165 lines
17 KiB
XML
165 lines
17 KiB
XML
<document id="D098D83AC2C63E162B486EC5C1E22491" ID-DOI="10.11646/zootaxa.5322.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8203657" ID-ZooBank="DD295EE6-AB3F-4377-900E-CC76F1C6BC2D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1690885128246" checkinUser="plazi" docAuthor="Klompen, Hans & Gerdeman, Beverly S." docDate="2023" docId="C55587C9FFDAB82A86C9CA50F3A1FAC2" docLanguage="en" docName="zootaxa.5322.1.1.pdf" docOrigin="Zootaxa 5322 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5322.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heterozerconidae Berlese 1892" docType="treatment" docVersion="5" lastPageNumber="21" masterDocId="396CFFB1FFCEB83E865EC861F07CFFD3" masterDocTitle="Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="21" updateTime="1699549639056" updateUser="plazi">
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<mods:titleInfo id="A894CE5AE1E67AA833DBBE03220BBE39">
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<mods:title id="68FDD14E4D4C5C2C34C45DF460FF2450">Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)</mods:title>
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<mods:name id="349F6CB0E912C30018FA39A8864D12FB" type="personal">
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<mods:namePart id="CCC4516B7B2A183462528F58E8D52597">Klompen, Hans</mods:namePart>
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<mods:nameIdentifier id="F279BF69FDDA6AA46D8011BEA6BB9110" type="email">klompen.1@osu.edu</mods:nameIdentifier>
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<mods:namePart id="D5A75209E70AC366E5F69B5C6EAAABF2">Gerdeman, Beverly S.</mods:namePart>
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<mods:nameIdentifier id="93DA75E67135A9D0D769987874CA0CE7" type="email">klompen.1@osu.edu</mods:nameIdentifier>
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<mods:title id="E3ADDC3FCD7D684D2F585C5F07AED059">Zootaxa</mods:title>
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<mods:part id="99D1D4876AD237866A1EA389E790F5EC">
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<mods:date id="D2796F86788A19AC45611F6429668FB5">2023</mods:date>
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<mods:number id="B2E50C770039EED24867537A2D78B8DB">2023-07-28</mods:number>
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<mods:detail id="A6A94CE13F4134F96A6EE4A104E86ACB" type="volume">
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<mods:number id="A27B33035EFBC79598CABE3AC1CAEE1F">5322</mods:number>
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<mods:start id="5596D1F2170084CA19CA595865E3A8AA">1</mods:start>
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<mods:end id="A43B73318ABEE71DCD780B041556F217">66</mods:end>
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<mods:url id="1028CC5ED3D071A5EA9BAE6F2A3207C6">http://dx.doi.org/10.11646/zootaxa.5322.1.1</mods:url>
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<mods:classification id="8B24A87D10B15FDEF0F1834C05707DA6">journal article</mods:classification>
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<mods:identifier id="B54668A992AD66CEEC37573EA4852A79" type="DOI">10.11646/zootaxa.5322.1.1</mods:identifier>
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<mods:identifier id="F1831A779D74F5AF9BFF24B853A35D50" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3BB5522D622E5498B9B7482D6E0241A8" type="Zenodo-Dep">8203657</mods:identifier>
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<treatment id="C55587C9FFDAB82A86C9CA50F3A1FAC2" ID-DOI="http://doi.org/10.5281/zenodo.8203502" ID-Zenodo-Dep="8203502" LSID="urn:lsid:plazi:treatment:C55587C9FFDAB82A86C9CA50F3A1FAC2" httpUri="http://treatment.plazi.org/id/C55587C9FFDAB82A86C9CA50F3A1FAC2" lastPageNumber="21" pageId="20" pageNumber="21">
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<subSubSection id="05E66554FFDAB82A86C9CA50F265FD98" box="[151,537,561,587]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="4D4336DFFFDAB82A86C9CA50F265FD98" blockId="20.[151,537,561,587]" box="[151,537,561,587]" pageId="20" pageNumber="21">
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<heading id="160B81B3FFDAB82A86C9CA50F265FD98" bold="true" box="[151,537,561,587]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="3">
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<taxonomicName id="8AFC4D5CFFDAB82A86C9CA50F265FD98" ID-CoL="624TK" ID-ENA="122028" authority="Berlese 1892" authorityName="Berlese" authorityYear="1892" box="[151,537,561,587]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">
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||
<emphasis id="7F88EACDFFDAB82A86C9CA50F265FD98" bold="true" box="[151,537,561,587]" pageId="20" pageNumber="21">
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Heterozerconidae
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<bibRefCitation id="296D4B2EFFDAB82A8725CA50F265FD98" author="Berlese, A." box="[379,537,561,587]" pageId="20" pageNumber="21" refId="ref37026" refString="Berlese, A. (1892) Acari, Myriapoda et Scorpiones hucusque in Italia reperta. Ordo Mesostigmata (Gamasidae). Tipografia del Seminario, Padua, 143 pp., XI pls." type="book" year="1892">Berlese 1892</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="05E66554FFDAB82A86C9CA11F180FD5A" box="[151,508,624,649]" pageId="20" pageNumber="21" type="reference_group">
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||
<paragraph id="4D4336DFFFDAB82A86C9CA11F180FD5A" blockId="20.[151,1077,624,649]" box="[151,508,624,649]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="6DEC11F1FFDAB82A86C9CA11F180FD5A" box="[151,508,624,649]" pageId="20" pageNumber="21">
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||
<treatmentCitation id="CC5D10CEFFDAB82A86C9CA11F180FD5A" author="Berlese, A." box="[151,508,624,649]" page="97" pageId="20" pageNumber="21" year="1892">
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<taxonomicName id="8AFC4D5CFFDAB82A86C9CA11F180FD5A" ID-CoL="624TK" authority="Berlese 1892: 97" authorityName="Berlese" authorityPageNumber="97" authorityYear="1892" box="[151,508,624,649]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">
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Heterozerconidae
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<bibRefCitation id="296D4B2EFFDAB82A870ECA11F180FD5A" author="Berlese, A." box="[336,508,624,649]" pageId="20" pageNumber="21" refId="ref37026" refString="Berlese, A. (1892) Acari, Myriapoda et Scorpiones hucusque in Italia reperta. Ordo Mesostigmata (Gamasidae). Tipografia del Seminario, Padua, 143 pp., XI pls." type="book" year="1892">Berlese 1892: 97</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="05E66554FFDAB82A845CCA10F449FD5A" box="[514,1077,624,649]" pageId="20" pageNumber="21" type="type_taxon">
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<paragraph id="4D4336DFFFDAB82A845CCA10F449FD5A" blockId="20.[151,1077,624,649]" box="[514,1077,624,649]" pageId="20" pageNumber="21">
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[
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<typeStatus id="9247887DFFDAB82A8456CA10F24AFD5A" box="[520,566,625,649]" pageId="20" pageNumber="21">type</typeStatus>
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genus:
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<taxonomicName id="8AFC4D5CFFDAB82A84DBCA10F3E5FD5A" authority="Berlese 1892" authorityName="Berlese" authorityYear="1892" box="[645,921,624,649]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A84DBCA10F371FD5B" box="[645,781,625,648]" italics="true" pageId="20" pageNumber="21">Heterozercon</emphasis>
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<bibRefCitation id="296D4B2EFFDAB82A854DCA11F3E5FD5A" author="Berlese, A." box="[787,921,624,649]" pageId="20" pageNumber="21" refId="ref37026" refString="Berlese, A. (1892) Acari, Myriapoda et Scorpiones hucusque in Italia reperta. Ordo Mesostigmata (Gamasidae). Tipografia del Seminario, Padua, 143 pp., XI pls." type="book" year="1892">Berlese 1892</bibRefCitation>
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</taxonomicName>
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by monotypy].
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</paragraph>
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</subSubSection>
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<subSubSection id="05E66554FFDAB82A86C9CAD6F219FC7A" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="4D4336DFFFDAB82A86C9CAD6F219FC7A" blockId="20.[151,1436,695,1045]" pageId="20" pageNumber="21">
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Diagnosis. Movable digit of the chelicera in the female elongate (length> 6x basal width) (3); spermatodactyl present on fixed digit of the male (6); axial outgrowth of palp trochanter in female distinct and membranous (8); palp femur seta
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<emphasis id="7F88EACDFFDAB82A8714CA9EF108FCCA" box="[330,372,767,793]" italics="true" pageId="20" pageNumber="21">pd1</emphasis>
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added (10); lateral lips enlarged to form a trough (19); distance from setae
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<emphasis id="7F88EACDFFDAB82A82E2CA9EF48EFCCA" box="[1212,1266,767,793]" italics="true" pageId="20" pageNumber="21">hyp3</emphasis>
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to
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<emphasis id="7F88EACDFFDAB82A8348CA9EF551FCCB" box="[1302,1325,767,792]" italics="true" pageId="20" pageNumber="21">sc</emphasis>
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distinctly larger than distance from
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<emphasis id="7F88EACDFFDAB82A87E9CB42F191FCEE" box="[439,493,803,829]" italics="true" pageId="20" pageNumber="21">hyp3</emphasis>
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to
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<emphasis id="7F88EACDFFDAB82A844FCB42F23BFCEE" box="[529,583,803,829]" italics="true" pageId="20" pageNumber="21">hyp2</emphasis>
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(21.2, reversed in
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<taxonomicName id="8AFC4D5CFFDAB82A8544CB42F387FCEE" authorityName="Gerdeman & Klompen" authorityYear="2003" box="[794,1019,803,829]" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A8544CB42F387FCEE" box="[794,1019,803,829]" italics="true" pageId="20" pageNumber="21">Narceoheterozercon</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="8AFC4D5CFFDAB82A8257CB42F4D7FCEF" box="[1033,1195,803,828]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A8257CB42F4D7FCEF" box="[1033,1195,803,828]" italics="true" pageId="20" pageNumber="21">Maracazercon</emphasis>
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</taxonomicName>
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, many
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<taxonomicName id="8AFC4D5CFFDAB82A82A1CB42F5E9FCEF" authorityName="Berlese" authorityYear="1892" box="[1279,1429,803,828]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A82A1CB42F5E9FCEF" box="[1279,1429,803,828]" italics="true" pageId="20" pageNumber="21">Heterozercon</emphasis>
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</taxonomicName>
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, and some
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<taxonomicName id="8AFC4D5CFFDAB82A8752CB26F1B1FCB2" authorityName="Fain" authorityYear="1989" box="[268,461,839,865]" class="Arachnida" family="Heterozerconidae" genus="Afroheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A8752CB26F1B1FCB2" box="[268,461,839,865]" italics="true" pageId="20" pageNumber="21">Afroheterozercon</emphasis>
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</taxonomicName>
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); setal base-like structure on hypostome present (25); opisthogastral suckers in adults present, posterior to ventral shield (63.1, reversed in
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<taxonomicName id="8AFC4D5CFFDAB82A8480CB0AF30EFC56" authorityName="Klompen & Gerdeman" authorityYear="2023" box="[734,882,875,901]" class="Arachnida" family="Heterozerconidae" genus="Amyzozercon" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A8480CB0AF30EFC56" box="[734,882,875,901]" italics="true" pageId="20" pageNumber="21">Amyzozercon</emphasis>
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</taxonomicName>
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); marginal opisthosomal setae all setiform (77.0); femora I seta
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<emphasis id="7F88EACDFFDAB82A876ECBEEF136FC7B" box="[304,330,911,936]" italics="true" pageId="20" pageNumber="21">av</emphasis>
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in male spine-like (104).
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</paragraph>
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</subSubSection>
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<subSubSection id="05E66554FFDAB82A8699CBD2F3A1FAC2" pageId="20" pageNumber="21" type="discussion">
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<paragraph id="4D4336DFFFDAB82A8699CBD2F140FBC6" blockId="20.[151,1436,695,1045]" pageId="20" pageNumber="21">
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Remarks. The family is well characterized by a broad set of modifications of the female and male chelicera and hypostome and, possibly, by the presence of a pair of large ventral suckers posterior to the ventral shield (63 but see
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<taxonomicName id="8AFC4D5CFFDAB82A86C9CB9AF157FBC6" authorityName="Klompen & Gerdeman" authorityYear="2023" box="[151,299,1019,1045]" class="Arachnida" family="Heterozerconidae" genus="Amyzozercon" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A86C9CB9AF157FBC6" box="[151,299,1019,1045]" italics="true" pageId="20" pageNumber="21">Amyzozercon</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="4D4336DFFFDAB82A86C9CC22F3A1FAC2" blockId="20.[151,1436,1091,1297]" pageId="20" pageNumber="21">
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The grouping of
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<taxonomicName id="8AFC4D5CFFDAB82A870ACC22F194FB8E" authorityName="Klompen & Gerdeman" authorityYear="2023" box="[340,488,1091,1117]" class="Arachnida" family="Heterozerconidae" genus="Amyzozercon" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A870ACC22F194FB8E" box="[340,488,1091,1117]" italics="true" pageId="20" pageNumber="21">Amyzozercon</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="8AFC4D5CFFDAB82A8440CC22F2DDFB8F" authorityName="Klompen & Gerdeman" authorityYear="2023" box="[542,673,1091,1116]" class="Arachnida" family="Heterozerconidae" genus="Ecuazercon" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A8440CC22F2DDFB8F" box="[542,673,1091,1116]" italics="true" pageId="20" pageNumber="21">Ecuazercon</emphasis>
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</taxonomicName>
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is supported by a few unusual characters: basal part of fixed digit in female elongate, length width ratio 4–6 (
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<figureCitation id="D5C72A5AFFDAB82A8404CC06F2E1FB52" box="[602,669,1127,1153]" captionStart="FIGURES 1–3" captionStartId="19.[151,264,1241,1265]" captionTargetBox="[154,1433,309,1212]" captionTargetId="figure-64@19.[151,1436,306,1215]" captionTargetPageId="19" captionText="FIGURES 1–3. Chelicera females: 1, Discozerconidae, Discozercon derricki., female (OSAL 052739); 2, Heterozerconidae, Amyzozercon chocoensis, female (OSAL 106741); 3, Al. (Philippinozercon) makilingensis, female (OSAL 053222; blue arrow: brush like structure on inside movable digit). Abbreviations: pd: pilus dentilus. Red arrows: thin, strap-like extensions on the movable digit." figureDoi="http://doi.org/10.5281/zenodo.8203659" httpUri="https://zenodo.org/record/8203659/files/figure.png" pageId="20" pageNumber="21">Fig. 2</figureCitation>
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) (1); posterior dorsal setae
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<emphasis id="7F88EACDFFDAB82A8593CC06F46AFB52" box="[973,1046,1127,1153]" italics="true" pageId="20" pageNumber="21">Z2–Z4</emphasis>
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elongate (32); sternal setae
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<emphasis id="7F88EACDFFDAB82A8313CC06F512FB52" box="[1357,1390,1127,1153]" italics="true" pageId="20" pageNumber="21">st1</emphasis>
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and lyrifissures
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<emphasis id="7F88EACDFFDAB82A8744CCEAF141FB76" box="[282,317,1163,1189]" italics="true" pageId="20" pageNumber="21">
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iv
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<quantity id="8A049B3AFFDAB82A876FCCEAF141FB76" box="[305,317,1163,1189]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="21" unit="in" value="1.0">1</quantity>
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</emphasis>
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in female and males on separate shields or inserted in soft cuticle (36, 49); postero-marginal shields in female small, not extending lateral to opisthogastral suckers (80); femora I seta
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<emphasis id="7F88EACDFFDAB82A8214CCCEF418FB1B" box="[1098,1124,1199,1224]" italics="true" pageId="20" pageNumber="21">av</emphasis>
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in male large curved spine (104). This grouping is poorly supported in jackknife analyses with an arrangement of
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<taxonomicName id="8AFC4D5CFFDAB82A8238CCB2F486FB3E" authorityName="Klompen & Gerdeman" authorityYear="2023" box="[1126,1274,1235,1261]" class="Arachnida" family="Heterozerconidae" genus="Amyzozercon" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFDAB82A8238CCB2F486FB3E" box="[1126,1274,1235,1261]" italics="true" pageId="20" pageNumber="21">Amyzozercon</emphasis>
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</taxonomicName>
|
||
as sistergroup to the remaining
|
||
<taxonomicName id="8AFC4D5CFFDAB82A8708CC96F267FAC2" authorityName="Berlese" authorityYear="1892" box="[342,539,1271,1297]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
|
||
as the most common alternative option.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |