId="9.[101,771,1893,2002]" pageId="9" pageNumber="132">
<emphasis id="74D2E3094F248D58FFBFF8C6FF4F4552" bold="true" box="[101,166,1893,1914]" pageId="9" pageNumber="132">Fig. 5.</emphasis>
—Lateral view of the mandibles of
<taxonomicName id="81A644984F248D58FDDEF8C6FD814552" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[516,616,1893,1914]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="132" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F248D58FDDEF8C6FD814552" box="[516,616,1893,1914]" italics="true" pageId="9" pageNumber="132">Rhagomys</emphasis>
</taxonomicName>
: (
<emphasis id="74D2E3094F248D58FDA4F8C6FD664552" bold="true" box="[638,655,1893,1914]" pageId="9" pageNumber="132">A</emphasis>
)
<taxonomicName id="81A644984F248D58FD45F8C6FF1D45B0" authority="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson, 2021" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="132" phylum="Chordata" rank="species" species="septentrionalis" status="sp. nov.">
<emphasis id="74D2E3094F248D58FD45F8C6FF1D45B0" italics="true" pageId="9" pageNumber="132">Rhagomys septentrionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FE15E724F248D58FF24F820FEA545B0" box="[254,332,1923,1944]" pageId="9" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
(MEPN 10898, holotype); (
<emphasis id="74D2E3094F248D58FDB2F820FD9145B0" bold="true" box="[616,632,1923,1944]" pageId="9" pageNumber="132">B</emphasis>
)
<taxonomicName id="81A644984F248D58FD50F820FCEA45B0" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[650,771,1923,1944]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="132" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F248D58FD50F820FCEA45B0" box="[650,771,1923,1944]" italics="true" pageId="9" pageNumber="132">R. rufescens</emphasis>
</taxonomicName>
(MZUSP AB401; from Pavan and Leite 2011); and (
<emphasis id="74D2E3094F248D58FD81F803FD85459D" bold="true" box="[603,620,1952,1973]" pageId="9" pageNumber="132">C</emphasis>
)
<taxonomicName id="81A644984F248D58FDA0F803FCEA459D" authorityName="Luna and Patterson" authorityYear="2003" box="[634,771,1952,1973]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="132" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F248D58FDA0F803FCEA459D" box="[634,771,1952,1973]" italics="true" pageId="9" pageNumber="132">R. longilingua</emphasis>
</taxonomicName>
(MUSM 20009). Scale bar = 10 mm.
</paragraph>
</caption>
<caption id="12D96F934F248D58FCE9F820FBD845FA" ID-DOI="http://doi.org/10.5281/zenodo.10632407" ID-Zenodo-Dep="10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="9" pageNumber="132" startId="9.[819,854,1923,1944]" targetBox="[843,1464,1082,1910]" targetPageId="9" targetType="figure">
<paragraph id="46193F1B4F248D58FCE9F820FBD845FA" blockId="9.[819,1488,1923,2002]" pageId="9" pageNumber="132">
<emphasis id="74D2E3094F248D58FCE9F820FC9B45B0" bold="true" box="[819,882,1923,1944]" pageId="9" pageNumber="132">Fig. 6.</emphasis>
—Molars of
<taxonomicName id="81A644984F248D58FC34F820FB0E45B0" authority="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson, 2021" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1006,1255,1923,1944]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="132" phylum="Chordata" rank="species" species="septentrionalis" status="sp. nov.">
<emphasis id="74D2E3094F248D58FC34F820FB0E45B0" box="[1006,1255,1923,1944]" italics="true" pageId="9" pageNumber="132">Rhagomys septentrionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FE15E724F248D58FB37F820FAD145B0" box="[1261,1336,1923,1944]" pageId="9" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
(MEPN 10898, holotype). (
<emphasis id="74D2E3094F248D58FC7EF803FC5C459D" bold="true" box="[932,949,1952,1973]" pageId="9" pageNumber="132">A</emphasis>
) Occlusal view of upper right molars; (
<emphasis id="74D2E3094F248D58FAE7F803FAA4459D" bold="true" box="[1341,1357,1952,1973]" pageId="9" pageNumber="132">B</emphasis>
) Lower right molars. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="46193F1B4F278D5BFF71FE2AFE9D4128" blockId="10.[144,814,201,2016]" pageId="10" pageNumber="133">
The procingulum of m1 presents the anterolingual conulid in the most anterior part of m1 that extends near the labial end of the procingulum (
<figureCitation id="DE9D239E4F278D5BFEA1FE6AFE5443C8" box="[379,445,457,480]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="10" pageNumber="133">Fig. 6</figureCitation>
). The m1 lacks an anteromedial flexid. However, a protolophid is present and it ends in a conspicuous protostylid. The anterior murid is conspicuous and connects to the anterior margin of the protoconid, continuing from the posterior margin of the protoconid towards the median muride. There is no anterolophid. The mesolophid is fused with a conspicuous mesostylid at the lingual edge. The entolophulid is fused with the trailing edge of the mesolophid. The posterolophid is long and reaches the posterior margin of the lingual cingulum.
</paragraph>
<paragraph id="46193F1B4F278D5BFF71FCAAFE6441C8" blockId="10.[144,814,201,2016]" pageId="10" pageNumber="133">The entoflexid and posteroflexid join to encircle the entoconid almost completely. The mesolophid on m2 is fused with the anterior edge of the entoconid at its lingual end. The median murid connects with the anterior margin of the hypoconid. There is a short posterocrista (Luna and Patterson 2003) that extends from the posterior part of the hypoconid, reaching the posterior margin of m2.</paragraph>
<paragraph id="46193F1B4F278D5BFF71FC4AFDAC4728" blockId="10.[144,814,201,2016]" pageId="10" pageNumber="133">
The m2 and m3 differ in size and shape. The m3 is subtriangular with well-defined protoconid and metaconid; posteriorly, the hypoconid and entoconid are fused at their internal margins. The m2 and m3 present a mesolophid that on m2 joins with the entoconid, but on m3 does not reach the entoconid and continues towards the lingual cingulum of the molar (
<figureCitation id="DE9D239E4F278D5BFD07FB2AFCF44688" box="[733,797,1161,1184]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="10" pageNumber="133">Fig. 6</figureCitation>
). In addition, the hypoconid of m3 does not connect with the mesolophid to form a median murid and a posterocrista is lacking on the posterior cingulum of m3.
</paragraph>
<paragraph id="46193F1B4F278D5BFF71FAAAFDEE47A8" blockId="10.[144,814,201,2016]" pageId="10" pageNumber="133">
<emphasis id="74D2E3094F278D5BFF71FAAAFEB24708" box="[171,347,1289,1312]" italics="true" pageId="10" pageNumber="133">Comparisons.—</emphasis>
<taxonomicName id="81A644984F278D5BFE80FAAAFDF44708" box="[346,541,1289,1312]" pageId="10" pageNumber="133">
<emphasis id="74D2E3094F278D5BFE80FAAAFE204708" box="[346,457,1289,1312]" italics="true" pageId="10" pageNumber="133">Rhagomys</emphasis>
species
</taxonomicName>
are very similar in color, but the spiny coat of
<taxonomicName id="81A644984F278D5BFE51FA8AFE134768" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[395,506,1321,1344]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F278D5BFE51FA8AFE134768" box="[395,506,1321,1344]" italics="true" pageId="10" pageNumber="133">Rhagomys</emphasis>
</taxonomicName>
in the Andean foothills of
<collectingCountry id="3EB17F8B4F278D5BFF4AFAEAFF024748" box="[144,235,1353,1376]" name="Ecuador" pageId="10" pageNumber="133">Ecuador</collectingCountry>
,
<collectingCountry id="3EB17F8B4F278D5BFF20FAEAFEC74748" box="[250,302,1353,1376]" name="Peru" pageId="10" pageNumber="133">Peru</collectingCountry>
, and
<collectingCountry id="3EB17F8B4F278D5BFEB5FAEAFE294748" box="[367,448,1353,1376]" name="Bolivia" pageId="10" pageNumber="133">Bolivia</collectingCountry>
, clearly distinguishes them from
<taxonomicName id="81A644984F278D5BFF4AFACAFEFA47A8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[144,275,1385,1408]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F278D5BFF4AFACAFEFA47A8" box="[144,275,1385,1408]" italics="true" pageId="10" pageNumber="133">R. rufescens</emphasis>
</taxonomicName>
, which has a soft coat.
</paragraph>
<paragraph id="46193F1B4F278D5BFF71FA2AFD384468" blockId="10.[144,814,201,2016]" pageId="10" pageNumber="133">
<emphasis id="74D2E3094F278D5BFF71FA2AFE894788" box="[171,352,1417,1440]" italics="true" pageId="10" pageNumber="133">Comparisonwith</emphasis>
<taxonomicName id="81A644984F278D5BFEB8FA2AFE1F4788" authorityName="Luna and Patterson" authorityYear="2003" box="[354,502,1417,1440]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">R.longilingua</taxonomicName>
.
<emphasis id="74D2E3094F278D5BFE27FA2AFCC44788" box="[509,813,1417,1440]" italics="true" pageId="10" pageNumber="133">―Rhagomysseptentrionalis</emphasis>
is similar in size to
<taxonomicName id="81A644984F278D5BFEBAFA0AFE1047E8" authorityName="Luna and Patterson" authorityYear="2003" box="[352,505,1449,1472]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFEBAFA0AFE1047E8" box="[352,505,1449,1472]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
(
<tableCitation id="0B240AA04F278D5BFDD3FA0AFDBE47E8" box="[521,599,1449,1472]" captionStart="Table 1" captionStartId="4.[126,182,506,527]" captionTargetPageId="4" captionText="Table 1.—External, vibrissal, and craniodental measurements (mm), and age class, for specimens of Rhagomys. Acronyms are: Natural History Museum, London (BMNH); Colección Boliviana de Fauna, La Paz (CBF); Field Museum of Natural History, Chicago (FMNH); Laboratory of Vertebrates, Universidade Federal de Ouro Preto (LZV-UFOP) with the accession number R; Instituto de Ciencias Biológicas, Escuela Politécnica Nacional, Quito (MEPN); Museu Nacional, Universidade Federal do Rio de Janeiro (MN); Museu de Zoologia, Universidade de Sao Paulo (MZUSP). The tabulated measurements were previously published, except for the MEPN holotype. Sex: Female (F), Male (M), unknown (?)." pageId="10" pageNumber="133">Table 1</tableCitation>
). Both species have spine-like hairs on the dorsum and belly. In both species, the length of the tail (individuals between ages 2 and 4) is almost the same as the length of the head-and-body, uniform in color, and covered with short hairs that are longer on the tip.
</paragraph>
<paragraph id="46193F1B4F278D5BFF71F9EAFBF34668" blockId="10.[144,814,201,2016]" lastBlockId="10.[861,1531,201,2016]" pageId="10" pageNumber="133">
The cranium of
<taxonomicName id="81A644984F278D5BFEB9F9EAFDCD4448" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[355,548,1609,1632]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFEB9F9EAFDCD4448" box="[355,548,1609,1632]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
is rounded but is more oval in
<taxonomicName id="81A644984F278D5BFF3BF9CAFE9244A8" authorityName="Luna and Patterson" authorityYear="2003" box="[225,379,1641,1664]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFF3BF9CAFE9244A8" box="[225,379,1641,1664]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
. The orbital region of
<taxonomicName id="81A644984F278D5BFDAAF9CAFCC744A8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[624,814,1641,1664]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFDAAF9CAFCC744A8" box="[624,814,1641,1664]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
is narrower anteriorly, as the maxillary root of the zygomatic arch hardly projects laterally, whereas in
<taxonomicName id="81A644984F278D5BFD9FF90AFD3744E8" authorityName="Luna and Patterson" authorityYear="2003" box="[581,734,1705,1728]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFD9FF90AFD3744E8" box="[581,734,1705,1728]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
the anterior part of the orbit is wider (
<figureCitation id="DE9D239E4F278D5BFE3CF96AFDCC44C8" box="[486,549,1737,1760]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="10" pageNumber="133">Fig. 3</figureCitation>
). The interorbital region in
<taxonomicName id="81A644984F278D5BFF74F94AFEA14528" authorityName="Luna and Patterson" authorityYear="2003" box="[174,328,1769,1792]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFF74F94AFEA14528" box="[174,328,1769,1792]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
is wide and converges anteriorly into a “
<collectionCode id="20B7A7DE4F278D5BFCD7F94AFCCA4528" box="[781,803,1769,1792]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="133" type="Museum">V</collectionCode>
” shape, with its narrowest point just behind the lacrimal bones. In contrast, in
<taxonomicName id="81A644984F278D5BFEF7F88AFE034568" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[301,490,1833,1856]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFEF7F88AFE034568" box="[301,490,1833,1856]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
, the interorbital region is narrowest more posteriorly and flares anteriorly towards the lacrimals, so that the region appears hourglass-shaped.
<taxonomicName id="81A644984F278D5BFD65F8CAFEC64588" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFD65F8CAFEC64588" italics="true" pageId="10" pageNumber="133">Rhagomys septentrionalis</emphasis>
</taxonomicName>
has inconspicuous supraorbital cristae, unlike those of
<taxonomicName id="81A644984F278D5BFEFAF80AFE5245E8" authorityName="Luna and Patterson" authorityYear="2003" box="[288,443,1961,1984]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFEFAF80AFE5245E8" box="[288,443,1961,1984]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
, which are evident even in young specimens (
<bibRefCitation id="223742EA4F278D5BFEC9F86AFDFB45C8" author="VILLALPANDO, G. &amp; J. VARGAS &amp; J. SALAZAR-BRAVO" box="[275,530,1993,2016]" pageId="10" pageNumber="133" pagination="143 - 149" refId="ref14774" refString="VILLALPANDO, G., J. VARGAS, AND J. SALAZAR-BRAVO. 2006. First record of Rhagomys (Mammalia: Sigmodontinae) in Bolivia. Mastozoologia Neotropical 13: 143 - 149." type="journal article" year="2006">Villalpando et al. 2006</bibRefCitation>
). The postorbital process in
<taxonomicName id="81A644984F278D5BFCA7FF6AFBD742C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[893,1086,201,224]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFCA7FF6AFBD742C8" box="[893,1086,201,224]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
is shorter and less conspicuous than in
<taxonomicName id="81A644984F278D5BFC87FF4AFC114328" authorityName="Luna and Patterson" authorityYear="2003" box="[861,1016,233,256]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFC87FF4AFC114328" box="[861,1016,233,256]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
. Relative to skull length,
<taxonomicName id="81A644984F278D5BFACBFF4AFA264328" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1297,1487,233,256]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFACBFF4AFA264328" box="[1297,1487,233,256]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
has a larger interparietal bone than
<taxonomicName id="81A644984F278D5BFB10FEAAFA804308" authorityName="Luna and Patterson" authorityYear="2003" box="[1226,1385,265,288]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFB10FEAAFA804308" box="[1226,1385,265,288]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
viewed dorsally. The rostrum in
<taxonomicName id="81A644984F278D5BFB8FFE8AFAFE4368" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1109,1303,297,320]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFB8FFE8AFAFE4368" box="[1109,1303,297,320]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
is narrower than in
<taxonomicName id="81A644984F278D5BFC87FEEAFC154348" authorityName="Luna and Patterson" authorityYear="2003" box="[861,1020,329,352]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFC87FEEAFC154348" box="[861,1020,329,352]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
, especially at the anterior nasal-premaxillary suture. Relative to cranial length, the rostrum is slightly longer in
<taxonomicName id="81A644984F278D5BFC0EFE2AFB704388" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[980,1177,393,416]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFC0EFE2AFB704388" box="[980,1177,393,416]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
(28%) than in
<taxonomicName id="81A644984F278D5BFA80FE2AFA124388" authorityName="Luna and Patterson" authorityYear="2003" box="[1370,1531,393,416]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFA80FE2AFA124388" box="[1370,1531,393,416]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
(26%). The diastema in
<taxonomicName id="81A644984F278D5BFBAFFE0AFADE43E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1141,1335,425,448]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFBAFFE0AFADE43E8" box="[1141,1335,425,448]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
also is enlarged, with the procingulum of M1 located just behind the posterior margin of the zygomatic plate, whereas in
<taxonomicName id="81A644984F278D5BFAEAFE4AFA224028" authorityName="Luna and Patterson" authorityYear="2003" box="[1328,1483,489,512]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFAEAFE4AFA224028" box="[1328,1483,489,512]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
, the procingulum of M1 lies far behind the posterior margin of the zygomatic plate (by the length of M2). Its incisive foramen is enlarged (
<collectionCode id="20B7A7DE4F278D5BFC3FFDEAFBE64048" box="[997,1039,585,608]" pageId="10" pageNumber="133">IFL</collectionCode>
=
<quantity id="815E92FE4F278D5BFBEAFDEAFB6F4048" box="[1072,1158,585,608]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="133" unit="mm" value="4.2">4.2 mm</quantity>
) relative to that of
<taxonomicName id="81A644984F278D5BFA85FDEAFA124048" authorityName="Luna and Patterson" authorityYear="2003" box="[1375,1531,585,608]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFA85FDEAFA124048" box="[1375,1531,585,608]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
(
<collectionCode id="20B7A7DE4F278D5BFCBFFDCAFC6640A8" box="[869,911,617,640]" pageId="10" pageNumber="133">IFL</collectionCode>
=
<quantity id="815E92FE4F278D5BFC74FDCAFBFA40A8" box="[942,1043,617,640]" metricMagnitude="-3" metricUnit="m" metricValue="3.76" pageId="10" pageNumber="133" unit="mm" value="3.76">3.76 mm</quantity>
;
<tableCitation id="0B240AA04F278D5BFBC5FDCAFB8640A8" box="[1055,1135,617,640]" captionStart="Table 1" captionStartId="4.[126,182,506,527]" captionTargetPageId="4" captionText="Table 1.—External, vibrissal, and craniodental measurements (mm), and age class, for specimens of Rhagomys. Acronyms are: Natural History Museum, London (BMNH); Colección Boliviana de Fauna, La Paz (CBF); Field Museum of Natural History, Chicago (FMNH); Laboratory of Vertebrates, Universidade Federal de Ouro Preto (LZV-UFOP) with the accession number R; Instituto de Ciencias Biológicas, Escuela Politécnica Nacional, Quito (MEPN); Museu Nacional, Universidade Federal do Rio de Janeiro (MN); Museu de Zoologia, Universidade de Sao Paulo (MZUSP). The tabulated measurements were previously published, except for the MEPN holotype. Sex: Female (F), Male (M), unknown (?)." pageId="10" pageNumber="133">Table 1</tableCitation>
). However, its palate is shorter than in
<taxonomicName id="81A644984F278D5BFCA7FD2AFBF04088" authorityName="Luna and Patterson" authorityYear="2003" box="[893,1049,649,672]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFCA7FD2AFBF04088" box="[893,1049,649,672]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
because the incisive foramina reach posteriorly to the procingulum of M1 (
<figureCitation id="DE9D239E4F278D5BFB02FD0AFAF040E8" box="[1240,1305,681,704]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="10" pageNumber="133">Fig. 3</figureCitation>
). The parapterygoid fossae in
<taxonomicName id="81A644984F278D5BFC19FD6AFB6940C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[963,1152,713,736]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFC19FD6AFB6940C8" box="[963,1152,713,736]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
are shallow and the fontanelles are rounded, while in
<taxonomicName id="81A644984F278D5BFBFFFD4AFB294128" authorityName="Luna and Patterson" authorityYear="2003" box="[1061,1216,745,768]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFBFFFD4AFB294128" box="[1061,1216,745,768]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
the parapterygoid fossae are deeper and the fontanelles are ovate. The posterior opening of the alisphenoid canal lies beside the oval foramen, because the posterior part of the pterygoid plate, which separates these two foramina in ventral view in all
<taxonomicName id="81A644984F278D5BFB64FCCAFAC441A8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1214,1325,873,896]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F278D5BFB64FCCAFAC441A8" box="[1214,1325,873,896]" italics="true" pageId="10" pageNumber="133">Rhagomys</emphasis>
</taxonomicName>
, is not as wide in
<taxonomicName id="81A644984F278D5BFC87FC2AFBF44188" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[861,1053,905,928]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFC87FC2AFBF44188" box="[861,1053,905,928]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
as in
<taxonomicName id="81A644984F278D5BFBBFFC2AFAE84188" authorityName="Luna and Patterson" authorityYear="2003" box="[1125,1281,905,928]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFBBFFC2AFAE84188" box="[1125,1281,905,928]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
. The auditory bulla in
<taxonomicName id="81A644984F278D5BFC87FC0AFBF341E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[861,1050,937,960]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFC87FC0AFBF341E8" box="[861,1050,937,960]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
is larger than in
<taxonomicName id="81A644984F278D5BFB17FC0AFA8F41E8" authorityName="Luna and Patterson" authorityYear="2003" box="[1229,1382,937,960]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFB17FC0AFA8F41E8" box="[1229,1382,937,960]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
, but its eustachian tubes are shorter, even compared with young specimens of
<taxonomicName id="81A644984F278D5BFC5AFC4AFAAD4628" authority="(Villalpando et al. 2006)" baseAuthorityName="Villalpando" baseAuthorityYear="2006" box="[896,1348,1001,1024]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFC5AFC4AFBF64628" box="[896,1055,1001,1024]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
(
<bibRefCitation id="223742EA4F278D5BFBEEFC4AFAD14628" author="VILLALPANDO, G. &amp; J. VARGAS &amp; J. SALAZAR-BRAVO" box="[1076,1336,1001,1024]" pageId="10" pageNumber="133" pagination="143 - 149" refId="ref14774" refString="VILLALPANDO, G., J. VARGAS, AND J. SALAZAR-BRAVO. 2006. First record of Rhagomys (Mammalia: Sigmodontinae) in Bolivia. Mastozoologia Neotropical 13: 143 - 149." type="journal article" year="2006">Villalpando et al. 2006</bibRefCitation>
)
</taxonomicName>
. The eustachian tubes are wide and short and located near the hamular process of the pterygoids.
</paragraph>
<paragraph id="46193F1B4F278D5BFCA2FBEAFC214708" blockId="10.[861,1531,201,2016]" pageId="10" pageNumber="133">
In lateral view, the cranium of
<taxonomicName id="81A644984F278D5BFB11FBEAFA634648" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1227,1418,1097,1120]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFB11FBEAFA634648" box="[1227,1418,1097,1120]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
exhibits a smaller postglenoid foramen than even young specimens of
<taxonomicName id="81A644984F278D5BFC87FB2AFC1F4688" box="[861,1014,1161,1184]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFC87FB2AFC1F4688" box="[861,1014,1161,1184]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
(
<figureCitation id="DE9D239E4F278D5BFBDFFB2AFBAA4688" box="[1029,1091,1161,1184]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="10" pageNumber="133">Fig. 3</figureCitation>
). The supraorbital crest in
<taxonomicName id="81A644984F278D5BFAB8FB2AFA124688" box="[1378,1531,1161,1184]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFAB8FB2AFA124688" box="[1378,1531,1161,1184]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
can be seen in lateral view of the cranium (including on young specimens), especially along the frontals where it arises on its anterior edge, but in
<taxonomicName id="81A644984F278D5BFBE4FB4AFB154728" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1086,1276,1257,1280]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFBE4FB4AFB154728" box="[1086,1276,1257,1280]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
this crest is not seen in side view.
</paragraph>
<paragraph id="46193F1B4F278D5BFCA2FA8AFB0947C8" blockId="10.[861,1531,201,2016]" pageId="10" pageNumber="133">
The maxillary cusps in the molars of
<taxonomicName id="81A644984F278D5BFAEDFA8AFA124768" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1335,1531,1321,1344]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFAEDFA8AFA124768" box="[1335,1531,1321,1344]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
are oriented posteriorly, while the cusps of the lower molars are oriented forward more than in
<taxonomicName id="81A644984F278D5BFB38FACAFA9647A8" box="[1250,1407,1385,1408]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFB38FACAFA9647A8" box="[1250,1407,1385,1408]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
. The maxillary toothrow in
<taxonomicName id="81A644984F278D5BFBF2FA2AFB014788" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1064,1256,1417,1440]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFBF2FA2AFB014788" box="[1064,1256,1417,1440]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
is similar in size to that of
<taxonomicName id="81A644984F278D5BFCA5FA0AFBF447E8" box="[895,1053,1449,1472]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFCA5FA0AFBF447E8" box="[895,1053,1449,1472]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
, although the toothrows of
<taxonomicName id="81A644984F278D5BFA87FA0AFA1247E8" box="[1373,1531,1449,1472]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFA87FA0AFA1247E8" box="[1373,1531,1449,1472]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
slightly converge anteriorly (
<figureCitation id="DE9D239E4F278D5BFB48FA6AFB3947C8" box="[1170,1232,1481,1504]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="10" pageNumber="133">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="46193F1B4F278D5AFCA2FA4AFD4E4028" blockId="10.[861,1531,201,2016]" lastBlockId="11.[101,772,201,2016]" lastPageId="11" lastPageNumber="134" pageId="10" pageNumber="133">
The M1 of
<taxonomicName id="81A644984F278D5BFBD3FA4AFB244428" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1033,1229,1513,1536]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFBD3FA4AFB244428" box="[1033,1229,1513,1536]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
is long (
<quantity id="815E92FE4F278D5BFA9BFA4AFA424428" box="[1345,1451,1513,1536]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="10" pageNumber="133" unit="mm" value="2.03">2.03 mm</quantity>
) as in
<taxonomicName id="81A644984F278D5BFC87F9AAFBEC4408" box="[861,1029,1545,1568]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFC87F9AAFBEC4408" box="[861,1029,1545,1568]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
, but its procingulum comprises a large anterolabial conule that occupies the entire anterior margin of M1 (
<figureCitation id="DE9D239E4F278D5BFC4BF9EAFC274448" box="[913,974,1609,1632]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="10" pageNumber="133">Fig. 6</figureCitation>
) and an inconspicuous anterolingual conule, making the anteromedial flexus minute. On the other hand, all Peruvian and Bolivian specimens of
<taxonomicName id="81A644984F278D5BFBACF92AFAE44488" box="[1142,1293,1673,1696]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFBACF92AFAE44488" box="[1142,1293,1673,1696]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
have a well-developed anterolingual conule and a deep anteromedial flexus (Luna and Patterson 2003;
<bibRefCitation id="223742EA4F278D5BFBD2F96AFB1344C8" author="VILLALPANDO, G. &amp; J. VARGAS &amp; J. SALAZAR-BRAVO" box="[1032,1274,1737,1760]" pageId="10" pageNumber="133" pagination="143 - 149" refId="ref14774" refString="VILLALPANDO, G., J. VARGAS, AND J. SALAZAR-BRAVO. 2006. First record of Rhagomys (Mammalia: Sigmodontinae) in Bolivia. Mastozoologia Neotropical 13: 143 - 149." type="journal article" year="2006">Villalpando et al. 2006</bibRefCitation>
;
<bibRefCitation id="223742EA4F278D5BFAD2F96AFA2644C8" author="MEDINA, C. E. &amp; D. R. DIAZ &amp; K. PINO &amp; A. PARI &amp; H. ZEBALLOS" box="[1288,1487,1737,1760]" pageId="10" pageNumber="133" pagination="2136" refId="ref13384" refString="MEDINA, C. E., D. R. DIAZ, K. PINO, A. PARI, AND H. ZEBALLOS. 2017. New locality records of Rhagomys longilingua Luna &amp; Patterson, 2003 (Rodentia: Cricetidae) in Peru. Check List 13: 2136." type="journal article" year="2017">Medina et al. 2017</bibRefCitation>
) although this flexus is smaller in the recently reported Brazilian specimen (
<bibRefCitation id="223742EA4F278D5BFC09F8AAFB2E4508" author="PERCEQUILLO, A. R. &amp; J. DALAPICOLLA &amp; E. F. ABREU-JUNIOR &amp; P. R. O. ROTH &amp; K. M. P. M. B. FERRAZ &amp; E. A. CHIQUITO" box="[979,1223,1801,1824]" pageId="10" pageNumber="133" pagination="4071" refId="ref13873" refString="PERCEQUILLO, A. R., J. DALAPICOLLA, E. F. ABREU-JUNIOR, P. R. O. ROTH, K. M. P. M. B. FERRAZ, AND E. A. CHIQUITO. 2017. How many species of mammals are there in Brazil? New records of rare rodents (Rodentia: Cricetidae: Sigmodontinae) from Amazonia raise the current known diversity. Peerj 5: e 4071." type="journal article" year="2017">Percequillo et al. 2017</bibRefCitation>
). The anterolabial conule of M
<quantity id="815E92FE4F278D5BFCA3F88AFC774568" box="[889,926,1833,1856]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="133" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="81A644984F278D5BFC7FF88AFB8B4568" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[933,1122,1833,1856]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFC7FF88AFB8B4568" box="[933,1122,1833,1856]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
lacks an anteromedial crista, which is present in Peruvian specimens but lacking in the Bolivian specimen of
<taxonomicName id="81A644984F278D5BFC69F8CAFAB845A8" authority="(Villalpando et al. 2006)" baseAuthorityName="Villalpando" baseAuthorityYear="2006" box="[947,1361,1897,1920]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFC69F8CAFBA345A8" box="[947,1098,1897,1920]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
(
<bibRefCitation id="223742EA4F278D5BFB82F8CAFAAF45A8" author="VILLALPANDO, G. &amp; J. VARGAS &amp; J. SALAZAR-BRAVO" box="[1112,1350,1897,1920]" pageId="10" pageNumber="133" pagination="143 - 149" refId="ref14774" refString="VILLALPANDO, G., J. VARGAS, AND J. SALAZAR-BRAVO. 2006. First record of Rhagomys (Mammalia: Sigmodontinae) in Bolivia. Mastozoologia Neotropical 13: 143 - 149." type="journal article" year="2006">Villalpando et al. 2006</bibRefCitation>
)
</taxonomicName>
. The anteroloph of M
<quantity id="815E92FE4F278D5BFC43F82AFC294588" box="[921,960,1929,1952]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="133" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="81A644984F278D5BFC13F82AFB604588" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[969,1161,1929,1952]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F278D5BFC13F82AFB604588" box="[969,1161,1929,1952]" italics="true" pageId="10" pageNumber="133">R. septentrionalis</emphasis>
</taxonomicName>
is attached directly to the rear of the anterolabial conule, whereas in
<taxonomicName id="81A644984F278D5BFB05F80AFA9045E8" box="[1247,1401,1961,1984]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F278D5BFB05F80AFA9045E8" box="[1247,1401,1961,1984]" italics="true" pageId="10" pageNumber="133">R. longilingua</emphasis>
</taxonomicName>
the two are connected via a secondary connection, the anterolophule (Luna and Patterson 2003). In
<taxonomicName id="81A644984F268D5AFEA9FF6AFDDA42C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[371,563,201,224]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFEA9FF6AFDDA42C8" box="[371,563,201,224]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
, the anterior mure of M1 and M2 is directly connected to the anterior margin of the protocone and exits the posterior margin of the protocone to form the median mure; the paracone of both molars lacks a connection with the median mure, while in
<taxonomicName id="81A644984F268D5AFDBCFEEAFCEA4348" box="[614,771,329,352]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F268D5AFDBCFEEAFCEA4348" box="[614,771,329,352]" italics="true" pageId="11" pageNumber="134">R. longilingua</emphasis>
</taxonomicName>
the paracone and protocone of M1 and M2 join the median mure by small connections. A posteroflexus is lacking on the M1 and M2 of
<taxonomicName id="81A644984F268D5AFEDCFE0AFE2B43E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[262,450,425,448]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFEDCFE0AFE2B43E8" box="[262,450,425,448]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
because the posteroloph joins the metacone labially, whereas Luna and Patterson (2003) described a small, shallow posteroflexus on M1 and M2.
</paragraph>
<paragraph id="46193F1B4F268D5AFF5AFDAAFD0440C8" blockId="11.[101,772,201,2016]" pageId="11" pageNumber="134">
In
<taxonomicName id="81A644984F268D5AFF7CFDAAFE854008" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[166,364,521,544]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFF7CFDAAFE854008" box="[166,364,521,544]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
, the hypoconid and the entoconid are fused along the posterior margin of m3 (
<figureCitation id="DE9D239E4F268D5AFD8BFD8AFD7B4068" box="[593,658,553,576]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="11" pageNumber="134">Fig. 6</figureCitation>
), whereas a median murid separates the hypoconid and entoconid in
<taxonomicName id="81A644984F268D5AFFBFFDCAFF1440A8" box="[101,253,617,640]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F268D5AFFBFFDCAFF1440A8" box="[101,253,617,640]" italics="true" pageId="11" pageNumber="134">R. longilingua</emphasis>
</taxonomicName>
. In
<taxonomicName id="81A644984F268D5AFEFEFDCAFE3640A8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[292,479,617,640]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFEFEFDCAFE3640A8" box="[292,479,617,640]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
, a mesolophid is evident on m3 that joins neither with the entoconid nor with the hypoconid and does not form a median murid. However,
<taxonomicName id="81A644984F268D5AFD99FD0AFD3340E8" box="[579,730,681,704]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F268D5AFD99FD0AFD3340E8" box="[579,730,681,704]" italics="true" pageId="11" pageNumber="134">R. longilingua</emphasis>
</taxonomicName>
has a median murid and a very conspicuous posterocrista on m3.
</paragraph>
<paragraph id="46193F1B4F268D5AFF5AFD4AFCEA4628" blockId="11.[101,772,201,2016]" pageId="11" pageNumber="134">
<emphasis id="74D2E3094F268D5AFF5AFD4AFED24128" box="[128,315,745,768]" italics="true" pageId="11" pageNumber="134">Comparison with</emphasis>
<taxonomicName id="81A644984F268D5AFE9EFD4AFE224128" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[324,459,745,768]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">R. rufescens</taxonomicName>
.
<emphasis id="74D2E3094F268D5AFE0BFD4AFCEA4128" box="[465,771,745,768]" italics="true" pageId="11" pageNumber="134">
<taxonomicName id="81A644984F268D5AFE36FD4AFCEA4128" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[492,771,745,768]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">Rhagomys septentrionalis</taxonomicName>
</emphasis>
also resembles
<taxonomicName id="81A644984F268D5AFED4FCAAFE7A4108" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[270,403,777,800]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFED4FCAAFE7A4108" box="[270,403,777,800]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
in size (
<tableCitation id="0B240AA04F268D5AFE2CFCAAFDAE4108" box="[502,583,777,800]" captionStart="Table 1" captionStartId="4.[126,182,506,527]" captionTargetPageId="4" captionText="Table 1.—External, vibrissal, and craniodental measurements (mm), and age class, for specimens of Rhagomys. Acronyms are: Natural History Museum, London (BMNH); Colección Boliviana de Fauna, La Paz (CBF); Field Museum of Natural History, Chicago (FMNH); Laboratory of Vertebrates, Universidade Federal de Ouro Preto (LZV-UFOP) with the accession number R; Instituto de Ciencias Biológicas, Escuela Politécnica Nacional, Quito (MEPN); Museu Nacional, Universidade Federal do Rio de Janeiro (MN); Museu de Zoologia, Universidade de Sao Paulo (MZUSP). The tabulated measurements were previously published, except for the MEPN holotype. Sex: Female (F), Male (M), unknown (?)." pageId="11" pageNumber="134">Table 1</tableCitation>
), but some adult specimens of
<taxonomicName id="81A644984F268D5AFF26FC8AFE684168" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[252,385,809,832]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFF26FC8AFE684168" box="[252,385,809,832]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
exceed the size apparently reached by
<taxonomicName id="81A644984F268D5AFF5CFCEAFEAB4148" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[134,322,841,864]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFF5CFCEAFEAB4148" box="[134,322,841,864]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
. Tail length is similar in
<taxonomicName id="81A644984F268D5AFD9DFCEAFCEA4148" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[583,771,841,864]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFD9DFCEAFCEA4148" box="[583,771,841,864]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
and
<taxonomicName id="81A644984F268D5AFF40FCCAFECC41A8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[154,293,873,896]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFF40FCCAFECC41A8" box="[154,293,873,896]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
. Externally, the main difference between these two species is that
<taxonomicName id="81A644984F268D5AFE5EFC2AFDE24188" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[388,523,905,928]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFE5EFC2AFDE24188" box="[388,523,905,928]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
lacks flattened spines in its pelage; rather,
<taxonomicName id="81A644984F268D5AFE8AFC0AFE3E41E8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[336,471,937,960]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFE8AFC0AFE3E41E8" box="[336,471,937,960]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
has soft and dense pelage (Luna and Patterson 2003;
<bibRefCitation id="223742EA4F268D5AFE4FFC6AFD7841C8" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[405,657,969,992]" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
), whereas
<taxonomicName id="81A644984F268D5AFFBFFC4AFEC84628" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[101,289,1001,1024]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFFBFFC4AFEC84628" box="[101,289,1001,1024]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
has spiny hairs covering its dorsum and belly.
</paragraph>
<paragraph id="46193F1B4F268D5AFF5AFBAAFD454448" blockId="11.[101,772,201,2016]" pageId="11" pageNumber="134">
Both species have an enlarged cranium with a longer, narrower rostrum than in
<taxonomicName id="81A644984F268D5AFEB8FB8AFE164668" box="[354,511,1065,1088]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F268D5AFEB8FB8AFE164668" box="[354,511,1065,1088]" italics="true" pageId="11" pageNumber="134">R. longilingua</emphasis>
</taxonomicName>
. However, the cranium in
<taxonomicName id="81A644984F268D5AFF5FFBEAFEE54648" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[133,268,1097,1120]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFF5FFBEAFEE54648" box="[133,268,1097,1120]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
is bigger (
<figureCitation id="DE9D239E4F268D5AFE51FBEAFE274648" box="[395,462,1097,1120]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="11" pageNumber="134">Fig. 3</figureCitation>
). The interorbital region of both species is narrowed at a midpoint, but the interorbital region in
<taxonomicName id="81A644984F268D5AFF11FB2AFE614688" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[203,392,1161,1184]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFF11FB2AFE614688" box="[203,392,1161,1184]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
is more sharply narrowed, whereas
<taxonomicName id="81A644984F268D5AFFBFFB0AFF0446E8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[101,237,1193,1216]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFFBFFB0AFF0446E8" box="[101,237,1193,1216]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
has an interorbital region shaped like a parenthesis and is more smoothly tapered (Pavan and Leite 2011). Both species have nasals that extend far beyond the premaxillary, and are slightly downturned at the anterior end. The zygomatic notch is deeper and wider in adult
<taxonomicName id="81A644984F268D5AFD9DFA8AFD254768" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[583,716,1321,1344]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFD9DFA8AFD254768" box="[583,716,1321,1344]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
than in
<taxonomicName id="81A644984F268D5AFF58FAEAFEA84748" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[130,321,1353,1376]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFF58FAEAFEA84748" box="[130,321,1353,1376]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
; although the zygomatic notch is tapered in younger specimens of
<taxonomicName id="81A644984F268D5AFEAFFACAFE1147A8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[373,504,1385,1408]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFEAFFACAFE1147A8" box="[373,504,1385,1408]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
(
<bibRefCitation id="223742EA4F268D5AFDD3FACAFD1247A8" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[521,763,1385,1408]" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
;
<bibRefCitation id="223742EA4F268D5AFFBFFA2AFEBF4788" author="PASSAMANI, M. &amp; R. A. SERPA CERBONCINI &amp; J. E. DE OLIVEIRA" box="[101,342,1417,1440]" pageId="11" pageNumber="134" pagination="827 - 831" refId="ref13707" refString="PASSAMANI, M., R. A. SERPA CERBONCINI, AND J. E. DE OLIVEIRA. 2011. Distribution extension of Phaenomys ferrugineus (Thomas, 1894), and new data on Abrawayaomys ruschii Cunha and Cruz, 1979 and Rhagomys rufescens (Thomas, 1886), three rare species of rodents (Rodentia: Cricetidae) in Minas Gerais, Brazil. Check List 7: 827 - 831" type="journal article" year="2011">Passamani et al. 2011</bibRefCitation>
), it remains more conspicuous than in
<taxonomicName id="81A644984F268D5AFFBFFA0AFEC547E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[101,300,1449,1472]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFFBFFA0AFEC547E8" box="[101,300,1449,1472]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
. Both species have a faint supraorbital crista along the interorbital region, squarely angled but not overhanging the margin (
<bibRefCitation id="223742EA4F268D5AFEA5FA4AFD924428" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[383,635,1513,1536]" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
). The interparietal in
<taxonomicName id="81A644984F268D5AFF0DF9AAFEB34408" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[215,346,1545,1568]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFF0DF9AAFEB34408" box="[215,346,1545,1568]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
is larger in the sagittal plane than in the transversal one as in
<taxonomicName id="81A644984F268D5AFE91F98AFDE04468" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[331,521,1577,1600]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFE91F98AFDE04468" box="[331,521,1577,1600]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
, and in neither species does it reach the squamosal bone at its lateral margins.
</paragraph>
<paragraph id="46193F1B4F268D5AFF5AF9CAFAE240C8" blockId="11.[101,772,201,2016]" lastBlockId="11.[819,1488,201,2016]" pageId="11" pageNumber="134">
The incisive foramen in
<taxonomicName id="81A644984F268D5AFE4FF9CAFDF544A8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[405,540,1641,1664]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFE4FF9CAFDF544A8" box="[405,540,1641,1664]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
is short (57%) relative to diastema length (
<bibRefCitation id="223742EA4F268D5AFEACF92AFD984488" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[374,625,1673,1696]" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
), whereas in
<emphasis id="74D2E3094F268D5AFFBFF90AFEC144E8" box="[101,296,1705,1728]" italics="true" pageId="11" pageNumber="134">
<taxonomicName id="81A644984F268D5AFFBFF90AFECA44E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[101,291,1705,1728]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">R. septentrionalis</taxonomicName>
,
</emphasis>
the incisive foramen is longer (64%), and its posterior margin reaches nearly the level of the M1 cingulum. The region of the masseteric tubercle in
<taxonomicName id="81A644984F268D5AFDF2F94AFD014528" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[552,744,1769,1792]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFDF2F94AFD014528" box="[552,744,1769,1792]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
is smooth, whereas in
<taxonomicName id="81A644984F268D5AFEE6F8AAFE564508" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[316,447,1801,1824]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFEE6F8AAFE564508" box="[316,447,1801,1824]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
, small fossae are evident near the anterior root of M1 (
<bibRefCitation id="223742EA4F268D5AFEBEF88AFDB94568" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[356,592,1833,1856]" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
). The zygomatic plate in
<taxonomicName id="81A644984F268D5AFF1DF8EAFE604548" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[199,393,1865,1888]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFF1DF8EAFE604548" box="[199,393,1865,1888]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
is narrower than in
<taxonomicName id="81A644984F268D5AFDA1F8EAFCEA4548" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[635,771,1865,1888]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFDA1F8EAFCEA4548" box="[635,771,1865,1888]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
(
<collectionCode id="20B7A7DE4F268D5AFFB7F8CAFF7645A8" box="[109,159,1897,1920]" country="South Africa" lsid="urn:lsid:biocol.org:col:15282" name="Herbarium Soutpansbergensis" pageId="11" pageNumber="134" type="Herbarium">ZPB</collectionCode>
=
<quantity id="815E92FE4F268D5AFF18F8CAFECD45A8" box="[194,292,1897,1920]" metricMagnitude="-3" metricUnit="m" metricValue="2.86" pageId="11" pageNumber="134" unit="mm" value="2.86">2.86 mm</quantity>
versus
<quantity id="815E92FE4F268D5AFEA6F8CAFE3C45A8" box="[380,469,1897,1920]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="11" pageNumber="134" unit="mm" value="3.3">3.3 mm</quantity>
;
<tableCitation id="0B240AA04F268D5AFE38F8CAFDDD45A8" box="[482,564,1897,1920]" captionStart="Table 1" captionStartId="4.[126,182,506,527]" captionTargetPageId="4" captionText="Table 1.—External, vibrissal, and craniodental measurements (mm), and age class, for specimens of Rhagomys. Acronyms are: Natural History Museum, London (BMNH); Colección Boliviana de Fauna, La Paz (CBF); Field Museum of Natural History, Chicago (FMNH); Laboratory of Vertebrates, Universidade Federal de Ouro Preto (LZV-UFOP) with the accession number R; Instituto de Ciencias Biológicas, Escuela Politécnica Nacional, Quito (MEPN); Museu Nacional, Universidade Federal do Rio de Janeiro (MN); Museu de Zoologia, Universidade de Sao Paulo (MZUSP). The tabulated measurements were previously published, except for the MEPN holotype. Sex: Female (F), Male (M), unknown (?)." pageId="11" pageNumber="134">Table 1</tableCitation>
), and in both species the anterior edge of the zygomatic plate is straight and lacks a zygomatic spine. However, in
<emphasis id="74D2E3094F268D5AFDDDF80AFD7A45E8" box="[519,659,1961,1984]" italics="true" pageId="11" pageNumber="134">
<taxonomicName id="81A644984F268D5AFDDDF80AFD6645E8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[519,655,1961,1984]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">R. rufescens</taxonomicName>
,
</emphasis>
the zygomatic plate forms an acute angle over its antero-dorsal margin. The length of the orbital fossa in
<taxonomicName id="81A644984F268D5AFB7EFF6AFA8A42C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1188,1379,201,224]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFB7EFF6AFA8A42C8" box="[1188,1379,201,224]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
is shorter (
<collectionCode id="20B7A7DE4F268D5AFCE0FF4AFC874328" box="[826,878,233,256]" pageId="11" pageNumber="134">LOF</collectionCode>
=
<quantity id="815E92FE4F268D5AFC5FFF4AFC0A4328" box="[901,995,233,256]" metricMagnitude="-3" metricUnit="m" metricValue="9.35" pageId="11" pageNumber="134" unit="mm" value="9.35">9.35 mm</quantity>
) than in
<taxonomicName id="81A644984F268D5AFBE3FF4AFB504328" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1081,1209,233,256]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFBE3FF4AFB504328" box="[1081,1209,233,256]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
(
<collectionCode id="20B7A7DE4F268D5AFB1EFF4AFB114328" box="[1220,1272,233,256]" pageId="11" pageNumber="134">LOF</collectionCode>
=
<quantity id="815E92FE4F268D5AFACAFF4AFAB34328" box="[1296,1370,233,256]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="134" unit="mm" value="10.0">10 mm</quantity>
,
<emphasis id="74D2E3094F268D5AFAB9FF4AFA6D4328" box="[1379,1412,233,256]" italics="true" pageId="11" pageNumber="134">SD</emphasis>
= 0.39, range = 9.3410.65), resembling specimens of
<taxonomicName id="81A644984F268D5AFAEBFEAAFA5C4308" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1329,1461,265,288]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFAEBFEAAFA5C4308" box="[1329,1461,265,288]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
at age 1 (
<tableCitation id="0B240AA04F268D5AFC5BFE8AFC3A4368" box="[897,979,297,320]" captionStart="Table 1" captionStartId="4.[126,182,506,527]" captionTargetPageId="4" captionText="Table 1.—External, vibrissal, and craniodental measurements (mm), and age class, for specimens of Rhagomys. Acronyms are: Natural History Museum, London (BMNH); Colección Boliviana de Fauna, La Paz (CBF); Field Museum of Natural History, Chicago (FMNH); Laboratory of Vertebrates, Universidade Federal de Ouro Preto (LZV-UFOP) with the accession number R; Instituto de Ciencias Biológicas, Escuela Politécnica Nacional, Quito (MEPN); Museu Nacional, Universidade Federal do Rio de Janeiro (MN); Museu de Zoologia, Universidade de Sao Paulo (MZUSP). The tabulated measurements were previously published, except for the MEPN holotype. Sex: Female (F), Male (M), unknown (?)." pageId="11" pageNumber="134">Table 1</tableCitation>
). The mesopterygoid foramen of
<taxonomicName id="81A644984F268D5AFA90FE8AFA394368" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1354,1488,297,320]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFA90FE8AFA394368" box="[1354,1488,297,320]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
is narrow (
<bibRefCitation id="223742EA4F268D5AFC68FEEAFB5D4348" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[946,1204,329,352]" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
; Pavan and Leite 2011), whereas the mesopterygoid foramen
<emphasis id="74D2E3094F268D5AFB0BFECAFA5A43A8" box="[1233,1459,361,384]" italics="true" pageId="11" pageNumber="134">
in
<taxonomicName id="81A644984F268D5AFB2BFECAFA5A43A8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1265,1459,361,384]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">R. septentrionalis</taxonomicName>
</emphasis>
is wide and square-shaped at its anterior end. Both species present sphenopalatine vacuities, but those in
<taxonomicName id="81A644984F268D5AFB38FE0AFA4D43E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1250,1444,425,448]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFB38FE0AFA4D43E8" box="[1250,1444,425,448]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
are smaller. The basioccipital of
<taxonomicName id="81A644984F268D5AFBAFFE6AFADC43C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1141,1333,457,480]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFBAFFE6AFADC43C8" box="[1141,1333,457,480]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
is wider posteriorly than in
<taxonomicName id="81A644984F268D5AFC39FE4AFB804028" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[995,1129,489,512]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFC39FE4AFB804028" box="[995,1129,489,512]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
(
<figureCitation id="DE9D239E4F268D5AFBA6FE4AFB574028" box="[1148,1214,489,512]" captionStart="Fig" captionStartId="7.[101,136,1365,1386]" captionTargetBox="[114,1475,200,1352]" captionTargetId="figure-6@7.[114,1475,200,1352]" captionTargetPageId="7" captionText="Fig. 3.—Skulls of Rhagomys to the same scale. (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype), showing dorsal, ventral, and lateral, view of the cranium and lateral view of the mandible; (B) Rhagomys rufescens (MZUSP 65545), in dorsal and ventral view; (C) Rhagomys longilingua (MUSM 20009), showing dorsal, ventral, and lateral, view of the cranium and lateral view of mandible. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632401" httpUri="https://zenodo.org/record/10632401/files/figure.png" pageId="11" pageNumber="134">Fig. 3</figureCitation>
). The auditory bullae of
<taxonomicName id="81A644984F268D5AFCE9FDAAFC064008" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[819,1007,521,544]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFCE9FDAAFC064008" box="[819,1007,521,544]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
are wider and longer with shorter Eustachian tubes. The occipital condyles of
<taxonomicName id="81A644984F268D5AFB4DFD8AFAF24068" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1175,1307,553,576]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFB4DFD8AFAF24068" box="[1175,1307,553,576]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
are farther from the upper molars in lateral view of cranium (
<bibRefCitation id="223742EA4F268D5AFA8DFDEAFC4740A8" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
) than the ones in
<emphasis id="74D2E3094F268D5AFB51FDCAFABC40A8" box="[1163,1365,617,640]" italics="true" pageId="11" pageNumber="134">
<taxonomicName id="81A644984F268D5AFB51FDCAFAB840A8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1163,1361,617,640]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">R. septentrionalis</taxonomicName>
.
</emphasis>
The squamosal hamular process in
<taxonomicName id="81A644984F268D5AFB81FD2AFAF54088" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1115,1308,649,672]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFB81FD2AFAF54088" box="[1115,1308,649,672]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
is narrow as in
<taxonomicName id="81A644984F268D5AFCE9FD0AFC5040E8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[819,953,681,704]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFCE9FD0AFC5040E8" box="[819,953,681,704]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
; however, in
<taxonomicName id="81A644984F268D5AFB8EFD0AFAFC40E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1108,1301,681,704]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFB8EFD0AFAFC40E8" box="[1108,1301,681,704]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
, it is longer and contacts with the mastoid at medium height.
</paragraph>
<paragraph id="46193F1B4F268D5AFC97FD4AFBF94668" blockId="11.[819,1488,201,2016]" pageId="11" pageNumber="134">
On the mandible, the mental foramen of both
<taxonomicName id="81A644984F268D5AFAC2FD4AFA394128" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1304,1488,745,768]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFAC2FD4AFA394128" box="[1304,1488,745,768]" italics="true" pageId="11" pageNumber="134">R.septentrionalis</emphasis>
</taxonomicName>
and
<taxonomicName id="81A644984F268D5AFCB9FCAAFC174108" box="[867,1022,777,800]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F268D5AFCB9FCAAFC174108" box="[867,1022,777,800]" italics="true" pageId="11" pageNumber="134">R. longilingua</emphasis>
</taxonomicName>
lies at a level about midway beneath m1; in
<taxonomicName id="81A644984F268D5AFC88FC8AFC314168" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[850,984,809,832]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFC88FC8AFC314168" box="[850,984,809,832]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
, the mental foramen is located ventral to the anterior margin of m1 (
<figureCitation id="DE9D239E4F268D5AFBECFCEAFB9F4148" box="[1078,1142,841,864]" captionStart="Fig" captionStartId="9.[101,136,1893,1914]" captionTargetBox="[126,747,1216,1880]" captionTargetId="figure-6@9.[126,747,1216,1880]" captionTargetPageId="9" captionText="Fig. 5.—Lateral view of the mandibles of Rhagomys: (A) Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype); (B) R. rufescens (MZUSP AB401; from Pavan and Leite 2011); and (C) R. longilingua (MUSM 20009). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10632405" httpUri="https://zenodo.org/record/10632405/files/figure.png" pageId="11" pageNumber="134">Fig. 5</figureCitation>
). The upper masseteric crest in
<taxonomicName id="81A644984F268D5AFCE9FCCAFC5D41A8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[819,948,873,896]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFCE9FCCAFC5D41A8" box="[819,948,873,896]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
evident in figures 1012 of Pavan and Leite (2011) is more conspicuous and straighter than in
<taxonomicName id="81A644984F268D5AFAD1FC2AFA234188" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1291,1482,905,928]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFAD1FC2AFA234188" box="[1291,1482,905,928]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
, where it is less conspicuous and more bowed. The capsular process of the lower incisor in
<taxonomicName id="81A644984F268D5AFB5CFC6AFAAC41C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1158,1349,969,992]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFB5CFC6AFAAC41C8" box="[1158,1349,969,992]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
is located at the middle of the angular notch of the mandible, whereas in
<taxonomicName id="81A644984F268D5AFCE9FBAAFC5D4608" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[819,948,1033,1056]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFCE9FBAAFC5D4608" box="[819,948,1033,1056]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
this process is located in the posterior third portion of the angular notch.
</paragraph>
<paragraph id="46193F1B4F268D5AFC97FBEAFA8746C8" blockId="11.[819,1488,201,2016]" pageId="11" pageNumber="134">
The dental maxillary cusps of
<taxonomicName id="81A644984F268D5AFB76FBEAFA874648" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1196,1390,1097,1120]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFB76FBEAFA874648" box="[1196,1390,1097,1120]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
are narrower than in
<taxonomicName id="81A644984F268D5AFC1EFBCAFBAF46A8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[964,1094,1129,1152]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFC1EFBCAFBAF46A8" box="[964,1094,1129,1152]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
. The labial cusps of the upper molars are slightly posterior to the lingual cusps in
<taxonomicName id="81A644984F268D5AFAC8FB2AFA394688" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1298,1488,1161,1184]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFAC8FB2AFA394688" box="[1298,1488,1161,1184]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
(
<figureCitation id="DE9D239E4F268D5AFCE6FB0AFC9246E8" box="[828,891,1193,1216]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="11" pageNumber="134">Fig. 6</figureCitation>
), whereas in
<taxonomicName id="81A644984F268D5AFBC8FB0AFB7F46E8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1042,1174,1193,1216]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFBC8FB0AFB7F46E8" box="[1042,1174,1193,1216]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
, the labial and lingual cusps are located at the same level (
<bibRefCitation id="223742EA4F268D5AFBB5FB6AFAB746C8" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[1135,1374,1225,1248]" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
).
</paragraph>
<paragraph id="46193F1B4F268D5AFC97FB4AFBE64508" blockId="11.[819,1488,201,2016]" pageId="11" pageNumber="134">
The anteromedial flexus of
<taxonomicName id="81A644984F268D5AFB49FB4AFAB14728" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1171,1368,1257,1280]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFB49FB4AFAB14728" box="[1171,1368,1257,1280]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
in M1 is short and shallow, whereas in
<taxonomicName id="81A644984F268D5AFB5FFAAAFAE54708" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1157,1292,1289,1312]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFB5FFAAAFAE54708" box="[1157,1292,1289,1312]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
it is well-defined and deep (
<bibRefCitation id="223742EA4F268D5AFC73FA8AFB4D4768" author="PERCEQUILLO, A. R. &amp; P. R. GONCALVES &amp; J. A. OLIVEIRA" box="[937,1188,1321,1344]" pageId="11" pageNumber="134" pagination="238 - 257" refId="ref13955" refString="PERCEQUILLO, A. R., P. R. GONCALVES, AND J. A. OLIVEIRA. 2004. The rediscovery of Rhagomys rufescens (Thomas, 1886), with a morphological redescription and comments on its systematic relationships based on morphological and molecular (cytochrome b) characters. Mammalian Biology 69: 238 - 257." type="journal article" year="2004">Percequillo et al. 2004</bibRefCitation>
). The anterolingual conule in
<taxonomicName id="81A644984F268D5AFC89FAEAFBFC4748" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[851,1045,1353,1376]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFC89FAEAFBFC4748" box="[851,1045,1353,1376]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
is smaller than the anterolabial conule (
<figureCitation id="DE9D239E4F268D5AFCE6FACAFC6847A8" box="[828,897,1385,1408]" captionStart="Fig" captionStartId="9.[819,854,1923,1944]" captionTargetBox="[843,1464,1082,1910]" captionTargetId="figure-54@9.[843,1464,1082,1910]" captionTargetPageId="9" captionText="Fig. 6.—Molars of Rhagomys septentrionalis sp. nov. (MEPN 10898, holotype). (A) Occlusal view of upper right molars; (B) Lower right molars. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10632407" httpUri="https://zenodo.org/record/10632407/files/figure.png" pageId="11" pageNumber="134">Fig. 6</figureCitation>
), whereas in
<taxonomicName id="81A644984F268D5AFBFDFACAFB5847A8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1063,1201,1385,1408]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFBFDFACAFB5847A8" box="[1063,1201,1385,1408]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
the anterolingual conule is similar in size to the anterolabial conule. The m1 of
<taxonomicName id="81A644984F268D5AFCE9FA0AFC0647E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[819,1007,1449,1472]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFCE9FA0AFC0647E8" box="[819,1007,1449,1472]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
presents the metaconid close to the posterior margin of the anterolingual conulid that reduces and narrows the metaflexid, while in
<taxonomicName id="81A644984F268D5AFB9AFA4AFB2C4428" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[1088,1221,1513,1536]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFB9AFA4AFB2C4428" box="[1088,1221,1513,1536]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
, the metaflexid is wider and more conspicuous. In the m1 and m2 of
<taxonomicName id="81A644984F268D5AFAD7F9AAFA234408" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1293,1482,1545,1568]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFAD7F9AAFA234408" box="[1293,1482,1545,1568]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
, the mesolophid adheres to the former margin of the entoconid, and does not allow the entoflexid to reach the lingual side, whereas in
<taxonomicName id="81A644984F268D5AFC71F9CAFBC444A8" baseAuthorityName="Thomas" baseAuthorityYear="1886" box="[939,1069,1641,1664]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="74D2E3094F268D5AFC71F9CAFBC444A8" box="[939,1069,1641,1664]" italics="true" pageId="11" pageNumber="134">R. rufescens</emphasis>
</taxonomicName>
the mesolophid of m2 clearly separates the mesoflexid and the entoflexid (
<bibRefCitation id="223742EA4F268D5AFB76F92AFA484488" author="PERCEQUILLO, A. R. &amp; J. DALAPICOLLA &amp; E. F. ABREU-JUNIOR &amp; P. R. O. ROTH &amp; K. M. P. M. B. FERRAZ &amp; E. A. CHIQUITO" box="[1196,1441,1673,1696]" pageId="11" pageNumber="134" pagination="4071" refId="ref13873" refString="PERCEQUILLO, A. R., J. DALAPICOLLA, E. F. ABREU-JUNIOR, P. R. O. ROTH, K. M. P. M. B. FERRAZ, AND E. A. CHIQUITO. 2017. How many species of mammals are there in Brazil? New records of rare rodents (Rodentia: Cricetidae: Sigmodontinae) from Amazonia raise the current known diversity. Peerj 5: e 4071." type="journal article" year="2017">Percequillo et al. 2017</bibRefCitation>
). In both species, the m3 is very similar with a reduced entoconid that is joined with the hypoconid, so that the posterior margin of m3 is more triangular than square (as in
<taxonomicName id="81A644984F268D5AFB2FF94AFA654528" box="[1269,1420,1769,1792]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F268D5AFB2FF94AFA654528" box="[1269,1420,1769,1792]" italics="true" pageId="11" pageNumber="134">R. longilingua</emphasis>
</taxonomicName>
; Luna and Patterson 2003).
</paragraph>
<paragraph id="46193F1B4F268D5DFC97F88AFE704168" blockId="11.[819,1488,201,2016]" lastBlockId="12.[144,814,201,2016]" lastPageId="12" lastPageNumber="135" pageId="11" pageNumber="134">
<emphasis id="74D2E3094F268D5AFC97F88AFC1E4568" box="[845,1015,1833,1856]" italics="true" pageId="11" pageNumber="134">Natural history.</emphasis>
—The
<typeStatus id="991D81B94F268D5AFB9AF88AFB874568" box="[1088,1134,1833,1856]" pageId="11" pageNumber="134">type</typeStatus>
locality of
<taxonomicName id="81A644984F268D5AFB32F88AFA4C4568" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1256,1445,1833,1856]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F268D5AFB32F88AFA4C4568" box="[1256,1445,1833,1856]" italics="true" pageId="11" pageNumber="134">R. septentrionalis</emphasis>
</taxonomicName>
lies in “El Zarza” (Wildlife Shelter “El Zarza”), located in Canton Yantzatza of Zamora-Chinchipe Province. This is a well preserved forest. Owing to peculiarities of local topography, humidity, and the atmosphere, the vegetation is characteristic of medium-height forests with canopy gaps. Some trees are covered by epiphytes. The forest in the area of San Antonio is very dense especially for the shrubby vegetation at the margins of Rio Blanco. The forest is periodically flooded in some areas close to the river. Great quantities of mosses and herbs cover the ground. According to Aguirre et al. (2013), El Zarza is part of the Evergreen foothill forest ecosystem of the mountain ranges of the Cóndor-Kutukú, characterized by dense vegetation. The representative trees species are:
<taxonomicName id="81A644984F218D5DFE2AFE0AFD9A43E8" box="[496,627,425,448]" class="Magnoliopsida" family="Lauraceae" genus="Aniba" kingdom="Plantae" order="Laurales" pageId="12" pageNumber="135" phylum="Tracheophyta" rank="species" species="muca">
<emphasis id="74D2E3094F218D5DFE2AFE0AFD9A43E8" box="[496,627,425,448]" italics="true" pageId="12" pageNumber="135">Aniba muca</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFD58FE0AFCCF43E8" box="[642,806,425,448]" class="Magnoliopsida" family="Moraceae" genus="Brosimum" kingdom="Plantae" order="Rosales" pageId="12" pageNumber="135" phylum="Tracheophyta" rank="species" species="utile">
<emphasis id="74D2E3094F218D5DFD58FE0AFCCF43E8" box="[642,806,425,448]" italics="true" pageId="12" pageNumber="135">Brosimum utile</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFF4AFE6AFE9943C8" box="[144,368,457,480]" class="Magnoliopsida" family="Urticaceae" genus="Cecropia" kingdom="Plantae" order="Rosales" pageId="12" pageNumber="135" phylum="Tracheophyta" rank="species" species="marginalis">
<emphasis id="74D2E3094F218D5DFF4AFE6AFE9943C8" box="[144,368,457,480]" italics="true" pageId="12" pageNumber="135">Cecropia marginalis</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFE5AFE6AFDF443C8" box="[384,541,457,480]" class="Magnoliopsida" family="Cannabaceae" genus="Celtis" kingdom="Plantae" order="Rosales" pageId="12" pageNumber="135" phylum="Tracheophyta" rank="species" species="schippii">
<emphasis id="74D2E3094F218D5DFE5AFE6AFDF443C8" box="[384,541,457,480]" italics="true" pageId="12" pageNumber="135">Celtis schippii</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFDF4FE6AFCCE43C8" box="[558,807,457,480]" class="Magnoliopsida" family="Rubiaceae" genus="Chimarrhis" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="135" phylum="Tracheophyta" rank="species" species="glabriflora">
<emphasis id="74D2E3094F218D5DFDF4FE6AFCCE43C8" box="[558,807,457,480]" italics="true" pageId="12" pageNumber="135">Chimarrhis glabriflora</emphasis>
</taxonomicName>
, and
<taxonomicName id="81A644984F218D5DFF64FE4AFD524028" authority="(Aguirre et al. 2013)" baseAuthorityName="Aguirre" baseAuthorityYear="2013" box="[190,699,489,512]" class="Polypodiopsida" family="Dryopteridaceae" genus="Elaphoglossum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="135" phylum="Tracheophyta" rank="species" species="latifolium">
<emphasis id="74D2E3094F218D5DFF64FE4AFE3A4028" box="[190,467,489,512]" italics="true" pageId="12" pageNumber="135">Elaphoglossum latifolium</emphasis>
(Aguirre et al. 2013)
</taxonomicName>
. This vegetation is characteristic of very humid forests located in the eastern foothills of the Andes in Zamora Chinchipe province (Aguirre et al. 2013), where many streams descend gentle slopes between 1,400 and
<quantity id="815E92FE4F218D5DFE86FDCAFE5C40A8" box="[348,437,617,640]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="135" unit="m" value="1800.0">1,800 m</quantity>
. The specimen of
<taxonomicName id="81A644984F218D5DFDA9FDCAFCC740A8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[627,814,617,640]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F218D5DFDA9FDCAFCC740A8" box="[627,814,617,640]" italics="true" pageId="12" pageNumber="135">R. septentrionalis</emphasis>
</taxonomicName>
was captured in one such stream in the “El Zarza,” opportunistically scooped out of the water with a hand fishing net. The habitat of the
<taxonomicName id="81A644984F218D5DFEF4FD6AFE0740C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[302,494,713,736]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F218D5DFEF4FD6AFE0740C8" box="[302,494,713,736]" italics="true" pageId="12" pageNumber="135">R. septentrionalis</emphasis>
</taxonomicName>
specimen was similar to the very humid habitats where
<taxonomicName id="81A644984F218D5DFE69FD4AFDA54128" box="[435,588,745,768]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="longilingua">
<emphasis id="74D2E3094F218D5DFE69FD4AFDA54128" box="[435,588,745,768]" italics="true" pageId="12" pageNumber="135">R. longilingua</emphasis>
</taxonomicName>
was found in central and southern
<collectingCountry id="3EB17F8B4F218D5DFEFFFCAAFEB04108" box="[293,345,777,800]" name="Peru" pageId="12" pageNumber="135">Peru</collectingCountry>
, northern
<collectingCountry id="3EB17F8B4F218D5DFE13FCAAFDF34108" box="[457,538,777,800]" name="Bolivia" pageId="12" pageNumber="135">Bolivia</collectingCountry>
, and Brazilian Amazonia (
<bibRefCitation id="223742EA4F218D5DFF43FC8AFE604168" author="PERCEQUILLO, A. R. &amp; J. DALAPICOLLA &amp; E. F. ABREU-JUNIOR &amp; P. R. O. ROTH &amp; K. M. P. M. B. FERRAZ &amp; E. A. CHIQUITO" box="[153,393,809,832]" pageId="12" pageNumber="135" pagination="4071" refId="ref13873" refString="PERCEQUILLO, A. R., J. DALAPICOLLA, E. F. ABREU-JUNIOR, P. R. O. ROTH, K. M. P. M. B. FERRAZ, AND E. A. CHIQUITO. 2017. How many species of mammals are there in Brazil? New records of rare rodents (Rodentia: Cricetidae: Sigmodontinae) from Amazonia raise the current known diversity. Peerj 5: e 4071." type="journal article" year="2017">Percequillo et al. 2017</bibRefCitation>
).
</paragraph>
<paragraph id="46193F1B4F218D5DFF71FCEAFE484628" blockId="12.[144,814,201,2016]" pageId="12" pageNumber="135">
Other species of small mammals that
<collectionCode id="20B7A7DE4F218D5DFD9FFCEAFD664148" box="[581,655,841,864]" pageId="12" pageNumber="135">MEPN</collectionCode>
surveys documented in the area of San Antonio are
<taxonomicName id="81A644984F218D5DFDE3FCCAFCCE41A8" authorityName="Linnaeus" authorityYear="1758" box="[569,807,873,896]" class="Mammalia" family="Didelphidae" genus="Didelphis" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="marsupialis">
<emphasis id="74D2E3094F218D5DFDE3FCCAFCCE41A8" box="[569,807,873,896]" italics="true" pageId="12" pageNumber="135">Didelphis marsupialis</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFF4AFC2AFE834188" authorityName="Ferreira, Oliveira, Lima-Silva &amp; Rossi" authorityYear="2020" box="[144,362,905,928]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="murina">
<emphasis id="74D2E3094F218D5DFF4AFC2AFE834188" box="[144,362,905,928]" italics="true" pageId="12" pageNumber="135">Marmosops murina</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFE5AFC2AFD884188" baseAuthorityName="Thomas" baseAuthorityYear="1897" box="[384,609,905,928]" class="Mammalia" family="Didelphidae" genus="Monodelphis" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="adusta">
<emphasis id="74D2E3094F218D5DFE5AFC2AFD884188" box="[384,609,905,928]" italics="true" pageId="12" pageNumber="135">Monodelphis adusta</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFDADFC2AFCC04188" box="[631,809,905,928]" class="Mammalia" family="Emballonuridae" genus="Saccopteryx" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="74D2E3094F218D5DFDADFC2AFD104188" box="[631,761,905,928]" italics="true" pageId="12" pageNumber="135">Saccopteryx</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFF4AFC0AFED641E8" baseAuthorityName="E. Geoffroy" baseAuthorityYear="1818" box="[144,319,937,960]" class="Mammalia" family="Phyllostomidae" genus="Anoura" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="caudifer">
<emphasis id="74D2E3094F218D5DFF4AFC0AFED641E8" box="[144,319,937,960]" italics="true" pageId="12" pageNumber="135">Anoura caudifer</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFE91FC0AFE2541E8" box="[331,460,937,960]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="74D2E3094F218D5DFE91FC0AFE4C41E8" box="[331,421,937,960]" italics="true" pageId="12" pageNumber="135">Artibeus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFE0DFC0AFD4441E8" baseAuthorityName="Schinz" baseAuthorityYear="1821" box="[471,685,937,960]" class="Mammalia" family="Phyllostomidae" genus="Carollia" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="brevicauda">
<emphasis id="74D2E3094F218D5DFE0DFC0AFD4441E8" box="[471,685,937,960]" italics="true" pageId="12" pageNumber="135">Carollia brevicauda</emphasis>
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFD60FC0AFCC041E8" box="[698,809,937,960]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="74D2E3094F218D5DFD60FC0AFCEA41E8" box="[698,771,937,960]" italics="true" pageId="12" pageNumber="135">Myotis</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFF4AFC6AFEF441C8" box="[144,285,969,992]" class="Mammalia" family="Molossidae" genus="Molossus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="74D2E3094F218D5DFF4AFC6AFF1C41C8" box="[144,245,969,992]" italics="true" pageId="12" pageNumber="135">Molossus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFEF0FC6AFE0241C8" box="[298,491,969,992]" class="Mammalia" family="Cricetidae" genus="Euryoryzomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="74D2E3094F218D5DFEF0FC6AFE2A41C8" box="[298,451,969,992]" italics="true" pageId="12" pageNumber="135">Euryoryzomys</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="81A644984F218D5DFE22FC6AFD1141C8" baseAuthorityName="Tomes" baseAuthorityYear="1860" box="[504,760,969,992]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="albigularis">
<emphasis id="74D2E3094F218D5DFE22FC6AFD1141C8" box="[504,760,969,992]" italics="true" pageId="12" pageNumber="135">Nephelomys albigularis</emphasis>
</taxonomicName>
, and
<taxonomicName id="81A644984F218D5DFF4AFC4AFE734628" box="[144,410,1001,1024]" class="Mammalia" family="Sciuridae" genus="Hadrosciurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="igniventris">
<emphasis id="74D2E3094F218D5DFF4AFC4AFE734628" box="[144,410,1001,1024]" italics="true" pageId="12" pageNumber="135">Hadrosciurus igniventris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="46193F1B4F218D5DFF71FBAAFE264428" blockId="12.[144,814,201,2016]" pageId="12" pageNumber="135">
<emphasis id="74D2E3094F218D5DFF71FBAAFEF94608" box="[171,272,1033,1056]" italics="true" pageId="12" pageNumber="135">Remarks.</emphasis>
—Our morphological and genetic results confidently recover the new mouse as a species of
<taxonomicName id="81A644984F218D5DFD5CFB8AFD1C4668" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[646,757,1065,1088]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFD5CFB8AFD1C4668" box="[646,757,1065,1088]" italics="true" pageId="12" pageNumber="135">Rhagomys</emphasis>
</taxonomicName>
. The genetic results corroborate the position of that genus in the tribe
<taxonomicName id="81A644984F218D5DFF4AFBCAFED846A8" authorityName="Steadman and Ray" authorityYear="1982" box="[144,305,1129,1152]" class="Mammalia" family="Cricetidae" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="tribe" tribe="Thomasomyini">Thomasomyini</taxonomicName>
proposed by D´Elía et al. (2006) and
<bibRefCitation id="223742EA4F218D5DFD0FFBCAFEF84688" author="PACHECO, V., J. L. &amp; PATTON, AND &amp; G. D'ELIA" pageId="12" pageNumber="135" refId="ref13544" refString="PACHECO, V., J. L. PATTON, AND G. D'ELIA. 2015. Tribe Thomasomyini Steadman and Ray, 1982. Pp. 571 - 574 in Mammals of South America. Vol. 2. Rodents (J. L. Patton, U. F. J. Pardinas, and G. D´Elia, eds.). The University of Chicago Press. Chicago, Illinois." type="book" year="2015">Pacheco et al. (2015)</bibRefCitation>
. Other members of the tribe include
<taxonomicName id="81A644984F218D5DFD4FFB2AFCE84688" authorityName="Thomas" authorityYear="1898" box="[661,769,1161,1184]" class="Mammalia" family="Cricetidae" genus="Aepeomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFD4FFB2AFCE84688" box="[661,769,1161,1184]" italics="true" pageId="12" pageNumber="135">Aepeomys</emphasis>
</taxonomicName>
and
<taxonomicName id="81A644984F218D5DFF4AFB0AFF1C46E8" authorityName="Thomas" authorityYear="1897" box="[144,245,1193,1216]" class="Mammalia" family="Cricetidae" genus="Chilomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFF4AFB0AFF1C46E8" box="[144,245,1193,1216]" italics="true" pageId="12" pageNumber="135">Chilomys</emphasis>
</taxonomicName>
(both endemic to the Northern Andes),
<taxonomicName id="81A644984F218D5DFD78FB0AFCC446E8" authorityName="Coues" authorityYear="1884" box="[674,813,1193,1216]" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFD78FB0AFCC446E8" box="[674,813,1193,1216]" italics="true" pageId="12" pageNumber="135">Thomasomys</emphasis>
</taxonomicName>
(endemic to the Northern and Central Andes), and the widespread genus
<taxonomicName id="81A644984F218D5DFEC5FB4AFE744728" authorityName="Tschudi" authorityYear="1845" box="[287,413,1257,1280]" class="Mammalia" family="Cricetidae" genus="Rhipidomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFEC5FB4AFE744728" box="[287,413,1257,1280]" italics="true" pageId="12" pageNumber="135">Rhipidomys</emphasis>
</taxonomicName>
, found across most of the Neotropics. The discovery of
<taxonomicName id="81A644984F218D5DFE8BFAAAFD8E4708" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[337,615,1289,1312]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F218D5DFE8BFAAAFD8E4708" box="[337,615,1289,1312]" italics="true" pageId="12" pageNumber="135">Rhagomys septentrionalis</emphasis>
</taxonomicName>
extends the range of that genus
<quantity id="815E92FE4F218D5DFEF6FA8AFE684768" box="[300,385,1321,1344]" metricMagnitude="5" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="135" unit="km" value="700.0">700 km</quantity>
northward from the records in central
<collectingCountry id="3EB17F8B4F218D5DFF4AFAEAFF284748" box="[144,193,1353,1376]" name="Peru" pageId="12" pageNumber="135">Peru</collectingCountry>
at Santa Rita Alta,
<collectingRegion id="8462F1F94F218D5DFE72FAEAFDE14748" box="[424,520,1353,1376]" country="Peru" name="Huanuco" pageId="12" pageNumber="135">Huánuco</collectingRegion>
(
<bibRefCitation id="223742EA4F218D5DFDC6FAEAFD1B4748" author="MEDINA, C. E. &amp; D. R. DIAZ &amp; K. PINO &amp; A. PARI &amp; H. ZEBALLOS" box="[540,754,1353,1376]" pageId="12" pageNumber="135" pagination="2136" refId="ref13384" refString="MEDINA, C. E., D. R. DIAZ, K. PINO, A. PARI, AND H. ZEBALLOS. 2017. New locality records of Rhagomys longilingua Luna &amp; Patterson, 2003 (Rodentia: Cricetidae) in Peru. Check List 13: 2136." type="journal article" year="2017">Medina et al. 2017</bibRefCitation>
) and
<quantity id="815E92FE4F218D5DFF4AFACAFF1547A8" box="[144,252,1385,1408]" metricMagnitude="6" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="135" unit="km" value="1400.0">1,400 km</quantity>
westward from the record near Jirau, Rondônia (
<bibRefCitation id="223742EA4F218D5DFF43FA2AFE7F4788" author="PERCEQUILLO, A. R. &amp; J. DALAPICOLLA &amp; E. F. ABREU-JUNIOR &amp; P. R. O. ROTH &amp; K. M. P. M. B. FERRAZ &amp; E. A. CHIQUITO" box="[153,406,1417,1440]" pageId="12" pageNumber="135" pagination="4071" refId="ref13873" refString="PERCEQUILLO, A. R., J. DALAPICOLLA, E. F. ABREU-JUNIOR, P. R. O. ROTH, K. M. P. M. B. FERRAZ, AND E. A. CHIQUITO. 2017. How many species of mammals are there in Brazil? New records of rare rodents (Rodentia: Cricetidae: Sigmodontinae) from Amazonia raise the current known diversity. Peerj 5: e 4071." type="journal article" year="2017">Percequillo et al. 2017</bibRefCitation>
). Despite the paucity of records for
<taxonomicName id="81A644984F218D5DFF4AFA0AFF1647E8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[144,255,1449,1472]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFF4AFA0AFF1647E8" box="[144,255,1449,1472]" italics="true" pageId="12" pageNumber="135">Rhagomys</emphasis>
</taxonomicName>
, its geographic range now rivals that of all 44 species of
<taxonomicName id="81A644984F218D5DFF2BFA6AFE9547C8" authorityName="Coues" authorityYear="1884" box="[241,380,1481,1504]" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFF2BFA6AFE9547C8" box="[241,380,1481,1504]" italics="true" pageId="12" pageNumber="135">Thomasomys</emphasis>
</taxonomicName>
(mammaldiversity.org) in geographic spread and biome occupation.
</paragraph>
<paragraph id="46193F1B4F218D5DFF71F9AAFEF445C8" blockId="12.[144,814,201,2016]" pageId="12" pageNumber="135">
The sigmodontine rodents appear to have diversified in the late Miocene (9.36 million years ago (mya);
<bibRefCitation id="223742EA4F218D5DFD7CF98AFF2C4448" author="VILELA, J. F. &amp; B. MELLO &amp; C. M. VOLOCH &amp; C. G. SCHRAGO" pageId="12" pageNumber="135" pagination="249 - 256" refId="ref14719" refString="VILELA, J. F., B. MELLO, C. M. VOLOCH, AND C. G. SCHRAGO. 2013. Sigmodontine rodents diversified in South America prior to the complete rise of the Panamanian Isthmus. Journal of Zoological Systematics and Evolutionary Research 52: 249 - 256." type="journal article" year="2013">Vilela et al. 2013</bibRefCitation>
;
<bibRefCitation id="223742EA4F218D5DFF09F9EAFE664448" author="PARADA, A. &amp; G. D'ELIA &amp; R. E. PALMA" box="[211,399,1609,1632]" pageId="12" pageNumber="135" pagination="172" refId="ref13667" refString="PARADA, A., G. D'ELIA, AND R. E. PALMA. 2015. The influence of ecological and geographical context in the radiation of Neotropical sigmodontine rodents. BMC Evolutionary Biology 15: 172." type="journal article" year="2015">Parada et al. 2015</bibRefCitation>
). From their origination 11.48.5 mya, sigmodontines began to colonize South America 7.966.49 mya (Steppan and Schenk 2017). The
<taxonomicName id="81A644984F218D5DFDF4F92AFD264488" authorityName="Steadman and Ray" authorityYear="1982" box="[558,719,1673,1696]" class="Mammalia" family="Cricetidae" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="137" phylum="Chordata" rank="tribe" tribe="Thomasomyini">Thomasomyini</taxonomicName>
radiated in northern South America and quickly colonized the Andean zones and adapted to Andean ecosystems (
<bibRefCitation id="223742EA4F218D5DFD8CF96AFCF444C8" author="MAESTRI, R. &amp; N. S. UPHAM &amp; B. D. PATTERSON" box="[598,797,1737,1760]" pageId="12" pageNumber="135" pagination="683 - 695" refId="ref13344" refString="MAESTRI, R., N. S. UPHAM, AND B. D. PATTERSON. 2019. Tracing the diversification history of a Neogene rodent invasion into South America. Ecography 42: 683 - 695." type="journal article" year="2019">Maestri et al. 2019</bibRefCitation>
); the divergence of
<taxonomicName id="81A644984F218D5DFE80F94AFE204528" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[346,457,1769,1792]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFE80F94AFE204528" box="[346,457,1769,1792]" italics="true" pageId="12" pageNumber="135">Rhagomys</emphasis>
</taxonomicName>
from other thomasomyines was dated at 6.5 million years by Schenk and Steppan (2018). Subsequently,
<taxonomicName id="81A644984F218D5DFEE3F88AFE414568" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[313,424,1833,1856]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFEE3F88AFE414568" box="[313,424,1833,1856]" italics="true" pageId="12" pageNumber="135">Rhagomys</emphasis>
</taxonomicName>
has acquired many remarkable apomorphies (Luna and Patterson 2003;
<bibRefCitation id="223742EA4F218D5DFDB3F8EAFF3B45A8" author="PERCEQUILLO, A. R. &amp; F. P. TIRELLI &amp; F. MICHALSKI &amp; E. EIZIRIK" pageId="12" pageNumber="135" pagination="195 - 199" refId="ref14017" refString="PERCEQUILLO, A. R., F. P. TIRELLI, F. MICHALSKI, AND E. EIZIRIK. 2011 a. The genus Rhagomys (Thomas 1917) (Rodentia, Cricetidae, Sigmodontinae) in South America: morphological considerations, geographic distribution and zoogeographic comments. Mammalia 75: 195 - 199." type="journal article" year="2011">Percequillo et al. 2011a</bibRefCitation>
), including its elongate facial vibrissae, a nail on the hallux, transverse grooves on digits tipped by crescent-shaped depressions with short, embedded claws, and its molar cusp arrangement.
</paragraph>
<paragraph id="46193F1B4F218D5DFCA2FF6AFBE543C8" blockId="12.[861,1532,201,1344]" pageId="12" pageNumber="135">
The new record of
<taxonomicName id="81A644984F218D5DFB97FF6AFB5542C8" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1101,1212,201,224]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFB97FF6AFB5542C8" box="[1101,1212,201,224]" italics="true" pageId="12" pageNumber="135">Rhagomys</emphasis>
</taxonomicName>
in
<collectingCountry id="3EB17F8B4F218D5DFB39FF6AFAD542C8" box="[1251,1340,201,224]" name="Ecuador" pageId="12" pageNumber="135">Ecuador</collectingCountry>
calls attention to the need for additional field surveys and taxonomic studies on rodents and other mammal groups. The Ecuadorian Andes are one of the most diverse areas for
<taxonomicName id="81A644984F218D5DFB03FE8AFA944368" authorityName="Steadman and Ray" authorityYear="1982" box="[1241,1405,297,320]" class="Mammalia" family="Cricetidae" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="137" phylum="Chordata" rank="tribe" tribe="Thomasomyini">Thomasomyini</taxonomicName>
, as well as for other tribes of
<taxonomicName id="81A644984F218D5DFBC1FEEAFB554348" box="[1051,1212,329,352]" pageId="12" pageNumber="123" rank="subFamily" subFamily="Sigmodontinae">Sigmodontinae</taxonomicName>
(Maestri and Patterson 2016). In addition, the Ecuadorean Andes are a locus of high endemism and represent the continents “hottest” hotspot for rodent species with highly restricted geographic ranges (Maestri and Patterson 2016).
</paragraph>
<paragraph id="46193F1B4F218D5DFCA2FE4AFBD44628" blockId="12.[861,1532,201,1344]" pageId="12" pageNumber="135">
The natural history and ecology of
<taxonomicName id="81A644984F218D5DFB2DFE4AFA5F4028" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1271,1462,489,512]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F218D5DFB2DFE4AFA5F4028" box="[1271,1462,489,512]" italics="true" pageId="12" pageNumber="135">R. septentrionalis</emphasis>
</taxonomicName>
is uncertain because only a single specimen is known, thus there is a lack of knowledge about most of its habits. Its locomotory habits likely resemble those of other species of
<taxonomicName id="81A644984F218D5DFA5EFDEAFA1D4048" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[1412,1524,585,608]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFA5EFDEAFA1D4048" box="[1412,1524,585,608]" italics="true" pageId="12" pageNumber="135">Rhagomys</emphasis>
</taxonomicName>
, because it shares the same characteristics of the hands, feet, and tail (Luna and Patterson 2003;
<bibRefCitation id="223742EA4F218D5DFB3FFD2AFA034088" author="PERCEQUILLO, A. R. &amp; M. WEKSLER &amp; L. P. COSTA" box="[1253,1514,649,672]" pageId="12" pageNumber="135" pagination="357 - 390" refId="ref14076" refString="PERCEQUILLO, A. R., M. WEKSLER, AND L. P. COSTA. 2011 b. A new genus and species of rodent from the Brazilian Atlantic Forest (Rodentia, Cricetidae, Sigmodontinae, Oryzomyini), with comments on oryzomyine biogeography. Zoological Journal of the Linnean Society 161: 357 - 390." type="journal article" year="2011">Percequillo et al. 2011b</bibRefCitation>
). This genus has short, straight claws—not curved ones—and a narrow metatarsus, as in the arboreal-climbing rodents
<taxonomicName id="81A644984F218D5DFC87FD4AFC354128" authorityName="Tschudi" authorityYear="1845" box="[861,988,745,768]" class="Mammalia" family="Cricetidae" genus="Rhipidomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFC87FD4AFC354128" box="[861,988,745,768]" italics="true" pageId="12" pageNumber="135">Rhipidomys</emphasis>
</taxonomicName>
and
<taxonomicName id="81A644984F218D5DFBC2FD4AFA384128" authority="(Rivas-Rodriguez et al. 2010)" baseAuthorityName="Rivas-Rodriguez" baseAuthorityYear="2010" box="[1048,1489,745,768]" class="Mammalia" family="Cricetidae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFBC2FD4AFB924128" box="[1048,1147,745,768]" italics="true" pageId="12" pageNumber="135">Oecomys</emphasis>
(
<bibRefCitation id="223742EA4F218D5DFB54FD4AFA2F4128" author="RIVAS-RODRIGUEZ, B. A. &amp; G. D'ELIA &amp; O. LINARES" box="[1166,1478,745,768]" pageId="12" pageNumber="135" pagination="97 - 109" refId="ref14175" refString="RIVAS-RODRIGUEZ, B. A., G. D'ELIA, AND O. LINARES. 2010. Diferenciacion morfologica en sigmodontinos (Rodentia: Cricetidae) de las guayanas venezolanas con relacion a su locomocion y habitat. Mastozoologia Neotropical 17: 97 - 109." type="journal article" year="2010">Rivas-Rodríguez et al. 2010</bibRefCitation>
)
</taxonomicName>
. Its short legs and bulging finger tips without claws resemble those of some genera of arboreal marsupials and primates.
<taxonomicName id="81A644984F218D5DFC87FCEAFC244148" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[861,973,841,864]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="74D2E3094F218D5DFC87FCEAFC244148" box="[861,973,841,864]" italics="true" pageId="12" pageNumber="135">Rhagomys</emphasis>
</taxonomicName>
also has been captured in traps placed on vegetation a meter or two aboveground (Luna and Patterson 2003). However, all these
<taxonomicName id="81A644984F218D5DFBF4FC2AFB1A4188" box="[1070,1267,905,928]" pageId="12" pageNumber="135">
<emphasis id="74D2E3094F218D5DFBF4FC2AFB774188" box="[1070,1182,905,928]" italics="true" pageId="12" pageNumber="135">Rhagomys</emphasis>
species
</taxonomicName>
are likely to have much broader ecological habits and ranges than are currently documented (
<bibRefCitation id="223742EA4F218D5DFC67FC6AFB5741C8" author="PERCEQUILLO, A. R. &amp; F. P. TIRELLI &amp; F. MICHALSKI &amp; E. EIZIRIK" box="[957,1214,969,992]" pageId="12" pageNumber="135" pagination="195 - 199" refId="ref14017" refString="PERCEQUILLO, A. R., F. P. TIRELLI, F. MICHALSKI, AND E. EIZIRIK. 2011 a. The genus Rhagomys (Thomas 1917) (Rodentia, Cricetidae, Sigmodontinae) in South America: morphological considerations, geographic distribution and zoogeographic comments. Mammalia 75: 195 - 199." type="journal article" year="2011">Percequillo et al. 2011a</bibRefCitation>
), simply because we have so few records of them.
</paragraph>
</subSubSection>
<subSubSection id="0EBC6C904F218D5DFCA2FBAAFA154768" pageId="12" pageNumber="135" type="biology_ecology">
<paragraph id="46193F1B4F218D5DFCA2FBAAFA154768" blockId="12.[861,1532,201,1344]" pageId="12" pageNumber="135">
<taxonomicName id="81A644984F218D5DFCA2FBAAFB674608" authorityName="Moreno Cárdenas &amp; Tinoco &amp; Albuja &amp; Patterson" authorityYear="2021" box="[888,1166,1033,1056]" class="Mammalia" family="Cricetidae" genus="Rhagomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="74D2E3094F218D5DFCA2FBAAFB674608" box="[888,1166,1033,1056]" italics="true" pageId="12" pageNumber="135">Rhagomys septentrionalis</emphasis>
</taxonomicName>
<materialsCitation id="F6CE35464F218D5DFB4CFBAAFA114768" collectionCode="MEPN" pageId="12" pageNumber="135" specimenCode="MEPN 10898" specimenCount="1" typeStatus="holotype">
was found living in a very mesic habitat crossed by many streams, raising the possibility that it might move in rivers. The
<typeStatus id="991D81B94F218D5DFBA8FBEAFB264648" box="[1138,1231,1097,1120]" pageId="12" pageNumber="135" type="holotype">holotype</typeStatus>
was actually captured in the middle of a river, although it lacks structures (interdigital webbing or stiff hairs along the margins of the pes or on the tail) typically present in semiaquatic rodents (
<bibRefCitation id="223742EA4F218D5DFAF4FB0AFA4846E8" author="VOSS, R. S." box="[1326,1441,1193,1216]" pageId="12" pageNumber="135" pagination="259 - 493" refId="ref14810" refString="VOSS, R. S. 1988. Systematics and ecology of ichthyomyine rodents (Muroidea): patterns of morphological evolution in a small adaptive radiation. Bulletin of the American Museum of Natural History 188: 259 - 493." type="journal article" year="1988">Voss 1988</bibRefCitation>
; Kerbis Peterhans and Patterson 1995). Although it could be natatorial, it seems equally probable that
<specimenCode id="160097604F218D5DFB7FFB4AFAD04728" box="[1189,1337,1257,1280]" collectionCode="MEPN" pageId="12" pageNumber="135">MEPN 10898</specimenCode>
might have fallen into the river accidentally, while climbing on riparian vegetation. Only additional observations can clarify its natural history
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>