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<document id="4A9CDB2DD14A40D0A493BDA2CB6CC804" ID-DOI="10.11646/zootaxa.3889.3.3" ID-GBIF-Dataset="0f472ebf-c9c3-441b-96cc-510a76de5b4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="224919" ID-ZooBank="F5294390-C1C6-4011-89F7-76BBCE641919" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460989340508" checkinUser="plazi" docAuthor="Lewis, John G. E." docDate="2014" docId="C95387E94071F15BB8D70C27C60DFD07" docLanguage="en" docName="zt03889p413.pdf" docOrigin="Zootaxa 3889 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otostigmus foveolatus Verhoeff 1937" docType="treatment" docVersion="8" lastPageNumber="395" masterDocId="356AFF914074F15CB8400A29C734FFBF" masterDocTitle="A review of the orientalis group of the Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)" masterLastPageNumber="413" masterPageNumber="388" pageNumber="393" updateTime="1698561154455" updateUser="plazi">
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<mods:title id="CBB32B15BF6FC3973D972815D6E17B8D">A review of the orientalis group of the Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)</mods:title>
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<mods:namePart id="87C60F0A16890E0F84E4DF4D7E4F3D99">Lewis, John G. E.</mods:namePart>
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<mods:date id="A9AE01EDF1C661E4FA983815C64FB217">2014</mods:date>
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<treatment id="C95387E94071F15BB8D70C27C60DFD07" ID-DOI="http://doi.org/10.5281/zenodo.5661005" ID-GBIF-Taxon="119596222" ID-Zenodo-Dep="5661005" LSID="urn:lsid:plazi:treatment:C95387E94071F15BB8D70C27C60DFD07" httpUri="http://treatment.plazi.org/id/C95387E94071F15BB8D70C27C60DFD07" lastPageId="7" lastPageNumber="395" pageId="5" pageNumber="393">
<subSubSection id="09E065744071F159B8D70C27C613F9F5" pageId="5" pageNumber="393" type="nomenclature">
<paragraph id="414536FF4071F159B8D70C27C554F997" blockId="5.[151,608,1550,1610]" box="[151,608,1550,1576]" pageId="5" pageNumber="393">
<heading id="1A0D81934071F159B8D70C27C554F997" bold="true" box="[151,608,1550,1576]" fontSize="11" level="1" pageId="5" pageNumber="393" reason="1">
<taxonomicName id="86FA4D7C4071F159B8D70C27C554F997" ID-CoL="4B7VN" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[151,608,1550,1576]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="738EEAED4071F159B8D70C27C554F997" bold="true" box="[151,608,1550,1576]" pageId="5" pageNumber="393">
<emphasis id="738EEAED4071F159B8D70C27C697F997" bold="true" box="[151,419,1550,1576]" italics="true" pageId="5" pageNumber="393">Otostigmus foveolatus</emphasis>
<bibRefCitation id="256B4B0E4071F159B9EA0C27C554F997" author="Verhoeff" box="[426,608,1550,1576]" pageId="5" pageNumber="393" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270. [+ plates 13 - 24.]" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="414536FF4071F159B8D70C18C613F9F5" blockId="5.[151,608,1550,1610]" box="[151,295,1585,1610]" pageId="5" pageNumber="393">(Figs 2126)</paragraph>
</subSubSection>
<subSubSection id="09E065744071F159B8D70C52C598F90A" pageId="5" pageNumber="393" type="reference_group">
<paragraph id="414536FF4071F159B8D70C52C598F90A" blockId="5.[151,1182,1658,1717]" pageId="5" pageNumber="393">
<taxonomicName id="86FA4D7C4071F159B8D70C52C597F92F" ID-CoL="4B7VN" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[151,675,1658,1680]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="foveolatus" subGenus="Otostigmus">
<emphasis id="738EEAED4071F159B8D70C52C537F92F" box="[151,515,1658,1680]" italics="true" pageId="5" pageNumber="393">Otostigmus (Otostigmus) foveolatus</emphasis>
<bibRefCitation id="256B4B0E4071F159BA490C53C597F92F" author="Verhoeff" box="[521,675,1658,1680]" pageId="5" pageNumber="393" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270. [+ plates 13 - 24.]" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
</taxonomicName>
, 217, Pl. 13, Fig. 11.
<collectingCountry id="39ED766F4071F159BB3B0C53C4E5F92F" box="[891,977,1658,1680]" name="Malaysia" pageId="5" pageNumber="393">Malacca</collectingCountry>
[
<collectingCountry id="39ED766F4071F159BB9D0C53C31AF92F" box="[989,1070,1658,1680]" name="Malaysia" pageId="5" pageNumber="393">Melaka</collectingCountry>
]
<collectingCountry id="39ED766F4071F159BC7A0C53C3AEF92F" box="[1082,1178,1658,1680]" name="Malaysia" pageId="5" pageNumber="393">Malaysia</collectingCountry>
.
<taxonomicName id="86FA4D7C4071F159B8D70C89C514F90A" ID-CoL="4B7VN" authority="Verhoeff, 1942" authorityName="Verhoeff" authorityYear="1942" box="[151,544,1695,1717]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="738EEAED4071F159B8D70C89C64CF90A" box="[151,376,1695,1717]" italics="true" pageId="5" pageNumber="393">Otostigmus foveolatus</emphasis>
:
<bibRefCitation id="256B4B0E4071F159B9C60CB6C514F90A" author="Verhoeff" box="[390,544,1695,1717]" pageId="5" pageNumber="393" refString="Verhoeff, K. W. (1942) Otostigmen der Mandschurei, Zoologische Anz eiger, 138, 181 - 189." type="journal article" year="1942">Verhoeff, 1942</bibRefCitation>
</taxonomicName>
, 185 (in key).
</paragraph>
</subSubSection>
<subSubSection id="09E065744071F159B8D70CC0C254F8D1" pageId="5" pageNumber="393" type="diagnosis">
<paragraph id="414536FF4071F159B8D70CC0C254F8D1" blockId="5.[151,1437,1769,2015]" pageId="5" pageNumber="393">
<emphasis id="738EEAED4071F159B8D70CC0C625F8BD" bold="true" box="[151,273,1769,1794]" pageId="5" pageNumber="393">Diagnosis.</emphasis>
Length
<quantity id="86029B1A4071F159B9320CC0C6F1F8BE" box="[370,453,1769,1793]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="393" unit="mm" value="37.0">37 mm</quantity>
. Antennae with 17 articles, the basal 2.5 glabrous. Forcipular coxosternal tooth-plates with 4+4 small teeth. Tergites without paramedian sutures, marginate from 10. Sternites also without paramedian sutures. Coxopleuron with a single apical spine, no lateral or dorsal spine. Legs 1 to 3 with two tarsal spurs. Ultimate legs lacking tarsal spur. Ultimate leg prefemur with VL 2, VM
<quantity id="86029B1A4071F159BB810D7CC4D3F8D1" box="[961,999,1877,1902]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="393" unit="in" value="1.0">1 in</quantity>
proximal half and a corner spine.
</paragraph>
</subSubSection>
<subSubSection id="09E065744071F159B8870D50C205F860" pageId="5" pageNumber="393" type="materials_examined">
<paragraph id="414536FF4071F159B8870D50C205F860" blockId="5.[151,1437,1769,2015]" pageId="5" pageNumber="393">
<emphasis id="738EEAED4071F159B8870D50C682F82D" bold="true" box="[199,438,1913,1938]" pageId="5" pageNumber="393">Material examined.</emphasis>
<collectionCode id="27EBAE3A4071F159B9820D50C6CBF82E" box="[450,511,1913,1937]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="393">ZSM</collectionCode>
. Presumed
<typeStatus id="9E41885D4071F159BAC90D50C5DBF82D" box="[649,751,1913,1938]" pageId="5" pageNumber="393" type="holotype">holotype</typeStatus>
. Two slides, the first with head, antennae, mandibles and maxillae and first four segments, the second with segments 20 and 21 and two detached ultimate legs. Trunk segments 7 to 19 missing. Each slide labelled
<taxonomicName id="86FA4D7C4071F159BADA0DEEC4ECF860" authority="Verh." authorityName="Verh." box="[666,984,1990,2015]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="738EEAED4071F159BADA0DEEC4A0F861" box="[666,916,1990,2014]" italics="true" pageId="5" pageNumber="393">Otostigmus foveolatus</emphasis>
Verh.
</taxonomicName>
[Penang Hill] Perak,
<collectingCountry id="39ED766F4071F159BC8A0DEFC219F860" box="[1226,1325,1990,2015]" name="Malaysia" pageId="5" pageNumber="393">Malacca</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="158566774072F15AB8D70D36C3AFF851" httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="6" pageNumber="394" targetBox="[257,1332,188,1789]" targetPageId="6">
<paragraph id="414536FF4072F15AB8D70D36C3AFF851" blockId="6.[151,1437,1823,2030]" pageId="6" pageNumber="394">
<emphasis id="738EEAED4072F15AB8D70D36C679F88A" bold="true" box="[151,333,1823,1846]" pageId="6" pageNumber="394">FIGURES 1220</emphasis>
.
<taxonomicName id="86FA4D7C4072F15AB9190D08C56CF889" box="[345,600,1824,1846]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="394" phylum="Arthropoda" rank="species" species="brevilabiatus">
<emphasis id="738EEAED4072F15AB9190D08C56CF889" box="[345,600,1824,1846]" italics="true" pageId="6" pageNumber="394">Otostigmus brevilabiatus</emphasis>
</taxonomicName>
Holotype.
<emphasis id="738EEAED4072F15ABA880D09C5D5F88A" bold="true" box="[712,737,1824,1845]" pageId="6" pageNumber="394">12</emphasis>
. Antennal article 3 dorsal.
<emphasis id="738EEAED4072F15ABBB40D09C327F88A" bold="true" box="[1012,1043,1824,1845]" pageId="6" pageNumber="394">13.</emphasis>
Distal articles of left second maxillary telopodite.
<emphasis id="738EEAED4072F15AB94A0D17C617F8EC" bold="true" box="[266,291,1854,1875]" pageId="6" pageNumber="394">14</emphasis>
. Pretarsus of right second maxillary telopodite.
<emphasis id="738EEAED4072F15ABB550D17C41AF8EC" bold="true" box="[789,814,1854,1875]" pageId="6" pageNumber="394">15</emphasis>
. Pretarsus of left second maxillary telopodite.
<emphasis id="738EEAED4072F15ABD510D17C21FF8EC" bold="true" box="[1297,1323,1854,1875]" pageId="6" pageNumber="394">16</emphasis>
Forcipular coxosternal tooth-plates.
<emphasis id="738EEAED4072F15AB9D50D74C69AF8CD" bold="true" box="[405,430,1885,1906]" pageId="6" pageNumber="394">17</emphasis>
. Right forcipular trochanteroprefemoral process.
<emphasis id="738EEAED4072F15ABBE20D74C48FF8CD" bold="true" box="[930,955,1885,1906]" pageId="6" pageNumber="394">18</emphasis>
. Left forcipular trochanteroprefemoral process.
<emphasis id="738EEAED4072F15AB8D70D55C784F82E" bold="true" box="[151,176,1916,1937]" pageId="6" pageNumber="394">19</emphasis>
. Right coxopleural process.
<emphasis id="738EEAED4072F15AB9960D55C6DBF82E" bold="true" box="[470,495,1916,1937]" pageId="6" pageNumber="394">20</emphasis>
. Prefemur left ultimate leg ventral.
<emphasis id="738EEAED4072F15ABB210D52C32CF82E" bold="true" box="[865,1048,1915,1938]" pageId="6" pageNumber="394">FIGURES 2126</emphasis>
.
<taxonomicName id="86FA4D7C4072F15ABC660D55C23DF82D" box="[1062,1289,1916,1938]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="394" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="738EEAED4072F15ABC660D55C23DF82D" box="[1062,1289,1916,1938]" italics="true" pageId="6" pageNumber="394">Otostigmus foveolatus</emphasis>
</taxonomicName>
Holotype.
<emphasis id="738EEAED4072F15ABD3C0D55C2A1F82E" bold="true" box="[1404,1429,1916,1937]" pageId="6" pageNumber="394">21</emphasis>
. Antennal articles 13 ventral.
<emphasis id="738EEAED4072F15AB99C0DB3C6C1F810" bold="true" box="[476,501,1946,1967]" pageId="6" pageNumber="394">22</emphasis>
. Antennal article 3 dorsal.
<emphasis id="738EEAED4072F15ABB600DB3C40DF810" bold="true" box="[800,825,1946,1967]" pageId="6" pageNumber="394">23</emphasis>
. Pretarsus of second maxillary telopodite.
<emphasis id="738EEAED4072F15ABD480DB3C215F810" bold="true" box="[1288,1313,1946,1967]" pageId="6" pageNumber="394">24</emphasis>
. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process.
<emphasis id="738EEAED4072F15ABBEE0D90C4F3F871" bold="true" box="[942,967,1977,1998]" pageId="6" pageNumber="394">25</emphasis>
. Right coxopleural process.
<emphasis id="738EEAED4072F15ABCB30D90C238F871" bold="true" box="[1267,1292,1977,1998]" pageId="6" pageNumber="394">26</emphasis>
. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs15, 16, 20, 21 &amp; 24 = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="09E065744073F15BB8870ABEC3C4FD91" pageId="7" pageNumber="395" type="description">
<paragraph id="414536FF4073F15BB8870ABEC52CFE84" blockId="7.[151,1437,151,696]" pageId="7" pageNumber="395">
<emphasis id="738EEAED4073F15BB8870ABEC66DFF0F" bold="true" box="[199,345,151,176]" pageId="7" pageNumber="395">Description.</emphasis>
(Verhoeffs (1937) data in parentheses where relevant). (Length
<quantity id="86029B1A4073F15BBC6C0AB1C34BFF0F" box="[1068,1151,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="395" unit="mm" value="37.0">37 mm</quantity>
). Antennae of 17 articles, the basal 2.5 glabrous dorsally and ventrally (Fig. 21, 22). Pretarsus of second maxilla without accessory spines (Figs 23), a poorly sclerotised spur on article 2. Forcipular coxosternal tooth-plates with 4+4 teeth, the inner and outer two on each side well separated (Fig. 24), the dorsal surface with several setae. Trochanteroprefemoral process with three small denticles.
</paragraph>
<paragraph id="414536FF4073F15BB8870B6CC544FE57" blockId="7.[151,1437,151,696]" pageId="7" pageNumber="395">(Tergites weakly marginate from 10, lacking [paramedian] sutures but with a median ridge delineated by two shallow sulci, otherwise smooth). Most sternites with a posterior deep rounded depression, with a very weak depression on each side, also a weak longitudinal groove in the middle but sutures absent. Sternite of ultimate legbearing segment posteriorly attenuated and broadly emarginate). Coxopleural process short, with a single apical spine, no lateral or dorsal spines (Fig. 25).</paragraph>
<paragraph id="414536FF4073F15BB8870BDAC3C4FD91" blockId="7.[151,1437,151,696]" pageId="7" pageNumber="395">
(Leg 1 with one tibial spur, two tarsal spurs only on legs 13). One tarsal spur on legs 19 and 20, 21 without. Ultimate leg prefemoral spines VL 2, VM
<quantity id="86029B1A4073F15BBA34083FC5ABFD91" box="[628,671,533,558]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="395" unit="in" value="1.0">1 in</quantity>
proximal half, the spines on swollen bases (Fig. 26).
</paragraph>
</subSubSection>
<subSubSection id="09E065744073F15BB887081EC60DFD07" pageId="7" pageNumber="395" type="discussion">
<paragraph id="414536FF4073F15BB887081EC60DFD07" blockId="7.[151,1437,151,696]" pageId="7" pageNumber="395">
<emphasis id="738EEAED4073F15BB887081EC600FDEF" bold="true" box="[199,308,567,592]" pageId="7" pageNumber="395">Remarks</emphasis>
.
<bibRefCitation id="256B4B0E4073F15BB977081EC6C4FDEF" author="Verhoeff" box="[311,496,567,592]" pageId="7" pageNumber="395" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270. [+ plates 13 - 24.]" type="journal article" year="1937">Verhoeff (1937)</bibRefCitation>
commented that this species comes near to
<taxonomicName id="86FA4D7C4073F15BBBB10811C39AFDEF" authority="Kraepelin" authorityName="Kraepelin" box="[1009,1198,567,592]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="395" phylum="Arthropoda" rank="species" species="oatesi">
<emphasis id="738EEAED4073F15BBBB10811C300FDF0" box="[1009,1076,568,591]" italics="true" pageId="7" pageNumber="395">oatesi</emphasis>
Kraepelin
</taxonomicName>
, adding, perhaps the forms should be joined as races. However, it may be differentiated from
<taxonomicName id="86FA4D7C4073F15BBB850872C31FFDCD" box="[965,1067,603,626]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="395" phylum="Arthropoda" rank="species" species="oatesi">
<emphasis id="738EEAED4073F15BBB850872C31FFDCD" box="[965,1067,603,626]" italics="true" pageId="7" pageNumber="395">O. oatesi</emphasis>
</taxonomicName>
by its lack of tergite and sternite paramedian sutures, the short coxopleural process with a single apical spine and ultimate prefemoral spines on swollen bases.
</paragraph>
</subSubSection>
</treatment>
</document>