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<document ID-DOI="http://dx.doi.org/10.3897/alpento.6.80349" ID-GBIF-Dataset="6a6dbf8e-2456-4ac3-b685-4509c0dbff9e" ID-Pensoft-Pub="2535-0889-6-13" ID-Pensoft-UUID="7266BBB11F3C52A397FD698561E728F0" ID-ZooBank="CAAB885C7DB84C4B9D0153B07656840B" ModsDocID="2535-0889-6-13" checkinTime="1652359712937" checkinUser="pensoft" docAuthor="Reyes-Hernandez, Jose L., Hansen, Aslak Kappel &amp; Solodovnikov, Alexey" docDate="2022" docId="24C4CA6F71AD58349C4DE6144886BE14" docLanguage="en" docName="AlpEntomol 6: 13-18" docOrigin="Alpine Entomology 6" docPubDate="2022-05-11" docSource="http://dx.doi.org/10.3897/alpento.6.80349" docTitle="Ikaros navarretei Reyes-Hernández &amp; Hansen &amp; Solodovnikov 2022, sp. nov." docType="treatment" docUuid="2C1E9003-82DD-4CF6-900D-16451E23B9E4" docUuidSource="ZooBank" docVersion="2" id="7266BBB11F3C52A397FD698561E728F0" lastPageNumber="13" masterDocId="7266BBB11F3C52A397FD698561E728F0" masterDocTitle="Ikaros navarretei (Coleoptera, Staphylinidae, Staphylininae), a new apterous rove beetle species from high elevations in Colombia" masterLastPageNumber="18" masterPageNumber="13" pageNumber="13" updateTime="1652360334029" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Ikaros navarretei (Coleoptera, Staphylinidae, Staphylininae), a new apterous rove beetle species from high elevations in Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reyes-Hernandez, Jose L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4726-4439</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, Copenhagen, 2100, Denmark</mods:affiliation>
<mods:nameIdentifier type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hansen, Aslak Kappel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2089-7233</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, Copenhagen, 2100, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Solodovnikov, Alexey</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2031-849X</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, Copenhagen, 2100, Denmark</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Alpine Entomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-05-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>18</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/alpento.6.80349</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/alpento.6.80349</mods:identifier>
<mods:identifier type="Pensoft-Pub">2535-0889-6-13</mods:identifier>
<mods:identifier type="ZooBank">CAAB885C7DB84C4B9D0153B07656840B</mods:identifier>
<mods:identifier type="Pensoft-UUID">7266BBB11F3C52A397FD698561E728F0</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:2C1E9003-82DD-4CF6-900D-16451E23B9E4" httpUri="http://treatment.plazi.org/id/24C4CA6F71AD58349C4DE6144886BE14" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection pageId="0" pageNumber="13" type="nomenclature">
<paragraph pageId="0" pageNumber="13">
<taxonomicName LSID="http://zoobank.org/2C1E9003-82DD-4CF6-900D-16451E23B9E4" authority="Reyes-Hernández &amp; Hansen &amp; Solodovnikov, 2022" authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei" status="sp. nov.">Ikaros navarretei</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="type locality">
<paragraph pageId="0" pageNumber="13">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="13">
Colombia, Cauca, Silvia,
<geoCoordinate degrees="2.5889" direction="north" orientation="latitude" precision="5" value="2.5889">2.5889</geoCoordinate>
,
<geoCoordinate degrees="76.24886" direction="west" orientation="longitude" precision="1" value="-76.24886">-76.24886</geoCoordinate>
, 3400 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="generic placement">
<paragraph pageId="0" pageNumber="13">Generic placement.</paragraph>
<paragraph pageId="0" pageNumber="13">
Our specimen fully agrees with the generic diagnosis and description provided by
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2021.744.1303" author="Chatzimanolis, S" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="13" pagination="67 - 82" refId="B1" refString="Chatzimanolis, S, Brunke, AJ, 2021. A new apterous rove beetle genus (Coleoptera: Staphylinidae) from the Northern Andes with an assessment of its phylogenetic position. European Journal of Taxonomy 744: 67 - 82, DOI: https://doi.org/10.5852/ejt.2021.744.1303" title="A new apterous rove beetle genus (Coleoptera: Staphylinidae) from the Northern Andes with an assessment of its phylogenetic position." url="https://doi.org/10.5852/ejt.2021.744.1303" volume="744" year="2021">Chatzimanolis and Brunke (2021)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="diagnosis">
<paragraph pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov. can be distinguished from all other species in this genus by the combination of the presence of an arch-like carina on terga III-V; the meshed to isodiametric microsculpture on the dorsal surface of the head, thorax, and mesoscutellum; antennomeres with crown-like macrosetae shorter than the length of antennomere.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="etymology">
<paragraph pageId="0" pageNumber="13">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="13">
The species epithet is in recognition (patronymic) of
<normalizedToken originalValue="José">Jose</normalizedToken>
Luis Navarrete Heredia for his contribution to the knowledge of the family
<taxonomicName class="Insecta" family="Staphylinidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
and training of many coleopterologists in Latin America.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="type material">
<paragraph pageId="0" pageNumber="13">Type material.</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">
<emphasis italics="true" pageId="0" pageNumber="13">Holotype</emphasis>
.
</emphasis>
Male, point mounted, with genitalia in a separate microvial, with labels as follows: &quot;COLOMB. [COLOMBIA], 20 km E Silvia, Cauca, [~
<geoCoordinate degrees="2.5889" direction="north" orientation="latitude" precision="5" value="2.5889">2.5889</geoCoordinate>
,
<geoCoordinate degrees="76.24886" direction="west" orientation="longitude" precision="1" value="-76.24886">-76.24886</geoCoordinate>
, 3400 m a.s.l.] VII.16.1970 [16 June 1970], II, 000' J.M. Campbell / Xantopygina sic? gen. det. Newton 1998 / HOLOTYPE
<taxonomicName authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
<normalizedToken originalValue="Reyes-Hernández">Reyes-Hernandez</normalizedToken>
, Hansen and Solodovnikov, des.
<normalizedToken originalValue="Reyes-Hernández">Reyes-Hernandez</normalizedToken>
, Hansen and Solodovnikov 2022&quot; CNC.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="description">
<paragraph pageId="0" pageNumber="13">Description.</paragraph>
<paragraph pageId="0" pageNumber="13">
Habitus as in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1A</figureCitation>
. Total body length 10.06 mm. Forebody length 5.25 mm long.
</paragraph>
<paragraph pageId="0" pageNumber="13">Coloration of body reddish-brown, with abdomen having undertones of metallic green-brown.</paragraph>
<paragraph pageId="0" pageNumber="13">
Head subrectangular, slightly wider than long, HW/HL ratio = 1.1. Epicranium with numerous large punctures, except impunctate center; punctures not contiguous, the distance between punctures typically equals the width of 1-2 punctures; with transversely meshed to isodiametric microsculpture (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1B</figureCitation>
). Labial palpus with palpomere 3 (apical) widest before the apex. Antennomeres with crown-like macrosetae shorter than length of antennomere.
</paragraph>
<caption doi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 1.</emphasis>
Dorsal view of
<taxonomicName authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="13">A.</emphasis>
Habitus;
<emphasis bold="true" pageId="0" pageNumber="13">B.</emphasis>
Close up of head punctation and meshed to isodiametric microsculpture;
<emphasis bold="true" pageId="0" pageNumber="13">C.</emphasis>
Close up of pronotum punctation and meshed to isodiametric microsculpture;
<emphasis bold="true" pageId="0" pageNumber="13">D.</emphasis>
Close up of mesoscutellum and elytra punctation and microsculpture;
<emphasis bold="true" pageId="0" pageNumber="13">E.</emphasis>
Close up of abdominal tergite III with arch-like carina highlighted by white dashed line.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="13">
Pronotum longer than wide, PW/PL ratio = 0.9; surface of pronotum with a median impunctate area as wide as 3-5 punctures; with multiple rows of irregular punctures in addition to rows flanking impunctate center; with meshed to isodiametric microsculpture (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="13">
Elytra shorter than pronotum, EL/PL ratio = 0.83. Elytra with large, deep punctures, the distance between punctures equals to width of 0.5-2 punctures; only with isodiametric microsculpture at mesoscutellum, disc of elytra with micropunctuation (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="13">
Abdominal terga III-V with arch-like carina (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1E</figureCitation>
), arch-like carina on terga IV and V nearly straight. Male secondary sexual structures with very shallow emargination on sternum VII; with deep, broad V-shaped emargination on sternum VIII; borders of emargination on sterna VII and VIII appearing
<normalizedToken originalValue="shaved">'shaved'</normalizedToken>
(with no setae); lateral tergal sclerites IX subcylindrical and the same length as sternum IX; tergum X subtruncate medio-apically; sternum IX with basal portion symmetrical, about 0.57
<normalizedToken originalValue="×">x</normalizedToken>
as long as distal portion and subtruncate apically.
</paragraph>
<paragraph pageId="0" pageNumber="13">
Aedeagus as in Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus of Ikaros navarretei sp. nov. A. Aedeagus in parameral view; B. Paramere in antiparameral view with peg setae; C. Aedeagus in lateral view. Scale bar: 1 mm." figureDoi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13">2</figureCitation>
; in parameral view apex of paramere nearly reaching apex of median lobe (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus of Ikaros navarretei sp. nov. A. Aedeagus in parameral view; B. Paramere in antiparameral view with peg setae; C. Aedeagus in lateral view. Scale bar: 1 mm." figureDoi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13">2A</figureCitation>
); paramere broadest at the middle of its length, converging to rounded tip (violin-like shape), characteristic arrangement of peg setae in two sublateral rows containing 4-5 setae with an additional single setae closer to apex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus of Ikaros navarretei sp. nov. A. Aedeagus in parameral view; B. Paramere in antiparameral view with peg setae; C. Aedeagus in lateral view. Scale bar: 1 mm." figureDoi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13">2B</figureCitation>
); in lateral view paramere narrower apically; median lobe in dorsal view narrowing to small, rounded apex; in lateral view, median lobe becoming narrower near apex, with one subapical tooth (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus of Ikaros navarretei sp. nov. A. Aedeagus in parameral view; B. Paramere in antiparameral view with peg setae; C. Aedeagus in lateral view. Scale bar: 1 mm." figureDoi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13">2C</figureCitation>
).
</paragraph>
<caption doi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 2.</emphasis>
Aedeagus of
<taxonomicName authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="13">A.</emphasis>
Aedeagus in parameral view;
<emphasis bold="true" pageId="0" pageNumber="13">B.</emphasis>
Paramere in antiparameral view with peg setae;
<emphasis bold="true" pageId="0" pageNumber="13">C.</emphasis>
Aedeagus in lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="distribution">
<paragraph pageId="0" pageNumber="13">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="13">
Known only from the type locality near Silvia in the Cauca Department, Colombia (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of Ikaros. Elevation colored from green (low) across brown (middle) to white (high). Black square (■) Ikaros apterus. Yellow star (★) Ikaros navarretei sp. nov .. Black circle (●) Ikaros paramo. Black triangle (▲) Ikaros polygonos." figureDoi="10.3897/alpento.6.80349.figure3" httpUri="https://binary.pensoft.net/fig/682342" pageId="0" pageNumber="13">3</figureCitation>
).
</paragraph>
<caption doi="10.3897/alpento.6.80349.figure3" httpUri="https://binary.pensoft.net/fig/682342" pageId="0" pageNumber="13" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 3.</emphasis>
Distribution of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ikaros." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ikaros." order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros.</emphasis>
</taxonomicName>
Elevation colored from green (low) across brown (middle) to white (high). Black square (■)
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros apterus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="apterus">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros apterus</emphasis>
</taxonomicName>
. Yellow star (★)
<taxonomicName authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov.. Black circle (●)
<taxonomicName authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros paramo" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="paramo">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros paramo</emphasis>
</taxonomicName>
. Black triangle (▲)
<taxonomicName authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros polygonos" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="polygonos">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros polygonos</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="notes">
<paragraph pageId="0" pageNumber="13">Note.</paragraph>
<paragraph pageId="0" pageNumber="13">
In
<taxonomicName authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov. a structure which we call the arch-like carina on the abdominal terga for the sake of compatibility with other literature on
<taxonomicName authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
, appears to be somewhat similar to a structure usually called in the
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="0" pageNumber="13" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
literature as the posterior basal tergal carina (PBTC). This is a very different condition from other species of
<taxonomicName authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Ikaros</emphasis>
</taxonomicName>
and, as far as we are aware, all other
<taxonomicName authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
where the arch-like carina is more of a curved fragment that does not reach the spiracles as in the PBTC.
</paragraph>
</subSubSection>
</treatment>
</document>