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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. &amp; Marek, Paul E." docDate="2021" docId="E76C06AC898355BE918A94BDDEEE05AE" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria laminata Hoffman 1949" docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Means, Jackson C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hennen, Derek A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marek, Paul E.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1030</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181004703" LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE" httpUri="http://treatment.plazi.org/id/E76C06AC898355BE918A94BDDEEE05AE" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="45M4R" ID-ENA="2675608" LSID="E76C06AC-8983-55BE-918A-94BDDEEE05AE" authority="Hoffman, 1949" authorityName="Hoffman" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria laminata" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="laminata">Nannaria laminata Hoffman, 1949</taxonomicName>
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Nannaria laminata holotype ♂ (NMNH, Type 1806) left gonopod A anterior view B medial view C posterior view; red arrow indicates reduced, acuminated prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure48" httpUri="https://binary.pensoft.net/fig/531151" pageId="0" pageNumber="1">Figs 48</figureCitation>
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Nannaria laminata non-type ♂ (VTEC, MPE 02392) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure49" httpUri="https://binary.pensoft.net/fig/531152" pageId="0" pageNumber="1">, 49 Vernacular name: &quot;The Laminate Twisted-Claw Millipede&quot;</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Hoffman" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria laminata" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="laminata">Nannaria laminata</taxonomicName>
Hoffman, 1949: 383, figs 11, 12.
<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.212" author="Chamberlin, RV" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="1 - 236" refId="B22" refString="Chamberlin, RV, Hoffman, RL, 1958. Checklist of the millipeds of North America. United States National Museum Bulletin 212: 1 - 236, DOI: https://doi.org/10.5479/si.03629236.212" title="Checklist of the millipeds of North America." url="https://doi.org/10.5479/si.03629236.212" volume="212" year="1958">Chamberlin and Hoffman 1958</bibRefCitation>
: 41.
<bibRefCitation author="Hoffman, RL" journalOrPublisher="Virginia Museum of Natural History Special Publication Number" pageId="0" pageNumber="1" pagination="1 - 584" refId="B33" refString="Hoffman, RL, 1999. Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publication Number 8: 1 - 584" title="Checklist of the millipeds of North and Middle America." volume="8" year="1999">Hoffman 1999</bibRefCitation>
: 367.
<bibRefCitation author="Marek, PE" journalOrPublisher="Virginia Museum of Natural History Publications" pageId="0" pageNumber="1" pagination="1 - 117" refId="B46" refString="Marek, PE, Tanabe, T, Sierwald, P, 2014. A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida). Virginia Museum of Natural History Publications 17: 1 - 117" title="A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida)." volume="17" year="2014">Marek et al. 2014</bibRefCitation>
: 37.
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. 2021</bibRefCitation>
: S70.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355720" collectingDate="1947-07-12" collectorName="H. H. Hobbs, C. M. Wilson, Type" country="U. S." county="Mercer County" latitude="37.355" location="Route" longLatPrecision="6" longitude="-80.8982" municipality="Glen Lyn" specimenCount="♂" stateProvince="West Virginia" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
United States -
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
</emphasis>
<specimenCount></specimenCount>
;
<collectingCounty>Mercer County</collectingCounty>
, ravine beside
<collectingCountry name="United States of America">U.S.</collectingCountry>
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:CE5879BDE3977D3BC0A2AC95EB069482" country="U. S." county="Mercer County" latitude="37.355" longLatPrecision="6" longitude="-80.8982" municipality="Glen Lyn" name="Route" stateProvince="West Virginia">Route</location>
460, ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.218688" unit="mi" value="2.0">2 miles</quantity>
south of
<collectingMunicipality>Glen Lyn</collectingMunicipality>
,
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
; [
<geoCoordinate degrees="37.3550" direction="north" orientation="latitude" precision="5" value="37.355">37.3550°N</geoCoordinate>
, -
<geoCoordinate degrees="80.8982" direction="west" orientation="longitude" precision="5" value="-80.8982">80.8982°W</geoCoordinate>
];
<collectingDate value="1947-07-12">12 July 1947</collectingDate>
;
<collectorName>H. H. Hobbs</collectorName>
,
<collectorName>C. M. Wilson</collectorName>
leg.; NMNH
<collectorName>Type</collectorName>
#1806.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355366" collectingDate="1999-11-13" collectorName="Survey" country="United States" county="Amherst County" latitude="37.7665" location="Tarjacket Ridge" longLatPrecision="6" longitude="-79.1863" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
</emphasis>
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Amherst County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:3A79AFEFEF80CB9EDAA2A7DCF5FB9721" country="United States" county="Amherst County" latitude="37.7665" longLatPrecision="6" longitude="-79.1863" name="Tarjacket Ridge" stateProvince="Virginia">Tarjacket Ridge</location>
FS 1167, 3500';
<geoCoordinate degrees="37.7665" direction="north" orientation="latitude" precision="5" value="37.7665">37.7665°N</geoCoordinate>
, -
<geoCoordinate degrees="79.1863" direction="west" orientation="longitude" precision="5" value="-79.1863">79.1863°W</geoCoordinate>
;
<collectingDate value="1999-11-13">13 Nov. 1999</collectingDate>
; VMNH
<collectorName>Survey</collectorName>
leg.; VMNH NAN0332
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355773" accessionNumber="MPE02401" collectingDate="2017-03-18" collectorName="J. Means" country="United States" county="Augusta County" elevation="828" latitude="37.89643" location="Barnwood Cabin" longLatPrecision="4" longitude="-79.0007" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Augusta County</collectingCounty>
, across creek from
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:B791F8FFAE486FA0334A98FE516F0CCA" country="United States" county="Augusta County" latitude="37.89643" longLatPrecision="4" longitude="-79.0007" name="Barnwood Cabin" stateProvince="Virginia">Barnwood Cabin</location>
, where old road flattens out by small stream;
<geoCoordinate degrees="37.89643" direction="north" orientation="latitude" precision="1" value="37.89643">37.89643°N</geoCoordinate>
, -
<geoCoordinate degrees="79.0007" direction="west" orientation="longitude" precision="5" value="-79.0007">79.0007°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.28" unit="m" value="828.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.28" unit="m" value="828.0">828 m</elevation>
</quantity>
;
<collectingDate value="2017-03-18">18 Mar. 2017</collectingDate>
;
<collectorName>J. Means</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02401">MPE02401</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355723" accessionNumber="MPE03679" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means" country="United States" county="Augusta County" elevation="828" latitude="37.89643" location="Barnwood Cabin" longLatPrecision="4" longitude="-79.0007" specimenCount="1" specimenCount-female="1" stateProvince="Virginia">
<specimenCount type="female">1 ♀</specimenCount>
; same collection data as preceding; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03679">MPE03679</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355613" accessionNumber="MPE02392" collectingDate="2017-03-19" collectorName="J. Means" country="United States" county="Augusta County" elevation="607" latitude="37.9139" location="upper Sherando Lake" longLatPrecision="6" longitude="-79.0207" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Augusta County</collectingCounty>
, far corner of
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:640C339B667DB7F892C2ABD538FC837E" country="United States" county="Augusta County" latitude="37.9139" longLatPrecision="6" longitude="-79.0207" name="upper Sherando Lake" stateProvince="Virginia">upper Sherando Lake</location>
, bank side by marsh;
<geoCoordinate degrees="37.9139" direction="north" orientation="latitude" precision="5" value="37.9139">37.9139°N</geoCoordinate>
, -
<geoCoordinate degrees="79.0207" direction="west" orientation="longitude" precision="5" value="-79.0207">79.0207°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.07" unit="m" value="607.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.07" unit="m" value="607.0">607 m</elevation>
</quantity>
;
<collectingDate value="2017-03-19">19 Mar. 2017</collectingDate>
;
<collectorName>J. Means</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02392">MPE02392</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355309" collectingDate="1986-10-27" collectorName="J. Mitchell" country="United States" county="Bedford County" latitude="37.4498" location="Peaks of Otter" longLatPrecision="6" longitude="-79.5834" municipality="Flattop Mtn." specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Bedford County</collectingCounty>
,
<collectingMunicipality>Flattop Mtn.</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:C399F280980D450124ADCEA874960106" country="United States" county="Bedford County" latitude="37.4498" longLatPrecision="6" longitude="-79.5834" municipality="Flattop Mtn." name="Peaks of Otter" stateProvince="Virginia">Peaks of Otter</location>
nr MP 82.8;
<geoCoordinate degrees="37.4498" direction="north" orientation="latitude" precision="5" value="37.4498">37.4498°N</geoCoordinate>
, -
<geoCoordinate degrees="79.5834" direction="west" orientation="longitude" precision="5" value="-79.5834">79.5834°W</geoCoordinate>
;
<collectingDate value="1986-10-27">27 Oct. 1986</collectingDate>
;
<collectorName>J. Mitchell</collectorName>
leg.; VMNH NAN0330
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355505" collectingDate="1980-05-16" collectorName="R. Hoffman" country="United States" county="Bland County" latitude="37.7032" location="Hamilton's Cave" longLatPrecision="6" longitude="-78.1627" specimenCount="3" specimenCount-male="3" stateProvince="Virginia">
<specimenCount type="male">3 ♂♂</specimenCount>
;
<collectingCounty>Bland County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:BBAC01576BCED69A5C99213DA4AA3794" country="United States" county="Bland County" latitude="37.7032" longLatPrecision="6" longitude="-78.1627" name="Hamilton's Cave" stateProvince="Virginia">
<normalizedToken originalValue="Hamiltons">Hamilton's</normalizedToken>
Cave
</location>
, ca.
<locationDeviation location="Mechanicsville">
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
ENE of Mechanicsville
</locationDeviation>
, rich wooded hillside;
<geoCoordinate degrees="37.7032" direction="north" orientation="latitude" precision="5" value="37.7032">37.7032°N</geoCoordinate>
, -
<geoCoordinate degrees="78.1627" direction="west" orientation="longitude" precision="5" value="-78.1627">78.1627°W</geoCoordinate>
;
<collectingDate value="1980-05-16">16 May 1980</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0150
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355335" collectingDate="1989-10-21" collectorName="J. Mitchell" country="United States" county="Botetourt County" latitude="37.4577" location="Blue Ridge Parkway" longLatPrecision="6" longitude="-79.6175" municipality="Harkening Hill" specimenCount="2" specimenCount-male="2" stateProvince="Virginia">
<specimenCount type="male">2 ♂♂</specimenCount>
;
<collectingCounty>Botetourt County</collectingCounty>
,
<collectingMunicipality>Harkening Hill</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:AEC6F4F40A434860717DE4804B1FCDDC" country="United States" county="Botetourt County" latitude="37.4577" longLatPrecision="6" longitude="-79.6175" municipality="Harkening Hill" name="Blue Ridge Parkway" stateProvince="Virginia">Blue Ridge Parkway</location>
;
<geoCoordinate degrees="37.4577" direction="north" orientation="latitude" precision="5" value="37.4577">37.4577°N</geoCoordinate>
, -
<geoCoordinate degrees="79.6175" direction="west" orientation="longitude" precision="5" value="-79.6175">79.6175°W</geoCoordinate>
;
<collectingDate value="1989-10-21">21 Oct. 1989</collectingDate>
;
<collectorName>J. Mitchell</collectorName>
leg.; VMNH NAN0333
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355684" collectingDate="1973-10-13" collectorName="R. Hoffman" country="United States" county="Botetourt County" latitude="37.5411" location="Arcadia" longLatPrecision="6" longitude="-79.5866" municipality="North Creek" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Botetourt County</collectingCounty>
,
<collectingMunicipality>North Creek</collectingMunicipality>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.218688" metricValueMax="4.828032" metricValueMin="1.609344" unit="mi" value="2.0" valueMax="3.0" valueMin="1.0">1-3 mi.</quantity>
east of
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:F4EB0BE796E68E898DB4F9EE1351B912" country="United States" county="Botetourt County" latitude="37.5411" longLatPrecision="6" longitude="-79.5866" municipality="North Creek" name="Arcadia" stateProvince="Virginia">Arcadia</location>
;
<geoCoordinate degrees="37.5411" direction="north" orientation="latitude" precision="5" value="37.5411">37.5411°N</geoCoordinate>
, -
<geoCoordinate degrees="79.5866" direction="west" orientation="longitude" precision="5" value="-79.5866">79.5866°W</geoCoordinate>
;
<collectingDate value="1973-10-13">13 Oct. 1973</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0334
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355615" collectingDate="1950-10-14" collectorName="L. Hubricht" country="United States" county="Campbell County" latitude="37.3176" location="4 miles NW of Rustburg" longLatPrecision="6" longitude="-79.1526" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Campbell County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:4E7EDDC37AD3421758D8A30ED1CE3181" country="United States" county="Campbell County" latitude="37.3176" longLatPrecision="6" longitude="-79.1526" name="4 miles NW of Rustburg" stateProvince="Virginia">
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.437376" unit="mi" value="4.0">4 miles</quantity>
NW of Rustburg
</location>
;
<geoCoordinate degrees="37.3176" direction="north" orientation="latitude" precision="5" value="37.3176">37.3176°N</geoCoordinate>
, -
<geoCoordinate degrees="79.1526" direction="west" orientation="longitude" precision="5" value="-79.1526">79.1526°W</geoCoordinate>
;
<collectingDate value="1950-10-14">14 Oct. 1950</collectingDate>
;
<collectorName>L. Hubricht</collectorName>
leg.; VMNH NAN0331
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355383" collectingDate="1990-05-17" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Cumberland County</collectingCounty>
, clearcut north DF site 2 ca
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:9E5F4B80C8635CFAB2D89E04B0BA7CD4" country="United States" county="Cumberland County" latitude="37.7361" longLatPrecision="6" longitude="-78.1714" name="2 km SSW of Columbia" stateProvince="Virginia">
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
SSW of Columbia
</location>
;
<geoCoordinate degrees="37.7361" direction="north" orientation="latitude" precision="5" value="37.7361">37.7361°N</geoCoordinate>
, -
<geoCoordinate degrees="78.1714" direction="west" orientation="longitude" precision="5" value="-78.1714">78.1714°W</geoCoordinate>
;
<collectingDate value="1990-05-17">17 May 1990</collectingDate>
;
<collectorName>J. Mitchell</collectorName>
leg.; VMNH NAN0253
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355671" collectingDate="1989-11" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="3" specimenCount-male="3" stateProvince="Virginia">
<specimenCount type="male">3 ♂♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1989-11">Nov. 1989</collectingDate>
; VMNH NAN0259
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355455" collectingDate="1990-04-02" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1990-04-02">2 Apr. 1990</collectingDate>
; VMNH NAN0261
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355454" collectingDate="1989-12-03" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1989-12-03">3 Dec. 1989</collectingDate>
; VMNH NAN0263
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355507" collectingDate="1990-02-15" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1990-02-15">15 Feb. 1990</collectingDate>
; VMNH NAN0264
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355694" collectingDate="1989-10-19" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="19" specimenCount-male="19" stateProvince="Virginia">
<specimenCount type="male">19 ♂♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1989-10-19">19 Oct. 1989</collectingDate>
VMNH NAN0266
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355609" collectingDate="1989-11-16" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="7" specimenCount-male="7" stateProvince="Virginia">
<specimenCount type="male">7 ♂♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1989-11-16">16 Nov. 1989</collectingDate>
; VMNH NAN0267
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355444" collectingDate="1989-11" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="11" specimenCount-male="11" stateProvince="Virginia">
<specimenCount type="male">11 ♂</specimenCount>
<specimenCount></specimenCount>
; same collection data as preceding;
<collectingDate value="1989-11">Nov. 1989</collectingDate>
; VMNH NAN0361
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355700" collectingDate="1990-05-01" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="6" specimenCount-male="6" stateProvince="Virginia">
<specimenCount type="male">6 ♂♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1990-05-01">1 May 1990</collectingDate>
; VMNH NAN0255
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355514" collectingDate="1990-05-01" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1715" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Cumberland County</collectingCounty>
, hardwood site 1 (north),
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:B9F1C5981EABFA2219B8C9960EE0B9C2" country="United States" county="Cumberland County" latitude="37.7361" longLatPrecision="6" longitude="-78.1715" name="2 km SSW of Columbia" stateProvince="Virginia">
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
SSW of Columbia
</location>
;
<geoCoordinate degrees="37.7361" direction="north" orientation="latitude" precision="5" value="37.7361">37.7361°N</geoCoordinate>
, -
<geoCoordinate degrees="78.1715" direction="west" orientation="longitude" precision="5" value="-78.1715">78.1715°W</geoCoordinate>
;
<collectingDate value="1990-05-01">1 May 1990</collectingDate>
;
<collectorName>J. Mitchell</collectorName>
leg.; VMNH NAN0254
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355646" collectingDate="1990-03-17" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW of Columbia" longLatPrecision="6" longitude="-78.1715" specimenCount="24" specimenCount-male="24" stateProvince="Virginia">
<specimenCount type="male">24 ♂♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1990-03-17">17 Mar. 1990</collectingDate>
; VMNH NAN0268
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355703" collectingDate="1989-10-05" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.6953" location="7 km SSW of Columbia" longLatPrecision="6" longitude="-78.1959" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Cumberland County</collectingCounty>
, hardwood site 4 (south),
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:60E419FF247EBEC881F2DA628D60A6E6" country="United States" county="Cumberland County" latitude="37.6953" longLatPrecision="6" longitude="-78.1959" name="7 km SSW of Columbia" stateProvince="Virginia">
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
SSW of Columbia
</location>
;
<geoCoordinate degrees="37.6953" direction="north" orientation="latitude" precision="5" value="37.6953">37.6953°N</geoCoordinate>
, -
<geoCoordinate degrees="78,1959" direction="west" orientation="longitude" precision="5" value="-78.1959">78,1959°W</geoCoordinate>
;
<collectingDate value="1989-10-05">5 Oct. 1989</collectingDate>
;
<collectorName>J. Mitchell</collectorName>
; VMNH NAN0257
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355437" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="United States" county="Cumberland County" latitude="37.7361" location="2 km SSW Columbia" longLatPrecision="6" longitude="-78.1714" specimenCount="3" specimenCount-male="3" stateProvince="Virginia">
<specimenCount type="male">3 ♂♂</specimenCount>
;
<collectingCounty>Cumberland County</collectingCounty>
, hardwood site 2 - N
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:452C92DCC254E15322064F22CABB49B7" country="United States" county="Cumberland County" latitude="37.7361" longLatPrecision="6" longitude="-78.1714" name="2 km SSW Columbia" stateProvince="Virginia">
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
SSW Columbia
</location>
;
<geoCoordinate degrees="37.7361" direction="north" orientation="latitude" precision="5" value="37.7361">37.7361°N</geoCoordinate>
, -
<geoCoordinate degrees="78.1714" direction="west" orientation="longitude" precision="5" value="-78.1714">78.1714°W</geoCoordinate>
; VMNH NAN0258
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355495" collectingDate="1990-05-17" collectorName="J. Mitchell" country="United States" county="Cumberland County" latitude="37.7048" location="5.5 km SSW of Columbia" longLatPrecision="6" longitude="-78.1889" specimenCount="2" specimenCount-male="2" stateProvince="Virginia">
<specimenCount type="male">2 ♂♂</specimenCount>
;
<collectingCounty>Cumberland County</collectingCounty>
, pinewoods DF site 5, ca.
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:3D9EBFB72AB46AE4D42E904866C84E9F" country="United States" county="Cumberland County" latitude="37.7048" longLatPrecision="6" longitude="-78.1889" name="5.5 km SSW of Columbia" stateProvince="Virginia">
<quantity metricMagnitude="3" metricUnit="m" metricValue="5.5" unit="km" value="5.5">5.5 km</quantity>
SSW of Columbia
</location>
;
<geoCoordinate degrees="37.7048" direction="north" orientation="latitude" precision="5" value="37.7048">37.7048°N</geoCoordinate>
, -
<geoCoordinate degrees="78.1889" direction="west" orientation="longitude" precision="5" value="-78.1889">78.1889°W</geoCoordinate>
;
<collectingDate value="1990-05-17">17 May 1990</collectingDate>
;
<collectorName>J. Mitchell</collectorName>
leg.; VMNH NAN0532
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355468" accessionNumber="MPE00891" collectingDate="2015-10-21" collectorName="D. Hennen" country="United States" county="Giles County" elevation="544" latitude="37.35" location="Cascades Day Use Area" longLatPrecision="6" longitude="-80.5835" municipality="Jefferson National Forest" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Giles County</collectingCounty>
,
<collectingMunicipality>Jefferson National Forest</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:59EEE3DB60FC116356084BAD958E404D" country="United States" county="Giles County" latitude="37.35" longLatPrecision="6" longitude="-80.5835" municipality="Jefferson National Forest" name="Cascades Day Use Area" stateProvince="Virginia">Cascades Day Use Area</location>
, within a few hundred meters of the start of the
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:EDD36789B0518FFB2492F4DB9F2D2282" country="United States" county="Giles County" latitude="37.35" longLatPrecision="6" longitude="-80.5835" municipality="Jefferson National Forest" name="Cascades Trail" stateProvince="Virginia">Cascades Trail</location>
, near parking lot;
<geoCoordinate degrees="37.3500" direction="north" orientation="latitude" precision="5" value="37.35">37.3500°N</geoCoordinate>
, -
<geoCoordinate degrees="80.5835" direction="west" orientation="longitude" precision="5" value="-80.5835">80.5835°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" unit="m" value="544.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" unit="m" value="544.0">544 m</elevation>
</quantity>
;
<collectingDate value="2015-10-21">21 Oct. 2015</collectingDate>
; hand collected;
<collectorName>D. Hennen</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00891">MPE00891</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355710" accessionNumber="MPE03807" collectingDate="2018-02-18" collectorName="J. Means" country="United States" county="Giles County" elevation="799" latitude="37.3613" location="Cascades" longLatPrecision="6" longitude="-80.5875" municipality="Upper Trail" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Giles County</collectingCounty>
, side of
<collectingMunicipality>Upper Trail</collectingMunicipality>
on way to
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:C873EAAA7843BB623E8DD40606C02E0D" country="United States" county="Giles County" latitude="37.3613" longLatPrecision="6" longitude="-80.5875" municipality="Upper Trail" name="Cascades" stateProvince="Virginia">Cascades</location>
;
<geoCoordinate degrees="37.3613" direction="north" orientation="latitude" precision="5" value="37.3613">37.3613°N</geoCoordinate>
, -
<geoCoordinate degrees="80.5875" direction="west" orientation="longitude" precision="5" value="-80.5875">80.5875°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.99" unit="m" value="799.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.99" unit="m" value="799.0">799 m</elevation>
</quantity>
;
<collectingDate value="2018-02-18">18 Feb. 2018</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03807">MPE03807</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355370" accessionNumber="MPE02124" collectingDate="2016-10-03" collectorName="J. Means, D. Hennen, V. Wong" country="United States" county="Giles County" elevation="1062" latitude="37.4209" location="Mountain Lake Biological Station" longLatPrecision="6" longitude="-80.5093" municipality="Jefferson National Forest" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Giles County</collectingCounty>
,
<collectingMunicipality>Jefferson National Forest</collectingMunicipality>
past
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:430A444C8E9CFE62E95500C249B3585B" country="United States" county="Giles County" latitude="37.4209" longLatPrecision="6" longitude="-80.5093" municipality="Jefferson National Forest" name="Mountain Lake Biological Station" stateProvince="Virginia">Mountain Lake Biological Station</location>
, past
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:B940DFE0B6A913F0926AA0178BE26AB4" country="United States" county="Giles County" latitude="37.4209" longLatPrecision="6" longitude="-80.5093" municipality="Jefferson National Forest" name="Mini Ball Hill on Rt" stateProvince="Virginia">Mini Ball Hill on Rt</location>
613;
<geoCoordinate degrees="37.4209" direction="north" orientation="latitude" precision="5" value="37.4209">37.4209°N</geoCoordinate>
, -
<geoCoordinate degrees="80.5093" direction="west" orientation="longitude" precision="5" value="-80.5093">80.5093°W</geoCoordinate>
; elev.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.062" unit="m" value="1062.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.062" unit="m" value="1062.0">1062 m</elevation>
</quantity>
;
<collectingDate value="2016-10-03">3 Oct. 2016</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>D. Hennen</collectorName>
,
<collectorName>V. Wong</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02124">MPE02124</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355597" accessionNumber="MPE02136" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means &amp; D. Hennen &amp; V. Wong" country="United States" county="Giles County" elevation="1062" latitude="37.4209" location="Mountain Lake Biological Station" longLatPrecision="6" longitude="-80.5093" municipality="Jefferson National Forest" specimenCount="2" specimenCount-female="2" stateProvince="Virginia">
<specimenCount type="female">2 ♀♀</specimenCount>
; same collection data as preceding; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02136">MPE02136</accessionNumber>
</materialsCitation>
, 2142 •
<materialsCitation ID-GBIF-Occurrence="3071355361" collectingDate="1988-06-28" collectorName="W. Shear" country="United States" county="Prince Edward County" latitude="37.3019" location="Price Drive" longLatPrecision="6" longitude="-78.3922" municipality="Farmville" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Prince Edward County</collectingCounty>
,
<collectingMunicipality>Farmville</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:D4CD62F5459CB1D01AFEEA645E2E9770" country="United States" county="Prince Edward County" latitude="37.3019" longLatPrecision="6" longitude="-78.3922" municipality="Farmville" name="Price Drive" stateProvince="Virginia">Price Drive</location>
;
<geoCoordinate degrees="37.3019" direction="north" orientation="latitude" precision="5" value="37.3019">37.3019°N</geoCoordinate>
, -
<geoCoordinate degrees="78.3922" direction="west" orientation="longitude" precision="5" value="-78.3922">78.3922°W</geoCoordinate>
;
<collectingDate value="1988-06-28">28 June 1988</collectingDate>
;
<collectorName>W. Shear</collectorName>
leg.; VMNH NAN0256
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355545" collectingDate="1975-02-22" collectorName="W. Shear" country="United States" county="Prince Edward County" latitude="37.1134" location="13 mi S Farmville" longLatPrecision="6" longitude="-78.3922" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Prince Edward County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:617FCF33E71399C1B0EEA9123F5DAB7F" country="United States" county="Prince Edward County" latitude="37.1134" longLatPrecision="6" longitude="-78.3922" name="13 mi S Farmville" stateProvince="Virginia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.0921472" unit="mi" value="13.0">13 mi</quantity>
S Farmville
</location>
, dug up
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.2" unit="cm" value="12.0">12 cm</quantity>
under surface of clay;
<geoCoordinate degrees="37.1134" direction="north" orientation="latitude" precision="5" value="37.1134">37.1134°N</geoCoordinate>
, -
<geoCoordinate degrees="78.3922" direction="west" orientation="longitude" precision="5" value="-78.3922">78.3922°W</geoCoordinate>
;
<collectingDate value="1975-02-22">22 Feb. 1975</collectingDate>
;
<collectorName>W. Shear</collectorName>
leg.; VMNH NAN0260
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355555" collectingDate="1981-06-17" collectingMethod="pitfall trap" collectorName="R. Bellinger" country="United States" county="Prince Edward County" latitude="37.275" location="Rice" longLatPrecision="6" longitude="-78.2916" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Prince Edward County</collectingCounty>
,
<collectingMethod>pitfall trap</collectingMethod>
, old field at
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:76747A58187311A9CD8E42A54D503A5E" country="United States" county="Prince Edward County" latitude="37.275" longLatPrecision="6" longitude="-78.2916" name="Rice" stateProvince="Virginia">Rice</location>
;
<geoCoordinate degrees="37.2750" direction="north" orientation="latitude" precision="5" value="37.275">37.2750°N</geoCoordinate>
, -
<geoCoordinate degrees="78.2916" direction="west" orientation="longitude" precision="5" value="-78.2916">78.2916°W</geoCoordinate>
;
<collectingDate value="1981-06-17">17 June 1981</collectingDate>
;
<collectorName>R. Bellinger</collectorName>
leg.; VMNH NAN0265; SCAU
</materialsCitation>
-
<materialsCitation ID-GBIF-Occurrence="3071355460" accessionNumber="MPE03474" collectingDate="2017-11-12" collectorName="J. Means" country="United States" county="Mercer County" elevation="507" latitude="37.3452" location="2 miles SW of Glen Lyn" longLatPrecision="6" longitude="-80.9105" specimenCount="1" specimenCount-female="1" stateProvince="West Virginia">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
</emphasis>
<specimenCount type="female">1 ♀</specimenCount>
;
<collectingCounty>Mercer County</collectingCounty>
, just over the WV state line on 460, ca.
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:96074694BF767442E89B9AE07A10EBB1" country="United States" county="Mercer County" latitude="37.3452" longLatPrecision="6" longitude="-80.9105" name="2 miles SW of Glen Lyn" stateProvince="West Virginia">
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.218688" unit="mi" value="2.0">2 miles</quantity>
SW of Glen Lyn
</location>
on 219/8;
<geoCoordinate degrees="37.3452" direction="north" orientation="latitude" precision="5" value="37.3452">37.3452°N</geoCoordinate>
, -
<geoCoordinate degrees="80.9105" direction="west" orientation="longitude" precision="5" value="-80.9105">80.9105°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.07" unit="m" value="507.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.07" unit="m" value="507.0">507 m</elevation>
</quantity>
;
<collectingDate value="2017-11-12">12 Nov. 2017</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03474">MPE03474</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355431" accessionNumber="MPE02520" collectingDate="2017-04-27" collectorName="J. Means, D. Hennen, P. Marek, P. Shorter, V. Wong" country="United States" county="Monroe County" elevation="934" latitude="37.4749" location="Linside" longLatPrecision="6" longitude="-80.5673" specimenCount="1" specimenCount-male="1" stateProvince="West Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Monroe County</collectingCounty>
, 5.8 air miles east of
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:44B37A6D92BD801BC7C327218816946A" country="United States" county="Monroe County" latitude="37.4749" longLatPrecision="6" longitude="-80.5673" name="Linside" stateProvince="West Virginia">Linside</location>
, up logging road;
<geoCoordinate degrees="37.4749" direction="north" orientation="latitude" precision="5" value="37.4749">37.4749°N</geoCoordinate>
, -
<geoCoordinate degrees="80.5673" direction="west" orientation="longitude" precision="5" value="-80.5673">80.5673°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.34" unit="m" value="934.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.34" unit="m" value="934.0">934 m</elevation>
</quantity>
;
<collectingDate value="2017-04-27">27 Apr. 2017</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>D. Hennen</collectorName>
,
<collectorName>P. Marek</collectorName>
,
<collectorName>P. Shorter</collectorName>
,
<collectorName>V. Wong</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02520">MPE02520</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355701" accessionNumber="MPE02578" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means &amp; D. Hennen &amp; P. Marek &amp; P. Shorter &amp; V. Wong" country="United States" county="Monroe County" elevation="934" latitude="37.4749" location="Linside" longLatPrecision="6" longitude="-80.5673" specimenCount="3" specimenCount-female="3" stateProvince="West Virginia">
<specimenCount type="female">3 ♀♀</specimenCount>
; same collection data as preceding; VTEC,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02578">MPE02578</accessionNumber>
</materialsCitation>
, 2579, 3675 •
<materialsCitation ID-GBIF-Occurrence="3071355373" accessionNumber="MPE02528" collectingDate="2016-04-27" collectorName="J. Means, D. Hennen, P. Marek, P. Shorter, V. Wong" country="United States" county="Monroe County" elevation="1107" latitude="37.4717" location="Monroe County" longLatPrecision="6" longitude="-80.562" specimenCount="2" specimenCount-male="2" stateProvince="West Virginia">
<specimenCount type="male">2 ♂♂</specimenCount>
;
<collectingCounty>Monroe County</collectingCounty>
, along logging road off VA rt 613, down in bowl and on each side of dirt and gravel road;
<geoCoordinate degrees="37.4717" direction="north" orientation="latitude" precision="5" value="37.4717">37.4717°N</geoCoordinate>
, -
<geoCoordinate degrees="80.5620" direction="west" orientation="longitude" precision="5" value="-80.562">80.5620°W</geoCoordinate>
; elev.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.107" unit="m" value="1107.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.107" unit="m" value="1107.0">1107 m</elevation>
</quantity>
;
<collectingDate value="2016-04-27">27 Apr. 2016</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>D. Hennen</collectorName>
,
<collectorName>P. Marek</collectorName>
,
<collectorName>P. Shorter</collectorName>
,
<collectorName>V. Wong</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02528">MPE02528</accessionNumber>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3071355321" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Suppl." country="United States" location="For" specimenCount="1" stateProvince="West Virginia">
2561.
<location LSID="urn:lsid:plazi:treatment:E76C06AC898355BE918A94BDDEEE05AE:1E2A5F97516F814CB1FCE0EA95FE1F26" country="United States" name="For" stateProvince="West Virginia">For</location>
detailed collection data see
<collectorName>Suppl.</collectorName>
material 7
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult males of
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. laminata" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="laminata">
<emphasis italics="true" pageId="0" pageNumber="1">N. laminata</emphasis>
</taxonomicName>
are distinct from other
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
and the nearby
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. solenas" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">N. solenas</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. wilsoni" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wilsoni">
<emphasis italics="true" pageId="0" pageNumber="1">N. wilsoni</emphasis>
</taxonomicName>
, based on the following combination of characters:
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
.
</emphasis>
Gonopodal acropodite semi-circular, curving dorsomedially with abrupt 90° curve after apex, not straight before abrupt 90° curve as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. solenas" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">N. solenas</emphasis>
</taxonomicName>
sp. nov. or with laminate corkscrew before apex as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. wilsoni" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wilsoni">
<emphasis italics="true" pageId="0" pageNumber="1">N. wilsoni</emphasis>
</taxonomicName>
. Acropodite tip simple, directed posteriorly, not directed anteriorly as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. solenas" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">N. solenas</emphasis>
</taxonomicName>
sp. nov. and not with triangular lateral flange as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. wilsoni" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wilsoni">
<emphasis italics="true" pageId="0" pageNumber="1">N. wilsoni</emphasis>
</taxonomicName>
. Tip terminating in sharp claw-like point, not blunt point as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. solenas" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">N. solenas</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. wilsoni" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wilsoni">
<emphasis italics="true" pageId="0" pageNumber="1">N. wilsoni</emphasis>
</taxonomicName>
. Height of telopodite basal zone&gt; 1/3 length of acropodite, not ca. 1/2 length as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. solenas" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">N. solenas</emphasis>
</taxonomicName>
sp. nov., or ca. 1/5 as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. wilsoni" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wilsoni">
<emphasis italics="true" pageId="0" pageNumber="1">N. wilsoni</emphasis>
</taxonomicName>
. Prefemoral process large, laminate and serpentine, with prefemoral spine reduced to small, acuminate projection (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Nannaria laminata holotype ♂ (NMNH, Type 1806) left gonopod A anterior view B medial view C posterior view; red arrow indicates reduced, acuminated prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure48" httpUri="https://binary.pensoft.net/fig/531151" pageId="0" pageNumber="1">48C</figureCitation>
, red arrow), not lacking as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. wilsoni" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wilsoni">
<emphasis italics="true" pageId="0" pageNumber="1">N. wilsoni</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
.
</emphasis>
Tergites with red paranotal spots (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Nannaria laminata non-type ♂ (VTEC, MPE 02392) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure49" httpUri="https://binary.pensoft.net/fig/531152" pageId="0" pageNumber="1">49</figureCitation>
). Black background. Dorsum of collum smooth with red margin.
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure48" httpUri="https://binary.pensoft.net/fig/531151" pageId="0" pageNumber="1" start="Figure 48" startId="F48">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 48.</emphasis>
<taxonomicName authorityName="Hoffman" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria laminata" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="laminata">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria laminata</emphasis>
</taxonomicName>
holotype ♂ (NMNH, Type #1806) left gonopod
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
medial view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior view; red arrow indicates reduced, acuminated prefemoral spine. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1030.62544.figure49" httpUri="https://binary.pensoft.net/fig/531152" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
<taxonomicName authorityName="Hoffman" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria laminata" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="laminata">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria laminata</emphasis>
</taxonomicName>
non-type ♂ (VTEC, MPE02392) coloration. Scale bar: 4.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="1">♂ holotype (NMNH, Type #1806): BL = 30.5, CW = 4.0, IW = 2.2, ISW = 0.9, B11W = 4.9, B11H = 3.4.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Hoffman" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria laminata" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="laminata">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria laminata</emphasis>
</taxonomicName>
occupies a relatively large geographic area,&gt; 10,000 km2, and displays a fair amount of morphological variation; however, the general shape of the acropodite and laminate prefemoral process remain constant throughout. Variation is seen in the sharpness and curve of the acropodite tip; some individuals display a more medially directed, blunt tip than the holotype. Additionally, the width of the prefemoral process (when viewed anteriorly or posteriorly) varies widely between individuals.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known from a horizontal strip from central Virginia to just north of the border into West Virginia (Virginia: Amherst, Augusta, Bedford, Botetourt, Campbell, Cumberland, Giles, and Prince Edward counties; West Virginia: Mercer and Monroe counties, Suppl. material 7; Fig.
<figureCitation captionStart="Figure 126" captionStartId="F126" captionText="Figure 126. Distribution map of Nannaria minor species group collection localities in the central Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure126" httpUri="https://binary.pensoft.net/fig/531229" pageId="0" pageNumber="1">126</figureCitation>
). Distribution area: 9,297 km2; status: SRE.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Individuals of
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. laminata" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="laminata">
<emphasis italics="true" pageId="0" pageNumber="1">N. laminata</emphasis>
</taxonomicName>
have been collected from mesic deciduous forests dominated by maple, oak, rhododendron, and some pine, often under 1-2 cm soil and/or leaf litter.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.5479/si.00963801.99-3244.371" author="Hoffman, RL" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" pagination="371 - 389" refId="B29" refString="Hoffman, RL, 1949. Nine new xystodesmid millipedes from Virginia and West Virginia, with records of established species. Proceedings of the United States National Museum 99: 371 - 389, DOI: https://doi.org/10.5479/si.00963801.99-3244.371" title="Nine new xystodesmid millipedes from Virginia and West Virginia, with records of established species." url="https://doi.org/10.5479/si.00963801.99-3244.371" volume="99" year="1949">Hoffman (1949)</bibRefCitation>
gives no explanation for the name
<taxonomicName authorityName="Hoffman" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria laminata" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="laminata">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria laminata</emphasis>
</taxonomicName>
in his description of the species, however it is reasonable to assume that the name refers to the laminate prefemoral process, the major distinguishing morphological character.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
United States, West Virginia, Mercer County, ravine beside U.S. Route 460, ca. 2 miles south of Glen Lyn, Virginia; [
<geoCoordinate degrees="37.3550" direction="north" orientation="latitude" precision="5" value="37.355">37.3550°N</geoCoordinate>
, -
<geoCoordinate degrees="80.8982" direction="west" orientation="longitude" precision="5" value="-80.8982">80.8982°W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
In the original publication,
<bibRefCitation DOI="https://doi.org/10.5479/si.00963801.99-3244.371" author="Hoffman, RL" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" pagination="371 - 389" refId="B29" refString="Hoffman, RL, 1949. Nine new xystodesmid millipedes from Virginia and West Virginia, with records of established species. Proceedings of the United States National Museum 99: 371 - 389, DOI: https://doi.org/10.5479/si.00963801.99-3244.371" title="Nine new xystodesmid millipedes from Virginia and West Virginia, with records of established species." url="https://doi.org/10.5479/si.00963801.99-3244.371" volume="99" year="1949">Hoffman (1949</bibRefCitation>
: 384) designates a male holotype (NMNH Type #1806) collected by H. Hobbs and C. Wilson on July 12, 1947.
</paragraph>
</subSubSection>
</treatment>
</document>