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<document id="277CA8AC055D57ADF2140EA710DD5881" ID="10.11646/zootaxa.4168.3.10" ID-DOI="10.11646/zootaxa.4168.3.10" ID-GBIF-Dataset="570048e9-7021-4281-87b9-426846734613" ID-ISSN="1175-5326" ID-Zenodo-Dep="262052" ID-ZooBank="908C77AF-951F-477B-93AF-83EA1EF98F33" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474010193695" checkinUser="plazi" docAuthor="Sanborn, Allen F. &amp; Heath, Maxine S." docDate="2016" docId="03A884555B2EFF8EFF6DFF6EC40D5DCA" docLanguage="en" docName="zootaxa.4168.3.10.pdf" docOrigin="Zootaxa 4168 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megatibicen Sanborn &amp; Heath, 2016, n. gen." docType="treatment" docVersion="6" lastPageNumber="580" masterDocId="FF91FC2D5B2FFF8DFFFAFFB3C759587E" masterDocTitle="Megatibicen n. gen., a new North American cicada genus (Hemiptera: Cicadidae: Cicadinae: Cryptotympanini)" masterLastPageNumber="582" masterPageNumber="577" pageNumber="578" updateTime="1698386108734" updateUser="plazi">
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<mods:title id="4317F7ED535092692F4BA75286E1192C">Megatibicen n. gen., a new North American cicada genus (Hemiptera: Cicadidae: Cicadinae: Cryptotympanini)</mods:title>
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<mods:namePart id="B08407CE713020191D7E70E9FFD20725">Sanborn, Allen F.</mods:namePart>
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<mods:namePart id="45BC19DF15A1ECDB590F4FEFB9BE4FC6">Heath, Maxine S.</mods:namePart>
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<mods:date id="81BB8E56560D875D1BFCC2EAED3E1DA7">2016</mods:date>
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<treatment id="03A884555B2EFF8EFF6DFF6EC40D5DCA" ID-DOI="http://doi.org/10.5281/zenodo.6058684" ID-GBIF-Taxon="159862508" ID-Zenodo-Dep="6058684" LSID="urn:lsid:plazi:treatment:03A884555B2EFF8EFF6DFF6EC40D5DCA" httpUri="http://treatment.plazi.org/id/03A884555B2EFF8EFF6DFF6EC40D5DCA" lastPageId="3" lastPageNumber="580" pageId="1" pageNumber="578">
<subSubSection id="C31B66C85B2EFF8CFF6DFF6EC6845889" box="[151,477,220,247]" pageId="1" pageNumber="578" type="nomenclature">
<paragraph id="8BBE35435B2EFF8CFF6DFF6EC6845889" blockId="1.[151,1437,152,2013]" box="[151,477,220,247]" pageId="1" pageNumber="578">
<emphasis id="B975E9515B2EFF8CFF6DFF6EC6845889" bold="true" box="[151,477,220,247]" pageId="1" pageNumber="578">
Genus
<taxonomicName id="4C014EC05B2EFF8CFF14FF6FC6DD5888" ID-CoL="8HM53" box="[238,388,220,246]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975E9515B2EFF8CFF14FF6FC6DD5888" bold="true" box="[238,388,220,246]" italics="true" pageId="1" pageNumber="578">Megatibicen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246542A5B2EFF8CFE71FF6EC6845889" box="[395,477,221,247]" pageId="1" pageNumber="578" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31B66C85B2EFF8CFF6DFEB3C2C25969" box="[151,1435,256,279]" pageId="1" pageNumber="578" type="type_taxon">
<paragraph id="8BBE35435B2EFF8CFF6DFEB3C2C25969" blockId="1.[151,1437,152,2013]" box="[151,1435,256,279]" pageId="1" pageNumber="578">
<emphasis id="B975E9515B2EFF8CFF6DFEB3C6735969" bold="true" box="[151,298,256,279]" pageId="1" pageNumber="578">
<typeStatus id="54BA8BE15B2EFF8CFF6DFEB3C7965969" box="[151,207,256,279]" pageId="1" pageNumber="578">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C014EC05B2EFF8CFECAFEB3C4535969" authority="Davis 1915: 162" authorityName="Davis" authorityPageNumber="162" authorityYear="1915" box="[304,778,256,279]" class="Insecta" family="Cicadellidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="subSpecies" species="marginata" subSpecies="dealbata">
<emphasis id="B975E9515B2EFF8CFECAFEB3C50C5969" box="[304,597,256,279]" italics="true" pageId="1" pageNumber="578">Cicada marginata dealbata</emphasis>
<bibRefCitation id="EF9048B25B2EFF8CFDA1FEB3C4535969" author="Davis" box="[603,778,256,279]" firstAuthor="Davis" pageId="1" pageNumber="578" pagination="161 - 164" refId="ref4109" refString="Davis, W. T. (1915) A new variety of Cicada resembling C. dorsata. Journal of the New York Entomological Society, 23, 161 - 164." type="journal article" year="1915">Davis 1915: 162</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="49C5FBA15B2EFF8CFCEDFEB3C4255969" box="[791,892,256,279]" country="United States of America" name="Colorado" pageId="1" pageNumber="578">Colorado</collectingRegion>
Springs, El Paso County,
<collectingRegion id="49C5FBA15B2EFF8CFB6BFEB3C3AE5969" box="[1169,1271,256,279]" country="United States of America" name="Colorado" pageId="1" pageNumber="578">Colorado</collectingRegion>
,
<collectingCountry id="F31675D35B2EFF8CFAFBFEB3C2615969" box="[1281,1336,256,279]" name="United States of America" pageId="1" pageNumber="578">USA</collectingCountry>
) (
<figureCitation id="133A29C65B2EFF8CFAB7FEB3C2D25969" box="[1357,1419,256,279]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B66C85B2EFF8CFF3DFE91C55559BA" pageId="1" pageNumber="578" type="nomenclature">
<paragraph id="8BBE35435B2EFF8CFF3DFE91C55559BA" blockId="1.[151,1437,152,2013]" pageId="1" pageNumber="578">
<emphasis id="B975E9515B2EFF8CFF3DFE91C6D35947" bold="true" box="[199,394,290,313]" pageId="1" pageNumber="578">Species included.</emphasis>
The genus is represented currently by the following nine species and one subspecies:
<taxonomicName id="4C014EC05B2EFF8CFAD6FE91C6675922" ID-CoL="6QXM6" authority="Germar, 1834" authorityName="Germar" authorityYear="1834" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="auletes" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFAD6FE91C2C25947" box="[1324,1435,290,313]" italics="true" pageId="1" pageNumber="578">M. auletes</emphasis>
(Germar, 1834)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFEBCFEF6C6F05922" bold="true" box="[326,425,325,348]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFEBCFEF6C6F05922" box="[326,425,325,348]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFE43FEF6C5A35922" authority="Davis, 1915" authorityName="Davis" authorityYear="1915" box="[441,762,325,348]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="cultriformis" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFE43FEF6C5055922" box="[441,604,325,348]" italics="true" pageId="1" pageNumber="578">M. cultriformis</emphasis>
(
<bibRefCitation id="EF9048B25B2EFF8CFD96FEF6C5AB5922" author="Davis" box="[620,754,325,348]" firstAuthor="Davis" pageId="1" pageNumber="578" pagination="161 - 164" refId="ref4109" refString="Davis, W. T. (1915) A new variety of Cicada resembling C. dorsata. Journal of the New York Entomological Society, 23, 161 - 164." type="journal article" year="1915">Davis, 1915</bibRefCitation>
)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFCF8FEF6C43C5922" bold="true" box="[770,869,325,348]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFCF8FEF6C43C5922" box="[770,869,325,348]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFC8FFEF6C3F95922" authority="Davis, 1915" authorityName="Davis" authorityYear="1915" box="[885,1184,325,348]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="dealbatus" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFC8FFEF6C35B5922" box="[885,1026,325,348]" italics="true" pageId="1" pageNumber="578">M. dealbatus</emphasis>
(
<bibRefCitation id="EF9048B25B2EFF8CFBE8FEF6C3CE5922" author="Davis" box="[1042,1175,325,348]" firstAuthor="Davis" pageId="1" pageNumber="578" pagination="161 - 164" refId="ref4109" refString="Davis, W. T. (1915) A new variety of Cicada resembling C. dorsata. Journal of the New York Entomological Society, 23, 161 - 164." type="journal article" year="1915">Davis, 1915</bibRefCitation>
)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFB52FEF6C2525922" bold="true" box="[1192,1291,325,348]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFB52FEF6C2525922" box="[1192,1291,325,348]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFAE1FEF6C64F5901" authority="Say, 1825" authorityName="Say" authorityYear="1825" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="dorsatus" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFAE1FEF6C2C55922" box="[1307,1436,325,348]" italics="true" pageId="1" pageNumber="578">M. dorsatus</emphasis>
(Say, 1825)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFEDBFEDBC6DF5901" bold="true" box="[289,390,360,383]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFEDBFEDBC6DF5901" box="[289,390,360,383]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFE6DFEDBC5975901" authority="Walker, 1858" authorityName="Walker" authorityYear="1858" box="[407,718,360,383]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="figuratus" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFE6DFEDBC5465901" box="[407,543,360,383]" italics="true" pageId="1" pageNumber="578">M. figuratus</emphasis>
(Walker, 1858)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFD23FEDBC4675901" bold="true" box="[729,830,360,383]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFD23FEDBC4675901" box="[729,830,360,383]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFCAAFEDBC4BD5901" box="[848,996,360,383]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="pronotalis">
<emphasis id="B975E9515B2EFF8CFCAAFEDBC4BD5901" box="[848,996,360,383]" italics="true" pageId="1" pageNumber="578">M. pronotalis</emphasis>
</taxonomicName>
<taxonomicName id="4C014EC05B2EFF8CFC15FEDBC3A75901" authority="Davis, 1938" authorityName="Davis" authorityYear="1938" box="[1007,1278,360,383]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="pronotalis" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFC15FEDBC3025901" box="[1007,1115,360,383]" italics="true" pageId="1" pageNumber="578">pronotalis</emphasis>
(Davis, 1938)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFAF2FEDBC2375901" bold="true" box="[1288,1390,360,383]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFAF2FEDBC2375901" box="[1288,1390,360,383]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFA85FEDBC65D59DF" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="pronotalis">
<emphasis id="B975E9515B2EFF8CFA85FEDBC65D59DF" italics="true" pageId="1" pageNumber="578">M. pronotalis</emphasis>
</taxonomicName>
<taxonomicName id="4C014EC05B2EFF8CFEF0FE39C55E59DF" authority="Metcalf, 1955" authorityName="Metcalf" authorityYear="1955" box="[266,519,394,417]" class="Insecta" family="Cicadidae" genus="Tibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="walkeri" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFEF0FE39C60359DF" box="[266,346,394,417]" italics="true" pageId="1" pageNumber="578">walkeri</emphasis>
(Metcalf, 1955)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFDF7FE39C53759DF" bold="true" box="[525,622,394,417]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFDF7FE39C53759DF" box="[525,622,394,417]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFD81FE39C4CB59DF" authority="Haldeman, 1852" authorityName="Haldeman" authorityYear="1852" box="[635,914,394,417]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="resh" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFD81FE39C59259DF" box="[635,715,394,417]" italics="true" pageId="1" pageNumber="578">M. resh</emphasis>
(Haldeman, 1852)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFC63FE39C4A059DF" bold="true" box="[921,1017,394,417]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFC63FE39C4A059DF" box="[921,1017,394,417]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFBFCFE39C27659DF" authority="Walker, 1850" authorityName="Walker" authorityYear="1850" box="[1030,1327,394,417]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="resonans" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFBFCFE39C3D059DF" box="[1030,1161,394,417]" italics="true" pageId="1" pageNumber="578">M. resonans</emphasis>
(Walker, 1850)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFACEFE39C2CC59DF" bold="true" box="[1332,1429,394,417]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFACEFE39C2CC59DF" box="[1332,1429,394,417]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C014EC05B2EFF8CFF6DFE1EC6FD59BA" authority="Cole, 2008" authorityName="Cole" authorityYear="2008" box="[151,420,429,452]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="tremulus" status="comb. nov.">
<emphasis id="B975E9515B2EFF8CFF6DFE1EC64F59BA" box="[151,278,429,452]" italics="true" pageId="1" pageNumber="578">M. tremulus</emphasis>
(Cole, 2008)
</taxonomicName>
<emphasis id="B975E9515B2EFF8CFE51FE1EC55559BA" bold="true" box="[427,524,429,452]" pageId="1" pageNumber="578">
<taxonomicNameLabel id="A246542A5B2EFF8CFE51FE1EC55559BA" box="[427,524,429,452]" pageId="1" pageNumber="578">n. comb.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31B66C85B2EFF8CFF3DFE63C34B5A77" pageId="1" pageNumber="578" type="etymology">
<paragraph id="8BBE35435B2EFF8CFF3DFE63C34B5A77" blockId="1.[151,1437,152,2013]" pageId="1" pageNumber="578">
<emphasis id="B975E9515B2EFF8CFF3DFE63C61E5999" bold="true" box="[199,327,464,487]" pageId="1" pageNumber="578">Etymology.</emphasis>
The name is a combination of
<emphasis id="B975E9515B2EFF8CFD74FE63C5915999" box="[654,712,464,487]" italics="true" pageId="1" pageNumber="578">mega</emphasis>
(Gr. large) and
<taxonomicName id="4C014EC05B2EFF8CFC94FE63C4EF5999" box="[878,950,464,487]" class="Insecta" family="Cicadidae" genus="Tibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="tibicen">
<emphasis id="B975E9515B2EFF8CFC94FE63C4EF5999" box="[878,950,464,487]" italics="true" pageId="1" pageNumber="578">tibicen</emphasis>
</taxonomicName>
(L. flute player) in reference to the large size, acoustic abilities and previous classification of the species. The genus is masculine.
</paragraph>
</subSubSection>
<subSubSection id="C31B66C85B2EFF8EFF3DFDA6C36F5A0F" lastPageId="3" lastPageNumber="580" pageId="1" pageNumber="578" type="description">
<paragraph id="8BBE35435B2EFF8CFF3DFDA6C4A05F0B" blockId="1.[151,1437,152,2013]" pageId="1" pageNumber="578">
<emphasis id="B975E9515B2EFF8CFF3DFDA6C60B5A52" bold="true" box="[199,338,533,556]" pageId="1" pageNumber="578">Description.</emphasis>
Large bodied cicadas (body length 32.543.0 mm, expanse of fore wings 90.0129.0 mm); the abdomen represents about one-half the body length in females but is generally slightly longer in males because of the extended male genitalia; body length is at least two and one-half times the width of the mesonotum.
<emphasis id="B975E9515B2EFF8CFB38FDE9C3A55A0F" box="[1218,1276,602,625]" italics="true" pageId="1" pageNumber="578">Head</emphasis>
: width of head including eyes about as broad or broader than mesonotum (
<figureCitation id="133A29C65B2EFF8CFCE2FDCEC4095AEA" box="[792,848,637,660]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
B); eyes extend slightly beyond the anterior margin of the pronotum; head not as wide as lateral pronotal collar; vertex of epicranium horizontal with distance between supraantennal plate and eye about equal to or greater than length of antennal plate; frons nearly vertical, forming a right or sharply obtuse angle with the dorsum of the postclypeus; distance between ocelli less than distance between lateral ocelli and eyes; postclypeus ranges from more or less truncate, in a continuous line with the supra-antennal plates, to moderately tumid; dorsum of postclypeus about as long or longer than frons; length of rostrum species specific, ranging from middle coxae to beyond posterior coxae.
<emphasis id="B975E9515B2EFF8CFD7FFCFEC58B5B1A" box="[645,722,845,868]" italics="true" pageId="1" pageNumber="578">Thorax</emphasis>
: pronotum ampliated laterally forming definite lateral fissures (
<figureCitation id="133A29C65B2EFF8CFA8CFCFEC7FB5BF9" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
B), lateral edges smooth, not toothed, occasionally spinose; pronotal collar moderately ampliate laterally, width equal to or slightly less than head width across eyes; length of pronotum shorter then mesonotum; cruciform elevation depression between anterior arms diminishing towards extremities of arms, cruciform elevation much longer than wide; mesonotum obscuring central portion of metanotum; basisternum 3 bulbous; opercula in males large, extending to rim of the tympanal cavity, overlapping medially (
<figureCitation id="133A29C65B2EFF8CFDADFC49C5CB5C6F" box="[599,658,1018,1041]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
D); meracanthus triangular; female operculum with sinuate posterior margin (
<figureCitation id="133A29C65B2EFF8CFF15FBAEC67C5C4A" box="[239,293,1053,1076]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
E).
<emphasis id="B975E9515B2EFF8CFEB4FBAEC6D95C4A" box="[334,384,1053,1076]" italics="true" pageId="1" pageNumber="578">Legs</emphasis>
: anterior femora with two prominent spines beneath, sometimes with a third smaller spine distally; primary spine erect.
<emphasis id="B975E9515B2EFF8CFE85FBF3C5EB5C29" box="[383,690,1088,1111]" italics="true" pageId="1" pageNumber="578">Fore wings and hind wings</emphasis>
(
<figureCitation id="133A29C65B2EFF8CFD3FFBF3C5A65C29" box="[709,767,1088,1111]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
A): hyaline; radial and radiomedial crossveins in fore wing oblique, virtually parallel, and frequently infuscated; length of radius posterior between the radial and radiomedial crossveins less than radius posterior between the radial crossvein and the ambient vein; medial and cubital veins arising separately from the arculus; stem of median vein shorter than one half length of discal cell; nodus central; costal margin of the fore wing with at least a few minute spines; radius anterior 1 and subcostal vein not diverging in subapical area; hind wing cubital cell 1 distal end width about equal to or slightly wider than width of cubital cell 2, cubital cell 2 width at distal end greater than anal cell 1, hind wing margin not excessively broad.
<emphasis id="B975E9515B2EFF8CFBFAFAA3C33C5D59" box="[1024,1125,1296,1319]" italics="true" pageId="1" pageNumber="578">Abdomen</emphasis>
: first abdominal segment in males containing timbal organs; lateral tergites straight or weakly convex; epipleurites reflexed to ventral surface; tergites 2 and 3 about twice as wide as tergites 37; second male tergite produced anteriorly on each side into a complete, slightly domed timbal cover, fully rounded dorsally, extending to metathorax, and tightly enclosing timbal cavity, lower margin extending anteriorly from or very near auditory capsule (
<figureCitation id="133A29C65B2EFF8CFCB2FA29C4275DCF" box="[840,894,1434,1457]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
C); male sternites IVVII convex in cross-section, sternite VIII U-shaped in cross-section (
<taxonomicName id="4C014EC05B2EFF8CFDBBFA0EC5C95DAB" box="[577,656,1469,1493]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="auletes">
<emphasis id="B975E9515B2EFF8CFDBBFA0EC5C95DAB" box="[577,656,1469,1493]" italics="true" pageId="1" pageNumber="578">auletes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C014EC05B2EFF8CFD67FA0EC45B5DAB" box="[669,770,1469,1493]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="figuratus">
<emphasis id="B975E9515B2EFF8CFD67FA0EC45B5DAB" box="[669,770,1469,1493]" italics="true" pageId="1" pageNumber="578">figuratus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C014EC05B2EFF8CFCCDFA0DC4C55DAB" box="[823,924,1470,1493]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="resonans">
<emphasis id="B975E9515B2EFF8CFCCDFA0DC4C55DAB" box="[823,924,1470,1493]" italics="true" pageId="1" pageNumber="578">resonans</emphasis>
</taxonomicName>
), much less than the maximum width of sternite VII, or V-shaped in cross-section as wide or about as wide as maximum width of sternite VII.
<emphasis id="B975E9515B2EFF8CFB78FA52C2475D86" box="[1154,1310,1505,1528]" italics="true" pageId="1" pageNumber="578">Male genitalia</emphasis>
: pygofer of males considerably broadened towards top presenting a triangular shape or with more or less parallel sided in ventral view, distal shoulders broad, rounded, either straight, turned inwards or turned outwards, upper pygofer lobes ill-defined, broadly rounded, internal and lightly sclerotized, basal lobes moderately developed, exposed in lateral view, of moderate length and apically rounded or very long (at least four times longer than wide) and apically pointed, may be against pygofer margin or entirely exposed, dorsal beak well developed, acute and spine-like or absent of poorly developed with broadly rounded basal portion (
<figureCitation id="133A29C65B2EFF8CFE10F902C5785EB6" box="[490,545,1713,1736]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
F); uncus retractable, short, very broad at base tapering evenly to rounded apex or tapering to an expanded apex with large, rounded, downward turned lateral lobes (
<figureCitation id="133A29C65B2EFF8CFBE5F967C30E5E95" box="[1055,1111,1748,1771]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
G); claspers absent; aedeagus restrained by tubular encapsulation prior to ventral surface of uncus; theca apex parallel sided or almost parallel sided, sclerotized, with apical serrations on rim, distal half of theca straight or weakly curved upwards with apical portion slightly turned downwards or curved downwards in an arc but kinked near base of curve.
<emphasis id="B975E9515B2EFF8CFBA6F88FC24B5F2D" box="[1116,1298,1852,1875]" italics="true" pageId="1" pageNumber="578">Female genitalia</emphasis>
: sternite VII of female smoothly bilobed (
<figureCitation id="133A29C65B2EFF8CFE30F8EDC5585F0B" box="[458,513,1886,1909]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">
Fig
<date id="FFBF13835B2EFF8CFE0CF8EDC5585F0B" box="[502,513,1886,1909]" pageId="1" pageNumber="578">1</date>
</figureCitation>
I), ovipositor longer than dorsal beak (
<figureCitation id="133A29C65B2EFF8CFC67F8EDC48A5F0B" box="[925,979,1886,1909]" captionStart="FIGURE 1" captionStartId="2.[151,244,1956,1978]" captionTargetBox="[175,1407,194,1923]" captionTargetId="figure@2.[175,1408,194,1923]" captionTargetPageId="2" captionText="FIGURE 1. Megatibicen dealbatus n. comb.: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; B E, 5 mm; F G, 2 mm; H I, 5 mm." httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="1" pageNumber="578">Fig 1</figureCitation>
H).
</paragraph>
<paragraph id="8BBE35435B2EFF8CFF3DF832C5325FA3" blockId="1.[151,1437,152,2013]" pageId="1" pageNumber="578">
<emphasis id="B975E9515B2EFF8CFF3DF832C6E25FE6" bold="true" box="[199,443,1921,1944]" pageId="1" pageNumber="578">Measurements (mm).</emphasis>
Length of body: 32.543.0; length of fore wing: 40.858.3; width of fore wing: 12.221.5; length of head: 5.37.1; width of head including eyes: 12.118.8; width of pronotum including suprahumeral plates: 14.519.6; width of mesonotum: 11.817.1.
</paragraph>
<caption id="DF7E65CB5B2DFF8FFF6DF817C4F55F8A" httpUri="https://zenodo.org/record/262053/files/figure.png" pageId="2" pageNumber="579" targetBox="[175,1407,194,1923]" targetPageId="2">
<paragraph id="8BBE35435B2DFF8FFF6DF817C4F55F8A" blockId="2.[151,1437,1956,2036]" pageId="2" pageNumber="579">
<emphasis id="B975E9515B2DFF8FFF6DF817C6555FC4" bold="true" box="[151,268,1956,1978]" pageId="2" pageNumber="579">FIGURE 1.</emphasis>
<taxonomicName id="4C014EC05B2DFF8FFEEBF817C6B55FC7" box="[273,492,1956,1977]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="579" phylum="Arthropoda" rank="species" species="dealbatus" status="comb. nov.">
<emphasis id="B975E9515B2DFF8FFEEBF817C6B55FC7" box="[273,492,1956,1977]" italics="true" pageId="2" pageNumber="579">Megatibicen dealbatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246542A5B2DFF8FFE0BF817C5115FC7" box="[497,584,1956,1978]" pageId="2" pageNumber="579">
<emphasis id="B975E9515B2DFF8FFE0BF817C51B5FC4" bold="true" box="[497,578,1956,1978]" pageId="2" pageNumber="579">n. comb</emphasis>
.
</taxonomicNameLabel>
: A, male and female habitus; B, male dorsum; C, male timbal cover; D, male operculum. E, female operculum; F, male lateral view of genitalia; G, male posterior view of genitalia; H, female lateral view of genitalia; I, female ventral view of genitalia. Scale bar: A, 2 cm; BE, 5 mm; FG, 2 mm; HI, 5 mm.
</paragraph>
</caption>
<paragraph id="8BBE35435B2CFF8EFF3DFF2BC36F5A0F" blockId="3.[151,1436,152,1460]" pageId="3" pageNumber="580">
<emphasis id="B975E9515B2CFF8EFF3DFF2BC6BF58D1" bold="true" box="[199,486,152,175]" pageId="3" pageNumber="580">Diagnostic charateristics.</emphasis>
The characters described by
<bibRefCitation id="EF9048B25B2CFF8EFCE6FF2BC49658D1" author="Hill" box="[796,975,152,175]" etAl="et al." firstAuthor="Hill" pageId="3" pageNumber="580" pagination="219 - 251" refId="ref4325" refString="Hill, K. B. R., Marshall, D. C., Moulds, M. S. &amp; Simon, C. (2015) Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae). Zootaxa, 3985 (2), 219 - 251." type="journal article" year="2015">
Hill
<emphasis id="B975E9515B2CFF8EFCB6FF2BC4DB58D1" box="[844,898,152,175]" italics="true" pageId="3" pageNumber="580">et al.</emphasis>
(2015)
</bibRefCitation>
as unique to
<taxonomicName id="4C014EC05B2CFF8EFBA6FF2BC39658D1" box="[1116,1231,152,175]" class="Insecta" family="Cicadidae" genus="Neotibicen" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFBA6FF2BC39658D1" box="[1116,1231,152,175]" italics="true" pageId="3" pageNumber="580">Neotibicen</emphasis>
</taxonomicName>
are also diagnostic for
<taxonomicName id="4C014EC05B2CFF8EFF46FF09C61858AF" box="[188,321,186,209]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975E9515B2CFF8EFF46FF09C61858AF" box="[188,321,186,209]" italics="true" pageId="3" pageNumber="580">Megatibicen</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFEB2FF09C6CA58AF" bold="true" box="[328,403,186,209]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFEB2FF09C6CA58AF" box="[328,403,186,209]" pageId="3" pageNumber="580" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
to separate the new genus from the other
<taxonomicName id="4C014EC05B2CFF8EFCB5FF09C35058AF" box="[847,1033,186,209]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="tribe" tribe="Cryptotympanini">Cryptotympanini</taxonomicName>
: timbal covers fully rounded dorsally, extending to metathorax and tightly closing the timbal cavity; male abdominal tergites 2 and 3 much larger than tergites 4-7; bulbous male basisternum 3; aedeagal basal plat bent downward in lateral view, and aedeagus restrained by tubular encapsulation prior to the ventral surface. We can add the following to distinguish
<taxonomicName id="4C014EC05B2CFF8EFBF3FE91C3D75947" box="[1033,1166,290,313]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975E9515B2CFF8EFBF3FE91C3D75947" box="[1033,1166,290,313]" italics="true" pageId="3" pageNumber="580">Megatibicen</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFB6EFE91C3B95947" bold="true" box="[1172,1248,290,313]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFB6EFE91C3B95947" box="[1172,1248,290,313]" pageId="3" pageNumber="580" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C014EC05B2CFF8EFADBFE91C2CC5947" box="[1313,1429,290,313]" class="Insecta" family="Cicadidae" genus="Neotibicen" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFADBFE91C2CC5947" box="[1313,1429,290,313]" italics="true" pageId="3" pageNumber="580">Neotibicen</emphasis>
</taxonomicName>
: head not as wide as lateral pronotal collar male opercula terminating at rim of tympanal cavity, male sternite VIII Ushaped or V-shaped and as wide as sternite VII in cross-section, pygofer considerably broadened towards top presenting a triangular shape or more or less parallel sided in ventral view, pygofer dorsal beak absent or poorly developed with basal portion broadly rounded or well developed, acute and spine-like, pygofer distal shoulder straight, turned inwards or turned outwards, pygofer basal lobes very long and apically pointed or of moderate length and apically rounded, uncus short, very broad at base tapering evenly to rounded apex or tapering to an expanded apex with large, rounded, downward turned lateral lobes, theca apex parallel sided or almost parallel sided, sclerotized, with apical serrations on rim, distal half of theca straight or weakly curved upwards with apical portion slightly turned downwards or curved downwards in an arc but kinked near base of curve, ovipositor longer than dorsal beak.
</paragraph>
</subSubSection>
<subSubSection id="C31B66C85B2CFF8EFF3DFDCEC4FB5D79" pageId="3" pageNumber="580" type="diagnosis">
<paragraph id="8BBE35435B2CFF8EFF3DFDCEC2CD5BD7" blockId="3.[151,1436,152,1460]" pageId="3" pageNumber="580">
<emphasis id="B975E9515B2CFF8EFF3DFDCEC6655AEA" bold="true" box="[199,316,637,660]" pageId="3" pageNumber="580">Diagnosis.</emphasis>
Within the
<taxonomicName id="4C014EC05B2CFF8EFE41FDCEC52D5AEA" box="[443,628,637,660]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="tribe" tribe="Cryptotympanini">Cryptotympanini</taxonomicName>
,
<taxonomicName id="4C014EC05B2CFF8EFD85FDCEC45D5AEA" box="[639,772,637,660]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975E9515B2CFF8EFD85FDCEC45D5AEA" box="[639,772,637,660]" italics="true" pageId="3" pageNumber="580">Megatibicen</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFCF6FDCEC4015AEA" bold="true" box="[780,856,637,660]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFCF6FDCEC4015AEA" box="[780,856,637,660]" pageId="3" pageNumber="580" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
can be distinguished from the Neartic genera (
<taxonomicName id="4C014EC05B2CFF8EFAA9FDCEC6775AC9" authority="Moulds, 2015" authorityName="Moulds" authorityYear="2015" class="Insecta" family="Cicadidae" genus="Hadoa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFAA9FDCEC2C55AEA" box="[1363,1436,637,660]" italics="true" pageId="3" pageNumber="580">Hadoa</emphasis>
<bibRefCitation id="EF9048B25B2CFF8EFF6DFD13C6775AC9" author="Moulds" box="[151,302,672,695]" pageId="3" pageNumber="580" pagination="219 - 251" refId="ref4325" refString="Hill, K. B. R., Marshall, D. C., Moulds, M. S. &amp; Simon, C. (2015) Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae). Zootaxa, 3985 (2), 219 - 251." type="journal article" year="2015">Moulds, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C014EC05B2CFF8EFEC3FD13C5725AC9" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" box="[313,555,672,695]" class="Insecta" family="Cicadidae" genus="Cacama" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFEC3FD13C6CB5AC9" box="[313,402,672,695]" italics="true" pageId="3" pageNumber="580">Cacama</emphasis>
Distant, 1904
</taxonomicName>
,
<taxonomicName id="4C014EC05B2CFF8EFDCCFD13C4675AC9" authority="Davis, 1944" authorityName="Davis" authorityYear="1944" box="[566,830,672,695]" class="Insecta" family="Cicadidae" genus="Cornuplura" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFDCCFD13C5ED5AC9" box="[566,692,672,695]" italics="true" pageId="3" pageNumber="580">Cornuplura</emphasis>
Davis, 1944
</taxonomicName>
,
<taxonomicName id="4C014EC05B2CFF8EFCB3FD13C3105AC9" authority="Stal, 1870" authorityName="Stal" authorityYear="1870" box="[841,1097,672,695]" class="Insecta" family="Cicadidae" genus="Diceroprocta" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFCB3FD13C48E5AC9" box="[841,983,672,695]" italics="true" pageId="3" pageNumber="580">Diceroprocta</emphasis>
Stål, 1870
</taxonomicName>
and
<taxonomicName id="4C014EC05B2CFF8EFB87FD13C7975AA7" authority="Hill &amp; Moulds, 2015" authorityName="Hill &amp; Moulds" authorityYear="2015" class="Insecta" family="Cicadidae" genus="Neotibicen" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFB87FD13C3A95AC9" box="[1149,1264,672,695]" italics="true" pageId="3" pageNumber="580">Neotibicen</emphasis>
<bibRefCitation id="EF9048B25B2CFF8EFB02FD13C7975AA7" author="Hill &amp; Moulds" firstAuthor="Hill" pageId="3" pageNumber="580" pagination="219 - 251" refId="ref4325" refString="Hill, K. B. R., Marshall, D. C., Moulds, M. S. &amp; Simon, C. (2015) Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae). Zootaxa, 3985 (2), 219 - 251." type="journal article" year="2015">Hill &amp; Moulds, 2015</bibRefCitation>
</taxonomicName>
) of the tribe with a few characters. Species of
<taxonomicName id="4C014EC05B2CFF8EFD31FD71C44D5AA7" box="[715,788,706,729]" class="Insecta" family="Cicadidae" genus="Hadoa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFD31FD71C44D5AA7" box="[715,788,706,729]" italics="true" pageId="3" pageNumber="580">Hadoa</emphasis>
</taxonomicName>
can be distinguished by the head being about as wide as or narrower than the mesonotum and the flat basisternum 3. The head is narrower than the mesonotum and flattened anteriorly, the mesonotum overlaps the metanotum and the five apical cells in the hind wing are unique to species of
<taxonomicName id="4C014EC05B2CFF8EFF6DFC99C7A95B3F" box="[151,240,810,833]" class="Insecta" family="Cicadidae" genus="Cacama" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFF6DFC99C7A95B3F" box="[151,240,810,833]" italics="true" pageId="3" pageNumber="580">Cacama</emphasis>
</taxonomicName>
. Species of
<taxonomicName id="4C014EC05B2CFF8EFE8BFC99C6B65B3F" box="[369,495,810,833]" class="Insecta" family="Cicadidae" genus="Cornuplura" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFE8BFC99C6B65B3F" box="[369,495,810,833]" italics="true" pageId="3" pageNumber="580">Cornuplura</emphasis>
</taxonomicName>
can be distinguished by spines extending from the ventral margin of the pygofer and the hooks on sternite VIII combined with the black basal cell of the fore wing. The radial crossvein in the fore wing is approximately in the middle of apical cell 2, the bifurcated, wishbone-shaped uncus and the acute lateral lobes of the pygofer of males and the sinuate posterior margin to sternite VII of the females can distinguish species of
<taxonomicName id="4C014EC05B2CFF8EFAFBFC21C2D75BD7" box="[1281,1422,914,937]" class="Insecta" family="Cicadidae" genus="Diceroprocta" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFAFBFC21C2D75BD7" box="[1281,1422,914,937]" italics="true" pageId="3" pageNumber="580">Diceroprocta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBE35435B2CFF8EFF3DFC06C4FB5D79" blockId="3.[151,1436,152,1460]" pageId="3" pageNumber="580">
<taxonomicName id="4C014EC05B2CFF8EFF3DFC06C6635BB2" box="[199,314,949,972]" class="Insecta" family="Cicadidae" genus="Neotibicen" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFF3DFC06C6635BB2" box="[199,314,949,972]" italics="true" pageId="3" pageNumber="580">Neotibicen</emphasis>
</taxonomicName>
is most similar to the new genus as it is the source of the species for the new genus.
<taxonomicName id="4C014EC05B2CFF8EFB20FC06C2145BB2" box="[1242,1357,949,972]" class="Insecta" family="Cicadidae" genus="Neotibicen" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B975E9515B2CFF8EFB20FC06C2145BB2" box="[1242,1357,949,972]" italics="true" pageId="3" pageNumber="580">Neotibicen</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C014EC05B2CFF8EFE93FC6BC6B75B91" box="[361,494,984,1007]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975E9515B2CFF8EFE93FC6BC6B75B91" box="[361,494,984,1007]" italics="true" pageId="3" pageNumber="580">Megatibicen</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFE0CFC6BC51B5B91" bold="true" box="[502,578,984,1007]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFE0CFC6BC51B5B91" box="[502,578,984,1007]" pageId="3" pageNumber="580" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
by the following characters: head as wide or wider than lateral pronotal collar, male opercula protruding past the rim of tympanal cavity, male sternite VIII U-shaped (V-shaped in
<taxonomicName id="4C014EC05B2CFF8EFADEFC48C23E5C6D" box="[1316,1383,1019,1043]" class="Insecta" family="Cicadidae" genus="Tibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="B975E9515B2CFF8EFADEFC48C23E5C6D" box="[1316,1383,1019,1043]" italics="true" pageId="3" pageNumber="580">davisi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C014EC05B2CFF8EFF6DFBADC7A05C48" box="[151,249,1054,1078]" class="Insecta" family="Cicadidae" genus="Tibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="similaris">
<emphasis id="B975E9515B2CFF8EFF6DFBADC7A05C48" box="[151,249,1054,1078]" italics="true" pageId="3" pageNumber="580">similaris</emphasis>
</taxonomicName>
but less than width of sternite VII), pygofer more or less parallel sided in ventral view, pygofer dorsal beak well developed, acute and spine-like, pygofer distal shoulder straight or turned inwards not turned outwards, pygofer basal lobes moderately developed, exposed in lateral view, of moderate length and apically rounded, pointed or bilobed, uncus long and broad with parallel sides throughout its length, arched in cross-section, apex straight in dorsal view or short, wide at base and tapering to a distal finger-like extension subapically with a pair of large ventral spikes, theca apex flared and not sclerotized, distal half of theca evenly curved downwards in an arc possibly kinked near base of curve, ovipositor not as long as dorsal beak (except in
<taxonomicName id="4C014EC05B2CFF8EFD69FB43C4CC5D79" authority="Degeer, 1773" authorityName="Degeer" authorityYear="1773" box="[659,917,1264,1287]" class="Insecta" family="Cicadidae" genus="Neotibicen" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="lyricen">
<emphasis id="B975E9515B2CFF8EFD69FB43C5A45D79" box="[659,765,1264,1287]" italics="true" pageId="3" pageNumber="580">N. lyricen</emphasis>
Degeer, 1773
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B66C85B2CFF8EFF3DFAA1C40D5DCA" pageId="3" pageNumber="580" type="distribution">
<paragraph id="8BBE35435B2CFF8EFF3DFAA1C40D5DCA" blockId="3.[151,1436,152,1460]" pageId="3" pageNumber="580">
<emphasis id="B975E9515B2CFF8EFF3DFAA1C6005D57" bold="true" box="[199,345,1298,1321]" pageId="3" pageNumber="580">Distribution.</emphasis>
Species of the genus
<taxonomicName id="4C014EC05B2CFF8EFDB1FAA1C5895D57" box="[587,720,1298,1321]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975E9515B2CFF8EFDB1FAA1C5895D57" box="[587,720,1298,1321]" italics="true" pageId="3" pageNumber="580">Megatibicen</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFD23FAA1C47F5D57" bold="true" box="[729,806,1298,1321]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFD23FAA1C47F5D57" box="[729,806,1298,1321]" pageId="3" pageNumber="580" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
expand across the
<collectingCountry id="F31675D35B2CFF8EFC03FAA1C3705D57" box="[1017,1065,1298,1321]" name="United States of America" pageId="3" pageNumber="580">U.S.</collectingCountry>
east of the Rocky Mountains and
<collectingCountry id="F31675D35B2CFF8EFF6DFA86C7B25D32" box="[151,235,1333,1356]" name="Canada" pageId="3" pageNumber="580">Canada</collectingCountry>
. They show the greatest diversity in the central plains with
<taxonomicName id="4C014EC05B2CFF8EFC84FA86C35D5D32" box="[894,1028,1333,1356]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="figuratus" status="comb. nov.">
<emphasis id="B975E9515B2CFF8EFC84FA86C35D5D32" box="[894,1028,1333,1356]" italics="true" pageId="3" pageNumber="580">M. figuratus</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFBF4FA86C32B5D32" bold="true" box="[1038,1138,1333,1356]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFBF4FA86C32B5D32" box="[1038,1138,1333,1356]" pageId="3" pageNumber="580">n. comb.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C014EC05B2CFF8EFB50FA86C2765D32" box="[1194,1327,1333,1356]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="resonans" status="comb. nov.">
<emphasis id="B975E9515B2CFF8EFB50FA86C2765D32" box="[1194,1327,1333,1356]" italics="true" pageId="3" pageNumber="580">M. resonans</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFAC2FA86C2C55D32" bold="true" box="[1336,1436,1333,1356]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFAC2FA86C2C55D32" box="[1336,1436,1333,1356]" pageId="3" pageNumber="580">n. comb.</taxonomicNameLabel>
</emphasis>
restricted to the southern and southeastern
<collectingCountry id="F31675D35B2CFF8EFD9DFAEBC5C15D11" box="[615,664,1368,1391]" name="United States of America" pageId="3" pageNumber="580">U.S.</collectingCountry>
, and
<taxonomicName id="4C014EC05B2CFF8EFD29FAEBC41A5D11" box="[723,835,1368,1391]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="auletes" status="comb. nov.">
<emphasis id="B975E9515B2CFF8EFD29FAEBC41A5D11" box="[723,835,1368,1391]" italics="true" pageId="3" pageNumber="580">M. auletes</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFCB6FAEBC4F75D11" bold="true" box="[844,942,1368,1391]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFCB6FAEBC4F75D11" box="[844,942,1368,1391]" pageId="3" pageNumber="580">n. comb.</taxonomicNameLabel>
</emphasis>
having the most expansive distribution being found in 31
<collectingCountry id="F31675D35B2CFF8EFEE2FAC9C6115DEF" box="[280,328,1402,1425]" name="United States of America" pageId="3" pageNumber="580">U.S.</collectingCountry>
states (
<bibRefCitation id="EF9048B25B2CFF8EFE6DFAC9C5E15DEF" author="Sanborn, A. F. &amp; Phillips, P. K." box="[407,696,1402,1425]" firstAuthor="Sanborn" journalOrPublisher="Diversity" pageId="3" pageNumber="580" pagination="166 - 239" part="5" refId="ref5148" refString="Sanborn, A. F. &amp; Phillips, P. K. (2013) Biogeography of the cicadas (Hemiptera: Cicadidae) of North America, North of Mexico. Diversity, 5, 166 - 239. http: // dx. doi. org / 10.3390 / d 5020166" title="Biogeography of the cicadas (Hemiptera: Cicadidae) of North America, North of Mexico" type="journal article" year="2013">Sanborn and Phillips 2013</bibRefCitation>
).
<taxonomicName id="4C014EC05B2CFF8EFD30FAC9C48A5DEF" box="[714,979,1402,1425]" class="Insecta" family="Cicadidae" genus="Megatibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="cultriformis" status="comb. nov.">
<emphasis id="B975E9515B2CFF8EFD30FAC9C48A5DEF" box="[714,979,1402,1425]" italics="true" pageId="3" pageNumber="580">Megatibicen cultriformis</emphasis>
</taxonomicName>
<emphasis id="B975E9515B2CFF8EFC20FAC9C3655DEF" bold="true" box="[986,1084,1402,1425]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A246542A5B2CFF8EFC20FAC9C3655DEF" box="[986,1084,1402,1425]" pageId="3" pageNumber="580">n. comb.</taxonomicNameLabel>
</emphasis>
has the westernmost distribution being found in AZ, NM and
<collectingCountry id="F31675D35B2CFF8EFE33FA2EC5435DCA" box="[457,538,1437,1460]" name="Mexico" pageId="3" pageNumber="580">
<collectingRegion id="49C5FBA15B2CFF8EFE33FA2EC5435DCA" box="[457,538,1437,1460]" country="Mexico" name="Mexico" pageId="3" pageNumber="580">Mexico</collectingRegion>
</collectingCountry>
(
<bibRefCitation id="EF9048B25B2CFF8EFDD2FA2EC4115DCA" author="Sanborn, A. F. &amp; Phillips, P. K." box="[552,840,1437,1460]" firstAuthor="Sanborn" journalOrPublisher="Diversity" pageId="3" pageNumber="580" pagination="166 - 239" part="5" refId="ref5148" refString="Sanborn, A. F. &amp; Phillips, P. K. (2013) Biogeography of the cicadas (Hemiptera: Cicadidae) of North America, North of Mexico. Diversity, 5, 166 - 239. http: // dx. doi. org / 10.3390 / d 5020166" title="Biogeography of the cicadas (Hemiptera: Cicadidae) of North America, North of Mexico" type="journal article" year="2013">Sanborn and Phillips 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>