233 lines
32 KiB
XML
233 lines
32 KiB
XML
<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="B63074948C6A5F269BAA673F0A21EAEF" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole beanka Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="4465D6B7-A7E3-4C86-A4DA-45D197B2731F" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
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<mods:mods id="7131C3438CC359EE6F60B9CC6E58FCF2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="18F0C23BC511615996241D050A3BC750">
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<mods:title id="4B0987CC72BAC5BB6E822124D141D4D5">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="568BCA71BCBE70AEBF246DF6DE188029" type="personal">
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<mods:role id="83B2FB0347F597AF5345128ACD874775">
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<mods:roleTerm id="AD18907669B3B33AD40E295419601E4F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A449CD25B1A331101D53FF3488267702">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="ABFCB46880BDC4B0817DED39B9EAD461">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="9F564B6A924ED74051C77D2BBDF7F788" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier id="1313EC603366BE65C91BA13BA840071F" type="email">sdsalata@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="285826FA0831654DA3B460BA6003AAB1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="44850647FCEC02D8CB497E727AA38257">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="0103405B9EA805659F51DC24D33EE460">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="880992FC15F9DB906949A50497DEC500" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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<mods:typeOfResource id="83BFF4623EBFEA1E4EDF48E67979EDE1">text</mods:typeOfResource>
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<mods:titleInfo id="113E66E3A1977E57AD3735F724434E61">
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<mods:title id="C1F57954DB4845A15E5E99E7437EC00A">ZooKeys</mods:title>
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<mods:part id="8A1F03AD161CEF19CA73BC536279591A">
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<mods:date id="6F6DCC9CB6700A026136B8743AF3B237">2020</mods:date>
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<mods:detail id="7F08B73D528E9A7F0FF9219ABFFC26CB" type="volume">
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<mods:number id="6C7AF7C369D9F1FF20985C1271DBD562">949</mods:number>
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<mods:start id="50310E5159868370C8F9CE1DEFCA0A8C">1</mods:start>
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<mods:end id="F3514071E1F23FF3F7E7482D82B635B7">185</mods:end>
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<mods:location id="33A7C26B53E5F74717D81BDE50B0F306">
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<mods:url id="AF2C7D2287A604C31373C34417D7B3CF">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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<mods:classification id="79FAEE76ACB0261610176AFBCD4DE367">journal article</mods:classification>
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<mods:identifier id="AB45EC3F1DC6D70CE16DD7F41E40BCC5" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier id="DC5E133F0BFBE6BE9F9886A78718C6E9" type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier id="4F3CC2FA33CAFD63B32CC1C11A6BA14E" type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<treatment id="B63074948C6A5F269BAA673F0A21EAEF" ID-GBIF-Taxon="165090171" LSID="urn:lsid:zoobank.org:act:4465D6B7-A7E3-4C86-A4DA-45D197B2731F" httpUri="http://treatment.plazi.org/id/B63074948C6A5F269BAA673F0A21EAEF" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="4AE7952758B3027868D6B03F9980F450" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="386516F0EF505DD07D5581EA5BB5CDFA" pageId="0" pageNumber="1">
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<taxonomicName id="DC606D2B166D6648EAB67B24BFE28D68" ID-CoL="CMKWH" LSID="B6307494-8C6A-5F26-9BAA-673F0A21EAEF" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole beanka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">Pheidole beanka</taxonomicName>
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<taxonomicNameLabel id="902447E8DDB469C7AFFD2F12B67D174D" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="F76BA99FEB3D7994452A9F9E86C91ACD" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">Figs 26A-F</figureCitation>
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<figureCitation id="AEE6FFE99A511B1A2EF98341EFF36F76" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63H</figureCitation>
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<figureCitation id="9E59761D32F8E6C0AE3B20C1F25D60D3" captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F44113C6B49948F5815742E012A6F23A" pageId="0" pageNumber="1" type="type material">
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<paragraph id="A557F1F904696C3BD1E45EFFFDF54046" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="4C492DD48408EE22F2A7A97C8AB41303" pageId="0" pageNumber="1">
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<emphasis id="92933EB35BEBB864A0D64E124876227F" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Mahajanga;
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<normalizedToken id="284D1460AB94E361D34232F71A3E48DB" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="64D1E70DD0562AD4F6F864F63C5018AA" originalValue="forestière">forestiere</normalizedToken>
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Beanka, 53.6 km E Maintirano;
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<geoCoordinate id="A52DEDC9729951039E817C29AB944D2A" degrees="18.04014" direction="south" orientation="latitude" precision="1" value="-18.04014">-18.04014</geoCoordinate>
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,
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<geoCoordinate id="A4E731C15EA3F1FCCEAA9F043B5DA044" degrees="44.53394" direction="east" orientation="longitude" precision="1" value="44.53394">44.53394</geoCoordinate>
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; alt. 272 m; 25 Oct 2009; B. L. Fisher et al. leg.; tropical dry forest on tsingy, ex rotten log; BLF22970; CASENT0156684 (CASC).
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<emphasis id="A9042B051045A8C687B6E018A39CC0FF" bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype, CASENT0923276 (CASC).
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</paragraph>
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<caption id="71826C9193C665A956FF686AB8853095" doi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1" start="Figure 26" startId="F26">
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<paragraph id="7155DD9263BF1453B120D505F27B747E" pageId="0" pageNumber="1">
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<emphasis id="BEFE16EC624D1F5046DF0B8C97C6D717" bold="true" pageId="0" pageNumber="1">Figure 26.</emphasis>
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<taxonomicName id="0DCDDB8656E962E1CB38395896855323" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole beanka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="beanka">
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<emphasis id="B4BE44ACFA82A2DAF4CAB65E9FB2C54C" italics="true" pageId="0" pageNumber="1">Pheidole beanka</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis id="B5290A51EDFD8EC22B278746F3D1B2CC" bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis id="6B27ED9CBB8907B77D1A28BB5E120190" bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis id="60FF8EE7A3C197BFBD273D97B996B4A6" bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923276) and full-face view (
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<emphasis id="FAA02BC7E70EC8D51593DC4C8843A18D" bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis id="31CC71AA3F4B5FEE6D920D95E1339AC4" bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis id="4CEE98A276E2CBB2677D589FE4E118B8" bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0156684).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="2161BE8296BEF80C26ED05074FC9AD10" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="08AB209CF350FD74BC66FCA866E1FBE3" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="5E6F83C7594A562A666BBC67B5A71DAE" pageId="0" pageNumber="1">
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Minute species.
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<emphasis id="930BB37D5ADA9EC13223B08B99AA1C00" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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HL <0.9 mm and WL <0.7 mm; head in full-face view elongate, not widening posteriorly with posterior sides slightly convex, in lateral view sub-oval and short with convex dorsal and ventral sides; body ocherous; sides of head with moderately dense, moderately short, suberect to erect pilosity; entire head distinctly sculptured, medial part of frons with thick and longitudinal rugae and distinctly rugulate and never smooth interspaces, occipital lobes with dense and irregular rugae with distinctly rugoreticulate interspaces; scape, when laid back, exceeding the midlength of head by one-fifth of its length; lateral sides of promesonotum distinctly foveolate without additional sparse and indistinct rugulae; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, approximately the same height and width as inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity; base of first gastral tergite shagreened.
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<emphasis id="DB411E17A08E60979AA7F96FEBB99ADA" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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HL <0.45 mm and WL <0.5 mm, scape, when laid back, surpassing the posterior head margin by one-fifth of its length; propodeal spines reduced to small tubercles; head relatively rectangular; body yellowish brown; lateral sides of frons smooth; vertex with transverse, sparse, and moderately thick, short rugulae; mesosoma smooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="100CED459B109C8D443ED7D63250DE83" pageId="0" pageNumber="1" type="description">
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<paragraph id="160B38E28B2F153AD2B4D405E24E00B6" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="106957441D796D958B313D97AF277876" pageId="0" pageNumber="1">
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<emphasis id="77EDB2E7E6ED77A81DFCD1029F18C33B" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="A864685F32EBAE2555CC286221185565" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 0.86; HW: 0.74; SL: 0.45; EL: 0.1; WL: 0.7; PSL: 0.09; MTL: 0.38; PNW: 0.39; PTW: 0.12; PPW: 0.3; CI: 115.3; SI: 60.3; PSLI: 10.5; PPI: 40.4; PNI: 51.9; MTI: 51.1.
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</paragraph>
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<paragraph id="394DE2023B23F6C5FFB7230090BCBFBC" pageId="0" pageNumber="1">
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<emphasis id="EB11EFEBB5F420E0DBC1ADDDA8DFCAC9" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view elongate, not widening posteriorly, with anterior and posterior sides slightly convex (Fig.
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<figureCitation id="3AAAEA99BCAF48A859FD86558AD15717" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately dense, moderately short, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, longitudinal, and dense rugae and distinctly rugulate interspaces, rugae more irregular and with denser sculpture of interspaces on posteromedial part; lateral sides with thick, dense and longitudinal rugae with distinctly rugofoveolate interspaces, rugae more irregular on posterolateral parts. Occipital lobes with dense, irregular rugae and distinctly rugoreticulate interspaces. Area posterolateral from eyes with dense rugofoveolae. Gena with relatively sparse and thick, longitudinal rugae and indistinctly rugoreticulate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (Fig.
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<figureCitation id="0F32769C11D5118BE236AEE8D7D69FA9" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26B, D</figureCitation>
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). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, approximately the same height and width as inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity (Fig.
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<figureCitation id="CFCE672392A9B52D93D3E2FE169E29B5" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63H</figureCitation>
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).
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<emphasis id="D189569556B4E1006C0261BF1337B0B7" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum moderately steep, mesonotal process very indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines very small, with wide base and acute apex; humeral area produced (Fig.
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<figureCitation id="5206F7CFA863D1FA55BE4C63AF0E858E" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26D</figureCitation>
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). Surface shiny and distinctly foveolate; katepisternum with sparser foveolae. Pilosity sparse, moderately long, and erect (Fig.
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<figureCitation id="4A9F56F5A9943D9FD48908F5A3DB61BD" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26D, F</figureCitation>
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).
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<emphasis id="18007FDDD3D050A194DF9345B6410271" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with dense foveolae; node finely foveolate, triangular, with rounded and thin apex, in rear view node dorsoventrally concave; pilosity moderately sparse and erect (Fig.
|
||
<figureCitation id="AEBAB224FED6329524D3700828FDC3B8" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26D, F</figureCitation>
|
||
).
|
||
<emphasis id="1DAF08F15F9D9CF2D18BE32E527B73C8" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
||
Shiny and foveolate; dorsum with reduced sculpture; in dorsal view oval, lateral margins medially with two distinct dentate projections; pilosity long, moderately sparse, and erect (Fig.
|
||
<figureCitation id="30C5272A5B69E5AC0431B254DE66CE15" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26D, F</figureCitation>
|
||
).
|
||
<emphasis id="6E6135E916761CE0888F125420D95F3C" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
Shiny and smooth with shagreened base; pilosity moderately dense, moderately short, and erect (Fig.
|
||
<figureCitation id="B990637807D3E2A718E6CABE070A85D5" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26D, F</figureCitation>
|
||
).
|
||
<emphasis id="A0D36F793C109171760F9322782EF4A7" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Ochreous with yellow legs and antenna (Fig.
|
||
<figureCitation id="7B9F95EC079CCF202FFFD3CC86BD694B" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="61025F38F77C7EA12F219CF4E3275A2D" pageId="0" pageNumber="1">
|
||
<emphasis id="C31A807EAE4343D773CA2C84AA43FEF2" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="1FA8F2E6A977D00FE82BC348D9D53CC4" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 1): HL: 0.44; HW: 0.39; SL: 0.39; EL: 0.1; WL: 0.48; PSL: 0.05; MTL: 0.3; PNW: 0.25; PTW: 0.06; PPW: 0.11; CI: 112.8; SI: 98.7; PSLI: 10.9; PPI: 48.7; PNI: 65.0; MTI: 77.0.
|
||
</paragraph>
|
||
<paragraph id="9C83511D4A0D3BE5735192627C89C8B3" pageId="0" pageNumber="1">
|
||
<emphasis id="C69CC02DA69353A08C8C9EDFCA58E5E0" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
Cephalic margin slightly concave (Fig.
|
||
<figureCitation id="63C040B3887FD53299E9A04323642EAE" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26A</figureCitation>
|
||
). Pilosity relatively sparse, moderately long, decumbent to subdecumbent. Sculpture shiny and smooth; vertex with transverse, sparse, and moderately thick, short rugulae; antennal sockets with few thick, curved outward rugae and smooth interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation id="9E318B11E6B42CB4577461138C703494" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26A, C</figureCitation>
|
||
).
|
||
<emphasis id="83C62BC5826D8289905E912EAB40660B" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum moderately high and short, box-like; promesonotal groove absent; metanotal groove distinct; propodeal spines reduced to small tubercles (Fig.
|
||
<figureCitation id="E1BE64D09BA549571A4F83803DA0C3D2" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26C</figureCitation>
|
||
). Sculpture shiny and smooth. Pilosity sparse, moderately long, and erect (Fig.
|
||
<figureCitation id="6A0ECF051793C4B8ECEC46FE6DE2D7AF" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26C, E</figureCitation>
|
||
).
|
||
<emphasis id="4437B0A70A6F0A3D1823A03EE9FC8C5D" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation id="39128EA14833C5B850EA03B19B6E593D" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26C, E</figureCitation>
|
||
).
|
||
<emphasis id="35354C5DD3FC62B0B912051377ECC0EE" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Yellow (Fig.
|
||
<figureCitation id="6ED659F0F8EE4DBE2E3FE49D4E97F5FF" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pheidole beanka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923276) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0156684)." figureDoi="10.3897/zookeys.949.51269.figure26" httpUri="https://binary.pensoft.net/fig/432195" pageId="0" pageNumber="1">26C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3B5BDC6DF79E1907FEAEFC4A3DFBB100" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="CE3E3E3A492A2319DDEF1D4FD0E11E24" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="A4578B1C36B934FE910B424EAF789454" pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A3182185803596B4EC237DF3D2144311" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="74EB46B028D9535D0C949482492A92B8" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="F3FA281203D1BF58DB24FEF25F15A3DD" pageId="0" pageNumber="1">The species was collected at 272 m in elevation, in tropical dry forest. Nest was located in rotten log.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="770DBFC16F778077E4084CB7995EC696" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="CA9F6AA22DFE73B6ACE586BB637551AE" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="1E4C407C92A10F25D33A4B43B90B7919" pageId="0" pageNumber="1">
|
||
<taxonomicName id="B61BAC8BE1F0A76A2DEC1AA232463B4D" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole beanka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="beanka">
|
||
<emphasis id="353A67524AAF49E667CBD7BDFA88A227" italics="true" pageId="0" pageNumber="1">Pheidole beanka</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. belongs to the group of species characterised by small body size (major workers: HL <1.05 mm, WL <0.9 mm and minor workers HL <0.5 mm, WL <0.6 mm), elongate and not widening posteriorly head in major workers, and minor workers with predominantly smooth and relatively rectangular head and yellow to brown body colouration. The group includes five species:
|
||
<taxonomicName id="F1DCC3FFD122E8E000D51EF819E582A3" lsidName="P. flavominuta" pageId="0" pageNumber="1" rank="species" species="flavominuta">
|
||
<emphasis id="D98C99F02A1C4775E5578A2B5F9D465B" italics="true" pageId="0" pageNumber="1">P. flavominuta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="D60783FCC006F7577AF9602806149291" lsidName="P. nitidobruna" pageId="0" pageNumber="1" rank="species" species="nitidobruna">
|
||
<emphasis id="53045A09B0A3A520EB7748A762C689E8" italics="true" pageId="0" pageNumber="1">P. nitidobruna</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="7AAFBF8CBF62C73365CF45A9DBC8D1EB" lsidName="P. mikros" pageId="0" pageNumber="1" rank="species" species="mikros">
|
||
<emphasis id="B8F7CE79DC8B0B906CFA55CDE53F4E1B" italics="true" pageId="0" pageNumber="1">P. mikros</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., and
|
||
<taxonomicName id="5E45DDC20C9ADA238E2EAA6C29007066" lsidName="P. beanka" pageId="0" pageNumber="1" rank="species" species="beanka">
|
||
<emphasis id="CBC09078BA720F2BF5AC699E0AB724EC" italics="true" pageId="0" pageNumber="1">P. beanka</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<taxonomicName id="889C8D958187878ED570B45C7FBA8BAA" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole beanka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="beanka">
|
||
<emphasis id="0183663134D6BD39813E3BBE300646C6" italics="true" pageId="0" pageNumber="1">Pheidole beanka</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., described from
|
||
<normalizedToken id="D6782FC315E2BE3D004BE66BD5C8CC1A" originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken id="33F1B3B8A64FC40E7CA3927F28D8DDE8" originalValue="forestière">forestiere</normalizedToken>
|
||
Beanka, is the only member of this group known from the Mahajanga and its distribution
|
||
<normalizedToken id="063845FE711C90D4E143A7CA4E80DB95" originalValue="doesn’t">doesn't</normalizedToken>
|
||
overlap with the remaining four taxa. However, morphology of
|
||
<taxonomicName id="56E66EB7553FC1FA7B93F0D631E2A153" lsidName="P. beanka" pageId="0" pageNumber="1" rank="species" species="beanka">
|
||
<emphasis id="1BED525D14B006106B23B2AB22E03550" italics="true" pageId="0" pageNumber="1">P. beanka</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName id="70BE2F71ADF2D979225C3622EE44CE00" lsidName="P. mikros" pageId="0" pageNumber="1" rank="species" species="mikros">
|
||
<emphasis id="C6F878DDB2FD8440F77637F91A868B3B" italics="true" pageId="0" pageNumber="1">P. mikros</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., which is distributed on the southernmost part of the Antsiranana prefecture. Major workers of
|
||
<taxonomicName id="66219F7F83B5BB110979C9EDBE82595C" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole beanka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="beanka">
|
||
<emphasis id="C0AB5854DB5820BAF4D9B18BF1767907" italics="true" pageId="0" pageNumber="1">Pheidole beanka</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differ from
|
||
<taxonomicName id="767837E354572D2803A24CC6E6010F29" lsidName="P. mikros" pageId="0" pageNumber="1" rank="species" species="mikros">
|
||
<emphasis id="09DCD8C35F8713FBF7599FCD20216ED2" italics="true" pageId="0" pageNumber="1">P. mikros</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in the combination of the following characters: medial part of frons with distinctly rugulate interspaces and very small propodeal spines; minor workers can be separated based on smooth lateral sides of frons and mesosoma, yellow body, and propodeal spines reduced to small tubercles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |