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<document id="1D2A61FBD74125D7C32167D46B9A1D75" ID-DOI="http://dx.doi.org/10.3897/mycokeys.50.32432" ID-GBIF-Dataset="059c89da-7ae5-4dbb-b876-bd1617110873" ID-PMC="PMC6477855" ID-Pensoft-Pub="1314-4049-50-1" ID-PubMed="31043855" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-50-1" ModsDocOrigin="MycoKeys 50" ModsDocTitle="Solving the taxonomic identity of Pseudotomentellatristis s.l. (Thelephorales, Basidiomycota) a multi-gene phylogeny and taxonomic review, integrating ecological and geographical data" checkinTime="1555333405629" checkinUser="pensoft" docAuthor="Svantesson, Sten, Larsson, Karl-Henrik, Koljalg, Urmas, W. May, Tom, Patrik Cangren,, Henrik Nilsson, R. &amp; Larsson, Ellen" docDate="2019" docId="B5ACA83EE7A150A0757CE4A6213CA371" docLanguage="en" docName="MycoKeys 50: 1-77" docOrigin="MycoKeys 50" docSource="http://dx.doi.org/10.3897/mycokeys.50.32432" docTitle="Pseudotomentella sciastra Svantesson &amp; Koljalg, sp. nov." docType="treatment" docVersion="5" lastPageNumber="38" masterDocId="BF413468CD59FFA53A5DFFC1FF9F272E" masterDocTitle="Solving the taxonomic identity of Pseudotomentellatristis s. l. (Thelephorales, Basidiomycota) - a multi-gene phylogeny and taxonomic review, integrating ecological and geographical data" masterLastPageNumber="77" masterPageNumber="1" pageNumber="36" updateTime="1732478795134" updateUser="ExternalLinkService">
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<mods:title id="878B93D3A78CA04D107CBD8773266DEB">Solving the taxonomic identity of Pseudotomentellatristis s. l. (Thelephorales, Basidiomycota) - a multi-gene phylogeny and taxonomic review, integrating ecological and geographical data</mods:title>
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<mods:namePart id="AA0095F8BE32F4B2413672B7BC771F3B">Svantesson, Sten</mods:namePart>
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<mods:namePart id="83DB273A44600F45B02FC51CAEDC847E">Larsson, Karl-Henrik</mods:namePart>
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<mods:namePart id="493C6F3E59CB5798C0EF699813D461BA">Koljalg, Urmas</mods:namePart>
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<mods:namePart id="13CF5CD1FF245088405B65BA7382826E">Patrik Cangren,</mods:namePart>
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<mods:namePart id="5167764BAA6F189FAFB5445419A71DB7">Henrik Nilsson, R.</mods:namePart>
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<treatment id="B5ACA83EE7A150A0757CE4A6213CA371" ID-GBIF-Taxon="156201981" LSID="urn:lsid:plazi:treatment:B5ACA83EE7A150A0757CE4A6213CA371" httpUri="http://treatment.plazi.org/id/B5ACA83EE7A150A0757CE4A6213CA371" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" scope_order="Thelephorales" scope_phylum="Basidiomycota">
<subSubSection id="8D1F29B02A75C6AA161CFDFE63459425" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="E9D69682D07FF0F408BE813AADBB17CE" pageId="35" pageNumber="36">
<taxonomicName id="F53966D78EB75F87F3B5D9F14B953719" ID-CoL="4PDRW" LSID="MB829025" authority="Svantesson &amp; Koljalg" class="Agaricomycetes" family="Thelephoraceae" genus="Pseudotomentella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Pseudotomentella sciastra" order="Thelephorales" pageId="35" pageNumber="36" phylum="Basidiomycota" rank="species" species="sciastra">
Pseudotomentella sciastra Svantesson &amp;
<normalizedToken id="A2F8EF65F00F09096FEA53ABCA40CB67" originalValue="Kõljalg">Koljalg</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="56F7978EA7914FCA06195E8DA3AD9EDA" pageId="35" pageNumber="36">sp. nov.</taxonomicNameLabel>
Fig. 15
</paragraph>
</subSubSection>
<subSubSection id="7BF18B2034F83974573E938E16BF53FD" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="C575E6D4702AE2751529C3BDEAE60331" pageId="35" pageNumber="36">Type.</paragraph>
<paragraph id="AFCC7E3A75B6CAEE98350DFDEDF9B3F1" pageId="35" pageNumber="36">
SWEDEN.
<normalizedToken id="A854DA6FF58AD77FBDA5656E4063E2CD" originalValue="Småland">Smaland</normalizedToken>
:
<normalizedToken id="770008855D3D8682417E2D1D30A7045A" originalValue="Jönköping">Joenkoeping</normalizedToken>
, Svarttorp, Ramlaklint, boreonemoral, mixed, old-growth forest, on soil with intermediate pH, 12 September 2016, S. Svantesson 359 (holotype: GB!, GenBank Acc. No. ITS: MK290686).
</paragraph>
</subSubSection>
<subSubSection id="00A758B27496BD0E6854E948A3C638D1" pageId="35" pageNumber="36" type="unite sh">
<paragraph id="F59DAD2834BD68236F75DF60E5283F24" pageId="35" pageNumber="36">UNITE SH.</paragraph>
<paragraph id="4DC080FE95F51E1BD5829D3BADF0DBA0" pageId="35" pageNumber="36">SH030554.07FU</paragraph>
</subSubSection>
<subSubSection id="1B401F2FC412388012EC2E1E76A1B8E9" pageId="35" pageNumber="36" type="etymology">
<paragraph id="8A27DCA91D77E9DF06D3766DB7034D17" pageId="35" pageNumber="36">Etymology.</paragraph>
<paragraph id="0925DED508F280A71DBC43316E83BB33" pageId="35" pageNumber="36">The name refers to the dark, star-like appearance of the spores.</paragraph>
</subSubSection>
<subSubSection id="8171D95537ED6082D5DC834DCF7245A3" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="8AC3F7F411EE37142DF97F6E6A0D7688" pageId="35" pageNumber="36">Description.</paragraph>
<paragraph id="C85969D4466E313ECBDE25A774606316" pageId="35" pageNumber="36">Basidiomata annual, resupinate, membranaceous, effused - often to several tens of centimetres in diameter. Mature parts continuous, with a cottony texture when fresh and a rather firm, fibrous and compact, yet quite soft and elastic texture when dried. Hymenium smooth, but sometimes strongly undulating; blue grey when fresh and brown with a pinkish hue when dried. Immature parts discontinuous, byssoid with a cottony texture, both when fresh and when dried. Subhymenium and hymenium of immature parts blue grey when fresh or occasionally green or even yellow; blue grey to brown grey when dried. Subiculum well developed, loose, fibrous, pale orange brown to brown; often forms the outer edge of basidiomata, extending noticeably beyond the hymenium.</paragraph>
<caption id="DC1742BEC20435C7E761BDCC0BFAAEE0" pageId="35" pageNumber="36">
<paragraph id="9CB9A7BA83E66A8647029B728E763410" pageId="35" pageNumber="36">
Figure 15. Morphological features of
<taxonomicName id="7C197BCA10B888F2DB17A2B265DA759A" lsidName="P. sciastra" pageId="35" pageNumber="36" rank="species" species="sciastra">P. sciastra</taxonomicName>
, mounted in KOH and macroscopically. A basidiospore in frontal face (O F110317) B in frontal and lateral faces (O F110317) C in lateral face (O F110317) D subicular hyphae (O F110317) E (SS420) F (SS312) mature basidiomata.
</paragraph>
</caption>
<paragraph id="95D277FEED52D8A7D03435BD8A8F8E8A" pageId="35" pageNumber="36">Hyphal cords lacking, but loose bundles of subicular hyphae sometimes present.</paragraph>
<paragraph id="C4E3CC2C624EFB530A4537989D00FD08" pageId="35" pageNumber="36">Hyphal system monomitic, clamp connections absent from all hyphae.</paragraph>
<paragraph id="A9B0631748CF8E0418236814B33C2601" pageId="35" pageNumber="36">
Subicular hyphae noticeably long and straight, thick-walled; forming a loose tissue. Individual hyphae (3.9-) 4.4-6.6 (-6.8)
<normalizedToken id="E3080A4947153C02C722F2D3FAC5E3D0" originalValue="μm">μm</normalizedToken>
wide, with a mean width of 5.0-5.8
<normalizedToken id="8C9C8F50BC2DD32D55030320DE52A6CF" originalValue="μm">μm</normalizedToken>
; brown to orange brown in KOH, orange brown in water.
</paragraph>
<paragraph id="E6DEE0FE21C687D2B681DF2761BC4297" pageId="35" pageNumber="36">
Subhymenial hyphae often somewhat sinuous, thin to thick-walled; forming a rather dense tissue. Individual hyphae 2.9-5.0 (-6.0)
<normalizedToken id="EB80B3C93EBCDFD750C20E49C621259E" originalValue="μm">μm</normalizedToken>
wide, with a mean width of 3.8-4.0
<normalizedToken id="2C984CE2A87E93A9C18085799D849819" originalValue="μm">μm</normalizedToken>
; hyaline to pale green in KOH, blue green in the presence of air; pale orange green to pale yellowish-green in water, with strongly granular contents.
</paragraph>
<paragraph id="0C464B20D08898492668561C8F1A0EBF" pageId="35" pageNumber="36">Encrustation granular, probably amyloid (hard to observe due to the colour); blackish-brown in KOH, dark blue green in the presence of air; blackish-brown in water; scattered in occurrence on the upper parts of subhymenial hyphae and on the lower parts of basidia.</paragraph>
<paragraph id="D1E02A1A8C3FABA7910635F2F30EBF13" pageId="36" pageNumber="37">
<pageBreakToken id="17B9497E53B42B823F17356F8E444D98" pageId="36" pageNumber="37" start="start">Basidia</pageBreakToken>
with four slightly curved sterigmata, occasionally two-sterigmate; clavate to narrowly clavate, sometimes clavopedunculate, thin-walled, with one-three slight constrictions. Dimensions: 42-67 (-68)
<normalizedToken id="089C95625D92C7B31674EE0FE57554C7" originalValue="×">x</normalizedToken>
7.3-9.0 (-9.3)
<normalizedToken id="0EF57457CDEAA92E3D1BC621C73FFE88" originalValue="μm">μm</normalizedToken>
; mean dimensions: 54-55
<normalizedToken id="943AEE74CD55B5F234E5198858A14CC3" originalValue="×">x</normalizedToken>
7.8-8.1
<normalizedToken id="E57C8E2DEDACDCE8BAA70F1EF19279C7" originalValue="μm">μm</normalizedToken>
. Sterigmata (6.0-) 6.3-8.9 (-9.1)
<normalizedToken id="8C8BC9DB67CF9032DCC3D69967CB3738" originalValue="μm">μm</normalizedToken>
long, with a mean length of 7.4-7.9
<normalizedToken id="7FCBD0816F1C6D836AE08332C8115705" originalValue="μm">μm</normalizedToken>
. Colours and reactions the same as for the subhymenial hyphae, but in addition often with granular contents in KOH.
</paragraph>
<paragraph id="1800C9A0206600E7B0D3DBDFF7F9C277" pageId="36" pageNumber="37">Cystidial organs lacking.</paragraph>
<paragraph id="782B788F55FE9CF2E3867D909E7A5DA9" pageId="36" pageNumber="37">
Basidiospores in frontal face with a subcircular basic shape and a star- or cross-shaped, sometimes angular to nodulose outline, covered in bi- or trifurcate, sometimes singularly attached, echinuli. Nearly all spores with four-six distinct, rounded to more often square lobes or rarely corners; abnormally large spores originating from two-sterigmate basidia infrequently occurring. Frontal dimensions: (6.0-) 6.1-7.9 (-8.1)
<normalizedToken id="6C66450516E51C353489006C0C922A55" originalValue="×">x</normalizedToken>
6.3-8.2
<normalizedToken id="A56EE0A1EEB2E9C48538EDCE0A322857" originalValue="μm">μm</normalizedToken>
; mean dimensions: 6.6-7.3
<normalizedToken id="7B2ED591911DCC4BC544D175AAE68134" originalValue="×">x</normalizedToken>
6.7-7.7
<normalizedToken id="A061A82945295F70FF65F90E3FC064FC" originalValue="μm">μm</normalizedToken>
; Q-value: 0.9-1.1; mean Q-value: 1.0. Echinuli (0.5-) 0.6-1.2 (-1.4)
<normalizedToken id="68834F1476B5A6778F8F7168A4A06C36" originalValue="μm">μm</normalizedToken>
long, with a mean length of 0.8-0.9
<normalizedToken id="A00586BD942518B99447BD92027C41ED" originalValue="μm">μm</normalizedToken>
. Lateral face ellipsoid to ovoid, with evenly rounded edges or one-three lobes. Lateral dimensions: (6.2-) 6.5-7.7 (-8.0)
<normalizedToken id="C0D3D770DE8895DF25483DAD04846837" originalValue="×">x</normalizedToken>
(4.3-) 4.4-6.0 (-6.2)
<normalizedToken id="EC88EF6942AF83901A1E9E029488AC94" originalValue="μm">μm</normalizedToken>
; mean dimensions: 6.8-7.3
<normalizedToken id="70BF68788AAE6DAA20FDB233F4AA4614" originalValue="×">x</normalizedToken>
4.6-5.4
<normalizedToken id="BD2001A03C0A9A0D0E00ECC575D1A0E0" originalValue="μm">μm</normalizedToken>
; Q-value: 1.2-1.6 (-1.7); mean Q-value: 1.3-1.5. Colour in KOH brown to yellow brown, in the presence of air often with a green to blue green reaction; in water pale greenish to pale greenish-orange; occasionally amyloid.
</paragraph>
<paragraph id="262BA74F2B0D8CCF51A686FD576FC69B" pageId="36" pageNumber="37">Chlamydospores lacking.</paragraph>
</subSubSection>
<subSubSection id="2EB6F03708245E3B7A8B2BAFDCFCA6C1" pageId="36" pageNumber="37" type="habitat">
<paragraph id="F9A4A3921A42D971FEB7CDD014B25420" pageId="36" pageNumber="37">Habitat.</paragraph>
<paragraph id="9964CF69958DD58B9DB0B3753D6070CC" pageId="36" pageNumber="37">
Recent Scandinavian collections have been made in mature to old coniferous, deciduous or mixed forests on soil with intermediate to high pH.
<taxonomicName id="1A7C1788656A2C5EA273DE9A60A6A458" class="Agaricomycetes" family="Thelephoraceae" genus="Pseudotomentella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Pseudotomentella sciastra" order="Thelephorales" pageId="36" pageNumber="37" phylum="Basidiomycota" rank="species" species="sciastra">Pseudotomentella sciastra</taxonomicName>
has been found to form ectomycorrhiza with at least
<taxonomicName id="26E2F7BCF9AC61A23E7ACEBCFE022778" class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Castanea sativa" order="Fagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="sativa">Castanea sativa</taxonomicName>
,
<taxonomicName id="D7F1BC1AFAC8A708683C74986AB4B106" class="Pinopsida" family="Pinaceae" genus="Cedrus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cedrus libani" order="Pinales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="libani">Cedrus libani</taxonomicName>
,
<taxonomicName id="8922C1A596CB591BBC0EBE081EC66691" class="Liliopsida" family="Orchidaceae" genus="Neottia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Neottia ovata" order="Asparagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="ovata">Neottia ovata</taxonomicName>
,
<taxonomicName id="634887D1C9F34809998AC5C2C1247827" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea abies" order="Pinales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="abies">Picea abies</taxonomicName>
and
<taxonomicName id="58699A4E25E36E27B488126DF7CF72F7" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
sp. (
<bibRefCitation id="96BB677A790C62E5C8A3C3D854587187" author="Koljalg, U" journalOrPublisher="New Phytologist" pageId="61" pageNumber="62" pagination="1063 - 1068" title="UNITE: a database providing web-based methods for the molecular identification of ectomycorrhizal fungi." url="https://doi.org/10.1111/j.1469-8137.2005.01376.x" volume="166" year="2005">
<normalizedToken id="9BA3B1F9F0AADA3BC79C9FC0D12DD8C4" originalValue="Kõljalg">Koljalg</normalizedToken>
et al. 2005
</bibRefCitation>
,
<bibRefCitation id="14907AE8690A13B6C58A44525AC0D7D1" author="Nilsson, RH" journalOrPublisher="MycoKeys" pageId="62" pageNumber="63" url="https://doi.org/10.1093/nar/gky1022" year="2019">Nilsson et al. 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FF2CD7784B5D9835828C643F0E0C3486" pageId="36" pageNumber="37" type="distribution">
<paragraph id="6C3DC1D0F3EFB01180CE66676589AEF7" pageId="36" pageNumber="37">Distribution.</paragraph>
<paragraph id="1292D274053D170868A3589788838C6F" pageId="36" pageNumber="37">Basidiomata encountered in: Estonia, Finland, Norway, Sweden, Turkey and the United Kingdom. Soil or root tip samples confirm presence also in: the Czech Republic, Mexico, Portugal (Madeira) and the United States.</paragraph>
</subSubSection>
<subSubSection id="D4581343CEA39DB44FB36815FD19B596" pageId="36" pageNumber="37" type="remarks">
<paragraph id="946EB02192F954D118B77BC975E8A8D4" pageId="36" pageNumber="37">Remarks.</paragraph>
<paragraph id="43C66FDCF48A4FF379C7E9179B84FEA0" pageId="36" pageNumber="37">
All studied European specimens previously identified as
<taxonomicName id="53E36A418358C4B0764158766ECFE4C4" lsidName="P. atrofusca" pageId="36" pageNumber="37" rank="species" species="atrofusca">P. atrofusca</taxonomicName>
belong to
<taxonomicName id="C696BBF90DF7B1E770AA50D54BAC168A" lsidName="P. sciastra" pageId="36" pageNumber="37" rank="species" species="sciastra">P. sciastra</taxonomicName>
. The two species display considerable morphological differences (see key).
</paragraph>
<paragraph id="D23F95458BFCB19A6E0AA74E5582E18C" pageId="36" pageNumber="37">
Within the
<taxonomicName id="DF5076DEED72F61BE0EE33BB1A4E134A" lsidName="P. tristis" pageId="36" pageNumber="37" rank="species" species="tristis">P. tristis</taxonomicName>
group, the basidiomata of
<taxonomicName id="3276F7DEB9C7BA71B43AE30B8CBB677D" lsidName="P. sciastra" pageId="36" pageNumber="37" rank="species" species="sciastra">P. sciastra</taxonomicName>
are recognised by their lack of hyphal cords and skeletal hyphae and their soft, yet rather firm and compact and
<normalizedToken id="EE7F30553D2038C6FB2C4AEB96FD5B67" originalValue="±">+/-</normalizedToken>
elastic texture after drying, bluish to greenish colour of immature parts, wide subicular hyphae and small, star-shaped spores.
<taxonomicName id="E4D81A813D7657009D34893F8F36B6AF" class="Agaricomycetes" family="Thelephoraceae" genus="Pseudotomentella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Pseudotomentella pinophila" order="Thelephorales" pageId="36" pageNumber="37" phylum="Basidiomycota" rank="species" species="pinophila">Pseudotomentella pinophila</taxonomicName>
is similar, but has narrower subicular hyphae and larger spores.
</paragraph>
</subSubSection>
<subSubSection id="5AB9B7B6B737C38C3B7A44356DE54034" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="additional specimens studied">
<paragraph id="32D734D765313218EC8A546A40B2A0A7" pageId="36" pageNumber="37">Additional specimens studied.</paragraph>
<paragraph id="EC09A1F271296DF779ACC14574BD4E53" pageId="36" pageNumber="37">
ESTONIA. Ida-Virumaa: Illuka, Puhatu Nature Reserve, Poruni primeval forest, wetlands, 1 October 2006, U.
<normalizedToken id="0ADE3B2B355FCEE3CF30AAAB219CDE14" originalValue="Kõljalg">Koljalg</normalizedToken>
(TU 100644*); [Saare,] Saarema, Kihelkonna,
<normalizedToken id="B95E40C61AA713FD6135A0E5BC5CA20C" originalValue="Hülgera">Huelgera</normalizedToken>
, Sampling area G4422, 25 September 2015, A. Saitta (TU 124211*, TU 124213*);
</paragraph>
<paragraph id="B1CEA58887B306CC874A2490F4CA4863" pageId="36" pageNumber="37">
FINLAND.
<normalizedToken id="AF68B68AD48202E2D0531D930036C6F3" originalValue="Etelä-Häme">Etelae-Haeme</normalizedToken>
:
<normalizedToken id="D954FC98A575333B5840D65D3E093DDF" originalValue="Jyväskylä">Jyvaeskylae</normalizedToken>
, Korpilahti, Oittila, on dead trunk of
<taxonomicName id="8023397D3D8AC8CB50457C0D4424ACE7" class="Magnoliopsida" family="Ulmaceae" genus="Ulmus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ulmus glabra" order="Rosales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="glabra">Ulmus glabra</taxonomicName>
, 3 September 2014, U.
<normalizedToken id="B9A7AEA4B2077F798FEE5DF131A7E35E" originalValue="Söderholm">Soederholm</normalizedToken>
4755 (H 6052710);
<normalizedToken id="4A5F16FF971D3C2FD8C9EFFFDABE656D" originalValue="Kanta-Häme">Kanta-Haeme</normalizedToken>
: Lammi, Lammi Biological Station, Leib-rich forest, 12 September 2001, K.-H. Larsson (TU 108754*);
</paragraph>
<paragraph id="AA962E56D9596B9465F62208F80DE31A" pageId="37" pageNumber="38">
<pageBreakToken id="84675F4C949262BE07D7D50654B67D36" pageId="37" pageNumber="38" start="start">NORWAY</pageBreakToken>
. Oslo (county): Oslo (municipality),
<normalizedToken id="0D51B8D58CA645EE4B7D86F6988EDA6A" originalValue="Bygdøy">Bygdoy</normalizedToken>
,
<normalizedToken id="38A6269ED1A26A83AF5B1070E5772DF0" originalValue="Hengsåsen">Hengsasen</normalizedToken>
, boreonemoral, mixed forest on soil with high pH, 16 August 2010, K.-H. Larsson and N. Svensson (O F110317*);
<normalizedToken id="2C6099C85E76F2B9494AD5504F5AC8B0" originalValue="Østfold">Ostfold</normalizedToken>
: Moss,
<normalizedToken id="9DF700901A9751868F4E6A9B89CB1838" originalValue="Jeløya">Jeloya</normalizedToken>
, boreonemoral, mixed forest on soil with high pH, 26 September 2010, S. Svantesson and N. Svensson (O F110318*); Oppland: Dovre, Grimsdalen, Austre
<normalizedToken id="2710770D128A3DAC0DA7BFF9B721DBB3" originalValue="Stakkstosætra">Stakkstosaetra</normalizedToken>
,
<taxonomicName id="F2C7FC8AA16C402EE5DAA3FD1B6EE808" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus sylvestris" order="Pinales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="sylvestris">Pinus sylvestris</taxonomicName>
forest, 26 August 2010, K.-H. Larsson and S. Svantesson (O F110301); Vestfold: Larvik, Kvelde,
<normalizedToken id="243B60776B35BAD60A7C20F945485A99" originalValue="Jordstøyp">Jordstoyp</normalizedToken>
, boreonemoral, mixed forest on soil with intermediate pH, 1 October 2010, K.-H. Larsson (O F110302); Ibidem, on soil with high pH, 1 September 2010, K.-H. Larsson and S. Svantesson (O F110303, F110304); Aust-Agder,
<normalizedToken id="10D7959A0F87D71DB5253DFCFB7CEA51" originalValue="Risør">Risor</normalizedToken>
, Glupedalen, boreonemoral, mixed forest on soil with high pH, 10 September 2010, S. Svantesson and N. Svensson (O F110319, F110320, F110321); Aust-Agder: Tvedestrand, Eidbo, boreonemoral, mixed forest on soil with intermediate pH, 10 September 2010, S. Svantesson and N. Svensson (O F110322*); Oslo (county): Oslo (municipality), Gressholmen, boreonemoral, mixed forest on soil with high pH, 20 September 2010, S. Svantesson (O F110323, F110324); Oslo (county): Oslo (municipality), Killingen, boreonemoral, mixed forest on soil with high pH, 22 September 2010, S. Svantesson (O F110325); Buskerud: Ringerike, Ulltveit Nature Reserve, boreonemoral, coniferous forest on soil with high pH, 25 September 2010, S. Svantesson and N. Svensson (O F110326);
</paragraph>
<paragraph id="B5535A5AE9DFA1E184B7F16C244032D4" pageId="37" pageNumber="38">
SWEDEN.
<normalizedToken id="CCF55D815575220CEDFD91DC15C967D0" originalValue="Småland">Smaland</normalizedToken>
:
<normalizedToken id="7A61255617442210F3D2B9438B638A24" originalValue="Jönköping">Joenkoeping</normalizedToken>
, Svarttorp, Ramlaklint, boreonemoral, mixed, old-growth forest, on soil with intermediate pH, 12 September 2016, S. Svantesson 360 (GB);
<normalizedToken id="DDB62C74FE2E4D6283E290B8DCD432F3" originalValue="Bohuslän">Bohuslaen</normalizedToken>
: Tanum (municipality), Tanum (parish),
<normalizedToken id="708899AAFB5B4B6039712E5E070021F4" originalValue="Lammön">Lammoen</normalizedToken>
, boreonemoral, deciduous forest on soil with high pH, 6 September 2016, S. Svantesson 312* (GB);
<normalizedToken id="91A0AEB92C29836039EF0C6D77F4B055" originalValue="Västergötland">Vaestergoetland</normalizedToken>
:
<normalizedToken id="80500F01A6CC3DED7AB5F9512292791C" originalValue="Göteborg">Goeteborg</normalizedToken>
, Askim,
<normalizedToken id="0FD74E63EC40DD862D54A2920E0F1374" originalValue="Årekärrslunden">Arekaerrslunden</normalizedToken>
, 24 October 2015, K.-H. Larsson 17308b* (GB); Dalsland, Mellerud,
<normalizedToken id="3FDE3D5AFC91F3C21F7ACF59FE4143F6" originalValue="Skållerud">Skallerud</normalizedToken>
,
<normalizedToken id="2701A73F4A7537F41B679FBE11C449E1" originalValue="Österbo">Oesterbo</normalizedToken>
, mixed forest on soil with high pH, 20 September 2017, S. Svantesson 420* (GB); Ibidem, Norgekullen SW, coniferous forest on soil with high pH, 20 September 2017, S. Svantesson 423* (GB);
</paragraph>
<paragraph id="0F71651B78A0D159E64AAEF7109455F7" pageId="37" pageNumber="38">
TURKEY. [Antalya:
<normalizedToken id="DC7577CE0B3BA43E95FE65B972F72D46" originalValue="Elmalı">Elmali</normalizedToken>
,] Ciglikara, 2009, L. Tedersoo (TU 110153*); [Isparta:]
<normalizedToken id="4DBC7CD5A00D3E16375EECD478C4332B" originalValue="Yukan-Gökdere">Yukan-Goekdere</normalizedToken>
[=
<normalizedToken id="2D6F391A158A4F430B1E4585D441DCA1" originalValue="Yukarı">Yukari</normalizedToken>
<normalizedToken id="681B7D79F6AC144C650B6949A82915CA" originalValue="Gökdere">Goekdere</normalizedToken>
], 2009, L. Tedersoo (TU 110113*);
</paragraph>
<paragraph id="79E7253DCDBE2F55DFD13128D0619EA4" pageId="37" pageNumber="38">
UNITED KINGDOM. Scotland, Aberdeenshire: Inverurie, Burnhervie, in a small group of planted
<taxonomicName id="05E4F8FE11E1A401438EFB0A40571E6C" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
trees, 16 September 2005, I. J. Alexander (TAA 187322*).
</paragraph>
</subSubSection>
</treatment>
</document>