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<document id="5AEFBEC410163F5F042893131F5600E7" ID-CLB-Dataset="37307" ID-DOI="10.5281/zenodo.247206" ID-GBIF-Dataset="cd615cd7-ecb8-4cc3-b074-5ddfab94649a" ID-Zenodo-Dep="247206" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484571130511" checkinUser="plazi" docAuthor="Ivan Löbl &amp; Ryo Ogawa" docDate="2016" docId="0C70F70CD750FFC2FF7BAE34DC68474F" docLanguage="en" docName="LinzerbiolBeitr.48.2.1339-1492.pdf" docOrigin="Linzer biologische Beiträge 48 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Scaphosoma Heller 1917" docType="treatment" docVersion="10" lastPageNumber="1435" masterDocId="F0498F74D70FFFA2FFC9AB07DD0F4264" masterDocTitle="On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH" masterLastPageNumber="1492" masterPageNumber="1339" pageNumber="1434" updateTime="1698435770511" updateUser="ExternalLinkService">
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<mods:title id="D63513B0B28C18D25D21AD9A722FBAEA">On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH</mods:title>
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<mods:name id="1492DF2152D05B51A57685D6EE3A9F98" type="personal">
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<mods:namePart id="83B29C1E358CCFE8FBBCB36D005F81C8">Ivan Löbl</mods:namePart>
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<mods:namePart id="EEB9F4E20343CC0FFF3AAFE9D704D8AC">Ryo Ogawa</mods:namePart>
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<mods:title id="CDF88F5FA18C9502BF1087AFFCD6E9CE">Linzer biologische Beiträge</mods:title>
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<mods:date id="3F0332F11C0CC45E91E5282452A9160D">2016</mods:date>
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<mods:number id="3C3011F50C77EF028A7D8FE96487F889">48</mods:number>
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<mods:identifier id="6BC2112C3BFC3D56DD60901351999F25" type="DOI">10.5281/zenodo.247206</mods:identifier>
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<treatment id="0C70F70CD750FFC2FF7BAE34DC68474F" ID-DOI="http://doi.org/10.5281/zenodo.4390948" ID-GBIF-Taxon="127598749" ID-Zenodo-Dep="4390948" LSID="urn:lsid:plazi:treatment:0C70F70CD750FFC2FF7BAE34DC68474F" httpUri="http://treatment.plazi.org/id/0C70F70CD750FFC2FF7BAE34DC68474F" lastPageId="96" lastPageNumber="1435" pageId="95" pageNumber="1434">
<subSubSection id="CCC31591D750FFFDFF7BAE34DE7E472D" box="[178,881,1331,1353]" pageId="95" pageNumber="1434" type="nomenclature">
<paragraph id="8466461AD750FFFDFF7BAE34DE7E472D" blockId="95.[178,1086,1331,1603]" box="[178,881,1331,1353]" pageId="95" pageNumber="1434">
<emphasis id="B6AD9A08D750FFFDFF7BAE34DC04472C" bold="true" box="[178,267,1331,1352]" pageId="95" pageNumber="1434">5.2.21.3.</emphasis>
<taxonomicName id="43D93D99D750FFFDFED8AE34DF89472C" ID-CoL="8XPR8" authorityName="Heller" authorityYear="1917" box="[273,646,1331,1353]" class="Insecta" family="Staphylinidae" genus="Scaphosoma" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="1434" phylum="Arthropoda" rank="genus">
<emphasis id="B6AD9A08D750FFFDFED8AE34DF48472C" bold="true" box="[273,583,1331,1353]" pageId="95" pageNumber="1434">
<emphasis id="B6AD9A08D750FFFDFED8AE34DCE6472D" bold="true" box="[273,489,1331,1353]" italics="true" pageId="95" pageNumber="1434">Scaphosoma tricolor</emphasis>
HELLER
</emphasis>
,
<emphasis id="B6AD9A08D750FFFDFD9AAE34DF89472C" bold="true" box="[595,646,1331,1352]" pageId="95" pageNumber="1434">1917</emphasis>
</taxonomicName>
<emphasis id="B6AD9A08D750FFFDFD45AE34DE7E472D" bold="true" box="[652,881,1331,1353]" pageId="95" pageNumber="1434">(Figs 38, 39, 234-237)</emphasis>
</paragraph>
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<subSubSection id="CCC31591D750FFFDFF7BAE54DFB947A7" pageId="95" pageNumber="1434" type="reference_group">
<paragraph id="8466461AD750FFFDFF7BAE54DE9247F2" blockId="95.[178,1086,1331,1603]" pageId="95" pageNumber="1434">
<taxonomicName id="43D93D99D750FFFDFF7BAE54DCF0470C" ID-CoL="8XPR8" box="[178,511,1363,1384]" class="Insecta" family="Staphylinidae" genus="Scaphosoma" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="1434" phylum="Arthropoda" rank="genus">
<emphasis id="B6AD9A08D750FFFDFF7BAE54DC7F470C" box="[178,368,1363,1384]" italics="true" pageId="95" pageNumber="1434">Scaphosoma tricolor</emphasis>
HELLER, 1917
</taxonomicName>
: 46. Lectotype 6, SMTD; type locality: Luzon, Mt. Makiling. Lectotype designation:
<bibRefCitation id="E0483BEBD750FFFDFE44AE6DDF0A471A" author="LOBL I." box="[397,517,1385,1407]" journalOrPublisher="Mitt. Schweiz. ent. Ges." pageId="95" pageNumber="1434" pagination="125 - 132" part="43" refString="LOBL I. (1970 a): Uber einige Scaphidiidae (Coleoptera) aus der Sammlung des Museum National d'Histoire Naturelle de Paris. - Mitt. Schweiz. ent. Ges. 43: 125 - 132." title="Uber einige Scaphidiidae (Coleoptera) aus der Sammlung des Museum National d'Histoire Naturelle de Paris" type="journal article" year="1970" yearSuffix="a">LÖBL 1970a</bibRefCitation>
: 126; apical margin of elytra:
<bibRefCitation id="E0483BEBD750FFFDFCE1AE6DDE90471A" author="LOBL I." box="[808,927,1385,1407]" journalOrPublisher="Steenstrupia" pageId="95" pageNumber="1434" pagination="247 - 253" part="1" refString="LOBL I. (1971 a): Scaphidiidae der Noona Dan Expedition nach den Philippinen und Bismarck Inseln (Insecta, Coleoptera). - Steenstrupia 1: 247 - 253." title="Scaphidiidae der Noona Dan Expedition nach den Philippinen und Bismarck Inseln (Insecta, Coleoptera)" type="journal article" year="1971" yearSuffix="a">LÖBL 1971a</bibRefCitation>
: 249. Additional published records. Palawan: Pinigisan; Balabac: Dalawan Bay (
<bibRefCitation id="E0483BEBD750FFFDFD3AAE87DE9947F2" author="LOBL I." box="[755,918,1408,1430]" journalOrPublisher="Steenstrupia" pageId="95" pageNumber="1434" pagination="247 - 253" part="1" refString="LOBL I. (1971 a): Scaphidiidae der Noona Dan Expedition nach den Philippinen und Bismarck Inseln (Insecta, Coleoptera). - Steenstrupia 1: 247 - 253." title="Scaphidiidae der Noona Dan Expedition nach den Philippinen und Bismarck Inseln (Insecta, Coleoptera)" type="journal article" year="1971" yearSuffix="a">LÖBL 1971a: 250</bibRefCitation>
)
</paragraph>
<paragraph id="8466461AD750FFFDFC55AE87DFB947A7" blockId="95.[178,1086,1331,1603]" pageId="95" pageNumber="1434">
.
<taxonomicName id="43D93D99D750FFFDFF7BAE90DCF147C8" ID-CoL="6Y737" authority="PIC, 1920b: 3" authorityName="PIC" authorityPageNumber="3" authorityYear="1920" box="[178,510,1431,1453]" class="Insecta" family="Staphylinidae" genus="Scaphosoma" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="1434" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B6AD9A08D750FFFDFF7BAE90DC6947C8" box="[178,358,1431,1452]" italics="true" pageId="95" pageNumber="1434">Scaphosoma latum</emphasis>
<bibRefCitation id="E0483BEBD750FFFDFEBAAE90DCEE47C9" author="PIC M." box="[371,481,1431,1453]" journalOrPublisher="Melang. exot. - ent" pageId="95" pageNumber="1434" pagination="1 - 28" part="32" refString="PIC M. (1920 b): Nouveautes diverses. - Melang. exot. - ent. 32: 1 - 28." title="Nouveautes diverses" type="journal article" year="1920" yearSuffix="b">PIC, 1920b</bibRefCitation>
: 3
</taxonomicName>
. Lectotype 6, MNHN; type locality: Luzon, Los Banos. Lectotype designation and synonymy:
<bibRefCitation id="E0483BEBD750FFFDFDD8AEA9DF8947A6" author="LOBL I." box="[529,646,1453,1475]" journalOrPublisher="Mitt. Schweiz. ent. Ges." pageId="95" pageNumber="1434" pagination="125 - 132" part="43" refString="LOBL I. (1970 a): Uber einige Scaphidiidae (Coleoptera) aus der Sammlung des Museum National d'Histoire Naturelle de Paris. - Mitt. Schweiz. ent. Ges. 43: 125 - 132." title="Uber einige Scaphidiidae (Coleoptera) aus der Sammlung des Museum National d'Histoire Naturelle de Paris" type="journal article" year="1970" yearSuffix="a">LÖBL 1970a</bibRefCitation>
: 126.
</paragraph>
</subSubSection>
<subSubSection id="CCC31591D750FFC2FF7BAECADEEF41D8" lastPageId="96" lastPageNumber="1435" pageId="95" pageNumber="1434" type="materials_examined">
<paragraph id="8466461AD750FFC2FF7BAECADEEF41D8" blockId="95.[178,1086,1331,1603]" lastBlockId="96.[178,1086,214,1323]" lastPageId="96" lastPageNumber="1435" pageId="95" pageNumber="1434">
A d d i t i o n a l m a t e r i a l e x a m i n e d: Luzon, Lagunas Prov., Mt. Makiling, 4 0 0- 600 m, summit road, 19., 21-22., 26. and 28.XI.1995, leg. I. Löbl, 59 ex. (MHNG, BZLA); same but above Mad Springs, 400-700 m, 19-22.XI.1 995, leg. J. Kodada 14 ex. (MHNG); Luzon, Lagunas Prov., Los Banos, small river, 28.XI.1 995, J. Kodada &amp; B. Rigová, 1 ex. (MHNG); Luzon, Lagunas Prov., Mt. Banahaw above Kinabuhayan, 600-700 m, trail to Crystalino, 24- 25.XI.1995, leg. J. Kodada &amp; B. Rigová, 5 ex. (MHNG); Luzon, Laguna Prov., Mt. Makiling 4 km SE Los Banos, 4-12.V.197 7, leg. L.E. Watrous, 5 ex. (MHNG); Luzon, Los Banos, 1 ex. (SMTS); Luzon, Ifugao Prov., Mt. Polis, 1900 m, 4-5.VI.1977, leg. M. Satô, 1 ex. (MHNG); Palawan: Central, Cabayugan nr. Lion’s Cave, sea level, 1.XII.1995 I. Löbl, 1 ex. (MHNG); same but leg. J. Kodada, 2 ex. (MHNG); Palawan, Sabang, trail to Underground River, sea level, 30.XI.1995, leg. I. Löbl, 4 ex. (MHNG); Palawan, Sabang, 50-100 m, degraded forest on slope, leg. J. Kodada, 8 ex. (MHNG); Palawan, Sabang, N. of Mt. St. Paul, 11-13.VII.1977, leg. M. Satô, 3 ex. (MHNG); Palawan, Conception, sea level, 6-7.XII.1995, leg. I. Löbl, 6b ex. (MHNG); Palawan Central, Conception, large logs across Conception River, NE San Rafael, ca 20 m, 8.XII.1995, leg. J. Kodada &amp; B. Rigová, 4 ex. (MHNG); Palawan, Olangoan, 18 km NE San Rafael, sea level, 5- 6.XII.1995, leg. I. Löbl, 6 ex. (MHNG); same but 1.IX.1985, leg. M. Sakai, 5 ex. (EUMJ); Palawan, Binaluan, 1 ex. (SMTD); Palawan, Matalangao, 150 m, nr. Roxas, 29.VIII.1985, leg. M. Sakai, 1 ex. (EUMJ); Palawan, Roxas, region of Matalangao, 80 m, 5.IV.1983, leg. S. Nagai &amp; C. Lienhard, 1 ex. (MHNG); Palawan, Mantalingajan Range, S. slope Mt. Balabag, 2800 ft., 4- 17.V.47, leg. F.G. Werner, 3 ex. (FMNH); Palawan, Trident
<taxonomicName id="43D93D99D76FFFC2FD34A921DE224058" box="[765,813,550,572]" class="Magnoliopsida" family="Convolvulaceae" genus="Mina" higherTaxonomySource="CoL" kingdom="Plantae" order="Solanales" pageId="96" pageNumber="1435" phylum="Tracheophyta" rank="genus">Mina</taxonomicName>
, foot of Victoria Peak, 100 m, nr. Narra, 5.IX.1985, leg. M. Sakai, 2 ex. (EUMJ, MHNG); Leyte, Visca N Baybay, primary forest, 200-500 m, 22.II., 27.II. and 8.III.1991, leg. W. Schawaller et al., 6 ex. (SMNS, MHNG); same but secondary forest, 100-200 m, 27.II. and 1.III.1991, 2 ex. (SMNS, MHNG); Leyte, Tarragona, 30.VI.1945, leg. C.L. Remington, 1 ex. (FMNH); Masbate, Aroroy, 14.VIII.1918, leg. G. Böttcher, 2 ex. (ZMUB); Mindoro, 28 km S Calapan, Balete, 27-29.XI.2002, leg. H. Schillhammer, 1 ex. (NHMW); Bucas, Sorocco, X.1916, leg. G. Böttcher, 1 ex. (ZMUB); Mindanao, Surigao, 30-X.1915, leg. G. Böttcher, 3 ex. (ZMUB); Mindanao, Cotabato Prov., 50 km N. of Parang, 500 ft., 6.XII.1946, leg. F.G. Werner, 19 ex. (FMNH, MHNG); Mindanao, Cotabato Prov., Burungkot, Upi, 1500 ft., 6.I.1947, leg. F.G. Werner, 2 ex. (FMNH); Mindanao, Davao Prov., Sitio Taglawig, Maco, Tagum, nr. sea level, X.1946, leg. H. Hoogstraal &amp; D. Heyneman, 5 ex. (FMNH, MHNG); Mindanao, 25 km NW of Zamboanga, 800 m, Camp Susana, 28-30.IV.1996, leg. Bolm, 3 ex. (SMNS, MHNG); Mindanao, Bukidnon Prov., 5 km N of Malaybalay, 900 m, 12.V.1996, leg. Bolm, 1 ex (SMNS); Mindanao, 30 km E of Malaybalay, Busdi, 1000 m, 5-9.V.1996, leg. Bolm, 18 ex. (SMNS, MHNG); Mindanao, 15 km NW of Bislig, 6-7.VII.1977, leg. M. Satô, 4 ex. (MHNG); Mindanao, Todaya, 29.VII.1970, leg. M. Satô, 2 ex. (MHNG); Mindanao, Baracatan, 1500 m, 27-29.VI.1977, leg. M. Satô, 10 ex. (MHNG).
</paragraph>
</subSubSection>
<subSubSection id="CCC31591D76FFFC2FF7BA8CCDCC7419A" pageId="96" pageNumber="1435" type="distribution">
<paragraph id="8466461AD76FFFC2FF7BA8CCDCC7419A" blockId="96.[178,1086,214,1323]" pageId="96" pageNumber="1435">D i s t r i b u t i o n: Philippines: Luzon, Palawan, Balabac, Leyte, Masbate, Mindoro, Dinagat, Bucas, Mindanao.</paragraph>
</subSubSection>
<subSubSection id="CCC31591D76FFFC2FF7BAF09DC68474F" pageId="96" pageNumber="1435" type="diagnosis">
<paragraph id="8466461AD76FFFC2FF7BAF09DC68474F" blockId="96.[178,1086,214,1323]" pageId="96" pageNumber="1435">
C o m p a r a t i v e n o t e s: The almost entirely ochreous male pronotum with very narrowly darkened along basal margin is diagnostic for this species. The elytral colora- tion is somewhat more variable, and specimens from Palawan have the apical light area narrower than that exhibited by Luzon specimens. The aedeagal characters are for the first time illustrated in the present paper (
<figureCitation id="1CE25A9FD76FFFC2FDA3AF84DFF646FD" box="[618,761,1155,1177]" captionStart="Figs 234 - 237" captionStartId="148.[178,218,1490,1512]" captionTargetBox="[198,1056,240,1463]" captionTargetId="figure@148.[178,1085,212,1480]" captionTargetPageId="148" captionText="Figs 234 - 237: Aedeagus of Scaphisoma tricolor HELLER; (234), in dorsal view; (235), in lateral view, without proximal part of basal bulb, scale = 0.2 mm; (236, 237), internal sac in dorsal and lateral views, scale = 0.1 mm." httpUri="https://zenodo.org/record/247237/files/figure.png" pageId="96" pageNumber="1435">Figs 234-237</figureCitation>
). Though the number, position and shape of the sclerites of the internal sac are diagnostic, they may appear highly vari- able if internal sac not completely in repos. The shape of the long and robust ventral branch of the apical process of the median lobe and the internal sac with a small tuber- culate lobe are also diagnostic. The inner apical angle of female elytra is weakly crenu- lated, not dentate.
</paragraph>
</subSubSection>
</treatment>
</document>
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