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<mods:title id="B10B9C14F4F948F096076C9C00F7D9FC">The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea)</mods:title>
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<paragraph id="8BCBCF3FFFAE207CFE03F8FD1C1CF883" blockId="4.[488,1100,1827,1888]" box="[488,1100,1827,1854]" pageId="4" pageNumber="36">
<heading id="D0837853FFAE207CFE03F8FD1C1CF883" box="[488,1100,1827,1854]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="36" reason="2">
<taxonomicName id="4C74B4BCFFAE207CFE03F8FD1C1CF883" ID-CoL="3GVRN" authority="(Sandberger &amp; Sandberger, 1851)" baseAuthorityName="Sandberger &amp; Sandberger" baseAuthorityYear="1851" box="[488,1100,1827,1854]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="4" pageNumber="36" phylum="Mollusca" rank="species" species="typus">
<emphasis id="B900132DFFAE207CFE03F8FD1AE9F880" bold="true" box="[488,697,1827,1853]" italics="true" pageId="4" pageNumber="36">Tornoceras typus</emphasis>
(Sandberger &amp; Sandberger, 1851)
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<paragraph id="8BCBCF3FFFAE207CFD2EF8981B3FF8DD" blockId="4.[488,1100,1827,1888]" box="[709,879,1862,1888]" pageId="4" pageNumber="36">
<figureCitation id="134FD3BAFFAE207CFD2EF8981B5CF8DD" box="[709,780,1862,1888]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[259,1329,1528,1886]" captionTargetId="figure-352@5.[249,1339,1527,1889]" captionTargetPageId="5" captionText="Fig. 4. Tornoceras typus (Sandberger &amp; Sandberger, 1851), reproduction of the lectotype illustrations by Sandberger &amp; Sandberger (18501856, pl. 10 fig. 14) and the photography by M.R. House in Becker (1993, pl. 3 figs 12). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6373971" httpUri="https://zenodo.org/record/6373971/files/figure.png" pageId="4" pageNumber="36">Fig. 4</figureCitation>
;
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<treatmentCitation id="0AD5E92EFFAE207CFF56F8561CD9F81F" author="Sandberger &amp; Sandberger" box="[189,1161,1927,1954]" page="109" pageId="4" pageNumber="36" year="1851">
<taxonomicName id="4C74B4BCFFAE207CFF56F8561BB1F81F" authority="Sandberger &amp; Sandberger, 1851: 109" authorityName="Sandberger &amp; Sandberger" authorityPageNumber="109" authorityYear="1851" box="[189,993,1927,1954]" class="Cephalopoda" family="Goniatitidae" genus="Goniatites" kingdom="Animalia" order="Ammonoidea" pageId="4" pageNumber="36" phylum="Mollusca" rank="variety" species="retrorsus" variety="typus">
<emphasis id="B900132DFFAE207CFF56F85619FBF81C" box="[189,427,1928,1954]" italics="true" pageId="4" pageNumber="36">Goniatites retrorsus</emphasis>
var.
<emphasis id="B900132DFFAE207CFE0EF8561A74F81C" box="[485,548,1928,1953]" italics="true" pageId="4" pageNumber="36">
<typeStatus id="54CF719DFFAE207CFE0EF8561A74F81C" box="[485,548,1928,1953]" pageId="4" pageNumber="36">typus</typeStatus>
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Sandberger &amp; Sandberger, 1851: 109
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, pl. 10 fig. 14.
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<taxonomicName id="4C74B4BCFFAE207CFF56F8141A7BF85E" ID-CoL="57G52" authority="Mut." authorityName="Mut." baseAuthorityName="Sandberger &amp; Sandberger" baseAuthorityYear="1851" box="[189,555,1993,2019]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="4" pageNumber="36" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B900132DFFAE207CFF56F81419F4F85E" box="[189,420,1993,2019]" italics="true" pageId="4" pageNumber="36">Tornoceras simplex</emphasis>
Mut.
<emphasis id="B900132DFFAE207CFE07F8141A7BF85E" box="[492,555,1994,2019]" italics="true" pageId="4" pageNumber="36">
<typeStatus id="54CF719DFFAE207CFE07F8141A7BF85E" box="[492,555,1994,2019]" pageId="4" pageNumber="36">typus</typeStatus>
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<treatmentCitation id="0AD5E92EFFAE207CFDA3F8171B7DF85E" author="Holzapfel E." box="[584,813,1993,2020]" page="98" pageId="4" pageNumber="36" year="1895">
<bibRefCitation id="EFE5B2CEFFAE207CFDA3F8171B7DF85E" author="Holzapfel E." box="[584,813,1993,2020]" pageId="4" pageNumber="36" pagination="1 - 460" refId="ref8581" refString="Holzapfel E. 1895. Das Obere Miteldevon (Schichten mit Stringocephalus Burtini und Maeneceras terebratum) im Rheinischen Gebirge. Abhandlungen der Koniglich Preussischen geologischen Landesanstalt, Neue Folge 16: 1 - 460." type="journal article" year="1895">Holzapfel 1895: 98</bibRefCitation>
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, text-fig. 6, pl. 6 fig. 11, pl. 7 fig. 9.
<taxonomicName id="4C74B4BCFFAF207DFF56FED419D9FE9E" ID-CoL="3GVRN" baseAuthorityName="Sandberger &amp; Sandberger" baseAuthorityYear="1851" box="[189,393,266,291]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="5" pageNumber="37" phylum="Mollusca" rank="species" species="typus">
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Tornoceras
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<bibRefCitation id="EFE5B2CEFFAF207DFE4DFED41A22FE9E" author="Korn D." box="[422,626,265,292]" pageId="5" pageNumber="37" pagination="147 - 167" refId="ref8927" refString="Korn D. 2021 b. Revision of Tornoceras typus (Sandberger &amp; Sandberger, 1851) - an iconic Devonian ammonoid of a clade with slow morphological evolution. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 302: 147 - 167. https: // doi. org / 10.1127 / njgpa / 2021 / 1026" type="journal article" year="2021">Korn 2021b: 149</bibRefCitation>
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, text-figs 25, 6a.
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<treatmentCitationGroup id="AB64E811FFAF207DFF56FEF31D4BFEFB" box="[189,1307,300,327]" pageId="5" pageNumber="37">
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<emphasis id="B900132DFFAF207DFF56FEF319C2FEFB" box="[189,402,301,326]" italics="true" pageId="5" pageNumber="37">Tornoceras typum</emphasis>
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<treatmentCitation id="0AD5E92EFFAF207DFE5BFEF21AD4FEFB" author="Becker R. T." box="[432,644,300,326]" page="182" pageId="5" pageNumber="37" year="1993">
<bibRefCitation id="EFE5B2CEFFAF207DFE5BFEF21AD4FEFB" author="Becker R. T." box="[432,644,300,326]" pageId="5" pageNumber="37" pagination="1 - 405" refId="ref7804" refString="Becker R. T. 1993. Stratigraphische Gliederung und Ammonoideen-Faunen im Nehdenium (Oberdevon II) von Europa und Nord-Afrika. Courier Forschungsinstitut Senckenberg 155: 1 - 405." type="journal article" year="1993">Becker 1993: 182</bibRefCitation>
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, pl. 3 figs 12. —
<treatmentCitation id="0AD5E92EFFAF207DFCB7FEF31C29FEFA" author="Korn D. &amp; Klug C." box="[860,1145,300,327]" page="156" pageId="5" pageNumber="37" year="2002">
<bibRefCitation id="EFE5B2CEFFAF207DFCB7FEF31C29FEFA" author="Korn D. &amp; Klug C." box="[860,1145,300,327]" pageId="5" pageNumber="37" refId="ref9031" refString="Korn D. &amp; Klug C. 2002. Ammoneae Devonicae. Backhuys Publishers, Leiden." type="book" year="2002">Korn &amp; Klug 2002: 156</bibRefCitation>
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, text-fig. 139.
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</paragraph>
<paragraph id="8BCBCF3FFFAF207DFF56FE8E1CB9FED7" blockId="5.[189,1307,265,432]" box="[189,1257,335,362]" pageId="5" pageNumber="37">
<treatmentCitationGroup id="AB64E811FFAF207DFF56FE8E1CB9FED7" box="[189,1257,335,362]" pageId="5" pageNumber="37">
?
<taxonomicName id="4C74B4BCFFAF207DFF3AFE8E1A68FED4" authorityName="Sandberger &amp; Sandberger" authorityYear="1851" box="[209,568,336,362]" class="Cephalopoda" family="Goniatitidae" genus="Goniatites" kingdom="Animalia" order="Ammonoidea" pageId="5" pageNumber="37" phylum="Mollusca" rank="variety" species="retrorsus" variety="typus">
<emphasis id="B900132DFFAF207DFF3AFE8E1990FED4" box="[209,448,336,362]" italics="true" pageId="5" pageNumber="37">Goniatites retrorsus</emphasis>
var.
<emphasis id="B900132DFFAF207DFE12FE8E1A68FED4" box="[505,568,336,361]" italics="true" pageId="5" pageNumber="37">
<typeStatus id="54CF719DFFAF207DFE12FE8E1A68FED4" box="[505,568,336,361]" pageId="5" pageNumber="37">typus</typeStatus>
</emphasis>
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<treatmentCitation id="0AD5E92EFFAF207DFDBEFE911CB9FED7" author="Sandberger &amp; Sandberger" box="[597,1257,335,362]" pageId="5" pageNumber="37" year="1851">Sandberger &amp; Sandberger 1851: pl. 10a figs 36, 1011.</treatmentCitation>
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</paragraph>
<paragraph id="8BCBCF3FFFAF207DFF56FEAD1C4BFE30" blockId="5.[189,1307,265,432]" box="[189,1051,370,397]" pageId="5" pageNumber="37">
<treatmentCitationGroup id="AB64E811FFAF207DFF56FEAD1C4BFE30" box="[189,1051,370,397]" pageId="5" pageNumber="37">
?
<taxonomicName id="4C74B4BCFFAF207DFF3AFEAD19E9FE31" ID-CoL="57G52" box="[209,441,370,396]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="5" pageNumber="37" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B900132DFFAF207DFF3AFEAD19E9FE31" box="[209,441,370,396]" italics="true" pageId="5" pageNumber="37">Tornoceras simplex</emphasis>
</taxonomicName>
<treatmentCitation id="0AD5E92EFFAF207DFE3CFEAC1AA5FE31" author="Foord A. H. &amp; Crick G. C." box="[471,757,370,397]" page="69" pageId="5" pageNumber="37" year="1897">
<bibRefCitation id="EFE5B2CEFFAF207DFE3CFEAC1AA5FE31" author="Foord A. H. &amp; Crick G. C." box="[471,757,370,397]" pageId="5" pageNumber="37" refId="ref8457" refString="Foord A. H. &amp; Crick G. C. 1897. Catalogue of the Fossil Cephalopoda in the British Museum (Natural History), Part III, Containing the Bactritidae and Part of the Suborder Ammonoidea. Printed by order of the Trustees, London." type="book" year="1897">Foord &amp; Crick 1897: 69</bibRefCitation>
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, 107, text-figs 25df, 48.
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</paragraph>
<paragraph id="8BCBCF3FFFAF207DFF56FE481BE9FE0D" blockId="5.[189,1307,265,432]" box="[189,953,406,432]" pageId="5" pageNumber="37">
<treatmentCitationGroup id="AB64E811FFAF207DFF56FE481BE9FE0D" box="[189,953,406,432]" pageId="5" pageNumber="37">
?
<taxonomicName id="4C74B4BCFFAF207DFF3AFE4919A6FE0D" ID-CoL="57G52" authority="mut." authorityName="Mut." box="[209,502,406,432]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="5" pageNumber="36" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B900132DFFAF207DFF3AFE4919E9FE0D" box="[209,441,406,432]" italics="true" pageId="5" pageNumber="37">Tornoceras simplex</emphasis>
mut.
</taxonomicName>
<emphasis id="B900132DFFAF207DFE16FE491A6FFE0D" box="[509,575,407,432]" italics="true" pageId="5" pageNumber="37">ovata</emphasis>
<treatmentCitation id="0AD5E92EFFAF207DFDB7FE481B4FFE0D" author="Frech F." box="[604,799,406,432]" page="47" pageId="5" pageNumber="37" year="1902">
<bibRefCitation id="EFE5B2CEFFAF207DFDB7FE481B4FFE0D" author="Frech F." box="[604,799,406,432]" pageId="5" pageNumber="37" pagination="172 - 176" refId="ref8557" refString="Frech F. 1902 b. Ueber Epitornoceras und Tornoceras. Centralblatt fur Mineralogie, Geologie und Palaeontologie 1902: 172 - 176." type="journal article" year="1902">Frech 1902b: 47</bibRefCitation>
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, pl. 3 fig. 21.
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<treatmentCitationGroup id="AB64E811FFAF207DFF56FE061B34FDA8" pageId="5" pageNumber="37">
non
<taxonomicName id="4C74B4BCFFAF207DFF18FE06199BFE4C" ID-CoL="57G56" baseAuthorityName="Sandberger &amp; Sandberger" baseAuthorityYear="1851" box="[243,459,472,497]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="5" pageNumber="37" phylum="Mollusca" rank="species" species="typum">
<emphasis id="B900132DFFAF207DFF18FE06199BFE4C" box="[243,459,472,497]" italics="true" pageId="5" pageNumber="37">Tornoceras typum</emphasis>
</taxonomicName>
<treatmentCitation id="0AD5E92EFFAF207DFE05FE091A96FE4C" author="Becker R. T." box="[494,710,471,497]" page="182" pageId="5" pageNumber="37" year="1993">
<bibRefCitation id="EFE5B2CEFFAF207DFE05FE091A96FE4C" author="Becker R. T." box="[494,710,471,497]" pageId="5" pageNumber="37" pagination="1 - 405" refId="ref7804" refString="Becker R. T. 1993. Stratigraphische Gliederung und Ammonoideen-Faunen im Nehdenium (Oberdevon II) von Europa und Nord-Afrika. Courier Forschungsinstitut Senckenberg 155: 1 - 405." type="journal article" year="1993">Becker 1993: 182</bibRefCitation>
</treatmentCitation>
, pl. 3 figs 34, text-fig. 68a. — Becker
<emphasis id="B900132DFFAF207DFB40FE061CB9FE4C" box="[1195,1257,471,497]" italics="true" pageId="5" pageNumber="37">et al.</emphasis>
2000: pl. 2 figs 78. —
<treatmentCitation id="0AD5E92EFFAF207DFE90FE251AC8FDA8" author="Korn D. &amp; Klug C." box="[379,664,506,533]" page="156" pageId="5" pageNumber="37" year="2002">
<bibRefCitation id="EFE5B2CEFFAF207DFE90FE251AC8FDA8" author="Korn D. &amp; Klug C." box="[379,664,506,533]" pageId="5" pageNumber="37" refId="ref9031" refString="Korn D. &amp; Klug C. 2002. Ammoneae Devonicae. Backhuys Publishers, Leiden." type="book" year="2002">Korn &amp; Klug 2002: 156</bibRefCitation>
</treatmentCitation>
, text-fig. 143ac.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCBCF3FFFAF207DFF56FDC0191AFDE6" blockId="5.[189,1399,471,603]" pageId="5" pageNumber="37">
<treatmentCitationGroup id="AB64E811FFAF207DFF56FDC0191AFDE6" pageId="5" pageNumber="37">
non
<taxonomicName id="4C74B4BCFFAF207DFF18FDC0198EFD85" ID-CoL="57G52" box="[243,478,542,568]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="5" pageNumber="37" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B900132DFFAF207DFF18FDC0198EFD85" box="[243,478,542,568]" italics="true" pageId="5" pageNumber="37">Tornoceras simplex</emphasis>
</taxonomicName>
Frech 1902: 47, pl. 5 fig. 8. —
<treatmentCitation id="0AD5E92EFFAF207DFC6EFDC01CD1FD85" author="Wedekind R." box="[901,1153,541,568]" page="135" pageId="5" pageNumber="37" year="1918">
<bibRefCitation id="EFE5B2CEFFAF207DFC6EFDC01CD1FD85" author="Wedekind R." box="[901,1153,541,568]" pageId="5" pageNumber="37" pagination="85 - 184" refId="ref9227" refString="Wedekind R. 1918. Die Genera der Palaeoammonoidea (Goniatiten). Mit Ausschluss der Mimoceratidae, Glyphioceratidae und Prolecanitidae. Palaeontographica 62: 85 - 184." type="journal article" year="1918">Wedekind 1918: 135</bibRefCitation>
</treatmentCitation>
, pl. 16 fig. 12, textfig. 40a.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36E9CB4FFAF207DFF56FD5C1B0CFC7C" pageId="5" pageNumber="37" type="diagnosis">
<paragraph id="8BCBCF3FFFAF207DFF56FD5C1967FD21" blockId="5.[189,311,642,668]" box="[189,311,642,668]" pageId="5" pageNumber="37">
<heading id="D0837853FFAF207DFF56FD5C1967FD21" bold="true" box="[189,311,642,668]" fontSize="11" level="3" pageId="5" pageNumber="37" reason="3">
<emphasis id="B900132DFFAF207DFF56FD5C1967FD21" bold="true" box="[189,311,642,668]" pageId="5" pageNumber="37">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCBCF3FFFAF207DFF56FD6C1B0CFC7C" blockId="5.[188,1399,689,961]" pageId="5" pageNumber="37">
<taxonomicName id="4C74B4BCFFAF207DFF56FD6C1913FD76" authorityName="Verae Frech" authorityYear="1902" box="[189,323,690,715]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="5" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis id="B900132DFFAF207DFF56FD6C1913FD76" box="[189,323,690,715]" italics="true" pageId="5" pageNumber="37">Tornoceras</emphasis>
</taxonomicName>
reaching about
<quantity id="4C8C62DAFFAF207DFDFBFD6F1A39FD76" box="[528,617,689,715]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="37" unit="mm" value="90.0">90 mm</quantity>
conch diameter. Thickly pachyconic, subevolute early juvenile stage (ww/dm ~ 0.80; uw/dm ~ 0.35 at
<quantity id="4C8C62DAFFAF207DFD7EFD0A1A8AFD52" box="[661,730,724,751]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="37" unit="mm" value="2.0">2 mm</quantity>
dm), thickly discoidal and involute subadult stage (ww/ dm ~ 0.58; uw/dm ~ 0.08 at
<quantity id="4C8C62DAFFAF207DFDC9FD291A28FCAF" box="[546,632,759,786]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="37" unit="mm" value="10.0">10 mm</quantity>
dm), thinly discoidal and involute adult stage (ww/dm ~ 0.42; umbilicus closed at
<quantity id="4C8C62DAFFAF207DFE42FCC519ADFC88" box="[425,509,795,821]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="37" unit="mm" value="30.0">30 mm</quantity>
dm). Whorl profile as wide as high in the subadult stage (ww/wh ~ 1.00 at
<quantity id="4C8C62DAFFAF207DFF56FCE01940FCE5" box="[189,272,830,856]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="37" unit="mm" value="10.0">10 mm</quantity>
dm) and strongly compressed in the adult stage (ww/wh ~ 0.75 at
<quantity id="4C8C62DAFFAF207DFBF7FCE01C3EFCE5" box="[1052,1134,830,856]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="37" unit="mm" value="30.0">30 mm</quantity>
dm); whorl expansion rate very high in the subadult and adult stage (WER = 2.30-2.45). Flanks convex, convergent, umbilical margin continuously rounded. Without any ventrolateral grooves. Suture line with broadly rounded, asymmetric adventive lobe and high ventrolateral saddle.
</paragraph>
</subSubSection>
<subSubSection id="C36E9CB4FFAF207DFF56FC371AE7FAC9" pageId="5" pageNumber="37" type="materials_examined">
<paragraph id="8BCBCF3FFFAF207DFF56FC37193DFBBF" blockId="5.[189,365,1000,1027]" box="[189,365,1000,1027]" pageId="5" pageNumber="37">
<heading id="D0837853FFAF207DFF56FC37193DFBBF" bold="true" box="[189,365,1000,1027]" fontSize="11" level="3" pageId="5" pageNumber="37" reason="3">
<emphasis id="B900132DFFAF207DFF56FC37193DFBBF" bold="true" box="[189,365,1000,1027]" pageId="5" pageNumber="37">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BCBCF3FFFAF207DFF01FBC91D27FB26" blockId="5.[186,1399,1047,1180]" pageId="5" pageNumber="37">
<materialsCitation id="3B1CC562FFAF207DFF01FBC91D23FB26" ID-GBIF-Occurrence="3714208321" country="Germany" location="Oberscheld" municipality="Rhenish Mountains" pageId="5" pageNumber="37" specimenCount="1" typeStatus="lectotype">
<emphasis id="B900132DFFAF207DFF01FBC91937FB8C" bold="true" box="[234,359,1047,1073]" pageId="5" pageNumber="37">
<typeStatus id="54CF719DFFAF207DFF01FBC91937FB8C" box="[234,359,1047,1073]" pageId="5" pageNumber="37" type="lectotype">Lectotype</typeStatus>
</emphasis>
<collectingCountry id="F3638FAFFFAF207DFF56FBE41900FBE9" box="[189,336,1082,1108]" name="Germany" pageId="5" pageNumber="37">GERMANY</collectingCountry>
<collectingMunicipality id="6BAF5545FFAF207DFE8FFBE41A1AFBE9" box="[356,586,1082,1108]" pageId="5" pageNumber="37">Rhenish Mountains</collectingMunicipality>
,
<location id="8EAB99E4FFAF207DFDB9FBE41A8BFBE9" LSID="urn:lsid:plazi:treatment:03DD7E29FFAE207EFE03F8FD1912FBE9:8EAB99E4FFAF207DFDB9FBE41A8BFBE9" box="[594,731,1082,1108]" country="Germany" municipality="Rhenish Mountains" name="Oberscheld" pageId="5" pageNumber="37">Oberscheld</location>
;
<geologicalTimeScale id="F9544F94FFAF207DFD0EFBE41CA0FBE8" box="[741,1264,1082,1109]" pageId="5" pageNumber="37">probably middle Frasnian (Adorf Limestone)</geologicalTimeScale>
; designated by
<bibRefCitation id="EFE5B2CEFFAF207DFF34FB8319DAFBC5" author="Becker R. T." box="[223,394,1117,1144]" pageId="5" pageNumber="37" pagination="1 - 405" refId="ref7804" refString="Becker R. T. 1993. Stratigraphische Gliederung und Ammonoideen-Faunen im Nehdenium (Oberdevon II) von Europa und Nord-Afrika. Courier Forschungsinstitut Senckenberg 155: 1 - 405." type="journal article" year="1993">Becker (1993)</bibRefCitation>
; figured by
<bibRefCitation id="EFE5B2CEFFAF207DFDFFFB831BC2FBC5" author="Sandberger G. &amp; Sandberger F." box="[532,914,1117,1144]" pageId="5" pageNumber="37" refId="ref9187" refString="Sandberger G. &amp; Sandberger F. 1850 - 1856. Die Versteinerungen des rheinischen Schichtensystems in Nassau. Mit einer kurzgefassten Geognosie dieses Gebietes und mit steter Berucksichtigung analoger Schichten anderer Lander. Kreidel &amp; Niedner, Wiesbaden." type="book" year="1850">Sandberger &amp; Sandberger (1850</bibRefCitation>
1856: pl. 10 fig. 14),
<bibRefCitation id="EFE5B2CEFFAF207DFB7EFB831D68FBC5" author="Becker R. T." box="[1173,1336,1117,1144]" pageId="5" pageNumber="37" pagination="1 - 405" refId="ref7804" refString="Becker R. T. 1993. Stratigraphische Gliederung und Ammonoideen-Faunen im Nehdenium (Oberdevon II) von Europa und Nord-Afrika. Courier Forschungsinstitut Senckenberg 155: 1 - 405." type="journal article" year="1993">Becker (1993</bibRefCitation>
: pl. 3 figs 13) and
<bibRefCitation id="EFE5B2CEFFAF207DFEB7FB5F19A7FB26" author="Korn D." box="[348,503,1153,1179]" pageId="5" pageNumber="37" pagination="147 - 167" refId="ref8927" refString="Korn D. 2021 b. Revision of Tornoceras typus (Sandberger &amp; Sandberger, 1851) - an iconic Devonian ammonoid of a clade with slow morphological evolution. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 302: 147 - 167. https: // doi. org / 10.1127 / njgpa / 2021 / 1026" type="journal article" year="2021">Korn (2021b</bibRefCitation>
: text-fig. 2), re-illustrated here in
<figureCitation id="134FD3BAFFAF207DFC94FB5F1B96FB26" box="[895,966,1153,1179]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[259,1329,1528,1886]" captionTargetId="figure-352@5.[249,1339,1527,1889]" captionTargetPageId="5" captionText="Fig. 4. Tornoceras typus (Sandberger &amp; Sandberger, 1851), reproduction of the lectotype illustrations by Sandberger &amp; Sandberger (18501856, pl. 10 fig. 14) and the photography by M.R. House in Becker (1993, pl. 3 figs 12). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6373971" httpUri="https://zenodo.org/record/6373971/files/figure.png" pageId="5" pageNumber="37">Fig. 4</figureCitation>
; specimen 53a; Wiesbaden Museum
</materialsCitation>
.
</paragraph>
<paragraph id="8BCBCF3FFFAF207DFF56FB1C1A68FB61" blockId="5.[189,568,1218,1244]" box="[189,568,1218,1244]" pageId="5" pageNumber="37">
<heading id="D0837853FFAF207DFF56FB1C1A68FB61" bold="true" box="[189,568,1218,1244]" fontSize="11" level="3" pageId="5" pageNumber="37" reason="3">
<emphasis id="B900132DFFAF207DFF56FB1C1A68FB61" bold="true" box="[189,568,1218,1244]" pageId="5" pageNumber="37">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCBCF3FFFAF207DFF56FB2F1AE7FAC9" blockId="5.[188,1400,1265,1397]" pageId="5" pageNumber="37">
<materialsCitation id="3B1CC562FFAF207DFF56FB2F19C1FA93" ID-GBIF-Occurrence="3714208309" collectionCode="MB" collectorName="Erbreich" country="Germany" county="Frasnian" location="Rhenish Mountains" municipality="Oberscheld" pageId="5" pageNumber="37" specimenCode="MB.C.4466" specimenCount="1">
<collectingCountry id="F3638FAFFFAF207DFF56FB2F1900FAB6" box="[189,336,1265,1291]" name="Germany" pageId="5" pageNumber="37">GERMANY</collectingCountry>
<specimenCount id="9D7204B6FFAF207DFE87FB2F1992FAB6" box="[364,450,1265,1291]" pageId="5" pageNumber="37" type="generic">1 spec.</specimenCount>
;
<location id="8EAB99E4FFAF207DFE24FB2F1AE9FAB6" LSID="urn:lsid:plazi:treatment:03DD7E29FFAE207EFE03F8FD1912FBE9:8EAB99E4FFAF207DFE24FB2F1AE9FAB6" box="[463,697,1265,1291]" country="Germany" county="Frasnian" municipality="Oberscheld" name="Rhenish Mountains" pageId="5" pageNumber="37">Rhenish Mountains</location>
,
<collectingMunicipality id="6BAF5545FFAF207DFD2EFB2F1B1EFAB6" box="[709,846,1265,1291]" pageId="5" pageNumber="37">Oberscheld</collectingMunicipality>
;
<geologicalTimeScale id="F9544F94FFAF207DFCB0FB2F1D55FAB1" box="[859,1285,1265,1292]" pageId="5" pageNumber="37">middle Frasnian (Adorf Limestone)</geologicalTimeScale>
;
<collectorName id="2681AAE9FFAF207DFAFAFB2F1D26FAB6" box="[1297,1398,1265,1291]" pageId="5" pageNumber="37">Erbreich</collectorName>
Coll.;
<specimenCode id="DBD26744FFAF207DFEEDFACA19C1FA93" box="[262,401,1300,1326]" pageId="5" pageNumber="37">MB.C.4466</specimenCode>
</materialsCitation>
<materialsCitation id="3B1CC562FFAF207DFE4CFACA19CFFAEC" ID-GBIF-Occurrence="3714208308" collectionCode="MB" collectorName="Koch" country="Germany" county="Frasnian" location="Rhenish Mountains" municipality="Oberscheld" pageId="5" pageNumber="37" specimenCode="MB.C.30409" specimenCount="1">
<specimenCount id="9D7204B6FFAF207DFE4CFACA19ABFA93" box="[423,507,1300,1326]" pageId="5" pageNumber="37" type="generic">1 spec.</specimenCount>
;
<location id="8EAB99E4FFAF207DFDEDFACA1ABDFA93" LSID="urn:lsid:plazi:treatment:03DD7E29FFAE207EFE03F8FD1912FBE9:8EAB99E4FFAF207DFDEDFACA1ABDFA93" box="[518,749,1300,1326]" country="Germany" county="Frasnian" municipality="Oberscheld" name="Rhenish Mountains" pageId="5" pageNumber="37">Rhenish Mountains</location>
,
<collectingMunicipality id="6BAF5545FFAF207DFD1CFACA1BD0FA93" box="[759,896,1300,1326]" pageId="5" pageNumber="37">Oberscheld</collectingMunicipality>
;
<geologicalTimeScale id="F9544F94FFAF207DFC60FACA1D7EFA92" box="[907,1326,1300,1327]" pageId="5" pageNumber="37">middle Frasnian (Adorf Limestone)</geologicalTimeScale>
;
<collectorName id="2681AAE9FFAF207DFAD3FACA1D28FA93" box="[1336,1400,1300,1326]" pageId="5" pageNumber="37">Koch</collectorName>
Coll.;
<specimenCode id="DBD26744FFAF207DFEEDFAE919CFFAEC" box="[262,415,1335,1361]" pageId="5" pageNumber="37">MB.C.30409</specimenCode>
</materialsCitation>
<materialsCitation id="3B1CC562FFAF207DFE52FAE91AE3FAC9" ID-GBIF-Occurrence="3714208314" collectionCode="MB" country="Germany" county="Frasnian" location="Rhenish Mountains" municipality="Eiserne Hand" pageId="5" pageNumber="37" specimenCode="MB.C.30410" specimenCount="1">
<specimenCount id="9D7204B6FFAF207DFE52FAE91A5EFAEF" box="[441,526,1335,1362]" pageId="5" pageNumber="37" type="generic">1 spec.</specimenCount>
;
<location id="8EAB99E4FFAF207DFDF1FAE91B53FAEC" LSID="urn:lsid:plazi:treatment:03DD7E29FFAE207EFE03F8FD1912FBE9:8EAB99E4FFAF207DFDF1FAE91B53FAEC" box="[538,771,1335,1361]" country="Germany" county="Frasnian" municipality="Eiserne Hand" name="Rhenish Mountains" pageId="5" pageNumber="37">Rhenish Mountains</location>
,
<location id="8EAB99E4FFAF207DFCE4FAE91BC1FAEC" LSID="urn:lsid:plazi:treatment:03DD7E29FFAE207EFE03F8FD1912FBE9:8EAB99E4FFAF207DFCE4FAE91BC1FAEC" box="[783,913,1335,1361]" country="Germany" county="Frasnian" municipality="Eiserne Hand" name="Dillenburg" pageId="5" pageNumber="37">Dillenburg</location>
,
<collectingMunicipality id="6BAF5545FFAF207DFC76FAE91C6CFAEC" box="[925,1084,1335,1361]" pageId="5" pageNumber="37">Eiserne Hand</collectingMunicipality>
(Grube Friedrich);
<geologicalTimeScale id="F9544F94FFAF207DFACFFAE91A5AFAC8" pageId="5" pageNumber="37">middle Frasnian (Adorf Limestone)</geologicalTimeScale>
;
<specimenCode id="DBD26744FFAF207DFDFDFA841AE3FAC9" box="[534,691,1370,1396]" pageId="5" pageNumber="37">MB.C.30410</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0B9FB7FFAF207DFF56F8561A89F854" ID-DOI="http://doi.org/10.5281/zenodo.6373971" ID-Zenodo-Dep="6373971" httpUri="https://zenodo.org/record/6373971/files/figure.png" pageId="5" pageNumber="37" startId="5.[189,232,1928,1954]" targetBox="[259,1329,1528,1886]" targetPageId="5">
<paragraph id="8BCBCF3FFFAF207DFF56F8561A89F854" blockId="5.[189,1399,1928,2025]" pageId="5" pageNumber="37">
<emphasis id="B900132DFFAF207DFF56F856195DF81F" bold="true" box="[189,269,1928,1954]" pageId="5" pageNumber="37">Fig. 4.</emphasis>
<taxonomicName id="4C74B4BCFFAF207DFEFDF8571B2EF81E" authority="(Sandberger &amp; Sandberger, 1851)" baseAuthorityName="Sandberger &amp; Sandberger" baseAuthorityYear="1851" box="[278,894,1928,1955]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="5" pageNumber="37" phylum="Mollusca" rank="species" species="typus">
<emphasis id="B900132DFFAF207DFEFDF85719B3F81F" box="[278,483,1929,1954]" italics="true" pageId="5" pageNumber="37">Tornoceras typus</emphasis>
(Sandberger &amp; Sandberger, 1851)
</taxonomicName>
, reproduction of the lectotype illustrations by
<bibRefCitation id="EFE5B2CEFFAF207DFF09F8721A0BF87B" author="Sandberger G. &amp; Sandberger F." box="[226,603,1963,1990]" pageId="5" pageNumber="37" refId="ref9187" refString="Sandberger G. &amp; Sandberger F. 1850 - 1856. Die Versteinerungen des rheinischen Schichtensystems in Nassau. Mit einer kurzgefassten Geognosie dieses Gebietes und mit steter Berucksichtigung analoger Schichten anderer Lander. Kreidel &amp; Niedner, Wiesbaden." type="book" year="1850">Sandberger &amp; Sandberger (1850</bibRefCitation>
1856, pl. 10 fig. 14) and the photography by M.R. House in
<bibRefCitation id="EFE5B2CEFFAF207DFAC8F8721954F854" author="Becker R. T." pageId="5" pageNumber="37" pagination="1 - 405" refId="ref7804" refString="Becker R. T. 1993. Stratigraphische Gliederung und Ammonoideen-Faunen im Nehdenium (Oberdevon II) von Europa und Nord-Afrika. Courier Forschungsinstitut Senckenberg 155: 1 - 405." type="journal article" year="1993">Becker (1993</bibRefCitation>
, pl. 3 figs 12). Scale bar units = 1 mm.
</paragraph>
</caption>
<caption id="DF0B9FB7FFAC207EFF56FED719DAFEFA" ID-Table-UUID="DF0B9FB7FFAC207EFF56FED719DAFEFA" httpUri="http://table.plazi.org/id/DF0B9FB7FFAC207EFF56FED719DAFEFA" pageId="6" pageNumber="38" startId="6.[189,259,265,291]" targetBox="[215,1358,366,430]" targetIsTable="true" targetPageId="6">
<paragraph id="8BCBCF3FFFAC207EFF56FED719DAFEFA" blockId="6.[189,1399,265,327]" pageId="6" pageNumber="38">
<emphasis id="B900132DFFAC207EFF56FED71970FE9E" bold="true" box="[189,288,265,291]" pageId="6" pageNumber="38">Table 1.</emphasis>
Conch dimensions and ratios of
<taxonomicName id="4C74B4BCFFAC207EFD4BFED41CAFFE99" authority="(Sandberger &amp; Sandberger, 1851)" baseAuthorityName="Sandberger &amp; Sandberger" baseAuthorityYear="1851" box="[672,1279,265,292]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Ammonoidea" pageId="6" pageNumber="38" phylum="Mollusca" rank="species" species="typus">
<emphasis id="B900132DFFAC207EFD4BFED41B3CFE9E" box="[672,876,266,291]" italics="true" pageId="6" pageNumber="38">Tornoceras typus</emphasis>
(Sandberger &amp; Sandberger, 1851)
</taxonomicName>
, lectotype (Wiesbaden 53a).
</paragraph>
</caption>
<paragraph id="8BCBCF3FFFAC207EFF3CFEB01D1EFE13" pageId="6" pageNumber="38">
<table id="F9743D9FFFACDF87FF3CFEB01D1EFE13" box="[215,1358,366,430]" gridcols="11" gridrows="2" pageId="6" pageNumber="38">
<tr id="3544CD7DFFACDF87FF3CFEB01D1EFE3A" box="[215,1358,366,391]" gridrow="0" pageId="6" pageNumber="38">
<th id="7695A401FFACDF87FF3CFEB01929FE3A" box="[215,377,366,391]" gridcol="0" gridrow="0" pageId="6" pageNumber="38">Specimen</th>
<td id="7695A401FFACDF87FE46FEB0198CFE3A" box="[429,476,366,391]" gridcol="1" gridrow="0" pageId="6" pageNumber="38">dm</td>
<td id="7695A401FFACDF87FDEFFEB01A63FE3A" box="[516,563,366,391]" gridcol="2" gridrow="0" pageId="6" pageNumber="38">ww</td>
<td id="7695A401FFACDF87FD89FEB01AC1FE3A" box="[610,657,366,391]" gridcol="3" gridrow="0" pageId="6" pageNumber="38">wh</td>
<td id="7695A401FFACDF87FD23FEB01ABAFE3A" box="[712,746,366,391]" gridcol="4" gridrow="0" pageId="6" pageNumber="38">uw</td>
<td id="7695A401FFACDF87FCF4FEB01B1EFE3A" box="[799,846,366,391]" gridcol="5" gridrow="0" pageId="6" pageNumber="38">ah</td>
<td id="7695A401FFACDF87FC9BFEB01B93FE3A" box="[880,963,366,391]" gridcol="6" gridrow="0" pageId="6" pageNumber="38">ww/dm</td>
<td id="7695A401FFACDF87FC36FEB01C7DFE3A" box="[989,1069,366,391]" gridcol="7" gridrow="0" pageId="6" pageNumber="38">ww/wh</td>
<td id="7695A401FFACDF87FBA3FEB01CC7FE3A" box="[1096,1175,366,391]" gridcol="8" gridrow="0" pageId="6" pageNumber="38">uw/dm</td>
<td id="7695A401FFACDF87FB5DFEB01CA6FE3A" box="[1206,1270,366,391]" gridcol="9" gridrow="0" pageId="6" pageNumber="38">
<collectionCode id="ED6557FAFFAC207EFB5DFEB11CA6FE3A" box="[1206,1270,367,391]" pageId="6" pageNumber="38">WER</collectionCode>
</td>
<td id="7695A401FFACDF87FAF4FEB01D1EFE3A" box="[1311,1358,366,391]" gridcol="10" gridrow="0" pageId="6" pageNumber="38">
<emphasis id="B900132DFFAC207EFAF4FEB11D1EFE3A" bold="true" box="[1311,1358,367,391]" pageId="6" pageNumber="38">IZR</emphasis>
</td>
</tr>
<tr id="3544CD7DFFACDF87FF3CFE481D1EFE13" box="[215,1358,406,430]" gridrow="1" pageId="6" pageNumber="38">
<th id="7695A401FFACDF87FF3CFE481929FE13" box="[215,377,406,430]" gridcol="0" gridrow="1" pageId="6" pageNumber="38">Wiesbaden 53a</th>
<td id="7695A401FFACDF87FE46FE48198CFE13" box="[429,476,406,430]" gridcol="1" gridrow="1" pageId="6" pageNumber="38">46.0</td>
<td id="7695A401FFACDF87FDEFFE481A63FE13" box="[516,563,406,430]" gridcol="2" gridrow="1" pageId="6" pageNumber="38">18.3</td>
<td id="7695A401FFACDF87FD89FE481AC1FE13" box="[610,657,406,430]" gridcol="3" gridrow="1" pageId="6" pageNumber="38">29.7</td>
<td id="7695A401FFACDF87FD23FE481ABAFE13" box="[712,746,406,430]" gridcol="4" gridrow="1" pageId="6" pageNumber="38">0.0</td>
<td id="7695A401FFACDF87FCF4FE481B1EFE13" box="[799,846,406,430]" gridcol="5" gridrow="1" pageId="6" pageNumber="38">15.2</td>
<td id="7695A401FFACDF87FC9BFE481B93FE13" box="[880,963,406,430]" gridcol="6" gridrow="1" pageId="6" pageNumber="38">0.40</td>
<td id="7695A401FFACDF87FC36FE481C7DFE13" box="[989,1069,406,430]" gridcol="7" gridrow="1" pageId="6" pageNumber="38">0.62</td>
<td id="7695A401FFACDF87FBA3FE481CC7FE13" box="[1096,1175,406,430]" gridcol="8" gridrow="1" pageId="6" pageNumber="38">0.00</td>
<td id="7695A401FFACDF87FB5DFE481CA6FE13" box="[1206,1270,406,430]" gridcol="9" gridrow="1" pageId="6" pageNumber="38">2.23</td>
<td id="7695A401FFACDF87FAF4FE481D1EFE13" box="[1311,1358,406,430]" gridcol="10" gridrow="1" pageId="6" pageNumber="38">0.49</td>
</tr>
</table>
</paragraph>
<subSubSection id="C36E9CB4FFAC207EFF56FDC61C6DFCC7" pageId="6" pageNumber="38" type="description">
<paragraph id="8BCBCF3FFFAC207EFF56FDC6191FFD8F" blockId="6.[189,335,536,562]" box="[189,335,536,562]" pageId="6" pageNumber="38">
<heading id="D0837853FFAC207EFF56FDC6191FFD8F" bold="true" box="[189,335,536,562]" fontSize="11" level="3" pageId="6" pageNumber="38" reason="3">
<emphasis id="B900132DFFAC207EFF56FDC6191FFD8F" bold="true" box="[189,335,536,562]" pageId="6" pageNumber="38">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCBCF3FFFAC207EFF01FD991AF4FDDC" blockId="6.[188,1400,583,891]" box="[234,676,583,609]" pageId="6" pageNumber="38">
<emphasis id="B900132DFFAC207EFF01FD9919C9FDDC" bold="true" box="[234,409,583,609]" pageId="6" pageNumber="38">
<typeStatus id="54CF719DFFAC207EFF01FD991936FDDC" box="[234,358,583,609]" pageId="6" pageNumber="38" type="lectotype">Lectotype</typeStatus>
53a
</emphasis>
(Wiesbaden Museum)
</paragraph>
<paragraph id="8BCBCF3FFFAC207EFF56FDB41C6DFCC7" blockId="6.[188,1400,583,891]" pageId="6" pageNumber="38">
Fairly well-preserved specimen with
<quantity id="4C8C62DAFFAC207EFD93FDB41A9DFD38" box="[632,717,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="6" pageNumber="38" unit="mm" value="46.0">46 mm</quantity>
conch diameter in red limestone; it is largely chambered and the last half volution belongs to the body chamber (
<figureCitation id="134FD3BAFFAC207EFC84FD531BECFD1A" box="[879,956,653,679]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[259,1329,1528,1886]" captionTargetId="figure-352@5.[249,1339,1527,1889]" captionTargetPageId="5" captionText="Fig. 4. Tornoceras typus (Sandberger &amp; Sandberger, 1851), reproduction of the lectotype illustrations by Sandberger &amp; Sandberger (18501856, pl. 10 fig. 14) and the photography by M.R. House in Becker (1993, pl. 3 figs 12). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6373971" httpUri="https://zenodo.org/record/6373971/files/figure.png" pageId="6" pageNumber="38">Fig. 4</figureCitation>
). It is a thinly discoidal conch (ww/ dm = 0.40) with a closed umbilicus. The body chamber is obviously slightly distorted, resulting in a stronger convergence of the flanks (as shown in the original illustration by
<bibRefCitation id="EFE5B2CEFFAC207EFBAEFD0A18AAFCAF" author="Sandberger G. &amp; Sandberger F." pageId="6" pageNumber="38" refId="ref9187" refString="Sandberger G. &amp; Sandberger F. 1850 - 1856. Die Versteinerungen des rheinischen Schichtensystems in Nassau. Mit einer kurzgefassten Geognosie dieses Gebietes und mit steter Berucksichtigung analoger Schichten anderer Lander. Kreidel &amp; Niedner, Wiesbaden." type="book" year="1850">Sandberger &amp; Sandberger 1850</bibRefCitation>
1856). Parts of the specimen are covered by shell material, which, however, is not well preserved and does not allow a detailed examination of the ornamentation. The suture line has a small, V-shaped outer lobe and a broadly rounded ventrolateral saddle. The adventitious lobe is twice as deep as the outer lobe and as deep as it is wide; it is broadly rounded and slightly asymmetric.
</paragraph>
</subSubSection>
<subSubSection id="C36E9CB4FFAC207EFF56FC7C1912FBE9" pageId="6" pageNumber="38" type="discussion">
<paragraph id="8BCBCF3FFFAC207EFF56FC7C1960FC01" blockId="6.[189,304,930,956]" box="[189,304,930,956]" pageId="6" pageNumber="38">
<heading id="D0837853FFAC207EFF56FC7C1960FC01" bold="true" box="[189,304,930,956]" fontSize="11" level="3" pageId="6" pageNumber="38" reason="3">
<emphasis id="B900132DFFAC207EFF56FC7C1960FC01" bold="true" box="[189,304,930,956]" pageId="6" pageNumber="38">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCBCF3FFFAC207EFF56FC0E1912FBE9" blockId="6.[189,1400,976,1108]" pageId="6" pageNumber="38">
The species has been discussed several times in the literature. A detailed revision and discussion was carried out by
<bibRefCitation id="EFE5B2CEFFAC207EFE87FC2A1A41FBB2" author="Korn D." box="[364,529,1012,1039]" pageId="6" pageNumber="38" pagination="147 - 167" refId="ref8927" refString="Korn D. 2021 b. Revision of Tornoceras typus (Sandberger &amp; Sandberger, 1851) - an iconic Devonian ammonoid of a clade with slow morphological evolution. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 302: 147 - 167. https: // doi. org / 10.1127 / njgpa / 2021 / 1026" type="journal article" year="2021">Korn (2021b)</bibRefCitation>
; in this paper both the taxonomic problems and the conch morphology as well as the ontogenetic development are presented on the basis of further material from the Rhenish Mountains.
</paragraph>
</subSubSection>
</treatment>
</document>