treatments-xml/data/E3/12/95/E3129514090DFF974093FB24FBBEFB0A.xml
2024-06-21 12:55:18 +02:00

2174 lines
215 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="CADE6B0A34B6682E71534ECADBC9B2F4" ID-CLB-Dataset="39259" ID-DOI="10.11646/zootaxa.4058.2.4" ID-GBIF-Dataset="b6aeffc9-06f4-4e5d-bd0e-cf480b5e06d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="253690" ID-ZooBank="1FDCAFAB-1E21-43A7-9543-E9071EA48FA7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461358880208" checkinUser="plazi" docAuthor="Vassilieva, Anna B." docDate="2015" docId="E3129514090DFF974093FB24FBBEFB0A" docLanguage="en" docName="zt04058p226.pdf" docOrigin="Zootaxa 4058 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oligodon arenarius Vassilieva, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="222" masterDocId="1F2BED6C090CFF9B4004FFD5FFC9FFAA" masterDocTitle="A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from coastal southern Vietnam" masterLastPageNumber="226" masterPageNumber="211" pageNumber="212" updateTime="1698661651792" updateUser="ExternalLinkService">
<mods:mods id="5ECA8C9433C5D9E4BCABE307073B78CB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A084D9373CA51505CB768BF6912D227F">
<mods:title id="41641B505B302D8EDE3B67593C79C0DB">A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from coastal southern Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="431E0BD3F3D79686AC187A7DB791DC54" type="personal">
<mods:role id="BC9CD741C46058FB4C8AD5743F4D2437">
<mods:roleTerm id="243C74A7AA5138A2434BAC020C6345D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="569B07A27349EED5EE0D09509965AD5B">Vassilieva, Anna B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="8C62007188C148D7031EA198F0549AC3">text</mods:typeOfResource>
<mods:relatedItem id="94B82F7D3DF7E299E740AB5C383764A3" type="host">
<mods:titleInfo id="F5408C0BE3EB6B66603C97DEAC36FBF6">
<mods:title id="03D5FFDA8CB5DE3490FEF8DAFF2BC362">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A96523EE94B2DB863B41DFCFEFB3951B">
<mods:date id="40A27CD89D345F10FA12894F7A1D62C7">2015</mods:date>
<mods:detail id="EE5F6F1CA1B85F9887C1351FA6D632E2" type="volume">
<mods:number id="E4DCD6A935DC83B328F7A2236D295F7A">4058</mods:number>
</mods:detail>
<mods:detail id="3E53E66BC78AFA680609C3578366E260" type="issue">
<mods:number id="7E80B770F61CB2D892CFE9623ED82B05">2</mods:number>
</mods:detail>
<mods:extent id="560AB936AC63A2682488CA7AD71872B9" unit="page">
<mods:start id="F5EF0705B0DE85167C8E889E799AC3FD">211</mods:start>
<mods:end id="1C6F8077CC8887F65DF814669B93F20D">226</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="080765997E0799FF3B5810F7E1908DF2">journal article</mods:classification>
<mods:identifier id="A88221386C2FB3A1A938ABC7C54B7EA6" type="CLB-Dataset">39259</mods:identifier>
<mods:identifier id="B92FDE47700D41D08913B149DBB54222" type="DOI">10.11646/zootaxa.4058.2.4</mods:identifier>
<mods:identifier id="EAC07F791C03057ACFD485B73E2B056D" type="GBIF-Dataset">b6aeffc9-06f4-4e5d-bd0e-cf480b5e06d4</mods:identifier>
<mods:identifier id="E65E927C1AA0249B622C8F7B2351B1EE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BEFFE2689D36DE5CC0450F012517C6A5" type="Zenodo-Dep">253690</mods:identifier>
<mods:identifier id="59085F5057674FF7219555F10AC75414" type="ZooBank">1FDCAFAB-1E21-43A7-9543-E9071EA48FA7</mods:identifier>
</mods:mods>
<treatment id="E3129514090DFF974093FB24FBBEFB0A" ID-DOI="http://doi.org/10.5281/zenodo.6103018" ID-GBIF-Taxon="119647833" ID-Zenodo-Dep="6103018" LSID="urn:lsid:plazi:treatment:E3129514090DFF974093FB24FBBEFB0A" httpUri="http://treatment.plazi.org/id/E3129514090DFF974093FB24FBBEFB0A" lastPageId="12" lastPageNumber="222" pageId="1" pageNumber="212">
<subSubSection id="23A17789090DFF9A4093FB24FE20FAA1" box="[151,489,1265,1291]" pageId="1" pageNumber="212" type="nomenclature">
<paragraph id="6B042402090DFF9A4093FB24FE20FAA1" blockId="1.[151,489,1265,1291]" box="[151,489,1265,1291]" pageId="1" pageNumber="212">
<heading id="304C936E090DFF9A4093FB24FE20FAA1" bold="true" box="[151,489,1265,1291]" fontSize="11" level="1" pageId="1" pageNumber="212" reason="1">
<emphasis id="59CFF810090DFF9A4093FB24FE20FAA1" bold="true" box="[151,489,1265,1291]" pageId="1" pageNumber="212">
<taxonomicName id="ACBB5F81090DFF9A4093FB24FE4AFAA1" box="[151,387,1265,1291]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="212" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF810090DFF9A4093FB24FE4AFAA1" bold="true" box="[151,387,1265,1291]" italics="true" pageId="1" pageNumber="212">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B090DFF9A418FFB24FE20FAA1" box="[395,489,1265,1291]" pageId="1" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="23A17789090DFF9A4093FAE2FB75F912" pageId="1" pageNumber="212" type="materials_examined">
<paragraph id="6B042402090DFF9A4093FAE2FB4DFA32" blockId="1.[151,1437,1335,2009]" pageId="1" pageNumber="212">
<typeStatus id="B4009AA0090DFF9A4093FAE2FECDFAFA" box="[151,260,1335,1360]" pageId="1" pageNumber="212" type="holotype">
<emphasis id="59CFF810090DFF9A4093FAE2FECDFAFA" bold="true" box="[151,260,1335,1360]" pageId="1" pageNumber="212">Holotype</emphasis>
</typeStatus>
.
<collectionCode id="0DAABCC7090DFF9A4113FAEDFEA7FAE5" box="[279,366,1336,1359]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="212">ZMMU</collectionCode>
<collectionCode id="0DAABCC7090DFF9A417FFAE2FE5EFAFA" box="[379,407,1335,1360]" pageId="1" pageNumber="212">R-</collectionCode>
14503 (field
<collectionCode id="0DAABCC7090DFF9A4236FAEDFD86FAE5" box="[562,591,1336,1359]" pageId="1" pageNumber="212">ID</collectionCode>
<collectionCode id="0DAABCC7090DFF9A4258FAEDFD7AFAE5" box="[604,691,1336,1359]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="212">ZMMU</collectionCode>
<collectionCode id="0DAABCC7090DFF9A42C4FAEDFD34FAFA" box="[704,765,1336,1360]" pageId="1" pageNumber="212">ABV</collectionCode>
00813), adult male from Binh ChauPhuoc Buu Nature Reserve, Xuen Moc District, Ba RiaVung
<collectingCountry id="13AC6492090DFF9A429CFA88FD0BFADE" box="[664,706,1373,1396]" name="China" pageId="1" pageNumber="212">Tau</collectingCountry>
Province, southern
<collectingCountry id="13AC6492090DFF9A43B4FA89FBC6FADF" box="[944,1039,1372,1397]" name="Vietnam" pageId="1" pageNumber="212">Vietnam</collectingCountry>
(
<figureCitation id="F3803887090DFF9A4426FA89FBA1FADE" box="[1058,1128,1372,1397]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[151,1413,445,1844]" captionTargetId="figure@2.[151,1436,445,1897]" captionTargetPageId="2" captionText="FIGURE 1. The type locality and known distribution of Oligodon arenarius sp. nov. (marked by red triangle) in Binh Chau Phuoc Buu Nature Reserve, southern Vietnam." httpUri="https://zenodo.org/record/253691/files/figure.png" pageId="1" pageNumber="212">Fig. 1</figureCitation>
), coordinates
<geoCoordinate id="0E8F42C5090DFF9A4515FA89FA51FADF" box="[1297,1432,1372,1397]" direction="north" orientation="latitude" pageId="1" pageNumber="212" precision="15" value="10.496111">10°29'46&quot;N</geoCoordinate>
,
<geoCoordinate id="0E8F42C5090DFF9A4093FAAAFEE1FA32" box="[151,296,1407,1432]" direction="east" orientation="longitude" pageId="1" pageNumber="212" precision="15" value="107.465">107°27'54&quot;E</geoCoordinate>
, elevation
<quantity id="AC4389E7090DFF9A41A5FAAAFE05FA3D" box="[417,460,1407,1432]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="212" unit="m" value="5.0">5 m</quantity>
a.s.l., collected by Anna B. Vassilieva on
<date id="1F0502C2090DFF9A43A0FA55FB48FA32" box="[932,1153,1407,1432]" pageId="1" pageNumber="212" value="2014-11-14">14 November 2014</date>
.
</paragraph>
<paragraph id="6B042402090DFF9A40C3FA71FB75F912" blockId="1.[151,1437,1335,2009]" pageId="1" pageNumber="212">
<typeStatus id="B4009AA0090DFF9A40C3FA71FEF6FA17" box="[199,319,1444,1469]" pageId="1" pageNumber="212" type="paratype">
<emphasis id="59CFF810090DFF9A40C3FA71FEF6FA17" bold="true" box="[199,319,1444,1469]" pageId="1" pageNumber="212">Paratypes</emphasis>
</typeStatus>
.
<collectionCode id="0DAABCC7090DFF9A4149FA70FE6DFA16" box="[333,420,1445,1468]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="212">ZMMU</collectionCode>
<collectionCode id="0DAABCC7090DFF9A41AFFA71FE0EFA16" box="[427,455,1444,1468]" pageId="1" pageNumber="212">R-</collectionCode>
14002 (field
<collectionCode id="0DAABCC7090DFF9A425CFA70FDBCFA16" box="[600,629,1445,1468]" pageId="1" pageNumber="212">ID</collectionCode>
<collectionCode id="0DAABCC7090DFF9A4279FA70FD1DFA16" box="[637,724,1445,1468]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="212">ZMMU</collectionCode>
<collectionCode id="0DAABCC7090DFF9A42DFFA71FCD6FA16" box="[731,799,1444,1468]" pageId="1" pageNumber="212">NAP-</collectionCode>
03137), adult male from
<typeStatus id="B4009AA0090DFF9A4431FA70FBACFA17" box="[1077,1125,1445,1469]" pageId="1" pageNumber="212">type</typeStatus>
locality: Binh ChauPhuoc Buu Nature Reserve, Xuen Moc Disctrict, Ba RiaVung
<collectingCountry id="13AC6492090DFF9A4368FA1DFC5FFA75" box="[876,918,1480,1503]" name="China" pageId="1" pageNumber="212">Tau</collectingCountry>
Province, southern
<collectingCountry id="13AC6492090DFF9A4493FA12FB3FFA4A" box="[1175,1270,1479,1504]" name="Vietnam" pageId="1" pageNumber="212">Vietnam</collectingCountry>
(
<geoCoordinate id="0E8F42C5090DFF9A4515FA1DFA5EFA4A" box="[1297,1431,1480,1504]" direction="north" orientation="latitude" pageId="1" pageNumber="212" precision="15" value="10.538334">10°32'18&quot;N</geoCoordinate>
,
<geoCoordinate id="0E8F42C5090DFF9A4093FA39FEE1F9AE" box="[151,296,1516,1540]" direction="east" orientation="longitude" pageId="1" pageNumber="212" precision="15" value="107.476944">107°28'37&quot;E</geoCoordinate>
,
<quantity id="AC4389E7090DFF9A413CFA39FEBFF9AE" box="[312,374,1516,1540]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="212" unit="m" value="43.0">43 m</quantity>
a.s.l.), collected on
<date id="1F0502C2090DFF9A426FFA39FCF6F9AE" box="[619,831,1516,1541]" pageId="1" pageNumber="212" value="2010-11-08">8 November 2010</date>
by Anna B. Vassilieva;
<collectionCode id="0DAABCC7090DFF9A4461FA38FB75F9AE" box="[1125,1212,1517,1540]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="212">ZMMU</collectionCode>
<collectionCode id="0DAABCC7090DFF9A44CCFA39FB2DF9AE" box="[1224,1252,1516,1540]" pageId="1" pageNumber="212">R-</collectionCode>
14002 (field
<collectionCode id="0DAABCC7090DFF9A457BFA38FA55F9AE" box="[1407,1436,1517,1540]" pageId="1" pageNumber="212">ID</collectionCode>
<collectionCode id="0DAABCC7090DFF9A4093F9C5FF27F98D" box="[151,238,1552,1575]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="212">ZMMU</collectionCode>
<collectionCode id="0DAABCC7090DFF9A40F8F9C5FE89F982" box="[252,320,1552,1576]" pageId="1" pageNumber="212">NAP-</collectionCode>
03138), adult female from
<typeStatus id="B4009AA0090DFF9A4280F9C5FD7DF982" box="[644,692,1552,1576]" pageId="1" pageNumber="212">type</typeStatus>
locality (
<geoCoordinate id="0E8F42C5090DFF9A4328F9DAFC7AF982" box="[812,947,1551,1576]" direction="north" orientation="latitude" pageId="1" pageNumber="212" precision="15" value="10.497223">10°29'50&quot;N</geoCoordinate>
,
<geoCoordinate id="0E8F42C5090DFF9A43C1F9DAFB9FF982" box="[965,1110,1551,1576]" direction="east" orientation="longitude" pageId="1" pageNumber="212" precision="15" value="107.46611">107°27'58&quot;E</geoCoordinate>
,
<quantity id="AC4389E7090DFF9A446CF9DAFB53F98D" box="[1128,1178,1551,1576]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="212" unit="m" value="5.0">5 m</quantity>
a.s.l.) collected on
<date id="1F0502C2090DFF9A458AF9C5FE86F9E6" pageId="1" pageNumber="212" value="2010-11-09">9 November 2010</date>
by Anna B. Vassilieva;
<collectionCode id="0DAABCC7090DFF9A426AF9E0FD0DF9E6" box="[622,708,1589,1612]" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="1" pageNumber="212">VNMN</collectionCode>
0 4724 (field
<collectionCode id="0DAABCC7090DFF9A4361F9E0FC18F9E6" box="[869,977,1588,1612]" pageId="1" pageNumber="212">ID NAP-</collectionCode>
03884), adult female from
<typeStatus id="B4009AA0090DFF9A450DF9E0FAF0F9E7" box="[1289,1337,1589,1613]" pageId="1" pageNumber="212">type</typeStatus>
locality, environs of the resort Ho Coc (
<geoCoordinate id="0E8F42C5090DFF9A41F6F982FDB0F9DA" box="[498,633,1623,1648]" direction="north" orientation="latitude" pageId="1" pageNumber="212" precision="15" value="10.498334">10°29'54&quot;N</geoCoordinate>
,
<geoCoordinate id="0E8F42C5090DFF9A4281F98DFCDEF9DA" box="[645,791,1624,1648]" direction="east" orientation="longitude" pageId="1" pageNumber="212" precision="15" value="107.46889">107°28'08&quot;E</geoCoordinate>
;
<quantity id="AC4389E7090DFF9A4326F98DFC84F9C5" box="[802,845,1624,1648]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="212" unit="m" value="8.0">8 m</quantity>
a.s.l.), captured on
<date id="1F0502C2090DFF9A442EF98DFB72F9DA" box="[1066,1211,1623,1648]" pageId="1" pageNumber="212" value="2012-07-10">10 July 2012</date>
by local people and collected by Anna B. Vassilieva and Nikolay Poyarkov;
<collectionCode id="0DAABCC7090DFF9A434CF9A8FC56F93E" box="[840,927,1661,1684]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="212">ZMMU</collectionCode>
<collectionCode id="0DAABCC7090DFF9A43A9F9A9FC03F93F" box="[941,970,1660,1685]" pageId="1" pageNumber="212">R-</collectionCode>
14504, adult female from
<typeStatus id="B4009AA0090DFF9A450EF9A8FAF3F93F" box="[1290,1338,1661,1685]" pageId="1" pageNumber="212">type</typeStatus>
locality (
<geoCoordinate id="0E8F42C5090DFF9A409BF975FEEFF912" box="[159,294,1696,1720]" direction="north" orientation="latitude" pageId="1" pageNumber="212" precision="15" value="10.494722">10°29'41&quot;N</geoCoordinate>
,
<geoCoordinate id="0E8F42C5090DFF9A4136F94AFE0DF912" box="[306,452,1695,1720]" direction="east" orientation="longitude" pageId="1" pageNumber="212" precision="15" value="107.462776">107°27'46&quot;E</geoCoordinate>
;
<quantity id="AC4389E7090DFF9A41CBF975FE33F91D" box="[463,506,1696,1719]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="212" unit="m" value="7.0">7 m</quantity>
a.s.l.), collected on
<date id="1F0502C2090DFF9A42D8F94AFC7DF912" box="[732,948,1695,1720]" pageId="1" pageNumber="212" value="2014-11-16">16 November 2014</date>
by Anna B. Vassilieva.
</paragraph>
</subSubSection>
<subSubSection id="23A17789090DFF9940C3F911FD46FE22" lastPageId="2" lastPageNumber="213" pageId="1" pageNumber="212" type="diagnosis">
<paragraph id="6B042402090DFF9A40C3F911FD6CF83A" blockId="1.[151,1437,1335,2009]" pageId="1" pageNumber="212">
<emphasis id="59CFF810090DFF9A40C3F911FE88F977" bold="true" box="[199,321,1732,1757]" pageId="1" pageNumber="212">Diagnosis.</emphasis>
The species is allocated to the genus
<taxonomicName id="ACBB5F81090DFF9A42E9F911FC9EF976" box="[749,855,1732,1756]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="212" phylum="Chordata" rank="genus">
<emphasis id="59CFF810090DFF9A42E9F911FC9EF976" box="[749,855,1732,1756]" italics="true" pageId="1" pageNumber="212">Oligodon</emphasis>
</taxonomicName>
based on the following characters considered to be diagnostic for the genus (
<bibRefCitation id="0F2A59F3090DFF9A41B1F932FD8CF8AA" author="Smith" box="[437,581,1767,1792]" pageId="1" pageNumber="212" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo - chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp." type="book" year="1943">Smith, 1943</bibRefCitation>
): posterior maxillary teeth strongly enlarged and compressed; palatine teeth well developed or vestigial; head short, not distinct from neck, usually displaying a specific coloration pattern; head scalation complete or reduced; eye moderate, with rounded pupil; rostral scale enlarged, extending on to the upper surface of the snout and partly separating the internasals; body cylindrical; scales smooth; ventrals rounded or obtusely keeled laterally; subcaudals paired.
</paragraph>
<paragraph id="6B042402090DFF9940C3F849FD46FE22" blockId="1.[151,1437,1335,2009]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="213" pageId="1" pageNumber="212">
The new species is distinguishable from other species of the genus by a combination of the following features: (1) medium size in adults (TL
<quantity id="AC4389E7090DFF9A41FBF815FD52F872" box="[511,667,1984,2009]" metricMagnitude="-1" metricUnit="m" metricValue="3.625" metricValueMax="3.8899999999999997" metricValueMin="3.36" pageId="1" pageNumber="212" unit="mm" value="362.5" valueMax="389.0" valueMin="336.0">336389 mm</quantity>
); (2) 17(19)1715 dorsal scale rows; (3) 68 maxillary teeth, the posterior three being enlarged; (4) head scalation lacking a loreal but including a presubocular (except one specimen), (5) nasal divided; (6) two postoculars; (7) 131
<emphasis id="59CFF810090EFF994321FF68FCFAFF7E" box="[805,819,189,212]" italics="true" pageId="2" pageNumber="213"></emphasis>
144 ventrals; (8) 3660 subcaudals (
<quantity id="AC4389E7090EFF9944D4FF69FAFFFF7F" box="[1232,1334,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" metricValueMax="10.16" metricValueMin="9.144" pageId="2" pageNumber="213" unit="in" value="38.0" valueMax="40.0" valueMin="36.0">3640 in</quantity>
females,
<quantity id="AC4389E7090EFF994093FF0AFF36FF52" box="[151,255,223,248]" metricMagnitude="0" metricUnit="m" metricValue="1.4986" metricValueMax="1.524" metricValueMin="1.4731999999999998" pageId="2" pageNumber="213" unit="in" value="59.0" valueMax="60.0" valueMin="58.0">5860 in</quantity>
males); (9) anal plate entire; (10) tail quite long for the genus in males (TaL/TL = 0.260.28) and moderate in females (TaL/TL = 0.130.17); (11) unforked hemipenes without spines; (12) head coloration pattern including ocular band, temporal bands and chevron-shaped mark on nape; (13) dorsal coloration without specific pattern, uniform or with dark speckling; (14) ventrals pinkish in life, immaculate; (15) iris bicolored with silvery upper third and dark brown lower two thirds.
</paragraph>
</subSubSection>
<caption id="3FC4748A090EFF994093F8ABFDB8F818" httpUri="https://zenodo.org/record/253691/files/figure.png" pageId="2" pageNumber="213" targetBox="[151,1413,445,1844]" targetPageId="2">
<paragraph id="6B042402090EFF994093F8ABFDB8F818" blockId="2.[151,1436,1918,1970]" pageId="2" pageNumber="213">
<emphasis id="59CFF810090EFF994093F8ABFEDAF839" bold="true" box="[151,275,1918,1940]" pageId="2" pageNumber="213">FIGURE 1.</emphasis>
The type locality and known distribution of
<taxonomicName id="ACBB5F81090EFF9942DAF8ABFC6EF83E" box="[734,935,1918,1940]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="213" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF810090EFF9942DAF8ABFC6EF83E" box="[734,935,1918,1940]" italics="true" pageId="2" pageNumber="213">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B090EFF9943AAF8ABFC36F839" box="[942,1023,1918,1939]" pageId="2" pageNumber="213" rank="species">
<emphasis id="59CFF810090EFF9943AAF8ABFC36F839" bold="true" box="[942,1023,1918,1939]" pageId="2" pageNumber="213">sp. nov.</emphasis>
</taxonomicNameLabel>
(marked by red triangle) in Binh Chau Phuoc Buu Nature Reserve, southern Vietnam.
</paragraph>
</caption>
<caption id="3FC4748A090FFF984093FD0EFDE3FCA5" httpUri="https://zenodo.org/record/253692/files/figure.png" pageId="3" pageNumber="214" targetBox="[151,1436,193,709]" targetPageId="3">
<paragraph id="6B042402090FFF984093FD0EFDE3FCA5" blockId="3.[151,1436,731,783]" pageId="3" pageNumber="214">
<emphasis id="59CFF810090FFF984093FD0EFEDAFD5A" bold="true" box="[151,275,731,753]" pageId="3" pageNumber="214">FIGURE 2.</emphasis>
Holotype
<taxonomicName id="ACBB5F81090FFF984185FD0EFD80FD5B" box="[385,585,731,753]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="214" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF810090FFF984185FD0EFD80FD5B" box="[385,585,731,753]" italics="true" pageId="3" pageNumber="214">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B090FFF984254FD0EFD6BFD5A" box="[592,674,731,752]" pageId="3" pageNumber="214" rank="species">
<emphasis id="59CFF810090FFF984254FD0EFD6BFD5A" bold="true" box="[592,674,731,752]" pageId="3" pageNumber="214">sp. nov.</emphasis>
</taxonomicNameLabel>
(ZMMU R-14503, field ID ABV-00813) in preservative; (A) dorsal view, (B) ventral view. Photos by V. Trounov.
</paragraph>
</caption>
<subSubSection id="23A17789090FFF9240C3FCE2FD62FD02" lastPageId="9" lastPageNumber="220" pageId="3" pageNumber="214" type="description">
<paragraph id="6B042402090FFF9840C3FCE2FBF9FCE5" blockId="3.[151,1437,823,2036]" box="[199,1072,823,848]" pageId="3" pageNumber="214">
<emphasis id="59CFF810090FFF9840C3FCE2FE2DFCFA" bold="true" box="[199,484,823,848]" pageId="3" pageNumber="214">
Description of
<typeStatus id="B4009AA0090FFF984172FCE2FE16FCFA" box="[374,479,823,848]" pageId="3" pageNumber="214" type="holotype">holotype</typeStatus>
.
</emphasis>
Main morphometric characters are given in
<tableCitation id="263911B9090FFF9843D2FCE2FBE3FCE5" box="[982,1066,823,848]" captionStart="TABLE 1" captionStartId="6.[119,206,594,616]" captionText="TABLE 1. Pholidosis (A) and other morphological features (B) of the type series of Oligodon arenarius sp. nov." pageId="3" pageNumber="214">Table 1</tableCitation>
.
</paragraph>
<paragraph id="6B042402090FFF9840C3FC8EFEF2FB82" blockId="3.[151,1437,823,2036]" pageId="3" pageNumber="214">
Body (
<figureCitation id="F3803887090FFF984110FC8EFE91FCD9" box="[276,344,859,884]" captionStart="FIGURE 2" captionStartId="3.[151,250,731,753]" captionTargetBox="[151,1436,193,709]" captionTargetId="figure@3.[151,1436,193,709]" captionTargetPageId="3" captionText="FIGURE 2. Holotype Oligodon arenarius sp. nov. (ZMMU R- 14503, field ID ABV- 00813) in preservative; (A) dorsal view, (B) ventral view. Photos by V. Trounov." httpUri="https://zenodo.org/record/253692/files/figure.png" pageId="3" pageNumber="214">Fig. 2</figureCitation>
) cylindrical, rather robust, tail long for the genus (TaL/TL = 0.26), slender, with acuminated tip; shallow artificial groove caused by the temporary (removed) ligature is seen at the base of tail; head (
<figureCitation id="F3803887090FFF984514FCAAFA9AFC3D" box="[1296,1363,895,920]" captionStart="FIGURE 3" captionStartId="4.[151,250,1386,1408]" captionTargetBox="[184,1403,211,1354]" captionTargetId="figure@4.[168,1419,193,1364]" captionTargetPageId="4" captionText="FIGURE 3. Holotype Oligodon arenarius sp. nov. head view (in preservative) and head scalation; (A) lateral view, (B) dorsal view, and (C) ventral view. Photos and drawings by V. Trounov." httpUri="https://zenodo.org/record/253693/files/figure.png" pageId="3" pageNumber="214">Fig. 3</figureCitation>
) short (HL/SVL = 0.04), ovoid in dorsal view, somewhat triangular in profile view, faintly distinct from neck, somewhat depressed dorsoventrally; snout narrowed, triangle with blunt tip; snout tip strongly extending beyond the lower jaw; large oval nostril piercing the central part of nasal; eye rather small, approximately 0.17 times the head length; pupil rounded.
</paragraph>
<paragraph id="6B042402090FFF9840C3FBE1FE7AFB3E" blockId="3.[151,1437,823,2036]" pageId="3" pageNumber="214">
<emphasis id="59CFF810090FFF9840C3FBE1FEF8FBE1" box="[199,305,1076,1099]" italics="true" pageId="3" pageNumber="214">Dentition</emphasis>
. Maxillary teeth: 8 functional teeth on the right and 7 on the left side, the posterior three teeth being strongly enlarged, slightly curved posteriorly, dagger-shaped; no distinct gap before the enlarged teeth. Palatine dentition well developed.
</paragraph>
<paragraph id="6B042402090FFF9840C3FB4AFDF3FB55" blockId="3.[151,1437,823,2036]" pageId="3" pageNumber="214">
<emphasis id="59CFF810090FFF9840C3FB4AFEBBFB1D" box="[199,370,1183,1207]" italics="true" pageId="3" pageNumber="214">Body scalation</emphasis>
. Dorsal scale rows 17
<emphasis id="59CFF810090FFF98427CFB75FD48FB1D" box="[632,641,1184,1207]" italics="true" pageId="3" pageNumber="214">-</emphasis>
17
<emphasis id="59CFF810090FFF984299FB75FD6FFB1D" box="[669,678,1184,1207]" italics="true" pageId="3" pageNumber="214">-</emphasis>
15, all smooth; outermost dorsal scale row slightly enlarged; 134 ventrals, obtusely keeled laterally; preventrals absent; anal plate entire; 58 subcaudals, all paired; terminal caudal scale forming an acuminated tail cap.
</paragraph>
<paragraph id="6B042402090FFF9840C3FADEFA54F85E" blockId="3.[151,1437,823,2036]" pageId="3" pageNumber="214">
<emphasis id="59CFF810090FFF9840C3FADEFEBBFA89" box="[199,370,1291,1315]" italics="true" pageId="3" pageNumber="214">Head scalation</emphasis>
. In lateral view (
<figureCitation id="F3803887090FFF98423CFADEFDB2FA89" box="[568,635,1291,1316]" captionStart="FIGURE 3" captionStartId="4.[151,250,1386,1408]" captionTargetBox="[184,1403,211,1354]" captionTargetId="figure@4.[168,1419,193,1364]" captionTargetPageId="4" captionText="FIGURE 3. Holotype Oligodon arenarius sp. nov. head view (in preservative) and head scalation; (A) lateral view, (B) dorsal view, and (C) ventral view. Photos and drawings by V. Trounov." httpUri="https://zenodo.org/record/253693/files/figure.png" pageId="3" pageNumber="214">Fig. 3</figureCitation>
A), head scalation comprising a subrectangular nasal, pierced by large nostril and divided almost vertically by a prominent suture running through the posterior half of nostril; single rectangular preocular, slightly higher than wide, separated from nasal by the lateral part of the prefrontal; single small subrectangular presubocular; two small postoculars, almost equal in size; eight supralabials: dorsally, I in contact with nasal, II in contact with nasal and prefrontal, III in contact with prefrontal and presubocular, IV in contact with presubocular and eye, V in contact with eye, VI in contact with lower postocular and anterior temporal, the largest VII in contact with anterior temporal, VIII in contact with lower posterior temporal; 1+2 temporals, the anterior one subrectangular, elongated and narrow, the upper posterior smoothly pentagonal, approximately equal to the anterior one, elongated, the lower posterior the smallest, rhomboid. In dorsal view (
<figureCitation id="F3803887090FFF98456BF9FEFF6FF9CD" captionStart="FIGURE 2" captionStartId="3.[151,250,731,753]" captionTargetBox="[151,1436,193,709]" captionTargetId="figure@3.[151,1436,193,709]" captionTargetPageId="3" captionText="FIGURE 2. Holotype Oligodon arenarius sp. nov. (ZMMU R- 14503, field ID ABV- 00813) in preservative; (A) dorsal view, (B) ventral view. Photos by V. Trounov." httpUri="https://zenodo.org/record/253692/files/figure.png" pageId="3" pageNumber="214">Fig. 2</figureCitation>
B), head scalation comprising a rostral, two internasals, two prefrontals, two supraoculars, a frontal, and two parietals. Rostral large, wider than high, extending on to the dorsal surface of the snout and inserting deeply between internasals. Internasals wider than long, narrowing and slightly curving back towards the median suture. Prefrontals large, pentagonal, approximately 1.5 times wider than internasals, curving down laterally to the loreal area; lateral portions of prefrontals contacting posteriorly with the upper preocular and ventrally with the II
<emphasis id="59CFF810090FFF984576F935FA49F95D" box="[1394,1408,1760,1783]" italics="true" pageId="3" pageNumber="214"></emphasis>
III supralabials. Supraoculars large, elongated, widening posteriorly, approximately half as wide as long. Frontal large, smoothly hexagonal, slightly longer than wide, posterior angle rather acute. Parietals irregularly pentagonal, slightly longer and wider than frontal, bordered laterally by the first and the upper second temporals and posteriorly by four small scales; no enlarged nuchal scales present. In ventral view (
<figureCitation id="F3803887090FFF9843C8F8BAFBD8F82D" box="[972,1041,1903,1928]" captionStart="FIGURE 2" captionStartId="3.[151,250,731,753]" captionTargetBox="[151,1436,193,709]" captionTargetId="figure@3.[151,1436,193,709]" captionTargetPageId="3" captionText="FIGURE 2. Holotype Oligodon arenarius sp. nov. (ZMMU R- 14503, field ID ABV- 00813) in preservative; (A) dorsal view, (B) ventral view. Photos by V. Trounov." httpUri="https://zenodo.org/record/253692/files/figure.png" pageId="3" pageNumber="214">Fig. 2</figureCitation>
C), nine infralabials on the left and seven on the right side, III+IV and VII+VIII being fused on the right side; I medially in contact with each other and rostrally with mental, II
<emphasis id="59CFF810090FFF9841ABF86DFE74F865" box="[431,445,1976,1999]" italics="true" pageId="3" pageNumber="214"></emphasis>
V in contact with two enlarged chin shields; mental small, triangular; two enlarged, elongated chin shields, the anterior pair being approximately twice longer than the posterior pair along the midline.
</paragraph>
<caption id="3FC4748A0908FF9F4093FABFFCD6FA34" httpUri="https://zenodo.org/record/253693/files/figure.png" pageId="4" pageNumber="215" targetBox="[184,1403,211,1354]" targetPageId="4">
<paragraph id="6B0424020908FF9F4093FABFFCD6FA34" blockId="4.[151,1436,1386,1438]" pageId="4" pageNumber="215">
<emphasis id="59CFF8100908FF9F4093FABFFEDAFAD5" bold="true" box="[151,275,1386,1408]" pageId="4" pageNumber="215">FIGURE 3.</emphasis>
Holotype
<taxonomicName id="ACBB5F810908FF9F417AFABFFD8FFA2A" box="[382,582,1386,1408]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="215" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100908FF9F417AFABFFD8FFA2A" box="[382,582,1386,1408]" italics="true" pageId="4" pageNumber="215">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0908FF9F4248FABFFD54FAD5" box="[588,669,1386,1407]" pageId="4" pageNumber="215" rank="species">
<emphasis id="59CFF8100908FF9F4248FABFFD54FAD5" bold="true" box="[588,669,1386,1407]" pageId="4" pageNumber="215">sp. nov.</emphasis>
</taxonomicNameLabel>
head view (in preservative) and head scalation; (A) lateral view, (B) dorsal view, and (C) ventral view. Photos and drawings by V. Trounov.
</paragraph>
</caption>
<caption id="3FC4748A0908FF9F4093F847FE46F86D" httpUri="https://zenodo.org/record/253694/files/figure.png" pageId="4" pageNumber="215" targetBox="[151,1436,1488,1917]" targetPageId="4">
<paragraph id="6B0424020908FF9F4093F847FE46F86D" blockId="4.[151,1437,1938,1991]" pageId="4" pageNumber="215">
<emphasis id="59CFF8100908FF9F4093F847FEDCF80D" bold="true" box="[151,277,1938,1960]" pageId="4" pageNumber="215">FIGURE 4.</emphasis>
Hemipenis of the holotype
<taxonomicName id="ACBB5F810908FF9F423AF847FCC1F802" box="[574,776,1938,1960]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="215" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100908FF9F423AF847FCC1F802" box="[574,776,1938,1960]" italics="true" pageId="4" pageNumber="215">Oligodon arenarius</emphasis>
</taxonomicName>
<emphasis id="59CFF8100908FF9F4316F846FCA5F802" bold="true" box="[786,876,1939,1960]" pageId="4" pageNumber="215">
<taxonomicNameLabel id="42FC456B0908FF9F4316F846FCAEF802" box="[786,871,1939,1960]" pageId="4" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
(A) everted, before preservation, (B) in preservative. Photos by A. Vassilieva.
</paragraph>
</caption>
<caption id="3FC4748A0909FF9E4093FC09FD1FFBBB" httpUri="https://zenodo.org/record/253695/files/figure.png" pageId="5" pageNumber="216" targetBox="[151,1436,193,966]" targetPageId="5">
<paragraph id="6B0424020909FF9E4093FC09FD1FFBBB" blockId="5.[151,1436,988,1041]" pageId="5" pageNumber="216">
<emphasis id="59CFF8100909FF9E4093FC09FEDDFC58" bold="true" box="[151,276,988,1010]" pageId="5" pageNumber="216">FIGURE 5.</emphasis>
Life coloration of the holotype
<taxonomicName id="ACBB5F810909FF9E4265FC09FCE3FC58" box="[609,810,988,1010]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="216" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100909FF9E4265FC09FCE3FC58" box="[609,810,988,1010]" italics="true" pageId="5" pageNumber="216">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0909FF9E4336FC08FC4AFC58" box="[818,899,989,1010]" pageId="5" pageNumber="216" rank="species">
<emphasis id="59CFF8100909FF9E4336FC08FC4AFC58" bold="true" box="[818,899,989,1010]" pageId="5" pageNumber="216">sp. nov.</emphasis>
</taxonomicNameLabel>
: (A) total view in situ, (B) portrait view, (C) dorsal coloration, (D) ventral coloration. Photos by V. Trounov.
</paragraph>
</caption>
<paragraph id="6B0424020909FF9E40C3FBEDFF1FFB33" blockId="5.[151,1436,1080,2005]" pageId="5" pageNumber="216">
<emphasis id="59CFF8100909FF9E40C3FBEDFE8EFBFB" bold="true" box="[199,327,1080,1105]" pageId="5" pageNumber="216">Hemipenis</emphasis>
(
<figureCitation id="F38038870909FF9E4151FBEDFE52FBFB" box="[341,411,1080,1105]" captionStart="FIGURE 4" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[151,1436,1488,1917]" captionTargetId="figure@4.[151,1436,1488,1917]" captionTargetPageId="4" captionText="FIGURE 4. Hemipenis of the holotype Oligodon arenarius sp. nov., (A) everted, before preservation, (B) in preservative. Photos by A. Vassilieva." httpUri="https://zenodo.org/record/253694/files/figure.png" pageId="5" pageNumber="216">Fig. 4</figureCitation>
). Hemipenis rather short, the partly everted organ hardly reaching IV subcaudals; unforked, without spines or obvious papillae, proximal part with smooth surface, distal part with flat and smooth circular folds.
</paragraph>
<paragraph id="6B0424020909FF9E40C3FB71FA93F88B" blockId="5.[151,1436,1080,2005]" pageId="5" pageNumber="216">
<emphasis id="59CFF8100909FF9E40C3FB71FE86FB17" bold="true" box="[199,335,1188,1213]" pageId="5" pageNumber="216">Coloration.</emphasis>
In life (
<figureCitation id="F38038870909FF9E41AAFB71FE38FB17" box="[430,497,1188,1213]" captionStart="FIGURE 5" captionStartId="5.[151,250,988,1010]" captionTargetBox="[151,1436,193,966]" captionTargetId="figure@5.[151,1436,193,967]" captionTargetPageId="5" captionText="FIGURE 5. Life coloration of the holotype Oligodon arenarius sp. nov.: (A) total view in situ, (B) portrait view, (C) dorsal coloration, (D) ventral coloration. Photos by V. Trounov." httpUri="https://zenodo.org/record/253695/files/figure.png" pageId="5" pageNumber="216">Fig. 5</figureCitation>
AC), dorsal coloration pale sandy-gray, with slightly more saturated yellowish tint on 4
<emphasis id="59CFF8100909FF9E40C3FB1CFF1CFB4A" box="[199,213,1225,1248]" italics="true" pageId="5" pageNumber="216"></emphasis>
5 upper dorsal and vertebral scale rows; all dorsal scales scattered with minute dark brown spots. Some dorsal scales (except of three lowerest rows on each side) edged by dark brown forming a diffuse speckled or dashed pattern on dorsum. On nape two pairs of irregular dark spots formed by brown-colored halves of 2
<emphasis id="59CFF8100909FF9E450CFAC4FADFFA82" box="[1288,1302,1297,1320]" italics="true" pageId="5" pageNumber="216"></emphasis>
3 scales are most visible along the paravertebral zone. Dark speckling less conspicuous on the caudal part of dorsum. On tail, small brownish speckles fuse into two very faint diffuse lines along two adjacent vertebral scales rows edging a slightly paler vertebral line. Ventrals immaculate; gular and anterior ventral scales creamy-white, belly salmonpink with more intensive coloration caudally. Subcaudals salmon-pink in the anterior part of the tail, gradually whitening toward the tail tip. Head (
<figureCitation id="F38038870909FF9E4244FA11FD4CFA77" box="[576,645,1476,1501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1386,1408]" captionTargetBox="[184,1403,211,1354]" captionTargetId="figure@4.[168,1419,193,1364]" captionTargetPageId="4" captionText="FIGURE 3. Holotype Oligodon arenarius sp. nov. head view (in preservative) and head scalation; (A) lateral view, (B) dorsal view, and (C) ventral view. Photos and drawings by V. Trounov." httpUri="https://zenodo.org/record/253693/files/figure.png" pageId="5" pageNumber="216">Fig. 3</figureCitation>
AB, 5B) ground coloration sandy-gray with yellowish tint on snout; supralabials slightly paler. Darker head pattern include a faintly visible brown crescent-shaped interocular band partly covering the internasals, prefrontals, supraoculars and frontal scales; the anterior margin of the interocular bar diffuse, the posterior rather clear, edged with dark brown; two diffuse brown markings on the posterior part of each supraocular; short brown oblique subocular stripe extending from the lower margin of each orbit towards the mouth and covering parts of V and VI supralabials; brown oblique temporal bars with sharp, dark-edged anterior end and diffuse posterior one, starting close to the anterior margins of parietals and running down towards the neck just behind the jaw angle; large arrow-shaped chevron, brown with clear darker edges, extending from the center of frontal and medial part of parietals to the occiput and nape, reaching the second paraventral scale row on each side laterally. Iris bicolor, with silvery upper third and dark brown lower two thirds; tongue pinkish-red (
<figureCitation id="F38038870909FF9E44F1F8DDFAF0F88B" box="[1269,1337,1800,1825]" captionStart="FIGURE 5" captionStartId="5.[151,250,988,1010]" captionTargetBox="[151,1436,193,966]" captionTargetId="figure@5.[151,1436,193,967]" captionTargetPageId="5" captionText="FIGURE 5. Life coloration of the holotype Oligodon arenarius sp. nov.: (A) total view in situ, (B) portrait view, (C) dorsal coloration, (D) ventral coloration. Photos by V. Trounov." httpUri="https://zenodo.org/record/253695/files/figure.png" pageId="5" pageNumber="216">Fig. 5</figureCitation>
B).
</paragraph>
<paragraph id="6B0424020909FF9E40C3F8F8FD0CF827" blockId="5.[151,1436,1080,2005]" pageId="5" pageNumber="216">
In preservative (
<figureCitation id="F38038870909FF9E4184F8F9FE0AF8EF" box="[384,451,1836,1861]" captionStart="FIGURE 2" captionStartId="3.[151,250,731,753]" captionTargetBox="[151,1436,193,709]" captionTargetId="figure@3.[151,1436,193,709]" captionTargetPageId="3" captionText="FIGURE 2. Holotype Oligodon arenarius sp. nov. (ZMMU R- 14503, field ID ABV- 00813) in preservative; (A) dorsal view, (B) ventral view. Photos by V. Trounov." httpUri="https://zenodo.org/record/253692/files/figure.png" pageId="5" pageNumber="216">Fig. 2</figureCitation>
AB, 3A
<emphasis id="59CFF8100909FF9E421FF8F8FDE0F8EE" box="[539,553,1837,1860]" italics="true" pageId="5" pageNumber="216"></emphasis>
C), all brown and yellow tints fade and turn slightly to gray or cream. Ventral surface yellowish-cream, very faint small grayish spots are visible in the middle of the anterior subcaudals and along the lateral margins of the posterior ventrals.
</paragraph>
<paragraph id="6B0424020909FF9E40C3F84DFCD2F87F" blockId="5.[151,1436,1080,2005]" pageId="5" pageNumber="216">
<emphasis id="59CFF8100909FF9E40C3F84DFCC4F81B" bold="true" box="[199,781,1944,1969]" pageId="5" pageNumber="216">
Description and variation of the
<typeStatus id="B4009AA00909FF9E4254F84DFD75F81B" box="[592,700,1944,1969]" pageId="5" pageNumber="216" type="paratype">paratype</typeStatus>
series.
</emphasis>
The most important characters of the four other available specimens, all considered
<typeStatus id="B4009AA00909FF9E41B9F868FDE2F87F" box="[445,555,1981,2005]" pageId="5" pageNumber="216" type="paratype">paratypes</typeStatus>
, are given in
<tableCitation id="263911B90909FF9E42C5F869FCDCF87F" box="[705,789,1980,2005]" captionStart="TABLE 1" captionStartId="6.[119,206,594,616]" captionText="TABLE 1. Pholidosis (A) and other morphological features (B) of the type series of Oligodon arenarius sp. nov." pageId="5" pageNumber="216">Table 1</tableCitation>
.
</paragraph>
<caption id="3FC4748A090AFF9D4073FD87FAC9FDC2" box="[119,1280,593,616]" pageId="6" pageNumber="217">
<paragraph id="6B042402090AFF9D4073FD87FAC9FDC2" blockId="6.[119,1863,593,1010]" box="[119,1280,593,616]" pageId="6" pageNumber="217">
<emphasis id="59CFF810090AFF9D4073FD87FF2EFDC2" bold="true" box="[119,231,594,616]" pageId="6" pageNumber="217">TABLE 1.</emphasis>
Pholidosis (A) and other morphological features (B) of the type series of
<taxonomicName id="ACBB5F81090AFF9D43DFFD84FB6FFDC2" box="[987,1190,593,616]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="217" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF810090AFF9D43DFFD84FB6FFDC2" box="[987,1190,593,616]" italics="true" pageId="6" pageNumber="217">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B090AFF9D44A8FD87FAC9FDC2" box="[1196,1280,594,616]" pageId="6" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="6B042402090AFF9D409FFDABFF64FD3E" blockId="6.[119,1863,593,1010]" box="[155,173,638,660]" pageId="6" pageNumber="217">A</paragraph>
<paragraph id="6B042402090AFF9D4073FD79F88EFD68" blockId="6.[119,1863,593,1010]" box="[119,1863,684,706]" pageId="6" pageNumber="217">
Specimen ASR
<collectingCountry id="13AC6492090AFF9D4191FD79FE02FD68" box="[405,459,684,706]" name="Egypt" pageId="6" pageNumber="217">MSR</collectingCountry>
PSR SL
<collectingCountry id="13AC6492090AFF9D4290FD79FD65FD68" box="[660,684,684,706]" name="Iceland" pageId="6" pageNumber="217">IL</collectingCountry>
Teeth Nasal Anal PreOc PrSbOc PostOc Temp VEN SC SL contacting eye
</paragraph>
<paragraph id="6B042402090AFF9D4094FD06F961FD43" blockId="6.[119,1863,593,1010]" box="[144,1704,723,745]" pageId="6" pageNumber="217">
14002 17 17
<date id="1F0502C2090AFF9D41EAFD06FD68FD43" box="[494,673,723,745]" pageId="6" pageNumber="217" value="1915-09-07">15 7 9</date>
D E 131 60 IV
</paragraph>
<paragraph id="6B042402090AFF9D42DEFD24FAB5FCAD" blockId="6.[119,1863,593,1010]" box="[730,1404,753,775]" pageId="6" pageNumber="217">7/6 1 1 2/1 1+2</paragraph>
<paragraph id="6B042402090AFF9D4094FCDBF90EFC8E" blockId="6.[119,1863,593,1010]" box="[144,1735,781,804]" pageId="6" pageNumber="217">
14002 19 17
<date id="1F0502C2090AFF9D41EAFCDBFD68FC8E" box="[494,673,782,804]" pageId="6" pageNumber="217" value="1915-09-08">15 8 9</date>
D E 142 40 IV
<emphasis id="59CFF810090AFF9D46ADFCD8F97FFC8E" box="[1705,1718,781,804]" italics="true" pageId="6" pageNumber="217"></emphasis>
V
</paragraph>
<paragraph id="6B042402090AFF9D42DEFCF9FAB5FCE8" blockId="6.[119,1863,593,1010]" box="[730,1404,812,834]" pageId="6" pageNumber="217">8/6 1 1 2 1+2</paragraph>
<paragraph id="6B042402090AFF9D4073FC9CF90EFCF5" blockId="6.[119,1863,593,1010]" box="[119,1735,840,863]" pageId="6" pageNumber="217">
VNMN 0 4724 17 17
<date id="1F0502C2090AFF9D41EAFC9CFD68FCF5" box="[494,673,841,863]" pageId="6" pageNumber="217" value="1915-09-08">15 8 9</date>
D E 143 36 IV
<emphasis id="59CFF810090AFF9D46ADFC9DF97FFCF5" box="[1705,1718,840,863]" italics="true" pageId="6" pageNumber="217"></emphasis>
V
</paragraph>
<paragraph id="6B042402090AFF9D42DEFCB3FAB5FCD6" blockId="6.[119,1863,593,1010]" box="[730,1404,870,892]" pageId="6" pageNumber="217">7/7 1 no 2 1+2</paragraph>
<paragraph id="6B042402090AFF9D4095FC51FAB5FC1D" blockId="6.[119,1863,593,1010]" pageId="6" pageNumber="217">
14503 17 17
<date id="1F0502C2090AFF9D41EAFC51FD56FC30" box="[494,671,900,922]" pageId="6" pageNumber="217" value="1915-08-08">15 8 8</date>
/9 D E 134
<date id="1F0502C2090AFF9D462BFC51F90EFC30" box="[1583,1735,899,922]" pageId="6" pageNumber="217" value="1958-04" valueMax="1958-05-31" valueMin="1958-04-00">
58 IV
<emphasis id="59CFF810090AFF9D46ADFC56F97FFC30" box="[1705,1718,899,922]" italics="true" pageId="6" pageNumber="217"></emphasis>
V
</date>
<typeStatus id="B4009AA0090AFF9D4073FC74FF19FC1D" box="[119,208,929,951]" pageId="6" pageNumber="217" type="holotype">holotype</typeStatus>
8/7 1 1 2 1+2
</paragraph>
<paragraph id="6B042402090AFF9D4095FC6AF90EFC7F" blockId="6.[119,1863,593,1010]" box="[145,1735,958,981]" pageId="6" pageNumber="217">
14504 19 17
<date id="1F0502C2090AFF9D41EAFC6AFD68FC7F" box="[494,673,959,981]" pageId="6" pageNumber="217" value="1915-09-08">15 8 9</date>
D E 144 37 IV
<emphasis id="59CFF810090AFF9D46ADFC6BF97FFC7F" box="[1705,1718,958,981]" italics="true" pageId="6" pageNumber="217"></emphasis>
V
</paragraph>
<paragraph id="6B042402090AFF9D42DEFC09FAB5FC58" blockId="6.[119,1863,593,1010]" box="[730,1404,988,1010]" pageId="6" pageNumber="217">6/7 1 1 2 1+2</paragraph>
<paragraph id="6B042402090AFF9D4073FBA6F903F991" pageId="6" pageNumber="217">
<table id="19BBD6A2090A00644073FBA6F88EF991" box="[119,1863,1139,1595]" gridcols="15" gridrows="6" pageId="6" pageNumber="217">
<tr id="D58B2640090A00644073FBA6F88EFB23" box="[119,1863,1139,1161]" gridrow="0" pageId="6" pageNumber="217">
<th id="965A4F3C090A00644073FBA6FEC5FB23" box="[119,268,1139,1161]" gridcol="0" gridrow="0" pageId="6" pageNumber="217">Specimen</th>
<th id="965A4F3C090A00644132FBA6FEB2FB23" box="[310,379,1139,1161]" gridcol="1" gridrow="0" pageId="6" pageNumber="217">Sex</th>
<th id="965A4F3C090A0064419EFBA6FE1DFB23" box="[410,468,1139,1161]" gridcol="2" gridrow="0" pageId="6" pageNumber="217">TL</th>
<th id="965A4F3C090A006441F2FBA6FDF8FB23" box="[502,561,1139,1161]" gridcol="3" gridrow="0" pageId="6" pageNumber="217">SVL</th>
<th id="965A4F3C090A00644249FBA6FDB2FB23" box="[589,635,1139,1161]" gridcol="4" gridrow="0" pageId="6" pageNumber="217">TaL</th>
<th id="965A4F3C090A006442A0FBA6FD3FFB23" box="[676,758,1139,1161]" gridcol="5" gridrow="0" pageId="6" pageNumber="217">TaL/TL</th>
<th id="965A4F3C090A00644310FBA6FC8BFB23" box="[788,834,1139,1161]" gridcol="6" gridrow="0" pageId="6" pageNumber="217">HW</th>
<th id="965A4F3C090A00644360FBA6FC58FB23" box="[868,913,1139,1161]" gridcol="7" gridrow="0" pageId="6" pageNumber="217">HL</th>
<th id="965A4F3C090A006443BFFBA6FC28FB23" box="[955,993,1139,1161]" gridcol="8" gridrow="0" pageId="6" pageNumber="217">HH</th>
<th id="965A4F3C090A00644405FBA6FBE8FB23" box="[1025,1057,1139,1161]" gridcol="9" gridrow="0" pageId="6" pageNumber="217">IO</th>
<th id="965A4F3C090A00644443FBA6FBA0FB23" box="[1095,1129,1139,1161]" gridcol="10" gridrow="0" pageId="6" pageNumber="217">ED</th>
<th id="965A4F3C090A00644488FBA6FB71FB23" box="[1164,1208,1139,1161]" gridcol="11" gridrow="0" pageId="6" pageNumber="217">SnL</th>
<th id="965A4F3C090A006444D0FBA6FB3FFB23" box="[1236,1270,1139,1161]" gridcol="12" gridrow="0" pageId="6" pageNumber="217">EN</th>
<th id="965A4F3C090A0064451CFBA6FAF1FB23" box="[1304,1336,1139,1161]" gridcol="13" gridrow="0" pageId="6" pageNumber="217">IN</th>
<th id="965A4F3C090A00644559FBA6F88EFB23" box="[1373,1863,1139,1161]" gridcol="14" gridrow="0" pageId="6" pageNumber="217">Dorsal coloration</th>
</tr>
<tr id="D58B2640090A00644073FB65F88EFB6C" box="[119,1863,1200,1222]" gridrow="1" pageId="6" pageNumber="217">
<th id="965A4F3C090A00644073FB65FEC5FB6C" box="[119,268,1200,1222]" gridcol="0" gridrow="1" pageId="6" pageNumber="217">R-14002</th>
<td id="965A4F3C090A00644132FB65FEB2FB6C" box="[310,379,1200,1222]" gridcol="1" gridrow="1" pageId="6" pageNumber="217">male</td>
<td id="965A4F3C090A0064419EFB65FE1DFB6C" box="[410,468,1200,1222]" gridcol="2" gridrow="1" pageId="6" pageNumber="217">355.0</td>
<td id="965A4F3C090A006441F2FB65FDF8FB6C" box="[502,561,1200,1222]" gridcol="3" gridrow="1" pageId="6" pageNumber="217">257.0</td>
<td id="965A4F3C090A00644249FB65FDB2FB6C" box="[589,635,1200,1222]" gridcol="4" gridrow="1" pageId="6" pageNumber="217">98.0</td>
<td id="965A4F3C090A006442A0FB65FD3FFB6C" box="[676,758,1200,1222]" gridcol="5" gridrow="1" pageId="6" pageNumber="217">0.28</td>
<td id="965A4F3C090A00644310FB65FC8BFB6C" box="[788,834,1200,1222]" gridcol="6" gridrow="1" pageId="6" pageNumber="217">8.6</td>
<td id="965A4F3C090A00644360FB65FC58FB6C" box="[868,913,1200,1222]" gridcol="7" gridrow="1" pageId="6" pageNumber="217">10.6</td>
<td id="965A4F3C090A006443BFFB65FC28FB6C" box="[955,993,1200,1222]" gridcol="8" gridrow="1" pageId="6" pageNumber="217">6.9</td>
<td id="965A4F3C090A00644405FB65FBE8FB6C" box="[1025,1057,1200,1222]" gridcol="9" gridrow="1" pageId="6" pageNumber="217">4.7</td>
<td id="965A4F3C090A00644443FB65FBA0FB6C" box="[1095,1129,1200,1222]" gridcol="10" gridrow="1" pageId="6" pageNumber="217">1.8</td>
<td id="965A4F3C090A00644488FB65FB71FB6C" box="[1164,1208,1200,1222]" gridcol="11" gridrow="1" pageId="6" pageNumber="217">4.5</td>
<td id="965A4F3C090A006444D0FB65FB3FFB6C" box="[1236,1270,1200,1222]" gridcol="12" gridrow="1" pageId="6" pageNumber="217">2.1</td>
<td id="965A4F3C090A0064451CFB65FAF1FB6C" box="[1304,1336,1200,1222]" gridcol="13" gridrow="1" pageId="6" pageNumber="217">2.6</td>
<td id="965A4F3C090A00644559FB65F88EFB6C" box="[1373,1863,1200,1222]" gridcol="14" gridrow="1" pageId="6" pageNumber="217">sandy-orange without clear speckling</td>
</tr>
<tr id="D58B2640090A00644073FB39F88EFA89" box="[119,1863,1260,1315]" gridrow="2" pageId="6" pageNumber="217">
<th id="965A4F3C090A00644073FB39FEC5FA89" box="[119,268,1260,1315]" gridcol="0" gridrow="2" pageId="6" pageNumber="217">R-14002</th>
<td id="965A4F3C090A00644132FB39FEB2FA89" box="[310,379,1260,1315]" gridcol="1" gridrow="2" pageId="6" pageNumber="217">female</td>
<td id="965A4F3C090A0064419EFB39FE1DFA89" box="[410,468,1260,1315]" gridcol="2" gridrow="2" pageId="6" pageNumber="217">336.0</td>
<td id="965A4F3C090A006441F2FB39FDF8FA89" box="[502,561,1260,1315]" gridcol="3" gridrow="2" pageId="6" pageNumber="217">286.0</td>
<td id="965A4F3C090A00644249FB39FDB2FA89" box="[589,635,1260,1315]" gridcol="4" gridrow="2" pageId="6" pageNumber="217">50.0</td>
<td id="965A4F3C090A006442A0FB39FD3FFA89" box="[676,758,1260,1315]" gridcol="5" gridrow="2" pageId="6" pageNumber="217">0.15</td>
<td id="965A4F3C090A00644310FB39FC8BFA89" box="[788,834,1260,1315]" gridcol="6" gridrow="2" pageId="6" pageNumber="217">8.1</td>
<td id="965A4F3C090A00644360FB39FC58FA89" box="[868,913,1260,1315]" gridcol="7" gridrow="2" pageId="6" pageNumber="217">10.1</td>
<td id="965A4F3C090A006443BFFB39FC28FA89" box="[955,993,1260,1315]" gridcol="8" gridrow="2" pageId="6" pageNumber="217">6.3</td>
<td id="965A4F3C090A00644405FB39FBE8FA89" box="[1025,1057,1260,1315]" gridcol="9" gridrow="2" pageId="6" pageNumber="217">4.9</td>
<td id="965A4F3C090A00644443FB39FBA0FA89" box="[1095,1129,1260,1315]" gridcol="10" gridrow="2" pageId="6" pageNumber="217">1.9</td>
<td id="965A4F3C090A00644488FB39FB71FA89" box="[1164,1208,1260,1315]" gridcol="11" gridrow="2" pageId="6" pageNumber="217">4.0</td>
<td id="965A4F3C090A006444D0FB39FB3FFA89" box="[1236,1270,1260,1315]" gridcol="12" gridrow="2" pageId="6" pageNumber="217">2.1</td>
<td id="965A4F3C090A0064451CFB39FAF1FA89" box="[1304,1336,1260,1315]" gridcol="13" gridrow="2" pageId="6" pageNumber="217">2.9</td>
<td id="965A4F3C090A00644559FB39F88EFA89" box="[1373,1863,1260,1315]" gridcol="14" gridrow="2" pageId="6" pageNumber="217">saturated orange with evenly distributed black speckling</td>
</tr>
<tr id="D58B2640090A00644073FA9CF88EFA2B" box="[119,1863,1353,1409]" gridrow="3" pageId="6" pageNumber="217">
<th id="965A4F3C090A00644073FA9CFEC5FA2B" box="[119,268,1353,1409]" gridcol="0" gridrow="3" pageId="6" pageNumber="217">VNMN 0 4724</th>
<td id="965A4F3C090A00644132FA9CFEB2FA2B" box="[310,379,1353,1409]" gridcol="1" gridrow="3" pageId="6" pageNumber="217">female</td>
<td id="965A4F3C090A0064419EFA9CFE1DFA2B" box="[410,468,1353,1409]" gridcol="2" gridrow="3" pageId="6" pageNumber="217">372.0</td>
<td id="965A4F3C090A006441F2FA9CFDF8FA2B" box="[502,561,1353,1409]" gridcol="3" gridrow="3" pageId="6" pageNumber="217">318.0</td>
<td id="965A4F3C090A00644249FA9CFDB2FA2B" box="[589,635,1353,1409]" gridcol="4" gridrow="3" pageId="6" pageNumber="217">54.0</td>
<td id="965A4F3C090A006442A0FA9CFD3FFA2B" box="[676,758,1353,1409]" gridcol="5" gridrow="3" pageId="6" pageNumber="217">0.17</td>
<td id="965A4F3C090A00644310FA9CFC8BFA2B" box="[788,834,1353,1409]" gridcol="6" gridrow="3" pageId="6" pageNumber="217">10.1</td>
<td id="965A4F3C090A00644360FA9CFC58FA2B" box="[868,913,1353,1409]" gridcol="7" gridrow="3" pageId="6" pageNumber="217">11.3</td>
<td id="965A4F3C090A006443BFFA9CFC28FA2B" box="[955,993,1353,1409]" gridcol="8" gridrow="3" pageId="6" pageNumber="217">7.9</td>
<td id="965A4F3C090A00644405FA9CFBE8FA2B" box="[1025,1057,1353,1409]" gridcol="9" gridrow="3" pageId="6" pageNumber="217">5.6</td>
<td id="965A4F3C090A00644443FA9CFBA0FA2B" box="[1095,1129,1353,1409]" gridcol="10" gridrow="3" pageId="6" pageNumber="217">1.7</td>
<td id="965A4F3C090A00644488FA9CFB71FA2B" box="[1164,1208,1353,1409]" gridcol="11" gridrow="3" pageId="6" pageNumber="217">4.9</td>
<td id="965A4F3C090A006444D0FA9CFB3FFA2B" box="[1236,1270,1353,1409]" gridcol="12" gridrow="3" pageId="6" pageNumber="217">2.4</td>
<td id="965A4F3C090A0064451CFA9CFAF1FA2B" box="[1304,1336,1353,1409]" gridcol="13" gridrow="3" pageId="6" pageNumber="217">3.8</td>
<td id="965A4F3C090A00644559FA9CF88EFA2B" box="[1373,1863,1353,1409]" gridcol="14" gridrow="3" pageId="6" pageNumber="217">yellowish-gray with evenly distributed black speckling, short paravertebral streaks on neck</td>
</tr>
<tr id="D58B2640090A00644073FA73F88EFA74" box="[119,1863,1446,1502]" gridrow="4" pageId="6" pageNumber="217">
<th id="965A4F3C090A00644073FA73FEC5FA74" box="[119,268,1446,1502]" gridcol="0" gridrow="4" pageId="6" pageNumber="217">R-14503 holotype</th>
<td id="965A4F3C090A00644132FA73FEB2FA74" box="[310,379,1446,1502]" gridcol="1" gridrow="4" pageId="6" pageNumber="217">male</td>
<td id="965A4F3C090A0064419EFA73FE1DFA74" box="[410,468,1446,1502]" gridcol="2" gridrow="4" pageId="6" pageNumber="217">351.0</td>
<td id="965A4F3C090A006441F2FA73FDF8FA74" box="[502,561,1446,1502]" gridcol="3" gridrow="4" pageId="6" pageNumber="217">260.0</td>
<td id="965A4F3C090A00644249FA73FDB2FA74" box="[589,635,1446,1502]" gridcol="4" gridrow="4" pageId="6" pageNumber="217">91.0</td>
<td id="965A4F3C090A006442A0FA73FD3FFA74" box="[676,758,1446,1502]" gridcol="5" gridrow="4" pageId="6" pageNumber="217">0.26</td>
<td id="965A4F3C090A00644310FA73FC8BFA74" box="[788,834,1446,1502]" gridcol="6" gridrow="4" pageId="6" pageNumber="217">8.9</td>
<td id="965A4F3C090A00644360FA73FC58FA74" box="[868,913,1446,1502]" gridcol="7" gridrow="4" pageId="6" pageNumber="217">10.3</td>
<td id="965A4F3C090A006443BFFA73FC28FA74" box="[955,993,1446,1502]" gridcol="8" gridrow="4" pageId="6" pageNumber="217">6.7</td>
<td id="965A4F3C090A00644405FA73FBE8FA74" box="[1025,1057,1446,1502]" gridcol="9" gridrow="4" pageId="6" pageNumber="217">4.8</td>
<td id="965A4F3C090A00644443FA73FBA0FA74" box="[1095,1129,1446,1502]" gridcol="10" gridrow="4" pageId="6" pageNumber="217">1.8</td>
<td id="965A4F3C090A00644488FA73FB71FA74" box="[1164,1208,1446,1502]" gridcol="11" gridrow="4" pageId="6" pageNumber="217">4.1</td>
<td id="965A4F3C090A006444D0FA73FB3FFA74" box="[1236,1270,1446,1502]" gridcol="12" gridrow="4" pageId="6" pageNumber="217">1.9</td>
<td id="965A4F3C090A0064451CFA73FAF1FA74" box="[1304,1336,1446,1502]" gridcol="13" gridrow="4" pageId="6" pageNumber="217">3.1</td>
<td id="965A4F3C090A00644559FA73F88EFA74" box="[1373,1863,1446,1502]" gridcol="14" gridrow="4" pageId="6" pageNumber="217">sandy-gray with faint dark speckling, clearer on neck</td>
</tr>
<tr id="D58B2640090A00644073F9D6F88EF991" box="[119,1863,1539,1595]" gridrow="5" pageId="6" pageNumber="217">
<th id="965A4F3C090A00644073F9D6FEC5F991" box="[119,268,1539,1595]" gridcol="0" gridrow="5" pageId="6" pageNumber="217">R-14504</th>
<td id="965A4F3C090A00644132F9D6FEB2F991" box="[310,379,1539,1595]" gridcol="1" gridrow="5" pageId="6" pageNumber="217">female</td>
<td id="965A4F3C090A0064419EF9D6FE1DF991" box="[410,468,1539,1595]" gridcol="2" gridrow="5" pageId="6" pageNumber="217">389.0</td>
<td id="965A4F3C090A006441F2F9D6FDF8F991" box="[502,561,1539,1595]" gridcol="3" gridrow="5" pageId="6" pageNumber="217">340.0</td>
<td id="965A4F3C090A00644249F9D6FDB2F991" box="[589,635,1539,1595]" gridcol="4" gridrow="5" pageId="6" pageNumber="217">49.0</td>
<td id="965A4F3C090A006442A0F9D6FD3FF991" box="[676,758,1539,1595]" gridcol="5" gridrow="5" pageId="6" pageNumber="217">0.13</td>
<td id="965A4F3C090A00644310F9D6FC8BF991" box="[788,834,1539,1595]" gridcol="6" gridrow="5" pageId="6" pageNumber="217">10.7</td>
<td id="965A4F3C090A00644360F9D6FC58F991" box="[868,913,1539,1595]" gridcol="7" gridrow="5" pageId="6" pageNumber="217">11.2</td>
<td id="965A4F3C090A006443BFF9D6FC28F991" box="[955,993,1539,1595]" gridcol="8" gridrow="5" pageId="6" pageNumber="217">6.8</td>
<td id="965A4F3C090A00644405F9D6FBE8F991" box="[1025,1057,1539,1595]" gridcol="9" gridrow="5" pageId="6" pageNumber="217">5.1</td>
<td id="965A4F3C090A00644443F9D6FBA0F991" box="[1095,1129,1539,1595]" gridcol="10" gridrow="5" pageId="6" pageNumber="217">1.9</td>
<td id="965A4F3C090A00644488F9D6FB71F991" box="[1164,1208,1539,1595]" gridcol="11" gridrow="5" pageId="6" pageNumber="217">4.6</td>
<td id="965A4F3C090A006444D0F9D6FB3FF991" box="[1236,1270,1539,1595]" gridcol="12" gridrow="5" pageId="6" pageNumber="217">2.3</td>
<td id="965A4F3C090A0064451CF9D6FAF1F991" box="[1304,1336,1539,1595]" gridcol="13" gridrow="5" pageId="6" pageNumber="217">3.7</td>
<td id="965A4F3C090A00644559F9D6F88EF991" box="[1373,1863,1539,1595]" gridcol="14" gridrow="5" pageId="6" pageNumber="217">gray with yellowish vertebral line and discrete paravertebral rows of dark speckles</td>
</tr>
</table>
</paragraph>
<caption id="3FC4748A090BFF9C4093F9F2FC69F9D0" httpUri="https://zenodo.org/record/253696/files/figure.png" pageId="7" pageNumber="218" targetBox="[323,1263,193,1553]" targetPageId="7">
<paragraph id="6B042402090BFF9C4093F9F2FC69F9D0" blockId="7.[151,1436,1575,1658]" pageId="7" pageNumber="218">
<emphasis id="59CFF810090BFF9C4093F9F2FEDFF996" bold="true" box="[151,278,1575,1597]" pageId="7" pageNumber="218">FIGURE 6.</emphasis>
Variation in life coloration of
<taxonomicName id="ACBB5F81090BFF9C4266F9F2FCE7F997" box="[610,814,1575,1597]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF810090BFF9C4266F9F2FCE7F997" box="[610,814,1575,1597]" italics="true" pageId="7" pageNumber="218">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B090BFF9C433CF9F2FC45F996" box="[824,908,1575,1596]" pageId="7" pageNumber="218" rank="species">
<emphasis id="59CFF810090BFF9C433CF9F2FC45F996" bold="true" box="[824,908,1575,1596]" pageId="7" pageNumber="218">sp. nov.</emphasis>
</taxonomicNameLabel>
: (A) male paratype ZMMU R-14002, (B) female paratype ZMMU R-14002, (C) female paratype VNMN 0 4724, (D) female paratype ZMMU R-14504 in situ (while swallowing a frog). Photos by V. Trounov (A,B), N. Poyarkov (C) and A. Vassilieva (D).
</paragraph>
</caption>
<paragraph id="6B042402090BFF9C40C3F977FADDF911" blockId="7.[151,1436,1698,2011]" box="[199,1300,1698,1723]" pageId="7" pageNumber="218">
Morphological characters mostly agree with those described for the
<typeStatus id="B4009AA0090BFF9C43C6F977FBEDF911" box="[962,1060,1698,1723]" pageId="7" pageNumber="218" type="holotype">holotype</typeStatus>
with few exceptions.
</paragraph>
<paragraph id="6B042402090BFF9C40C3F913FD1BF8A9" blockId="7.[151,1436,1698,2011]" pageId="7" pageNumber="218">
<emphasis id="59CFF810090BFF9C40C3F913FE47F974" box="[199,398,1734,1758]" italics="true" pageId="7" pageNumber="218">Body proportions</emphasis>
. Relative tail length differs significantly in the male
<typeStatus id="B4009AA0090BFF9C4405F912FBA8F975" box="[1025,1121,1735,1759]" pageId="7" pageNumber="218" type="paratype">paratype</typeStatus>
(TaL/TL 0.28) and female
<typeStatus id="B4009AA0090BFF9C4093F93EFECBF8A9" box="[151,258,1771,1795]" pageId="7" pageNumber="218" type="paratype">paratypes</typeStatus>
(average TaL/TL 0.15, three specimens).
</paragraph>
<paragraph id="6B042402090BFF9C40C3F8DAFD0CF8E1" blockId="7.[151,1436,1698,2011]" pageId="7" pageNumber="218">
<emphasis id="59CFF810090BFF9C40C3F8DAFEF8F88C" box="[199,305,1807,1830]" italics="true" pageId="7" pageNumber="218">Dentition</emphasis>
. 68 functional maxillary teeth in all specimens, the posterior three being strongly enlarged and dagger-shaped; palatine dentition well developed.
</paragraph>
<paragraph id="6B042402090BFF9C40C3F883FB3DF871" blockId="7.[151,1436,1698,2011]" pageId="7" pageNumber="218">
<emphasis id="59CFF810090BFF9C40C3F883FEA6F8C4" box="[199,367,1878,1902]" italics="true" pageId="7" pageNumber="218">Body scalation</emphasis>
. All scales are smooth, small, ovoid; first and second rows slightly enlarged, the vertebral row does not differ in size from other median dorsal rows; ASR varies 17/19,
<collectingCountry id="13AC6492090BFF9C43CFF8AFFBCFF838" box="[971,1030,1914,1938]" name="Egypt" pageId="7" pageNumber="218">MSR</collectingCountry>
and PSR are equal in all specimens, 17 and 15, respectively. Ventrals obtusely keeled laterally,
<quantity id="AC4389E7090BFF9C432DF84AFCB9F81D" box="[809,880,1950,1975]" metricMagnitude="0" metricUnit="m" metricValue="3.3274" pageId="7" pageNumber="218" unit="in" value="131.0">131 in</quantity>
one male (
<quantity id="AC4389E7090BFF9C43E8F84AFBFDF81D" box="[1004,1076,1950,1975]" metricMagnitude="0" metricUnit="m" metricValue="3.4036" pageId="7" pageNumber="218" unit="in" value="134.0">134 in</quantity>
the
<typeStatus id="B4009AA0090BFF9C4461F84BFB02F81D" box="[1125,1227,1950,1975]" pageId="7" pageNumber="218" type="holotype">holotype</typeStatus>
),
<quantity id="AC4389E7090BFF9C44DAF84AFA94F81D" box="[1246,1373,1950,1975]" metricMagnitude="0" metricUnit="m" metricValue="3.6322" metricValueMax="3.6576" metricValueMin="3.6068" pageId="7" pageNumber="218" unit="in" value="143.0" valueMax="144.0" valueMin="142.0">142144 in</quantity>
three females; anal plate entire, subcaudals paired,
<quantity id="AC4389E7090BFF9C4297F817FD04F871" box="[659,717,1986,2011]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="218" unit="in" value="60.0">60 in</quantity>
one male (
<quantity id="AC4389E7090BFF9C434CF817FC4BF871" box="[840,898,1986,2011]" metricMagnitude="0" metricUnit="m" metricValue="1.4731999999999998" pageId="7" pageNumber="218" unit="in" value="58.0">58 in</quantity>
the
<typeStatus id="B4009AA0090BFF9C43B6F817FBD1F871" box="[946,1048,1986,2011]" pageId="7" pageNumber="218" type="holotype">holotype</typeStatus>
),
<quantity id="AC4389E7090BFF9C4428F817FB46F871" box="[1068,1167,1986,2011]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" metricValueMax="10.16" metricValueMin="9.144" pageId="7" pageNumber="218" unit="in" value="38.0" valueMax="40.0" valueMin="36.0">3640 in</quantity>
females.
</paragraph>
<caption id="3FC4748A0904FF934093F907FECCF8AD" httpUri="https://zenodo.org/record/253697/files/figure.png" pageId="8" pageNumber="219" targetBox="[151,1436,193,1723]" targetPageId="8">
<paragraph id="6B0424020904FF934093F907FECCF8AD" blockId="8.[151,1436,1746,1799]" pageId="8" pageNumber="219">
<emphasis id="59CFF8100904FF934093F907FEDCF94D" bold="true" box="[151,277,1746,1768]" pageId="8" pageNumber="219">FIGURE 7.</emphasis>
Natural habitats of
<taxonomicName id="ACBB5F810904FF9341EFF907FD7FF942" box="[491,694,1746,1768]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100904FF9341EFF907FD7FF942" box="[491,694,1746,1768]" italics="true" pageId="8" pageNumber="219">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0904FF9342BAF907FCDBF94D" box="[702,786,1746,1767]" pageId="8" pageNumber="219" rank="species">
<emphasis id="59CFF8100904FF9342BAF907FCDBF94D" bold="true" box="[702,786,1746,1767]" pageId="8" pageNumber="219">sp. nov.</emphasis>
</taxonomicNameLabel>
: (A) coastal dunes, (B) light dipterocarp forest. Photos by A. Vassilieva.
</paragraph>
</caption>
<paragraph id="6B0424020904FF9240C3F8E6FE1CFF7F" blockId="8.[151,1437,1843,2012]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="220" pageId="8" pageNumber="219">
<emphasis id="59CFF8100904FF9340C3F8E6FEB8F8E1" box="[199,369,1843,1867]" italics="true" pageId="8" pageNumber="219">Head scalation</emphasis>
. Generally as for
<typeStatus id="B4009AA00904FF93423EF8E6FD69F8E6" box="[570,672,1843,1868]" pageId="8" pageNumber="219" type="holotype">holotype</typeStatus>
. In all specimens the nasal is divided through the nostril by a suture; loreal absent, preocular single, presubocular present in three specimens (and in the
<typeStatus id="B4009AA00904FF934454F882FB7DF8DA" box="[1104,1204,1879,1904]" pageId="8" pageNumber="219" type="holotype">holotype</typeStatus>
) and absent on both sides in one specimen (VNMN 04724); two postoculars present with one exception (ZMMU R-14002 male), when the asymmetrical condition is observed (2/1). Eight supralabials (VI and V contacting eye) with one exception (ZMMU R-14002 male), when the IV supralabial (occupying the place of VI and V supralabials of other specimens) is unusually large and the single to contact the eye. Nine infralabials in all
<typeStatus id="B4009AA00905FF924486FF4DFB38FF1A" box="[1154,1265,152,176]" pageId="9" pageNumber="220" type="paratype">paratypes</typeStatus>
. No variability observed in temporal scales.
</paragraph>
<paragraph id="6B0424020905FF9240C3FF35FD23FE5F" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">
<emphasis id="59CFF8100905FF9240C3FF35FE40FF52" box="[199,393,224,248]" italics="true" pageId="9" pageNumber="220">Coloration in life</emphasis>
. Dorsal background coloration varies from sandy-orange (ZMMU R-14002 male,
<figureCitation id="F38038870905FF924525FF0AFAABFF52" box="[1313,1378,223,248]" captionStart="FIGURE 6" captionStartId="7.[151,250,1575,1597]" captionTargetBox="[323,1263,193,1553]" captionTargetId="figure@7.[323,1263,193,1554]" captionTargetPageId="7" captionText="FIGURE 6. Variation in life coloration of Oligodon arenarius sp. nov.: (A) male paratype ZMMU R- 14002, (B) female paratype ZMMU R- 14002, (C) female paratype VNMN 0 4724, (D) female paratype ZMMU R- 14504 in situ (while swallowing a frog). Photos by V. Trounov (A, B), N. Poyarkov (C) and A. Vassilieva (D)." httpUri="https://zenodo.org/record/253696/files/figure.png" pageId="9" pageNumber="220">Fig. 6</figureCitation>
A) to orange (ZMMU R-14002 female,
<figureCitation id="F38038870905FF924225FED1FDAEFEB7" box="[545,615,260,285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1575,1597]" captionTargetBox="[323,1263,193,1553]" captionTargetId="figure@7.[323,1263,193,1554]" captionTargetPageId="7" captionText="FIGURE 6. Variation in life coloration of Oligodon arenarius sp. nov.: (A) male paratype ZMMU R- 14002, (B) female paratype ZMMU R- 14002, (C) female paratype VNMN 0 4724, (D) female paratype ZMMU R- 14504 in situ (while swallowing a frog). Photos by V. Trounov (A, B), N. Poyarkov (C) and A. Vassilieva (D)." httpUri="https://zenodo.org/record/253696/files/figure.png" pageId="9" pageNumber="220">Fig. 6</figureCitation>
B), yellowish-gray (VNMN 0 4724,
<figureCitation id="F38038870905FF924401FED1FB83FEB7" box="[1029,1098,260,285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1575,1597]" captionTargetBox="[323,1263,193,1553]" captionTargetId="figure@7.[323,1263,193,1554]" captionTargetPageId="7" captionText="FIGURE 6. Variation in life coloration of Oligodon arenarius sp. nov.: (A) male paratype ZMMU R- 14002, (B) female paratype ZMMU R- 14002, (C) female paratype VNMN 0 4724, (D) female paratype ZMMU R- 14504 in situ (while swallowing a frog). Photos by V. Trounov (A, B), N. Poyarkov (C) and A. Vassilieva (D)." httpUri="https://zenodo.org/record/253696/files/figure.png" pageId="9" pageNumber="220">Fig. 6</figureCitation>
C) or gray (ABV-00814,
<figureCitation id="F38038870905FF924574FED1FF6FFEEA" captionStart="FIGURE 6" captionStartId="7.[151,250,1575,1597]" captionTargetBox="[323,1263,193,1553]" captionTargetId="figure@7.[323,1263,193,1554]" captionTargetPageId="7" captionText="FIGURE 6. Variation in life coloration of Oligodon arenarius sp. nov.: (A) male paratype ZMMU R- 14002, (B) female paratype ZMMU R- 14002, (C) female paratype VNMN 0 4724, (D) female paratype ZMMU R- 14504 in situ (while swallowing a frog). Photos by V. Trounov (A, B), N. Poyarkov (C) and A. Vassilieva (D)." httpUri="https://zenodo.org/record/253696/files/figure.png" pageId="9" pageNumber="220">Fig. 6</figureCitation>
D); orange or yellow tint always being more saturated on the dorsum (sometimes forming a faint and diffuse vertebral line, as in female
<typeStatus id="B4009AA00905FF9241C1FE98FDECFECF" box="[453,549,333,357]" pageId="9" pageNumber="220" type="paratype">paratype</typeStatus>
ZMMU R-14504) and gradually turning paler to lower flanks. Blackish or dark brown speckles on dorsum are almost absent (ZMMU R-14002 male) or quite numerous and evident, evenly dispersed (VNMN 04724) or forming two more discrete paravertebral rows (ZMMU R-14504). On neck these dark speckles are always more discrete and may form two short paravertebral streaks (VNMN 04724), on tail they may fuse into faint paravertebral lines (ZMMU R-14504).
</paragraph>
<paragraph id="6B0424020905FF9240C3FE2AFB47FDCA" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">
Head pattern similar in all specimens, including faintly visible intraocular band and more clear brown-colored temporal bars and chevron. Ventrals and subcaudals immaculate in all specimens, in
<typeStatus id="B4009AA00905FF92445CFDF0FB70FD97" box="[1112,1209,549,573]" pageId="9" pageNumber="220" type="paratype">paratype</typeStatus>
female ABV-00814 salmon-pink as in
<typeStatus id="B4009AA00905FF924161FD92FE02FDCA" box="[357,459,583,608]" pageId="9" pageNumber="220" type="holotype">holotype</typeStatus>
; vital belly coloration for other
<typeStatus id="B4009AA00905FF92432AFD9DFC50FDCA" box="[814,921,584,608]" pageId="9" pageNumber="220" type="paratype">paratypes</typeStatus>
was not documented.
</paragraph>
<paragraph id="6B0424020905FF9240C3FDB8FD62FD02" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">In preservative, coloration fades slightly, especially the orange and yellow tints, which change to beige. Ventrals and subcaudals creamy-white or ivory.</paragraph>
</subSubSection>
<subSubSection id="23A177890905FF9240C3FD61FE17FA6A" pageId="9" pageNumber="220" type="distribution">
<paragraph id="6B0424020905FF9240C3FD61FADEFD66" blockId="9.[151,1437,151,2013]" box="[199,1303,692,717]" pageId="9" pageNumber="220">
<emphasis id="59CFF8100905FF9240C3FD61FEA9FD67" bold="true" box="[199,352,692,717]" pageId="9" pageNumber="220">Distribution.</emphasis>
To date known only from the
<typeStatus id="B4009AA00905FF9242B0FD60FD2DFD67" box="[692,740,693,717]" pageId="9" pageNumber="220">type</typeStatus>
locality (Binh ChauPhuoc Buu Nature Reserve).
</paragraph>
<paragraph id="6B0424020905FF9240C3FD02FCC0FC47" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">
<emphasis id="59CFF8100905FF9240C3FD02FDD1FD5A" bold="true" box="[199,536,727,752]" pageId="9" pageNumber="220">Habitat and natural history</emphasis>
. Four specimens (ZMMU R-14002 female, VNMN 0 4724, ZMMU R-14503, ZMMU R-14504) were collected in the coastal area along the Loc AnBinh Chau secondary road. The landscape comprises gently sloping low sand dunes covered with more or less sparse trees or high bushes (
<figureCitation id="F38038870905FF9244C1FCCAFACEFC92" box="[1221,1287,799,824]" captionStart="FIGURE 7" captionStartId="8.[151,250,1746,1768]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@8.[151,1436,193,1725]" captionTargetPageId="8" captionText="FIGURE 7. Natural habitats of Oligodon arenarius sp. nov.: (A) coastal dunes, (B) light dipterocarp forest. Photos by A. Vassilieva." httpUri="https://zenodo.org/record/253697/files/figure.png" pageId="9" pageNumber="220">Fig. 7</figureCitation>
A), including
<taxonomicName id="ACBB5F810905FF924093FC91FD9FFCF7" box="[151,598,836,861]" class="Gastropoda" family="Fissurellidae" genus="Buchanania" kingdom="Animalia" order="Sapindales" pageId="9" pageNumber="220" phylum="Mollusca" rank="species" species="siamensis">
<emphasis id="59CFF8100905FF924093FC91FE5AFCF6" box="[151,403,836,860]" italics="true" pageId="9" pageNumber="220">Buchanania siamensis</emphasis>
(Anacardiaceae)
</taxonomicName>
,
<taxonomicName id="ACBB5F810905FF924266FC91FBE2FCF7" box="[610,1067,836,861]" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="equisetifolia">
<emphasis id="59CFF8100905FF924266FC91FCA2FCF6" box="[610,875,836,860]" italics="true" pageId="9" pageNumber="220">Casuarina equisetifolia</emphasis>
(Casuarinaceae)
</taxonomicName>
,
<emphasis id="59CFF8100905FF924432FC91FAEBFCF6" box="[1078,1314,836,860]" italics="true" pageId="9" pageNumber="220">Blachiaja trophifolia</emphasis>
,
<taxonomicName id="ACBB5F810905FF924536FC91FE18FC2A" class="Magnoliopsida" family="Euphorbiaceae" genus="Suregada" kingdom="Plantae" order="Malpighiales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="multiflora">
<emphasis id="59CFF8100905FF924536FC91FECEFC2A" italics="true" pageId="9" pageNumber="220">Suregada multiflora</emphasis>
(Euphorbiaceae)
</taxonomicName>
and
<taxonomicName id="ACBB5F810905FF924219FCBDFC59FC2A" box="[541,912,871,896]" class="Aves" family="Bernieridae" genus="Randia" kingdom="Animalia" order="Passeriformes" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="dasycarpa">
<emphasis id="59CFF8100905FF924219FCBDFD3CFC2A" box="[541,757,872,896]" italics="true" pageId="9" pageNumber="220">Randia dasycarpa</emphasis>
(Rubiaceae)
</taxonomicName>
, with the locally present herbs
<taxonomicName id="ACBB5F810905FF924528FCBDFE2EFC0F" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="odoratissimus">
<emphasis id="59CFF8100905FF924528FCBDFEFDFC0E" italics="true" pageId="9" pageNumber="220">Pandanus odoratissimus</emphasis>
(Pandanaceae)
</taxonomicName>
,
<taxonomicName id="ACBB5F810905FF9241F3FC58FCA6FC0F" box="[503,879,908,933]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="rotundifolia">
<emphasis id="59CFF8100905FF9241F3FC58FD09FC0E" box="[503,704,908,932]" italics="true" pageId="9" pageNumber="220">Vitex rotundifolia</emphasis>
(Verbenaceae)
</taxonomicName>
,
<taxonomicName id="ACBB5F810905FF924384FC59FB77FC0E" box="[896,1214,908,932]" class="Liliopsida" family="Poaceae" genus="Spinifex" kingdom="Plantae" order="Poales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="littoreus">
<emphasis id="59CFF8100905FF924384FC59FB8BFC0E" box="[896,1090,908,932]" italics="true" pageId="9" pageNumber="220">Spinifex littoreus</emphasis>
(Poaceae)
</taxonomicName>
and
<taxonomicName id="ACBB5F810905FF9244FBFC58FE7FFC62" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" kingdom="Plantae" order="Solanales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="pescaprae">
<emphasis id="59CFF8100905FF9244FBFC58FF2CFC62" italics="true" pageId="9" pageNumber="220">Ipomoea pescaprae</emphasis>
(Convolvulaceae)
</taxonomicName>
. The area's natural character has been severely degraded by human activities, and most of the coastal area is occupied by resorts or private homes.
</paragraph>
<paragraph id="6B0424020905FF9240C3FC2DFB28FB0A" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">
One specimen (ZMMU R-14002 male) was collected approximately
<quantity id="AC4389E70905FF9243D5FC2DFBC2FBBA" box="[977,1035,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="220" unit="km" value="4.0">4 km</quantity>
from the seacoast, on the edge of a lowland forest (
<figureCitation id="F38038870905FF924154FBC9FE51FB9E" box="[336,408,1052,1077]" captionStart="FIGURE 7" captionStartId="8.[151,250,1746,1768]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@8.[151,1436,193,1725]" captionTargetPageId="8" captionText="FIGURE 7. Natural habitats of Oligodon arenarius sp. nov.: (A) coastal dunes, (B) light dipterocarp forest. Photos by A. Vassilieva." httpUri="https://zenodo.org/record/253697/files/figure.png" pageId="9" pageNumber="220">Fig. 7</figureCitation>
B), where open sandy areas are present and the co-dominant species are
<taxonomicName id="ACBB5F810905FF9244FEFBC8FEDBFBF2" class="Magnoliopsida" family="Dipterocarpaceae" genus="Dipterocarpus" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="chartaceus">
<emphasis id="59CFF8100905FF9244FEFBC8FEDBFBF2" italics="true" pageId="9" pageNumber="220">Dipterocarpus chartaceus</emphasis>
</taxonomicName>
,
<taxonomicName id="ACBB5F810905FF924121FB95FE3EFBF2" box="[293,503,1088,1112]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Shorea" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="roxburghii">
<emphasis id="59CFF8100905FF924121FB95FE3EFBF2" box="[293,503,1088,1112]" italics="true" pageId="9" pageNumber="220">Shorea roxburghii</emphasis>
</taxonomicName>
,
<taxonomicName id="ACBB5F810905FF92420EFB95FCB6FBF2" box="[522,895,1087,1112]" class="Gastropoda" family="Fissurellidae" genus="Buchanania" kingdom="Animalia" order="Sapindales" pageId="9" pageNumber="220" phylum="Mollusca" rank="species" species="siamensis">
<emphasis id="59CFF8100905FF92420EFB95FD50FBFD" box="[522,665,1088,1111]" italics="true" pageId="9" pageNumber="220">S. siamensis</emphasis>
(Dipterocarpaceae)
</taxonomicName>
,
<taxonomicName id="ACBB5F810905FF924394FB95FAAFFBF2" box="[912,1382,1087,1112]" class="Magnoliopsida" family="Anacardiaceae" genus="Melanorrhoea" higherTaxonomySource="CoL" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="laccifera">
<emphasis id="59CFF8100905FF924394FB95FB6BFBF2" box="[912,1186,1088,1112]" italics="true" pageId="9" pageNumber="220">Melanorrhoea laccifera</emphasis>
(Anacardiaceae)
</taxonomicName>
and
<taxonomicName id="ACBB5F810905FF924093FBB0FDDBFBD7" box="[151,530,1124,1149]" class="Magnoliopsida" family="Irvingiaceae" genus="Irvingia" kingdom="Plantae" order="Malpighiales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="malayana">
<emphasis id="59CFF8100905FF924093FBB0FEA1FBD6" box="[151,360,1124,1148]" italics="true" pageId="9" pageNumber="220">Irvingia malayana</emphasis>
(Irvingiaceae)
</taxonomicName>
, with
<taxonomicName id="ACBB5F810905FF92425EFBB1FC0EFBD6" box="[602,967,1124,1148]" class="Magnoliopsida" family="Myrtaceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" order="Myrtales" pageId="9" pageNumber="220" phylum="Tracheophyta" rank="species" species="cajuputi">
<emphasis id="59CFF8100905FF92425EFBB1FCFCFBD6" box="[602,821,1124,1148]" italics="true" pageId="9" pageNumber="220">Melaleuca cajuputi</emphasis>
(Myrtaceae)
</taxonomicName>
dominating in seasonally wet areas. The natural character of the area has been profoundly altered by selective logging and cattle pasturing.
</paragraph>
<paragraph id="6B0424020905FF9240C3FB79FB33FAFF" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">
All specimens were collected on the ground. They were active during the day (one specimen) or after dark (20:0022:00, three specimens). In attempts to avoid capture two of the specimens dived into the loose sand and moved swiftly under the sand layer, reappearing on the surface after
<quantity id="AC4389E70905FF92439FFB21FC33FAA6" box="[923,1018,1268,1293]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="9" pageNumber="220" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">
1.5
<specimenCount id="7DBDEF8B0905FF9243C8FB21FC33FAA6" box="[972,1018,1268,1293]" pageId="9" pageNumber="220" type="generic">2 m</specimenCount>
</quantity>
. The specimen ZMMU R-14504 was found while in the process of swallowing an adult microhylid frog
<taxonomicName id="ACBB5F810905FF924392FACDFB3EFA9A" authority="Hallowell" authorityName="Hallowell" box="[918,1271,1303,1328]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="pulchra">
<emphasis id="59CFF8100905FF924392FACDFBA0FA9A" box="[918,1129,1304,1328]" italics="true" pageId="9" pageNumber="220">Microhyla pulchra</emphasis>
(Hallowell)
</taxonomicName>
. The digestive tract of other specimens contained some amounts of sand, implying that they foraged on the ground.
</paragraph>
<paragraph id="6B0424020905FF9240C3FAB4FE17FA6A" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">
In the Binh ChauPhuoc Buu Nature Reserve,
<taxonomicName id="ACBB5F810905FF9242F7FAB5FC1CFADD" box="[755,981,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100905FF9242F7FAB5FC1CFADD" box="[755,981,1376,1400]" italics="true" pageId="9" pageNumber="220">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0905FF9243E4FA8AFBF7FAD2" box="[992,1086,1375,1400]" pageId="9" pageNumber="220" rank="species">
<emphasis id="59CFF8100905FF9243E4FA8AFBF7FAD2" bold="true" box="[992,1086,1375,1400]" pageId="9" pageNumber="220">sp. nov.</emphasis>
</taxonomicNameLabel>
coexists with other
<taxonomicName id="ACBB5F810905FF924536FAB5FA55FAD2" box="[1330,1436,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="genus">
<emphasis id="59CFF8100905FF924536FAB5FA55FAD2" box="[1330,1436,1376,1400]" italics="true" pageId="9" pageNumber="220">Oligodon</emphasis>
</taxonomicName>
species:
<taxonomicName id="ACBB5F810905FF9240F2FA51FEA8FA36" box="[246,353,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="deuvei">
<emphasis id="59CFF8100905FF9240F2FA51FEA8FA36" box="[246,353,1412,1436]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF9240F2FA51FEC4FA31" box="[246,269,1412,1435]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. deuvei
</emphasis>
</taxonomicName>
(data of A. Vassilieva and N. Poyarkov), as well as
<taxonomicName id="ACBB5F810905FF9243A0FA51FBF6FA36" box="[932,1087,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="fasciolatus">
<emphasis id="59CFF8100905FF9243A0FA51FBF6FA36" box="[932,1087,1412,1436]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF9243A0FA51FC72FA31" box="[932,955,1412,1435]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. fasciolatus
</emphasis>
</taxonomicName>
and one unidentified
<taxonomicName id="ACBB5F810905FF924536FA51FA55FA36" box="[1330,1436,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="genus">
<emphasis id="59CFF8100905FF924536FA51FA55FA36" box="[1330,1436,1412,1436]" italics="true" pageId="9" pageNumber="220">Oligodon</emphasis>
</taxonomicName>
sp. (
<bibRefCitation id="0F2A59F30905FF9240C1FA7DFE1BFA6A" author="Nguyen" box="[197,466,1448,1473]" pageId="9" pageNumber="220" refString="Nguyen, N. H. &amp; Hoang, M. D. (2013) The species composition of Amphibian and Reptile in Binh Chau - Phuoc Buu Nature Reserve, Ba Ria Vung Tau Province. In: Proceedings of the Fifth National Conference on Ecology and Biological Resources. Agriculture Publishing House, Hanoi, pp. 504 - 513." type="book chapter" year="2013">Nguyen &amp; Hoang 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="23A177890905FF9140C3FA19FD0FFDB2" lastPageId="10" lastPageNumber="221" pageId="9" pageNumber="220" type="etymology">
<paragraph id="6B0424020905FF9240C3FA19FA55F987" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">
<emphasis id="59CFF8100905FF9240C3FA19FE8FFA4F" bold="true" box="[199,326,1484,1509]" pageId="9" pageNumber="220">Etymology</emphasis>
. The new species name is an adjective in the nominative case, masculine gender, derived from the Latin word &quot;arena&quot; meaning &quot;sand&quot;, &quot;sandy land&quot;. The name is intended to reflect the evident preference by the new species for sandy coastal habitats and the snake's remarkable ability to plunge into the sand and move under it.
</paragraph>
<paragraph id="6B0424020905FF9240C3F9E2FB4EF917" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="220">
<emphasis id="59CFF8100905FF9240C3F9E2FEA4F9FA" bold="true" box="[199,365,1591,1616]" pageId="9" pageNumber="220">Comparisons.</emphasis>
<taxonomicName id="ACBB5F810905FF92417FF9EDFDA9F9E5" box="[379,608,1592,1616]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100905FF92417FF9EDFDA9F9E5" box="[379,608,1592,1616]" italics="true" pageId="9" pageNumber="220">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0905FF92426AF9E2FD06F9FA" box="[622,719,1591,1616]" pageId="9" pageNumber="220" rank="species">
<emphasis id="59CFF8100905FF92426AF9E2FD06F9FA" bold="true" box="[622,719,1591,1616]" pageId="9" pageNumber="220">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from all other species of the genus by the unique combination of the following characters: 17 DSR, the absence of a loreal scale, unforked hemipenis without spines or obvious papillae, the absence of a specific dorsal coloration pattern (blotches, crossbars or longitudinal lines) and the expressed sexual dimorphism in the relative tail length and number of subcaudals.
</paragraph>
<paragraph id="6B0424020905FF9140C3F912FD0FFDB2" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="221" pageId="9" pageNumber="220">
The absence of a loreal is a rather rare feature in kukri snakes. In particular, among the 22
<taxonomicName id="ACBB5F810905FF9244DDF91DFA8AF94A" box="[1241,1347,1736,1760]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="genus">
<emphasis id="59CFF8100905FF9244DDF91DFA8AF94A" box="[1241,1347,1736,1760]" italics="true" pageId="9" pageNumber="220">Oligodon</emphasis>
</taxonomicName>
species known from Indochina, this element of head scalation is consistently absent in only three species:
<taxonomicName id="ACBB5F810905FF9244F0F939FF27F882" authority="Leviton" authorityName="Leviton" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="59CFF8100905FF9244F0F939FA55F8AE" box="[1268,1436,1772,1796]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF9244F0F939FAC2F8A9" box="[1268,1291,1772,1795]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. annamensis
</emphasis>
Leviton
</taxonomicName>
(
<collectingCountry id="13AC64920905FF924106F8DAFEACF882" box="[258,357,1807,1832]" name="Vietnam" pageId="9" pageNumber="220">Vietnam</collectingCountry>
,
<collectingCountry id="13AC64920905FF924172F8DAFE27F882" box="[374,494,1807,1832]" name="Cambodia" pageId="9" pageNumber="220">Cambodia</collectingCountry>
),
<taxonomicName id="ACBB5F810905FF924202F8C5FD3FF882" authority="Blyth" authorityName="Blyth" box="[518,758,1807,1832]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="catenatus">
<emphasis id="59CFF8100905FF924202F8C5FD51F882" box="[518,664,1808,1832]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF924202F8C5FDD4F88D" box="[518,541,1808,1831]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. catenatus
</emphasis>
(Blyth)
</taxonomicName>
(
<collectingCountry id="13AC64920905FF92430EF8DAFCA4F882" box="[778,877,1807,1832]" name="Vietnam" pageId="9" pageNumber="220">Vietnam</collectingCountry>
,
<collectingCountry id="13AC64920905FF92437AF8DAFC2DF882" box="[894,996,1807,1832]" name="Thailand" pageId="9" pageNumber="220">Thailand</collectingCountry>
,
<collectingCountry id="13AC64920905FF9243F1F8C5FBADF882" box="[1013,1124,1808,1832]" name="Myanmar" pageId="9" pageNumber="220">Myanmar</collectingCountry>
) and
<taxonomicName id="ACBB5F810905FF9244A8F8C5FF25F8E6" authority="Angel et Bourret" authorityName="Angel et Bourret" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="lacroixi">
<emphasis id="59CFF8100905FF9244A8F8C5FAE3F882" box="[1196,1322,1808,1832]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF9244A8F8C5FB0AF88D" box="[1196,1219,1808,1831]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. lacroixi
</emphasis>
Angel et Bourret
</taxonomicName>
(
<collectingCountry id="13AC64920905FF9240FAF8E1FEA8F8E7" box="[254,353,1844,1869]" name="Vietnam" pageId="9" pageNumber="220">Vietnam</collectingCountry>
,
<collectingCountry id="13AC64920905FF924174F8E1FE71F8E7" box="[368,440,1844,1869]" name="China" pageId="9" pageNumber="220">China</collectingCountry>
); also, two Indochinese species occurring in
<collectingCountry id="13AC64920905FF9243C6F8E1FBEDF8E7" box="[962,1060,1844,1869]" name="Vietnam" pageId="9" pageNumber="220">Vietnam</collectingCountry>
,
<taxonomicName id="ACBB5F810905FF924436F8E1FA8CF8E7" authority="Boulenger" authorityName="Boulenger" box="[1074,1349,1844,1869]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="59CFF8100905FF924436F8E1FB7AF8E6" box="[1074,1203,1844,1868]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF924436F8E1FB80F8E1" box="[1074,1097,1844,1867]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. mouhoti
</emphasis>
(Boulenger)
</taxonomicName>
and
<taxonomicName id="ACBB5F810905FF924585F8E1FEA1F8DA" authority="Angel" authorityName="Angel" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="59CFF8100905FF924585F8E1FECAF8DA" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF924585F8E1FA51F8E1" box="[1409,1432,1844,1867]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. macrurus
</emphasis>
(Angel)
</taxonomicName>
, lack the loreal scale facultatively.
<taxonomicName id="ACBB5F810905FF924312F88DFC30F8C5" box="[790,1017,1880,1904]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100905FF924312F88DFC30F8C5" box="[790,1017,1880,1904]" italics="true" pageId="9" pageNumber="220">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0905FF924401F882FBACF8DA" box="[1029,1125,1879,1904]" pageId="9" pageNumber="220" rank="species">
<emphasis id="59CFF8100905FF924401F882FBACF8DA" bold="true" box="[1029,1125,1879,1904]" pageId="9" pageNumber="220">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from them by the following features: from
<taxonomicName id="ACBB5F810905FF9241B7F8A9FD92F83E" box="[435,603,1916,1940]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="59CFF8100905FF9241B7F8A9FD92F83E" box="[435,603,1916,1940]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF9241B7F8A9FE03F839" box="[435,458,1916,1939]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. annamensis
</emphasis>
</taxonomicName>
,
<taxonomicName id="ACBB5F810905FF92426EF8A9FD30F83E" box="[618,761,1916,1940]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="catenatus">
<emphasis id="59CFF8100905FF92426EF8A9FD30F83E" box="[618,761,1916,1940]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF92426EF8A9FD48F839" box="[618,641,1916,1939]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. catenatus
</emphasis>
</taxonomicName>
and
<taxonomicName id="ACBB5F810905FF924336F8A9FC62F83E" box="[818,939,1916,1940]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="lacroixi">
<emphasis id="59CFF8100905FF924336F8A9FC62F83E" box="[818,939,1916,1940]" italics="true" pageId="9" pageNumber="220">
<collectingCountry id="13AC64920905FF924336F8A9FC80F839" box="[818,841,1916,1939]" name="Iceland" pageId="9" pageNumber="220">O</collectingCountry>
. lacroixi
</emphasis>
</taxonomicName>
, by a greater DSR count (17 vs. 13, 13 and 15, respectively); usually present presubocular; 8(7) supralabials vs. 6(5), 6 and 5, respectively; 9(8) infralabials vs. 6, 7 and 6, respectively; a lesser number of ventrals (131144 vs. 148170, 179212 and 162178, respectively) and a dorsal coloration pattern (uniform or with dark speckling vs. thin light crossbars in
<taxonomicName id="ACBB5F810906FF91448AFF4DFAFFFF05" box="[1166,1334,152,175]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="59CFF8100906FF91448AFF4DFAFFFF05" box="[1166,1334,152,175]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91448AFF4DFB6CFF05" box="[1166,1189,152,175]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. annamensis
</emphasis>
</taxonomicName>
, chained vertebral line in
<taxonomicName id="ACBB5F810906FF91414BFF69FE14FF7E" box="[335,477,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="catenatus">
<emphasis id="59CFF8100906FF91414BFF69FE14FF7E" box="[335,477,188,212]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91414BFF69FEAFFF79" box="[335,358,188,211]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. catenatus
</emphasis>
</taxonomicName>
and four indistinct longitudinal lines in
<taxonomicName id="ACBB5F810906FF914399FF69FBDFFF7E" box="[925,1046,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="lacroixi">
<emphasis id="59CFF8100906FF914399FF69FBDFFF7E" box="[925,1046,188,212]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914399FF69FC7DFF79" box="[925,948,188,211]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. lacroixi
</emphasis>
</taxonomicName>
); additionally,
<taxonomicName id="ACBB5F810906FF9144BBFF69FA55FF7E" box="[1215,1436,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100906FF9144BBFF69FA55FF7E" box="[1215,1436,188,212]" italics="true" pageId="10" pageNumber="221">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0906FF914093FF0AFF38FF52" box="[151,241,223,248]" pageId="10" pageNumber="221" rank="species">
<emphasis id="59CFF8100906FF914093FF0AFF38FF52" bold="true" box="[151,241,223,248]" pageId="10" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="ACBB5F810906FF914181FF35FDE5FF5D" box="[389,556,224,247]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="59CFF8100906FF914181FF35FDE5FF5D" box="[389,556,224,247]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914181FF35FE55FF5D" box="[389,412,224,247]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. annamensis
</emphasis>
</taxonomicName>
and
<taxonomicName id="ACBB5F810906FF914267FF35FD12FF52" box="[611,731,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="lacroixi">
<emphasis id="59CFF8100906FF914267FF35FD12FF52" box="[611,731,224,248]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914267FF35FDB3FF5D" box="[611,634,224,247]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. lacroixi
</emphasis>
</taxonomicName>
by a greater number of subcaudals (5860 vs.
<quantity id="AC4389E70906FF9144EDFF0AFA85FF52" box="[1257,1356,223,248]" metricMagnitude="0" metricUnit="m" metricValue="1.143" metricValueMax="1.1684" metricValueMin="1.1176" pageId="10" pageNumber="221" unit="in" value="45.0" valueMax="46.0" valueMin="44.0">4446 in</quantity>
males, 3640 vs.
<quantity id="AC4389E70906FF91410EFED1FE8AFEB7" box="[266,323,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="221" unit="in" value="30.0">30 in</quantity>
females of
<taxonomicName id="ACBB5F810906FF9141C1FED1FDA2FEB6" box="[453,619,260,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="59CFF8100906FF9141C1FED1FDA2FEB6" box="[453,619,260,284]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9141C1FED1FE15FEB1" box="[453,476,260,283]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. annamensis
</emphasis>
</taxonomicName>
,
<quantity id="AC4389E70906FF91427DFED1FD15FEB7" box="[633,732,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="8.001" metricValueMax="8.636" metricValueMin="7.366" pageId="10" pageNumber="221" unit="in" value="31.5" valueMax="34.0" valueMin="29.0">2934 in</quantity>
<taxonomicName id="ACBB5F810906FF9142E6FED1FC93FEB6" box="[738,858,260,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="lacroixi">
<emphasis id="59CFF8100906FF9142E6FED1FC93FEB6" box="[738,858,260,284]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9142E6FED1FD30FEB1" box="[738,761,260,283]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. lacroixi
</emphasis>
</taxonomicName>
) and from
<taxonomicName id="ACBB5F810906FF9143D2FED1FBADFEB6" box="[982,1124,260,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="catenatus">
<emphasis id="59CFF8100906FF9143D2FED1FBADFEB6" box="[982,1124,260,284]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9143D2FED1FC24FEB1" box="[982,1005,260,283]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. catenatus
</emphasis>
</taxonomicName>
and
<taxonomicName id="ACBB5F810906FF91449EFED1FADBFEB6" box="[1178,1298,260,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="lacroixi">
<emphasis id="59CFF8100906FF91449EFED1FADBFEB6" box="[1178,1298,260,284]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91449EFED1FB78FEB1" box="[1178,1201,260,283]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. lacroixi
</emphasis>
</taxonomicName>
by an entire anal plate instead of divided one;
<taxonomicName id="ACBB5F810906FF914219FEFDFD35FE95" box="[541,764,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100906FF914219FEFDFD35FE95" box="[541,764,296,320]" italics="true" pageId="10" pageNumber="221">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0906FF914301FEF2FCA9FEEA" box="[773,864,295,320]" pageId="10" pageNumber="221" rank="species">
<emphasis id="59CFF8100906FF914301FEF2FCA9FEEA" bold="true" box="[773,864,295,320]" pageId="10" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="ACBB5F810906FF9143FFFEFDFBB2FEEA" box="[1019,1147,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="59CFF8100906FF9143FFFEFDFBB2FEEA" box="[1019,1147,296,320]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9143FFFEFDFBDBFE95" box="[1019,1042,296,319]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. mouhoti
</emphasis>
</taxonomicName>
by a longer tail in males (0.260.28 vs. 0.170.19) and a greater number of subcaudals in males (5860 vs. 3943), as well as by dorsal coloration features (vertebral stripe and two large dark blotches on tail in
<taxonomicName id="ACBB5F810906FF91441AFEA5FB6AFE22" box="[1054,1187,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="59CFF8100906FF91441AFEA5FB6AFE22" box="[1054,1187,368,392]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91441AFEA5FBFCFE2D" box="[1054,1077,368,391]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. mouhoti
</emphasis>
</taxonomicName>
) and the hemipenial morphology (see below); from
<taxonomicName id="ACBB5F810906FF9141F2FE41FD4DFE06" box="[502,644,404,428]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="59CFF8100906FF9141F2FE41FD4DFE06" box="[502,644,404,428]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9141F2FE41FDC4FE01" box="[502,525,404,427]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. macrurus
</emphasis>
</taxonomicName>
, by a relatively shorter tail in males (TaL/TL 0.260.28 vs. 0.340.37, calculated after the data of
<bibRefCitation id="0F2A59F30906FF9141C3FE62FD6AFE7A" author="Geissler" box="[455,675,439,464]" pageId="10" pageNumber="221" refString="Geissler, P., Krohn, A. R. &amp; Rennert, A. (2011) Herpetofaunal records in coastal dune areas, Binh Thuan Province, Southern Vietnam, with the rediscovery of Oligodon macrurus Angel, 1927. Russian Journal of Herpetology, 18 (4), 317 - 324." type="journal article" year="2011">
Geissler
<emphasis id="59CFF8100906FF91422EFE6CFDAAFE7A" box="[554,611,440,464]" italics="true" pageId="10" pageNumber="221">et al.</emphasis>
2011
</bibRefCitation>
, or
<quantity id="AC4389E70906FF9142C9FE6DFCD2FE7A" box="[717,795,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="7.874" pageId="10" pageNumber="221" unit="in" value="0.31">0.31 in</quantity>
one specimen, P. David, pers. comm.), a markedly lesser number of subcaudals in males (5860 vs. 7394), a slightly lesser number of ventrals in males (131134 vs. 139 145) and the hemipenial morphology (see below).
</paragraph>
</subSubSection>
<subSubSection id="23A177890906FF9740C3FDF1FBBEFB0A" lastPageId="12" lastPageNumber="223" pageId="10" pageNumber="221" type="reference_group">
<paragraph id="6B0424020906FF9140C3FDF1FEE7FBBA" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="221">
<taxonomicName id="ACBB5F810906FF9140C3FDF1FE60FD96" box="[199,425,548,572]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100906FF9140C3FDF1FE60FD96" box="[199,425,548,572]" italics="true" pageId="10" pageNumber="221">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0906FF9141B2FDF1FDDFFD97" box="[438,534,548,573]" pageId="10" pageNumber="221" rank="species">
<emphasis id="59CFF8100906FF9141B2FDF1FDDFFD97" bold="true" box="[438,534,548,573]" pageId="10" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
also differs from many congeners by its hemipenial morphology (unforked hemipenis devoid of spines and papillae). In particular, it differs from all the members of the informal
<taxonomicName id="ACBB5F810906FF914517FD9DFA55FDF5" box="[1299,1436,584,607]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="taeniatus">
<emphasis id="59CFF8100906FF914517FD9DFA55FDF5" box="[1299,1436,584,607]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914517FD9DFAE3FDF5" box="[1299,1322,584,607]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. taeniatus
</emphasis>
</taxonomicName>
group (sensu
<bibRefCitation id="0F2A59F30906FF914137FDB9FE73FD2E" author="Smith" box="[307,442,620,645]" pageId="10" pageNumber="221" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo - chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
,
<bibRefCitation id="0F2A59F30906FF9141CCFDB9FD6BFD2F" author="David" box="[456,674,620,645]" pageId="10" pageNumber="221" refString="David, P., Vogel, G. &amp; Rooijen, J. van (2008 b) A revision of the Oligodon taeniatus (Gunther, 1861) (Squamata: Colubridae) group, with the description of three new species from the Indochinese Region. Zootaxa, 1965, 1 - 49." type="journal article" year="2008" yearSuffix="b">
David
<emphasis id="59CFF8100906FF914212FDB8FD98FD2E" box="[534,593,620,644]" italics="true" pageId="10" pageNumber="221">et al.</emphasis>
2008b
</bibRefCitation>
) with Indochinese distribution, namely
<taxonomicName id="ACBB5F810906FF914468FDB9FABAFD2F" authority="Gunther" authorityName="Gunther" box="[1132,1395,620,645]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="taeniatus">
<emphasis id="59CFF8100906FF914468FDB9FB31FD2E" box="[1132,1272,620,644]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914468FDB9FB4AFD29" box="[1132,1155,620,643]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. taeniatus
</emphasis>
(Günther)
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF914585FDB9FD62FD02" authority="David, Vogel &amp; Van Rooijen" authorityName="David, Vogel &amp; Van Rooijen" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="pseudotaeniatus">
<emphasis id="59CFF8100906FF914585FDB9FE84FD02" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914585FDB9FA51FD29" box="[1409,1432,620,643]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. pseudotaeniatus
</emphasis>
David, Vogel &amp; Van Rooijen
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF9142BDFD45FB4EFD02" authority="David, Vogel &amp; Van Rooijen" authorityName="David, Vogel &amp; Van Rooijen" box="[697,1159,655,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="deuvei">
<emphasis id="59CFF8100906FF9142BDFD45FCE1FD02" box="[697,808,656,680]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9142BDFD45FD19FD0D" box="[697,720,656,679]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. deuvei
</emphasis>
David, Vogel &amp; Van Rooijen
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF914491FD45FABBFD02" authority="Smith" authorityName="Smith" box="[1173,1394,655,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="barroni">
<emphasis id="59CFF8100906FF914491FD45FAC6FD02" box="[1173,1295,656,680]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914491FD45FB65FD0D" box="[1173,1196,656,679]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. barroni
</emphasis>
(Smith)
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF914585FD45FD88FD67" authority="David, Vogel &amp; Van Rooijen" authorityName="David, Vogel &amp; Van Rooijen" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="moricei">
<emphasis id="59CFF8100906FF914585FD45FF25FD66" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914585FD45FA51FD0D" box="[1409,1432,656,679]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. moricei
</emphasis>
David, Vogel &amp; Van Rooijen
</taxonomicName>
and
<taxonomicName id="ACBB5F810906FF91427FFD61FD35FD66" box="[635,764,692,716]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="59CFF8100906FF91427FFD61FD35FD66" box="[635,764,692,716]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91427FFD61FD5BFD61" box="[635,658,692,715]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. mouhoti
</emphasis>
</taxonomicName>
discussed above, which possess deeply forked hemipenes. Additionally,
<taxonomicName id="ACBB5F810906FF914134FD0DFDC4FD45" box="[304,525,728,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100906FF914134FD0DFDC4FD45" box="[304,525,728,752]" italics="true" pageId="10" pageNumber="221">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0906FF914217FD02FDA5FD5A" box="[531,620,727,752]" pageId="10" pageNumber="221" rank="species">
<emphasis id="59CFF8100906FF914217FD02FDA5FD5A" bold="true" box="[531,620,727,752]" pageId="10" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from them externally by the dorsal coloration (all members of the
<taxonomicName id="ACBB5F810906FF914093FD29FEE8FCBE" box="[151,289,764,788]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="taeniatus">
<emphasis id="59CFF8100906FF914093FD29FEE8FCBE" box="[151,289,764,788]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914093FD29FF67FCB9" box="[151,174,764,787]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. taeniatus
</emphasis>
</taxonomicName>
group possessing vertebral stripes or blotches) and by the following characters: from
<taxonomicName id="ACBB5F810906FF9144ECFD29FAB8FCBE" box="[1256,1393,764,788]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="taeniatus">
<emphasis id="59CFF8100906FF9144ECFD29FAB8FCBE" box="[1256,1393,764,788]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9144ECFD29FB36FCB9" box="[1256,1279,764,787]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. taeniatus
</emphasis>
</taxonomicName>
, by 17 vs. 19 DSR and a greater subcaudals count in males (5860 vs. 3848); from
<taxonomicName id="ACBB5F810906FF91443AFCF5FAD1FC92" box="[1086,1304,800,824]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="pseudotaeniatus">
<emphasis id="59CFF8100906FF91443AFCF5FAD1FC92" box="[1086,1304,800,824]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91443AFCF5FB9CFC9D" box="[1086,1109,800,823]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. pseudotaeniatus
</emphasis>
</taxonomicName>
, by 9 vs. 8 infralabials and a greater subcaudals count in males (5860 vs. 4446); from
<taxonomicName id="ACBB5F810906FF9143F0FC91FB96FCF6" box="[1012,1119,836,860]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="deuvei">
<emphasis id="59CFF8100906FF9143F0FC91FB96FCF6" box="[1012,1119,836,860]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9143F0FC91FBC2FCF1" box="[1012,1035,836,859]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. deuvei
</emphasis>
</taxonomicName>
, by a slightly lesser ventrals count (131134 vs.
<quantity id="AC4389E70906FF91417DFCBDFE33FC2A" box="[377,506,871,896]" metricMagnitude="0" metricUnit="m" metricValue="3.6449" metricValueMax="3.7338" metricValueMin="3.556" pageId="10" pageNumber="221" unit="in" value="143.5" valueMax="147.0" valueMin="140.0">140147 in</quantity>
males, 142144 vs.
<quantity id="AC4389E70906FF9142E2FCB2FCAFFC2A" box="[742,870,871,896]" metricMagnitude="0" metricUnit="m" metricValue="3.8354" metricValueMax="3.937" metricValueMin="3.7338" pageId="10" pageNumber="221" unit="in" value="151.0" valueMax="155.0" valueMin="147.0">147155 in</quantity>
females) and a greater subcaudals count in males (5860 vs. 3647); from
<taxonomicName id="ACBB5F810906FF9141C7FC59FDF7FC0E" box="[451,574,908,932]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="barroni">
<emphasis id="59CFF8100906FF9141C7FC59FDF7FC0E" box="[451,574,908,932]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9141C7FC59FE13FC09" box="[451,474,908,931]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. barroni
</emphasis>
</taxonomicName>
, by a relatively longer tail in males (0.260.28 vs. 0.170.19), a greater subcaudals count in males (5860 vs. 3648) and females (3640 vs. 2835) and the immaculate belly coloration; from
<taxonomicName id="ACBB5F810906FF9140D3FC01FE98FC46" box="[215,337,980,1004]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="moricei">
<emphasis id="59CFF8100906FF9140D3FC01FE98FC46" box="[215,337,980,1004]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9140D3FC01FF27FC41" box="[215,238,980,1003]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. moricei
</emphasis>
</taxonomicName>
, by a lesser ventrals count (131144 vs. 175), 68 maxillary teeth vs. 12 and a usually present presubocular.
</paragraph>
<paragraph id="6B0424020906FF9140C3FBC9FB7FF9DE" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="221">
<taxonomicName id="ACBB5F810906FF9140C3FBC9FE6CFB9E" box="[199,421,1052,1076]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100906FF9140C3FBC9FE6CFB9E" box="[199,421,1052,1076]" italics="true" pageId="10" pageNumber="221">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0906FF9141A9FBC9FDC1FB9F" box="[429,520,1052,1077]" pageId="10" pageNumber="221" rank="species">
<emphasis id="59CFF8100906FF9141A9FBC9FDC1FB9F" bold="true" box="[429,520,1052,1077]" pageId="10" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
also differs by its hemipenial morphology from all the members of the informal
<taxonomicName id="ACBB5F810906FF914093FB95FEA3FBF2" box="[151,362,1088,1112]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="59CFF8100906FF914093FB95FEA3FBF2" box="[151,362,1088,1112]" italics="true" pageId="10" pageNumber="221">Oligodon cyclurus</emphasis>
</taxonomicName>
group (sensu
<bibRefCitation id="0F2A59F30906FF914217FBEAFD52FBF2" author="Smith" box="[531,667,1087,1112]" pageId="10" pageNumber="221" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo - chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
,
<bibRefCitation id="0F2A59F30906FF9142AEFBEAFC4EFBF2" author="David" box="[682,903,1087,1112]" pageId="10" pageNumber="221" refString="David, P., Vogel, G. &amp; Pauwels, O. S. G. (2008 a) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from southern Vietnam and Cambodia. Zootaxa, 1939, 19 - 37." type="journal article" year="2008" yearSuffix="a">
David
<emphasis id="59CFF8100906FF9142FDFB94FCFCFBF2" box="[761,821,1088,1112]" italics="true" pageId="10" pageNumber="221">et al.</emphasis>
2008a
</bibRefCitation>
, Green
<emphasis id="59CFF8100906FF9143E1FB94FBE9FBF2" box="[997,1056,1088,1112]" italics="true" pageId="10" pageNumber="221">et al.</emphasis>
2010). The species of
<taxonomicName id="ACBB5F810906FF914536FB95FF3FFBD6" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="59CFF8100906FF914536FB95FF3FFBD6" italics="true" pageId="10" pageNumber="221">Oligodon cyclurus</emphasis>
</taxonomicName>
group are distributed in Indochina,
<collectingCountry id="13AC64920906FF91428EFBB1FD04FBD7" box="[650,717,1124,1149]" name="China" pageId="10" pageNumber="221">China</collectingCountry>
and
<collectingCountry id="13AC64920906FF914300FBB1FC89FBD7" box="[772,832,1124,1149]" name="India" pageId="10" pageNumber="221">India</collectingCountry>
, and possess deeply forked hemipenes without spines but sometimes with papillae; these features were documented for
<taxonomicName id="ACBB5F810906FF914397FB5DFDFAFB6F" authority="Vassilieva, Geissler, Galoyan, Poyarkov, Van Devender &amp; Bohme" authorityName="Vassilieva, Geissler, Galoyan, Poyarkov, Van Devender &amp; Bohme" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cattienensis">
<emphasis id="59CFF8100906FF914397FB5DFBF7FB35" box="[915,1086,1160,1183]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914397FB5DFC63FB35" box="[915,938,1160,1183]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. cattienensis
</emphasis>
Vassilieva, Geissler, Galoyan, Poyarkov, Van Devender &amp; Böhme
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF914245FB79FC83FB6F" authority="Gunther" authorityName="Gunther" box="[577,842,1196,1221]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="59CFF8100906FF914245FB79FD07FB6E" box="[577,718,1196,1220]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914245FB79FD91FB69" box="[577,600,1196,1219]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. chinensis
</emphasis>
(Günther)
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF91435DFB79FB80FB6E" authority="Cantor" authorityName="Cantor" box="[857,1097,1196,1220]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="59CFF8100906FF91435DFB79FC14FB6E" box="[857,989,1196,1220]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91435DFB79FCB9FB69" box="[857,880,1196,1219]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. cyclurus
</emphasis>
(Cantor)
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF91445CFB79FABBFB6F" authority="Gunther" authorityName="Gunther" box="[1112,1394,1196,1221]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="fasciolatus">
<emphasis id="59CFF8100906FF91445CFB79FB3EFB6E" box="[1112,1271,1196,1220]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91445CFB79FBA6FB69" box="[1112,1135,1196,1219]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. fasciolatus
</emphasis>
(Günther)
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF914585FB79FE5BFB42" authority="Gunther" authorityName="Gunther" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="59CFF8100906FF914585FB79FED3FB42" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914585FB79FA51FB69" box="[1409,1432,1196,1219]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. formosanus
</emphasis>
(Günther)
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF914199FB05FD45FB42" authority="Wall" authorityName="Wall" box="[413,652,1231,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="juglandifer">
<collectingCountry id="13AC64920906FF914199FB05FE78FB4D" box="[413,433,1232,1255]" name="Iceland" pageId="10" pageNumber="221">
<emphasis id="59CFF8100906FF914199FB05FE78FB4D" box="[413,433,1232,1255]" italics="true" pageId="10" pageNumber="221">O</emphasis>
</collectingCountry>
.
<emphasis id="59CFF8100906FF9141BBFB05FDF2FB42" box="[447,571,1232,1256]" italics="true" pageId="10" pageNumber="221">juglandifer</emphasis>
(Wall)
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF91429CFB05FB51FB42" authority="Neang, Grismer &amp; Daltry" authorityName="Neang, Grismer &amp; Daltry" box="[664,1176,1231,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="kampucheaensis">
<emphasis id="59CFF8100906FF91429CFB05FCB8FB42" box="[664,881,1232,1256]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91429CFB05FD66FB4D" box="[664,687,1232,1255]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. kampucheaensis
</emphasis>
Neang, Grismer &amp; Daltry
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF9144A7FB05FA51FB42" authority="Morice" authorityName="Morice" box="[1187,1432,1231,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="59CFF8100906FF9144A7FB05FAE3FB42" box="[1187,1322,1232,1256]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9144A7FB05FB73FB4D" box="[1187,1210,1232,1255]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. ocellatus
</emphasis>
(Morice)
</taxonomicName>
,
<taxonomicName id="ACBB5F810906FF914093FB21FD94FAA7" authority="David, Vogel &amp; Pauwels" authorityName="David, Vogel &amp; Pauwels" box="[151,605,1268,1293]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="saintgironsi">
<emphasis id="59CFF8100906FF914093FB21FEF7FAA6" box="[151,318,1268,1292]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914093FB21FF67FAA1" box="[151,174,1268,1291]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. saintgironsi
</emphasis>
David, Vogel &amp; Pauwels
</taxonomicName>
, and
<taxonomicName id="ACBB5F810906FF914293FB21FCEFFAA6" box="[663,806,1268,1292]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="59CFF8100906FF914293FB21FCEFFAA6" box="[663,806,1268,1292]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914293FB21FD67FAA1" box="[663,686,1268,1291]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. macrurus
</emphasis>
</taxonomicName>
(discussed above). Additionally, from the species of the
<taxonomicName id="ACBB5F810906FF914093FACDFEA2FA9A" box="[151,363,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="59CFF8100906FF914093FACDFEA2FA9A" box="[151,363,1304,1328]" italics="true" pageId="10" pageNumber="221">Oligodon cyclurus</emphasis>
</taxonomicName>
group occurring in Indochina,
<taxonomicName id="ACBB5F810906FF9142D9FACDFC76FA85" box="[733,959,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100906FF9142D9FACDFC76FA85" box="[733,959,1304,1328]" italics="true" pageId="10" pageNumber="221">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0906FF9143CEFAC2FBE1FA9A" box="[970,1064,1303,1328]" pageId="10" pageNumber="221" rank="species">
<emphasis id="59CFF8100906FF9143CEFAC2FBE1FA9A" bold="true" box="[970,1064,1303,1328]" pageId="10" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
differs externally by the dorsal coloration features (all members of the
<taxonomicName id="ACBB5F810906FF914255FAE9FCE8FAFE" box="[593,801,1340,1364]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="59CFF8100906FF914255FAE9FCE8FAFE" box="[593,801,1340,1364]" italics="true" pageId="10" pageNumber="221">Oligodon cyclurus</emphasis>
</taxonomicName>
group except
<taxonomicName id="ACBB5F810906FF9143C5FAE9FB86FAFE" box="[961,1103,1340,1364]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="59CFF8100906FF9143C5FAE9FB86FAFE" box="[961,1103,1340,1364]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9143C5FAE9FC11FAF9" box="[961,984,1340,1363]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. macrurus
</emphasis>
</taxonomicName>
possess blotches, reticulation or crossbars) and by the following characters: from
<taxonomicName id="ACBB5F810906FF9142E1FAB5FC45FADD" box="[741,908,1376,1399]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cattienensis">
<emphasis id="59CFF8100906FF9142E1FAB5FC45FADD" box="[741,908,1376,1399]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9142E1FAB5FD35FADD" box="[741,764,1376,1399]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. cattienensis
</emphasis>
</taxonomicName>
, by the immaculate belly and a lesser ventrals count (131144 vs. 167178); from
<taxonomicName id="ACBB5F810906FF91422AFA51FD71FA36" box="[558,696,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="59CFF8100906FF91422AFA51FD71FA36" box="[558,696,1412,1436]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91422AFA51FD8CFA31" box="[558,581,1412,1435]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. chinensis
</emphasis>
</taxonomicName>
, by a much lesser ventrals count (131144 vs. 170206); from
<taxonomicName id="ACBB5F810906FF914585FA51FED9FA6A" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="fasciolatus">
<emphasis id="59CFF8100906FF914585FA51FED9FA6A" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914585FA51FA51FA31" box="[1409,1432,1412,1435]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. fasciolatus
</emphasis>
</taxonomicName>
, by DSR 17 vs. 2123 and by a lesser ventrals count (131144 vs. 147210); from
<taxonomicName id="ACBB5F810906FF9144CFFA7DFAB8FA6A" box="[1227,1393,1448,1472]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="59CFF8100906FF9144CFFA7DFAB8FA6A" box="[1227,1393,1448,1472]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9144CFFA7DFB2BFA15" box="[1227,1250,1448,1471]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. formosanus
</emphasis>
</taxonomicName>
, by DSR 17 vs. 19 and a lesser ventrals count (131144 vs. 154189); from
<taxonomicName id="ACBB5F810906FF9143BBFA19FB51FA4E" box="[959,1176,1484,1508]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="kampucheaensis">
<emphasis id="59CFF8100906FF9143BBFA19FB51FA4E" box="[959,1176,1484,1508]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9143BBFA19FC1FFA49" box="[959,982,1484,1507]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. kampucheaensis
</emphasis>
</taxonomicName>
, by DSR 17 vs. 15 and a lesser ventrals count (131144 vs. 165); from
<taxonomicName id="ACBB5F810906FF9142BAFA25FC81F9A2" box="[702,840,1520,1544]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="59CFF8100906FF9142BAFA25FC81F9A2" box="[702,840,1520,1544]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9142BAFA25FD1CF9AD" box="[702,725,1520,1543]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. ocellatus
</emphasis>
</taxonomicName>
, by DSR 17 vs. 19, a relatively longer tail in males (TaL/TL 0.260.28 vs. 0.110.14) and a greater subcaudals count (5860 vs.
<quantity id="AC4389E70906FF914401F9C1FBA0F987" box="[1029,1129,1556,1581]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" metricValueMax="11.176" metricValueMin="8.128" pageId="10" pageNumber="221" unit="in" value="38.0" valueMax="44.0" valueMin="32.0">3244 in</quantity>
males, 3640 vs.
<quantity id="AC4389E70906FF91453CF9C1FA55F987" box="[1336,1436,1556,1581]" metricMagnitude="-1" metricUnit="m" metricValue="7.493" metricValueMax="8.382" metricValueMin="6.604" pageId="10" pageNumber="221" unit="in" value="29.5" valueMax="33.0" valueMin="26.0">2633 in</quantity>
females); from
<taxonomicName id="ACBB5F810906FF91414EF9EDFE3CF9E5" box="[330,501,1592,1615]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="saintgironsi">
<emphasis id="59CFF8100906FF91414EF9EDFE3CF9E5" box="[330,501,1592,1615]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91414EF9EDFEA8F9E5" box="[330,353,1592,1615]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. saintgironsi
</emphasis>
</taxonomicName>
, by a lesser ventrals count (131144 vs. 166184), a lesser subcaudals count in females (3640 vs. 53) and a relatively longer tail in males (TaL/TL 0.260.28 vs. 0.190.20).
</paragraph>
<paragraph id="6B0424020906FF9040C3F955FC99FE7A" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1436,151,2013]" lastPageId="11" lastPageNumber="222" pageId="10" pageNumber="221">
From the members of the informal
<taxonomicName id="ACBB5F810906FF91424BF955FCD4F93D" box="[591,797,1664,1688]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="59CFF8100906FF91424BF955FCD4F93D" box="[591,797,1664,1688]" italics="true" pageId="10" pageNumber="221">Oligodon cinereus</emphasis>
</taxonomicName>
group (sensu
<bibRefCitation id="0F2A59F30906FF9143BEF9AAFBF6F932" author="Smith" box="[954,1087,1663,1688]" pageId="10" pageNumber="221" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo - chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
, Green
<emphasis id="59CFF8100906FF914491F954FB04F932" box="[1173,1229,1664,1688]" italics="true" pageId="10" pageNumber="221">et al.</emphasis>
2010,
<bibRefCitation id="0F2A59F30906FF91451DF9AAFF18F916" author="David" pageId="10" pageNumber="221" refString="David, P., Das, I. &amp; Vogel, G. (2011) On some taxonomic and nomenclatural problems in Indian species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae). Zootaxa, 2799, 1 - 14." type="journal article" year="2011">
David
<emphasis id="59CFF8100906FF914560F954FA55F932" box="[1380,1436,1664,1688]" italics="true" pageId="10" pageNumber="221">et al.</emphasis>
2011
</bibRefCitation>
, 2012,
<bibRefCitation id="0F2A59F30906FF914121F970FE3CF916" author="Neang" box="[293,501,1700,1725]" pageId="10" pageNumber="221" refString="Neang, T., Grismer, L. L. &amp; Daltry, J. C. (2012) A new species of kukri snake (Colubridae: Oligodon Fitzinger, 1826) from the Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3388, 41 - 55." type="journal article" year="2012">
Neang
<emphasis id="59CFF8100906FF914173F970FE78F916" box="[375,433,1700,1724]" italics="true" pageId="10" pageNumber="221">et al.</emphasis>
2012
</bibRefCitation>
), which are distributed in Indochina,
<collectingCountry id="13AC64920906FF9143A5F971FC2EF917" box="[929,999,1700,1725]" name="China" pageId="10" pageNumber="221">China</collectingCountry>
,
<collectingCountry id="13AC64920906FF9143F0F971FBF9F917" box="[1012,1072,1700,1725]" name="India" pageId="10" pageNumber="221">India</collectingCountry>
, Andaman and
<collectingCountry id="13AC64920906FF9144E0F971FA51F917" box="[1252,1432,1700,1725]" name="India" pageId="10" pageNumber="221">Nicobar islands</collectingCountry>
, and are characterized by the unforked hemipenes, not spinose but with papillae,
<taxonomicName id="ACBB5F810906FF914416F91DFB27F975" box="[1042,1262,1736,1760]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100906FF914416F91DFB27F975" box="[1042,1262,1736,1760]" italics="true" pageId="10" pageNumber="221">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0906FF9144F0F912FA84F94A" box="[1268,1357,1735,1760]" pageId="10" pageNumber="221" rank="species">
<emphasis id="59CFF8100906FF9144F0F912FA84F94A" bold="true" box="[1268,1357,1735,1760]" pageId="10" pageNumber="221">sp. nov.</emphasis>
</taxonomicNameLabel>
differs by the absence of a loreal (present in all group members except
<taxonomicName id="ACBB5F810906FF914385F939FB45F8AF" authority="Wall" authorityName="Wall" box="[897,1164,1772,1797]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="melanozonatus">
<emphasis id="59CFF8100906FF914385F939FB87F8AE" box="[897,1102,1772,1796]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914385F939FC51F8A9" box="[897,920,1772,1795]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. melanozonatus
</emphasis>
Wall
</taxonomicName>
from
<collectingCountry id="13AC64920906FF9144D1F939FAD9F8AF" box="[1237,1296,1772,1797]" name="India" pageId="10" pageNumber="221">India</collectingCountry>
) and by the following features: from
<taxonomicName id="ACBB5F810906FF9141ABF8C5FD6EF882" authority="Gunther" authorityName="Gunther" box="[431,679,1807,1832]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="59CFF8100906FF9141ABF8C5FDE6F88D" box="[431,559,1808,1831]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9141ABF8C5FE0FF88D" box="[431,454,1808,1831]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. cinereus
</emphasis>
(Günther)
</taxonomicName>
, by a lesser ventrals count (131144 vs. 155186) and by the dorsal coloration pattern (from all subspecies having blotches, reticulations or crossbars except the uniformly colored
<emphasis id="59CFF8100906FF914585F8E1FEDBF8C5" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914585F8E1FA51F8E1" box="[1409,1432,1844,1867]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. c.
<taxonomicName id="ACBB5F810906FF9140B0F88DFEDBF8C5" box="[180,274,1880,1903]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="cinereus">cinereus</taxonomicName>
</emphasis>
); from
<taxonomicName id="ACBB5F810906FF914169F88DFDB6F8DA" authority="Cantor" authorityName="Cantor" box="[365,639,1880,1904]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="albocinctus">
<emphasis id="59CFF8100906FF914169F88DFDDDF8DA" box="[365,532,1880,1904]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914169F88DFE4DF8C5" box="[365,388,1880,1903]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. albocinctus
</emphasis>
(Cantor)
</taxonomicName>
, by a lesser ventrals count (131144 vs. 177207) and by the dorsal coloration pattern (light crossbars in
<taxonomicName id="ACBB5F810906FF914235F8A9FD1DF83E" box="[561,724,1916,1940]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="albocinctus">
<emphasis id="59CFF8100906FF914235F8A9FD1DF83E" box="[561,724,1916,1940]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914235F8A9FD81F839" box="[561,584,1916,1939]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. albocinctus
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810906FF91432CF8A9FB8FF83F" authority="Boulenger" authorityName="Boulenger" box="[808,1094,1916,1941]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="59CFF8100906FF91432CF8A9FC7DF83E" box="[808,948,1916,1940]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91432CF8A9FCF6F839" box="[808,831,1916,1939]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. inornatus
</emphasis>
(Boulenger)
</taxonomicName>
, by DSR 17 vs. 15 and a lesser ventrals count (131144 vs. 171174); from
<taxonomicName id="ACBB5F810906FF91428FF874FCA2F813" authority="Smith" authorityName="Smith" box="[651,875,1952,1977]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="joynsoni">
<emphasis id="59CFF8100906FF91428FF874FCC5F812" box="[651,780,1953,1976]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF91428FF874FD6BF812" box="[651,674,1953,1976]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. joynsoni
</emphasis>
(Smith)
</taxonomicName>
, by a lesser ventrals count (131144 vs. 186195) and the dorsal coloration pattern (blotches and reticulated crossbars in
<taxonomicName id="ACBB5F810906FF9143ACF811FBE1F876" box="[936,1064,1988,2012]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="joynsoni">
<emphasis id="59CFF8100906FF9143ACF811FBE1F876" box="[936,1064,1988,2012]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF9143ACF811FC76F871" box="[936,959,1988,2011]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. joynsoni
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810906FF914478F811FABFF877" authority="Taylor" authorityName="Taylor" box="[1148,1398,1988,2013]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="59CFF8100906FF914478F811FAD8F876" box="[1148,1297,1988,2012]" italics="true" pageId="10" pageNumber="221">
<collectingCountry id="13AC64920906FF914478F811FB5AF871" box="[1148,1171,1988,2011]" name="Iceland" pageId="10" pageNumber="221">O</collectingCountry>
. maculatus
</emphasis>
(Taylor)
</taxonomicName>
, by a lesser ventrals count (131144 vs. 156
<emphasis id="59CFF8100907FF904261FF4CFDBAFF1A" box="[613,627,153,176]" italics="true" pageId="11" pageNumber="222"></emphasis>
164) and the dorsal coloration pattern (dark blotches in
<taxonomicName id="ACBB5F810907FF9044F0FF4DFA42FF1A" box="[1268,1419,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="59CFF8100907FF9044F0FF4DFA42FF1A" box="[1268,1419,152,176]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9044F0FF4DFAC2FF05" box="[1268,1291,152,175]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. maculatus
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810907FF9040D0FF69FE56FF7E" box="[212,415,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="melanozonatus">
<emphasis id="59CFF8100907FF9040D0FF69FE56FF7E" box="[212,415,188,212]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9040D0FF69FF22FF79" box="[212,235,188,211]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. melanozonatus
</emphasis>
</taxonomicName>
, by a lesser ventrals count (131144 vs. 171173) and the dorsal coloration pattern (blackedged or black crossbars in
<taxonomicName id="ACBB5F810907FF9041DEFF35FD6FFF52" box="[474,678,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="melanozonatus">
<emphasis id="59CFF8100907FF9041DEFF35FD6FFF52" box="[474,678,224,248]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9041DEFF35FE38FF5D" box="[474,497,224,247]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. melanozonatus
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810907FF904304FF35FF23FEB7" authority="David, Nguyen, Nguyen, Jiang, Chen, Teynie &amp; Ziegler" authorityName="David, Nguyen, Nguyen, Jiang, Chen, Teynie &amp; Ziegler" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="nagao">
<emphasis id="59CFF8100907FF904304FF35FCA2FF52" box="[768,875,224,248]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904304FF35FCDEFF5D" box="[768,791,224,247]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. nagao
</emphasis>
David, Nguyen, Nguyen, Jiang, Chen, Teynié &amp; Ziegler
</taxonomicName>
, by a lesser ventrals count (131144 vs. 184193), a relatively longer tail in males (TaL/TL 0.260.28 vs. 0.140.15) and the dorsal coloration (butterfly-shaped blotches in
<taxonomicName id="ACBB5F810907FF904390FEFDFC36FEEA" box="[916,1023,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="nagao">
<emphasis id="59CFF8100907FF904390FEFDFC36FEEA" box="[916,1023,296,320]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904390FEFDFC62FE95" box="[916,939,296,319]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. nagao
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810907FF90445EFEFDFABBFEEA" authority="Gunther" authorityName="Gunther" box="[1114,1394,295,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="splendidus">
<emphasis id="59CFF8100907FF90445EFEFDFB3EFEEA" box="[1114,1271,296,320]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90445EFEFDFBB8FE95" box="[1114,1137,296,319]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. splendidus
</emphasis>
(Günther)
</taxonomicName>
, by DSR 17 vs. 21, a single pair of both prefrontals and internasals (up to four pairs of each in
<taxonomicName id="ACBB5F810907FF904492FE99FAF8FECE" box="[1174,1329,332,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="splendidus">
<emphasis id="59CFF8100907FF904492FE99FAF8FECE" box="[1174,1329,332,356]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904492FE99FB64FEC9" box="[1174,1197,332,355]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. splendidus
</emphasis>
</taxonomicName>
), a lesser ventrals count (131144 vs. 169193) and the dorsal coloration (large spots on dorsum in
<taxonomicName id="ACBB5F810907FF90448BFEA5FAE3FE22" box="[1167,1322,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="splendidus">
<emphasis id="59CFF8100907FF90448BFEA5FAE3FE22" box="[1167,1322,368,392]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90448BFEA5FB6FFE2D" box="[1167,1190,368,391]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. splendidus
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810907FF904585FEA5FE56FE07" authority="Sclater" authorityName="Sclater" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="woodmansoni">
<emphasis id="59CFF8100907FF904585FEA5FEFAFE06" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904585FEA5FA51FE2D" box="[1409,1432,368,391]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. woodmansoni
</emphasis>
(Sclater)
</taxonomicName>
, by 8(7) supralabials vs. 6, a lesser ventrals count (131144 vs. 180190) and pale belly coloration (dark brown with white edges in
<taxonomicName id="ACBB5F810907FF904285FE6DFC89FE7A" box="[641,832,440,464]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="woodmansoni">
<emphasis id="59CFF8100907FF904285FE6DFC89FE7A" box="[641,832,440,464]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904285FE6DFD51FE65" box="[641,664,440,463]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. woodmansoni
</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="6B0424020907FF9040C3FE09FA96FC92" blockId="11.[151,1436,151,2013]" pageId="11" pageNumber="222">
The members of the informal
<taxonomicName id="ACBB5F810907FF90421EFE09FD5EFE5E" box="[538,663,476,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="59CFF8100907FF90421EFE09FD5EFE5E" box="[538,663,476,500]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90421EFE09FDF8FE59" box="[538,561,476,499]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. dorsalis
</emphasis>
</taxonomicName>
group (sensu
<bibRefCitation id="0F2A59F30907FF904333FE09FC72FE5E" author="Smith" box="[823,955,476,501]" pageId="11" pageNumber="222" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo - chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
) are distributed in Indochina,
<collectingCountry id="13AC64920907FF904515FE09FA84FE5F" box="[1297,1357,476,501]" name="India" pageId="11" pageNumber="222">India</collectingCountry>
,
<collectingCountry id="13AC64920907FF90455DFE09FA55FE5F" box="[1369,1436,476,501]" name="Nepal" pageId="11" pageNumber="222">Nepal</collectingCountry>
and
<collectingCountry id="13AC64920907FF9040CDFDD5FEF3FDB2" box="[201,314,512,536]" name="Myanmar" pageId="11" pageNumber="222">Myanmar</collectingCountry>
; the species with documented hemipenial morphology are characterized by unforked and usually spinose hemipenes. From all congeners belonging to this group, except
<taxonomicName id="ACBB5F810907FF9043D8FDF1FB91FD96" box="[988,1112,548,572]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="lacroixi">
<emphasis id="59CFF8100907FF9043D8FDF1FB91FD96" box="[988,1112,548,572]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9043D8FDF1FC3AFD91" box="[988,1011,548,571]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. lacroixi
</emphasis>
</taxonomicName>
and
<taxonomicName id="ACBB5F810907FF904491FDF1FAEFFD96" box="[1173,1318,548,572]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="catenatus">
<emphasis id="59CFF8100907FF904491FDF1FAEFFD96" box="[1173,1318,548,572]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904491FDF1FB65FD91" box="[1173,1196,548,571]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. catenatus
</emphasis>
</taxonomicName>
discussed above,
<taxonomicName id="ACBB5F810907FF9040EAFD9DFE18FDF5" box="[238,465,584,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100907FF9040EAFD9DFE18FDF5" box="[238,465,584,608]" italics="true" pageId="11" pageNumber="222">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0907FF9041DBFD92FDF6FDCA" box="[479,575,583,608]" pageId="11" pageNumber="222" rank="species">
<emphasis id="59CFF8100907FF9041DBFD92FDF6FDCA" bold="true" box="[479,575,583,608]" pageId="11" pageNumber="222">sp. nov.</emphasis>
</taxonomicNameLabel>
differs by the dorsal coloration (longitudinal stripes or crossbars in all members of the
<taxonomicName id="ACBB5F810907FF90414BFDB9FE05FD2E" box="[335,460,620,644]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="59CFF8100907FF90414BFDB9FE05FD2E" box="[335,460,620,644]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90414BFDB9FEAFFD29" box="[335,358,620,643]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. dorsalis
</emphasis>
</taxonomicName>
group) and an entire anal plate vs. divided, as well as by the following features of the pholidosis: from
<taxonomicName id="ACBB5F810907FF90415CFD45FD05FD02" authority="Gray &amp; Hardwicke" authorityName="Gray &amp; Hardwicke" box="[344,716,655,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="59CFF8100907FF90415CFD45FE1CFD02" box="[344,469,656,680]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90415CFD45FEA6FD0D" box="[344,367,656,679]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. dorsalis
</emphasis>
(Gray &amp; Hardwicke)
</taxonomicName>
, by DSR 17 vs. 15, the absence of a loreal and a lesser ventrals count (131144 vs. 162188); from
<taxonomicName id="ACBB5F810907FF90422BFD61FCFFFD67" authority="Pellegrin" authorityName="Pellegrin" box="[559,822,692,717]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="eberhardti">
<emphasis id="59CFF8100907FF90422BFD61FD0EFD66" box="[559,711,692,716]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90422BFD61FD8FFD61" box="[559,582,692,715]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. eberhardti
</emphasis>
Pellegrin
</taxonomicName>
, by DSR 17 vs. 13, 8(7) supralabials vs. 6 and a lesser ventrals count (131144 vs. 165191); from
<taxonomicName id="ACBB5F810907FF904292FD0DFC19FD5A" authority="Boulenger" authorityName="Boulenger" box="[662,976,727,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="59CFF8100907FF904292FD0DFC98FD5A" box="[662,849,728,752]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904292FD0DFD64FD45" box="[662,685,728,751]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. erythrogaster
</emphasis>
Boulenger
</taxonomicName>
, by a lesser ventrals count (131144 vs. 163186); from
<taxonomicName id="ACBB5F810907FF904149FD29FD9CFCBF" authority="Boulenger" authorityName="Boulenger" box="[333,597,764,789]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="hamptoni">
<emphasis id="59CFF8100907FF904149FD29FE11FCBE" box="[333,472,764,788]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904149FD29FEADFCB9" box="[333,356,764,787]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. hamptoni
</emphasis>
Boulenger
</taxonomicName>
, by DSR 17 vs. 15, 8(7) supralabials vs. 5 and a lesser ventrals count (131 144 vs. 160175); from
<taxonomicName id="ACBB5F810907FF9041A2FCF5FD4DFC92" authority="Wall" authorityName="Wall" box="[422,644,799,824]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="mcdougalli">
<emphasis id="59CFF8100907FF9041A2FCF5FD8FFC92" box="[422,582,800,824]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9041A2FCF5FE74FC9D" box="[422,445,800,823]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. mcdougalli
</emphasis>
Wall
</taxonomicName>
, by DSR 17 vs. 13 and a lesser ventrals count (131144 vs. 199).
</paragraph>
<paragraph id="6B0424020907FF9040C3FC90FF3BFBD7" blockId="11.[151,1436,151,2013]" pageId="11" pageNumber="222">
In the species of the informal
<taxonomicName id="ACBB5F810907FF904217FC91FD56FCF6" box="[531,671,836,860]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="59CFF8100907FF904217FC91FD56FCF6" box="[531,671,836,860]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904217FC91FDE3FCF1" box="[531,554,836,859]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. torquatus
</emphasis>
</taxonomicName>
group (sensu
<bibRefCitation id="0F2A59F30907FF904338FC91FC08FCF6" author="Smith" box="[828,961,836,861]" pageId="11" pageNumber="222" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo - chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
, Green
<emphasis id="59CFF8100907FF904412FC90FB87FCF6" box="[1046,1102,836,860]" italics="true" pageId="11" pageNumber="222">et al.</emphasis>
2010) with the distribution in
<collectingCountry id="13AC64920907FF904093FCB2FF1AFC2A" box="[151,211,871,896]" name="India" pageId="11" pageNumber="222">India</collectingCountry>
,
<collectingCountry id="13AC64920907FF9040E5FCBDFE87FC2A" box="[225,334,872,896]" name="Myanmar" pageId="11" pageNumber="222">Myanmar</collectingCountry>
and
<collectingCountry id="13AC64920907FF90418CFCB2FE27FC2A" box="[392,494,871,896]" name="Thailand" pageId="11" pageNumber="222">Thailand</collectingCountry>
, hemipenes are unforked with apical papillae.
<taxonomicName id="ACBB5F810907FF904402FCBDFB2CFCD5" box="[1030,1253,872,896]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100907FF904402FCBDFB2CFCD5" box="[1030,1253,872,896]" italics="true" pageId="11" pageNumber="222">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0907FF9044EAFCB2FA83FC2A" box="[1262,1354,871,896]" pageId="11" pageNumber="222" rank="species">
<emphasis id="59CFF8100907FF9044EAFCB2FA83FC2A" bold="true" box="[1262,1354,871,896]" pageId="11" pageNumber="222">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the species of this group by the dorsal coloration features (longitudinal stripes in the
<taxonomicName id="ACBB5F810907FF9044C4FC59FA99FC0E" box="[1216,1360,908,932]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="59CFF8100907FF9044C4FC59FA99FC0E" box="[1216,1360,908,932]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9044C4FC59FB1EFC09" box="[1216,1239,908,931]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. torquatus
</emphasis>
</taxonomicName>
group except
<taxonomicName id="ACBB5F810907FF9040ECFC65FEBEFC62" box="[232,375,944,968]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="planiceps">
<emphasis id="59CFF8100907FF9040ECFC65FEBEFC62" box="[232,375,944,968]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9040ECFC65FF36FC6D" box="[232,255,944,967]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. planiceps
</emphasis>
</taxonomicName>
), an entire anal plate vs. divided and additionally by the following external features: from
<taxonomicName id="ACBB5F810907FF904585FC65FE4FFC47" authority="Gunther" authorityName="Gunther" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="cruentatus">
<emphasis id="59CFF8100907FF904585FC65FEC4FC46" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904585FC65FA51FC6D" box="[1409,1432,944,967]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. cruentatus
</emphasis>
(Günther)
</taxonomicName>
, by a lesser ventrals count (131144 vs. 148173); from
<taxonomicName id="ACBB5F810907FF904403FC01FADAFC47" authority="Boulenger" authorityName="Boulenger" box="[1031,1299,980,1005]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="planiceps">
<emphasis id="59CFF8100907FF904403FC01FB5FFC46" box="[1031,1174,980,1004]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904403FC01FBD7FC41" box="[1031,1054,980,1003]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. planiceps
</emphasis>
Boulenger
</taxonomicName>
, by DSR 17 vs. 13, 8(7) supralabials vs. 5(4), a greater subcaudals count (3660 vs. 2227) and the immaculate ventrals (ventrals with paired dark blotches in
<taxonomicName id="ACBB5F810907FF904247FBC9FD19FB9E" box="[579,720,1052,1076]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="planiceps">
<emphasis id="59CFF8100907FF904247FBC9FD19FB9E" box="[579,720,1052,1076]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904247FBC9FD93FB99" box="[579,602,1052,1075]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. planiceps
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810907FF904323FBC9FBE4FB9F" authority="Gunther" authorityName="Gunther" box="[807,1069,1052,1077]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="theobaldi">
<emphasis id="59CFF8100907FF904323FBC9FC7DFB9E" box="[807,948,1052,1076]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904323FBC9FCF7FB99" box="[807,830,1052,1075]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. theobaldi
</emphasis>
(Günther)
</taxonomicName>
, by the absence of a loreal and a lesser ventrals count (131144 vs. 164180); from
<taxonomicName id="ACBB5F810907FF9042D0FB95FC3BFBF2" authority="Boulenger" authorityName="Boulenger" box="[724,1010,1087,1112]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="59CFF8100907FF9042D0FB95FCA8FBF2" box="[724,865,1088,1112]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9042D0FB95FD22FBFD" box="[724,747,1088,1111]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. torquatus
</emphasis>
(Boulenger)
</taxonomicName>
, by DSR 17 vs. 15 and the absence of a loreal.
</paragraph>
<paragraph id="6B0424020907FF9040C3FB5CFCDDF932" blockId="11.[151,1436,151,2013]" pageId="11" pageNumber="222">
In the species of the informal
<emphasis id="59CFF8100907FF904226FB5DFC9EFB0A" box="[546,855,1160,1184]" italics="true" pageId="11" pageNumber="222">
<taxonomicName id="ACBB5F810907FF904226FB5DFD6EFB0A" box="[546,679,1160,1184]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="venustus">
<collectingCountry id="13AC64920907FF904226FB5DFDF0FB35" box="[546,569,1160,1183]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. venustus
</taxonomicName>
<taxonomicName id="ACBB5F810907FF9042B1FB5DFC9EFB0A" box="[693,855,1160,1184]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="taeniolatus">
<collectingCountry id="13AC64920907FF9042B1FB5DFD05FB35" box="[693,716,1160,1183]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. taeniolatus
</taxonomicName>
</emphasis>
group (sensu
<bibRefCitation id="0F2A59F30907FF9043FEFB52FB48FB0A" author="Smith" box="[1018,1153,1159,1184]" pageId="11" pageNumber="222" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo - chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
, Green
<emphasis id="59CFF8100907FF9044D8FB5CFAD9FB0A" box="[1244,1296,1160,1184]" italics="true" pageId="11" pageNumber="222">et al</emphasis>
. 2010) with distribution in
<collectingCountry id="13AC64920907FF90414CFB79FE4DFB6F" box="[328,388,1196,1221]" name="India" pageId="11" pageNumber="222">India</collectingCountry>
,
<collectingCountry id="13AC64920907FF904192FB79FDC4FB6F" box="[406,525,1196,1221]" name="Sri Lanka" pageId="11" pageNumber="222">Sri Lanka</collectingCountry>
,
<collectingCountry id="13AC64920907FF90421BFB79FD62FB6F" box="[543,683,1196,1221]" name="Afghanistan" pageId="11" pageNumber="222">Afghanistan</collectingCountry>
,
<collectingCountry id="13AC64920907FF9042B8FB79FCD7FB6F" box="[700,798,1196,1221]" name="Pakistan" pageId="11" pageNumber="222">Pakistan</collectingCountry>
,
<collectingCountry id="13AC64920907FF90432BFB79FCBBFB6F" box="[815,882,1196,1221]" name="Nepal" pageId="11" pageNumber="222">Nepal</collectingCountry>
and
<collectingCountry id="13AC64920907FF9043B2FB79FBF2FB6F" box="[950,1083,1196,1221]" name="Bangladesh" pageId="11" pageNumber="222">Bangladesh</collectingCountry>
, hemipenes are unforked and spinose.
<taxonomicName id="ACBB5F810907FF9040FAFB05FE29FB4D" box="[254,480,1232,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100907FF9040FAFB05FE29FB4D" box="[254,480,1232,1256]" italics="true" pageId="11" pageNumber="222">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0907FF9041E8FB1AFD82FB42" box="[492,587,1231,1256]" pageId="11" pageNumber="222" rank="species">
<emphasis id="59CFF8100907FF9041E8FB1AFD82FB42" bold="true" box="[492,587,1231,1256]" pageId="11" pageNumber="222">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the species of this group by the absence of spines on the hemipenis, the dorsal coloration pattern (crossbars or longitudinal stripes or blotches in all species of the
<emphasis id="59CFF8100907FF904585FB21FE61FA9A" italics="true" pageId="11" pageNumber="222">
<taxonomicName id="ACBB5F810907FF904585FB21FF3EFA9A" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="venustus">
<collectingCountry id="13AC64920907FF904585FB21FA51FAA1" box="[1409,1432,1268,1291]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. venustus
</taxonomicName>
<taxonomicName id="ACBB5F810907FF904101FACDFE61FA9A" box="[261,424,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="taeniolatus">
<collectingCountry id="13AC64920907FF904101FACDFED5FA85" box="[261,284,1304,1327]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. taeniolatus
</taxonomicName>
</emphasis>
group), an entire anal plate vs. divided, and the following external characters: from
<taxonomicName id="ACBB5F810907FF904585FACDFE8AFAFF" authority="Gunther" authorityName="Gunther" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="affinis">
<emphasis id="59CFF8100907FF904585FACDFF14FAFE" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904585FACDFA51FA85" box="[1409,1432,1304,1327]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. affinis
</emphasis>
Günther
</taxonomicName>
, by the immaculate belly (with black subrectangular spots in
<taxonomicName id="ACBB5F810907FF9043FEFAE9FBABFAFE" box="[1018,1122,1340,1364]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="affinis">
<emphasis id="59CFF8100907FF9043FEFAE9FBABFAFE" box="[1018,1122,1340,1364]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9043FEFAE9FBD8FAF9" box="[1018,1041,1340,1363]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. affinis
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810907FF9044BEFAE9FA5EFAFF" authority="Shaw" authorityName="Shaw" box="[1210,1431,1340,1365]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="arnensis">
<emphasis id="59CFF8100907FF9044BEFAE9FAF5FAFE" box="[1210,1340,1340,1364]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9044BEFAE9FB18FAF9" box="[1210,1233,1340,1363]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. arnensis
</emphasis>
(Shaw)
</taxonomicName>
, by a lesser ventrals count (131144 vs. 164202); from
<taxonomicName id="ACBB5F810907FF904321FAB5FBE6FAD2" authority="Gunther" authorityName="Gunther" box="[805,1071,1375,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="brevicauda">
<emphasis id="59CFF8100907FF904321FAB5FC0EFAD2" box="[805,967,1376,1400]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904321FAB5FCF5FADD" box="[805,828,1376,1399]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. brevicauda
</emphasis>
Günther
</taxonomicName>
, by the presence of internasals, DSR 17 vs. 15, a lesser ventrals count (131144 vs. 158173) and a greater subcaudals count (3660 vs. 2529); from
<taxonomicName id="ACBB5F810907FF9040D0FA7DFE31FA6A" authority="Linnaeus" authorityName="Linnaeus" box="[212,504,1447,1472]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="calamarius">
<emphasis id="59CFF8100907FF9040D0FA7DFEBCFA6A" box="[212,373,1448,1472]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9040D0FA7DFF22FA15" box="[212,235,1448,1471]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. calamarius
</emphasis>
(Linnaeus)
</taxonomicName>
, by the absence of a loreal and a greater subcaudals count (3660 vs. 2034); from
<taxonomicName id="ACBB5F810907FF904093FA19FE44FA4F" authority="Wall" authorityName="Wall" box="[151,397,1484,1509]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="erythrorachis">
<emphasis id="59CFF8100907FF904093FA19FE99FA4E" box="[151,336,1484,1508]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904093FA19FF67FA49" box="[151,174,1484,1507]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. erythrorachis
</emphasis>
Wall
</taxonomicName>
, by DSR 17 vs. 15, 9(8) infralabials vs. 4 and a lesser ventrals count (131144 vs. 154); from
<taxonomicName id="ACBB5F810907FF904093FA25FEADF9A2" authority="Wall" authorityName="Wall" box="[151,356,1519,1544]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="melaneus">
<emphasis id="59CFF8100907FF904093FA25FEECF9A2" box="[151,293,1520,1544]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904093FA25FF67F9AD" box="[151,174,1520,1543]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. melaneus
</emphasis>
Wall
</taxonomicName>
, by DSR 17 vs. 15, the absence of a loreal and a lesser ventrals count (131144 vs. 152160); from
<taxonomicName id="ACBB5F810907FF9040D0F9C1FD7FF987" authority="Dumeril, Bibron &amp; Dumeril" authorityName="Dumeril, Bibron &amp; Dumeril" box="[212,694,1556,1581]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="sublineatus">
<emphasis id="59CFF8100907FF9040D0F9C1FEBCF986" box="[212,373,1556,1580]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9040D0F9C1FF22F981" box="[212,235,1556,1579]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. sublineatus
</emphasis>
Duméril, Bibron &amp; Duméril
</taxonomicName>
, by DSR 17 vs. 15 and the absence of a loreal; from
<taxonomicName id="ACBB5F810907FF9044FAF9C1FF3EF9FA" authority="Jerdon" authorityName="Jerdon" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="taeniolatus">
<emphasis id="59CFF8100907FF9044FAF9C1FA55F986" box="[1278,1436,1556,1580]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9044FAF9C1FADCF981" box="[1278,1301,1556,1579]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. taeniolatus
</emphasis>
(Jerdon)
</taxonomicName>
, by DSR 17 vs. 15, the absence of a loreal and a lesser ventrals count (131144 vs. 158218); from
<taxonomicName id="ACBB5F810907FF904585F9EDFE62F9DF" authority="Beddome" authorityName="Beddome" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="travancoricus">
<emphasis id="59CFF8100907FF904585F9EDFEFAF9DE" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904585F9EDFA51F9E5" box="[1409,1432,1592,1615]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. travancoricus
</emphasis>
Beddome
</taxonomicName>
, by a lesser ventrals count (131144 vs. 154155); from
<taxonomicName id="ACBB5F810907FF904456F989FA8DF9DF" authority="Jerdon" authorityName="Jerdon" box="[1106,1348,1628,1653]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="venustus">
<emphasis id="59CFF8100907FF904456F989FB11F9DE" box="[1106,1240,1628,1652]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904456F989FBA0F9D9" box="[1106,1129,1628,1651]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. venustus
</emphasis>
(Jerdon)
</taxonomicName>
, by the immaculate belly (with large black spots in
<taxonomicName id="ACBB5F810907FF904285F955FCCAF932" box="[641,771,1664,1688]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="venustus">
<emphasis id="59CFF8100907FF904285F955FCCAF932" box="[641,771,1664,1688]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904285F955FD51F93D" box="[641,664,1664,1687]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. venustus
</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="6B0424020907FF9740C3F970FBBEFB0A" blockId="11.[151,1436,151,2013]" lastBlockId="12.[151,1436,151,1184]" lastPageId="12" lastPageNumber="223" pageId="11" pageNumber="222">
From other species of the genus, which are not clearly assigned to any specific groups of species,
<taxonomicName id="ACBB5F810907FF904536F971FECAF975" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="arenarius" status="sp. nov.">
<emphasis id="59CFF8100907FF904536F971FECAF975" italics="true" pageId="11" pageNumber="222">Oligodon arenarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="42FC456B0907FF904108F912FEA1F94A" box="[268,360,1735,1760]" pageId="11" pageNumber="222" rank="species">
<emphasis id="59CFF8100907FF904108F912FEA1F94A" bold="true" box="[268,360,1735,1760]" pageId="11" pageNumber="222">sp. nov.</emphasis>
</taxonomicNameLabel>
differs by the following characters: from
<taxonomicName id="ACBB5F810907FF904349F91DFBF8F94A" authority="Girard" authorityName="Girard" box="[845,1073,1735,1760]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="ancorus">
<emphasis id="59CFF8100907FF904349F91DFC02F94A" box="[845,971,1736,1760]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904349F91DFCADF975" box="[845,868,1736,1759]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. ancorus
</emphasis>
(Girard)
</taxonomicName>
(the
<collectingCountry id="13AC64920907FF90446BF912FB3BF94A" box="[1135,1266,1735,1760]" name="Philippines" pageId="11" pageNumber="222">Philippines</collectingCountry>
), by the dorsal coloration (large dorsal spots in
<taxonomicName id="ACBB5F810907FF904206F939FDB7F8AE" box="[514,638,1772,1796]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="ancorus">
<emphasis id="59CFF8100907FF904206F939FDB7F8AE" box="[514,638,1772,1796]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904206F939FDD0F8A9" box="[514,537,1772,1795]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. ancorus
</emphasis>
</taxonomicName>
), the absence of a loreal and a lesser ventrals count (131144 vs. 147 173); from
<taxonomicName id="ACBB5F810907FF904119F8C5FDF2F882" authority="Boulenger" authorityName="Boulenger" box="[285,571,1807,1832]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="annulifer">
<emphasis id="59CFF8100907FF904119F8C5FE60F882" box="[285,425,1808,1832]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904119F8C5FEFDF88D" box="[285,308,1808,1831]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. annulifer
</emphasis>
(Boulenger)
</taxonomicName>
(
<collectingCountry id="13AC64920907FF904249F8DAFD70F882" box="[589,697,1807,1832]" name="Malaysia" pageId="11" pageNumber="222">Malaysia</collectingCountry>
: Borneo), by the dorsal coloration (crossbands in
<taxonomicName id="ACBB5F810907FF9044FBF8C5FA45F882" box="[1279,1420,1808,1832]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="annulifer">
<emphasis id="59CFF8100907FF9044FBF8C5FA45F882" box="[1279,1420,1808,1832]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9044FBF8C5FADFF88D" box="[1279,1302,1808,1831]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. annulifer
</emphasis>
</taxonomicName>
), DSR 17 vs. 15 and the absence of a loreal; from
<taxonomicName id="ACBB5F810907FF9042BEF8E1FBEBF8E7" authority="Boie" authorityName="Boie" box="[698,1058,1844,1869]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="bitorquatus">
<emphasis id="59CFF8100907FF9042BEF8E1FC95F8E6" box="[698,860,1844,1868]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF9042BEF8E1FD18F8E1" box="[698,721,1844,1867]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. bitorquatus
</emphasis>
Boie (
<collectingCountry id="13AC64920907FF9043A3F8E1FBD3F8E7" box="[935,1050,1844,1869]" name="Indonesia" pageId="11" pageNumber="222">Indonesia</collectingCountry>
)
</taxonomicName>
, by the dorsal coloration (red and yellow dots in
<taxonomicName id="ACBB5F810907FF904142F88DFE25F8DA" box="[326,492,1880,1904]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="bitorquatus">
<emphasis id="59CFF8100907FF904142F88DFE25F8DA" box="[326,492,1880,1904]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904142F88DFE94F8C5" box="[326,349,1880,1903]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. bitorquatus
</emphasis>
</taxonomicName>
) and the immaculate belly (with black spots in
<taxonomicName id="ACBB5F810907FF90441AF88DFB0AF8DA" box="[1054,1219,1880,1904]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="bitorquatus">
<emphasis id="59CFF8100907FF90441AF88DFB0AF8DA" box="[1054,1219,1880,1904]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90441AF88DFBFCF8C5" box="[1054,1077,1880,1903]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. bitorquatus
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810907FF90451BF88DFEA9F83F" authority="Leong &amp; Grismer" authorityName="Leong &amp; Grismer" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="booliati">
<emphasis id="59CFF8100907FF90451BF88DFA55F8DA" box="[1311,1436,1880,1904]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90451BF88DFAFFF8C5" box="[1311,1334,1880,1903]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. booliati
</emphasis>
Leong &amp; Grismer
</taxonomicName>
(Peninsular
<collectingCountry id="13AC64920907FF9041EBF8A9FD93F83F" box="[495,602,1916,1941]" name="Malaysia" pageId="11" pageNumber="222">Malaysia</collectingCountry>
), by the dorsal coloration (crossbands in
<taxonomicName id="ACBB5F810907FF904428F8A9FB6FF83E" box="[1068,1190,1916,1940]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="booliati">
<emphasis id="59CFF8100907FF904428F8A9FB6FF83E" box="[1068,1190,1916,1940]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904428F8A9FB8AF839" box="[1068,1091,1916,1939]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. booliati
</emphasis>
</taxonomicName>
) and 9(8) infralabials vs. 7; from
<taxonomicName id="ACBB5F810907FF90411CF874FDC1F813" authority="Boulenger" authorityName="Boulenger" box="[280,520,1952,1977]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="everetti">
<emphasis id="59CFF8100907FF90411CF874FE47F812" box="[280,398,1953,1976]" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF90411CF874FEE6F812" box="[280,303,1953,1976]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. everetti
</emphasis>
Boulenger
</taxonomicName>
(
<collectingCountry id="13AC64920907FF90421CF875FD49F813" box="[536,640,1952,1977]" name="Malaysia" pageId="11" pageNumber="222">Malaysia</collectingCountry>
and
<collectingCountry id="13AC64920907FF9042B3F875FCE1F813" box="[695,808,1952,1977]" name="Indonesia" pageId="11" pageNumber="222">Indonesia</collectingCountry>
), by DSR 17 vs. 15 and the absence of a loreal; from
<taxonomicName id="ACBB5F810907FF904585F874FE37F877" authority="(Boulenger) (Indonesia)" authorityName="Boulenger" baseAuthorityName="Boulenger" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="222" phylum="Chordata" rank="species" species="forbesi">
<emphasis id="59CFF8100907FF904585F874FF2CF876" italics="true" pageId="11" pageNumber="222">
<collectingCountry id="13AC64920907FF904585F874FA51F812" box="[1409,1432,1953,1976]" name="Iceland" pageId="11" pageNumber="222">O</collectingCountry>
. forbesi
</emphasis>
(Boulenger) (
<collectingCountry id="13AC64920907FF904187F811FE3FF877" box="[387,502,1988,2013]" name="Indonesia" pageId="11" pageNumber="222">Indonesia</collectingCountry>
)
</taxonomicName>
, by the absence of a loreal, a lesser ventrals count (131144 vs. 150170) and the immaculate belly (ventrals with brown spots in
<taxonomicName id="ACBB5F810900FF9742D3FF4DFC84FF1A" box="[727,845,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="forbesi">
<emphasis id="59CFF8100900FF9742D3FF4DFC84FF1A" box="[727,845,152,176]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9742D3FF4DFD27FF05" box="[727,750,152,175]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. forbesi
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810900FF9743ABFF4DFE59FF7F" authority="Pauwels, Wallach, David &amp; Chanhome" authorityName="Pauwels, Wallach, David &amp; Chanhome" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="jintakunei">
<emphasis id="59CFF8100900FF9743ABFF4DFB8EFF1A" box="[943,1095,152,176]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9743ABFF4DFC0FFF05" box="[943,966,152,175]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. jintakunei
</emphasis>
Pauwels, Wallach, David &amp; Chanhome (
<collectingCountry id="13AC64920900FF97411BFF69FE41FF7F" box="[287,392,188,213]" name="Thailand" pageId="12" pageNumber="223">Thailand</collectingCountry>
)
</taxonomicName>
, by the dorsal coloration (crossbars in
<taxonomicName id="ACBB5F810900FF97433BFF69FC18FF7E" box="[831,977,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="jintakunei">
<emphasis id="59CFF8100900FF97433BFF69FC18FF7E" box="[831,977,188,212]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF97433BFF69FC9FFF79" box="[831,854,188,211]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. jintakunei
</emphasis>
</taxonomicName>
), DSR 17 vs. 15, an entire anal plate and a lesser ventrals count (131144 vs. 189); from
<taxonomicName id="ACBB5F810900FF9742ABFF35FC34FF52" authority="Achardji &amp; Ray" authorityName="Achardji &amp; Ray" box="[687,1021,223,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="kheriensis">
<emphasis id="59CFF8100900FF9742ABFF35FC8AFF52" box="[687,835,224,248]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9742ABFF35FD0FFF5D" box="[687,710,224,247]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. kheriensis
</emphasis>
Achardji &amp; Ray
</taxonomicName>
(
<collectingCountry id="13AC64920900FF974408FF0AFB98FF52" box="[1036,1105,223,248]" name="Nepal" pageId="12" pageNumber="223">Nepal</collectingCountry>
and
<collectingCountry id="13AC64920900FF974483FF0AFB0CFF52" box="[1159,1221,223,248]" name="India" pageId="12" pageNumber="223">India</collectingCountry>
), by the absence of a loreal and a lesser ventrals count (131144 vs. 196); from
<taxonomicName id="ACBB5F810900FF97433CFED1FADDFEB7" authority="Zheng &amp; Huang" authorityName="Zheng &amp; Huang" box="[824,1300,260,285]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="lungshenensis">
<emphasis id="59CFF8100900FF97433CFED1FC31FEB6" box="[824,1016,260,284]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF97433CFED1FC86FEB1" box="[824,847,260,283]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. lungshenensis
</emphasis>
Zheng &amp; Huang (
<collectingCountry id="13AC64920900FF9744C0FED1FAC5FEB7" box="[1220,1292,260,285]" name="China" pageId="12" pageNumber="223">China</collectingCountry>
)
</taxonomicName>
, by DSR 17 vs. 15, 8(7) supralabials vs. 6, 9(8) infralabials vs. 6 and a lesser ventrals count (131144 vs. 163179); from
<taxonomicName id="ACBB5F810900FF974585FEFDFE73FECF" authority="Steindachner" authorityName="Steindachner" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="meyerinkii">
<emphasis id="59CFF8100900FF974585FEFDFEC4FECE" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974585FEFDFA51FE95" box="[1409,1432,296,319]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. meyerinkii
</emphasis>
(Steindachner)
</taxonomicName>
(the
<collectingCountry id="13AC64920900FF9741FEFE99FDB1FECF" box="[506,632,332,357]" name="Philippines" pageId="12" pageNumber="223">Philippines</collectingCountry>
and
<collectingCountry id="13AC64920900FF9742B0FE99FCD6FECF" box="[692,799,332,357]" name="Malaysia" pageId="12" pageNumber="223">Malaysia</collectingCountry>
), by the absence of a loreal and a lesser ventrals count (131144 vs. 154169); from
<taxonomicName id="ACBB5F810900FF9741E3FEA5FD1DFE22" authority="Gunther" authorityName="Gunther" box="[487,724,367,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="modestus">
<emphasis id="59CFF8100900FF9741E3FEA5FDB8FE22" box="[487,625,368,392]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9741E3FEA5FE37FE2D" box="[487,510,368,391]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. modestus
</emphasis>
Günther
</taxonomicName>
(the
<collectingCountry id="13AC64920900FF974309FEBAFC46FE22" box="[781,911,367,392]" name="Philippines" pageId="12" pageNumber="223">Philippines</collectingCountry>
), by DSR 17 vs. 15, 8(7) supralabials vs. 6 and a lesser ventrals count (131144 vs. 158176); from
<taxonomicName id="ACBB5F810900FF9742EFFE41FB5EFE07" authority="Whitaker &amp; Dattatri" authorityName="Whitaker &amp; Dattatri" box="[747,1175,404,429]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="nikhili">
<emphasis id="59CFF8100900FF9742EFFE41FC9CFE06" box="[747,853,404,428]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9742EFFE41FCCBFE01" box="[747,770,404,427]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. nikhili
</emphasis>
Whitaker &amp; Dattatri (
<collectingCountry id="13AC64920900FF974454FE41FB46FE07" box="[1104,1167,404,429]" name="India" pageId="12" pageNumber="223">India</collectingCountry>
)
</taxonomicName>
, by DSR 17 vs. 15, the dorsal coloration (longitudinal stripes in
<taxonomicName id="ACBB5F810900FF974270FE6DFD2BFE7A" box="[628,738,440,464]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="nikhili">
<emphasis id="59CFF8100900FF974270FE6DFD2BFE7A" box="[628,738,440,464]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974270FE6DFD42FE65" box="[628,651,440,463]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. nikhili
</emphasis>
</taxonomicName>
) and the immaculate belly (ventrals with black spots in
<taxonomicName id="ACBB5F810900FF974585FE6DFF16FE5E" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="nikhili">
<emphasis id="59CFF8100900FF974585FE6DFF16FE5E" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974585FE6DFA51FE65" box="[1409,1432,440,463]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. nikhili
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810900FF974133FE09FDFDFE5F" authority="Gunther" authorityName="Gunther" box="[311,564,476,501]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="notospilus">
<emphasis id="59CFF8100900FF974133FE09FE07FE5E" box="[311,462,476,500]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974133FE09FE87FE59" box="[311,334,476,499]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. notospilus
</emphasis>
Günther
</taxonomicName>
(the
<collectingCountry id="13AC64920900FF974275FE09FD3DFE5F" box="[625,756,476,501]" name="Philippines" pageId="12" pageNumber="223">Philippines</collectingCountry>
), by the absence of a loreal and the dorsal coloration (large rhombic yellow spots in
<taxonomicName id="ACBB5F810900FF9741ABFDD5FD8FFDB2" box="[431,582,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="notospilus">
<emphasis id="59CFF8100900FF9741ABFDD5FD8FFDB2" box="[431,582,512,536]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9741ABFDD5FE0FFDBD" box="[431,454,512,535]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. notospilus
</emphasis>
</taxonomicName>
); from
<taxonomicName id="ACBB5F810900FF974299FDD5FC1DFDB2" authority="Schneider" authorityName="Schneider" box="[669,980,511,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="octolineatus">
<emphasis id="59CFF8100900FF974299FDD5FC80FDB2" box="[669,841,512,536]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974299FDD5FD7DFDBD" box="[669,692,512,535]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. octolineatus
</emphasis>
(Schneider)
</taxonomicName>
(
<collectingCountry id="13AC64920900FF9743E7FE2AFB82FDB2" box="[995,1099,511,536]" name="Malaysia" pageId="12" pageNumber="223">Malaysia</collectingCountry>
and
<collectingCountry id="13AC64920900FF974480FE2AFB3CFDB2" box="[1156,1269,511,536]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
), by the dorsal coloration (longitudinal stripes in
<taxonomicName id="ACBB5F810900FF974215FDF1FD72FD96" box="[529,699,548,572]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="octolineatus">
<emphasis id="59CFF8100900FF974215FDF1FD72FD96" box="[529,699,548,572]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974215FDF1FDE1FD91" box="[529,552,548,571]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. octolineatus
</emphasis>
</taxonomicName>
), the absence of a loreal and a lesser number of ventrals (131144 vs. 155197); from
<taxonomicName id="ACBB5F810900FF974180FD9DFD7DFDCA" authority="Van Denburgh" authorityName="Van Denburgh" box="[388,692,583,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="59CFF8100900FF974180FD9DFDCBFDCA" box="[388,514,584,608]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974180FD9DFE52FDF5" box="[388,411,584,607]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. ornatus
</emphasis>
Van Denburgh
</taxonomicName>
(
<collectingCountry id="13AC64920900FF9742CDFD92FCC7FDCA" box="[713,782,583,608]" name="China" pageId="12" pageNumber="223">China</collectingCountry>
and
<collectingCountry id="13AC64920900FF97434AFD92FC6CFDCA" box="[846,933,583,608]" name="Taiwan" pageId="12" pageNumber="223">Taiwan</collectingCountry>
), by the dorsal coloration (crossbars in
<taxonomicName id="ACBB5F810900FF974585FD9DFF25FD2E" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="59CFF8100900FF974585FD9DFF25FD2E" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974585FD9DFA51FDF5" box="[1409,1432,584,607]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. ornatus
</emphasis>
</taxonomicName>
), DSR 17 vs. 15 and a lesser number of ventrals (131144 vs. 156182); from
<taxonomicName id="ACBB5F810900FF974480FDB9FAAEFD2F" authority="Taylor" authorityName="Taylor" box="[1156,1383,620,645]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="perkinsi">
<emphasis id="59CFF8100900FF974480FDB9FACBFD2E" box="[1156,1282,620,644]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974480FDB9FB52FD29" box="[1156,1179,620,643]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. perkinsi
</emphasis>
(Taylor)
</taxonomicName>
(the
<collectingCountry id="13AC64920900FF974093FD5AFED3FD02" box="[151,282,655,680]" name="Philippines" pageId="12" pageNumber="223">Philippines</collectingCountry>
), by the absence of a loreal and a lesser number of ventrals (131144 vs. 183188); from
<taxonomicName id="ACBB5F810900FF9744F8FD45FEA9FD67" authority="Roux" authorityName="Roux" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="petronellae">
<emphasis id="59CFF8100900FF9744F8FD45FA55FD02" box="[1276,1436,656,680]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9744F8FD45FADAFD0D" box="[1276,1299,656,679]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. petronellae
</emphasis>
Roux (
<collectingCountry id="13AC64920900FF9740E2FD61FE90FD67" box="[230,345,692,717]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
)
</taxonomicName>
, by the dorsal coloration (dark spots in
<taxonomicName id="ACBB5F810900FF974331FD61FC10FD66" box="[821,985,692,716]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="petronellae">
<emphasis id="59CFF8100900FF974331FD61FC10FD66" box="[821,985,692,716]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974331FD61FC85FD61" box="[821,844,692,715]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. petronellae
</emphasis>
</taxonomicName>
), DSR 17 vs. 15 and the absence of a loreal; from
<taxonomicName id="ACBB5F810900FF974126FD0DFD7EFD5A" authority="Werner" authorityName="Werner" box="[290,695,727,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="praefrontalis">
<emphasis id="59CFF8100900FF974126FD0DFE1CFD5A" box="[290,469,728,752]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974126FD0DFEF0FD45" box="[290,313,728,751]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. praefrontalis
</emphasis>
Werner (
<collectingCountry id="13AC64920900FF974238FD02FD66FD5A" box="[572,687,727,752]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
)
</taxonomicName>
, by the presence of internasals, an entire anal plate vs. divided and a lesser number of ventrals (131144 vs. 183193); from
<taxonomicName id="ACBB5F810900FF974318FD29FBBAFCBF" authority="Jan" authorityName="Jan" box="[796,1139,764,789]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="propinquus">
<emphasis id="59CFF8100900FF974318FD29FC77FCBE" box="[796,958,764,788]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974318FD29FCFAFCB9" box="[796,819,764,787]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. propinquus
</emphasis>
Jan (
<collectingCountry id="13AC64920900FF9743FCFD29FBA2FCBF" box="[1016,1131,764,789]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
)
</taxonomicName>
, by the absence of a loreal and DSR 17 vs. 15; from
<taxonomicName id="ACBB5F810900FF9741BDFCF5FC9CFC92" authority="Werner" authorityName="Werner" box="[441,853,799,824]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="59CFF8100900FF9741BDFCF5FDB8FC92" box="[441,625,800,824]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9741BDFCF5FE19FC9D" box="[441,464,800,823]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. pulcherrimus
</emphasis>
Werner (
<collectingCountry id="13AC64920900FF9742DEFCCAFC84FC92" box="[730,845,799,824]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
)
</taxonomicName>
, by the dorsal coloration (elongate spots on dorsum in
<taxonomicName id="ACBB5F810900FF9740BCFC91FEBAFCF6" box="[184,371,836,860]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="59CFF8100900FF9740BCFC91FEBAFCF6" box="[184,371,836,860]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9740BCFC91FF06FCF1" box="[184,207,836,859]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. pulcherrimus
</emphasis>
</taxonomicName>
), the absence of a loreal and a lesser number of ventrals (131144 vs. 152176); from
<taxonomicName id="ACBB5F810900FF974585FC91FE62FC2A" authority="Schlegel" authorityName="Schlegel" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="purpurascens">
<emphasis id="59CFF8100900FF974585FC91FEE6FC2A" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974585FC91FA51FCF1" box="[1409,1432,836,859]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. purpurascens
</emphasis>
(Schlegel)
</taxonomicName>
(
<collectingCountry id="13AC64920900FF9741BEFCB2FDEBFC2A" box="[442,546,871,896]" name="Thailand" pageId="12" pageNumber="223">Thailand</collectingCountry>
,
<collectingCountry id="13AC64920900FF97422BFCB2FD5CFC2A" box="[559,661,871,896]" name="Malaysia" pageId="12" pageNumber="223">Malaysia</collectingCountry>
and
<collectingCountry id="13AC64920900FF9742CAFCB2FCF6FC2A" box="[718,831,871,896]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
), by the absence of a loreal, DSR 17 vs. 1921 and a lesser number of ventrals (131144 vs. 150210); from
<taxonomicName id="ACBB5F810900FF97431AFC59FBD1FC0F" authority="Gunther" authorityName="Gunther" box="[798,1048,908,933]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="signatus">
<emphasis id="59CFF8100900FF97431AFC59FC69FC0E" box="[798,928,908,932]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF97431AFC59FCFCFC09" box="[798,821,908,931]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. signatus
</emphasis>
(Günther)
</taxonomicName>
(
<collectingCountry id="13AC64920900FF97442DFC59FB51FC0F" box="[1065,1176,908,933]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
and
<collectingCountry id="13AC64920900FF9744D0FC59FAF6FC0F" box="[1236,1343,908,933]" name="Malaysia" pageId="12" pageNumber="223">Malaysia</collectingCountry>
), by the dorsal coloration (yellow spots on dorsum in
<taxonomicName id="ACBB5F810900FF974294FC65FCC6FC6D" box="[656,783,944,967]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="signatus">
<emphasis id="59CFF8100900FF974294FC65FCC6FC6D" box="[656,783,944,967]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974294FC65FD6EFC6D" box="[656,679,944,967]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. signatus
</emphasis>
</taxonomicName>
) and the absence of a loreal; from
<taxonomicName id="ACBB5F810900FF974497FC65FE31FC47" authority="(Dumeril, Bibron &amp; Dumeril) (Indonesia)" authorityName="Dumeril, Bibron &amp; Dumeril" baseAuthorityName="Dumeril, Bibron &amp; Dumeril" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="trilineatus">
<emphasis id="59CFF8100900FF974497FC65FAE1FC62" box="[1171,1320,944,968]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974497FC65FB63FC6D" box="[1171,1194,944,967]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. trilineatus
</emphasis>
(Duméril, Bibron &amp; Duméril) (
<collectingCountry id="13AC64920900FF974179FC01FE39FC47" box="[381,496,980,1005]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
)
</taxonomicName>
, by the dorsal coloration (longitudinal stripes in
<taxonomicName id="ACBB5F810900FF974411FC01FB63FC46" box="[1045,1194,980,1004]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="trilineatus">
<emphasis id="59CFF8100900FF974411FC01FB63FC46" box="[1045,1194,980,1004]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974411FC01FBE5FC41" box="[1045,1068,980,1003]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. trilineatus
</emphasis>
</taxonomicName>
) and the absence of a loreal; from
<taxonomicName id="ACBB5F810900FF974122FC2DFD7DFBBA" authority="(Kopstein) (Indonesia)" authorityName="Kopstein" baseAuthorityName="Kopstein" box="[294,692,1015,1040]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="59CFF8100900FF974122FC2DFE60FBBA" box="[294,425,1016,1040]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF974122FC2DFEF4FBA5" box="[294,317,1016,1039]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. unicolor
</emphasis>
(Kopstein) (
<collectingCountry id="13AC64920900FF97423DFC22FD65FBBA" box="[569,684,1015,1040]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
)
</taxonomicName>
, by the absence of a loreal and a lesser number of ventrals (131 144 vs. 162); from
<taxonomicName id="ACBB5F810900FF97417FFBC9FD49FB9F" authority="Gunther" authorityName="Gunther" box="[379,640,1052,1077]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="59CFF8100900FF97417FFBC9FDD3FB9E" box="[379,538,1052,1076]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF97417FFBC9FE5BFB99" box="[379,402,1052,1075]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. vertebralis
</emphasis>
Günther
</taxonomicName>
(
<collectingCountry id="13AC64920900FF974296FBC9FCC8FB9F" box="[658,769,1052,1077]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
and
<collectingCountry id="13AC64920900FF97433AFBC9FC60FB9F" box="[830,937,1052,1077]" name="Malaysia" pageId="12" pageNumber="223">Malaysia</collectingCountry>
), by the dorsal coloration (white or yellow dorsal spots in
<taxonomicName id="ACBB5F810900FF97413AFB95FE11FBF2" box="[318,472,1088,1112]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="59CFF8100900FF97413AFB95FE11FBF2" box="[318,472,1088,1112]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF97413AFB95FE9CFBFD" box="[318,341,1088,1111]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. vertebralis
</emphasis>
</taxonomicName>
), DSR 17 vs. 15 and the absence of a loreal; from
<taxonomicName id="ACBB5F810900FF97440DFB95FA51FBF2" authority="(Bleeker) (Indonesia)" authorityName="Bleeker" baseAuthorityName="Bleeker" box="[1033,1432,1087,1112]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="waandersi">
<emphasis id="59CFF8100900FF97440DFB95FB57FBF2" box="[1033,1182,1088,1112]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF97440DFB95FBE9FBFD" box="[1033,1056,1088,1111]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. waandersi
</emphasis>
(Bleeker) (
<collectingCountry id="13AC64920900FF974519FBEAFA59FBF2" box="[1309,1424,1087,1112]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
)
</taxonomicName>
, by DSR 17 vs. 15 and a greater number of subcaudals (3660 vs. 1829); from
<taxonomicName id="ACBB5F810900FF97445EFBB1FED0FB0A" authority="David &amp; Vogel" authorityName="David &amp; Vogel" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="wagneri">
<emphasis id="59CFF8100900FF97445EFBB1FB17FBD6" box="[1114,1246,1124,1148]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF97445EFBB1FBB8FBD1" box="[1114,1137,1124,1147]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. wagneri
</emphasis>
David &amp; Vogel (
<collectingCountry id="13AC64920900FF97409AFB52FED8FB0A" box="[158,273,1159,1184]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
)
</taxonomicName>
, by DSR 17 vs. 15 and the dorsal coloration (white crossbars in
<taxonomicName id="ACBB5F810900FF9743EDFB5DFBAEFB35" box="[1001,1127,1160,1183]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="223" phylum="Chordata" rank="species" species="wagneri">
<emphasis id="59CFF8100900FF9743EDFB5DFBAEFB35" box="[1001,1127,1160,1183]" italics="true" pageId="12" pageNumber="223">
<collectingCountry id="13AC64920900FF9743EDFB5DFBC9FB35" box="[1001,1024,1160,1183]" name="Iceland" pageId="12" pageNumber="223">O</collectingCountry>
. wagneri
</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>