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<document id="FDC919AB5AA84807402E9A57F9A98C14" ID-DOI="10.5281/zenodo.6391684" ID-ISSN="1942-1354" ID-Zenodo-Dep="6391684" ID-ZooBank="CBD29188-143B-44DF-BE21-1654D50D8621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648501406612" checkinUser="felipe" docAuthor="Phillips, Robert A." docDate="2022" docId="E8511E53FF88EF296A8AFAAEFCE8FE64" docLanguage="en" docName="InsectaMundi.2022.0907.1-214.pdf" docOrigin="Insecta Mundi 2022 (907)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Culicoides (Selfia) moabensis Phillips G 2015" docType="treatment" docVersion="3" lastPageNumber="84" masterDocId="1468662BFFDCEF7C6A4AFFC3FFCCFFE4" masterDocTitle="Culicoides Latreille and Leptoconops Skuse biting midges of the southwestern United States with emphasis on the Canyonlands of southeastern Utah (Diptera: Ceratopogonidae)" masterLastPageNumber="214" masterPageNumber="1" pageNumber="83" updateTime="1699308763882" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="59628D3E6FA6B39BF02E71526A57AC0A">Culicoides Latreille and Leptoconops Skuse biting midges of the southwestern United States with emphasis on the Canyonlands of southeastern Utah (Diptera: Ceratopogonidae)</mods:title>
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<mods:name id="863FCED0B4BA8754A05A4D357A815340" type="personal">
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<mods:namePart id="77BF0EBF864EF678D3CA18672CC6B004">Phillips, Robert A.</mods:namePart>
<mods:affiliation id="DE3C616604B1913E5C1B0155F07EECAE">2962 Desert Road Moab, UT 84532 USA</mods:affiliation>
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<mods:title id="F122F358C656D7373AF5C94399A91355">Insecta Mundi</mods:title>
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<mods:part id="82FD092B38B2D3A08A8B54922C5A7BEF">
<mods:date id="192E52FD386EB4E30D572432BD3368BB">2022</mods:date>
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<mods:number id="9CAE04DA5490BB7971533C09B4BE48A8">2022-01-28</mods:number>
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<mods:identifier id="82A22AE77740B18595B7245F564F85B2" type="ISSN">1942-1354</mods:identifier>
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<treatment id="E8511E53FF88EF296A8AFAAEFCE8FE64" ID-DOI="http://doi.org/10.5281/zenodo.6391774" ID-Zenodo-Dep="6391774" LSID="urn:lsid:plazi:treatment:E8511E53FF88EF296A8AFAAEFCE8FE64" httpUri="http://treatment.plazi.org/id/E8511E53FF88EF296A8AFAAEFCE8FE64" lastPageId="85" lastPageNumber="84" pageId="84" pageNumber="83">
<subSubSection id="28E2FCCEFF88EF286A8AFAAEFD56FA6E" box="[192,666,1389,1418]" pageId="84" pageNumber="83" type="nomenclature">
<paragraph id="6047AF45FF88EF286A8AFAAEFD56FA6E" blockId="84.[192,666,1389,1453]" box="[192,666,1389,1418]" pageId="84" pageNumber="83">
<heading id="3B0F1829FF88EF286A8AFAAEFD56FA6E" box="[192,666,1389,1418]" fontSize="12" level="2" pageId="84" pageNumber="83" reason="2">
<taxonomicName id="A7F8D4C6FF88EF286A8AFAAEFD56FA6E" authority="Phillips" authorityName="Phillips G" authorityYear="2015" box="[192,666,1389,1418]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="84" pageNumber="83" phylum="Arthropoda" rank="species" species="moabensis" subGenus="Selfia">
<emphasis id="528C7357FF88EF286A8AFAAEFD56FA6E" bold="true" box="[192,666,1389,1418]" pageId="84" pageNumber="83">
<emphasis id="528C7357FF88EF286A8AFAAEFE88FA6E" bold="true" box="[192,324,1389,1418]" italics="true" pageId="84" pageNumber="83">Culicoides</emphasis>
(
<emphasis id="528C7357FF88EF286B1EFAAEFE55FA6E" bold="true" box="[340,409,1389,1418]" italics="true" pageId="84" pageNumber="83">Selfia</emphasis>
)
<emphasis id="528C7357FF88EF286BE0FAAEFDE3FA6E" bold="true" box="[426,559,1389,1418]" italics="true" pageId="84" pageNumber="83">moabensis</emphasis>
Phillips
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFF88EF286A8AFA51FE8FFA49" box="[192,323,1426,1453]" pageId="84" pageNumber="83" type="description">
<paragraph id="6047AF45FF88EF286A8AFA51FE8FFA49" blockId="84.[192,666,1389,1453]" box="[192,323,1426,1453]" pageId="84" pageNumber="83">
(
<figureCitation id="F8C3B3C0FF88EF286A82FA51FEF7FA49" box="[200,315,1426,1453]" captionStart="Figures 3336. 33" captionStartId="168.[193,280,1765,1791]" captionTargetBox="[221,1409,213,1734]" captionTargetId="figure-102@168.[199,1435,213,1736]" captionTargetPageId="168" captionText="Figures 3336. 33) Culicoides moabensis female (top) and unidentified brown C. subgenus Selfia female (bottom), lateral habitus (in alcohol). Male genitalia, ventral view focused on aedeagus. 34) C. brookmani, basal arm of paramere (Pa), distal tip of median process of paramere delineated (Pt), basal arm of aedeagus (Aa), tip of median process of aedeagus (At), gonocoxite (Gc), gonostylus (Gs). 35) C. moabensis, median process of paramere (Pmp) (paratype). 36) C. denningi, sternite 9 caudal lobes (S9cl)." figureDoi="http://doi.org/10.5281/zenodo.6391700" httpUri="https://zenodo.org/record/6391700/files/figure.png" pageId="84" pageNumber="83">Fig. 33, 35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFF88EF286AAAFA78FB9AFA17" pageId="84" pageNumber="83" type="reference_group">
<paragraph id="6047AF45FF88EF286AAAFA78FB9AFA17" blockId="84.[224,1440,1467,1523]" pageId="84" pageNumber="83">
<treatmentCitationGroup id="40E8886BFF88EF286AAAFA78FB9AFA17" pageId="84" pageNumber="83">
<treatmentCitation id="E1598954FF88EF286AAAFA78FD03FA37" author="Phillips RA" box="[224,719,1467,1491]" page="842" pageId="84" pageNumber="83" year="2015">
<taxonomicName id="A7F8D4C6FF88EF286AAAFA78FD03FA37" authority="Phillips, 2015: 842" authorityName="Phillips G" authorityPageNumber="842" authorityYear="2015" box="[224,719,1467,1491]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="84" pageNumber="83" phylum="Arthropoda" rank="species" species="moabensis" subGenus="Selfia">
<emphasis id="528C7357FF88EF286AAAFA78FE85FA37" box="[224,329,1467,1491]" italics="true" pageId="84" pageNumber="83">Culicoides</emphasis>
(
<emphasis id="528C7357FF88EF286B12FA78FE42FA37" box="[344,398,1467,1491]" italics="true" pageId="84" pageNumber="83">Selfia</emphasis>
)
<emphasis id="528C7357FF88EF286BD4FA78FDC5FA37" box="[414,521,1467,1491]" italics="true" pageId="84" pageNumber="83">moabensis</emphasis>
<materialsCitation id="D090A518FF88EF28685AFA78FD03FA37" box="[528,719,1467,1491]" county="Grand County" location="Grand County" pageId="84" pageNumber="83" specimenCount="1" stateProvince="Utah">
<bibRefCitation id="0469D2B4FF88EF28685AFA78FD03FA37" author="Phillips RA" box="[528,719,1467,1491]" pageId="84" pageNumber="83" pagination="842 - 849" refId="ref138233" refString="Phillips RA. 2015. A new species of Culicoides (Selfia) (Diptera: Ceratopogonidae) from southeastern Utah. Journal of Medical Entomology 52: 842 - 849." type="journal article" year="2015">Phillips, 2015: 842</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(key; diagnosis, male, female; fig. male head, thorax, wing, genitalia; female head, thorax, color, wing, genitalia; seasonal activity;
<collectingRegion id="A23C61A7FF88EF28693BFA19FC64FA16" box="[881,936,1498,1522]" country="United States of America" name="Utah" pageId="84" pageNumber="83">Utah</collectingRegion>
:
<collectingCounty id="8926D7C9FF88EF2869FBFA19FB87FA17" box="[945,1099,1498,1523]" pageId="84" pageNumber="83">Grand County</collectingCounty>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFF88EF286A8AF9C0FE8DF908" pageId="84" pageNumber="83" type="diagnosis">
<paragraph id="6047AF45FF88EF286A8AF9C0FE8DF908" blockId="84.[192,1442,1539,1772]" pageId="84" pageNumber="83">
<emphasis id="528C7357FF88EF286A8AF9C0FEF1F9F9" bold="true" box="[192,317,1539,1565]" pageId="84" pageNumber="83">Diagnosis.</emphasis>
(
<tableCitation id="2D7A9AFEFF88EF286B01F9C0FE74F9F9" box="[331,440,1539,1565]" captionStart="Table 14" captionStartId="151.[186,249,455,483]" captionText="Table 14. Quantitative characters of Culicoides. Entries to the right of the median dashed line apply only to females. Data in () are variations. Wing pattern: 0 none, + reduced, ++ extensive but faint or diffuse, +++ extensive and distinct. Eye separations are approximate in ommatidium diameters. Spermathecae “n” is the number of fully developed spermathecae, with a “v” indicating an additional vestigial spermatheca, and “ratio” is the sum of the largers width and length including any sclerotized neck divided by the sum of the smallers width and length including any sclerotized necks. Measurements and ratios are expressed as means. All data for species marked (r) and data for wing length and costal, antennal, palpal, and proboscis ratios are compiled from the description references in the species accounts, with emphasis on southwestern United States data, and supplemented or modified from borrowed and collected specimens." pageId="84" pageNumber="83">Tables 14</tableCitation>
,
<tableCitation id="2D7A9AFEFF88EF286B89F9C0FE12F9FA" box="[451,478,1539,1566]" captionStart="Table 15" captionStartId="154.[192,255,214,240]" captionText="Table 15. Pale leg-banding on Culicoides. ++ = distinct, + = faint or indistinct, ± = faint to absent,? = undetermined. Shading indicates darker pigmentation. Data for species marked (r) are compiled from the literature." pageId="84" pageNumber="83">15</tableCitation>
) Yellowish to reddish yellow; wing without pattern of pale spots; three unsclerotized or faintly sclerotized long fingerlike spermathecae; male scutellum with four setae; hind tarsomeres without apical spines; posterior margin of sternite 9 convex over base of aedeagus, without paired median lobes; basal third of gonocoxite abruptly expanded and bent mesally; gonocoxal apodeme simple, without hooklike process; aedeagus ~2× longer than wide, lateral arms not boomerang-shaped sclerites, apical third trifurcate into subparallel lobes&gt;2× longer than wide; parameres fused, as long as wide, median process elongate triangular tonguelike.
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFF88EF296A8AF93FFDFFFED0" lastPageId="85" lastPageNumber="84" pageId="84" pageNumber="83" type="distribution">
<paragraph id="6047AF45FF88EF286A8AF93FFD9FF8F2" blockId="84.[192,595,1788,1815]" box="[192,595,1788,1815]" pageId="84" pageNumber="83">
<emphasis id="528C7357FF88EF286A8AF93FFE90F8F2" bold="true" box="[192,348,1788,1814]" pageId="84" pageNumber="83">Distribution.</emphasis>
Grand County,
<collectingRegion id="A23C61A7FF88EF286859F93FFD83F8F2" box="[531,591,1788,1814]" country="United States of America" name="Utah" pageId="84" pageNumber="83">Utah</collectingRegion>
.
</paragraph>
<paragraph id="6047AF45FF88EF286A8AF8E4FDA1F862" blockId="84.[192,1440,1831,1927]" pageId="84" pageNumber="83">
<emphasis id="528C7357FF88EF286A8AF8E4FEB5F8A5" bold="true" box="[192,377,1831,1857]" pageId="84" pageNumber="83">Adult behavior.</emphasis>
The mandibular and lacinial teeth on the female and collection in CO
<subScript id="FC7CAD00FF88EF286EEFF8F5FB61F8A0" attach="right" box="[1189,1197,1846,1860]" fontSize="6" pageId="84" pageNumber="83">2</subScript>
-baited traps indicate it feeds on vertebrate blood; and though its hosts are unknown, the SCo presence on only the proximal flagellomeres suggests it is mammalophilic.
</paragraph>
<paragraph id="6047AF45FF89EF296B4AFF15FDFFFED0" blockId="85.[192,1440,214,308]" pageId="85" pageNumber="84">
<taxonomicName id="A7F8D4C6FF89EF296B4AFF15FE22FF14" authorityName="Phillips G" authorityYear="2015" box="[256,494,214,240]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="moabensis">
<emphasis id="528C7357FF89EF296B4AFF15FE22FF14" box="[256,494,214,240]" italics="true" pageId="85" pageNumber="84">Culicoides moabensis</emphasis>
</taxonomicName>
was collected from mid-April through October, with strong peaks in late May (week 21) and early October (week 41), suggesting it is bivoltine in Grand County, which may not be the case in other areas (see
<taxonomicName id="A7F8D4C6FF89EF296B78FED9FE06FED7" authorityName=": Khalaf" authorityYear="1954" box="[306,458,281,308]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="brookmani">
<emphasis id="528C7357FF89EF296B78FED9FE88FED0" box="[306,324,282,308]" italics="true" pageId="85" pageNumber="84">C</emphasis>
.
<emphasis id="528C7357FF89EF296B1AFEDAFE06FED7" box="[336,458,281,307]" italics="true" pageId="85" pageNumber="84">brookmani</emphasis>
</taxonomicName>
biology).
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFF89EF296A8AFE80FCE8FE64" pageId="85" pageNumber="84" type="discussion">
<paragraph id="6047AF45FF89EF296A8AFE80FCE8FE64" blockId="85.[192,1440,323,384]" pageId="85" pageNumber="84">
<emphasis id="528C7357FF89EF296A8AFE80FEFCFEB9" bold="true" box="[192,304,323,349]" pageId="85" pageNumber="84">Remarks.</emphasis>
<taxonomicName id="A7F8D4C6FF89EF296B7FFE80FDEEFEB9" authorityName="Phillips G" authorityYear="2015" box="[309,546,323,349]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="moabensis">
<emphasis id="528C7357FF89EF296B7FFE80FDEEFEB9" box="[309,546,323,349]" italics="true" pageId="85" pageNumber="84">Culicoides moabensis</emphasis>
</taxonomicName>
is more closely related to
<taxonomicName id="A7F8D4C6FF89EF296977FE87FC1AFEB9" authorityName=": Khalaf" authorityYear="1954" box="[829,982,323,350]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="brookmani">
<emphasis id="528C7357FF89EF296977FE87FC83FEBA" box="[829,847,324,350]" italics="true" pageId="85" pageNumber="84">C</emphasis>
.
<emphasis id="528C7357FF89EF296916FE80FC1AFEB9" box="[860,982,323,349]" italics="true" pageId="85" pageNumber="84">brookmani</emphasis>
</taxonomicName>
and
<taxonomicName id="A7F8D4C6FF89EF296E40FE87FB18FEB9" authorityName="Malloch" authorityYear="1915" box="[1034,1236,323,350]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="multipunctatus">
<emphasis id="528C7357FF89EF296E40FE87FBD0FEBA" box="[1034,1052,324,350]" italics="true" pageId="85" pageNumber="84">C</emphasis>
.
<emphasis id="528C7357FF89EF296E63FE80FB18FEB9" box="[1065,1236,323,349]" italics="true" pageId="85" pageNumber="84">multipunctatus</emphasis>
</taxonomicName>
of the Multipunctatus group than to other
<taxonomicName id="A7F8D4C6FF89EF296BA8FEA6FDD3FE9B" box="[482,543,357,383]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="167" phylum="Arthropoda" rank="subGenus" subGenus="Selfia">
<emphasis id="528C7357FF89EF296BA8FEA6FDD3FE9B" box="[482,543,357,383]" italics="true" pageId="85" pageNumber="84">Selfia</emphasis>
</taxonomicName>
species (
<bibRefCitation id="0469D2B4FF89EF2968C8FEA6FCD5FE64" author="Phillips RA" box="[642,793,357,384]" pageId="85" pageNumber="84" pagination="842 - 849" refId="ref138233" refString="Phillips RA. 2015. A new species of Culicoides (Selfia) (Diptera: Ceratopogonidae) from southeastern Utah. Journal of Medical Entomology 52: 842 - 849." type="journal article" year="2015">Phillips 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>