treatments-xml/data/5B/50/87/5B5087F3D82FFFD4FF617B62FB753195.xml

<document id="AD3A726E3AECC64DA78F49B9A0A97E3B" ID-DOI="10.11646/zootaxa.3110.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5246099" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629753679668" checkinUser="felipe" docAuthor="Volynkin, A. V. &amp; Matov, A. Yu." docDate="2011" docId="5B5087F3D82FFFD4FF617B62FB753195" docLanguage="en" docName="zt03110p052.pdf" docOrigin="Zootaxa 3110 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3110.1.4" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Syngrapha venustula Volynkin &amp; Matov 2011, sp.n." docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="A769FF8BD82FFFD2FFF67D43FFC73301" masterDocTitle="A new Syngrapha Hübner, [1821] from the Altai Mountain Country (Lepidoptera Noctuidae)" masterLastPageNumber="52" masterPageNumber="46" pageNumber="46" updateTime="1699106605093" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="331FE72A1AA4C036E760383D3D35177F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="626AE845AAB39FF395FC1AE19EC4B71D">
<mods:title id="CE1D0897B901D42C77B1BA34DB0DC74B">A new Syngrapha Hübner, [1821] from the Altai Mountain Country (Lepidoptera Noctuidae)</mods:title>
</mods:titleInfo>
<mods:name id="70DC4880CEFEBF1A15573DC415FF2F58" type="personal">
<mods:role id="E5F14BFA85ACA5C4EEFE6F93E22D9ED5">
<mods:roleTerm id="351CA1922499E2B5ED7F66CEA5FE00BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E34339C98E74C4376AEC5338A80B5C5">Volynkin, A. V.</mods:namePart>
<mods:affiliation id="28FB230E9ABDEEA992FEFCB995BF2957">Tigirek State Reserve, office 42, Nikitina str. 111, RF- 656043, Barnaul, Russia. E-mail: volynkin _ a @ mail. ru</mods:affiliation>
</mods:name>
<mods:name id="FC90F9D788B6329BC6E0657417E390C6" type="personal">
<mods:role id="295D58AA1A2D49169D2445BC7F9228DB">
<mods:roleTerm id="BF8A02838A813D7D63285108FF4911C8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="03326E6648A15F6942A08C272AA7FE06">Matov, A. Yu.</mods:namePart>
<mods:affiliation id="6EF6D618878DCDA3B83CF79BD7F044CC">Zoological Institute RAS, Universitetskaja emb. 1, RF- 199034, Saint-Petersburg, Russia. E-mail: noctua @ zin. ru</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7CA90EDB458CCC2C6182DB611F2A8D0D">text</mods:typeOfResource>
<mods:relatedItem id="2B275767814C37673F93116245AB3233" type="host">
<mods:titleInfo id="4F9CACA85D30D69ED7BA3648C1F8B80B">
<mods:title id="7FCD24FFE279889D0DFC3A9049401F8C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CA23480AEF0D450A310667F704E8B275">
<mods:date id="7F8A57339A11B5AF722991ACA3DAE7EB">2011</mods:date>
<mods:detail id="0A68018E049BBAECEF53CFEAF8A55890" type="pubDate">
<mods:number id="2D3A0BC01D1B86FC8827D9CD3A8E54C3">2011-11-25</mods:number>
</mods:detail>
<mods:detail id="0AA40AF355601BCDF29BCA95CD2293F2" type="volume">
<mods:number id="07F07020AC41DF7FE6D6E1B5A6F6A045">3110</mods:number>
</mods:detail>
<mods:detail id="F25AC727B556ABEDC527974A0D9715FA" type="issue">
<mods:number id="CEA43B7B7A4735B5CC4930DC61DC8A3E">1</mods:number>
</mods:detail>
<mods:extent id="61C02A6A2AA5E7ADD157DC97251EF159" unit="page">
<mods:start id="D09CC93A904A57CA6E223B4F3501A87F">46</mods:start>
<mods:end id="AC5CAB9B27B34AC0C5CF2BFA78DEDB97">52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="5F8045448CDDD40C52DF9D7FE0EEDBC7">
<mods:url id="37D6579FA6888017AD2225FFA4CA1FA0">https://biotaxa.org/Zootaxa/article/view/zootaxa.3110.1.4</mods:url>
</mods:location>
<mods:classification id="38279ADE9AF49E9DD91B20F123E0122F">journal article</mods:classification>
<mods:identifier id="7402D7F2227FB39E7DEB1FC2A7F38A53" type="DOI">10.11646/zootaxa.3110.1.4</mods:identifier>
<mods:identifier id="F71A17253DA5CD6D51C38ACD8EC4C01B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="21167D5DAD59FEB428616AD40D74DA14" type="Zenodo-Dep">5246099</mods:identifier>
</mods:mods>
<treatment id="5B5087F3D82FFFD4FF617B62FB753195" ID-DOI="http://doi.org/10.5281/zenodo.5245243" ID-Zenodo-Dep="5245243" LSID="urn:lsid:plazi:treatment:5B5087F3D82FFFD4FF617B62FB753195" httpUri="http://treatment.plazi.org/id/5B5087F3D82FFFD4FF617B62FB753195" lastPageId="6" lastPageNumber="52" pageId="0" pageNumber="46">
<subSubSection id="9BE3656ED82FFFD2FF617B62FD5B3538" box="[151,668,1569,1593]" pageId="0" pageNumber="46" type="nomenclature">
<paragraph id="D34636E5D82FFFD2FF617B62FD5B3538" blockId="0.[151,668,1569,1627]" box="[151,668,1569,1593]" pageId="0" pageNumber="46">
<heading id="880E8189D82FFFD2FF617B62FD5B3538" bold="true" box="[151,668,1569,1593]" fontSize="10" level="2" pageId="0" pageNumber="46" reason="2">
<emphasis id="E18DEAF7D82FFFD2FF617B62FD5B3538" bold="true" box="[151,668,1569,1593]" pageId="0" pageNumber="46">
<taxonomicName id="14F94D66D82FFFD2FF617B62FD9C3538" authority="Volynkin &amp; Matov" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[151,603,1569,1593]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="46" phylum="Arthropoda" rank="species" species="venustula" status="sp. nov.">
<emphasis id="E18DEAF7D82FFFD2FF617B62FE463538" bold="true" box="[151,385,1569,1593]" italics="true" pageId="0" pageNumber="46">Syngrapha venustula</emphasis>
Volynkin &amp; Matov
</taxonomicName>
, 
<taxonomicNameLabel id="FABE578CD82FFFD2FD917B62FD5B3538" box="[615,668,1569,1593]" pageId="0" pageNumber="46" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9BE3656ED82FFFD2FF617B07FE14355A" box="[151,467,1604,1627]" pageId="0" pageNumber="46" type="description">
<paragraph id="D34636E5D82FFFD2FF617B07FE14355A" blockId="0.[151,668,1569,1627]" box="[151,467,1604,1627]" pageId="0" pageNumber="46">
(
<figureCitation id="4BC22A60D82FFFD2FF697B07FEC2355A" box="[159,261,1604,1627]" captionStart="FIGURES 3–10" captionStartId="2.[151,269,1799,1823]" captionTargetBox="[160,1433,194,1775]" captionTargetId="figure-94@2.[153,1433,194,1775]" captionTargetPageId="2" captionText="FIGURES 3–10. Syngrapha spp., adults. 3, S. venustula, male, holotype, Russia, Tyva Rep., Sayansky pass; 4, S. venustula, male, paratype, Russia, Altai Rep., Katunsky Ridge; 5, S. venustula, male, paratype, E. Kazakhstan, Saur Ridge; 6, S. devergens, male, Switzerland, Valais; 7, S. rilaecacuminum, male, Bulgaria; 8, S. alticola, male, Canada, Alberta; 9, S. parilis, male, Russia, [E part of Chita region], Jablonevoj Ridge; 10, S. hochenwarthii cuprina, male, E. Kazakhstan, Saur Ridge." figureDoi="http://doi.org/10.5281/zenodo.5246103" httpUri="https://zenodo.org/record/5246103/files/figure.png" pageId="0" pageNumber="46">Figs. 3–5</figureCitation>
, 
<figureCitation id="4BC22A60D82FFFD2FEE67B07FE93355A" box="[272,340,1604,1627]" captionStart="FIGURES 11–18" captionStartId="3.[151,269,1805,1829]" captionTargetBox="[151,1435,196,1774]" captionTargetId="figure-105@3.[151,1436,194,1781]" captionTargetPageId="3" captionText="FIGURES 11–18. Syngrapha spp., male genitalia. 11, S. venustula, holotype, prep. Volynkin; 12 S. venustula, paratype, Altai Mts., Katunsky Ridge, prep. Volynkin; 13, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide 0259 Matov; 14, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 15, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 16, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 17, S. alticola, Canada, Alberta, Slide 0240 Matov; 18, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246105" httpUri="https://zenodo.org/record/5246105/files/figure.png" pageId="0" pageNumber="46">11–14</figureCitation>
, 
<figureCitation id="4BC22A60D82FFFD2FEA97B07FE62355A" box="[351,421,1604,1627]" captionStart="FIGURES 19–24" captionStartId="4.[151,269,690,714]" captionTargetBox="[151,1436,194,665]" captionTargetId="figure-661@4.[151,1436,194,666]" captionTargetPageId="4" captionText="FIGURES 19–24. Syngrapha spp., male genitalia: juxta and right harpe. 19, S. venustula, holotype, prep. Volynkin; 20, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 21, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 22, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 23, S. alticola, Canada, Alberta, Slide 0240 Matov; 24, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246107" httpUri="https://zenodo.org/record/5246107/files/figure.png" pageId="0" pageNumber="46">19–20</figureCitation>
, 
<figureCitation id="4BC22A60D82FFFD2FE597B07FE0C355A" box="[431,459,1604,1627]" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="0" pageNumber="46">27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9BE3656ED82FFFD2FF617BCAFB0E34B8" pageId="0" pageNumber="46" type="materials_examined">
<paragraph id="D34636E5D82FFFD2FF617BCAFB0E34B8" blockId="0.[151,1437,1673,1977]" pageId="0" pageNumber="46">
<emphasis id="E18DEAF7D82FFFD2FF617BCAFEB135A0" bold="true" box="[151,374,1673,1697]" inLineHeading="true" pageId="0" pageNumber="46" reason="1">Material examined.</emphasis>
<materialsCitation id="63913CB8D82FFFD2FE887BC9FB0E34B8" collectingDate="1904-06-13" collectingDateMax="1988-07-20" collectingDateMin="1904-06-13" collectionCode="SZMN, ZISP" collectorName="W. Sayan &amp; Yu. Korshunov &amp; A. Volynkin &amp; E. Katunsky Ridge &amp; O. Kosterin &amp; Tarbagatai &amp; Saur &amp; Mustau &amp; Ul'kun-Ulasty &amp; Coll. Tshetverikov &amp; Matov &amp; Uidene &amp; Volynkin" country="Russia" county="Koksu river" elevation="2300" latitude="0.70166665" location="Chagan-Obo watershed" longLatPrecision="21" longitude="0.885" municipality="Argut (Dirantai) river" pageId="0" pageNumber="46" specimenCount="67" specimenCount-female="29" specimenCount-male="38" stateProvince="Tyva" typeStatus="holotype">
<typeStatus id="0C428847D82FFFD2FE887BC9FE2635A0" box="[382,481,1674,1697]" pageId="0" pageNumber="46" type="holotype">Holotype</typeStatus>
: male, [
<collectingCountry id="ABEE7675D82FFFD2FDCF7BC9FD4235A0" box="[569,645,1674,1697]" name="Russia" pageId="0" pageNumber="46">Russia</collectingCountry>
, 
<collectingRegion id="113DF807D82FFFD2FD677BC9FCF335A0" box="[657,820,1674,1697]" country="Russia" name="Tyva" pageId="0" pageNumber="46">Tyva Republic</collectingRegion>
], 27.vii.[19]75, 
<collectorName id="7E0C5333D82FFFD2FC297BC9FB8135A0" box="[991,1094,1674,1697]" pageId="0" pageNumber="46">W. Sayan</collectorName>
[Sajlyg Ridge], Sayansky pass, [51 
<geoCoordinate id="B6CD5022D82FFFD2FF4C7BEFFEE635C7" box="[186,289,1708,1734]" degrees="0" direction="north" minutes="42" orientation="latitude" pageId="0" pageNumber="46" precision="15" seconds="6" value="0.70166665">
<superScript id="248C9BADD82FFFD2FF4C7BEFFF0635B8" attach="none" box="[186,193,1708,1721]" fontSize="5" pageId="0" pageNumber="46">0</superScript>
42’6’’ N
</geoCoordinate>
, 89 
<geoCoordinate id="B6CD5022D82FFFD2FEBA7BEFFE7535C7" box="[332,434,1708,1734]" degrees="0" direction="east" minutes="53" orientation="longitude" pageId="0" pageNumber="46" precision="15" seconds="6" value="0.885">
<superScript id="248C9BADD82FFFD2FEBA7BEFFE9435B8" attach="right" box="[332,339,1708,1721]" fontSize="5" pageId="0" pageNumber="46">0</superScript>
53’6’’ E
</geoCoordinate>
], 
<quantity id="14019B00D82FFFD2FE3E7BECFDD835C7" box="[456,543,1711,1734]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="0" pageNumber="46" unit="m" value="2300.0">
<elevation id="58D4D1D6D82FFFD2FE3E7BECFDD835C7" box="[456,543,1711,1734]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="0" pageNumber="46" unit="m" value="2300.0">2300 m</elevation>
</quantity>
, leg. 
<collectorName id="7E0C5333D82FFFD2FDA97BECFCC435C7" box="[607,771,1711,1734]" pageId="0" pageNumber="46">Yu. Korshunov</collectorName>
/ &quot; 
<taxonomicName id="14F94D66D82FFFD2FCDE7BECFAC235C7" authority="Hub., G. Zolotarenko" authorityName="Hub., G. Zolotarenko" box="[808,1285,1711,1734]" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="46" phylum="Arthropoda" rank="species" species="devergens">Caloplusia devergens Hub., G. Zolotarenko</taxonomicName>
[det.]&quot; [Coll. 
<collectionCode id="B5E8AE20D82FFFD2FF617B91FF2135E8" box="[151,230,1746,1769]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="0" pageNumber="46">SZMN</collectionCode>
], prep. 
<collectorName id="7E0C5333D82FFFD2FECE7B91FE7835E8" box="[312,447,1746,1769]" pageId="0" pageNumber="46">A. Volynkin</collectorName>
; 
<typeStatus id="0C428847D82FFFD2FE3A7B91FDF535E8" box="[460,562,1746,1769]" pageId="0" pageNumber="46" type="paratype">paratypes</typeStatus>
: 
<specimenCount id="C5FFFD6CD82FFFD2FDB47B91FD4835E8" box="[578,655,1746,1769]" pageId="0" pageNumber="46" type="male">1 male</specimenCount>
, 
<specimenCount id="C5FFFD6CD82FFFD2FD6A7B91FCCF35E8" box="[668,776,1746,1769]" pageId="0" pageNumber="46" type="female">2 females</specimenCount>
, with the same data as holotype, prep. 
<collectorName id="7E0C5333D82FFFD2FB437B91FAFE35E8" box="[1205,1337,1746,1769]" pageId="0" pageNumber="46">A. Volynkin</collectorName>
(female) [Coll. 
<collectionCode id="B5E8AE20D82FFFD2FF2B7BB7FEEA340A" box="[221,301,1780,1803]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="0" pageNumber="46">SZMN</collectionCode>
]; 
<specimenCount id="C5FFFD6CD82FFFD2FEB67BB7FE4A340A" box="[320,397,1780,1803]" pageId="0" pageNumber="46" type="male">1 male</specimenCount>
, [
<collectingCountry id="ABEE7675D82FFFD2FE567BB7FE2B340A" box="[416,492,1780,1803]" name="Russia" pageId="0" pageNumber="46">Russia</collectingCountry>
, 
<collectingRegion id="113DF807D82FFFD2FE0C7BB7FDF7340A" box="[506,560,1780,1803]" country="Russia" name="Altay" pageId="0" pageNumber="46">Altai</collectingRegion>
Rep.], 
<collectingRegion id="113DF807D82FFFD2FD747BB7FD7D340A" box="[642,698,1780,1803]" country="Russia" name="Altay" pageId="0" pageNumber="46">Altai</collectingRegion>
, 
<collectorName id="7E0C5333D82FFFD2FD317BB7FC5E340A" box="[711,921,1780,1803]" pageId="0" pageNumber="46">E. Katunsky Ridge</collectorName>
, left side of 
<collectingCounty id="3A274E69D82FFFD2FBD37BB7FB61340A" box="[1061,1190,1780,1803]" pageId="0" pageNumber="46">Koksu river</collectingCounty>
downstream, in 
<collectingMunicipality id="3322AC9FD82FFFD2FAA87BB7FEFB342F" pageId="0" pageNumber="46">Argut (Dirantai) river</collectingMunicipality>
headwater, 
<date id="A7471025D82FFFD2FE347A54FDF9342F" box="[450,574,1815,1838]" pageId="0" pageNumber="46" value="1988-07-20">
<collectingDate id="B703E9CDD82FFFD2FE347A54FDF9342F" box="[450,574,1815,1838]" pageId="0" pageNumber="46" value="1988-07-20">20.vii.1988</collectingDate>
</date>
, leg. 
<collectorName id="7E0C5333D82FFFD2FD8B7A54FCC7342F" box="[637,768,1815,1838]" pageId="0" pageNumber="46">O. Kosterin</collectorName>
, prep. 
<collectorName id="7E0C5333D82FFFD2FCBB7A54FC16342F" box="[845,977,1815,1838]" pageId="0" pageNumber="46">A. Volynkin</collectorName>
[Coll. 
<collectionCode id="B5E8AE20D82FFFD2FBD47A54FBB5342F" box="[1058,1138,1815,1838]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="0" pageNumber="46">SZMN</collectionCode>
]; 
<specimenCount id="C5FFFD6CD82FFFD2FB707A54FB14342F" box="[1158,1235,1815,1838]" pageId="0" pageNumber="46" type="male">1 male</specimenCount>
, 
<specimenCount id="C5FFFD6CD82FFFD2FB177A54FA84342F" box="[1249,1347,1815,1838]" pageId="0" pageNumber="46" type="female">1 female</specimenCount>
, [? NW 
<collectingCountry id="ABEE7675D82FFFD2FF617A79FF1B3450" box="[151,220,1850,1873]" name="China" pageId="0" pageNumber="46">China</collectingCountry>
], 
<collectorName id="7E0C5333D82FFFD2FF077A79FEA33450" box="[241,356,1850,1873]" pageId="0" pageNumber="46">Tarbagatai</collectorName>
, 
<collectorName id="7E0C5333D82FFFD2FE857A79FE613450" box="[371,422,1850,1873]" pageId="0" pageNumber="46">Saur</collectorName>
, 
<collectorName id="7E0C5333D82FFFD2FE427A79FDC33450" box="[436,516,1850,1873]" pageId="0" pageNumber="46">Mustau</collectorName>
riv., 
<collectorName id="7E0C5333D82FFFD2FDB57A79FD1D3450" box="[579,730,1850,1873]" pageId="0" pageNumber="46">Ul'kun-Ulasty</collectorName>
, near glacier, 
<date id="A7471025D82FFFD2FC8D7A79FC283450" box="[891,1007,1850,1873]" pageId="0" pageNumber="46" value="1904-06-13">
<collectingDate id="B703E9CDD82FFFD2FC8D7A79FC283450" box="[891,1007,1850,1873]" pageId="0" pageNumber="46" value="1904-06-13">13.vi.1904</collectingDate>
</date>
, 
<collectorName id="7E0C5333D82FFFD2FC087A79FB0D3450" box="[1022,1226,1850,1873]" pageId="0" pageNumber="46">Coll. Tshetverikov</collectorName>
, slide 0259 
<collectorName id="7E0C5333D82FFFD2FAA07A79FA5C3450" box="[1366,1435,1850,1873]" pageId="0" pageNumber="46">Matov</collectorName>
(male) [Coll. 
<collectionCode id="B5E8AE20D82FFFD2FEDD7A1FFEAF3472" box="[299,360,1884,1907]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="0" pageNumber="46">ZISP</collectionCode>
]; 
<specimenCount id="C5FFFD6CD82FFFD2FE8D7A1FFE153472" box="[379,466,1884,1907]" pageId="0" pageNumber="46" type="male">2 males</specimenCount>
, [E.- 
<collectingCountry id="ABEE7675D82FFFD2FDF77A1FFD4E3472" box="[513,649,1884,1907]" name="Kazakhstan" pageId="0" pageNumber="46">Kazakhstan</collectingCountry>
], 
<collectorName id="7E0C5333D82FFFD2FD6A7A1FFD2E3472" box="[668,745,1884,1907]" pageId="0" pageNumber="46">Uidene</collectorName>
and 
<location id="D626603ED82FFFD2FCD77A1FFBE63472" LSID="urn:lsid:plazi:treatment:5B5087F3D82FFFD4FF617B62FB753195:D626603ED82FFFD2FCD77A1FFBE63472" box="[801,1057,1884,1907]" country="Russia" county="Koksu river" latitude="0.70166665" longLatPrecision="21" longitude="0.885" municipality="Argut (Dirantai) river" name="Chagan-Obo watershed" pageId="0" pageNumber="46" stateProvince="Tyva">Chagan-Obo watershed</location>
, 
<collectorName id="7E0C5333D82FFFD2FBDB7A1FFB673472" box="[1069,1184,1884,1907]" pageId="0" pageNumber="46">Tarbagatai</collectorName>
, 
<collectorName id="7E0C5333D82FFFD2FB5B7A1FFB183472" box="[1197,1247,1884,1907]" pageId="0" pageNumber="46">Saur</collectorName>
, 
<date id="A7471025D82FFFD2FB1A7A1FFA9D3472" box="[1260,1370,1884,1907]" pageId="0" pageNumber="46" value="1904-07-04">
<collectingDate id="B703E9CDD82FFFD2FB1A7A1FFA9D3472" box="[1260,1370,1884,1907]" pageId="0" pageNumber="46" value="1904-07-04">4.vii.1904</collectingDate>
</date>
, 
<collectorName id="7E0C5333D82FFFD2FA917A1FFEE43497" pageId="0" pageNumber="46">Coll. Tshetverikov</collectorName>
, slide AV0004 
<collectorName id="7E0C5333D82FFFD2FE327A3CFDE23497" box="[452,549,1919,1942]" pageId="0" pageNumber="46">Volynkin</collectorName>
[Coll. 
<collectionCode id="B5E8AE20D82FFFD2FD867A3CFD6B3497" box="[624,684,1919,1942]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="0" pageNumber="46">ZISP</collectionCode>
]; 
<specimenCount id="C5FFFD6CD82FFFD2FD4B7A3CFCD83497" box="[701,799,1919,1942]" pageId="0" pageNumber="46" type="male">33 males</specimenCount>
, 
<specimenCount id="C5FFFD6CD82FFFD2FCDF7A3CFC583497" box="[809,927,1919,1942]" pageId="0" pageNumber="46" type="female">26 females</specimenCount>
, [E.- 
<collectingCountry id="ABEE7675D82FFFD2FC3A7A3CFB933497" box="[972,1108,1919,1942]" name="Kazakhstan" pageId="0" pageNumber="46">Kazakhstan</collectingCountry>
], 
<location id="D626603ED82FFFD2FB937A3CFB603497" LSID="urn:lsid:plazi:treatment:5B5087F3D82FFFD4FF617B62FB753195:D626603ED82FFFD2FB937A3CFB603497" box="[1125,1191,1919,1942]" country="Russia" county="Koksu river" latitude="0.70166665" longLatPrecision="21" longitude="0.885" municipality="Argut (Dirantai) river" name="Upper" pageId="0" pageNumber="46" stateProvince="Tyva">Upper</location>
reach of 
<location id="D626603ED82FFFD2FAFC7A3CFA523497" LSID="urn:lsid:plazi:treatment:5B5087F3D82FFFD4FF617B62FB753195:D626603ED82FFFD2FAFC7A3CFA523497" box="[1290,1429,1919,1942]" country="Russia" county="Koksu river" latitude="0.70166665" longLatPrecision="21" longitude="0.885" municipality="Argut (Dirantai) river" name="Uidene river" pageId="0" pageNumber="46" stateProvince="Tyva">Uidene river</location>
., 
<location id="D626603ED82FFFD2FF617AE1FECD34B8" LSID="urn:lsid:plazi:treatment:5B5087F3D82FFFD4FF617B62FB753195:D626603ED82FFFD2FF617AE1FECD34B8" box="[151,266,1954,1977]" country="Russia" county="Koksu river" latitude="0.70166665" longLatPrecision="21" longitude="0.885" municipality="Argut (Dirantai) river" name="Tarbagatai" pageId="0" pageNumber="46" stateProvince="Tyva">Tarbagatai</location>
, 
<location id="D626603ED82FFFD2FEE27AE1FE8134B8" LSID="urn:lsid:plazi:treatment:5B5087F3D82FFFD4FF617B62FB753195:D626603ED82FFFD2FEE27AE1FE8134B8" box="[276,326,1954,1977]" country="Russia" county="Koksu river" latitude="0.70166665" longLatPrecision="21" longitude="0.885" municipality="Argut (Dirantai) river" name="Saur" pageId="0" pageNumber="46" stateProvince="Tyva">Saur</location>
, 
<date id="A7471025D82FFFD2FEA77AE1FE7834B8" box="[337,447,1954,1977]" pageId="0" pageNumber="46" value="1904-07-03">
<collectingDate id="B703E9CDD82FFFD2FEA77AE1FE7834B8" box="[337,447,1954,1977]" pageId="0" pageNumber="46" value="1904-07-03">3.vii.1904</collectingDate>
</date>
, slide AV0005 
<location id="D626603ED82FFFD2FD957AE1FD0334B8" LSID="urn:lsid:plazi:treatment:5B5087F3D82FFFD4FF617B62FB753195:D626603ED82FFFD2FD957AE1FD0334B8" box="[611,708,1954,1977]" country="Russia" county="Koksu river" latitude="0.70166665" longLatPrecision="21" longitude="0.885" municipality="Argut (Dirantai) river" name="Volynkin" pageId="0" pageNumber="46" stateProvince="Tyva">Volynkin</location>
(female) [Coll. 
<collectionCode id="B5E8AE20D82FFFD2FC867AE1FC6A34B8" box="[880,941,1954,1977]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="0" pageNumber="46">ZISP</collectionCode>
]. 
<location id="D626603ED82FFFD2FC4B7AE1FC1834B8" LSID="urn:lsid:plazi:treatment:5B5087F3D82FFFD4FF617B62FB753195:D626603ED82FFFD2FC4B7AE1FC1834B8" box="[957,991,1954,1977]" country="Russia" county="Koksu river" latitude="0.70166665" longLatPrecision="21" longitude="0.885" municipality="Argut (Dirantai) river" name="All" pageId="0" pageNumber="46" stateProvince="Tyva">All</location>
labels are in Russian.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="8786666DD82EFFD3FF617A21FDEF349A" ID-DOI="http://doi.org/10.5281/zenodo.5246101" ID-Zenodo-Dep="5246101" httpUri="https://zenodo.org/record/5246101/files/figure.png" pageId="1" pageNumber="47" startId="1.[151,269,1890,1914]" targetBox="[151,1436,194,1866]" targetPageId="1">
<paragraph id="D34636E5D82EFFD3FF617A21FDEF349A" blockId="1.[151,1436,1890,1947]" pageId="1" pageNumber="47">
<emphasis id="E18DEAF7D82EFFD3FF617A21FE82347B" bold="true" box="[151,325,1890,1914]" pageId="1" pageNumber="47">FIGURES 1–2.</emphasis>
1, Borders of the Altai Mountain Country; 2, Distribution map of 
<taxonomicName id="14F94D66D82EFFD3FBDB7A21FB763478" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[1069,1201,1890,1913]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82EFFD3FBDB7A21FB763478" box="[1069,1201,1890,1913]" italics="true" pageId="1" pageNumber="47">S. venustula</emphasis>
</taxonomicName>
: blue star—the typelocality, blue points—other locations.
</paragraph>
</caption>
<caption id="8786666DD82DFFD0FF617A44FE5834A8" ID-DOI="http://doi.org/10.5281/zenodo.5246103" ID-Zenodo-Dep="5246103" httpUri="https://zenodo.org/record/5246103/files/figure.png" pageId="2" pageNumber="48" startId="2.[151,269,1799,1823]" targetBox="[160,1433,194,1775]" targetPageId="2">
<paragraph id="D34636E5D82DFFD0FF617A44FE5834A8" blockId="2.[151,1436,1799,1961]" pageId="2" pageNumber="48">
<emphasis id="E18DEAF7D82DFFD0FF617A44FE93341E" bold="true" box="[151,340,1799,1823]" pageId="2" pageNumber="48">FIGURES 3–10.</emphasis>
<taxonomicName id="14F94D66D82DFFD0FE967A4BFDCA341E" box="[352,525,1800,1823]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E18DEAF7D82DFFD0FE967A4BFE14341E" box="[352,467,1800,1823]" italics="true" pageId="2" pageNumber="48">Syngrapha</emphasis>
spp.
</taxonomicName>
, adults. 3, 
<taxonomicName id="14F94D66D82DFFD0FD7B7A4BFCD5341E" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[653,786,1800,1823]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82DFFD0FD7B7A4BFCD5341E" box="[653,786,1800,1823]" italics="true" pageId="2" pageNumber="48">S. venustula</emphasis>
</taxonomicName>
, male, holotype, Russia, Tyva Rep., Sayansky pass; 4, 
<taxonomicName id="14F94D66D82DFFD0FA7E7A4BFF3A3440" authorityName="Volynkin &amp; Matov" authorityYear="2011" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82DFFD0FA7E7A4BFF3A3440" italics="true" pageId="2" pageNumber="48">S. venustula</emphasis>
</taxonomicName>
, male, paratype, Russia, Altai Rep., Katunsky Ridge; 5, 
<taxonomicName id="14F94D66D82DFFD0FC817A69FC3C3440" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[887,1019,1834,1857]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82DFFD0FC817A69FC3C3440" box="[887,1019,1834,1857]" italics="true" pageId="2" pageNumber="48">S. venustula</emphasis>
</taxonomicName>
, male, paratype, E. Kazakhstan, Saur Ridge; 6, 
<taxonomicName id="14F94D66D82DFFD0FEF27A0EFE4B3465" box="[260,396,1869,1892]" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="devergens">
<emphasis id="E18DEAF7D82DFFD0FEF27A0EFE4B3465" box="[260,396,1869,1892]" italics="true" pageId="2" pageNumber="48">S. devergens</emphasis>
</taxonomicName>
, male, Switzerland, Valais; 7, 
<taxonomicName id="14F94D66D82DFFD0FD2B7A0EFC603465" authorityName="Varga &amp; Ronkay" authorityYear="1982" box="[733,935,1869,1892]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="rilaecacuminum">
<emphasis id="E18DEAF7D82DFFD0FD2B7A0EFC603465" box="[733,935,1869,1892]" italics="true" pageId="2" pageNumber="48">S. rilaecacuminum</emphasis>
</taxonomicName>
, male, Bulgaria; 8, 
<taxonomicName id="14F94D66D82DFFD0FB727A0EFB353465" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[1156,1266,1869,1892]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E18DEAF7D82DFFD0FB727A0EFB353465" box="[1156,1266,1869,1892]" italics="true" pageId="2" pageNumber="48">S. alticola</emphasis>
</taxonomicName>
, male, Canada, Alberta; 9, 
<taxonomicName id="14F94D66D82DFFD0FEE57A33FEB23486" baseAuthorityName="Hubner" baseAuthorityYear="1809" box="[275,373,1904,1927]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="E18DEAF7D82DFFD0FEE57A33FEB23486" box="[275,373,1904,1927]" italics="true" pageId="2" pageNumber="48">S. parilis</emphasis>
</taxonomicName>
, male, Russia, [E part of Chita region], Jablonevoj Ridge; 10, 
<taxonomicName id="14F94D66D82DFFD0FBD47A33FAF43486" baseAuthorityName="Warren" baseAuthorityYear="1913" box="[1058,1331,1904,1927]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="subSpecies" species="hochenwarthii" subSpecies="cuprina">
<emphasis id="E18DEAF7D82DFFD0FBD47A33FAF43486" box="[1058,1331,1904,1927]" italics="true" pageId="2" pageNumber="48">S. hochenwarthii cuprina</emphasis>
</taxonomicName>
, male, E. Kazakhstan, Saur Ridge.
</paragraph>
</caption>
<caption id="8786666DD82CFFD1FF617A4EFAB534AE" ID-DOI="http://doi.org/10.5281/zenodo.5246105" ID-Zenodo-Dep="5246105" httpUri="https://zenodo.org/record/5246105/files/figure.png" pageId="3" pageNumber="49" startId="3.[151,269,1805,1829]" targetBox="[151,1435,196,1774]" targetPageId="3">
<paragraph id="D34636E5D82CFFD1FF617A4EFAB534AE" blockId="3.[151,1436,1805,1967]" pageId="3" pageNumber="49">
<emphasis id="E18DEAF7D82CFFD1FF617A4EFE9B3424" bold="true" box="[151,348,1805,1829]" pageId="3" pageNumber="49">FIGURES 11–18.</emphasis>
<taxonomicName id="14F94D66D82CFFD1FE957A4DFDCB3425" box="[355,524,1805,1829]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E18DEAF7D82CFFD1FE957A4DFE113424" box="[355,470,1806,1829]" italics="true" pageId="3" pageNumber="49">Syngrapha</emphasis>
spp.
</taxonomicName>
, male genitalia. 11, 
<emphasis id="E18DEAF7D82CFFD1FD157A4DFCAD3424" box="[739,874,1806,1829]" italics="true" pageId="3" pageNumber="49">
<taxonomicName id="14F94D66D82CFFD1FD157A4DFCA13424" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[739,870,1806,1829]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="venustula">S. venustula</taxonomicName>
,
</emphasis>
holotype, prep. Volynkin; 12 
<taxonomicName id="14F94D66D82CFFD1FB5D7A4DFAEB3424" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[1195,1324,1806,1829]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82CFFD1FB5D7A4DFAEB3424" box="[1195,1324,1806,1829]" italics="true" pageId="3" pageNumber="49">S. venustula</emphasis>
</taxonomicName>
, paratype, Altai Mts., Katunsky Ridge, prep. Volynkin; 13, 
<taxonomicName id="14F94D66D82CFFD1FD5F7A73FCEC3446" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[681,811,1840,1863]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82CFFD1FD5F7A73FCEC3446" box="[681,811,1840,1863]" italics="true" pageId="3" pageNumber="49">S. venustula</emphasis>
</taxonomicName>
, paratype, E. Kazakhstan, Saur Ridge, slide 0259 Matov; 14, 
<taxonomicName id="14F94D66D82CFFD1FF317A10FE96346B" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[199,337,1875,1898]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82CFFD1FF317A10FE96346B" box="[199,337,1875,1898]" italics="true" pageId="3" pageNumber="49">S. venustula</emphasis>
</taxonomicName>
, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 15, 
<taxonomicName id="14F94D66D82CFFD1FB8E7A10FAC1346B" box="[1144,1286,1875,1898]" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="devergens">
<emphasis id="E18DEAF7D82CFFD1FB8E7A10FAC1346B" box="[1144,1286,1875,1898]" italics="true" pageId="3" pageNumber="49">S. devergens</emphasis>
</taxonomicName>
, Switzerland (“Helvetia”), slide 0162 Matov; 16, 
<taxonomicName id="14F94D66D82CFFD1FDC77A35FD39348C" authorityName="Varga &amp; Ronkay" authorityYear="1982" box="[561,766,1910,1933]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="rilaecacuminum">
<emphasis id="E18DEAF7D82CFFD1FDC77A35FD39348C" box="[561,766,1910,1933]" italics="true" pageId="3" pageNumber="49">S. rilaecacuminum</emphasis>
</taxonomicName>
, Bulgaria, slide AV0008 Volynkin; 17, 
<taxonomicName id="14F94D66D82CFFD1FB347A35FAF4348C" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[1218,1331,1910,1933]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E18DEAF7D82CFFD1FB347A35FAF4348C" box="[1218,1331,1910,1933]" italics="true" pageId="3" pageNumber="49">S. alticola</emphasis>
</taxonomicName>
, Canada, Alberta, Slide 0240 Matov; 18, 
<taxonomicName id="14F94D66D82CFFD1FE1F7ADBFD8D34AE" baseAuthorityName="Hubner" baseAuthorityYear="1809" box="[489,586,1944,1967]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="E18DEAF7D82CFFD1FE1F7ADBFD8D34AE" box="[489,586,1944,1967]" italics="true" pageId="3" pageNumber="49">S. parilis</emphasis>
</taxonomicName>
, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov.
</paragraph>
</caption>
<caption id="8786666DD82BFFD6FF617FF1FB5C3033" ID-DOI="http://doi.org/10.5281/zenodo.5246107" ID-Zenodo-Dep="5246107" httpUri="https://zenodo.org/record/5246107/files/figure.png" pageId="4" pageNumber="50" startId="4.[151,269,690,714]" targetBox="[151,1436,194,665]" targetPageId="4">
<paragraph id="D34636E5D82BFFD6FF617FF1FB5C3033" blockId="4.[151,1436,690,818]" pageId="4" pageNumber="50">
<emphasis id="E18DEAF7D82BFFD6FF617FF1FE9931CB" bold="true" box="[151,350,690,714]" pageId="4" pageNumber="50">FIGURES 19–24.</emphasis>
<taxonomicName id="14F94D66D82BFFD6FE937FF0FDC931CB" box="[357,526,691,714]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E18DEAF7D82BFFD6FE937FF0FE1F31CB" box="[357,472,691,714]" italics="true" pageId="4" pageNumber="50">Syngrapha</emphasis>
spp.
</taxonomicName>
, male genitalia: juxta and right harpe. 19, 
<emphasis id="E18DEAF7D82BFFD6FC257FF0FB9C31CB" box="[979,1115,691,714]" italics="true" pageId="4" pageNumber="50">
<taxonomicName id="14F94D66D82BFFD6FC257FF0FB9031CB" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[979,1111,691,714]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="venustula">S. venustula</taxonomicName>
,
</emphasis>
holotype, prep. Volynkin; 20, 
<taxonomicName id="14F94D66D82BFFD6FF617F96FEDF31ED" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[151,280,725,748]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82BFFD6FF617F96FEDF31ED" box="[151,280,725,748]" italics="true" pageId="4" pageNumber="50">S. venustula</emphasis>
</taxonomicName>
, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 21, 
<taxonomicName id="14F94D66D82BFFD6FC037F96FBBB31ED" box="[1013,1148,725,748]" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="devergens">
<emphasis id="E18DEAF7D82BFFD6FC037F96FBBB31ED" box="[1013,1148,725,748]" italics="true" pageId="4" pageNumber="50">S. devergens</emphasis>
</taxonomicName>
, Switzerland (“Helvetia”), slide 0162 Matov; 22, 
<taxonomicName id="14F94D66D82BFFD6FE717FBBFD88300E" authorityName="Varga &amp; Ronkay" authorityYear="1982" box="[391,591,760,783]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="rilaecacuminum">
<emphasis id="E18DEAF7D82BFFD6FE717FBBFD88300E" box="[391,591,760,783]" italics="true" pageId="4" pageNumber="50">S. rilaecacuminum</emphasis>
</taxonomicName>
, Bulgaria, slide AV0008 Volynkin; 23, 
<taxonomicName id="14F94D66D82BFFD6FC037FBBFBA6300E" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[1013,1121,760,783]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E18DEAF7D82BFFD6FC037FBBFBA6300E" box="[1013,1121,760,783]" italics="true" pageId="4" pageNumber="50">S. alticola</emphasis>
</taxonomicName>
, Canada, Alberta, Slide 0240 Matov; 24, 
<taxonomicName id="14F94D66D82BFFD6FEE47E58FEB43033" baseAuthorityName="Hubner" baseAuthorityYear="1809" box="[274,371,795,818]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="E18DEAF7D82BFFD6FEE47E58FEB43033" box="[274,371,795,818]" italics="true" pageId="4" pageNumber="50">S. parilis</emphasis>
</taxonomicName>
, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov.
</paragraph>
</caption>
<subSubSection id="9BE3656ED82BFFD6FF317E23FADF352D" pageId="4" pageNumber="50" type="diagnosis">
<paragraph id="D34636E5D82BFFD6FF317E23FADF352D" blockId="4.[151,1436,864,2031]" pageId="4" pageNumber="50">
<emphasis id="E18DEAF7D82BFFD6FF317E23FEFB3079" bold="true" box="[199,316,864,888]" inLineHeading="true" pageId="4" pageNumber="50" reason="1">Diagnosis.</emphasis>
The new species belongs to the 
<taxonomicName id="14F94D66D82BFFD6FD6C7E23FC083076" box="[666,975,864,887]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="devergens-parilis">
<emphasis id="E18DEAF7D82BFFD6FD6C7E23FC083076" box="[666,975,864,887]" italics="true" pageId="4" pageNumber="50">Syngrapha devergens-parilis</emphasis>
</taxonomicName>
species-complex, including four described species (
<bibRefCitation id="B7684B14D82BFFD6FF027EC0FE06309B" author="Goater, B. &amp; Ronkay, L. &amp; Fibiger, M." box="[244,449,899,922]" pageId="4" pageNumber="50" pagination="1 - 452" refId="ref2559" refString="Goater, B., Ronkay, L. &amp; Fibiger, M. (2003) Catocalinae &amp; Plusiinae. Noctuidae Europaeae, 10. Entomological Press, Soro, 1 - 452." type="book chapter" year="2003">
Goater 
<emphasis id="E18DEAF7D82BFFD6FEB17EC0FEB8309B" box="[327,383,899,922]" italics="true" pageId="4" pageNumber="50">et al.</emphasis>
2003
</bibRefCitation>
; 
<bibRefCitation id="B7684B14D82BFFD6FE3B7EC0FD61309B" author="Ronkay G. &amp; Ronkay L. &amp; Behounek, G." box="[461,678,899,922]" pageId="4" pageNumber="50" pagination="1 - 342" refId="ref2698" refString="Ronkay G., Ronkay L. &amp; Behounek, G. (2008) Plusiinae 1. A taxonomic atlas of the Eurasian and North African Noctuoidea, 1. Heterocera Press, Budapest, 1 - 342." type="book chapter" year="2008">
Ronkay 
<emphasis id="E18DEAF7D82BFFD6FDDF7EC0FDA4309B" box="[553,611,899,922]" italics="true" pageId="4" pageNumber="50">&amp; al.</emphasis>
2008
</bibRefCitation>
): 
<taxonomicName id="14F94D66D82BFFD6FD4C7EC0FBC5309B" authority="(Hubner, [1813])" baseAuthorityName="Hubner" baseAuthorityYear="1813" box="[698,1026,899,922]" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="devergens">
<emphasis id="E18DEAF7D82BFFD6FD4C7EC0FC85309B" box="[698,834,899,922]" italics="true" pageId="4" pageNumber="50">S. devergens</emphasis>
(Hübner, [1813])
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FBE57EC0FB91309B" box="[1043,1110,899,922]" captionStart="FIGURES 3–10" captionStartId="2.[151,269,1799,1823]" captionTargetBox="[160,1433,194,1775]" captionTargetId="figure-94@2.[153,1433,194,1775]" captionTargetPageId="2" captionText="FIGURES 3–10. Syngrapha spp., adults. 3, S. venustula, male, holotype, Russia, Tyva Rep., Sayansky pass; 4, S. venustula, male, paratype, Russia, Altai Rep., Katunsky Ridge; 5, S. venustula, male, paratype, E. Kazakhstan, Saur Ridge; 6, S. devergens, male, Switzerland, Valais; 7, S. rilaecacuminum, male, Bulgaria; 8, S. alticola, male, Canada, Alberta; 9, S. parilis, male, Russia, [E part of Chita region], Jablonevoj Ridge; 10, S. hochenwarthii cuprina, male, E. Kazakhstan, Saur Ridge." figureDoi="http://doi.org/10.5281/zenodo.5246103" httpUri="https://zenodo.org/record/5246103/files/figure.png" pageId="4" pageNumber="50">Fig. 6</figureCitation>
), 
<taxonomicName id="14F94D66D82BFFD6FB9B7EC0FEEB30BD" authority="Varga &amp; Ronkay, 1982" authorityName="Varga &amp; Ronkay" authorityYear="1982" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="rilaecacuminum">
<emphasis id="E18DEAF7D82BFFD6FB9B7EC0FAF0309B" box="[1133,1335,899,922]" italics="true" pageId="4" pageNumber="50">S. rilaecacuminum</emphasis>
Varga &amp; Ronkay, 1982
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FECB7EE6FEB830BD" box="[317,383,933,956]" captionStart="FIGURES 3–10" captionStartId="2.[151,269,1799,1823]" captionTargetBox="[160,1433,194,1775]" captionTargetId="figure-94@2.[153,1433,194,1775]" captionTargetPageId="2" captionText="FIGURES 3–10. Syngrapha spp., adults. 3, S. venustula, male, holotype, Russia, Tyva Rep., Sayansky pass; 4, S. venustula, male, paratype, Russia, Altai Rep., Katunsky Ridge; 5, S. venustula, male, paratype, E. Kazakhstan, Saur Ridge; 6, S. devergens, male, Switzerland, Valais; 7, S. rilaecacuminum, male, Bulgaria; 8, S. alticola, male, Canada, Alberta; 9, S. parilis, male, Russia, [E part of Chita region], Jablonevoj Ridge; 10, S. hochenwarthii cuprina, male, E. Kazakhstan, Saur Ridge." figureDoi="http://doi.org/10.5281/zenodo.5246103" httpUri="https://zenodo.org/record/5246103/files/figure.png" pageId="4" pageNumber="50">Fig. 7</figureCitation>
), 
<taxonomicName id="14F94D66D82BFFD6FE637EE6FD7930BD" authority="(Walker, [1858])" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[405,702,933,956]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E18DEAF7D82BFFD6FE637EE6FDC330BD" box="[405,516,933,956]" italics="true" pageId="4" pageNumber="50">S. alticola</emphasis>
(Walker, [1858])
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FD397EE6FCD730BD" box="[719,784,933,956]" captionStart="FIGURES 3–10" captionStartId="2.[151,269,1799,1823]" captionTargetBox="[160,1433,194,1775]" captionTargetId="figure-94@2.[153,1433,194,1775]" captionTargetPageId="2" captionText="FIGURES 3–10. Syngrapha spp., adults. 3, S. venustula, male, holotype, Russia, Tyva Rep., Sayansky pass; 4, S. venustula, male, paratype, Russia, Altai Rep., Katunsky Ridge; 5, S. venustula, male, paratype, E. Kazakhstan, Saur Ridge; 6, S. devergens, male, Switzerland, Valais; 7, S. rilaecacuminum, male, Bulgaria; 8, S. alticola, male, Canada, Alberta; 9, S. parilis, male, Russia, [E part of Chita region], Jablonevoj Ridge; 10, S. hochenwarthii cuprina, male, E. Kazakhstan, Saur Ridge." figureDoi="http://doi.org/10.5281/zenodo.5246103" httpUri="https://zenodo.org/record/5246103/files/figure.png" pageId="4" pageNumber="50">Fig. 8</figureCitation>
) and 
<taxonomicName id="14F94D66D82BFFD6FCA67EE6FB9830BD" authority="(Hubner, 1809)" baseAuthorityName="Hubner" baseAuthorityYear="1809" box="[848,1119,933,956]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="E18DEAF7D82BFFD6FCA67EE6FC7530BD" box="[848,946,933,956]" italics="true" pageId="4" pageNumber="50">S. parilis</emphasis>
(Hübner, 1809)
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FB997EE6FB7530BD" box="[1135,1202,933,956]" captionStart="FIGURES 3–10" captionStartId="2.[151,269,1799,1823]" captionTargetBox="[160,1433,194,1775]" captionTargetId="figure-94@2.[153,1433,194,1775]" captionTargetPageId="2" captionText="FIGURES 3–10. Syngrapha spp., adults. 3, S. venustula, male, holotype, Russia, Tyva Rep., Sayansky pass; 4, S. venustula, male, paratype, Russia, Altai Rep., Katunsky Ridge; 5, S. venustula, male, paratype, E. Kazakhstan, Saur Ridge; 6, S. devergens, male, Switzerland, Valais; 7, S. rilaecacuminum, male, Bulgaria; 8, S. alticola, male, Canada, Alberta; 9, S. parilis, male, Russia, [E part of Chita region], Jablonevoj Ridge; 10, S. hochenwarthii cuprina, male, E. Kazakhstan, Saur Ridge." figureDoi="http://doi.org/10.5281/zenodo.5246103" httpUri="https://zenodo.org/record/5246103/files/figure.png" pageId="4" pageNumber="50">Fig. 9</figureCitation>
). Externally it differs from other Eurasian congeners by greyish-brown ground colour of forewing, diffused forewing pattern and paler hindwing; from Nearctic 
<taxonomicName id="14F94D66D82BFFD6FE5E7EA8FDD13703" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[424,534,1003,1026]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E18DEAF7D82BFFD6FE5E7EA8FDD13703" box="[424,534,1003,1026]" italics="true" pageId="4" pageNumber="50">S. alticola</emphasis>
</taxonomicName>
by smaller size, paler forewing ground colour, more dentate subterminal line, paler hindwings with narrower black border. The male genitalia of 
<taxonomicName id="14F94D66D82BFFD6FCDC794EFC6C3725" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[810,939,1037,1060]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82BFFD6FCDC794EFC6C3725" box="[810,939,1037,1060]" italics="true" pageId="4" pageNumber="50">S. venustula</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FC4C794EFBFD3725" box="[954,1082,1037,1060]" captionStart="FIGURES 11–18" captionStartId="3.[151,269,1805,1829]" captionTargetBox="[151,1435,196,1774]" captionTargetId="figure-105@3.[151,1436,194,1781]" captionTargetPageId="3" captionText="FIGURES 11–18. Syngrapha spp., male genitalia. 11, S. venustula, holotype, prep. Volynkin; 12 S. venustula, paratype, Altai Mts., Katunsky Ridge, prep. Volynkin; 13, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide 0259 Matov; 14, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 15, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 16, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 17, S. alticola, Canada, Alberta, Slide 0240 Matov; 18, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246105" httpUri="https://zenodo.org/record/5246105/files/figure.png" pageId="4" pageNumber="50">Figs. 11–14</figureCitation>
, 
<figureCitation id="4BC22A60D82BFFD6FBB0794EFB4D3725" box="[1094,1162,1037,1060]" captionStart="FIGURES 19–24" captionStartId="4.[151,269,690,714]" captionTargetBox="[151,1436,194,665]" captionTargetId="figure-661@4.[151,1436,194,666]" captionTargetPageId="4" captionText="FIGURES 19–24. Syngrapha spp., male genitalia: juxta and right harpe. 19, S. venustula, holotype, prep. Volynkin; 20, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 21, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 22, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 23, S. alticola, Canada, Alberta, Slide 0240 Matov; 24, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246107" httpUri="https://zenodo.org/record/5246107/files/figure.png" pageId="4" pageNumber="50">19–20</figureCitation>
) differ from 
<taxonomicName id="14F94D66D82BFFD6FAE0794EFA5B3725" baseAuthorityName="Hubner" baseAuthorityYear="1813" box="[1302,1436,1037,1060]" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="devergens">
<emphasis id="E18DEAF7D82BFFD6FAE0794EFA5B3725" box="[1302,1436,1037,1060]" italics="true" pageId="4" pageNumber="50">S. devergens</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FF697973FF3F3746" box="[159,248,1072,1095]" captionStart="FIGURES 11–18" captionStartId="3.[151,269,1805,1829]" captionTargetBox="[151,1435,196,1774]" captionTargetId="figure-105@3.[151,1436,194,1781]" captionTargetPageId="3" captionText="FIGURES 11–18. Syngrapha spp., male genitalia. 11, S. venustula, holotype, prep. Volynkin; 12 S. venustula, paratype, Altai Mts., Katunsky Ridge, prep. Volynkin; 13, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide 0259 Matov; 14, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 15, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 16, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 17, S. alticola, Canada, Alberta, Slide 0240 Matov; 18, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246105" httpUri="https://zenodo.org/record/5246105/files/figure.png" pageId="4" pageNumber="50">Figs. 15</figureCitation>
, 
<figureCitation id="4BC22A60D82BFFD6FEF27973FEE73746" box="[260,288,1072,1095]" captionStart="FIGURES 19–24" captionStartId="4.[151,269,690,714]" captionTargetBox="[151,1436,194,665]" captionTargetId="figure-661@4.[151,1436,194,666]" captionTargetPageId="4" captionText="FIGURES 19–24. Syngrapha spp., male genitalia: juxta and right harpe. 19, S. venustula, holotype, prep. Volynkin; 20, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 21, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 22, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 23, S. alticola, Canada, Alberta, Slide 0240 Matov; 24, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246107" httpUri="https://zenodo.org/record/5246107/files/figure.png" pageId="4" pageNumber="50">21</figureCitation>
) by somewhat slender uncus, relatively shorter dorso-apical process of juxta, broader, apically non-arcuate and less twisted harpe, more tapered apical part of cucullus, distally broader aedeagus, broader basal and medial parts of vesica; from 
<taxonomicName id="14F94D66D82BFFD6FED77936FE2E378D" authorityName="Varga &amp; Ronkay" authorityYear="1982" box="[289,489,1141,1164]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="rilaecacuminum">
<emphasis id="E18DEAF7D82BFFD6FED77936FE2E378D" box="[289,489,1141,1164]" italics="true" pageId="4" pageNumber="50">S. rilaecacuminum</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FE017936FD97378D" box="[503,592,1141,1164]" captionStart="FIGURES 11–18" captionStartId="3.[151,269,1805,1829]" captionTargetBox="[151,1435,196,1774]" captionTargetId="figure-105@3.[151,1436,194,1781]" captionTargetPageId="3" captionText="FIGURES 11–18. Syngrapha spp., male genitalia. 11, S. venustula, holotype, prep. Volynkin; 12 S. venustula, paratype, Altai Mts., Katunsky Ridge, prep. Volynkin; 13, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide 0259 Matov; 14, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 15, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 16, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 17, S. alticola, Canada, Alberta, Slide 0240 Matov; 18, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246105" httpUri="https://zenodo.org/record/5246105/files/figure.png" pageId="4" pageNumber="50">Figs. 16</figureCitation>
, 
<figureCitation id="4BC22A60D82BFFD6FDAD7936FDB0378D" box="[603,631,1141,1164]" captionStart="FIGURES 19–24" captionStartId="4.[151,269,690,714]" captionTargetBox="[151,1436,194,665]" captionTargetId="figure-661@4.[151,1436,194,666]" captionTargetPageId="4" captionText="FIGURES 19–24. Syngrapha spp., male genitalia: juxta and right harpe. 19, S. venustula, holotype, prep. Volynkin; 20, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 21, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 22, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 23, S. alticola, Canada, Alberta, Slide 0240 Matov; 24, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246107" httpUri="https://zenodo.org/record/5246107/files/figure.png" pageId="4" pageNumber="50">22</figureCitation>
) by slender dorso-apical process of juxta, broader, apically non-arcuate and less twisted harpe, broader valva, distally broader aedeagus, broader basal and medial parts of vesica; from 
<taxonomicName id="14F94D66D82BFFD6FAD879DBFA5B37AE" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[1326,1436,1176,1199]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E18DEAF7D82BFFD6FAD879DBFA5B37AE" box="[1326,1436,1176,1199]" italics="true" pageId="4" pageNumber="50">S. alticola</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FF6979F8FF3037D3" box="[159,247,1211,1234]" captionStart="FIGURES 11–18" captionStartId="3.[151,269,1805,1829]" captionTargetBox="[151,1435,196,1774]" captionTargetId="figure-105@3.[151,1436,194,1781]" captionTargetPageId="3" captionText="FIGURES 11–18. Syngrapha spp., male genitalia. 11, S. venustula, holotype, prep. Volynkin; 12 S. venustula, paratype, Altai Mts., Katunsky Ridge, prep. Volynkin; 13, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide 0259 Matov; 14, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 15, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 16, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 17, S. alticola, Canada, Alberta, Slide 0240 Matov; 18, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246105" httpUri="https://zenodo.org/record/5246105/files/figure.png" pageId="4" pageNumber="50">Figs. 17</figureCitation>
, 
<figureCitation id="4BC22A60D82BFFD6FEF479F8FED937D3" box="[258,286,1211,1234]" captionStart="FIGURES 19–24" captionStartId="4.[151,269,690,714]" captionTargetBox="[151,1436,194,665]" captionTargetId="figure-661@4.[151,1436,194,666]" captionTargetPageId="4" captionText="FIGURES 19–24. Syngrapha spp., male genitalia: juxta and right harpe. 19, S. venustula, holotype, prep. Volynkin; 20, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 21, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 22, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 23, S. alticola, Canada, Alberta, Slide 0240 Matov; 24, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246107" httpUri="https://zenodo.org/record/5246107/files/figure.png" pageId="4" pageNumber="50">23</figureCitation>
) by longer and narrower valva, broader and shorter harpe and broader basal and medial parts of vesica; from 
<taxonomicName id="14F94D66D82BFFD6FF61799EFF3D37F5" baseAuthorityName="Hubner" baseAuthorityYear="1809" box="[151,250,1245,1268]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="E18DEAF7D82BFFD6FF61799EFF3D37F5" box="[151,250,1245,1268]" italics="true" pageId="4" pageNumber="50">S. parilis</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FEFD799EFEA037F5" box="[267,359,1245,1268]" captionStart="FIGURES 11–18" captionStartId="3.[151,269,1805,1829]" captionTargetBox="[151,1435,196,1774]" captionTargetId="figure-105@3.[151,1436,194,1781]" captionTargetPageId="3" captionText="FIGURES 11–18. Syngrapha spp., male genitalia. 11, S. venustula, holotype, prep. Volynkin; 12 S. venustula, paratype, Altai Mts., Katunsky Ridge, prep. Volynkin; 13, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide 0259 Matov; 14, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 15, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 16, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 17, S. alticola, Canada, Alberta, Slide 0240 Matov; 18, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246105" httpUri="https://zenodo.org/record/5246105/files/figure.png" pageId="4" pageNumber="50">Figs. 18</figureCitation>
, 
<figureCitation id="4BC22A60D82BFFD6FE83799EFE5637F5" box="[373,401,1245,1268]" captionStart="FIGURES 19–24" captionStartId="4.[151,269,690,714]" captionTargetBox="[151,1436,194,665]" captionTargetId="figure-661@4.[151,1436,194,666]" captionTargetPageId="4" captionText="FIGURES 19–24. Syngrapha spp., male genitalia: juxta and right harpe. 19, S. venustula, holotype, prep. Volynkin; 20, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 21, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 22, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 23, S. alticola, Canada, Alberta, Slide 0240 Matov; 24, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246107" httpUri="https://zenodo.org/record/5246107/files/figure.png" pageId="4" pageNumber="50">24</figureCitation>
) by shorter and broader valvae, narrower and longer harpe, broader basal and medial parts of vesica. The female genitalia of 
<taxonomicName id="14F94D66D82BFFD6FE077843FDB33616" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[497,628,1280,1303]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="venustula">
<emphasis id="E18DEAF7D82BFFD6FE077843FDB33616" box="[497,628,1280,1303]" italics="true" pageId="4" pageNumber="50">S. venustula</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FD737843FD123616" box="[645,725,1280,1303]" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="4" pageNumber="50">Fig. 27</figureCitation>
) differ from 
<taxonomicName id="14F94D66D82BFFD6FC907843FC293616" baseAuthorityName="Hubner" baseAuthorityYear="1813" box="[870,1006,1280,1303]" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="devergens">
<emphasis id="E18DEAF7D82BFFD6FC907843FC293616" box="[870,1006,1280,1303]" italics="true" pageId="4" pageNumber="50">S. devergens</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FC097843FB883616" box="[1023,1103,1280,1303]" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="4" pageNumber="50">Fig. 28</figureCitation>
) by horseshoe-shaped antrum, relatively shorter ductus bursae, larger and more conical appendix bursae with stronger sclerotisation; from 
<taxonomicName id="14F94D66D82BFFD6FA7E7860FE83365D" authorityName="Varga &amp; Ronkay" authorityYear="1982" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="rilaecacuminum">
<emphasis id="E18DEAF7D82BFFD6FA7E7860FE83365D" italics="true" pageId="4" pageNumber="50">S. rilaecacuminum</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FEA27806FE64365D" box="[340,419,1349,1372]" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="4" pageNumber="50">Fig. 29</figureCitation>
) by horseshoe-shaped antrum, relatively shorter ductus bursae, narrower junction with corpus bursae, somewhat larger appendix bursae with stronger sclerotisation; from 
<taxonomicName id="14F94D66D82BFFD6FC2F782BFB8F367E" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[985,1096,1384,1407]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E18DEAF7D82BFFD6FC2F782BFB8F367E" box="[985,1096,1384,1407]" italics="true" pageId="4" pageNumber="50">S. alticola</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FBAF782BFB6D367E" box="[1113,1194,1384,1407]" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="4" pageNumber="50">Fig. 30</figureCitation>
) by horseshoe-shaped antrum, shorter ductus bursae, larger appendix bursae and broader corpus bursae; from 
<taxonomicName id="14F94D66D82BFFD6FBBF78C8FB6C36A3" baseAuthorityName="Hubner" baseAuthorityYear="1809" box="[1097,1195,1419,1442]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="E18DEAF7D82BFFD6FBBF78C8FB6C36A3" box="[1097,1195,1419,1442]" italics="true" pageId="4" pageNumber="50">S. parilis</emphasis>
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FB4C78C8FACF36A3" box="[1210,1288,1419,1442]" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="4" pageNumber="50">Fig. 31</figureCitation>
) by narrower corpus bursae, smaller appendix bursae. The new species is sympatric with 
<taxonomicName id="14F94D66D82BFFD6FC2C78EEFA5B36C5" authority="(Warren, 1913)" baseAuthorityName="Warren" baseAuthorityYear="1913" box="[986,1436,1453,1476]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="hochenwarthii" subSpecies="cuprina">
<emphasis id="E18DEAF7D82BFFD6FC2C78EEFB2936C5" box="[986,1262,1453,1476]" italics="true" pageId="4" pageNumber="50">S. hochenwarthii cuprina</emphasis>
(Warren, 1913)
</taxonomicName>
(
<figureCitation id="4BC22A60D82BFFD6FF697893FF2936E6" box="[159,238,1488,1511]" captionStart="FIGURES 3–10" captionStartId="2.[151,269,1799,1823]" captionTargetBox="[160,1433,194,1775]" captionTargetId="figure-94@2.[153,1433,194,1775]" captionTargetPageId="2" captionText="FIGURES 3–10. Syngrapha spp., adults. 3, S. venustula, male, holotype, Russia, Tyva Rep., Sayansky pass; 4, S. venustula, male, paratype, Russia, Altai Rep., Katunsky Ridge; 5, S. venustula, male, paratype, E. Kazakhstan, Saur Ridge; 6, S. devergens, male, Switzerland, Valais; 7, S. rilaecacuminum, male, Bulgaria; 8, S. alticola, male, Canada, Alberta; 9, S. parilis, male, Russia, [E part of Chita region], Jablonevoj Ridge; 10, S. hochenwarthii cuprina, male, E. Kazakhstan, Saur Ridge." figureDoi="http://doi.org/10.5281/zenodo.5246103" httpUri="https://zenodo.org/record/5246103/files/figure.png" pageId="4" pageNumber="50">Fig. 10</figureCitation>
), but is smaller in size, having shorter gamma mark and more dentate subterminal line. The male genitalia (
<figureCitation id="4BC22A60D82BFFD6FA877893FF74350B" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="4" pageNumber="50">Fig. 25</figureCitation>
) differ by shorter and wider valva, triangular harpe, shorter aedeagus, and vesical shape. The female genitalia (
<figureCitation id="4BC22A60D82BFFD6FA8678B0FF74352D" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="4" pageNumber="50">Fig. 26</figureCitation>
) differ by shape of the antrum, shorter and not twisted appendix bursae, broader, sack-like corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="9BE3656ED82BFFD4FF317B7BFB80327E" lastPageId="6" lastPageNumber="52" pageId="4" pageNumber="50" type="description">
<paragraph id="D34636E5D82BFFD4FF317B7BFB80327E" blockId="4.[151,1436,864,2031]" lastBlockId="6.[151,1436,152,487]" lastPageId="6" lastPageNumber="52" pageId="4" pageNumber="50">
<emphasis id="E18DEAF7D82BFFD6FF317B7BFE953551" bold="true" box="[199,338,1592,1616]" inLineHeading="true" pageId="4" pageNumber="50" reason="1">Description.</emphasis>
External morphology of adults (
<figureCitation id="4BC22A60D82BFFD6FD397B7BFCF9354E" box="[719,830,1592,1615]" captionStart="FIGURES 3–10" captionStartId="2.[151,269,1799,1823]" captionTargetBox="[160,1433,194,1775]" captionTargetId="figure-94@2.[153,1433,194,1775]" captionTargetPageId="2" captionText="FIGURES 3–10. Syngrapha spp., adults. 3, S. venustula, male, holotype, Russia, Tyva Rep., Sayansky pass; 4, S. venustula, male, paratype, Russia, Altai Rep., Katunsky Ridge; 5, S. venustula, male, paratype, E. Kazakhstan, Saur Ridge; 6, S. devergens, male, Switzerland, Valais; 7, S. rilaecacuminum, male, Bulgaria; 8, S. alticola, male, Canada, Alberta; 9, S. parilis, male, Russia, [E part of Chita region], Jablonevoj Ridge; 10, S. hochenwarthii cuprina, male, E. Kazakhstan, Saur Ridge." figureDoi="http://doi.org/10.5281/zenodo.5246103" httpUri="https://zenodo.org/record/5246103/files/figure.png" pageId="4" pageNumber="50">Figs. 3–5</figureCitation>
). Wingspan 
<quantity id="14019B00D82BFFD6FC267B7BFB8B354E" box="[976,1100,1592,1615]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="4" pageNumber="50" unit="mm" value="24.5" valueMax="27.0" valueMin="22.0">22–27 mm</quantity>
, forewing length 
<quantity id="14019B00D82BFFD6FAED7B7BFA5E354E" box="[1307,1433,1592,1615]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="4" pageNumber="50" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
. Antennae filiform. Labial palps slightly longer than head capsule, upcurved, on 2nd segment covered with brown scales, on 3rd segment covered with dark-brown with violet scales; 3rd segment about one-third length of the 2nd. Head, thorax and abdomen dark brown. Ground colour of forewing greyish-brown, often with violet tint. Medial area between medial cell and terminal margin dark, blackish-brown. Wing pattern diffused. Subbasal line as short, indistinct pale stroke between costal and subcostal veins, curved from costa basally. Antemedial line evenly curved from dorsum to cell then sharply angled outwards to costa; indistinct in cell. Postmedial line sinuous, curved around cell. Both lines pale, darkedged. Reniform and orbicular pale-brown. Reniform medially constricted, edged with blackish-brown. Orbicular rounded, prominent. Gamma mark creamy-white, with short tail and short and hardly divergent arms. Subterminal area pale-brown. Subterminal line dark brown, strongly and irregularly dentate, shaded blackish-brown basally. Terminal line blackish-brown. Cilia brown. Hindwing yellow with narrow black border. Discal spot semilunar, smudged. Cilia yellowish-white, with thick dark band near base. Male genitalia (
<figureCitation id="4BC22A60D82BFFD6FCBC7AF6FC0E34CD" box="[842,969,1973,1996]" captionStart="FIGURES 11–18" captionStartId="3.[151,269,1805,1829]" captionTargetBox="[151,1435,196,1774]" captionTargetId="figure-105@3.[151,1436,194,1781]" captionTargetPageId="3" captionText="FIGURES 11–18. Syngrapha spp., male genitalia. 11, S. venustula, holotype, prep. Volynkin; 12 S. venustula, paratype, Altai Mts., Katunsky Ridge, prep. Volynkin; 13, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide 0259 Matov; 14, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 15, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 16, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 17, S. alticola, Canada, Alberta, Slide 0240 Matov; 18, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246105" httpUri="https://zenodo.org/record/5246105/files/figure.png" pageId="4" pageNumber="50">Figs. 11–14</figureCitation>
, 
<figureCitation id="4BC22A60D82BFFD6FC237AF6FBDC34CD" box="[981,1051,1973,1996]" captionStart="FIGURES 19–24" captionStartId="4.[151,269,690,714]" captionTargetBox="[151,1436,194,665]" captionTargetId="figure-661@4.[151,1436,194,666]" captionTargetPageId="4" captionText="FIGURES 19–24. Syngrapha spp., male genitalia: juxta and right harpe. 19, S. venustula, holotype, prep. Volynkin; 20, S. venustula, paratype, E. Kazakhstan, Saur Ridge, slide AV0004 Volynkin; 21, S. devergens, Switzerland (“Helvetia”), slide 0162 Matov; 22, S. rilaecacuminum, Bulgaria, slide AV0008 Volynkin; 23, S. alticola, Canada, Alberta, Slide 0240 Matov; 24, S. parilis, “Kola” [NW Russia, Murmansk area, Kola penninsula], slide 0258 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246107" httpUri="https://zenodo.org/record/5246107/files/figure.png" pageId="4" pageNumber="50">19–20</figureCitation>
). Uncus long, thick, distally slightly curved, apically pointed and hooked. Tegumen narrow, medium-long. Penicular lobes narrow, weak. Juxta with elliptical-rounded basal plate and strong dorso-apical process. Vinculum short, V-shaped. Valva narrow, medium-long, with mostly parallel dorsal and ventral margins medially. Cucullus triangular, with broad, evenly rounded finely tapered apex. Sacculus strongly sclerotised, short. Clavus weak, short, triangular, apically finely pointed and setose. Harpe positioned medially, medium-long, broad, strong, triangular, not reach costa. Aedeagus cylindrical, distally broader. Carina with stronger ventral plate and slender, short ventro-lateral bar. Vesica short, broadly tubular, gradually tapered distally, projected forward, distally upturned dorsally; basally with extensive dense fine spines. Proximally vesica broad, dorsally globular, with fine scobination, medially without scobination, distally densely, finely scobinate. Female genitalia (
<figureCitation id="4BC22A60D829FFD4FEF47D9EFE9433F5" box="[258,339,221,244]" captionStart="FIGURES 25–31" captionStartId="5.[151,269,1618,1642]" captionTargetBox="[151,1430,198,1591]" captionTargetId="figure-190@5.[151,1436,194,1594]" captionTargetPageId="5" captionText="FIGURES 25–31. Syngrapha spp., male and female genitalia. 25, S. hochenwarthii cuprina, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, S. hochenwarthii cuprina, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, S. venustula, female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, S. devergens, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, S. rilaecacuminum, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, S. alticola, female genitalia, after Lafontaine &amp; Poole 1991; 31. S. parilis, Lapponia, slide 0238 Matov." figureDoi="http://doi.org/10.5281/zenodo.5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="6" pageNumber="52">Fig. 27</figureCitation>
). Ovipositor short, subconical. Papillae anales quadrate, rounded, setose. Apophyses posteriores long, thin; apophyses anteriores shorter, thicker. Ostium bursae sclerotised, short and narrow, more or less quadrangular, strongly sclerotised, with lateral margins dorsally upturned. Ductus bursae short, tubular, weakly ribbed and finely verrucose, anteriorly somewhat twisted, broadened at junction with corpus bursae. Appendix bursae relatively small, rounded conical, membranous, finely sclerotised. Corpus bursae large, elliptical-saccate.
</paragraph>
</subSubSection>
<caption id="8786666DD82AFFD7FF617B11FD213417" ID-DOI="http://doi.org/10.5281/zenodo.5246109" ID-Zenodo-Dep="5246109" httpUri="https://zenodo.org/record/5246109/files/figure.png" pageId="5" pageNumber="51" startId="5.[151,269,1618,1642]" targetBox="[151,1430,198,1591]" targetPageId="5">
<paragraph id="D34636E5D82AFFD7FF617B11FD213417" blockId="5.[151,1436,1618,1815]" pageId="5" pageNumber="51">
<emphasis id="E18DEAF7D82AFFD7FF617B11FEA6356B" bold="true" box="[151,353,1618,1642]" pageId="5" pageNumber="51">FIGURES 25–31.</emphasis>
<taxonomicName id="14F94D66D82AFFD7FE9D7B11FDD03568" box="[363,535,1618,1641]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E18DEAF7D82AFFD7FE9D7B11FE193568" box="[363,478,1618,1641]" italics="true" pageId="5" pageNumber="51">Syngrapha</emphasis>
spp.
</taxonomicName>
, male and female genitalia. 25, 
<taxonomicName id="14F94D66D82AFFD7FC887B11FB543568" baseAuthorityName="Warren" baseAuthorityYear="1913" box="[894,1171,1618,1641]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="subSpecies" species="hochenwarthii" subSpecies="cuprina">
<emphasis id="E18DEAF7D82AFFD7FC887B11FB543568" box="[894,1171,1618,1641]" italics="true" pageId="5" pageNumber="51">S. hochenwarthii cuprina</emphasis>
</taxonomicName>
, male genitalia, Russia, Altai Mts., Slide AV0451 Volynkin; 26, 
<taxonomicName id="14F94D66D82AFFD7FDA77B36FCA3358D" baseAuthorityName="Warren" baseAuthorityYear="1913" box="[593,868,1653,1676]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="subSpecies" species="hochenwarthii" subSpecies="cuprina">
<emphasis id="E18DEAF7D82AFFD7FDA77B36FCA3358D" box="[593,868,1653,1676]" italics="true" pageId="5" pageNumber="51">S. hochenwarthii cuprina</emphasis>
</taxonomicName>
, female genitalia, Russia, Altai Mts., Slide AV0506 Volynkin; 27, 
<emphasis id="E18DEAF7D82AFFD7FECC7BDBFE0035AE" box="[314,455,1688,1711]" italics="true" pageId="5" pageNumber="51">
<taxonomicName id="14F94D66D82AFFD7FECC7BDBFE0335AE" authorityName="Volynkin &amp; Matov" authorityYear="2011" box="[314,452,1688,1711]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="venustula">S. venustula</taxonomicName>
,
</emphasis>
female genitalia, paratype, Russia, Tyva Rep., Sayansky pass., prep. Volynkin; 28, 
<taxonomicName id="14F94D66D82AFFD7FA7E7BDBFEC535D0" baseAuthorityName="Hubner" baseAuthorityYear="1813" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="devergens">
<emphasis id="E18DEAF7D82AFFD7FA7E7BDBFEC535D0" italics="true" pageId="5" pageNumber="51">S. devergens</emphasis>
</taxonomicName>
, female genitalia, Alps, slide RL7786 L. Ronkay (photo by L. Ronkay); 29, 
<taxonomicName id="14F94D66D82AFFD7FB827BF9FA8635D0" authorityName="Varga &amp; Ronkay" authorityYear="1982" box="[1140,1345,1722,1745]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="rilaecacuminum">
<emphasis id="E18DEAF7D82AFFD7FB827BF9FA8635D0" box="[1140,1345,1722,1745]" italics="true" pageId="5" pageNumber="51">S. rilaecacuminum</emphasis>
</taxonomicName>
, female genitalia, Balkans, slide RL7361 L. Ronkay (photo by L. Ronkay); 30, 
<taxonomicName id="14F94D66D82AFFD7FC6A7B9EFBCE35F5" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[924,1033,1757,1780]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E18DEAF7D82AFFD7FC6A7B9EFBCE35F5" box="[924,1033,1757,1780]" italics="true" pageId="5" pageNumber="51">S. alticola</emphasis>
</taxonomicName>
, female genitalia, after 
<bibRefCitation id="B7684B14D82AFFD7FAFA7B9FFED53417" author="Lafontaine, J. D. &amp; R. W. Poole" pageId="5" pageNumber="51" pagination="1 - 182" refId="ref2660" refString="Lafontaine, J. D. &amp; R. W. Poole (1991) Noctuoidea: Noctuidae (Part), Plusiinae. The moths of America north of Mexico, 25.1, 1 - 182." type="book chapter" year="1991">Lafontaine &amp; Poole 1991</bibRefCitation>
; 31. 
<taxonomicName id="14F94D66D82AFFD7FEB37A43FDDD3417" authority=", Lapponia" authorityName="Lapponia" box="[325,538,1791,1815]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="E18DEAF7D82AFFD7FEB37A43FE623416" box="[325,421,1792,1815]" italics="true" pageId="5" pageNumber="51">S. parilis</emphasis>
, Lapponia
</taxonomicName>
, slide 0238 Matov.
</paragraph>
</caption>
<subSubSection id="9BE3656ED829FFD4FF317CC9FB753195" pageId="6" pageNumber="52" type="distribution">
<paragraph id="D34636E5D829FFD4FF317CC9FE3132E6" blockId="6.[151,1436,152,487]" pageId="6" pageNumber="52">
<emphasis id="E18DEAF7D829FFD4FF317CC9FE9E32A3" bold="true" box="[199,345,394,418]" inLineHeading="true" pageId="6" pageNumber="52" reason="1">Distribution.</emphasis>
The new species is allopatric to its congeners. It is known from high altitudes of the Altai Mountain Country: Saur Ridge in E. 
<collectingCountry id="ABEE7675D829FFD4FE457CEEFDE832C5" box="[435,559,429,452]" name="Kazakhstan" pageId="6" pageNumber="52">Kazakhstan</collectingCountry>
(and probably NW 
<collectingCountry id="ABEE7675D829FFD4FCF37CEEFC8E32C5" box="[773,841,429,452]" name="China" pageId="6" pageNumber="52">China</collectingCountry>
), Katunsky and Sajlyg Ridges in the Russian part of the Altai Mountain Country (
<figureCitation id="4BC22A60D829FFD4FE517C93FE2E32E6" box="[423,489,464,487]" captionStart="FIGURES 1–2" captionStartId="1.[151,269,1890,1914]" captionTargetBox="[151,1436,194,1866]" captionTargetId="figure-41@1.[151,1436,194,1866]" captionTargetPageId="1" captionText="FIGURES 1–2. 1, Borders of the Altai Mountain Country; 2, Distribution map of S. venustula: blue star—the typelocality, blue points—other locations." figureDoi="http://doi.org/10.5281/zenodo.5246101" httpUri="https://zenodo.org/record/5246101/files/figure.png" pageId="6" pageNumber="52">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="D34636E5D829FFD4FF317F56FB753195" blockId="6.[151,1436,533,660]" pageId="6" pageNumber="52">
We thank László Ronkay (
<collectingRegion id="113DF807D829FFD4FE067F56FD90312D" box="[496,599,533,556]" country="Hungary" name="Budapest" pageId="6" pageNumber="52">Budapest</collectingRegion>
, 
<collectingCountry id="ABEE7675D829FFD4FD907F56FD02312D" box="[614,709,533,556]" name="Hungary" pageId="6" pageNumber="52">Hungary</collectingCountry>
) for valuable consultations and pictures of female genitalia of 
<taxonomicName id="14F94D66D829FFD4FA7E7F56FEC5314E" baseAuthorityName="Hubner" baseAuthorityYear="1813" class="Insecta" family="Noctuidae" genus="Caloplusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="devergens">
<emphasis id="E18DEAF7D829FFD4FA7E7F56FEC5314E" italics="true" pageId="6" pageNumber="52">S. devergens</emphasis>
</taxonomicName>
and 
<taxonomicName id="14F94D66D829FFD4FECF7F7BFDC5314E" authorityName="Varga &amp; Ronkay" authorityYear="1982" box="[313,514,568,591]" class="Insecta" family="Noctuidae" genus="Syngrapha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="rilaecacuminum">
<emphasis id="E18DEAF7D829FFD4FECF7F7BFDC5314E" box="[313,514,568,591]" italics="true" pageId="6" pageNumber="52">S. rilaecacuminum</emphasis>
</taxonomicName>
; 
<collectingRegion id="113DF807D829FFD4FDE47F7BFDB5314E" box="[530,626,568,591]" country="Russia" name="Vladimir" pageId="6" pageNumber="52">Vladimir</collectingRegion>
S. Kononenko (Vladivostok, 
<collectingCountry id="ABEE7675D829FFD4FC417F7BFBC7314E" box="[951,1024,568,591]" name="Russia" pageId="6" pageNumber="52">Russia</collectingCountry>
) and Roman V. Yakovlev (
<collectingRegion id="113DF807D829FFD4FAD17F7BFA5B314E" box="[1319,1436,568,591]" country="Russia" name="Altay" pageId="6" pageNumber="52">Altai State</collectingRegion>
University, South Siberian Botanical garden) for valuable suggestions. Volynkin was supported by a grant from RFBR Number 11-04-90769-mob_st; Matov was supported by a grant from RFBR Number 11-04-01119.
</paragraph>
</subSubSection>
</treatment>
</document>