156 lines
19 KiB
XML
156 lines
19 KiB
XML
<document id="4B78A175A175D2E07D1883B9473694E9" ID-CLB-Dataset="2794" ID-DOI="10.11646/zootaxa.5083.1.1" ID-GBIF-Dataset="8e076bdf-97f5-4fc4-8c63-6ecc98c3dc05" ID-ISSN="1175-5326" ID-Zenodo-Dep="5800630" ID-ZooBank="14FA7505-F1F8-47F3-878C-AC349B50671D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1640237905030" checkinUser="plazi" docAuthor="Angulo, Arturo" docDate="2021" docId="191E87DEFFB5FFACD58F8D50FC9EFCCC" docLanguage="en" docName="zootaxa.5083.1.1.pdf" docOrigin="Zootaxa 5083 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Rhencus macracanthus" docType="treatment" docVersion="6" lastPageNumber="10" masterDocId="E527FFA6FFBCFFA5D5188C1BFFAAFFD9" masterDocTitle="New records and range extensions to the Costa Rican freshwater fish fauna, with an updated checklist" masterLastPageNumber="72" masterPageNumber="1" pageNumber="10" updateTime="1699473083522" updateUser="ExternalLinkService">
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<mods:mods id="97D04A12EA4A6D302351E37FDA3A3D5B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EDB7C653F057B6EFA554456F4EE377D6">
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<mods:title id="8CD46680DA0A58719062CEF128F2405D">New records and range extensions to the Costa Rican freshwater fish fauna, with an updated checklist</mods:title>
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<mods:name id="394E9AE1302070D414C1F637C027D6E6" type="personal">
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<mods:namePart id="31D2863991F9C0040F92F2BCE0B2B6C7">Angulo, Arturo</mods:namePart>
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<mods:nameIdentifier id="4211F903FB158087019A4A46D0D59A7C" type="email">arturo.angulosibaja@ucr.ac.cr</mods:nameIdentifier>
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<mods:typeOfResource id="B9EEF5740602FB1314D5CF57B9E67937">text</mods:typeOfResource>
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<mods:title id="8D14B84BD5DDE73952FE183864B09B6C">Zootaxa</mods:title>
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<mods:part id="C69B8181070E4755985ABED0A0DEAD0A">
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<mods:date id="19553E720C4B2FD52222F182549DA53A">2021</mods:date>
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<mods:number id="C3F0DC0C588E70857248CE6E0BFDCBA6">2021-12-23</mods:number>
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<mods:number id="6A207AB2DC6D5832B486486C9F658A60">5083</mods:number>
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<mods:number id="8E07DB60BB5538AFA18A38EA5FF3983F">1</mods:number>
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<mods:classification id="9E4CEA2222EBC2570B963454966A3231">journal article</mods:classification>
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<mods:identifier id="1745C701191FAF5F1C1CE47FC443B625" type="CLB-Dataset">2794</mods:identifier>
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<mods:identifier id="2028DA317F345D17C38F51A04EBB4266" type="DOI">10.11646/zootaxa.5083.1.1</mods:identifier>
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<mods:identifier id="5C1909A7FDB55596BD684733B43C9C84" type="GBIF-Dataset">8e076bdf-97f5-4fc4-8c63-6ecc98c3dc05</mods:identifier>
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<mods:identifier id="6BC9544561E3F26142A624F1C6571EDF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9500FE2811F80EBC49855E8701F56C06" type="Zenodo-Dep">5800630</mods:identifier>
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<treatment id="191E87DEFFB5FFACD58F8D50FC9EFCCC" ID-DOI="http://doi.org/10.5281/zenodo.5818109" ID-GBIF-Taxon="191880251" ID-Zenodo-Dep="5818109" LSID="urn:lsid:plazi:treatment:191E87DEFFB5FFACD58F8D50FC9EFCCC" httpUri="http://treatment.plazi.org/id/191E87DEFFB5FFACD58F8D50FC9EFCCC" lastPageNumber="10" pageId="9" pageNumber="10">
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<subSubSection id="D9AD6543FFB5FFACD58F8D50FD2EFEBF" box="[151,644,331,358]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="910836C8FFB5FFACD58F8D50FD2EFEBF" blockId="9.[151,1437,330,789]" box="[151,644,331,358]" pageId="9" pageNumber="10">
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11)
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<taxonomicName id="56B74D4BFFB5FFACD5D98D50FDD5FEBF" authority="(Gunther, 1864)" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[193,639,331,358]" class="Actinopterygii" family="Haemulidae" genus="Rhencus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="macracanthus">
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<emphasis id="A3C3EADAFFB5FFACD5D98D50FE6EFEBC" box="[193,452,331,357]" italics="true" pageId="9" pageNumber="10">Rhencus macracanthus</emphasis>
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(Günther, 1864)
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</taxonomicName>
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;
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</paragraph>
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</subSubSection>
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||
<subSubSection id="D9AD6543FFB5FFACD7918D51FC8BFEBC" box="[649,801,330,357]" pageId="9" pageNumber="10" type="multiple">
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<paragraph id="910836C8FFB5FFACD7918D51FC8BFEBC" blockId="9.[151,1437,330,789]" box="[649,801,330,357]" pageId="9" pageNumber="10">
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<taxonomicName id="56B74D4BFFB5FFACD7918D51FCB0FEBD" authorityName="Gill" authorityYear="1885" box="[649,794,330,356]" class="Actinopterygii" family="Haemulidae" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="family">
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<emphasis id="A3C3EADAFFB5FFACD7918D51FCB0FEBD" bold="true" box="[649,794,330,356]" pageId="9" pageNumber="10">Haemulidae</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="D9AD6543FFB5FFACD63E8D50FE63FE50" pageId="9" pageNumber="10" type="vernacular_names">
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<paragraph id="910836C8FFB5FFACD63E8D50FE63FE50" blockId="9.[151,1437,330,789]" pageId="9" pageNumber="10">
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Longspine Grunt; Corocoro Coche, Ronco Espina Larga, Roncacho Gordo (
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<figureCitation id="098C2A4DFFB5FFACD47D8D74FE16FE50" box="[357,444,367,393]" captionStart="FIG" captionStartId="11.[151,198,1287,1311]" captionTargetBox="[408,1180,179,1262]" captionTargetId="figure-15@11.[408,1180,179,1262]" captionTargetPageId="11" captionText="FIG. 3. New country records. (A) Mugil incilis, specimen captured in the Río Pacuare, near the mouth of the river, Siquirres, Limón (10.219424, -83.284898), at 8 m.a.s.l., on July 22, 2004, by Jorge Picado (UCR 2835-005). (B) Oligoplites saliens, specimen captured in the Caño Palma, near the Caño Palma Biological Station, Pococí, Limón (10.5938, -83.52731), at 11 m.a.s.l., on February 22, 2019, by Nathan Lovejoy et al. (UCR 3311-001). (C) Oreochromis aureus, specimen captured the Río Pacuar (Térraba river basin), 4 km SE of San Isidro del General, under bridge on road San Isidro-Dominical, San José (09.35264, -83.72790), at 619 m.a.s.l., on March 25, 2013 by Carlos Garita (UCR 3018-001). (D) Pangasianodon hypophthalmus, specimen captured in the laguna San Sebastian, Caño Negro, Alajuela (10.8630, -84.7807), at 41 m.a.s.l., on September 29, 2021, by Bryan Castro and Jader Urbina. (E) Rhencus macracanthus, specimen captured in the Río Lagarto, Chomes, Puntarenas (10.08320, -84.92064), at 15 m.a.s.l., on September 1, 1979, by William Bussing et al (UCR 1268-025). (F) Thalassoma bifasciatum, specimen captured in the Quebrada Cocles (Matina river basin), 2.5 km SE of Puerto Viejo, near the bridge on El Tucán-Cocles road, Limón (9.64221, -82.73588), at 22 m.a.s.l., on August 20, 2020, by Walter Carranza. (G) Trinectes fluviatilis, specimen captured in the Quebrada Cocal Amarillo, 5 Km N of Pavones, on the Puerto Pilón-Pavones road, Puntarenas (8.43643, - 83.09660), at 26 m.a.s.l., on March 30, 2021, by Arturo Angulo et al. (H) Trinectes xanthurus, specimen captured in the Río Tempisque (Tempisque river basin), at Puerto Humo, Guanacaste (10.31653, -85.35365), at 9 m.a.s.l., on January 25, 1976, by William Bussing et al. (UCR 0970-005). Range extensions. (I) Achirus declivis, specimen captured in the Río Moín, at the Puerto Fluvial JAPDEVA, Limón (09.99745, -83.08077), at 7 m.a.s.l., on December 14, 2006, by Karina Rodríguez (UCR 2761-003). (J) Amatitlania myrnae, specimen captured in the Río Estrella, Valle La Estrella, Limón (9.77552, -82.93240), at 13 m.a.s.l., on April 6, 2014, by Arturo Angulo. (K) Amphilophus citrinellus, specimen captured in the Río Bebedero, near Paso Real, Cañas, Guanacaste (10.43318, -85.15697), at 32 m.a.s.l., on August 7, 2021, by Luis Bolivar Montero-Chacón. (L) Brachyrhaphis olomina, specimens captured in an unnamed pound in the Reserva Biológica Monteverde, Altos de San Luis, Monteverde, Puntarenas (10.31667, -84.80932), at 1486 m.a.s.l., on July 6, 2019, by Arturo Angulo." figureDoi="http://doi.org/10.5281/zenodo.5800638" httpUri="https://zenodo.org/record/5800638/files/figure.png" pageId="9" pageNumber="10">Fig. 3E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="D9AD6543FFB5FFACD5DF8D89FDA9FD70" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="910836C8FFB5FFACD5DF8D89FDA9FD70" blockId="9.[151,1437,330,789]" pageId="9" pageNumber="10">
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<emphasis id="A3C3EADAFFB5FFACD5DF8D89FEEEFE75" bold="true" box="[199,324,402,428]" pageId="9" pageNumber="10">Diagnosis:</emphasis>
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This species differs from the other species of the family occurring in the freshwaters of the country (Pacific) by the following combination of characters: pectoral fins very long, exceeding the end of the pelvic fins and reaching the level of the anus [vs. pectoral fins short, not reaching the end of the pelvic fins in
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<taxonomicName id="56B74D4BFFB5FFACD0058DC0FE00FDC0" authority="Bussing, 1993" authorityName="Bussing" authorityYear="1993" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="empherus">
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<emphasis id="A3C3EADAFFB5FFACD0058DC0FEAEFDC0" italics="true" pageId="9" pageNumber="10">Pomadasys empherus</emphasis>
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Bussing, 1993
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</taxonomicName>
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and
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<taxonomicName id="56B74D4BFFB5FFACD4C68DE4FD6BFDC0" baseAuthorityName="Jordan & Evermann" baseAuthorityYear="1898" box="[478,705,511,537]" class="Actinopterygii" family="Haemulidae" genus="Rhonciscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bayanus">
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<emphasis id="A3C3EADAFFB5FFACD4C68DE4FD6BFDC0" box="[478,705,511,537]" italics="true" pageId="9" pageNumber="10">Rhonciscus bayanus</emphasis>
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</taxonomicName>
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||
(
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<collectingCountry id="E9A07658FFB5FFACD7D78DE4FCB6FDC0" box="[719,796,511,537]" name="Jordan" pageId="9" pageNumber="10">Jordan</collectingCountry>
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& Evermann, 1898)]; lateral line scales 41–46 [vs.
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||
<quantity id="564F9B2DFFB5FFACD04E8DE5FF07FDE4" metricMagnitude="0" metricUnit="m" metricValue="1.3208" metricValueMax="1.3716" metricValueMin="1.27" pageId="9" pageNumber="10" unit="in" value="52.0" valueMax="54.0" valueMin="50.0">50–54 in</quantity>
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||
<taxonomicName id="56B74D4BFFB5FFACD5AE8E38FDD0FDE7" authority="(Gunther, 1864)" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[182,634,547,574]" class="Actinopterygii" family="Haemulidae" genus="Haemulopsis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="leuciscus">
|
||
<emphasis id="A3C3EADAFFB5FFACD5AE8E38FE13FDE4" box="[182,441,547,573]" italics="true" pageId="9" pageNumber="10">Haemulopsis leuciscus</emphasis>
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||
(Günther, 1864)
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||
</taxonomicName>
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||
,
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||
<quantity id="564F9B2DFFB5FFACD7908E39FD44FDE4" box="[648,750,546,573]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" metricValueMax="1.2953999999999999" metricValueMin="1.1938" pageId="9" pageNumber="10" unit="in" value="49.0" valueMax="51.0" valueMin="47.0">47–51 in</quantity>
|
||
<taxonomicName id="56B74D4BFFB5FFACD7EF8E38FB70FDE4" authority="(Steindachner, 1879)" baseAuthorityName="Steindachner" baseAuthorityYear="1879" box="[759,1242,547,573]" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="branickii">
|
||
<emphasis id="A3C3EADAFFB5FFACD7EF8E38FC4FFDE4" box="[759,997,547,573]" italics="true" pageId="9" pageNumber="10">Pomadasys branickii</emphasis>
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(Steindachner, 1879)
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||
</taxonomicName>
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||
and
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||
<quantity id="564F9B2DFFB5FFACD00E8E39FAD1FDE4" box="[1302,1403,546,573]" metricMagnitude="0" metricUnit="m" metricValue="1.2827" metricValueMax="1.2953999999999999" metricValueMin="1.27" pageId="9" pageNumber="10" unit="in" value="50.5" valueMax="51.0" valueMin="50.0">50–51 in</quantity>
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||
<taxonomicName id="56B74D4BFFB5FFACD09C8E38FF5EFDB8" baseAuthorityName="Jordan & Evermann" baseAuthorityYear="1898" class="Actinopterygii" family="Haemulidae" genus="Rhonciscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bayanus">
|
||
<emphasis id="A3C3EADAFFB5FFACD09C8E38FA3FFDE5" box="[1412,1429,547,572]" italics="true" pageId="9" pageNumber="10">R</emphasis>
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||
.
|
||
<emphasis id="A3C3EADAFFB5FFACD58F8E5CFF5EFDB8" box="[151,244,583,609]" italics="true" pageId="9" pageNumber="10">bayanus</emphasis>
|
||
</taxonomicName>
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||
); dorsal fin rays 13 (vs.
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||
<quantity id="564F9B2DFFB5FFACD70A8E5CFDE6FDB8" box="[530,588,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="9" pageNumber="10" unit="in" value="14.0">14 in</quantity>
|
||
<taxonomicName id="56B74D4BFFB5FFACD74D8E5CFD4AFDB8" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[597,736,583,609]" class="Actinopterygii" family="Haemulidae" genus="Haemulopsis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="leuciscus">
|
||
<emphasis id="A3C3EADAFFB5FFACD74D8E5CFD4AFDB8" box="[597,736,583,609]" italics="true" pageId="9" pageNumber="10">H. leuciscus</emphasis>
|
||
</taxonomicName>
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||
and
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||
<quantity id="564F9B2DFFB5FFACD6018E5CFCD7FDB8" box="[793,893,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="9" pageNumber="10" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">11–12 in</quantity>
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||
<emphasis id="A3C3EADAFFB5FFACD69E8E5CFB08FDB8" box="[902,1186,583,609]" italics="true" pageId="9" pageNumber="10">
|
||
<taxonomicName id="56B74D4BFFB5FFACD69E8E5CFBA1FDB8" baseAuthorityName="Steindachner" baseAuthorityYear="1879" box="[902,1035,583,609]" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="branickii">P. branickii</taxonomicName>
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||
,
|
||
<taxonomicName id="56B74D4BFFB5FFACD1008E5CFB08FDB8" authorityName="Bussing" authorityYear="1993" box="[1048,1186,583,609]" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="empherus">P. empherus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="56B74D4BFFB5FFACD1C48E5CFAF0FDB8" baseAuthorityName="Jordan & Evermann" baseAuthorityYear="1898" box="[1244,1370,583,609]" class="Actinopterygii" family="Haemulidae" genus="Rhonciscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bayanus">
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||
<emphasis id="A3C3EADAFFB5FFACD1C48E5CFB47FDB9" box="[1244,1261,583,608]" italics="true" pageId="9" pageNumber="10">R</emphasis>
|
||
.
|
||
<emphasis id="A3C3EADAFFB5FFACD1E58E5CFAF0FDB8" box="[1277,1370,583,609]" italics="true" pageId="9" pageNumber="10">bayanus</emphasis>
|
||
</taxonomicName>
|
||
); and second anal spine longer and thicker than the third anal spine (vs. second anal spine shorter and thinner than the third anal spine in
|
||
<taxonomicName id="56B74D4BFFB5FFACD4728E94FE59FD70" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[362,499,655,681]" class="Actinopterygii" family="Haemulidae" genus="Haemulopsis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="leuciscus">
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<emphasis id="A3C3EADAFFB5FFACD4728E94FE59FD70" box="[362,499,655,681]" italics="true" pageId="9" pageNumber="10">H. leuciscus</emphasis>
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||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="D9AD6543FFB5FFACD7138E95FA37FD14" pageId="9" pageNumber="10" type="materials_examined">
|
||
<paragraph id="910836C8FFB5FFACD7138E95FA37FD14" blockId="9.[151,1437,330,789]" pageId="9" pageNumber="10">
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<emphasis id="A3C3EADAFFB5FFACD7138E95FD5EFD70" bold="true" box="[523,756,654,681]" pageId="9" pageNumber="10">Basis of the record:</emphasis>
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||
Three specimens were captured in the río Lagarto, Chomes, Puntarenas (
|
||
<geoCoordinate id="F483500FFFB5FFACD4388EA8FE27FD14" box="[288,397,691,717]" degrees="10.08320" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="1" value="10.0832">10.08320</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F483500FFFB5FFACD48F8EA8FDA4FD17" box="[407,526,691,718]" degrees="84.92064" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="1" value="-84.92064">-84.92064</geoCoordinate>
|
||
), at
|
||
<specimenCount id="87B1FD41FFB5FFACD7228EA9FDD7FD14" box="[570,637,690,717]" pageId="9" pageNumber="10" type="generic">
|
||
<quantity id="564F9B2DFFB5FFACD7228EA9FDD3FD14" box="[570,633,690,717]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="m" value="15.0">15 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
a.s.l., on
|
||
<date id="E5091008FFB5FFACD7C38EA8FC05FD14" box="[731,943,691,717]" pageId="9" pageNumber="10" value="1979-09-01">September 1, 1979</date>
|
||
, by William Bussing
|
||
<emphasis id="A3C3EADAFFB5FFACD1838EA8FB78FD14" box="[1179,1234,691,717]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
(UCR 1268-025).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AD6543FFB5FFACD58F8ECCFC9EFCCC" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="910836C8FFB5FFACD58F8ECCFC9EFCCC" blockId="9.[151,1437,330,789]" pageId="9" pageNumber="10">
|
||
<emphasis id="A3C3EADAFFB5FFACD58F8ECCFEA7FD28" bold="true" box="[151,269,727,753]" pageId="9" pageNumber="10">Remarks:</emphasis>
|
||
This species inhabits estuarine areas and coastal environs; it could be considered as an opportunistic visitor of freshwater ecosystems (
|
||
<bibRefCitation id="F5264B39FFB5FFACD7178EE0FC83FCCC" author="Robertson, D. R. & Allen, G. R." box="[527,809,762,789]" pageId="9" pageNumber="10" refId="ref66239" refString="Robertson, D. R. & Allen, G. R. (2015) Shorefishes of the Tropical Eastern Pacific: online information system. Version 2.0. Smithsonian Tropical Research Institute, Balboa, Panama. Available from: https: // biogeodb. stri. si. edu / sftep / en / pages (accessed 27 August 2021)." type="url" year="2015">Robertson & Allen 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |