261 lines
34 KiB
XML
261 lines
34 KiB
XML
<document id="4F83A4E453970DB902A44CD8F043E1E4" ID-CLB-Dataset="9022" ID-DOI="10.11646/zootaxa.4868.1.2" ID-GBIF-Dataset="a5c421db-9d17-4a25-9431-378a79f940bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417203" ID-ZooBank="B869EB5E-073D-471D-A9F3-D390D57B25E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609804682391" checkinUser="plazi" docAuthor="Sankaran, Pradeep M. & Sebastian, Pothalil A." docDate="2020" docId="03B27A7EFFD10068FF445899FDCFFB4B" docLanguage="en" docName="zootaxa.4868.1.2.pdf" docOrigin="Zootaxa 4868 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carlogonus Demange 1961" docType="treatment" docVersion="8" lastPageNumber="29" masterDocId="FF8B0206FFD0006AFFD35E5EFFBBFFA2" masterDocTitle="A new species of Carlogonus Demange, 1961 from southern India (Spirostreptida Harpagophoridae, Harpagophorinae), with phylogenetic analysis and speciesgroup divisions in the genus" masterLastPageNumber="40" masterPageNumber="27" pageNumber="28" updateTime="1698852323459" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CE78E3A3E9663C66D6D9CEEF6951AC62">A new species of Carlogonus Demange, 1961 from southern India (Spirostreptida Harpagophoridae, Harpagophorinae), with phylogenetic analysis and speciesgroup divisions in the genus</mods:title>
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<mods:name id="E82F900C2CE2E8BC2BD4258D26908121" type="personal">
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<mods:namePart id="8598E19F11880005C064C7427FDF1CC0">Sankaran, Pradeep M.</mods:namePart>
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<mods:affiliation id="2137C622EE1D23BFDEAFAAC95F104214">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India.</mods:affiliation>
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<mods:namePart id="A5C8BDCD38BEA41B2D95D2ED6B04CFD2">Sebastian, Pothalil A.</mods:namePart>
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<mods:nameIdentifier id="EC9E80E64CAB7CA94790C5674C3B61EC" type="ORCID">0000-0002-4936-4310</mods:nameIdentifier>
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<mods:affiliation id="DBC6CC89DDDDDE43EC92CD70BD93006A">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India. & drpothalil @ rediffmail. com; https: // orcid. org / 0000 - 0002 - 4936 - 4310</mods:affiliation>
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<mods:nameIdentifier id="8FE66B56D24D160121B95126018B2487" type="email">drpothalil@rediffmail.com</mods:nameIdentifier>
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<mods:title id="4BF97EE75F8A964811567D8C9EC4177F">Zootaxa</mods:title>
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<mods:date id="46371793A72D1C9302F4602100E49FB9">2020</mods:date>
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<mods:number id="18387E3DB7D3C9444D09CA67C277686B">2020-10-23</mods:number>
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<mods:number id="9E875A13645F168628F86FF298040163">4868</mods:number>
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<treatment id="03B27A7EFFD10068FF445899FDCFFB4B" ID-DOI="http://doi.org/10.5281/zenodo.4417447" ID-GBIF-Taxon="176440894" ID-Zenodo-Dep="4417447" LSID="urn:lsid:plazi:treatment:03B27A7EFFD10068FF445899FDCFFB4B" httpUri="http://treatment.plazi.org/id/03B27A7EFFD10068FF445899FDCFFB4B" lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28">
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<subSubSection id="C30198E3FFD1006BFF445899FDF8F943" box="[151,579,1735,1762]" pageId="1" pageNumber="28" type="nomenclature">
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<paragraph id="8BA4CB68FFD1006BFF445899FDF8F943" blockId="1.[151,579,1735,1762]" box="[151,579,1735,1762]" pageId="1" pageNumber="28">
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<heading id="D0EC7C04FFD1006BFF445899FDF8F943" bold="true" box="[151,579,1735,1762]" fontSize="11" level="1" pageId="1" pageNumber="28" reason="1">
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<emphasis id="B96F177AFFD1006BFF445899FDF8F943" bold="true" box="[151,579,1735,1762]" pageId="1" pageNumber="28">
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Genus
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<taxonomicName id="4C1BB0EBFFD1006BFF3D5899FDF8F943" ID-CoL="7NW7G" authority="Demange, 1961" authorityName="Demange" authorityYear="1961" box="[238,579,1735,1762]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD1006BFF3D5899FEC7F940" bold="true" box="[238,380,1735,1762]" italics="true" pageId="1" pageNumber="28">Carlogonus</emphasis>
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<bibRefCitation id="EF8AB699FFD1006BFE575899FDF8F943" author="Demange, J. M." box="[388,579,1735,1761]" pageId="1" pageNumber="28" pagination="1 - 274" refId="ref7005" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae (Myriapodes-Diplopodes). Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie, 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30198E3FFD1006BFF445952FE4BF886" box="[151,496,1803,1828]" pageId="1" pageNumber="28" type="reference_group">
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<paragraph id="8BA4CB68FFD1006BFF445952FE4BF886" blockId="1.[151,496,1803,1828]" box="[151,496,1803,1828]" pageId="1" pageNumber="28">
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<treatmentCitationGroup id="AB0BEC46FFD1006BFF445952FE4BF886" box="[151,496,1803,1828]" pageId="1" pageNumber="28">
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<treatmentCitation id="0ABAED79FFD1006BFF445952FE57F886" author="Demange, J. M." box="[151,492,1803,1828]" page="232" pageId="1" pageNumber="28" year="1961">
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<taxonomicName id="4C1BB0EBFFD1006BFF445952FE57F886" ID-CoL="7NW7G" authority="Demange, 1961: 232" authorityName="Demange" authorityPageNumber="232" authorityYear="1961" box="[151,492,1803,1828]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD1006BFF445952FEB4F886" box="[151,271,1804,1828]" italics="true" pageId="1" pageNumber="28">Carlogonus</emphasis>
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<bibRefCitation id="EF8AB699FFD1006BFEC65952FE57F886" author="Demange, J. M." box="[277,492,1803,1828]" pageId="1" pageNumber="28" pagination="1 - 274" refId="ref7005" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae (Myriapodes-Diplopodes). Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie, 24, 1 - 274." type="journal article" year="1961">Demange, 1961: 232</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="C30198E3FFD1006BFF445917FC39F8C3" box="[151,898,1865,1889]" pageId="1" pageNumber="28" type="type_taxon">
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<paragraph id="8BA4CB68FFD1006BFF445917FC39F8C3" blockId="1.[151,898,1865,1889]" box="[151,898,1865,1889]" pageId="1" pageNumber="28">
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<emphasis id="B96F177AFFD1006BFF445917FE98F8C3" bold="true" box="[151,291,1865,1889]" pageId="1" pageNumber="28">
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<typeStatus id="54A075CAFFD1006BFF445917FF77F8C3" box="[151,204,1865,1889]" pageId="1" pageNumber="28">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C1BB0EBFFD1006BFEF95917FD3CF8C3" ID-CoL="7BZCJ" authority="Carl, 1941" authorityName="Carl" authorityYear="1941" box="[298,647,1865,1889]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="subvalidus">
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<emphasis id="B96F177AFFD1006BFEF95917FDA8F8C3" box="[298,531,1865,1889]" italics="true" pageId="1" pageNumber="28">Thyropygus subvalidus</emphasis>
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<bibRefCitation id="EF8AB699FFD1006BFDC95917FD3CF8C3" author="Carl, J." box="[538,647,1865,1889]" pageId="1" pageNumber="28" pagination="569 - 714" refId="ref6848" refString="Carl, J. (1941) Diplopoden aus Sudindien und Ceylon. 2. Teil: Nematophora und Juliformia. Revue suisse de Zoologie, 48, 569 - 714. https: // doi. org / 10.5962 / bhl. part. 154483" type="journal article" year="1941">Carl, 1941</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30198E3FFD10068FF4459D1FEF1FC9B" lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28" type="diagnosis">
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<paragraph id="8BA4CB68FFD10068FF4459D1FAC8FEBF" blockId="1.[151,1437,1935,2034]" lastBlockId="2.[151,1437,151,1257]" lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28">
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<emphasis id="B96F177AFFD1006BFF4459D1FEA9F80B" bold="true" box="[151,274,1935,1961]" pageId="1" pageNumber="28">Diagnosis.</emphasis>
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Moderately large to large-sized millipedes (60–150), with a prominent, acute caudal spine, evenly curved downwards. Antero-lateral border of collum abruptly sinuate. Sternite of gonopod short with globular posterior parts. Anterior coxal fold lamelliform with modifications on the lateral margin and an antero-mesal hook-like process. Mesal margin of posterior coxal fold internally folded (towards back) to lie between the anterior coxal fold and telopodite. Telopodite strongly spiralled with unbranched palette, with femoral and tibial spines. Palette with blepharochaetae (
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<bibRefCitation id="EF8AB699FFD20068FE8E5E81FE4FFF5B" author="Attems, C." box="[349,500,223,249]" pageId="2" pageNumber="29" pagination="133 - 323" refId="ref6732" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 133 - 323." type="journal article" year="1936">Attems 1936</bibRefCitation>
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: fig. 78e–g,
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<bibRefCitation id="EF8AB699FFD20068FD505E81FD43FF5B" author="Carl, J." box="[643,760,223,249]" pageId="2" pageNumber="29" pagination="569 - 714" refId="ref6848" refString="Carl, J. (1941) Diplopoden aus Sudindien und Ceylon. 2. Teil: Nematophora und Juliformia. Revue suisse de Zoologie, 48, 569 - 714. https: // doi. org / 10.5962 / bhl. part. 154483" type="journal article" year="1941">Carl 1941</bibRefCitation>
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: figs 143–147, 149–150,
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<bibRefCitation id="EF8AB699FFD20068FBCA5E81FB6EFF5B" author="Demange, J. - M." box="[1049,1237,223,249]" pageId="2" pageNumber="29" pagination="231 - 235" refId="ref7044" refString="Demange, J. - M. (1977 a) Harpagophoridae (Myriapodes, Diplopodes) de l'Inde nouveaux ou peu connus. Bulletin du Museum national d'Histoire naturelle, Serie 3, Zoologie, 231, 231 - 235." type="journal article" year="1977">Demange 1977a</bibRefCitation>
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: figs 2–5; 1977b: figs 1–8; 1981: figs 14–15, 18; 1983: figs 11–15 and herein
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<figureCitation id="1320D7EDFFD20068FCEB5F5DFCC6FEBF" box="[824,893,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,547,571]" captionTargetBox="[156,1430,279,518]" captionTargetId="figure-75@4.[151,1437,273,524]" captionTargetPageId="4" captionText="FIGURE 2. Field photographs of Carlogonus gayathri sp. nov.. A: Male from Pullodu in Trippalur, Palakkad (MILLI- ADSH0012). B: Female from Pullodu in Trippalur, Palakkad (MILLI-ADSH0013). Photo courtesy Jimmy Paul." figureDoi="http://doi.org/10.5281/zenodo.4417207" httpUri="https://zenodo.org/record/4417207/files/figure.png" pageId="2" pageNumber="29">Figs 2</figureCitation>
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A–B, 3A, C, E, M–N, 5A–D, 7A–H, 8A–H).
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</paragraph>
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<paragraph id="8BA4CB68FFD20068FF145F79FEF1FC9B" blockId="2.[151,1437,151,1257]" pageId="2" pageNumber="29">
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<taxonomicName id="4C1BB0EBFFD20068FF145F79FEF0FEE3" authorityName="Demange" authorityYear="1961" box="[199,331,295,321]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD20068FF145F79FEF0FEE3" box="[199,331,295,321]" italics="true" pageId="2" pageNumber="29">Carlogonus</emphasis>
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</taxonomicName>
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species can be distinguished from the species of
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<taxonomicName id="4C1BB0EBFFD20068FCA05F79FBA5FEE3" authorityName="Attems" authorityYear="1942" box="[883,1054,295,321]" class="Diplopoda" family="Harpagophoridae" genus="Gnomognathus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD20068FCA05F79FBA5FEE3" box="[883,1054,295,321]" italics="true" pageId="2" pageNumber="29">Gnomognathus</emphasis>
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</taxonomicName>
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by the presence of an unbranched palette of telopodite and single/paired, long/short, slightly curved/straight tibial spines (
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<taxonomicName id="4C1BB0EBFFD20068FBB35F15FAB0FEC7" authorityName="Attems" authorityYear="1942" box="[1120,1291,331,357]" class="Diplopoda" family="Harpagophoridae" genus="Gnomognathus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD20068FBB35F15FAB0FEC7" box="[1120,1291,331,357]" italics="true" pageId="2" pageNumber="29">Gnomognathus</emphasis>
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</taxonomicName>
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species have a gonopod telopodite with a branched/unbranched palette and single/more than one, short tibial spines); from the species of
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<taxonomicName id="4C1BB0EBFFD20068FEDF5FCDFE7EFE0E" authorityName="Attems" authorityYear="1936" box="[268,453,403,428]" class="Diplopoda" family="Harpagophoridae" genus="Harpurostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD20068FEDF5FCDFE7EFE0E" box="[268,453,403,428]" italics="true" pageId="2" pageNumber="29">Harpurostreptus</emphasis>
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</taxonomicName>
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by the modified lateral margin of anterior coxal fold, telopodite with tibial spine and the absence of short median spines on the inner surface of the telopodite (
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<taxonomicName id="4C1BB0EBFFD20068FC615FE9FBD0FE72" authorityName="Attems" authorityYear="1936" box="[946,1131,439,464]" class="Diplopoda" family="Harpagophoridae" genus="Harpurostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD20068FC615FE9FBD0FE72" box="[946,1131,439,464]" italics="true" pageId="2" pageNumber="29">Harpurostreptus</emphasis>
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</taxonomicName>
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species lack modifications on the lateral margin of the anterior coxal fold and the tibial spine on the telopodite and instead possess short median spines on the inner surface of the telopodite); from the species of
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<taxonomicName id="4C1BB0EBFFD20068FC7D5FA1FBD4FDBB" authorityName="Demange" authorityYear="1989" box="[942,1135,511,537]" class="Diplopoda" family="Harpagophoridae" genus="Janardananeptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD20068FC7D5FA1FBD4FDBB" box="[942,1135,511,537]" italics="true" pageId="2" pageNumber="29">Janardananeptus</emphasis>
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</taxonomicName>
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by the short sternite of the gonopod, weak indentations on the lateral margin of the anterior coxal fold and the telopodite with an unbranched palette (
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<taxonomicName id="4C1BB0EBFFD20068FF205C19FE0FFDC3" authorityName="Demange" authorityYear="1989" box="[243,436,583,609]" class="Diplopoda" family="Harpagophoridae" genus="Janardananeptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD20068FF205C19FE0FFDC3" box="[243,436,583,609]" italics="true" pageId="2" pageNumber="29">Janardananeptus</emphasis>
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</taxonomicName>
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species have long sternite of the gonopod, deep indentations on the lateral margin of the anterior coxal fold and a branched palette); and from the species of
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<taxonomicName id="4C1BB0EBFFD20068FC575C35FB88FD27" authorityName="Verhoeff" authorityYear="1936" box="[900,1075,619,645]" class="Diplopoda" family="Harpagophoridae" genus="Organognathus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B96F177AFFD20068FC575C35FB88FD27" box="[900,1075,619,645]" italics="true" pageId="2" pageNumber="29">Organognathus</emphasis>
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</taxonomicName>
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by the absence of an antero-mesal spine-like modification on the posterior coxal fold and an unbranched palette of the telopodite (
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<taxonomicName id="4C1BB0EBFFD20068FB3E5CD1FA27FD0B" authorityName="Verhoeff" authorityYear="1936" box="[1261,1436,655,681]" class="Diplopoda" family="Harpagophoridae" genus="Organognathus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96F177AFFD20068FB3E5CD1FA27FD0B" box="[1261,1436,655,681]" italics="true" pageId="2" pageNumber="29">Organognathus</emphasis>
|
||
</taxonomicName>
|
||
species have a strong antero-mesal spine-like modification on the posterior coxal fold and a branched palette of the telopodite) (compare
|
||
<figureCitation id="1320D7EDFFD20068FE615C89FE49FD53" box="[434,498,727,753]" captionStart="FIGURE 7" captionStartId="9.[151,250,1671,1695]" captionTargetBox="[259,1327,186,1634]" captionTargetId="figure-17@9.[246,1342,181,1647]" captionTargetPageId="9" captionText="FIGURE 7. Carlogonus gayathri sp. nov., male. A: Gonopods, ventral view. B: Same, dorsal view. C: Right gonopod after removing telopodite, dorsal view. D: Left gonopod, lateral view. E: Same, mesal view. F: Left telopodite, ventral view. G: Same, lateral view. H: Enlarged view of distal part of left telopodite, mesal view. I: Enlarged view of blepharochaetae, lateral view. Abbreviations: alpAC = antero-lateral process of anterior coxal fold, amhpAC = antero-mesal hook-like process of anterior coxal fold, amtAC = antero-mesal thumb-like process of anterior coxal fold, AC = anterior coxal fold, B = blepharochaetae, C = distal longitudinal crest, F = femur, FS = femoral spine, dtpP = distal thorne-like process of palette, P = palette, apPC = apical process of posterior coxal fold, blpPC = baso-lateral process of posterior coxal fold, imPC = internal mesal fold of posterior coxal fold, lpPC = lateral process of posterior coxal fold, mpPC = mesal process of posterior coxal fold, PC = posterior coxal fold, PX = paracoxite, ST = sternite, T = telopodite, TS = tibial spine, TT = tibiotarsus. Arrows indicate indentation on lateral margin of anterior coxal fold (1) and basal bifurcated seta on tibial spine (2). Scale bars: A–G = 1 mm; H–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4417222" httpUri="https://zenodo.org/record/4417222/files/figure.png" pageId="2" pageNumber="29">Fig. 7</figureCitation>
|
||
A–H with
|
||
<bibRefCitation id="EF8AB699FFD20068FDBB5C89FD47FD53" author="Attems, C." box="[616,764,727,753]" pageId="2" pageNumber="29" pagination="133 - 323" refId="ref6732" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 133 - 323." type="journal article" year="1936">Attems 1936</bibRefCitation>
|
||
: figs 59d, 77f,
|
||
<bibRefCitation id="EF8AB699FFD20068FC775C89FBACFD53" author="Carl, J." box="[932,1047,727,753]" pageId="2" pageNumber="29" pagination="569 - 714" refId="ref6848" refString="Carl, J. (1941) Diplopoden aus Sudindien und Ceylon. 2. Teil: Nematophora und Juliformia. Revue suisse de Zoologie, 48, 569 - 714. https: // doi. org / 10.5962 / bhl. part. 154483" type="journal article" year="1941">Carl 1941</bibRefCitation>
|
||
: figs 139, 168–169, 178, 190, 194, 196–197, 214, 217, 220, 222,
|
||
<bibRefCitation id="EF8AB699FFD20068FE375CA5FD29FCB7" author="Demange, J. M." box="[484,658,763,789]" pageId="2" pageNumber="29" pagination="1 - 274" refId="ref7005" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae (Myriapodes-Diplopodes). Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie, 24, 1 - 274." type="journal article" year="1961">Demange 1961</bibRefCitation>
|
||
: figs 322, 328–330; 1977a: figs 6, 8; 1989: figs 1, 3–4, 6 and
|
||
<bibRefCitation id="EF8AB699FFD20068FAE85CA5FF68FC9B" author="Verhoeff, K. W." pageId="2" pageNumber="29" pagination="103 - 123" refId="ref8042" refString="Verhoeff, K. W. (1936) Ueber einige indische Chilognathen gesammelt von Herrn S. Jones, Madras. Records of the Indian Museum, 38, 103 - 123." type="journal article" year="1936">Verhoeff 1936</bibRefCitation>
|
||
: figs 2–4).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30198E3FFD20068FF145D1CFDCFFB4B" pageId="2" pageNumber="29" type="distribution">
|
||
<paragraph id="8BA4CB68FFD20068FF145D1CFDC5FCFF" blockId="2.[151,1437,151,1257]" box="[199,638,834,861]" pageId="2" pageNumber="29">
|
||
<emphasis id="B96F177AFFD20068FF145D1CFEDAFCFE" bold="true" box="[199,353,834,860]" pageId="2" pageNumber="29">Distribution.</emphasis>
|
||
<collectingCountry id="F30C8BF8FFD20068FEB45D1DFE18FCFF" box="[359,419,835,861]" name="India" pageId="2" pageNumber="29">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="F30C8BF8FFD20068FE7C5D1DFDA6FCFF" box="[431,541,835,861]" name="Sri Lanka" pageId="2" pageNumber="29">Sri Lanka</collectingCountry>
|
||
(
|
||
<figureCitation id="1320D7EDFFD20068FDFF5D1DFDCBFCFF" box="[556,624,835,861]" captionStart="FIGURE 9" captionStartId="11.[151,250,1495,1519]" captionTargetBox="[158,1430,531,1464]" captionTargetId="figure-165@11.[151,1436,525,1471]" captionTargetPageId="11" captionText="FIGURE 9. Currently known distribution records of Carlogonus spp.. * Carlogonus acifer Demange, 1977, ♦ Carlogonus auriculus Demange, 1983, é Carlogonus chowdaiahi Demange, 1977, ■ Carlogonus exaratus (Attems, 1936), r Carlogonus gayathri sp. nov., ● Carlogonus palmatus Demange, 1977, □ Carlogonus robustior (Attems, 1936), ○ Carlogonus subvalidus (Carl, 1941), ▲ Carlogonus verhoeffi Demange, 1981 (record of Carlogonus spinicaudus (Gervais, 1847) and subsequent records of other species are omitted from the map due to the lack of locality details)." figureDoi="http://doi.org/10.5281/zenodo.4417228" httpUri="https://zenodo.org/record/4417228/files/figure.png" pageId="2" pageNumber="29">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA4CB68FFD20068FF145D38FDA0FBB3" blockId="2.[151,1437,151,1257]" pageId="2" pageNumber="29">
|
||
<emphasis id="B96F177AFFD20068FF145D38FE3AFC22" bold="true" box="[199,385,870,896]" pageId="2" pageNumber="29">Species-groups.</emphasis>
|
||
In total, there are 10 nominal
|
||
<taxonomicName id="4C1BB0EBFFD20068FD095D39FCE5FC23" authorityName="Demange" authorityYear="1961" box="[730,862,871,897]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96F177AFFD20068FD095D39FCE5FC23" box="[730,862,871,897]" italics="true" pageId="2" pageNumber="29">Carlogonus</emphasis>
|
||
</taxonomicName>
|
||
species, including the new species described here. Based on a single character of the gonopod telopodite and not on phylogenetic evidences, nine species can be assigned into two species-groups. The species
|
||
<taxonomicName id="4C1BB0EBFFD20068FD565DF1FBEAFC6B" authority="(Gervais, 1847)" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[645,1105,943,969]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="spinicaudus">
|
||
<emphasis id="B96F177AFFD20068FD565DF1FC2DFC6B" box="[645,918,943,969]" italics="true" pageId="2" pageNumber="29">Carlogonus spinicaudus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AB699FFD20068FC775DF1FBF9FC6B" author="Gervais, P." box="[932,1090,943,969]" pageId="2" pageNumber="29" refId="ref7269" refString="Gervais, P. (1847) Myriapodes. In: Walckenaer, C. A. & Gervais, P. (Eds.), Histoire naturelle des Insectes, Apteres. Vol. 4. Roret, Paris, pp. 1 - 623." type="book" year="1847">Gervais, 1847</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is omitted due to the unavailability of relevant character details. Moreover, this species may be a junior synonym of
|
||
<taxonomicName id="4C1BB0EBFFD20068FB455D8DFDACFBB3" authority="(Carl, 1941) (Mauries et al. 2001)" authorityName="Mauries et al." authorityYear="2001" baseAuthorityName="Carl" baseAuthorityYear="1941" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="subvalidus">
|
||
<emphasis id="B96F177AFFD20068FB455D8DFA27FC4F" box="[1174,1436,979,1005]" italics="true" pageId="2" pageNumber="29">Carlogonus subvalidus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AB699FFD20068FF4C5DA9FEA3FBB3" author="Carl, J." box="[159,280,1015,1041]" pageId="2" pageNumber="29" pagination="569 - 714" refId="ref6848" refString="Carl, J. (1941) Diplopoden aus Sudindien und Ceylon. 2. Teil: Nematophora und Juliformia. Revue suisse de Zoologie, 48, 569 - 714. https: // doi. org / 10.5962 / bhl. part. 154483" type="journal article" year="1941">Carl, 1941</bibRefCitation>
|
||
) (
|
||
<bibRefCitation id="EF8AB699FFD20068FEFC5DA9FDB5FBB3" author="Mauries, J. - P. & Golovatch, S. I. & Hoffman, R. L." box="[303,526,1015,1041]" pageId="2" pageNumber="29" pagination="579 - 589" refId="ref7567" refString="Mauries, J. - P., Golovatch, S. I. & Hoffman, R. L. (2001) On type material and the identity of several Iulus species described by Paul Gervais, in the collection of the Museum national d'Histoire naturelle in Paris (Diplopoda, Spirostreptida, Spirobolida). Zoosystema, 23, 579 - 589." type="journal article" year="2001">
|
||
Mauriès
|
||
<emphasis id="B96F177AFFD20068FE405DA9FE70FBB3" box="[403,459,1015,1041]" italics="true" pageId="2" pageNumber="29">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA4CB68FFD20068FF145A45FE2DFBDF" blockId="2.[151,1437,151,1257]" pageId="2" pageNumber="29">
|
||
<emphasis id="B96F177AFFD20068FF145A45FE00FB97" bold="true" box="[199,443,1051,1077]" pageId="2" pageNumber="29">
|
||
The
|
||
<taxonomicName id="4C1BB0EBFFD20068FF2D5A45FED9FB97" baseAuthorityName="Attems" baseAuthorityYear="1936" box="[254,354,1051,1077]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="exaratus">
|
||
<emphasis id="B96F177AFFD20068FF2D5A45FED9FB97" bold="true" box="[254,354,1051,1077]" italics="true" pageId="2" pageNumber="29">exaratus</emphasis>
|
||
</taxonomicName>
|
||
-group:
|
||
</emphasis>
|
||
Species-group with a single tibial spine on the gonopod telopodite. It includes five spe-cies—
|
||
<taxonomicName id="4C1BB0EBFFD20068FF0C5A61FD34FBFB" authority="Demange, 1983" authorityName="Demange" authorityYear="1983" box="[223,655,1087,1113]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="auriculus">
|
||
<emphasis id="B96F177AFFD20068FF0C5A61FE6FFBFB" box="[223,468,1087,1113]" italics="true" pageId="2" pageNumber="29">Carlogonus auriculus</emphasis>
|
||
<bibRefCitation id="EF8AB699FFD20068FE085A61FD34FBFB" author="Demange, J. - M." box="[475,655,1087,1113]" pageId="2" pageNumber="29" pagination="561 - 584" refId="ref7169" refString="Demange, J. - M. (1983) Donnees nouvelles sur la famille des Harpagophoridae (Myriapoda, Diplopoda). Bulletin du Museum national d'Histoire naturelle, Serie 4, Section A, 5 (2), 561 - 584." type="journal article" year="1983">Demange, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1BB0EBFFD20068FD495A61FB82FBF8" authority="(Attems, 1936)" baseAuthorityName="Attems" baseAuthorityYear="1936" box="[666,1081,1087,1114]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="exaratus">
|
||
<emphasis id="B96F177AFFD20068FD495A61FC3DFBFA" box="[666,902,1087,1113]" italics="true" pageId="2" pageNumber="29">Carlogonus exaratus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AB699FFD20068FC465A61FB89FBF8" author="Attems, C." box="[917,1074,1087,1114]" pageId="2" pageNumber="29" pagination="133 - 323" refId="ref6732" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 133 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1BB0EBFFD20068FB965A61FB7EFBFB" authority="Sankaran & Sebastian, 2020" authorityName="Sankaran & Sebastian" authorityYear="2020" box="[1093,1221,1087,1113]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="gayathri" status="sp. nov.">
|
||
<emphasis id="B96F177AFFD20068FB965A61FB7EFBFB" box="[1093,1221,1087,1113]" italics="true" pageId="2" pageNumber="29">C. gayathri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96F177AFFD20068FB1F5A61FA9EFBFB" bold="true" box="[1228,1317,1087,1113]" pageId="2" pageNumber="29">
|
||
<taxonomicNameLabel id="A25CAA01FFD20068FB1F5A61FA9EFBFB" box="[1228,1317,1087,1113]" pageId="2" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C1BB0EBFFD20068FAE05A61FF04FBDE" baseAuthorityName="Attems" baseAuthorityYear="1936" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="robustior">
|
||
<emphasis id="B96F177AFFD20068FAE05A61FF04FBDE" italics="true" pageId="2" pageNumber="29">C. robustior</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1BB0EBFFD20068FF255A3DFE34FBDF" authorityName="Mauries et al." authorityYear="2001" baseAuthorityName="Carl" baseAuthorityYear="1941" box="[246,399,1123,1149]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="subvalidus">
|
||
<emphasis id="B96F177AFFD20068FF255A3DFE34FBDF" box="[246,399,1123,1149]" italics="true" pageId="2" pageNumber="29">C. subvalidus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA4CB68FFD20068FF145AD9FDCFFB4B" blockId="2.[151,1437,151,1257]" pageId="2" pageNumber="29">
|
||
<emphasis id="B96F177AFFD20068FF145AD9FE23FB03" bold="true" box="[199,408,1158,1185]" pageId="2" pageNumber="29">
|
||
The
|
||
<taxonomicName id="4C1BB0EBFFD20068FF2F5AD8FE84FB02" authorityName="Demange" authorityYear="1977" box="[252,319,1158,1184]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="acifer">
|
||
<emphasis id="B96F177AFFD20068FF2F5AD8FE84FB02" bold="true" box="[252,319,1158,1184]" italics="true" pageId="2" pageNumber="29">acifer</emphasis>
|
||
</taxonomicName>
|
||
-group:
|
||
</emphasis>
|
||
Species-group with paired tibial spines on the gonopod telopodite. It includes four species—
|
||
<taxonomicName id="4C1BB0EBFFD20068FF445AF5FDA7FB67" authority="Demange, 1977" authorityName="Demange" authorityYear="1977" box="[151,540,1195,1221]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="acifer">
|
||
<emphasis id="B96F177AFFD20068FF445AF5FED9FB67" box="[151,354,1195,1221]" italics="true" pageId="2" pageNumber="29">Carlogonus acifer</emphasis>
|
||
Demange, 1977
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1BB0EBFFD20068FDF55AF5FC50FB67" authority="Demange, 1977" authorityName="Demange" authorityYear="1977" box="[550,1003,1195,1221]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="chowdaiahi">
|
||
<emphasis id="B96F177AFFD20068FDF55AF5FC88FB67" box="[550,819,1195,1221]" italics="true" pageId="2" pageNumber="29">Carlogonus chowdaiahi</emphasis>
|
||
Demange, 1977
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1BB0EBFFD20068FC265AF5FA27FB67" authority="Demange, 1977" authorityName="Demange" authorityYear="1977" box="[1013,1436,1195,1221]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="palmatus">
|
||
<emphasis id="B96F177AFFD20068FC265AF5FB5DFB67" box="[1013,1254,1195,1221]" italics="true" pageId="2" pageNumber="29">Carlogonus palmatus</emphasis>
|
||
Demange, 1977
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1BB0EBFFD20068FF145A91FDCBFB4B" authority="Demange, 1981" authorityName="Demange" authorityYear="1981" box="[199,624,1231,1257]" class="Diplopoda" family="Harpagophoridae" genus="Carlogonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="verhoeffi">
|
||
<emphasis id="B96F177AFFD20068FF145A91FE0EFB4B" box="[199,437,1231,1257]" italics="true" pageId="2" pageNumber="29">Carlogonus verhoeffi</emphasis>
|
||
<bibRefCitation id="EF8AB699FFD20068FE6F5A91FDCBFB4B" author="Demange, J. - M." box="[444,624,1231,1257]" pageId="2" pageNumber="29" pagination="63 - 80" refId="ref7139" refString="Demange, J. - M. (1981) Spirostreptida, Harpagophoridae (Myriapoda-Diplopoda) de Sri Lanka. Entomologica Scandinavica, Supplement 11, 63 - 80." type="journal article" year="1981">Demange, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |