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<document id="E818CDA96B406F02A8F2F839178E7526" ID-CLB-Dataset="7404" ID-DOI="10.11646/zootaxa.4951.2.3" ID-GBIF-Dataset="29a4191d-6969-4e29-8e4c-9177e61b17df" ID-ISSN="1175-5326" ID-Zenodo-Dep="4664047" ID-ZooBank="F9842659-4211-4346-B4B2-A61ABBBA7F18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617699682956" checkinUser="plazi" docAuthor="Dupérré, Nadine &amp; Tapia, Elicio" docDate="2021" docId="CD5187F1303A6C43FF14FACFFE5000CE" docLanguage="en" docName="zootaxa.4951.2.3.pdf" docOrigin="Zootaxa 4951 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclosternum gaujoni Simon 1889" docType="treatment" docVersion="5" lastPageNumber="279" masterDocId="3168FF89302A6C57FF83FFFAFFD7024B" masterDocTitle="The endless search for type specimens; illustrations of eleven spider (Araneae Mygalomorphae) species described by Eugène Simon" masterLastPageNumber="282" masterPageNumber="259" pageNumber="275" updateTime="1698935892348" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="555353CED1A825B157FC42AAFF5CF5B1">The endless search for type specimens; illustrations of eleven spider (Araneae Mygalomorphae) species described by Eugène Simon</mods:title>
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<mods:namePart id="949E780209C9AE06284A1BBDDB85829B">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="3289E9E30C54784AD3A5E050A0E1FF0B">Zoological Museum, Center of Natural History, Universität Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. &amp; Research Associate, American Museum of Natural History, New York, NY, U. S. A.</mods:affiliation>
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<mods:namePart id="AA9F1836441D4AA3197C7D81167B6105">Tapia, Elicio</mods:namePart>
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<mods:affiliation id="7AB65AC28CB703128C61E81511FEF408">Fundación OTONGA, Calle Rither y Bolivia Quito, Ecuador. &amp; https: // orcid. org / 0000 - 0001 - 9005 - 5328</mods:affiliation>
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<treatment id="CD5187F1303A6C43FF14FACFFE5000CE" ID-DOI="http://doi.org/10.5281/zenodo.4681789" ID-GBIF-Taxon="180999677" ID-Zenodo-Dep="4681789" LSID="urn:lsid:plazi:treatment:CD5187F1303A6C43FF14FACFFE5000CE" httpUri="http://treatment.plazi.org/id/CD5187F1303A6C43FF14FACFFE5000CE" lastPageId="20" lastPageNumber="279" pageId="16" pageNumber="275">
<subSubSection id="0DE2656C303A6C47FF14FACFFD93071B" box="[151,580,1333,1360]" pageId="16" pageNumber="275" type="nomenclature">
<paragraph id="454736E7303A6C47FF14FACFFD93071B" blockId="16.[151,581,1333,1396]" box="[151,580,1333,1360]" pageId="16" pageNumber="275">
<heading id="1E0F818B303A6C47FF14FACFFD93071B" bold="true" box="[151,580,1333,1360]" fontSize="11" level="1" pageId="16" pageNumber="275" reason="1">
<taxonomicName id="82F84D64303A6C47FF14FACFFD93071B" ID-CoL="32XW6" authority="Simon, 1889" authorityName="Simon" authorityYear="1889" box="[151,580,1333,1360]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="275" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF5303A6C47FF14FACFFD93071B" bold="true" box="[151,580,1333,1360]" pageId="16" pageNumber="275">
<emphasis id="778CEAF5303A6C47FF14FACFFE74071B" bold="true" box="[151,419,1333,1360]" italics="true" pageId="16" pageNumber="275">Cyclosternum gaujoni</emphasis>
Simon, 1889
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DE2656C303A6C47FF14FAA0FD92073F" box="[151,581,1370,1396]" pageId="16" pageNumber="275" type="description">
<paragraph id="454736E7303A6C47FF14FAA0FD92073F" blockId="16.[151,581,1333,1396]" box="[151,581,1370,1396]" pageId="16" pageNumber="275">
<figureCitation id="DDC32A62303A6C47FF14FAA0FEF0073F" box="[151,295,1370,1396]" captionStart="FIGURE 18" captionStartId="17.[151,250,1438,1462]" captionTargetBox="[163,1420,191,1409]" captionTargetId="figure-19@17.[151,1436,181,1414]" captionTargetPageId="17" captionText="FIGURE 18. Cyclosternum gaujoni Simon, 1889 female from Casitagua (Berland det. 1913). A. Carapace, dorsal view. B, C. Internal genitalia dorsal view. Scale bars: A: 1.0mm; B: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4664106" httpUri="https://zenodo.org/record/4664106/files/figure.png" pageId="16" pageNumber="275">Figs 18 A, B</figureCitation>
,
<figureCitation id="DDC32A62303A6C47FEB0FAA0FE5C073F" box="[307,395,1370,1396]" captionStart="FIGURE 19" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[173,1410,193,1905]" captionTargetId="figure-27@18.[151,1436,181,1913]" captionTargetPageId="18" captionText="FIGURE 19. Cyclosternum gaujoni Simon, 1889 female from Riobamba (Berland det. 1913). A. Habitus, dorsal view. B. Habitus, ventral view. Scale bars: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4664108" httpUri="https://zenodo.org/record/4664108/files/figure.png" pageId="16" pageNumber="275">19 A, B</figureCitation>
,
<figureCitation id="DDC32A62303A6C47FE14FAA0FE3C073F" box="[407,491,1370,1396]" captionStart="FIGURE 20" captionStartId="19.[151,250,586,610]" captionTargetBox="[156,1423,186,556]" captionTargetId="figure-19@19.[152,1436,181,562]" captionTargetPageId="19" captionText="FIGURE 20. Cyclosternum gaujoni Simon, 1889 female from Riobamba (Berland det. 1913). A, B. Internal genitalia, dorsal view. Scale bars: 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4664110" httpUri="https://zenodo.org/record/4664110/files/figure.png" pageId="16" pageNumber="275">20A, B</figureCitation>
.
<figureCitation id="DDC32A62303A6C47FE75FAA0FD92073F" box="[502,581,1370,1396]" captionStart="MAP" captionStartId="20.[151,208,1743,1767]" captionTargetBox="[163,1426,674,1712]" captionTargetId="figure-222@20.[151,1436,669,1720]" captionText="MAP. 1. Distribution of Cyclosternum gaujoni (red star, red dots), C. janthium (yellow star) and C. schmardae (green star)." figureDoi="http://doi.org/10.5281/zenodo.4681785" httpUri="https://zenodo.org/record/4681785/files/figure.png" pageId="16" pageNumber="275">Map. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0DE2656C303A6C47FF14FA64FCDA079E" pageId="16" pageNumber="275" type="reference_group">
<paragraph id="454736E7303A6C47FF14FA64FCF707FD" blockId="16.[151,800,1438,1493]" box="[151,800,1438,1462]" pageId="16" pageNumber="275">
<treatmentCitationGroup id="65E811C9303A6C47FF14FA64FCF707FD" box="[151,800,1438,1462]" pageId="16" pageNumber="275">
<treatmentCitation id="C45910F6303A6C47FF14FA64FD9707FD" author="Simon, E." box="[151,576,1438,1462]" page="403" pageId="16" pageNumber="275" year="1889">
<taxonomicName id="82F84D64303A6C47FF14FA64FD9707FD" ID-CoL="32XW6" authority="Simon, 1889 g: 403" authorityName="Simon" authorityYear="1889" box="[151,576,1438,1462]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="275" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF5303A6C47FF14FA64FEA007FD" box="[151,375,1438,1462]" italics="true" pageId="16" pageNumber="275">Cyclosternum gaujoni</emphasis>
<bibRefCitation id="21694B16303A6C47FEFDFA64FD9707FD" author="Simon, E." box="[382,576,1438,1462]" pageId="16" pageNumber="275" pagination="399 - 404" refId="ref7789" refString="Simon, E. (1889 g) Revision des Aviculariidae de la Republique de l'Ecuador. Actes de la Societe Linneenne de Bordeaux, 42, 399 - 404." type="journal article" year="1889">
Simon,
<quantity id="82009B02303A6C47FE4EFA64FDC707FD" box="[461,528,1438,1462]" metricMagnitude="0" metricUnit="kg" metricValue="1.889" pageId="16" pageNumber="275" unit="g" value="1889.0">1889g</quantity>
: 403
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(Description female).
</treatmentCitationGroup>
</paragraph>
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<taxonomicName id="82F84D64303A6C47FF14FA47FD96079E" ID-CoL="32XW6" authority="Berland, 1913 b: 83" authorityName="Berland" authorityPageNumber="83" authorityYear="1913" box="[151,577,1469,1493]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="275" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF5303A6C47FF14FA47FEA0079E" box="[151,375,1469,1493]" italics="true" pageId="16" pageNumber="275">Cyclosternum gaujoni</emphasis>
<bibRefCitation id="21694B16303A6C47FEFDFA47FDC7079E" author="Berland, L." box="[382,528,1469,1493]" pageId="16" pageNumber="275" pagination="78 - 119" refId="ref6401" refString="Berland, L. (1913) Araignees. In: Gauthier-Villars et Cie (Eds.), Mission du Service geographique de l'armee pour la mesure d'un arc du meridien equatorial en Amerique du Sud (1899 - 1906). Tome 10. Ministere de l'instruction publique, Paris, pp. 78 - 119." type="book chapter" year="1913">Berland, 1913</bibRefCitation>
b: 83
</taxonomicName>
(Description male).
</paragraph>
</subSubSection>
<subSubSection id="0DE2656C303A6C43FF14F9F9FE5000CE" lastPageId="20" lastPageNumber="279" pageId="16" pageNumber="275" type="materials_examined">
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<materialsCitation id="F5903CBA303A6C47FF14F9F9FE65040A" ID-GBIF-Occurrence="3071151309" country="Ecuador" location="Loja" pageId="16" pageNumber="275" specimenCount="1" specimenCount-female="1" stateProvince="Loja" typeStatus="holotype">
<emphasis id="778CEAF5303A6C47FF14F9F9FE920456" bold="true" box="[151,325,1539,1565]" pageId="16" pageNumber="275">
<typeStatus id="9A438845303A6C47FF14F9F9FF050456" box="[151,210,1539,1565]" pageId="16" pageNumber="275">Type</typeStatus>
material.
</emphasis>
<specimenCount id="53FEFD6E303A6C47FECEF9FEFEB40455" box="[333,355,1540,1566]" pageId="16" pageNumber="275" type="female"></specimenCount>
. Ceph.th. long.
<quantity id="82009B02303A6C47FD94F9F9FDA90455" box="[535,638,1539,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="16" pageNumber="275" unit="mm" value="13.5">13,5 mm</quantity>
; lat.
<quantity id="82009B02303A6C47FD37F9F9FCCC0455" box="[692,795,1539,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="16" pageNumber="275" unit="mm" value="11.2">11,2 mm</quantity>
. Abd. long.
<quantity id="82009B02303A6C47FC20F9F9FC210455" box="[931,1014,1539,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="275" unit="mm" value="17.0">17 mm</quantity>
; lat.
<quantity id="82009B02303A6C47FBAFF9F9FB440455" box="[1068,1171,1539,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="16" pageNumber="275" unit="mm" value="10.5">10,5 mm</quantity>
.
<collectingCountry id="3DEF7677303A6C47FB1DF9F9FB2C0456" box="[1182,1275,1539,1565]" name="Ecuador" pageId="16" pageNumber="275">Ecuador</collectingCountry>
mérid.:
<collectingRegion id="873CF805303A6C47FAE1F9F9FA4F0456" box="[1378,1432,1539,1565]" country="Ecuador" name="Loja" pageId="16" pageNumber="275">Loja</collectingRegion>
, Amaluza.
<emphasis id="778CEAF5303A6C47FE89F9DDFE65040A" bold="true" box="[266,434,1575,1601]" pageId="16" pageNumber="275">
<taxonomicName id="82F84D64303A6C47FE89F9DDFEE1040A" box="[266,310,1575,1601]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="275" phylum="Arthropoda" rank="class">Not</taxonomicName>
examined
</emphasis>
</materialsCitation>
<emphasis id="778CEAF5303A6C47FE31F9DDFE60040A" bold="true" box="[434,439,1575,1601]" pageId="16" pageNumber="275">.</emphasis>
</paragraph>
<paragraph id="454736E7303A6C47FF44F9B1FE66042E" blockId="16.[151,1436,1539,1782]" box="[199,433,1611,1637]" pageId="16" pageNumber="275">
<emphasis id="778CEAF5303A6C47FF44F9B1FE66042E" bold="true" box="[199,433,1611,1637]" pageId="16" pageNumber="275">Material examined.</emphasis>
</paragraph>
<paragraph id="454736E7303A6C47FF44F995FC3C04C2" blockId="16.[151,1436,1539,1782]" box="[199,1003,1647,1674]" pageId="16" pageNumber="275">
<specimenCount id="53FEFD6E303A6C47FF44F995FF3B04C2" box="[199,236,1647,1673]" pageId="16" pageNumber="275" type="female">1♀</specimenCount>
. Vial 1. label 1. Museum Paris AR 4640.
<taxonomicName id="82F84D64303A6C47FD3EF995FC3C04C2" authority="E. S." box="[701,1003,1647,1673]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="275" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF5303A6C47FD3EF995FC6104C2" box="[701,950,1647,1673]" italics="true" pageId="16" pageNumber="275">Cyclosternum gaujoni</emphasis>
E.S.
</taxonomicName>
</paragraph>
<paragraph id="454736E7303A6C47FF44F96EFC6704E6" blockId="16.[151,1436,1539,1782]" box="[199,944,1683,1710]" pageId="16" pageNumber="275">
L. Berland det. 1913.
<collectingCountry id="3DEF7677303A6C47FE3FF96EFDF404E5" box="[444,547,1684,1710]" name="Ecuador" pageId="16" pageNumber="275">Equateur</collectingCountry>
, Casitagua (
<quantity id="82009B02303A6C47FD28F969FD2B04E5" box="[683,764,1683,1710]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="275" unit="m" value="3600.0">3600m</quantity>
) G. Rivet 1903.
</paragraph>
<paragraph id="454736E7303A6C47FF44F94DFC23049A" blockId="16.[151,1436,1539,1782]" box="[199,1012,1719,1746]" pageId="16" pageNumber="275">
<specimenCount id="53FEFD6E303A6C47FF44F94DFF3B049A" box="[199,236,1719,1745]" pageId="16" pageNumber="275" type="female">1♀</specimenCount>
. Vial 2. Label 1. Museum Paris AR 4641.
<taxonomicName id="82F84D64303A6C47FD45F94DFC23049A" authority="E. S." box="[710,1012,1719,1745]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="275" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF5303A6C47FD45F94DFC68049A" box="[710,959,1719,1745]" italics="true" pageId="16" pageNumber="275">Cyclosternum gaujoni</emphasis>
E.S.
</taxonomicName>
</paragraph>
<paragraph id="454736E7303A6C47FF44F926FC6204BE" blockId="16.[151,1436,1539,1782]" box="[199,949,1755,1782]" pageId="16" pageNumber="275">
L. Berland det. 1913.
<collectingCountry id="3DEF7677303A6C47FE3FF926FDF404BD" box="[444,547,1756,1782]" name="Ecuador" pageId="16" pageNumber="275">Equateur</collectingCountry>
, Riobamba (
<quantity id="82009B02303A6C47FD33F921FCD604BD" box="[688,769,1755,1782]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="275" unit="m" value="3600.0">3600m</quantity>
) Dr. Rivet 1901
</paragraph>
<caption id="1187666F303B6C46FF14FA64FCC90791" ID-DOI="http://doi.org/10.5281/zenodo.4664106" ID-Zenodo-Dep="4664106" httpUri="https://zenodo.org/record/4664106/files/figure.png" pageId="17" pageNumber="276" startId="17.[151,250,1438,1462]" targetBox="[163,1420,191,1409]" targetPageId="17">
<paragraph id="454736E7303B6C46FF14FA64FCC90791" blockId="17.[151,1437,1438,1498]" pageId="17" pageNumber="276">
<emphasis id="778CEAF5303B6C46FF14FA64FEF707FD" bold="true" box="[151,288,1438,1462]" pageId="17" pageNumber="276">FIGURE 18.</emphasis>
<taxonomicName id="82F84D64303B6C46FEA4FA64FD4507FD" authority="Simon, 1889" authorityName="Simon" authorityYear="1889" box="[295,658,1438,1462]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="276" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF5303B6C46FEA4FA64FDDF07FD" box="[295,520,1438,1462]" italics="true" pageId="17" pageNumber="276">Cyclosternum gaujoni</emphasis>
Simon, 1889
</taxonomicName>
female from Casitagua (Berland det. 1913). A. Carapace, dorsal view. B, C. Internal genitalia dorsal view. Scale bars: A: 1.0mm; B: 0.5mm.
</paragraph>
</caption>
<caption id="1187666F30386C45FF14F86BFDF60585" ID-DOI="http://doi.org/10.5281/zenodo.4664108" ID-Zenodo-Dep="4664108" httpUri="https://zenodo.org/record/4664108/files/figure.png" pageId="18" pageNumber="277" startId="18.[151,250,1937,1961]" targetBox="[173,1410,193,1905]" targetPageId="18">
<paragraph id="454736E730386C45FF14F86BFDF60585" blockId="18.[151,1436,1937,1998]" pageId="18" pageNumber="277">
<emphasis id="778CEAF530386C45FF14F86BFEF705E1" bold="true" box="[151,288,1937,1962]" pageId="18" pageNumber="277">FIGURE 19.</emphasis>
<taxonomicName id="82F84D6430386C45FEA4F868FD4605E1" authority="Simon, 1889" authorityName="Simon" authorityYear="1889" box="[295,657,1938,1962]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="277" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530386C45FEA4F868FDD005E2" box="[295,519,1938,1962]" italics="true" pageId="18" pageNumber="277">Cyclosternum gaujoni</emphasis>
Simon, 1889
</taxonomicName>
female from Riobamba (Berland det. 1913). A. Habitus, dorsal view. B. Habitus, ventral view. Scale bars: 1.0mm.
</paragraph>
</caption>
<caption id="1187666F30396C44FF14FDB0FE4000CD" ID-DOI="http://doi.org/10.5281/zenodo.4664110" ID-Zenodo-Dep="4664110" httpUri="https://zenodo.org/record/4664110/files/figure.png" pageId="19" pageNumber="278" startId="19.[151,250,586,610]" targetBox="[156,1423,186,556]" targetPageId="19">
<paragraph id="454736E730396C44FF14FDB0FE4000CD" blockId="19.[151,1436,586,646]" pageId="19" pageNumber="278">
<emphasis id="778CEAF530396C44FF14FDB0FEF60029" bold="true" box="[151,289,586,610]" pageId="19" pageNumber="278">FIGURE 20.</emphasis>
<taxonomicName id="82F84D6430396C44FEABFDB0FD430029" authority="Simon, 1889" authorityName="Simon" authorityYear="1889" box="[296,660,586,610]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FEABFDB0FDDE0029" box="[296,521,586,610]" italics="true" pageId="19" pageNumber="278">Cyclosternum gaujoni</emphasis>
Simon, 1889
</taxonomicName>
female from Riobamba (Berland det. 1913). A, B. Internal genitalia, dorsal view. Scale bars: 0.1mm.
</paragraph>
</caption>
<paragraph id="454736E730396C44FF44FD4FFAC00779" blockId="19.[151,1437,692,2015]" pageId="19" pageNumber="278">
<emphasis id="778CEAF530396C44FF44FD4FFED10084" bold="true" box="[199,262,693,719]" pageId="19" pageNumber="278">Note.</emphasis>
In 1889, Simon described
<taxonomicName id="82F84D6430396C44FDB2FD4FFD7F0085" authorityName="Simon" authorityYear="1889" box="[561,680,693,719]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FDB2FD4FFD7F0085" box="[561,680,693,719]" italics="true" pageId="19" pageNumber="278">C. gaujoni</emphasis>
</taxonomicName>
based on a female specimen from Amazula,
<collectingRegion id="873CF80530396C44FB19FD4EFAED0085" box="[1178,1338,692,718]" country="Ecuador" name="Loja" pageId="19" pageNumber="278">Loja Province</collectingRegion>
, situated in southern
<collectingCountry id="3DEF767730396C44FE9DFD22FEAC00B9" box="[286,379,728,754]" name="Ecuador" pageId="19" pageNumber="278">Ecuador</collectingCountry>
(red star,
<figureCitation id="DDC32A6230396C44FE6CFD23FDE800B8" box="[495,575,729,755]" captionStart="MAP" captionStartId="20.[151,208,1743,1767]" captionTargetBox="[163,1426,674,1712]" captionTargetId="figure-222@20.[151,1436,669,1720]" captionText="MAP. 1. Distribution of Cyclosternum gaujoni (red star, red dots), C. janthium (yellow star) and C. schmardae (green star)." figureDoi="http://doi.org/10.5281/zenodo.4681785" httpUri="https://zenodo.org/record/4681785/files/figure.png" pageId="19" pageNumber="278">Map. 1</figureCitation>
). Unfortunately, the
<typeStatus id="9A43884530396C44FCAEFD23FC8A00B8" box="[813,861,729,755]" pageId="19" pageNumber="278">type</typeStatus>
specimen was on loan and could
<taxonomicName id="82F84D6430396C44FB5CFD23FAD400B8" box="[1247,1283,729,755]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="275" phylum="Arthropoda" rank="class">not</taxonomicName>
be imaged at this time but two other specimens determined as
<taxonomicName id="82F84D6430396C44FD46FD07FCEA015D" authorityName="Simon" authorityYear="1889" box="[709,829,765,791]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FD46FD07FCEA015D" box="[709,829,765,791]" italics="true" pageId="19" pageNumber="278">C. gaujoni</emphasis>
</taxonomicName>
by Lucien Berland from
<collectingCountry id="3DEF767730396C44FBDCFD06FB6B015D" box="[1119,1212,764,790]" name="Ecuador" pageId="19" pageNumber="278">Ecuador</collectingCountry>
were located in the collection. In 1913, Berland mentions a list of specimens he identified as
<taxonomicName id="82F84D6430396C44FC63FCDAFB0D0171" authorityName="Simon" authorityYear="1889" box="[992,1242,800,826]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FC63FCDAFB0D0171" box="[992,1242,800,826]" italics="true" pageId="19" pageNumber="278">Cyclosternum gaujoni</emphasis>
</taxonomicName>
including males. Berland does
<taxonomicName id="82F84D6430396C44FEB1FCBFFE810114" box="[306,342,837,863]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="275" phylum="Arthropoda" rank="class">not</taxonomicName>
describe the male formally, no illustrations or characters are mentioned in the text. Berland mentions only the following specimens and localities: Riobamba (
<specimenCount id="53FEFD6E30396C44FCC7FC93FCB301C8" box="[836,868,873,899]" pageId="19" pageNumber="278" type="female">1♀</specimenCount>
), Quito (
<specimenCount id="53FEFD6E30396C44FC48FC93FC3901C8" box="[971,1006,873,899]" pageId="19" pageNumber="278" type="male">1♂</specimenCount>
), El. Vinculo (
<specimenCount id="53FEFD6E30396C44FB13FC93FB6401C8" box="[1168,1203,873,899]" pageId="19" pageNumber="278" type="male">1♂</specimenCount>
), Casitagua (
<specimenCount id="53FEFD6E30396C44FAC4FC93FAB801C8" box="[1351,1391,873,899]" pageId="19" pageNumber="278" type="male">1♂</specimenCount>
<specimenCount id="53FEFD6E30396C44FAEDFC93FA5B01C8" box="[1390,1420,873,899]" pageId="19" pageNumber="278" type="female">1♀</specimenCount>
). In the
<collectionCode id="23E9AE2230396C44FF61FC77FEEF01EC" box="[226,312,909,935]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="278">MNHN</collectionCode>
collection we were able to locate only one female from Casitagua and one female from Riobamba (
<figureCitation id="DDC32A6230396C44FF23FC4AFF270180" box="[160,240,944,971]" captionStart="FIGURE 18" captionStartId="17.[151,250,1438,1462]" captionTargetBox="[163,1420,191,1409]" captionTargetId="figure-19@17.[151,1436,181,1414]" captionTargetPageId="17" captionText="FIGURE 18. Cyclosternum gaujoni Simon, 1889 female from Casitagua (Berland det. 1913). A. Carapace, dorsal view. B, C. Internal genitalia dorsal view. Scale bars: A: 1.0mm; B: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4664106" httpUri="https://zenodo.org/record/4664106/files/figure.png" pageId="19" pageNumber="278">Fig. 18</figureCitation>
and
<figureCitation id="DDC32A6230396C44FEABFC4AFEA90180" box="[296,382,944,971]" captionStart="FIGURE 19" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[173,1410,193,1905]" captionTargetId="figure-27@18.[151,1436,181,1913]" captionTargetPageId="18" captionText="FIGURE 19. Cyclosternum gaujoni Simon, 1889 female from Riobamba (Berland det. 1913). A. Habitus, dorsal view. B. Habitus, ventral view. Scale bars: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4664108" httpUri="https://zenodo.org/record/4664108/files/figure.png" pageId="19" pageNumber="278">Figs 19</figureCitation>
,
<figureCitation id="DDC32A6230396C44FE09FC4BFE710180" box="[394,422,945,971]" captionStart="FIGURE 20" captionStartId="19.[151,250,586,610]" captionTargetBox="[156,1423,186,556]" captionTargetId="figure-19@19.[152,1436,181,562]" captionTargetPageId="19" captionText="FIGURE 20. Cyclosternum gaujoni Simon, 1889 female from Riobamba (Berland det. 1913). A, B. Internal genitalia, dorsal view. Scale bars: 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4664110" httpUri="https://zenodo.org/record/4664110/files/figure.png" pageId="19" pageNumber="278">20</figureCitation>
respectively) identified by Berland, we assume that the remaining specimens may also be on loan. Because Berland (1931: 83) presented no text description and no illustration, the male of
<taxonomicName id="82F84D6430396C44FB55FC2FFA9B01A5" authorityName="Simon" authorityYear="1889" box="[1238,1356,981,1007]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FB55FC2FFA9B01A5" box="[1238,1356,981,1007]" italics="true" pageId="19" pageNumber="278">C. gaujoni</emphasis>
</taxonomicName>
should be considered as unknown (contra
<collectionCode id="23E9AE2230396C44FD9BFC03FD800658" box="[536,599,1017,1043]" country="0" httpUri="http://grbio.org/cool/cnv0-s4bi" name="Westfield State College, Museum and Herbarium" pageId="19" pageNumber="278">WSC</collectionCode>
, 2020). Mapping the distribution of the material identified by Berland from
<collectingCountry id="3DEF767730396C44FF14FBE6FF21067D" box="[151,246,1052,1078]" name="Ecuador" pageId="19" pageNumber="278">Ecuador</collectingCountry>
, and the
<typeStatus id="9A43884530396C44FEDBFBE7FE5F067C" box="[344,392,1053,1079]" pageId="19" pageNumber="278">type</typeStatus>
locality,
<taxonomicName id="82F84D6430396C44FE6DFBE7FDB3067D" authorityName="Simon" authorityYear="1889" box="[494,612,1053,1079]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FE6DFBE7FDB3067D" box="[494,612,1053,1079]" italics="true" pageId="19" pageNumber="278">C. gaujoni</emphasis>
</taxonomicName>
would be distributed from Northern to Southern
<collectingCountry id="3DEF767730396C44FB05FBE6FB34067D" box="[1158,1251,1052,1078]" name="Ecuador" pageId="19" pageNumber="278">Ecuador</collectingCountry>
covering almost completely the Ecuadorian Andes (red dots and red star,
<figureCitation id="DDC32A6230396C44FC89FBBBFC8F0610" box="[778,856,1089,1115]" captionStart="MAP" captionStartId="20.[151,208,1743,1767]" captionTargetBox="[163,1426,674,1712]" captionTargetId="figure-222@20.[151,1436,669,1720]" captionText="MAP. 1. Distribution of Cyclosternum gaujoni (red star, red dots), C. janthium (yellow star) and C. schmardae (green star)." figureDoi="http://doi.org/10.5281/zenodo.4681785" httpUri="https://zenodo.org/record/4681785/files/figure.png" pageId="19" pageNumber="278">Map. 1</figureCitation>
). Knowing the high diversity and presence of shortrange endemic in the Ecuadorian Andes (
<bibRefCitation id="21694B1630396C44FDD8FB9FFC8E0634" author="Duperre, N. &amp; Tapia, E." box="[603,857,1124,1151]" pageId="19" pageNumber="278" pagination="1 - 81" refId="ref6806" refString="Duperre, N. &amp; Tapia, E. (2020) Megadiverse Ecuador: a review of Mysmenopsis (Araneae, Mysmenidae) of Ecuador, with the description of twenty-one new kleptoparasitic spider species. Zootaxa, 4761 (1), 1 - 81. https: // doi. org / 10.11646 / zootaxa. 4761.1.1" type="journal article" year="2020">Dupérré &amp; Tapia 2020</bibRefCitation>
), and the fact that mygalomorphs have low dispersal capabilities (
<bibRefCitation id="21694B1630396C44FEA6FB72FDA906E8" author="Janowski-Bell, M. E. &amp; Horner, N. V." box="[293,638,1160,1187]" pageId="19" pageNumber="278" pagination="503 - 512" refId="ref7337" refString="Janowski-Bell, M. E. &amp; Horner, N. V. (1990) Movement of the male brown tarantula, Aphonopelma hentzi (Araneae, Theraphosidae), using radio telemetry. Journal of Arachnolgy, 27, 503 - 512." type="journal article" year="1990">Janowski-Bell &amp; Horner 1990</bibRefCitation>
;
<bibRefCitation id="21694B1630396C44FD09FB72FC9F06E8" author="Bond, J. E. &amp; Hedin, M. &amp; Ramirez, M. G. &amp; Opell, B. D." box="[650,840,1160,1187]" pageId="19" pageNumber="278" pagination="899 - 910" refId="ref6513" refString="Bond, J. E., Hedin, M. C, Ramirez, M. G. &amp; Opell, B. D. (2001) Deep molecular divergence in the absence of morphological and ecological change in the Californian coastal dune endemic trapdoor spider Aptostichus simus. Molecular Ecology, 10, 899 - 910. https: // doi. org / 10.1046 / j. 1365 - 294 x. 2001.01233. x" type="journal article" year="2001">
Bond
<emphasis id="778CEAF530396C44FD4DFB73FCD106E9" box="[718,774,1160,1186]" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
2001
</bibRefCitation>
;
<bibRefCitation id="21694B1630396C44FCD7FB72FC0306E8" author="Bond, J. E." box="[852,980,1160,1187]" pageId="19" pageNumber="278" pagination="361 - 376" refId="ref6460" refString="Bond, J. E. (2004) Systematics of the Californian euctenizine spider genus Apomastus (Araneae: Mygalomorphae: Cyrtaucheniidae): the relationship between molecular and morphological taxonomy. Invertebrate Systematics, 18, 361 - 376. https: // doi. org / 10.1071 / is 04008" type="journal article" year="2004">Bond 2004</bibRefCitation>
;
<bibRefCitation id="21694B1630396C44FC5CFB72FAD206E8" author="Hendrixson, B. E. &amp; Bond, J. E." box="[991,1285,1160,1187]" pageId="19" pageNumber="278" pagination="1" refId="ref7282" refString="Hendrixson, B. E., &amp; Bond, J. E. (2005) Two sympatric species ofAntrodiaetus from south- western North Carolina (Araneae, Mygalomorphae, Antrodiaetidae). Zootaxa, 872 (1), 1. https: // doi. org / 10.11646 / zootaxa. 872.1.1" type="journal article" year="2005">Hendrixson &amp; Bond 2005</bibRefCitation>
,
<bibRefCitation id="21694B1630396C44FA92FB72FF22068C" author="Stockman, A. K. &amp; Beamer, D. A. &amp; Bond, J. E." pageId="19" pageNumber="278" refId="ref8147" refString="Stockman, A. K., Beamer, D. A. &amp; Bond, J. E. (2005) An evaluation of a GARP model as an approach to predicting the spatial distribution of non-vagile invertebrate species." type="book" year="2005">
Stockman
<emphasis id="778CEAF530396C44FA0BFB73FF63068D" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
2005
</bibRefCitation>
;
<bibRefCitation id="21694B1630396C44FF7CFB57FE5F068C" author="Raven, R. J." box="[255,392,1197,1223]" pageId="19" pageNumber="278" pagination="11 - 17" refId="ref7639" refString="Raven, R. J. (2010) A review of the Mygalomorphae: biology, morphology and systematics. Book of Abstracts of the 18 th International Congress of Arachnology, Siedle, Poland, 11 - 17 July 2010. [unknown pagination]" type="book chapter" year="2010">Raven 2010</bibRefCitation>
;
<bibRefCitation id="21694B1630396C44FE12FB56FD84068C" author="Hedin, M. &amp; Starrett, J. &amp; Hayashi, C." box="[401,595,1196,1223]" pageId="19" pageNumber="278" pagination="508 - 526" refId="ref7143" refString="Hedin, M., Starrett, J. &amp; Hayashi, C. (2013) Crossing the uncrossable: novel trans-valley biogeographic patterns revealed in the genetic history of low-dispersal mygalo- morph spiders (Antrodiaetidae, Antrodiaetus) from California. Molecular Ecology, 22, 508 - 526. https: // doi. org / 10.1111 / mec. 12130" type="journal article" year="2013">
Hedin
<emphasis id="778CEAF530396C44FE5FFB57FDC5068D" box="[476,530,1196,1222]" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="21694B1630396C44FDDEFB56FCFA068C" author="Ferretti, N. &amp; Peirez-Miles, F. &amp; Gonzailez, A." box="[605,813,1196,1223]" pageId="19" pageNumber="278" pagination="1 - 10" refId="ref6866" refString="Ferretti, N., Peirez-Miles, F. &amp; Gonzailez, A. (2014) Historical relationships among Argentinean biogeographic provinces based on mygalomorph spider distribution data (Araneae: Mygalomorphae). Studies on Neotropical Fauna and Environment, 49, 1 - 10. https: // doi. org / 10.1080 / 01650521.2014.903616" type="journal article" year="2014">
Ferretti
<emphasis id="778CEAF530396C44FD36FB57FD3B068D" box="[693,748,1196,1222]" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
2014
</bibRefCitation>
), it would
<taxonomicName id="82F84D6430396C44FC23FB57FC13068C" box="[928,964,1197,1223]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="275" phylum="Arthropoda" rank="class">not</taxonomicName>
be surprising that the specimens identified as
<taxonomicName id="82F84D6430396C44FF35FB2BFEF906A1" authorityName="Simon" authorityYear="1889" box="[182,302,1233,1259]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FF35FB2BFEF906A1" box="[182,302,1233,1259]" italics="true" pageId="19" pageNumber="278">C. gaujoni</emphasis>
</taxonomicName>
by Berland represent in fact various species. To this day there are no valid illustration of
<taxonomicName id="82F84D6430396C44FAA6FB2BFA4A06A1" authorityName="Simon" authorityYear="1889" box="[1317,1437,1233,1259]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FAA6FB2BFA4A06A1" box="[1317,1437,1233,1259]" italics="true" pageId="19" pageNumber="278">C. gaujoni</emphasis>
</taxonomicName>
because the female
<typeStatus id="9A43884530396C44FEF0FB0FFE740744" box="[371,419,1269,1295]" pageId="19" pageNumber="278">type</typeStatus>
was never illustrated. The males from the Northern part of
<collectingCountry id="3DEF767730396C44FBBDFB0EFB4C0745" box="[1086,1179,1268,1294]" name="Ecuador" pageId="19" pageNumber="278">Ecuador</collectingCountry>
were never illustrated, but more importantly they match with the Southern female and long-range distribution is questionable.
</paragraph>
<paragraph id="454736E730396C43FF44FAC6FDA502B2" blockId="19.[151,1437,692,2015]" lastBlockId="20.[151,1437,151,645]" lastPageId="20" lastPageNumber="279" pageId="19" pageNumber="278">
The genus
<taxonomicName id="82F84D6430396C44FEC2FAC6FE0A071D" authorityName="Ausserer" authorityYear="1871" box="[321,477,1340,1366]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="778CEAF530396C44FEC2FAC6FE0A071D" box="[321,477,1340,1366]" italics="true" pageId="19" pageNumber="278">Cyclosternum</emphasis>
</taxonomicName>
needs a complete and thorough revision, the Ecuadorian species identity and distribution are
<taxonomicName id="82F84D6430396C44FF76FA9BFECE0730" box="[245,281,1377,1403]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="275" phylum="Arthropoda" rank="class">not</taxonomicName>
well established. For example,
<taxonomicName id="82F84D6430396C44FD02FA9AFB990730" authority="Ausserer, 1871" authorityName="Ausserer" authorityYear="1871" box="[641,1102,1376,1403]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="schmardae">
<emphasis id="778CEAF530396C44FD02FA9AFC480731" box="[641,927,1376,1402]" italics="true" pageId="19" pageNumber="278">Cyclosternum schmardae</emphasis>
<bibRefCitation id="21694B1630396C44FC26FA9BFB990730" author="Ausserer, A." box="[933,1102,1377,1403]" pageId="19" pageNumber="278" pagination="117 - 224" refId="ref6273" refString="Ausserer, A. (1871) Beitrage zur Kenntniss der Arachniden-Familie der Territelariae Thorell (Mygalidae Autor). Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 21, 117 - 224, pl. I." type="journal article" year="1871">Ausserer, 1871</bibRefCitation>
</taxonomicName>
was described based on a female specimen from “Cordilleren 4000-5000 ” (
<bibRefCitation id="21694B1630396C44FD30FA7FFCB707D4" author="Ausserer, A." box="[691,864,1413,1439]" pageId="19" pageNumber="278" pagination="117 - 224" refId="ref6273" refString="Ausserer, A. (1871) Beitrage zur Kenntniss der Arachniden-Familie der Territelariae Thorell (Mygalidae Autor). Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 21, 117 - 224, pl. I." type="journal article" year="1871">Ausserer, 1871</bibRefCitation>
; page 193). Simon for some reason seemed to think that the
<typeStatus id="9A43884530396C44FF72FA53FEF60788" box="[241,289,1449,1475]" pageId="19" pageNumber="278">type</typeStatus>
specimen came from Northern
<collectingCountry id="3DEF767730396C44FD00FA52FD350789" box="[643,738,1448,1474]" name="Ecuador" pageId="19" pageNumber="278">Ecuador</collectingCountry>
, Quito and that the species distribution extended until Cuenca,
<collectingRegion id="873CF80530396C44FF14FA37FE9D07AD" box="[151,330,1484,1511]" country="Ecuador" name="Azuay" pageId="19" pageNumber="278">Azuay Province</collectingRegion>
in southern
<collectingCountry id="3DEF767730396C44FE52FA36FDF907AD" box="[465,558,1484,1510]" name="Ecuador" pageId="19" pageNumber="278">Ecuador</collectingCountry>
(green star,
<figureCitation id="DDC32A6230396C44FD36FA37FD2B07AC" box="[693,764,1485,1511]" captionStart="MAP" captionStartId="20.[151,208,1743,1767]" captionTargetBox="[163,1426,674,1712]" captionTargetId="figure-222@20.[151,1436,669,1720]" captionText="MAP. 1. Distribution of Cyclosternum gaujoni (red star, red dots), C. janthium (yellow star) and C. schmardae (green star)." figureDoi="http://doi.org/10.5281/zenodo.4681785" httpUri="https://zenodo.org/record/4681785/files/figure.png" pageId="19" pageNumber="278">Map 1</figureCitation>
.).
<bibRefCitation id="21694B1630396C44FC9AFA36FC6D07AC" author="Simon, E." box="[793,954,1484,1511]" pageId="19" pageNumber="278" pagination="399 - 404" refId="ref7789" refString="Simon, E. (1889 g) Revision des Aviculariidae de la Republique de l'Ecuador. Actes de la Societe Linneenne de Bordeaux, 42, 399 - 404." type="journal article" year="1889">
Simon (
<quantity id="82009B0230396C44FCF3FA37FC6D07AC" box="[880,954,1485,1511]" metricMagnitude="0" metricUnit="kg" metricValue="1.889" pageId="19" pageNumber="278" unit="g" value="1889.0">1889g</quantity>
</bibRefCitation>
; 403) stated: “Décrit de Quito, nous en possedons plusieurs individus de la même localité, .... Los Puentes, Rumipamba aux environ de Quito.... et de Cuenca province de
<collectingRegion id="873CF80530396C44FE9DF9EFFEBC0464" box="[286,363,1557,1583]" country="Ecuador" name="Azuay" pageId="19" pageNumber="278">Azuay</collectingRegion>
.” Thereafter, Simon described the male without mentioning the exact locality and only illustrated the tibial mating spur (
<quantity id="82009B0230396C44FE17F9C3FE080418" box="[404,479,1593,1619]" metricMagnitude="0" metricUnit="kg" metricValue="1.892" pageId="19" pageNumber="278" unit="g" value="1892.0">1892g</quantity>
: 141: fig. 123).
<bibRefCitation id="21694B1630396C44FD13F9C2FCAF0418" author="Bucherl, W. &amp; Timotheo da Costa, A. &amp; Lucas, S." box="[656,888,1592,1619]" pageId="19" pageNumber="278" pagination="117 - 138" refId="ref6665" refString="Bucherl, W., Timotheo da Costa, A. &amp; Lucas, S. (1971) Revisao de alguns tipos de aranhas caranguejeiras (Orthognatha) estabelecidos por Candido de Mello-Leitao e depositados no Museu Nacional do Rio. Memorias do Instituto Butantan, 35, 117 - 138." type="journal article" year="1971">
Bücherl
<emphasis id="778CEAF530396C44FD6CF9C3FCF00419" box="[751,807,1592,1618]" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
(1971)
</bibRefCitation>
illustrated a male from
<collectingCountry id="3DEF767730396C44FB00F9C2FB230419" box="[1155,1268,1592,1618]" name="Colombia" pageId="19" pageNumber="278">Colombia</collectingCountry>
, clearly stating that they cannot be sure that the species is correct because the male is from
<collectingCountry id="3DEF767730396C44FC64F9A6FB8E043D" box="[999,1113,1628,1654]" name="Colombia" pageId="19" pageNumber="278">Colombia</collectingCountry>
; again they do
<taxonomicName id="82F84D6430396C44FA81F9A7FAF1043C" box="[1282,1318,1629,1655]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="275" phylum="Arthropoda" rank="class">not</taxonomicName>
mention a precise locality in
<collectingCountry id="3DEF767730396C44FEEBF97AFE0D04D1" box="[360,474,1664,1690]" name="Colombia" pageId="19" pageNumber="278">Colombia</collectingCountry>
. Gerschman &amp; Shiapelli (1973) also presented an illustration of a male and in fact illustrated a male based on a specimen from Paris (NO. 9985) determined by Simon (maybe the male used by Simon for his description and illustration, because of the illustration of the tibial mating spur match), they also illustrated a female
<collectionCode id="23E9AE2230396C44FF69F917FEE9054C" box="[234,318,1773,1799]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="19" pageNumber="278" type="Museum">MACN</collectionCode>
(No 6545) without mentioning a country of origin or locality. The illustrations presented by
<bibRefCitation id="21694B1630396C44FAC7F916FEC80560" author="Bucherl, W. &amp; Timotheo da Costa, A. &amp; Lucas, S." pageId="19" pageNumber="278" pagination="117 - 138" refId="ref6665" refString="Bucherl, W., Timotheo da Costa, A. &amp; Lucas, S. (1971) Revisao de alguns tipos de aranhas caranguejeiras (Orthognatha) estabelecidos por Candido de Mello-Leitao e depositados no Museu Nacional do Rio. Memorias do Instituto Butantan, 35, 117 - 138." type="journal article" year="1971">
Bücherl
<emphasis id="778CEAF530396C44FF14F8EBFF180561" box="[151,207,1808,1834]" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
(1971)
</bibRefCitation>
and Gerschman &amp; Shiapelli (1973) differ drastically, the male from
<collectingCountry id="3DEF767730396C44FB9AF8EAFB5D0561" box="[1049,1162,1808,1834]" name="Colombia" pageId="19" pageNumber="278">Colombia</collectingCountry>
, identified by
<bibRefCitation id="21694B1630396C44FAAAF8EAFED20504" author="Bucherl, W. &amp; Timotheo da Costa, A. &amp; Lucas, S." pageId="19" pageNumber="278" pagination="117 - 138" refId="ref6665" refString="Bucherl, W., Timotheo da Costa, A. &amp; Lucas, S. (1971) Revisao de alguns tipos de aranhas caranguejeiras (Orthognatha) estabelecidos por Candido de Mello-Leitao e depositados no Museu Nacional do Rio. Memorias do Instituto Butantan, 35, 117 - 138." type="journal article" year="1971">
Bücherl
<emphasis id="778CEAF530396C44FA0BF8EBFF630505" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
(1971)
</bibRefCitation>
as
<taxonomicName id="82F84D6430396C44FEA8F8CFFE750505" authorityName="Simon" authorityYear="1889" box="[299,418,1845,1871]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF530396C44FEA8F8CFFE750505" box="[299,418,1845,1871]" italics="true" pageId="19" pageNumber="278">C. gaujoni</emphasis>
</taxonomicName>
should be considered a doubtful identification, as they clearly stated. In 1996,
<bibRefCitation id="21694B1630396C44FA96F8CEFEF60538" author="Perez-Miles, F. &amp; Lucas, S. M. &amp; Silva Jr., P. I. da &amp; Bertani, R." pageId="19" pageNumber="278" pagination="33 - 68" refId="ref7507" refString="Perez-Miles, F., Lucas, S. M., Silva Jr., P. I. da &amp; Bertani, R. (1996) Systematic revision and cladistic analysis of Theraphosinae (Araneae: Theraphosidae). Mygalomorph, 1, 33 - 68." type="journal article" year="1996">
Pérez-Miles
<emphasis id="778CEAF530396C44FF14F8A3FF070539" box="[151,208,1880,1906]" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
(1996)
</bibRefCitation>
examined and illustrated a male and female (
<bibRefCitation id="21694B1630396C44FC9DF8A2FBFF0539" author="Perez-Miles, F. &amp; Lucas, S. M. &amp; Silva Jr., P. I. da &amp; Bertani, R." box="[798,1064,1880,1906]" pageId="19" pageNumber="278" pagination="33 - 68" refId="ref7507" refString="Perez-Miles, F., Lucas, S. M., Silva Jr., P. I. da &amp; Bertani, R. (1996) Systematic revision and cladistic analysis of Theraphosinae (Araneae: Theraphosidae). Mygalomorph, 1, 33 - 68." type="journal article" year="1996">
Pérez-Miles
<emphasis id="778CEAF530396C44FC2DF8A3FC310539" box="[942,998,1880,1906]" italics="true" pageId="19" pageNumber="278">et al.</emphasis>
1996
</bibRefCitation>
; 46: fig. 12, 13) of
<taxonomicName id="82F84D6430396C44FA82F8A2FEC605DD" authorityName="Ausserer" authorityYear="1871" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="schmardae">
<emphasis id="778CEAF530396C44FA82F8A2FEC605DD" italics="true" pageId="19" pageNumber="278">Cyclosternum schmardae</emphasis>
</taxonomicName>
from Paris (
<collectionCode id="23E9AE2230396C44FE19F887FE3B05DC" box="[410,492,1917,1943]" country="France" httpUri="http://grbio.org/cool/a5xg-x4h4" name="Museum d'Histoire Naturelle Perpignan" pageId="19" pageNumber="278">MHNP</collectionCode>
9885, the same male specimen examined and illustrated by Gerschman &amp;
<bibRefCitation id="21694B1630396C44FAAFF886FF0705F0" author="Schiapelli, R. D." pageId="19" pageNumber="278" pagination="43 - 77" refId="ref7000" refString="Gerschman de P., B. S. &amp; Schiapelli, R. D. (1973) La subfamilia Ischnocolinae (Araneae: Theraphosidae). Revista del Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Entomology, 4, 43 - 77." type="journal article" year="1973">Schiapelli 1973</bibRefCitation>
) and stated that the specimens were the
<typeStatus id="9A43884530396C44FD1EF85BFD0B05F0" box="[669,732,1953,1979]" pageId="19" pageNumber="278">types</typeStatus>
. Those illustrations of
<taxonomicName id="82F84D6430396C44FC5CF85BFBAC05F1" authorityName="Ausserer" authorityYear="1871" box="[991,1147,1952,1979]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="278" phylum="Arthropoda" rank="species" species="schmardae">
<emphasis id="778CEAF530396C44FC5CF85BFBAC05F1" box="[991,1147,1952,1979]" italics="true" pageId="19" pageNumber="278">C. schmardae</emphasis>
</taxonomicName>
do
<taxonomicName id="82F84D6430396C44FB24F85BFB1C05F0" box="[1191,1227,1953,1979]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="275" phylum="Arthropoda" rank="class">not</taxonomicName>
represent the
<typeStatus id="9A43884530396C44FAEFF85BFA4B05F0" box="[1388,1436,1953,1979]" pageId="19" pageNumber="278">type</typeStatus>
specimens of the species, because the species was described by Ausserer. Finally,
<bibRefCitation id="21694B1630396C44FBA6F83EFB1D0594" author="Gabriel, R." box="[1061,1226,1988,2015]" pageId="19" pageNumber="278" pagination="61 - 92" refId="ref6924" refString="Gabriel, R. (2016) Revised taxonomic placement of the species in the Central American genera Davus O. Pickard-Cambridge, 1892, Metriopelma Becker, 1878, and Schizopelma F. O. Pickard-Cambridge, 1897, with comments on species in related genera (Araneae: Theraphosidae). Arachnology, 17 (2), 61 - 92. https: // doi. org / 10.13156 / arac. 2006.17.2.61" type="journal article" year="2016">Gabriel (2016)</bibRefCitation>
presented the only valid illustration for the species, the genitalia of
<typeStatus id="9A438845303E6C43FD3CFF6DFD3802FA" box="[703,751,151,177]" pageId="20" pageNumber="279">type</typeStatus>
specimen described by Ausserer, “
<taxonomicName id="82F84D64303E6C43FBFCFF6DFE95029E" authority="Ausserer, 1871" authorityName="Ausserer" authorityYear="1871" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="schmardae">
<emphasis id="778CEAF5303E6C43FBFCFF6DFA4B02FA" box="[1151,1436,151,177]" italics="true" pageId="20" pageNumber="279">Cyclosternum schmardae</emphasis>
<bibRefCitation id="21694B16303E6C43FF14FF41FE95029E" author="Ausserer, A." box="[151,322,187,213]" pageId="20" pageNumber="279" pagination="117 - 224" refId="ref6273" refString="Ausserer, A. (1871) Beitrage zur Kenntniss der Arachniden-Familie der Territelariae Thorell (Mygalidae Autor). Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 21, 117 - 224, pl. I." type="journal article" year="1871">Ausserer, 1871</bibRefCitation>
</taxonomicName>
,
<collectionCode id="23E9AE22303E6C43FECDFF41FE74029E" box="[334,419,187,213]" pageId="20" pageNumber="279">NHMV</collectionCode>
110 Cordilleren von
<collectingCountry id="3DEF7677303E6C43FD12FF41FD22029E" box="[657,757,187,213]" name="Ecuador" pageId="20" pageNumber="279">Ecuador</collectingCountry>
”. All the male descriptions and illustrations of
<taxonomicName id="82F84D64303E6C43FA82FF41FEC602B2" authorityName="Ausserer" authorityYear="1871" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="schmardae">
<emphasis id="778CEAF5303E6C43FA82FF41FEC602B2" italics="true" pageId="20" pageNumber="279">Cyclosternum schmardae</emphasis>
</taxonomicName>
should be considered doubtful.
</paragraph>
<paragraph id="454736E7303E6C43FF44FEF9FEF2032E" blockId="20.[151,1437,151,645]" pageId="20" pageNumber="279">
Finally, in the same paper
<bibRefCitation id="21694B16303E6C43FE6DFEF9FD410356" author="Simon, E." box="[494,662,259,285]" pageId="20" pageNumber="279" pagination="399 - 404" refId="ref7789" refString="Simon, E. (1889 g) Revision des Aviculariidae de la Republique de l'Ecuador. Actes de la Societe Linneenne de Bordeaux, 42, 399 - 404." type="journal article" year="1889">
Simon (
<quantity id="82009B02303E6C43FDC5FEF9FD590356" box="[582,654,259,285]" metricMagnitude="0" metricUnit="kg" metricValue="1.889" pageId="20" pageNumber="279" unit="g" value="1889.0">1889g</quantity>
)
</bibRefCitation>
also described
<taxonomicName id="82F84D64303E6C43FCC0FEF9FB8F0356" baseAuthorityName="Simon" baseAuthorityYear="1889" box="[835,1112,259,285]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="janthinum">
<emphasis id="778CEAF5303E6C43FCC0FEF9FB8F0356" box="[835,1112,259,285]" italics="true" pageId="20" pageNumber="279">Cyclosternum janthinum</emphasis>
</taxonomicName>
based on a male from Quito, northern
<collectingCountry id="3DEF7677303E6C43FF7DFEDDFE89030A" box="[254,350,295,321]" name="Ecuador" pageId="20" pageNumber="279">Ecuador</collectingCountry>
. Fortunately, the species was re-illustrated by
<bibRefCitation id="21694B16303E6C43FCE6FEDDFBA90309" author="Perez-Miles, F. &amp; Lucas, S. M. &amp; Silva Jr., P. I. da &amp; Bertani, R." box="[869,1150,295,322]" pageId="20" pageNumber="279" pagination="33 - 68" refId="ref7507" refString="Perez-Miles, F., Lucas, S. M., Silva Jr., P. I. da &amp; Bertani, R. (1996) Systematic revision and cladistic analysis of Theraphosinae (Araneae: Theraphosidae). Mygalomorph, 1, 33 - 68." type="journal article" year="1996">
Pérez-Miles
<emphasis id="778CEAF5303E6C43FC70FEDDFBFB030A" box="[1011,1068,295,321]" italics="true" pageId="20" pageNumber="279">et al.</emphasis>
(1996)
</bibRefCitation>
based on the
<typeStatus id="9A438845303E6C43FA99FEDDFA9D030A" box="[1306,1354,295,321]" pageId="20" pageNumber="279">type</typeStatus>
deposited in Paris.
</paragraph>
<paragraph id="454736E7303E6C43FF44FE95FE5000CE" blockId="20.[151,1437,151,645]" pageId="20" pageNumber="279">
<taxonomicName id="82F84D64303E6C43FF44FE95FEB403C2" authorityName="Ausserer" authorityYear="1871" box="[199,355,367,393]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="778CEAF5303E6C43FF44FE95FEB403C2" box="[199,355,367,393]" italics="true" pageId="20" pageNumber="279">Cyclosternum</emphasis>
</taxonomicName>
is further confused by three species described with overlapping distribution in
<collectingCountry id="3DEF7677303E6C43FB55FE95FAE403C2" box="[1238,1331,367,393]" name="Ecuador" pageId="20" pageNumber="279">Ecuador</collectingCountry>
(
<figureCitation id="DDC32A62303E6C43FAC1FE95FA5C03C2" box="[1346,1419,367,393]" captionStart="MAP" captionStartId="20.[151,208,1743,1767]" captionTargetBox="[163,1426,674,1712]" captionTargetId="figure-222@20.[151,1436,669,1720]" captionText="MAP. 1. Distribution of Cyclosternum gaujoni (red star, red dots), C. janthium (yellow star) and C. schmardae (green star)." figureDoi="http://doi.org/10.5281/zenodo.4681785" httpUri="https://zenodo.org/record/4681785/files/figure.png" pageId="20" pageNumber="279">Map 1</figureCitation>
), with the opposite sexes often described by different authors, or
<taxonomicName id="82F84D64303E6C43FCE6FE69FC5E03E6" box="[869,905,403,429]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="275" phylum="Arthropoda" rank="class">not</taxonomicName>
described, from different locality or more than often without clear localities;
<taxonomicName id="82F84D64303E6C43FE60FE4DFD8E039B" authorityName="Simon" authorityYear="1889" box="[483,601,439,465]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="778CEAF5303E6C43FE60FE4DFD8E039B" box="[483,601,439,465]" italics="true" pageId="20" pageNumber="279">C. gaujoni</emphasis>
</taxonomicName>
(female
<typeStatus id="9A438845303E6C43FD38FE4DFD3C039A" box="[699,747,439,465]" pageId="20" pageNumber="279">type</typeStatus>
described by Simon, male unknown,
<typeStatus id="9A438845303E6C43FB0DFE4DFB69039A" box="[1166,1214,439,465]" pageId="20" pageNumber="279">type</typeStatus>
locality
<collectingRegion id="873CF805303E6C43FA9CFE4DFA82039A" box="[1311,1365,439,465]" country="Ecuador" name="Loja" pageId="20" pageNumber="279">Loja</collectingRegion>
, south
<collectingCountry id="3DEF7677303E6C43FF14FE21FF2E03BD" box="[151,249,475,502]" name="Ecuador" pageId="20" pageNumber="279">Ecuador</collectingCountry>
),
<taxonomicName id="82F84D64303E6C43FE8FFE21FE7003BE" authorityName="Ausserer" authorityYear="1871" box="[268,423,475,501]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="schmardae">
<emphasis id="778CEAF5303E6C43FE8FFE21FE7003BE" box="[268,423,475,501]" italics="true" pageId="20" pageNumber="279">C. schmardae</emphasis>
</taxonomicName>
(female
<typeStatus id="9A438845303E6C43FD89FE21FDED03BE" box="[522,570,475,501]" pageId="20" pageNumber="279">type</typeStatus>
described by Ausserer, male described by Simon without locality,
<typeStatus id="9A438845303E6C43FAA6FE21FA8203BE" box="[1317,1365,475,501]" pageId="20" pageNumber="279">type</typeStatus>
locality Cordilleren
<collectingCountry id="3DEF7677303E6C43FEC6FE05FE730051" box="[325,420,511,538]" name="Ecuador" pageId="20" pageNumber="279">Ecuador</collectingCountry>
) and
<taxonomicName id="82F84D64303E6C43FE66FE05FDAD0052" baseAuthorityName="Simon" baseAuthorityYear="1889" box="[485,634,511,537]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="janthinum">
<emphasis id="778CEAF5303E6C43FE66FE05FDAD0052" box="[485,634,511,537]" italics="true" pageId="20" pageNumber="279">C. janthinum</emphasis>
</taxonomicName>
(male
<typeStatus id="9A438845303E6C43FD49FE05FD2D0052" box="[714,762,511,537]" pageId="20" pageNumber="279">type</typeStatus>
described by Simon, female unknown,
<typeStatus id="9A438845303E6C43FB43FE05FB270052" box="[1216,1264,511,537]" pageId="20" pageNumber="279">type</typeStatus>
locality Quito, north
<collectingCountry id="3DEF7677303E6C43FF59FDD9FEEA0075" box="[218,317,547,574]" name="Ecuador" pageId="20" pageNumber="279">Ecuador</collectingCountry>
). To fully resolve the Ecuadorian
<taxonomicName id="82F84D64303E6C43FD3FFDD9FC8F0076" authorityName="Ausserer" authorityYear="1871" box="[700,856,547,573]" class="Arachnida" family="Theraphosidae" genus="Cyclosternum" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="778CEAF5303E6C43FD3FFDD9FC8F0076" box="[700,856,547,573]" italics="true" pageId="20" pageNumber="279">Cyclosternum</emphasis>
</taxonomicName>
puzzle, all
<typeStatus id="9A438845303E6C43FC58FDD9FBDC0076" box="[987,1035,547,573]" pageId="20" pageNumber="279">type</typeStatus>
specimens need to be re-illustrated and re-described, fresh specimens of males and females need to be collected at the
<typeStatus id="9A438845303E6C43FBC8FDBDFBAC002A" box="[1099,1147,583,609]" pageId="20" pageNumber="279">type</typeStatus>
localities to establish the correct sexes pairing.
</paragraph>
</subSubSection>
</treatment>
</document>