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<mods:title id="A53CC5A796FE1FCEE05ED12D873DAB60">A survey of Hersiliidae (Arachnida: Araneae) of Iran with description of one new genus and two new species</mods:title>
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<taxonomicName id="4C779FC9FFEFFFC3FE6EF9EDFC1DAB74" ID-CoL="6LRMP" authority="Marusik &amp; Fet, 2009" authorityName="Marusik &amp; Fet" authorityYear="2009" box="[404,910,1608,1634]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="1458" phylum="Arthropoda" rank="species" species="sternbergsi">
<emphasis id="B9033858FFEFFFC3FE6EF9EDFD0AAB74" bold="true" box="[404,665,1608,1634]" inLineHeading="true" italics="true" pageId="12" pageNumber="1458" reason="3">Hersiliola sternbergsi</emphasis>
<bibRefCitation id="EFE699BBFFEFFFC3FD59F9ECFC1DAB74" author="Marusik YM &amp; Fet V" box="[675,910,1609,1634]" pageId="12" pageNumber="1458" pagination="75 - 114" refId="ref4820" refString="Marusik YM, Fet V. 2009. A survey of East Palearctic Hersiliola Thorell, 1870 (Araneae: Hersiliidae), with description of three new genera. Zookeys. 16: 75 - 114." type="journal article" year="2009">Marusik &amp; Fet, 2009</bibRefCitation>
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<treatmentCitation id="0AD6C25BFFEFFFC3FF5AF9DBFD23AB80" author="Marusik YM &amp; Fet V" box="[160,688,1662,1686]" page="86" pageId="12" pageNumber="1458" year="2009">
<taxonomicName id="4C779FC9FFEFFFC3FF5AF9DBFD23AB80" ID-CoL="6LRMP" authority="Marusik &amp; Fet, 2009: 86" authorityName="Marusik &amp; Fet" authorityPageNumber="86" authorityYear="2009" box="[160,688,1662,1686]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="1458" phylum="Arthropoda" rank="species" species="sternbergsi">
<emphasis id="B9033858FFEFFFC3FF5AF9DBFE15AB80" box="[160,390,1662,1686]" italics="true" pageId="12" pageNumber="1458">Hersiliola sternbergsi</emphasis>
<bibRefCitation id="EFE699BBFFEFFFC3FE69F9DBFD23AB80" author="Marusik YM &amp; Fet V" box="[403,688,1662,1686]" pageId="12" pageNumber="1458" pagination="75 - 114" refId="ref4820" refString="Marusik YM, Fet V. 2009. A survey of East Palearctic Hersiliola Thorell, 1870 (Araneae: Hersiliidae), with description of three new genera. Zookeys. 16: 75 - 114." type="journal article" year="2009">Marusik &amp; Fet, 2009: 86</bibRefCitation>
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, figs 1.1, 2.1, 3.2, 4.3, 5.3, 6.4
<emphasis id="B9033858FFEFFFC3FBEEF9DBFBB2AB80" box="[1044,1057,1662,1686]" italics="true" pageId="12" pageNumber="1458"></emphasis>
5, 7.7
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9, 8.2 (
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<specimenCount id="9D712FC3FFEFFFC3FEFFF905FE8AABAE" box="[261,281,1696,1720]" pageId="12" pageNumber="1458" type="female"></specimenCount>
).
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</paragraph>
<paragraph id="8BC8E44AFFEEFFC2FF5AFF2BFCA4ADB0" blockId="13.[160,823,142,166]" box="[160,823,142,166]" pageId="13" pageNumber="1459">
<treatmentCitationGroup id="AB67C364FFEEFFC2FF5AFF2BFCA4ADB0" box="[160,823,142,166]" pageId="13" pageNumber="1459">
<treatmentCitation id="0AD6C25BFFEEFFC2FF5AFF2BFD51ADB0" author="Mirshamsi O &amp; Marusik YM &amp; Jamili E" box="[160,706,142,166]" page="348" pageId="13" pageNumber="1459" year="2013">
<taxonomicName id="4C779FC9FFEEFFC2FF5AFF2BFD51ADB0" ID-CoL="6LRMP" authority=": Mirshamsi et al., 2013: 348" authorityName="Mirshamsi" authorityPageNumber="348" authorityYear="2013" box="[160,706,142,166]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="species" species="sternbergsi">
<emphasis id="B9033858FFEEFFC2FF5AFF2BFE11ADB0" box="[160,386,142,166]" italics="true" pageId="13" pageNumber="1459">Hersiliola sternbergsi</emphasis>
:
<bibRefCitation id="EFE699BBFFEEFFC2FE6AFF2BFD51ADB0" author="Mirshamsi O &amp; Marusik YM &amp; Jamili E" box="[400,706,142,166]" pageId="13" pageNumber="1459" pagination="347 - 352" refId="ref4893" refString="Mirshamsi O, Marusik YM, Jamili E. 2013. The spider family Hersiliidae Thorell, 1870 (Arachnida: Araneae) in Iran. Zool Middle East. 59: 347 - 352." type="journal article" year="2013">Mirshamsi et al., 2013: 348</bibRefCitation>
</taxonomicName>
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, fig. 1 (
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).
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</paragraph>
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<subSubSection id="C36DB7C1FFEEFFC2FF5AFF78FC66AC9C" pageId="13" pageNumber="1459" type="discussion">
<paragraph id="8BC8E44AFFEEFFC2FF5AFF78FE8DADE0" blockId="13.[160,639,221,280]" box="[160,286,221,246]" pageId="13" pageNumber="1459">
<heading id="D0805326FFEEFFC2FF5AFF78FE8DADE0" box="[160,286,221,246]" fontSize="11" level="3" pageId="13" pageNumber="1459" reason="5">
<emphasis id="B9033858FFEEFFC2FF5AFF78FE8DADE0" bold="true" box="[160,286,221,246]" inLineHeading="true" italics="true" pageId="13" pageNumber="1459" reason="3">Comments</emphasis>
</heading>
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Well described by
<bibRefCitation id="EFE699BBFFEEFFC2FE8EFEA5FDE8AC0E" author="Marusik YM &amp; Fet V" box="[372,635,256,280]" pageId="13" pageNumber="1459" pagination="75 - 114" refId="ref4820" refString="Marusik YM, Fet V. 2009. A survey of East Palearctic Hersiliola Thorell, 1870 (Araneae: Hersiliidae), with description of three new genera. Zookeys. 16: 75 - 114." type="journal article" year="2009">Marusik and Fet (2009)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC8E44AFFEEFFC2FF5AFEEAFF6FAC7E" blockId="13.[160,1013,335,394]" box="[160,252,335,360]" pageId="13" pageNumber="1459">
<heading id="D0805326FFEEFFC2FF5AFEEAFF6FAC7E" box="[160,252,335,360]" fontSize="11" level="3" pageId="13" pageNumber="1459" reason="5">
<emphasis id="B9033858FFEEFFC2FF5AFEEAFF6FAC7E" bold="true" box="[160,252,335,360]" inLineHeading="true" italics="true" pageId="13" pageNumber="1459" reason="3">Records</emphasis>
</heading>
</paragraph>
<paragraph id="8BC8E44AFFEEFFC2FF5AFED7FC66AC9C" blockId="13.[160,1013,335,394]" box="[160,1013,370,394]" pageId="13" pageNumber="1459">
<collectingRegion id="49B32AA8FFEEFFC2FF5AFED7FE2DAC9C" box="[160,446,370,394]" country="Iran" name="Khorasan-e Jonubi" pageId="13" pageNumber="1459">South Khorasan Province</collectingRegion>
. No new specimens were examined in this study.
</paragraph>
</subSubSection>
<subSubSection id="C36DB7C1FFEEFFC1FF5AFE64FF6BACC8" lastPageId="14" lastPageNumber="1460" pageId="13" pageNumber="1459" type="distribution">
<paragraph id="8BC8E44AFFEEFFC2FF5AFE64FEBFACCC" blockId="13.[160,675,449,509]" box="[160,300,449,474]" pageId="13" pageNumber="1459">
<heading id="D0805326FFEEFFC2FF5AFE64FEBFACCC" box="[160,300,449,474]" fontSize="11" level="3" pageId="13" pageNumber="1459" reason="5">
<emphasis id="B9033858FFEEFFC2FF5AFE64FEBFACCC" bold="true" box="[160,300,449,474]" inLineHeading="true" italics="true" pageId="13" pageNumber="1459" reason="3">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC8E44AFFEEFFC2FF5AFE40FD30ACEB" blockId="13.[160,675,449,509]" box="[160,675,485,509]" pageId="13" pageNumber="1459">
<collectingCountry id="F360A4DAFFEEFFC2FF5AFE40FEAFACEB" box="[160,316,485,509]" name="Turkmenistan" pageId="13" pageNumber="1459">Turkmenistan</collectingCountry>
,
<collectingCountry id="F360A4DAFFEEFFC2FEBDFE40FE56ACEB" box="[327,453,485,509]" name="Uzbekistan" pageId="13" pageNumber="1459">Uzbekistan</collectingCountry>
, north-eastern
<collectingCountry id="F360A4DAFFEEFFC2FD88FE40FD0CACEB" box="[626,671,485,509]" name="Iran" pageId="13" pageNumber="1459">Iran</collectingCountry>
.
</paragraph>
<paragraph id="8BC8E44AFFEEFFC2FF5AFDEFFEADAF72" blockId="13.[160,318,586,612]" box="[160,318,586,612]" pageId="13" pageNumber="1459">
<emphasis id="B9033858FFEEFFC2FF5AFDEFFEADAF72" bold="true" box="[160,318,586,612]" pageId="13" pageNumber="1459">Conclusions</emphasis>
</paragraph>
<paragraph id="8BC8E44AFFEEFFC2FF5AFD25FC8AAE34" blockId="13.[160,1155,640,802]" pageId="13" pageNumber="1459">
According to the literature, the fauna of
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in
<collectingCountry id="F360A4DAFFEEFFC2FCF0FD25FCA6AF8E" box="[778,821,640,664]" name="Iran" pageId="13" pageNumber="1459">Iran</collectingCountry>
should be considered poorly studied (
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). The first records of these spiders in the Iranian Plateau have been recently provided by
<bibRefCitation id="EFE699BBFFEEFFC2FDDEFD60FCA1AFCB" author="Marusik YM &amp; Fet V" box="[548,818,709,733]" pageId="13" pageNumber="1459" pagination="75 - 114" refId="ref4820" refString="Marusik YM, Fet V. 2009. A survey of East Palearctic Hersiliola Thorell, 1870 (Araneae: Hersiliidae), with description of three new genera. Zookeys. 16: 75 - 114." type="journal article" year="2009">Marusik and Fet (2009)</bibRefCitation>
and
<bibRefCitation id="EFE699BBFFEEFFC2FC8EFD60FBECAFCB" author="Mirshamsi O &amp; Marusik YM &amp; Jamili E" box="[884,1151,709,733]" pageId="13" pageNumber="1459" pagination="347 - 352" refId="ref4893" refString="Mirshamsi O, Marusik YM, Jamili E. 2013. The spider family Hersiliidae Thorell, 1870 (Arachnida: Araneae) in Iran. Zool Middle East. 59: 347 - 352." type="journal article" year="2013">Mirshamsi et al. (2013)</bibRefCitation>
.
<bibRefCitation id="EFE699BBFFEEFFC2FF5AFD4DFE37AE16" author="Mirshamsi O &amp; Marusik YM &amp; Jamili E" box="[160,420,744,768]" pageId="13" pageNumber="1459" pagination="347 - 352" refId="ref4893" refString="Mirshamsi O, Marusik YM, Jamili E. 2013. The spider family Hersiliidae Thorell, 1870 (Arachnida: Araneae) in Iran. Zool Middle East. 59: 347 - 352." type="journal article" year="2013">Mirshamsi et al. (2013)</bibRefCitation>
listed five species from the country, which showed the highest species diversity of
<taxonomicName id="4C779FC9FFEEFFC2FE7AFCAFFE67AE34" authorityName="Thorell" authorityYear="1870" box="[384,500,778,802]" class="Arachnida" family="Hersiliidae" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="family">Hersiliidae</taxonomicName>
in the Palaearctic region.
</paragraph>
<paragraph id="8BC8E44AFFEEFFC2FF5AF9CEFE2FABAA" blockId="13.[160,1155,1642,1724]" pageId="13" pageNumber="1459">
<emphasis id="B9033858FFEEFFC2FF5AF9CEFF6CAB94" bold="true" box="[160,255,1643,1666]" pageId="13" pageNumber="1459">Figure 6.</emphasis>
Map of the distribution of
<taxonomicName id="4C779FC9FFEEFFC2FDDDF9CFFC97AB97" authority="Mirshamsi &amp; Zamani &amp; Marusik, 2016" authorityName="Mirshamsi &amp; Zamani &amp; Marusik" authorityYear="2016" box="[551,772,1642,1665]" class="Arachnida" family="Hersiliidae" genus="Bastanius" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="species" species="kermanensis" status="sp. nov.">
<emphasis id="B9033858FFEEFFC2FDDDF9CFFC97AB97" box="[551,772,1642,1665]" italics="true" pageId="13" pageNumber="1459">Bastanius kermanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2308523FFEEFFC2FCEAF9CEFCF1AB94" box="[784,866,1643,1666]" pageId="13" pageNumber="1459" rank="species">sp. nov.</taxonomicNameLabel>
(∇),
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<emphasis id="B9033858FFEEFFC2FC61F9CFFC63AB97" box="[923,1008,1642,1665]" italics="true" pageId="13" pageNumber="1459">B. foordi</emphasis>
</taxonomicName>
(♦),
<taxonomicName id="4C779FC9FFEEFFC2FBD6F9CFFE94AB88" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="species" species="sternbergsi">
<emphasis id="B9033858FFEEFFC2FBD6F9CFFE94AB88" italics="true" pageId="13" pageNumber="1459">Hersiliola sternbergsi</emphasis>
</taxonomicName>
(●),
<taxonomicName id="4C779FC9FFEEFFC2FEC1F922FE5BAB88" baseAuthorityName="Dufour" baseAuthorityYear="1831" box="[315,456,1671,1694]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="species" species="macullulata">
<emphasis id="B9033858FFEEFFC2FEC1F922FE5BAB88" box="[315,456,1671,1694]" italics="true" pageId="13" pageNumber="1459">H. macullulata</emphasis>
</taxonomicName>
(○),
<taxonomicName id="4C779FC9FFEEFFC2FDF9F922FD1EAB88" authority="Mirshamsi &amp; Zamani &amp; Marusik, 2016" authorityName="Mirshamsi &amp; Zamani &amp; Marusik" authorityYear="2016" box="[515,653,1671,1694]" class="Insecta" family="Tachinidae" genus="Hersilia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="species" species="talebii" status="sp. nov.">
<emphasis id="B9033858FFEEFFC2FDF9F922FD1EAB88" box="[515,653,1671,1694]" italics="true" pageId="13" pageNumber="1459">Hersilia talebii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2308523FFEEFFC2FD6FF92DFD77AB89" box="[661,740,1672,1695]" pageId="13" pageNumber="1459" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B9033858FFEEFFC2FD08F92DFD6CAB89" bold="true" box="[754,767,1672,1695]" italics="true" pageId="13" pageNumber="1459"></emphasis>
),
<taxonomicName id="4C779FC9FFEEFFC2FCEEF922FC26AB88" box="[788,949,1671,1694]" class="Arachnida" family="Hersiliidae" genus="Duninia" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="species" species="baehrae">
<emphasis id="B9033858FFEEFFC2FCEEF922FC26AB88" box="[788,949,1671,1694]" italics="true" pageId="13" pageNumber="1459">Duninia baehrae</emphasis>
</taxonomicName>
(□),
<taxonomicName id="4C779FC9FFEEFFC2FC0AF922FBC4AB88" authorityName="Mirshamsi &amp; Marusik" authorityYear="2013" box="[1008,1111,1671,1694]" class="Arachnida" family="Hersiliidae" genus="Duninia" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="species" species="darvishi">
<emphasis id="B9033858FFEEFFC2FC0AF922FBC4AB88" box="[1008,1111,1671,1694]" italics="true" pageId="13" pageNumber="1459">D. darvishi</emphasis>
</taxonomicName>
(▲) and
<taxonomicName id="4C779FC9FFEEFFC2FF34F900FED3ABAA" authorityName="Marusik &amp; Fet" authorityYear="2009" box="[206,320,1701,1724]" class="Arachnida" family="Hersiliidae" genus="Duninia" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="1459" phylum="Arthropoda" rank="species" species="rheimsae">
<emphasis id="B9033858FFEEFFC2FF34F900FED3ABAA" box="[206,320,1701,1724]" italics="true" pageId="13" pageNumber="1459">D. rheimsae</emphasis>
</taxonomicName>
(▼) in
<collectingCountry id="F360A4DAFFEEFFC2FE6AF900FE2BABAA" box="[400,440,1701,1724]" name="Iran" pageId="13" pageNumber="1459">Iran</collectingCountry>
.
</paragraph>
<paragraph id="8BC8E44AFFEDFFC1FF3AFF2BFBC7ADFA" blockId="14.[160,1156,142,478]" pageId="14" pageNumber="1460">
The results of this study increase the number of hersiliids of
<collectingCountry id="F360A4DAFFEDFFC1FC86FF2BFC34ADB0" box="[892,935,142,166]" name="Iran" pageId="14" pageNumber="1460">Iran</collectingCountry>
to eight species in four genera:
<taxonomicName id="4C779FC9FFEDFFC1FEC2FF14FDA0ADDF" authority="Mirshamsi &amp; Zamani &amp; Marusik, 2016" authorityName="Mirshamsi &amp; Zamani &amp; Marusik" authorityYear="2016" box="[312,563,177,201]" class="Arachnida" family="Hersiliidae" genus="Bastanius" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1460" phylum="Arthropoda" rank="species" species="kermanensis" status="sp. nov.">
<emphasis id="B9033858FFEDFFC1FEC2FF14FDA0ADDF" box="[312,563,177,201]" italics="true" pageId="14" pageNumber="1460">Bastanius kermanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2308523FFEDFFC1FDC4FF14FD08ADDF" box="[574,667,177,201]" pageId="14" pageNumber="1460" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="B9033858FFEDFFC1FD53FF14FE3FADFA" italics="true" pageId="14" pageNumber="1460">
<taxonomicName id="4C779FC9FFEDFFC1FD53FF14FC9AADDF" authorityName="Marusik &amp; Fet" authorityYear="2009" box="[681,777,177,201]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1460" phylum="Arthropoda" rank="species" species="foordi">B. foordi</taxonomicName>
,
<taxonomicName id="4C779FC9FFEDFFC1FCE3FF14FC47ADDF" box="[793,980,177,201]" class="Arachnida" family="Hersiliidae" genus="Duninia" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1460" phylum="Arthropoda" rank="species" species="baehrae">Duninia baehrae</taxonomicName>
,
<taxonomicName id="4C779FC9FFEDFFC1FC1EFF14FBCEADDF" authorityName="Mirshamsi &amp; Marusik" authorityYear="2013" box="[996,1117,177,201]" class="Arachnida" family="Hersiliidae" genus="Duninia" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1460" phylum="Arthropoda" rank="species" species="darvishi">D. darvishi</taxonomicName>
,
<taxonomicName id="4C779FC9FFEDFFC1FB96FF14FE97ADFA" authorityName="Marusik &amp; Fet" authorityYear="2009" class="Arachnida" family="Hersiliidae" genus="Duninia" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1460" phylum="Arthropoda" rank="species" species="rheimsae">D. rheimsae</taxonomicName>
,
<taxonomicName id="4C779FC9FFEDFFC1FEEBFF71FE3FADFA" authority="Mirshamsi &amp; Zamani &amp; Marusik, 2016" authorityName="Mirshamsi &amp; Zamani &amp; Marusik" authorityYear="2016" box="[273,428,212,236]" class="Insecta" family="Tachinidae" genus="Hersilia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1460" phylum="Arthropoda" rank="species" species="talebii" status="sp. nov.">Hersilia talebii</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A2308523FFEDFFC1FE4FFF71FD9CADFA" box="[437,527,212,236]" pageId="14" pageNumber="1460" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C779FC9FFEDFFC1FDE1FF71FD6EADFA" box="[539,765,212,236]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1460" phylum="Arthropoda" rank="species" species="sternbergsi">
<emphasis id="B9033858FFEDFFC1FDE1FF71FD6EADFA" box="[539,765,212,236]" italics="true" pageId="14" pageNumber="1460">Hersiliola sternbergsi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C779FC9FFEDFFC1FCC3FF71FC4AADFA" baseAuthorityName="Dufour" baseAuthorityYear="1831" box="[825,985,212,236]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1460" phylum="Arthropoda" rank="species" species="macullulata">
<emphasis id="B9033858FFEDFFC1FCC3FF71FC4AADFA" box="[825,985,212,236]" italics="true" pageId="14" pageNumber="1460">H. macullulata</emphasis>
</taxonomicName>
(Figure 6).
</paragraph>
<paragraph id="8BC8E44AFFEDFFC1FF3AFF53FF6BACC8" blockId="14.[160,1156,142,478]" pageId="14" pageNumber="1460">
Although all of the above species are morphologically distinct, the application of molecular markers, such as cytochrome c oxidase, subunit I (
<emphasis id="B9033858FFEDFFC1FC85FEBCFC35AC27" box="[895,934,281,305]" italics="true" pageId="14" pageNumber="1460">COI</emphasis>
), may lead to the discovery of more new species of this taxon in this geographic area. Furthermore, considering the fact that comprehensive faunistic studies on spiders in much of the Iranian Plateau and Zagros Mountains have not been conducted, any newly collected material from these regions is therefore valuable and may represent new records or species.
</paragraph>
</subSubSection>
</treatment>
</document>