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<document id="8E7765745AFB9F58821D99225CB30AD0" ID-CLB-Dataset="293563" ID-DOI="10.26515/rzsi/v122/i1/2022/154229" ID-GBIF-Dataset="59dd71f0-051d-46df-9099-2a8398d5c7fc" ID-ISSN="2581-8686" ID-Zenodo-Dep="10913849" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1712171461372" checkinUser="felipe" docAuthor="Ganesh, S. R. &amp; Channakeshava Murthy, B. H." docDate="2022" docId="E9598B66FFCFFFCEFF48C8B7FAE5FD32" docLanguage="en" docName="RecZoolSurvIndia.122.1.25-35.pdf" docOrigin="Records of the Zoological Survey of India 122 (1)" docStyle="DocumentStyle:30C78A88DFAFB68AE5A0604D5A6B2FFB.2:RecZoolSurvIndia.2017-.journal_article" docStyleId="30C78A88DFAFB68AE5A0604D5A6B2FFB" docStyleName="RecZoolSurvIndia.2017-.journal_article" docStyleVersion="2" docTitle="Teretrurus sanguineus" docType="treatment" docVersion="4" lastPageNumber="29" masterDocId="1560F31EFFCCFFCAFFE8CC38FFE6FFF1" masterDocTitle="Taxonomy of the shieldtail snake genus Teretrurus Beddome, 1886 (Serpentes: Uropeltidae) with a revised key and remarks on the geographic gaps in the Western Ghats, Peninsular India" masterLastPageNumber="35" masterPageNumber="25" pageNumber="28" updateTime="1712947598891" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7722F5D50AAD2320489D851ECF103B7F">Taxonomy of the shieldtail snake genus Teretrurus Beddome, 1886 (Serpentes: Uropeltidae) with a revised key and remarks on the geographic gaps in the Western Ghats, Peninsular India</mods:title>
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<mods:namePart id="04B90D8D77B1B0B4A4C17204F4C3A9F3">Ganesh, S. R.</mods:namePart>
<mods:affiliation id="BD28A2B37F980886A8314A962076B337">Chennai Snake Park, Rajbhavan Post, Chennai - 600022, Tamil Nadu, India; Email: snakeranglerr @ gmail. com</mods:affiliation>
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<mods:namePart id="2929F0BC662C40AF7418F48B81BCE307">Channakeshava Murthy, B. H.</mods:namePart>
<mods:affiliation id="F200F2C53217ED34FADC23088A6D5F48">Zoological Survey of India, Western Ghats Regional Station, Kozhikode - 673006, Kerala, India; Email: puttakeshava @ gmail. com</mods:affiliation>
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<mods:title id="EBDA45D9861E41EFBC79C6A9830B609B">Records of the Zoological Survey of India</mods:title>
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<mods:date id="877A2D8DF4789AE606988718F703FCFB">2022</mods:date>
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<treatment id="E9598B66FFCFFFCEFF48C8B7FAE5FD32" ID-DOI="http://doi.org/10.5281/zenodo.10966819" ID-Zenodo-Dep="10966819" LSID="urn:lsid:plazi:treatment:E9598B66FFCFFFCEFF48C8B7FAE5FD32" httpUri="http://treatment.plazi.org/id/E9598B66FFCFFFCEFF48C8B7FAE5FD32" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
<subSubSection id="29EA69FBFFCFFFC9FF48C8B7FD81FB5B" box="[160,615,1167,1194]" pageId="3" pageNumber="28" type="nomenclature">
<paragraph id="614F3A70FFCFFFC9FF48C8B7FD81FB5B" blockId="3.[160,615,1167,1229]" box="[160,615,1167,1194]" pageId="3" pageNumber="28">
<heading id="3A078D1CFFCFFFC9FF48C8B7FD81FB5B" box="[160,615,1167,1194]" fontSize="11" level="2" pageId="3" pageNumber="28" reason="4">
<taxonomicName id="A6F041F3FFCFFFC9FF48C8B7FD81FB5B" ID-CoL="55FFY" authority="(Beddome, 1867)" baseAuthorityName="Beddome" baseAuthorityYear="1867" box="[160,615,1167,1194]" class="Squamata" family="Uropeltidae" genus="Teretrurus" kingdom="Animalia" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="5384E662FFCFFFC9FF48C8B7FE79FB58" bold="true" box="[160,415,1167,1193]" italics="true" pageId="3" pageNumber="28">Teretrurus sanguineus</emphasis>
(
<bibRefCitation id="05614781FFCFFFC9FE45C8B7FDB9FB5B" author="Beddome, R. H." box="[429,607,1167,1194]" pageId="3" pageNumber="28" pagination="14 - 16" refId="ref6894" refString="Beddome, R. H. 1867. Descriptions and figures of five new snakes from the Madras Presidency, Madras Quarterly Journal of Medical Sciences, 11: 14 - 16." type="journal article" year="1867">Beddome, 1867</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="29EA69FBFFCFFFC9FF48C88BFEA2FB3C" box="[160,324,1203,1229]" pageId="3" pageNumber="28" type="description">
<paragraph id="614F3A70FFCFFFC9FF48C88BFEA2FB3C" blockId="3.[160,615,1167,1229]" box="[160,324,1203,1229]" pageId="3" pageNumber="28">
(
<figureCitation id="F9CB26F5FFCFFFC9FF4FC88BFEE3FB3C" box="[167,261,1203,1229]" captionStart="Figure 1" captionStartId="1.[160,236,1779,1805]" captionTargetBox="[160,797,1225,1747]" captionTargetId="figure-568@1.[160,797,1225,1747]" captionTargetPageId="1" captionText="Figure 1. Map of southern India showing the distribution of Teretrurus species, in the Western Ghats." figureDoi="http://doi.org/10.5281/zenodo.10913853" httpUri="https://zenodo.org/record/10913853/files/figure.png" pageId="3" pageNumber="28">Figure 1</figureCitation>
,
<figureCitation id="F9CB26F5FFCFFFC9FEFAC88BFEC7FB3C" box="[274,289,1203,1229]" captionStart="Figure 2" captionStartId="3.[161,237,1749,1775]" captionTargetBox="[160,797,1306,1727]" captionTargetId="figure-231@3.[160,797,1306,1727]" captionTargetPageId="3" captionText="Figure 2. Preserved specimens of Teretrurus species: (a) T. travancoricus ZSI/WGRC/2135-6, (b) T. hewstoni ZSI/WGRC/3124, (c) T. sanguineus CSPT/S-10, (d) T. rhodogaster CSPT/S-14. Photos: S. R. Ganesh." figureDoi="http://doi.org/10.5281/zenodo.10913855" httpUri="https://zenodo.org/record/10913855/files/figure.png" pageId="3" pageNumber="28">2</figureCitation>
,
<figureCitation id="F9CB26F5FFCFFFC9FEC5C88BFEDDFB3C" box="[301,315,1203,1229]" captionStart="Figure 3" captionStartId="4.[836,915,1729,1755]" captionTargetBox="[835,1470,1157,1701]" captionTargetId="figure-590@4.[835,1470,1157,1701]" captionTargetPageId="4" captionText="Figure 3. Holotype of Plecturus scabricauda ZSI 6996 coll. from Anamallays: (a) entire view, (b) jar label, (c) head right view, (d) head left view, (e) head ventral view, (f) tail ventral view. Photos: N.S. Achyuthan." figureDoi="http://doi.org/10.5281/zenodo.10913859" httpUri="https://zenodo.org/record/10913859/files/figure.png" pageId="3" pageNumber="28">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="29EA69FBFFCFFFC9FF48C8E7FC7CFEF6" pageId="3" pageNumber="28" type="reference_group">
<paragraph id="614F3A70FFCFFFC9FF48C8E7FC7CFEF6" blockId="3.[160,789,1247,1269]" lastBlockId="3.[861,1499,214,263]" pageId="3" pageNumber="28">
1867.
<taxonomicName id="A6F041F3FFCFFFC9FF3CC8E7FD3EFB04" authority="Beddome, Madras Quart. J.Med. Sci." box="[212,728,1247,1269]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="5384E662FFCFFFC9FF3CC8E7FE69FB04" box="[212,399,1247,1269]" italics="true" pageId="3" pageNumber="28">Plectrurus sanguineus</emphasis>
Beddome,
<emphasis id="5384E662FFCFFFC9FE1AC8E7FD3EFB04" box="[498,728,1247,1269]" italics="true" pageId="3" pageNumber="28">Madras Quart. J.Med. Sci.</emphasis>
</taxonomicName>
<emphasis id="5384E662FFCFFFC9FD3FC8D8FD3DFB04" box="[727,731,1248,1269]" italics="true" pageId="3" pageNumber="28">,</emphasis>
<emphasis id="5384E662FFCFFFC9FD32C8E7FD16FB05" bold="true" box="[730,752,1247,1268]" pageId="3" pageNumber="28">11</emphasis>
: 14. 1876.
<taxonomicName id="A6F041F3FFCFFFC9FC7DCCEDFA33FF1A" authority="Theobald, Descr. Catl. Reptiles Brit. Ind." box="[917,1493,213,235]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="scabricauda">
<emphasis id="5384E662FFCFFFC9FC7DCCEDFB87FF1B" box="[917,1121,213,234]" italics="true" pageId="3" pageNumber="28">Plectrurus scabricauda</emphasis>
Theobald,
<emphasis id="5384E662FFCFFFC9FB22CCEEFA36FF1B" box="[1226,1488,213,235]" italics="true" pageId="3" pageNumber="28">Descr. Catl. Reptiles Brit. Ind</emphasis>
.
</taxonomicName>
, p. 136.
</paragraph>
</subSubSection>
<subSubSection id="29EA69FBFFCFFFC9FCB5CD26FA9BFE55" pageId="3" pageNumber="28" type="materials_examined">
<paragraph id="614F3A70FFCFFFC9FCB5CD26FA9BFE55" blockId="3.[861,1499,286,421]" pageId="3" pageNumber="28">
<emphasis id="5384E662FFCFFFC9FCB5CD26FB72FEC8" box="[861,1172,286,313]" italics="true" pageId="3" pageNumber="28">Material examined (n=3)</emphasis>
:
<materialsCitation id="D198302DFFCFFFC9FB58CD27FB66FEAD" collectionCode="CSPT" collectorName="Anamalai" pageId="3" pageNumber="28" specimenCount="10" specimenCount-female="10">
<collectionCode id="07E1A2B5FFCFFFC9FB58CD27FB12FEC8" box="[1200,1268,287,313]" pageId="3" pageNumber="28">CSPT</collectionCode>
/S-
<specimenCount id="77F6F1F9FFCFFFC9FAFBCD27FA6BFEC9" box="[1299,1421,286,313]" count="10" pageId="3" pageNumber="28" type="female">10 female</specimenCount>
from
<collectorName id="CC055FA6FFCFFFC9FCB5CD7AFC2AFEAD" box="[861,972,322,348]" pageId="3" pageNumber="28">Anamalai</collectorName>
, date unknown
</materialsCitation>
;
<materialsCitation id="D198302DFFCFFFC9FB67CD7BFB24FE71" collectionCode="MAD" collectorName="Kavalai &amp; Cochindate" pageId="3" pageNumber="28" specimenCount="1">
<collectionCode id="07E1A2B5FFCFFFC9FB67CD7BFB36FEAC" box="[1167,1232,323,349]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="3" pageNumber="28">MAD</collectionCode>
no number male from
<collectorName id="CC055FA6FFCFFFC9FCB5CD5EFC54FE71" box="[861,946,358,384]" pageId="3" pageNumber="28">Kavalai</collectorName>
,
<collectorName id="CC055FA6FFCFFFC9FC2CCD5EFBA0FE71" box="[964,1094,358,384]" pageId="3" pageNumber="28">Cochindate</collectorName>
unknown
</materialsCitation>
;
<materialsCitation id="D198302DFFCFFFC9FB3CCD5EFA9FFE55" collectionCode="SACON" collectorName="S. Bhupathy &amp; Anamalai" pageId="3" pageNumber="28" specimenCount="46" specimenCount-female="46">
<collectionCode id="07E1A2B5FFCFFFC9FB3CCD5EFAD7FE71" box="[1236,1329,358,384]" country="India" httpUri="http://grbio.org/cool/fckk-x2vi" name="Salim Ali Centre for Ornithology and Natural History" pageId="3" pageNumber="28">SACON</collectionCode>
/VR-
<specimenCount id="77F6F1F9FFCFFFC9FA80CD5EFA3DFE71" box="[1384,1499,358,384]" count="46" pageId="3" pageNumber="28" type="female">46 female</specimenCount>
coll.
<collectorName id="CC055FA6FFCFFFC9FC7ACDB3FBFEFE55" box="[914,1048,394,421]" pageId="3" pageNumber="28">S. Bhupathy</collectorName>
from
<collectorName id="CC055FA6FFCFFFC9FBB4CDB2FB2DFE55" box="[1116,1227,394,420]" pageId="3" pageNumber="28">Anamalai</collectorName>
, date unknown
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="29EA69FBFFCFFFCEFCB5CD85FF2AFEE5" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28" type="discussion">
<paragraph id="614F3A70FFCFFFC9FCB5CD85FAE1FC76" blockId="3.[859,1501,444,903]" pageId="3" pageNumber="28">
<emphasis id="5384E662FFCFFFC9FCB5CD85FBDEFE27" box="[861,1080,445,470]" italics="true" pageId="3" pageNumber="28">Taxonomic History</emphasis>
: This species was first described as
<taxonomicName id="A6F041F3FFCFFFC9FCB5CDD8FB84FE0B" box="[861,1122,480,506]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="5384E662FFCFFFC9FCB5CDD8FB84FE0B" box="[861,1122,480,506]" italics="true" pageId="3" pageNumber="28">Plectrurus sanguineus</emphasis>
</taxonomicName>
by
<bibRefCitation id="05614781FFCFFFC9FB73CDD8FAB9FE0A" author="Beddome, R. H." box="[1179,1375,480,507]" pageId="3" pageNumber="28" pagination="14 - 16" refId="ref6894" refString="Beddome, R. H. 1867. Descriptions and figures of five new snakes from the Madras Presidency, Madras Quarterly Journal of Medical Sciences, 11: 14 - 16." type="journal article" year="1867">Beddome (1867)</bibRefCitation>
based on
<typeStatus id="BE4B84D2FFCFFFC9FCB5CE3CFC59FDEF" box="[861,959,516,542]" pageId="3" pageNumber="28" type="holotype">holotype</typeStatus>
from Anamalai forests above Ponachi [=Pollachi;
<bibRefCitation id="05614781FFCFFFC9FCB3CE10FBABFDB3" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." box="[859,1101,551,578]" pageId="3" pageNumber="28" refId="ref8891" refString="Wallach, V., Williams, K. L. and Boundy, J. 2014. Snakes of the World: A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, USA; p. 1237. https: // doi. org / 10.1201 / b 16901." type="journal volume" year="2014">
Wallach
<emphasis id="5384E662FFCFFFC9FC2CCE11FC1DFDB3" box="[964,1019,552,578]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
., 2014
</bibRefCitation>
] (10.446˚
<geoCoordinate id="04C45CB7FFCFFFC9FB55CE1FFAC9FDB3" box="[1213,1327,551,578]" degrees="76.984" direction="north" orientation="latitude" pageId="3" pageNumber="28" precision="55" value="76.984">N 76.984</geoCoordinate>
˚E;
<quantity id="A6089795FFCFFFC9FABECE10FA47FDB3" box="[1366,1441,552,578]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="28" unit="m" value="900.0">900 m</quantity>
asl).
<bibRefCitation id="05614781FFCFFFC9FCB5CE74FBFCFD96" author="Beddome, R. H." box="[861,1050,588,615]" pageId="3" pageNumber="28" pagination="3 - 33" refId="ref6982" refString="Beddome, R. H. 1886. An account of the earth-snakes of the peninsula of India and Ceylon, Annals and Magazines of Natural History, 17 (5): 3 - 33. https: // doi. org / 10.1080 / 00222938609460106." type="journal article" year="1886">Beddome (1886)</bibRefCitation>
whilst describing the genus
<taxonomicName id="A6F041F3FFCFFFC9FA8ACE75FA32FD97" box="[1378,1492,589,614]" class="Squamata" family="Uropeltidae" genus="Teretrurus" kingdom="Animalia" pageId="3" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="5384E662FFCFFFC9FA8ACE75FA32FD97" box="[1378,1492,589,614]" italics="true" pageId="3" pageNumber="28">Teretrurus</emphasis>
</taxonomicName>
, rightly allocated his
<taxonomicName id="A6F041F3FFCFFFC9FBA4CE49FB0CFD7B" box="[1100,1258,625,650]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="5384E662FFCFFFC9FBA4CE49FB0CFD7B" box="[1100,1258,625,650]" italics="true" pageId="3" pageNumber="28">P. sanguineus</emphasis>
</taxonomicName>
, as the type species, giving it the current name combination (
<bibRefCitation id="05614781FFCFFFC9FAD7CEACFA29FD5F" author="Smith, M. A." box="[1343,1487,660,686]" pageId="3" pageNumber="28" pagination="583" refId="ref8648" refString="Smith, M. A. 1943. Fauna of British India including Ceylon and Burma. Vol-III Serpentes. Taylor &amp; Francis, London, UK; p. 583." type="book chapter" year="1943">Smith, 1943</bibRefCitation>
). A subjective junior synonym
<taxonomicName id="A6F041F3FFCFFFC9FB21CE80FBEEFD07" authority="Theobald, 1876" authorityName="Theobald" authorityYear="1876" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="scabricauda">
<emphasis id="5384E662FFCFFFC9FB21CE80FA3CFD23" box="[1225,1498,696,722]" italics="true" pageId="3" pageNumber="28">Plectrurus scabricauda</emphasis>
<bibRefCitation id="05614781FFCFFFC9FCB5CEE4FBEEFD07" author="Theobald, W." box="[861,1032,732,758]" pageId="3" pageNumber="28" refId="ref8680" refString="Theobald, W. 1876. Descriptive catalogue of the reptiles of British India, Thacker, Spink and Co., Calcutta; p. xiii + 238. https: // doi. org / 10.5962 / bhl. title. 54001." type="book" year="1876">Theobald, 1876</bibRefCitation>
</taxonomicName>
described based on the
<typeStatus id="BE4B84D2FFCFFFC9FAF8CEE4FA94FD07" box="[1296,1394,732,758]" pageId="3" pageNumber="28" type="holotype">holotype</typeStatus>
<collectionCode id="07E1A2B5FFCFFFC9FA90CEE4FA79FD07" box="[1400,1439,732,758]" pageId="3" pageNumber="28">ZSI</collectionCode>
6996 (stated to be lost fide Das
<emphasis id="5384E662FFCFFFC9FB60CF39FB5FFCEB" box="[1160,1209,768,794]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
., 1998, but present in the collection N.S. Achyuthan pers. comm. in
<date id="154E1CB0FFCFFFC9FA8ACF1CFA33FCCF" box="[1378,1493,804,830]" pageId="3" pageNumber="28" value="2021-05">May 2021</date>
) from Anamallays exists (
<bibRefCitation id="05614781FFCFFFC9FB79CF70FAABFC93" author="Beddome, R. H." box="[1169,1357,840,866]" pageId="3" pageNumber="28" pagination="3 - 33" refId="ref6982" refString="Beddome, R. H. 1886. An account of the earth-snakes of the peninsula of India and Ceylon, Annals and Magazines of Natural History, 17 (5): 3 - 33. https: // doi. org / 10.1080 / 00222938609460106." type="journal article" year="1886">Beddome, 1886</bibRefCitation>
;
<bibRefCitation id="05614781FFCFFFC9FA8BCF70FC73FC77" author="Boulenger, G. A." pageId="3" pageNumber="28" refId="ref7122" refString="Boulenger, G. A. 1893. Catalogue of the snakes in the British Museum (Natural History). Volume I., containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae and Colubridae aglyphae, part. British Museum (Natural History), London; 448 pp." type="book" year="1893">Boulenger, 1893</bibRefCitation>
;
<bibRefCitation id="05614781FFCFFFC9FC48CF54FBC1FC77" author="Smith, M. A." box="[928,1063,876,902]" pageId="3" pageNumber="28" pagination="583" refId="ref8648" refString="Smith, M. A. 1943. Fauna of British India including Ceylon and Burma. Vol-III Serpentes. Taylor &amp; Francis, London, UK; p. 583." type="book chapter" year="1943">Smith, 1943</bibRefCitation>
;
<bibRefCitation id="05614781FFCFFFC9FBDACF54FB1AFC76" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." box="[1074,1276,876,903]" pageId="3" pageNumber="28" pagination="453 - 506" refId="ref8414" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V., and Somaweera, R. 2016. A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae), Zoosystema, 38 (4): 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2." type="journal article" year="2016">
Pyron
<emphasis id="5384E662FFCFFFC9FB95CF55FB48FC77" box="[1149,1198,876,902]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
., 2016
</bibRefCitation>
).
</paragraph>
<paragraph id="614F3A70FFCFFFCEFCB5CFA5FF2AFEE5" blockId="3.[860,1500,925,1888]" lastBlockId="4.[133,771,214,276]" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
<emphasis id="5384E662FFCFFFC9FCB5CFA5FC59FC46" box="[861,959,925,951]" italics="true" pageId="3" pageNumber="28">Remarks</emphasis>
:
<bibRefCitation id="05614781FFCFFFC9FC25CFA5FB77FC46" author="Boulenger, G. A." box="[973,1169,925,951]" pageId="3" pageNumber="28" refId="ref7122" refString="Boulenger, G. A. 1893. Catalogue of the snakes in the British Museum (Natural History). Volume I., containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae and Colubridae aglyphae, part. British Museum (Natural History), London; 448 pp." type="book" year="1893">Boulenger (1893)</bibRefCitation>
listed
<specimenCount id="77F6F1F9FFCFFFC9FB35CFA5FA9EFC46" box="[1245,1400,925,951]" count="6" pageId="3" pageNumber="28" type="generic">six specimens</specimenCount>
(
<specimenCount id="77F6F1F9FFCFFFC9FA62CFA5FA30FC46" box="[1418,1494,925,951]" count="1" pageId="3" pageNumber="28" type="male">1 male</specimenCount>
,
<specimenCount id="77F6F1F9FFCFFFC9FCB7CFF9FC2DFC2A" box="[863,971,961,987]" count="3" pageId="3" pageNumber="28" type="female">3 females</specimenCount>
,
<specimenCount id="77F6F1F9FFCFFFC9FC3FCFF9FBB4FC2D" box="[983,1106,961,988]" count="2" pageId="3" pageNumber="28" type="juvenile">2 juveniles</specimenCount>
) all with the locality data stated as “Anamallays” (sic) with a ventral range of 128-147. We agree with subsequent workers that as
<bibRefCitation id="05614781FFCFFFC9FAEDC831FA58FBD5" author="Beddome, R. H." box="[1285,1470,1033,1060]" pageId="3" pageNumber="28" pagination="14 - 16" refId="ref6894" refString="Beddome, R. H. 1867. Descriptions and figures of five new snakes from the Madras Presidency, Madras Quarterly Journal of Medical Sciences, 11: 14 - 16." type="journal article" year="1867">Beddome (1867)</bibRefCitation>
in his original description elaborated on only
<specimenCount id="77F6F1F9FFCFFFC9FAD7C816FA3FFBB6" box="[1343,1497,1069,1096]" count="1" pageId="3" pageNumber="28" type="generic">one specimen</specimenCount>
with 144 ventrals and 7 subcaudals, that specimen is the name-bearing type, the
<typeStatus id="BE4B84D2FFCFFFC9FB93C84DFB3BFB7E" box="[1147,1245,1141,1167]" pageId="3" pageNumber="28" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="05614781FFCFFFC9FB1AC84DFA36FB61" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." box="[1266,1488,1141,1168]" pageId="3" pageNumber="28" pagination="453 - 506" refId="ref8414" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V., and Somaweera, R. 2016. A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae), Zoosystema, 38 (4): 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2." type="journal article" year="2016">
Pyron
<emphasis id="5384E662FFCFFFC9FAACC84EFA9AFB7E" box="[1348,1404,1141,1167]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
., 2016
</bibRefCitation>
).
<bibRefCitation id="05614781FFCFFFC9FCB5C8A1FBA1FB42" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." box="[861,1095,1177,1204]" pageId="3" pageNumber="28" refId="ref8891" refString="Wallach, V., Williams, K. L. and Boundy, J. 2014. Snakes of the World: A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, USA; p. 1237. https: // doi. org / 10.1201 / b 16901." type="journal volume" year="2014">
Wallach
<emphasis id="5384E662FFCFFFC9FC57C8A2FC16FB42" box="[959,1008,1177,1203]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
. (2014)
</bibRefCitation>
had opined that specimen to be the lectotypebut the original description (
<bibRefCitation id="05614781FFCFFFC9FAF4C885FA32FB29" author="Beddome, R. H." box="[1308,1492,1213,1240]" pageId="3" pageNumber="28" pagination="14 - 16" refId="ref6894" refString="Beddome, R. H. 1867. Descriptions and figures of five new snakes from the Madras Presidency, Madras Quarterly Journal of Medical Sciences, 11: 14 - 16." type="journal article" year="1867">Beddome, 1867</bibRefCitation>
) makes no indication of any other specimen but for the described one and hence it was corrected by
<bibRefCitation id="05614781FFCFFFC9FA9FC93EFC2BFAB5" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." pageId="3" pageNumber="28" pagination="453 - 506" refId="ref8414" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V., and Somaweera, R. 2016. A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae), Zoosystema, 38 (4): 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2." type="journal article" year="2016">
Pyron
<emphasis id="5384E662FFCFFFC9FA2EC93EFC95FAB2" italics="true" pageId="3" pageNumber="28">et al</emphasis>
. (2016)
</bibRefCitation>
. In either case, the rest of the five specimensin that lot, with differing ventral counts, have no name-bearing status.
<bibRefCitation id="05614781FFCFFFC9FBE2C94AFB05FA7D" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." box="[1034,1251,1393,1420]" pageId="3" pageNumber="28" pagination="453 - 506" refId="ref8414" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V., and Somaweera, R. 2016. A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae), Zoosystema, 38 (4): 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2." type="journal article" year="2016">
Pyron
<emphasis id="5384E662FFCFFFC9FBBFC94AFB6DFA7A" box="[1111,1163,1393,1419]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
. (2016)
</bibRefCitation>
also commented that Boulengers ventral values of those
<specimenCount id="77F6F1F9FFCFFFC9FB1AC9ADFA98FA5E" box="[1266,1406,1429,1455]" count="6" pageId="3" pageNumber="28" type="generic">6 specimens</specimenCount>
did not match with that of Beddomes count (144), but herein the authorsopine that one specimens count (147) is the closest to Beddomes count. As explained above, owing to the large ventral count range (120-150) (
<bibRefCitation id="05614781FFCFFFC9FAFECA1DFA28F9CE" author="Boulenger, G. A." box="[1302,1486,1573,1599]" pageId="3" pageNumber="28" refId="ref7122" refString="Boulenger, G. A. 1893. Catalogue of the snakes in the British Museum (Natural History). Volume I., containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae and Colubridae aglyphae, part. British Museum (Natural History), London; 448 pp." type="book" year="1893">Boulenger, 1893</bibRefCitation>
), Boulengers statement that these specimens originate only from the Anamallays is most likelyincorrect. But the type specimen (with 144 or 147 ventrals) is consistent with what is generally the ventral value for the Anamalai population of
<taxonomicName id="A6F041F3FFCFFFC9FBEACAE2FB90F902" box="[1026,1142,1754,1779]" class="Squamata" family="Uropeltidae" genus="Teretrurus" kingdom="Animalia" pageId="3" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="5384E662FFCFFFC9FBEACAE2FB90F902" box="[1026,1142,1754,1779]" italics="true" pageId="3" pageNumber="28">Teretrurus</emphasis>
</taxonomicName>
.
<bibRefCitation id="05614781FFCFFFC9FB6ECAE1FAA2F905" author="Beddome, R. H." box="[1158,1348,1753,1780]" pageId="3" pageNumber="28" pagination="14 - 16" refId="ref6894" refString="Beddome, R. H. 1867. Descriptions and figures of five new snakes from the Madras Presidency, Madras Quarterly Journal of Medical Sciences, 11: 14 - 16." type="journal article" year="1867">Beddome (1867)</bibRefCitation>
stated in his original description, the dorsal scale rows count to be 17. Usually, it is understood as midbody count. But the value 17 was later reported as a miscount, the correct value of midbody count being 15 (
<bibRefCitation id="05614781FFC8FFCEFE55CCEEFD9AFF01" author="Boulenger, G. A." box="[445,636,214,240]" pageId="4" pageNumber="29" refId="ref7122" refString="Boulenger, G. A. 1893. Catalogue of the snakes in the British Museum (Natural History). Volume I., containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae and Colubridae aglyphae, part. British Museum (Natural History), London; 448 pp." type="book" year="1893">Boulenger, 1893</bibRefCitation>
;
<bibRefCitation id="05614781FFC8FFCEFD64CCEEFF59FEE5" author="Constable, J. D." pageId="4" pageNumber="29" pagination="59 - 160" refId="ref7307" refString="Constable, J. D. 1949. Reptiles from the Indian Peninsula in the Museum of Comparative Zoology, Bulletin of the Museum of Comparative Zoology, Harvard, 103: 59 - 160." type="journal article" year="1949">Constable, 1949</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="29EA69FBFFC8FFCEFF6DCD13FE7EFE27" pageId="4" pageNumber="29" type="diagnosis">
<paragraph id="614F3A70FFC8FFCEFF6DCD13FE7EFE27" blockId="4.[133,771,299,470]" pageId="4" pageNumber="29">
<emphasis id="5384E662FFC8FFCEFF6DCD13FE65FEB4" box="[133,387,299,325]" italics="true" pageId="4" pageNumber="29">Differential Diagnosis</emphasis>
: A species of
<taxonomicName id="A6F041F3FFC8FFCEFDCDCD14FD7FFEB4" box="[549,665,300,325]" class="Squamata" family="Uropeltidae" genus="Teretrurus" kingdom="Animalia" pageId="4" pageNumber="29" phylum="Chordata" rank="genus">
<emphasis id="5384E662FFC8FFCEFDCDCD14FD7FFEB4" box="[549,665,300,325]" italics="true" pageId="4" pageNumber="29">Teretrurus</emphasis>
</taxonomicName>
endemic to the Anamalai Hills, possessing a distinct supraocular scale (vs. lacking in
<taxonomicName id="A6F041F3FFC8FFCEFE9ACD4CFDFFFE7C" box="[370,537,371,397]" class="Squamata" family="Uropeltidae" genus="Brachyophidium" kingdom="Animalia" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="5384E662FFC8FFCEFE9ACD4CFDFFFE7C" box="[370,537,371,397]" italics="true" pageId="4" pageNumber="29">T. rhodogaster</emphasis>
</taxonomicName>
), with much higher ventral counts 142-150 (vs.
<quantity id="A6089795FFC8FFCEFE28CDAFFDDFFE40" box="[448,569,407,433]" metricMagnitude="0" metricUnit="m" metricValue="3.1496" metricValueMax="3.2512" metricValueMin="3.048" pageId="4" pageNumber="29" unit="in" value="124.0" valueMax="128.0" valueMin="120.0">120-128 in</quantity>
<taxonomicName id="A6F041F3FFC8FFCEFDABCDA0FD24FE40" box="[579,706,407,433]" class="Squamata" family="Uropeltidae" genus="Platyplectrurus" kingdom="Animalia" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="hewstoni">
<emphasis id="5384E662FFC8FFCEFDABCDA0FD24FE40" box="[579,706,407,433]" italics="true" pageId="4" pageNumber="29">T. hewstoni</emphasis>
</taxonomicName>
;
<quantity id="A6089795FFC8FFCEFD38CDAFFF2DFE24" metricMagnitude="0" metricUnit="m" metricValue="3.3781999999999996" metricValueMax="3.4543999999999997" metricValueMin="3.3019999999999996" pageId="4" pageNumber="29" unit="in" value="133.0" valueMax="136.0" valueMin="130.0">130- 136 in</quantity>
<taxonomicName id="A6F041F3FFC8FFCEFF39CD84FE6FFE24" box="[209,393,444,469]" class="Squamata" family="Uropeltidae" genus="Teretrurus" kingdom="Animalia" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="travancoricus">
<emphasis id="5384E662FFC8FFCEFF39CD84FE6FFE24" box="[209,393,444,469]" italics="true" pageId="4" pageNumber="29">T. travancoricus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="29EA69FBFFC8FFCEFF6DCDD5FF00F89D" pageId="4" pageNumber="29" type="description">
<paragraph id="614F3A70FFC8FFCEFF6DCDD5FEB2FB43" blockId="4.[133,773,492,1203]" pageId="4" pageNumber="29">
<emphasis id="5384E662FFC8FFCEFF6DCDD5FE4AFDF7" box="[133,428,492,518]" italics="true" pageId="4" pageNumber="29">Description and variation</emphasis>
:
<emphasis id="5384E662FFC8FFCEFE52CDD5FD4AFDF7" box="[442,684,493,518]" italics="true" pageId="4" pageNumber="29">Measurements in mm</emphasis>
: snoutvent length: 150-190; tail length: 2.5-6; head length: 4.5- 5.5; head width: 4-5; body width 4.5-6; eye diameter: 0.8; eye-snout distance: 1.8; inter-orbital distance: 2-2.5; inter-narial distance: 0.5-0.7. Habitus: Small-bodied, trunk cylindrical to slightly compressed, especially near tail; body fairly slender; head not distinct from neck, as wide as midbody; snout obtusely rounded in profile; tail small, suddenly tapering to a sharp, finepoint.
<emphasis id="5384E662FFC8FFCEFD7CCF34FD1BFCD7" box="[660,765,780,806]" italics="true" pageId="4" pageNumber="29">Scalation</emphasis>
: scales smooth, cycloid, without apical pits; head scales lacking pre-, post-oculars; supraocular and temporal present; eye covered under ocular scale, partially fused; mental groove absent; dorsal scales in 16:15:15 rows, with 16 --&gt;15 at 36-38th ventral scale; supralabials 4/4; infralabials 4/4; ventrals 144-149; subcaudals 7-9 pairs; terminal caudal scales rather smooth, with a single sharp projection in the end. Coloration: dorsum dark to coffee brown, rather uniform and unpatterned; venter rich red (turning creamy yellow in preservative) with a few black patches; eye black.
</paragraph>
<paragraph id="614F3A70FFC8FFCEFF6DC8F2FF00F89D" blockId="4.[132,772,1226,1900]" pageId="4" pageNumber="29">
<taxonomicName id="A6F041F3FFC8FFCEFF6DC8F2FE68FB15" box="[133,398,1226,1252]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="scabricauda">
<emphasis id="5384E662FFC8FFCEFF6DC8F2FE68FB15" box="[133,398,1226,1252]" italics="true" pageId="4" pageNumber="29">Plectrurus scabricauda</emphasis>
</taxonomicName>
<typeStatus id="BE4B84D2FFC8FFCEFE7EC8F2FE18FB15" box="[406,510,1226,1252]" pageId="4" pageNumber="29" type="holotype">Holotype</typeStatus>
<collectionCode id="07E1A2B5FFC8FFCEFDEEC8F2FDCBFB15" box="[518,557,1226,1252]" pageId="4" pageNumber="29">ZSI</collectionCode>
6996 Annamallay: Measurements in mm: Snout to vent length 159; tail length 13; head length 5; head width 3.5;
<emphasis id="5384E662FFC8FFCEFDAFC92AFD46FADD" box="[583,672,1298,1324]" italics="true" pageId="4" pageNumber="29">Habitus</emphasis>
: a rather soft and brittle specimen; body slightly damaged at mid portion due to preservation artifact; trunk subcylindrical to mildly dorso-ventrally depressed at places, owing to long-time preservation; head small, snout pointed to slightly ovoid in profile;
<emphasis id="5384E662FFC8FFCEFE2CC9FEFDC8FA11" box="[452,558,1478,1504]" italics="true" pageId="4" pageNumber="29">Scalation</emphasis>
: dorsal scale rows 13:15:13 scales smooth, cycloid, without apical pits; scales on head wrinkled, due to long-time preservation; eyes covered under ocular scales; supraocular and temporal present; preocular, postocular, loreal and mental groove absent; supralabials 4/4; infralabials 4/4; ventrals 141, subcaudals 11 pairs; terminal caudal scales equally wrinkled as head scales, due to preservation artifact.
<emphasis id="5384E662FFC8FFCEFF6DCADEFEE6F8F1" box="[133,256,1766,1792]" italics="true" pageId="4" pageNumber="29">Coloration</emphasis>
: dorsum drab greyish brown, venter creamy orange, except towards body extremities (mental and subcaudals) which present with patches of darker shades; eye grey.
</paragraph>
</subSubSection>
<subSubSection id="29EA69FBFFC8FFCEFCABCCEDFAE5FD32" pageId="4" pageNumber="29" type="biology_ecology">
<paragraph id="614F3A70FFC8FFCEFCABCCEDFAE5FD32" blockId="4.[833,1475,213,707]" pageId="4" pageNumber="29">
<emphasis id="5384E662FFC8FFCEFCABCCEDFB52FF1E" box="[835,1204,213,239]" italics="true" pageId="4" pageNumber="29">Distribution and Natural History</emphasis>
: In a relativelyrestricted sense (as re-appraised hereinincl.
<taxonomicName id="A6F041F3FFC8FFCEFB35CCC2FA6AFEE2" box="[1245,1420,249,275]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="scabricauda">
<emphasis id="5384E662FFC8FFCEFB35CCC2FA6AFEE2" box="[1245,1420,249,275]" italics="true" pageId="4" pageNumber="29">P. scabricauda</emphasis>
</taxonomicName>
),
<taxonomicName id="A6F041F3FFC8FFCEFA44CCC2FC27FEC6" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="5384E662FFC8FFCEFA44CCC2FC27FEC6" italics="true" pageId="4" pageNumber="29">T. sanguineus</emphasis>
</taxonomicName>
is definitely known only from Anamalai-Munnar ranges (
<bibRefCitation id="05614781FFC8FFCEFBE9CD79FB51FEAD" author="Beddome, R. H." box="[1025,1207,321,348]" pageId="4" pageNumber="29" pagination="14 - 16" refId="ref6894" refString="Beddome, R. H. 1867. Descriptions and figures of five new snakes from the Madras Presidency, Madras Quarterly Journal of Medical Sciences, 11: 14 - 16." type="journal article" year="1867">Beddome, 1867</bibRefCitation>
). It is perhaps the only congener that partly co-occurs with another congener
<taxonomicName id="A6F041F3FFC8FFCEFCABCDB2FC0FFE52" box="[835,1001,393,419]" class="Squamata" family="Uropeltidae" genus="Brachyophidium" kingdom="Animalia" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="5384E662FFC8FFCEFCABCDB2FC0FFE52" box="[835,1001,393,419]" italics="true" pageId="4" pageNumber="29">T. rhodogaster</emphasis>
</taxonomicName>
that inhabits a much easterly and higher elevation range (Palni Hills).
<bibRefCitation id="05614781FFC8FFCEFB6ACD95FAA2FE39" author="Rajendran, M. V." box="[1154,1348,429,456]" pageId="4" pageNumber="29" pagination="132" refId="ref8492" refString="Rajendran, M. V. 1985. Studies in Uropeltid snakes, Madurai Kamaraj University, Madurai, India; p. 132." type="book chapter" year="1985">Rajendran (1985)</bibRefCitation>
mentioned of having collected this species from Nyamakkad of Kannan Devan Hills (10.151˚N, 77.080˚E;
<quantity id="A6089795FFC8FFCEFADECDCDFA68FDFE" box="[1334,1422,501,527]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="4" pageNumber="29" unit="m" value="1870.0">1870 m</quantity>
asl) near Munnar. This apparently rare species has not been recently sighted anywhere in the Anamalai (museum holding records in
<bibRefCitation id="05614781FFC8FFCEFBC1CE59FB00FD8A" author="Boulenger, G. A." box="[1065,1254,609,635]" pageId="4" pageNumber="29" refId="ref7122" refString="Boulenger, G. A. 1893. Catalogue of the snakes in the British Museum (Natural History). Volume I., containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae and Colubridae aglyphae, part. British Museum (Natural History), London; 448 pp." type="book" year="1893">Boulenger, 1893</bibRefCitation>
;
<bibRefCitation id="05614781FFC8FFCEFB1FCE59FA52FD8A" author="Constable, J. D." box="[1271,1460,609,635]" pageId="4" pageNumber="29" pagination="59 - 160" refId="ref7307" refString="Constable, J. D. 1949. Reptiles from the Indian Peninsula in the Museum of Comparative Zoology, Bulletin of the Museum of Comparative Zoology, Harvard, 103: 59 - 160." type="journal article" year="1949">Constable, 1949</bibRefCitation>
), despite surveys that covered at least some parts of this massif (
<bibRefCitation id="05614781FFC8FFCEFC70CE91FBFCFD32" author="Roux, J." box="[920,1050,681,707]" pageId="4" pageNumber="29" pagination="439 - 471" refId="ref8517" refString="Roux, J. 1928. Reptiles et amphibiens de l'Inde meridionale, Revue Suisse de Zoologie, Geneve, 35: 439 - 471. [In French]. https: // doi. org / 10.5962 / bhl. part. 117624." type="journal article" year="1928">Roux, 1928</bibRefCitation>
;
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</paragraph>
</subSubSection>
</treatment>
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