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<document id="D88A2AF0B09A761022597B86AA146C77" ID-CLB-Dataset="299644" ID-DOI="10.3897/zse.100.117859" ID-GBIF-Dataset="f17d3dc6-151e-4b2c-bfe6-1e3978daa701" ID-ZooBank="15CA9ED4-C24C-4AD7-81AC-F699FA953FE8" ID-publisher-id="117859" URI-arpha="35315993-8E70-5007-A8AE-277D7EFC9889" URI-zoobank="http://zoobank.org/15CA9ED4-C24C-4AD7-81AC-F699FA953FE8" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1720542344403" checkinUser="pensoft" docAuthor="Yuan, Zi-Ming, Jiang, Wei &amp; Sha, Zhong-Li" docDate="2024" docId="9682DAD4848258339F5C7FD265643EF7" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 965-987" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/117859/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Gothus consobrinus Yuan &amp; Jiang &amp; Sha 2024, comb. nov." docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="987" masterDocId="353159938E705007A8AE277D7EFC9889" masterDocTitle="Morphological and molecular evidence for Gothus teemo gen. et sp. nov., a new xanthid crab (Crustacea, Brachyura, Xanthoidea) from coral reefs in the South China Sea, with a review of the taxonomy of Actaeodes consobrinus (A. Milne-Edwards, 1867)" masterLastPageNumber="987" masterPageNumber="965" pageNumber="965" updateTime="1720578290841" updateUser="ExternalLinkService">
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<mods:title id="2844573142DAA56C55CF82434CBB4983">Morphological and molecular evidence for Gothus teemo gen. et sp. nov., a new xanthid crab (Crustacea, Brachyura, Xanthoidea) from coral reefs in the South China Sea, with a review of the taxonomy of Actaeodes consobrinus (A. Milne-Edwards, 1867)</mods:title>
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<mods:namePart id="9618BEC95458443B7561B0833A2A4796">Yuan, Zi-Ming</mods:namePart>
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<mods:affiliation id="54DAA90670F94145726A6B24135BFE41">Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China &amp; Laoshan Laboratory, Qingdao 266237, China &amp; Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China</mods:affiliation>
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<mods:namePart id="237376F51C384DFFF3FD9A5C858FFF4D">Jiang, Wei</mods:namePart>
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<mods:affiliation id="22C3152DC1EF0A182BC343494284EFFE">Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China &amp; Laoshan Laboratory, Qingdao 266237, China</mods:affiliation>
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<treatment id="9682DAD4848258339F5C7FD265643EF7" ID-DOI="http://doi.org/10.5281/zenodo.12702518" ID-GBIF-Taxon="235025289" ID-Zenodo-Dep="12702518" ID-arpha="9682DAD4-8482-5833-9F5C-7FD265643EF7" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7" httpUri="http://treatment.plazi.org/id/9682DAD4848258339F5C7FD265643EF7">
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<taxonomicName id="9AC1431F81FD19454E421C4AF4751524" authority="(A. Milne-Edwards, 1873)" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus" status="comb. nov.">
<emphasis id="43B1A2330D373EFA4C9270BB99E5D435" italics="true">Gothus consobrinus</emphasis>
(A.
<bibRefCitation id="AC42E9ADBA0F944F0887163001CBD85D" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" journalOrPublisher="Journal des Museum Godeffroy" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustaces nouveaux ou peu connus provenant du musee de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253 - 264. [pls. 12 - 13] https: // doi. org / 10.5962 / bhl. title. 10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards, 1873</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="12DCBD46AB99BC256784489F8D24E1C7" rank="species">comb. nov.</taxonomicNameLabel>
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<paragraph id="6FDF28381D04FC45AE59F6C3FCC9A2FB">
<figureCitation id="3793C1092B615D8778996394AEB87D1B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">Figs 7</figureCitation>
,
<figureCitation id="67F742B95678CBB9960F4D982671A202" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8</figureCitation>
,
<figureCitation id="5AC3B241A2E8E6949B429CD50346788F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9</figureCitation>
,
<figureCitation id="9582E7F35B3FCAA61E5A269B4727F631" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Gothus consobrinus (A. Milne-Edwards, 1873). A. Male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336; B. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014, displaying live coloration. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure10" httpUri="https://binary.pensoft.net/fig/1088103">10</figureCitation>
</paragraph>
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<treatmentCitation id="A53F4CA746F89083DA59BAE7884CC0D3" author="Milne-Edwards A" page="255" year="1873">
<taxonomicName id="C0C43781058E2F79B5C4474CFCA19A2C" ID-CoL="CCTR4" authority="A. Milne-Edwards, 1873: 255" authorityName="A. Milne-Edwards" authorityPageNumber="255" authorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrina">
<emphasis id="987F4A1B142D39E1A4E1AE4DDFE84FAC" italics="true">Actaea consobrina</emphasis>
<comment id="406B50FB1FFD0B5A9F561A631058CC2E">
A.
<bibRefCitation id="884C10434C43FD38D87678CD23FAE7BA" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" journalOrPublisher="Journal des Museum Godeffroy" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustaces nouveaux ou peu connus provenant du musee de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253 - 264. [pls. 12 - 13] https: // doi. org / 10.5962 / bhl. title. 10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards, 1873: 255</bibRefCitation>
;
<treatmentCitation id="EABA7B84C88D66C783FF92F5C147D637" author="de Man JG" page="503" year="1896">
<bibRefCitation id="B4B06549A7EB15144A875679F2891C06" DOI="10.5962/bhl.title.16084" author="de Man JG" journalOrPublisher="Abteilung fur Systematik, Geographie und Biologie der Tiere" pagination="459-514" refId="B6" refString="de Man JG (1896) Bericht uber die von herrn schiffscapitan storm zu atjeh, an den westlichen kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Theil 4. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere 9: 459 - 514. https: // doi. org / 10.5962 / bhl. title. 16084" title="Bericht über die von herrn schiffscapitän storm zu atjeh, an den westlichen küsten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Theil 4. Zoologische Jahrbucher." volume="9" year="1896">de Man, 1896: 503</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="D5492514ADC58E238F39A2949F0E60C8" author="Odhner T" page="67" year="1925">
<bibRefCitation id="C8AF0D5A23775D46C7B255DB267ACEFA" author="Odhner T" refId="B31" refString="Odhner T (1925) Monographierte gattungen der krabben-familie Xanthidae. I. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar (4) 29 (1): 1 - 92. [figs 1 - 7, pls 1 - 5]" year="1925">Odhner, 1925: 67</bibRefCitation>
</treatmentCitation>
, pl. 4, fig. 14;
<treatmentCitation id="57E93A4F6BFE538516FA1A0AAD4323DB" author="Ward M" page="246" year="1933">
<bibRefCitation id="3E8D1484B84E8694EE4A5D415360E2C7" author="Ward M" issue="5" journalOrPublisher="Australian Zoologist, Sydney" pagination="237-255" refId="B52" refString="Ward M (1933) The true crabs of the Capricorn group, Queensland (Class Crustacea, Order Decapoda Brachyura). Part 1. Xanthidae. Australian Zoologist, Sydney 7 (5): 237 - 255." title="The true crabs of the Capricorn group, Queensland (Class Crustacea, Order Decapoda Brachyura). Part 1. Xanthidae." volume="7" year="1933">Ward, 1933: 246</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="69FA0867392B3E2D8E7AFB2AD1255454" author="Sakai T" page="491" year="1939">
<bibRefCitation id="D64CD079BC2D9DC16C3B2569BDAFC677" author="Sakai T" refId="B40" refString="Sakai T (1939) Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo Co., Tokyo, 365 - 741. [figs 1 - 129, pls 42 - 111, table 1]" year="1939">Sakai, 1939: 491</bibRefCitation>
</treatmentCitation>
, pl. 94, fig. 2;
<treatmentCitation id="9038E6786E86994AA4D2FB44B2D7E459" author="Tweedie MWF" page="118" year="1950">
<bibRefCitation id="14843232B49E9C0F30C0937FD6B3CAB0" author="Tweedie MWF" journalOrPublisher="Bulletin of the Raffles Museum, Singapore" pagination="105-148" refId="B50" refString="Tweedie MWF (1950) The fauna of the Cocos-Keeling Islands. Brachyura and Stomatopoda. Bulletin of the Raffles Museum, Singapore 22: 105 - 148. [figs 1 - 4, pls 16 - 17]" title="The fauna of the Cocos-Keeling Islands. Brachyura and Stomatopoda." volume="22" year="1950">Tweedie, 1950: 118</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="FDFF805001AF38F9160CD73D9D7741B2" author="Serène R &amp; Lang BT" page="291" year="1959">
<bibRefCitation id="8A95D4385FA341C72D9A770AA59E843C" author="Serène R &amp; Lang BT" journalOrPublisher="Annales de la Faculté des Sciences: Saigon" pagination="285-300" refId="B43" refString="Serene R, Lang BT (1959) Observations sur les premiers pleopodes males d'especes d'Actea (Brachyures) du Viet-Nam. Annales de la Faculte des Sciences: Saigon 43: 285 - 300." title="Observations sur les premiers pléopodes mâles despèces dActea (Brachyures) du Viêt-Nam." volume="43" year="1959">Serène &amp; Lang, 1959: 291</bibRefCitation>
</treatmentCitation>
, fig. 2, A 1 - A 3; Guinot, 1967 a: 260
</comment>
</taxonomicName>
</treatmentCitation>
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</paragraph>
<paragraph id="D77FF47AEDFA1F15B4B6752A174D667F">
<treatmentCitationGroup id="7AA63BB639C10B5150CAC0D702823460">
<treatmentCitation id="3695C3328D7415BD39FC548996C79467" author="Rathbun MJ" page="220" year="1911">
<taxonomicName id="DEA6EE531762D53142DE040711125320" ID-CoL="9S5P" authority="Rathbun, 1911: 220" authorityName="Rathbun" authorityPageNumber="220" authorityYear="1911" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="suffuscula">
<emphasis id="B0086886EDD9586E72DCA32F8FAD2E0A" italics="true">Actaea suffuscula</emphasis>
<comment id="35D06EAF20165AFF8C62D796BA057D07">
<bibRefCitation id="64B069F42A2B6CC7DEAB0E549F278B4D" author="Rathbun MJ" refId="B38" refString="Rathbun MJ (1911) Marine Brachyura. In: The Percy Sladen Trust expedition to the Indian Ocean in 1905, under the leadership of Mr J. Stanley Gardiner. Vol. III, part II, No. XI. Transactions of the Linnean Society of London: (Zool.), (2) 14 (2): 191 - 261. [figs 1 - 2, pls 15 - 20]" year="1911">Rathbun, 1911: 220</bibRefCitation>
, pl. 17, figs 1011;
<treatmentCitation id="8E7CCC7F8BAE7095C30B2037E84E17E4" author="Ward M" page="18" year="1934">
<bibRefCitation id="85C62F5D0059D09606B9CD5C6C5DCB81" author="Ward M" journalOrPublisher="Bulletin of the Raffles Museum, Singapore" pagination="5-28" refId="B53" refString="Ward M (1934) Notes on a collection of crabs from Christmas Island, Indian Ocean. Bulletin of the Raffles Museum, Singapore 9: 5 - 28. [pls 1 - 3]" title="Notes on a collection of crabs from Christmas Island, Indian Ocean." volume="9" year="1934">Ward, 1934: 18</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="212BAB4D8AC54FE9C75C967DB2C62C2C" author="Estampador EP" page="81" year="1959">
<bibRefCitation id="0F3B3DE873E4DAB7E754D950B074ECB2" author="Estampador EP" issue="1" journalOrPublisher="Natural and Applied Science Bulletin" pagination="1-127" refId="B7" refString="Estampador EP (1959) Revised check list of Philippine crustacean decapods. Natural and Applied Science Bulletin 17 (1): 1 - 127." title="Revised check list of Philippine crustacean decapods." volume="17" year="1959">Estampador, 1959: 81</bibRefCitation>
</treatmentCitation>
.
</comment>
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</treatmentCitation>
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</paragraph>
<paragraph id="CB15A7C62FB16F8766DD7CB07B0FFE73">
<treatmentCitationGroup id="07DFA488E51943D474F5F9390E1C12A8">
<treatmentCitation id="5867041F23386F4FAFD364ECCB1F2E2A" author="Guinot D" page="246" year="1976">
<taxonomicName id="A22A3A9AD281DAC84855D230321948DC" ID-CoL="9S6L" authority="Guinot, 1967 b: 561" authorityName="Guinot" authorityPageNumber="561" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="9A72885845B082718C37461E4242868B" italics="true">Actaeodes consobrinus</emphasis>
<comment id="6F034D3810BC87050BFD0E9152FA2B12">
Guinot, 1967 b: 561;
<bibRefCitation id="9D07751C44B1F0B6ECF06C2CFFD078BF" author="Guinot D" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustaces Decapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Memoires du Museum National d'Histoire Naturelle, Paris: A, 97, 1 - 308. [figs 1 - 47, pls 1 - 19]" year="1976">Guinot, 1976: 246</bibRefCitation>
, pl. 15, fig. 5;
<treatmentCitation id="F8601D6CF0176F9C61545F96A9012A05" author="Sakai T" page="448" year="1976">
<bibRefCitation id="1225831B81C76B87733CF3779134831D" author="Sakai T" refId="B41" refString="Sakai T (1976) Crabs of Japan and the adjacent seas. Kodansha Ltd, Tokyo, pp. i - xxix, 1 - 773. [figs 1 - 379; pls 1 - 251; 1 - 461, figs 1 - 2, 3 maps]" year="1976">Sakai, 1976: 448</bibRefCitation>
</treatmentCitation>
, pl. 158, fig. 3;
<treatmentCitation id="0BDB96C9563C614744DA39F2DFBEE5DC" author="Takeda M &amp; Miyake S" page="108" year="1976">
<bibRefCitation id="B5E2F8474FDBB88D5F33E9274F8F7951" DOI="10.18353/rcustacea.7.0_101" author="Takeda M &amp; Miyake S" journalOrPublisher="Researches on Crustacea, Tokyo" pagination="101-115" refId="B46" refString="Takeda M, Miyake S (1976) Crabs of the Ogasawara Island. Researches on Crustacea, Tokyo 7: 101 - 115. [fig. 1] https: // doi. org / 10.18353 / rcustacea. 7.0 _ 101" title="Crabs of the Ogasawara Island." volume="7" year="1976">Takeda &amp; Miyake, 1976: 108</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="8A6D426906899F77F33A92283B3D49A3" author="Serène R" page="133" year="1984">
<bibRefCitation id="01E465D870C274A274ACFEA73AB324CB" author="Serène R" refId="B42" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1 - 349. [figs A - C + 1 - 243, pls 1 - 48]" year="1984">Serène, 1984: 133</bibRefCitation>
</treatmentCitation>
(key), 134 (key), 135, pl. 18 C;
<treatmentCitation id="168088EC534826CAADC370DFCD080C8C" author="Galil BS &amp; Vannini M" page="37" year="1990">
<bibRefCitation id="4E281F8996970D62028763F759AC9C5C" DOI="10.1080/03946975.1990.10539447" author="Galil BS &amp; Vannini M" issue="1" journalOrPublisher="Tropical Zoology" pagination="21-56" refId="B9" refString="Galil BS, Vannini M (1990) Research on the coast of Somalia. Xanthidae, Trapeziidae, Carpiliidae, Menippidae (Crustacea, Brachyura). Tropical Zoology 3 (1): 21 - 56. https: // doi. org / 10.1080 / 03946975.1990. 10539447" title="Research on the coast of Somalia. Xanthidae, Trapeziidae, Carpiliidae, Menippidae (Crustacea, Brachyura)." volume="3" year="1990">Galil &amp; Vannini, 1990: 37</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="DE88FD7090ABD891A63157DC4E2BF87F">
<treatmentCitationGroup id="903AE78CABE7F4626E810B95B58627A5">
<treatmentCitation id="ACECCBD5D7838521F4FF4ECE75BAC7CC">
<taxonomicName id="F683520F5C1FA00E6DCC9E6FC58C5FD2" authority="Guinot, 1971 a: 1072" authorityName="Guinot" authorityPageNumber="1072" authorityYear="1971" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrina">
<emphasis id="D4B6C5E3ABE065BC6A639C0FFBD453FD" italics="true">Actaeodes consobrina</emphasis>
Guinot, 1971 a: 1072
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="18001E966B4840915FE81DDD35846982">
<treatmentCitationGroup id="58F01A24EFB60CB524E4CDCA5B57F9A4">
<comment id="B29407AAA47D15E2AFD3CE03EB0B8AEF">
Non
<treatmentCitation id="232875DD8747684BD35A69FE190F338F" author="Nobili MG" page="390" year="1907">
<taxonomicName id="A40ACE87B7CC4E38E977440CFC436B66" ID-CoL="CCTR4" authority="Nobili, 1907: 390" authorityName="Nobili" authorityPageNumber="390" authorityYear="1907" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrina">
<emphasis id="D0B5DDB3914402C26128FB75AB378BDD" italics="true">Actaea consobrina</emphasis>
<bibRefCitation id="23A64F7C18DCE840BBDE0E7AAB99B2D9" DOI="10.5962/bhl.title.53748" author="Nobili MG" issue="2" journalOrPublisher="Memorie della reale accademia delle scienze di torino" pagination="351-430" refId="B30" refString="Nobili MG (1907) Ricerche sui crostacei della Polinesia (Decapodi, Stomatopodi, Anisopodi e Isopodi). Memorie della reale accademia delle scienze di torino 57 (2): 351 - 430. [pls 1 - 3] https: // doi. org / 10.5962 / bhl. title. 53748" title="Ricerche sui crostacei della Polinesia (Decapodi, Stomatopodi, Anisopodi e Isopodi)." volume="57" year="1907">Nobili, 1907: 390</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="066AE3DC761650DB632D37AAD9C4321E">
<treatmentCitationGroup id="A947C04FD9F24AE7B7C1890902B9FFB9">
=
<treatmentCitation id="3E5FD88006352D9AA3244FA9E786977E">
<taxonomicName id="4957673F4C6AC033C01255687A6D0A6C" ID-CoL="4NZPQ" authority="(Odhner, 1925)" baseAuthorityName="Odhner" baseAuthorityYear="1925" class="Malacostraca" family="Xanthidae" genus="Pseudoliomera" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ruppellioides">
<emphasis id="92C51DC89565127EA7D10C72F17CC004" italics="true">Pseudoliomera ruppellioides</emphasis>
(
<bibRefCitation id="71E39D86EA24F2F19FB4677369D2B3C0" author="Odhner T" refId="B31" refString="Odhner T (1925) Monographierte gattungen der krabben-familie Xanthidae. I. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar (4) 29 (1): 1 - 92. [figs 1 - 7, pls 1 - 5]" year="1925">Odhner, 1925</bibRefCitation>
)
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETSCI" type="material">
<paragraph id="1891FDC7F629F6C39AFDA0786EADD751">
<heading id="A30F3375DD50E30D3BA2CB792043A134" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="92D4BAB91066113CA86659FCCB176539">
<materialsCitation id="D8975FBC4D13381E8424DFA1B9E6B371" ID-GBIF-Occurrence="4904612371" collectingDate="2022-05-06" collectorName="Ziming Yuan &amp; Yuli Sun &amp; Shaobo Ma" country="China" elevation="10" latitude="9.664435" location="Yongshu Reef" longLatPrecision="1" longitude="113.01472" municipality="Nansha Islands" specimenCount="1" specimenCount-male="1">
<collectingCountry id="0DE38D59FA2E22BC127B872786F9B27F" name="China">China</collectingCountry>
<specimenCount id="E3A0B81FA0D90B7452FF91A4B4D8AFF3" count="1" type="male">1 male</specimenCount>
;
<abbrev id="ABBRID0EZSCI" xlink_title="maximum carapace width">CW</abbrev>
3.0 mm,
<abbrev id="ABBRID0E4SCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="D60E6BF3E51FDAB5F5517929C29A5DB9" metricMagnitude="-3" metricUnit="m" metricValue="1.9" unit="mm" value="1.9">1.9 mm</quantity>
;
<location id="69616167728D6984DB98014BC7BC0ED6" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:69616167728D6984DB98014BC7BC0ED6" country="China" latitude="9.664435" longLatPrecision="1" longitude="113.01472" municipality="Nansha Islands" name="Yongshu Reef">Yongshu Reef</location>
,
<collectingMunicipality id="6DBDECD8A8B439C752AC1CD01A42A73A">Nansha Islands</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EETCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[113.014717,9.664436]}">
<geoCoordinate id="B1B7E0A10E79DF68221382605BB3F0B6" degrees="9" direction="north" minutes="39" orientation="latitude" precision="1" seconds="51.97" value="9.664435">9 ° 39 ' 51.97 &quot; N</geoCoordinate>
,
<geoCoordinate id="3FE62FB893D9F366480F589E9012C954" degrees="113" direction="east" minutes="0" orientation="longitude" precision="1" seconds="52.98" value="113.01472">113 ° 0 ' 52.98 &quot; E</geoCoordinate>
</named-content>
;
<quantity id="C480B503818290D695E24705C66F8DBD" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">
<elevation id="355119ECA2952A3132DBFD12C452DCC1" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</elevation>
</quantity>
;
<date id="C473DC9B44BAC891F140742E12AFF6C2" value="2022-05-06">
<collectingDate id="96A2D0CCDADFB0565655FB2EABDE0DAD" value="2022-05-06">6 May 2022</collectingDate>
</date>
;
<collectorName id="8EF73ED8B68EE50AA1E8545732788DC9">Ziming Yuan</collectorName>
,
<collectorName id="AD010F78F5BE455B5E0CFFFA03628674">Yuli Sun</collectorName>
,
<collectorName id="01F421C7AE00121F8DA109CD4C0061FC">Shaobo Ma</collectorName>
coll.;
<specimenCode id="4324BD66AC64855F4B82AD69859F1F5F">NS-YS- 2022-1226</specimenCode>
</materialsCitation>
<materialsCitation id="070FCF060E12ADD2C010F1F2AEAE6DD7" ID-GBIF-Occurrence="4904612365" collectingDate="2022-05-14" collectorName="Ziming Yuan &amp; Yuli Sun &amp; Shaobo Ma" country="China" elevation="10" latitude="9.664435" location="Yongshu Reef" longLatPrecision="1" longitude="113.01472" municipality="Nansha Islands" specimenCount="1" specimenCount-male="1">
<specimenCount id="64143DDE5A81371FA415BEF1F5793A1D" count="1" type="male">1 male</specimenCount>
;
<abbrev id="ABBRID0EJTCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="35EF27101060270DFAE6BFBBBE356465" metricMagnitude="-3" metricUnit="m" metricValue="7.6" unit="mm" value="7.6">7.6 mm</quantity>
,
<abbrev id="ABBRID0ENTCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="46FC9729617DA26011BB9BC3DC3C0095" metricMagnitude="-3" metricUnit="m" metricValue="5.2" unit="mm" value="5.2">5.2 mm</quantity>
; same collection data as for preceding;
<date id="64F4A3DD7766239980BA32DEE288824D" value="2022-05-14">
<collectingDate id="9B75DED4E615279C462EAFA2CCAAED84" value="2022-05-14">14 May 2022</collectingDate>
</date>
;
<specimenCode id="5DE70E070A87E3D9554FA0820F8A905C">NS-YS- 2022-1227</specimenCode>
</materialsCitation>
<materialsCitation id="4DAC3EE557DA2C2A9999BA6F86E330C0" ID-GBIF-Occurrence="4904612362" collectingDate="2022-05-14" collectorName="Ziming Yuan &amp; Yuli Sun &amp; Shaobo Ma" country="China" elevation="10" latitude="9.664435" location="Yongshu Reef" longLatPrecision="1" longitude="113.01472" municipality="Nansha Islands" specimenCount="2" specimenCount-juvenile="2">
<specimenCount id="1A9FA6A5EC7747CEF54BB8BC33EC7F37" count="2" type="juvenile">2 juveniles</specimenCount>
;
<abbrev id="ABBRID0ERTCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="799B82EF7B75A4FD38ED3C22E324F11A" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.12.2 mm</quantity>
,
<abbrev id="ABBRID0EVTCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="366ED183598F2B6FE22F23648ED764FA" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.51.6 mm</quantity>
; same collection data as for preceding;
<specimenCode id="0C36DA1DC0AB6A05FD6DCCD585D5852F">NS-YS- 2022-1263</specimenCode>
</materialsCitation>
<materialsCitation id="B2789EDE47864453ECC28177CBB45147" ID-GBIF-Occurrence="4904612355" collectingDate="2022-05-14" collectorName="Ziming Yuan &amp; Yuli Sun &amp; Shaobo Ma" country="China" elevation="10" latitude="9.664435" location="Yongshu Reef" longLatPrecision="1" longitude="113.01472" municipality="Nansha Islands" specimenCount="1" specimenCount-male="1">
<specimenCount id="D2B1BA381259C5A37338E5AE4603D5AE" count="1" type="male">1 male</specimenCount>
;
<abbrev id="ABBRID0EZTCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="D9511D1E9EF75B20ED515C7B9D27008B" metricMagnitude="-3" metricUnit="m" metricValue="6.7" unit="mm" value="6.7">6.7 mm</quantity>
,
<abbrev id="ABBRID0E4TCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="1B23E75FA4E89EB034D98CC799DC436C" metricMagnitude="-3" metricUnit="m" metricValue="4.4" unit="mm" value="4.4">4.4 mm</quantity>
; same collection data as for preceding;
<specimenCode id="C98FC8481906BD5347E13E183A4E8B09">NS-YS- 2022-1336</specimenCode>
</materialsCitation>
<materialsCitation id="38DCBDE710E380A3392E963CDDE8BF9D" ID-GBIF-Occurrence="4904612364" collectingDate="2022-05-07" country="China" elevation="8" latitude="9.877275" location="Meiji Reef" longLatPrecision="1" longitude="115.521416" municipality="Nansha Islands" specimenCount="1" specimenCount-juvenile="1">
<specimenCount id="5918638A789DEB5BCF2CA7683DA2E3E3" count="1" type="juvenile">1 juvenile</specimenCount>
;
<abbrev id="ABBRID0EBUCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="F9782292B97BB7621EE36F6036F19C9C" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
,
<abbrev id="ABBRID0EFUCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="902A300AA13A32D9AA39F04FCB1CD043" metricMagnitude="-3" metricUnit="m" metricValue="1.7" unit="mm" value="1.7">1.7 mm</quantity>
;
<location id="93EFD81921D799B4661E2ABBE12A1C79" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:93EFD81921D799B4661E2ABBE12A1C79" country="China" latitude="9.877275" longLatPrecision="1" longitude="115.521416" municipality="Nansha Islands" name="Meiji Reef">Meiji Reef</location>
,
<collectingMunicipality id="52CD706C8BCCFF50E5609E44B9BF9F82">Nansha Islands</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMUCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[115.521411,9.877275]}">
<geoCoordinate id="286A05E84F57F5DD4170FDEA7603B9E5" degrees="9" direction="north" minutes="52" orientation="latitude" precision="1" seconds="38.19" value="9.877275">9 ° 52 ' 38.19 &quot; N</geoCoordinate>
,
<geoCoordinate id="6EF4863E373FDE8A661AE7B89C7759CA" degrees="115" direction="east" minutes="31" orientation="longitude" precision="1" seconds="17.08" value="115.521416">115 ° 31 ' 17.08 &quot; E</geoCoordinate>
</named-content>
;
<quantity id="F0322E71DBD783A3195F012DB16E48D5" metricMagnitude="0" metricUnit="m" metricValue="8.0" unit="m" value="8.0">
<elevation id="E5A3C74605DB90A4A3E331957D0465E5" metricMagnitude="0" metricUnit="m" metricValue="8.0" unit="m" value="8.0">8 m</elevation>
</quantity>
;
<date id="CE54A75058CB6DB2E336CCCED61AFCC7" value="2022-05-07">
<collectingDate id="0B1434E3914B5CD20739E716550FE4DD" value="2022-05-07">7 May 2022</collectingDate>
</date>
;
<specimenCode id="BFEFB5922179037BEF2D53DD96668C92">NS-MJ- 2022-1438</specimenCode>
</materialsCitation>
<materialsCitation id="2B44967D06BF2CB8CF8DB5A35C12ADA8" ID-GBIF-Occurrence="4904612354" collectingDate="2022-05-10" country="China" elevation="10" latitude="9.8919" location="Meiji Reef" longLatPrecision="1" longitude="115.57279" municipality="Nansha Islands" specimenCount="1" specimenCount-juvenile="1">
<specimenCount id="99AE0D6377FA76D5102E44C8B091FB50" count="1" type="juvenile">1 juvenile</specimenCount>
;
<abbrev id="ABBRID0ERUCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="339D8A027BC22B2EFEA5E152A16F03E9" metricMagnitude="-3" metricUnit="m" metricValue="2.2" unit="mm" value="2.2">2.2 mm</quantity>
,
<abbrev id="ABBRID0EVUCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="302D6C63E64DB0DAC3A4EF9B20E0B5B7" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
; same collection data as for preceding but at
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3UCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[115.572792,9.891900]}">
<geoCoordinate id="860B9C25F90C370955194297EF8431F2" degrees="9" direction="north" minutes="53" orientation="latitude" precision="1" seconds="30.84" value="9.8919">9 ° 53 ' 30.84 &quot; N</geoCoordinate>
,
<geoCoordinate id="54C6649819BBF4C2AD8208A2AD357EFC" degrees="115" direction="east" minutes="34" orientation="longitude" precision="1" seconds="22.05" value="115.57279">115 ° 34 ' 22.05 &quot; E</geoCoordinate>
</named-content>
;
<quantity id="EDF23C3E79689C4CDE5250A95412CAAD" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">
<elevation id="349B6D27297A189C60CAC3F69AE19495" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</elevation>
</quantity>
;
<date id="B2C0AD28E18E2224E4C43CE5803AD8C4" value="2022-05-10">
<collectingDate id="3C151FC5CC708096D09C411B0522C034" value="2022-05-10">10 May 2022</collectingDate>
</date>
;
<specimenCode id="79F2170962CD69024832411B3D53B61A">NS-MJ- 2022-1789</specimenCode>
</materialsCitation>
<materialsCitation id="4A7F5534466BAE3E44C26A8DB1ED8704" ID-GBIF-Occurrence="4904612330" collectingDate="2022-05-19" country="China" elevation="10" latitude="16.967834" location="Qilianyu" longLatPrecision="1" longitude="112.26972" municipality="Xisha Islands" specimenCount="1" specimenCount-female="1">
<specimenCount id="4007FEADE9EFF3B95DD8176B68395088" count="1" type="female">1 female</specimenCount>
;
<abbrev id="ABBRID0EBVCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="3833E18C465A08C814492B1350AB87A7" metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
,
<abbrev id="ABBRID0EFVCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="9D9741B4F3F1979B126E065F3E474567" metricMagnitude="-3" metricUnit="m" metricValue="2.9" unit="mm" value="2.9">2.9 mm</quantity>
;
<location id="DFB0EF047D26FC279725AF3EF8CE560E" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:DFB0EF047D26FC279725AF3EF8CE560E" country="China" latitude="16.967834" longLatPrecision="1" longitude="112.26972" municipality="Xisha Islands" name="Qilianyu">Qilianyu</location>
,
<collectingMunicipality id="2484375DE2CEDF0CC339A701838FDA90">Xisha Islands</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMVCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[112.269722,16.967833]}">
<geoCoordinate id="99ED53A1D6EE12B2C45D19A30D7E859F" degrees="16" direction="north" minutes="58" orientation="latitude" precision="1" seconds="04.2" value="16.967834">16 ° 58 ' 04.2 &quot; N</geoCoordinate>
,
<geoCoordinate id="F781ECEF1AECC8E4448D5E337ED0D769" degrees="112" direction="east" minutes="16" orientation="longitude" precision="1" seconds="11.0" value="112.26972">112 ° 16 ' 11.0 &quot; E</geoCoordinate>
</named-content>
;
<quantity id="120F6DF5931C61E73E464FDB718CFC61" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">
<elevation id="D2FABCCED9197F6E7B5A65A57063AAFF" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</elevation>
</quantity>
;
<date id="648BD5E3EEC8E35BC5E10E1B6E365903" value="2022-05-19">
<collectingDate id="4748B161C5351B6FF47CBE3FA3124C15" value="2022-05-19">19 May 2022</collectingDate>
</date>
;
<specimenCode id="8A97733F6E55323E761F740609C96628">XS-QL- 2022-1014</specimenCode>
</materialsCitation>
<materialsCitation id="11AD43BFB175A6656DD1F7825972203F" ID-GBIF-Occurrence="4904612372" collectingDate="2023-05-08" collectorName="Aiyang Wang &amp; Bingqin Liu" country="China" latitude="17.116806" location="Bei Reef" longLatPrecision="1" longitude="111.534225" municipality="Xisha Islands" specimenCount="1" specimenCount-juvenile="1">
<specimenCount id="F88E0C0FD96933C6BA067B8F5AA69EA5" count="1" type="juvenile">1 juvenile</specimenCount>
;
<abbrev id="ABBRID0ERVCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="160B28CACDCF204D5370BAD1C2258274" metricMagnitude="-3" metricUnit="m" metricValue="2.9" unit="mm" value="2.9">2.9 mm</quantity>
,
<abbrev id="ABBRID0EVVCI" xlink_title="median carapace length">CL</abbrev>
2.0 mm;
<location id="89FC589C7140D44665E925A527DA86FF" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:89FC589C7140D44665E925A527DA86FF" country="China" latitude="17.116806" longLatPrecision="1" longitude="111.534225" municipality="Xisha Islands" name="Bei Reef">Bei Reef</location>
,
<collectingMunicipality id="1E960BFD0F67F08207B84514BD7E375C">Xisha Islands</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3VCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[111.534222,17.116806]}">
<geoCoordinate id="E6C4498B027DC5B4EB467D11601E0730" degrees="17" direction="north" minutes="07" orientation="latitude" precision="1" seconds="00.5" value="17.116806">17 ° 07 ' 00.5 &quot; N</geoCoordinate>
<geoCoordinate id="4FD9A3199E8C2D3A4F2D6C3D6AA48D29" degrees="111" direction="east" minutes="32" orientation="longitude" precision="1" seconds="03.2" value="111.534225">111 ° 32 ' 03.2 &quot; E</geoCoordinate>
</named-content>
;
<date id="6FFC2CC680CF829E0EABD53DEB04612A" value="2023-05-08">
<collectingDate id="92F071A1D6C47ECC34576032D02E5EE7" value="2023-05-08">8 May 2023</collectingDate>
</date>
;
<collectorName id="FBFDE507FEC1BF25C809EDD969330D45">Aiyang Wang</collectorName>
,
<collectorName id="8BBF981294B02B6697DA10C582DA5E7D">Bingqin Liu</collectorName>
coll.; 2305089358
</materialsCitation>
<materialsCitation id="12B106D8AA98B33CA7EC8061772D8E99" ID-GBIF-Occurrence="4904612340" collectingDate="2021-06-05" collectorName="Geng Qin" country="China" elevation="21" location="Zhongsha Islands" specimenCount="1" specimenCount-male="1">
<specimenCount id="D157A662096177599E278868E057A4CC" count="1" type="male">1 male</specimenCount>
; not measured;
<location id="31E1DF964A68B9FA2543D460DF1590CE" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:31E1DF964A68B9FA2543D460DF1590CE" country="China" name="Zhongsha Islands">Zhongsha Islands</location>
;
<quantity id="E081AF21B0B5B62591A784E290CD713F" metricMagnitude="1" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" unit="m" value="20.5" valueMax="23.0" valueMin="18.0">
<elevation id="BBCC07FE57040DA497B9CB0548131289" metricMagnitude="1" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" unit="m" value="20.5" valueMax="23.0" valueMin="18.0">1823 m</elevation>
</quantity>
; dead coral;
<date id="A65E4BBC2E13B9C21415EF85FB3238F9" value="2021-06-05">
<collectingDate id="E149C30519D7634D71E45B582456C97E" value="2021-06-05">5 Jun. 2021</collectingDate>
</date>
;
<collectorName id="C850BF7661600D11175416112060C09E">Geng Qin</collectorName>
coll.; C 13-5
</materialsCitation>
<materialsCitation id="21FBEF5680C13AF31B2F2CDC6AD43F5D" ID-GBIF-Occurrence="4904612325" collectingDate="2021-06-09" collectorName="Geng Qin" country="China" elevation="21" location="Zhongsha Islands" specimenCount="1" specimenCount-male="1">
<specimenCount id="7C499DD9EB7C9BF44DD1955668642575" count="1" type="male">1 male</specimenCount>
; not measured; same collection data as for preceding,
<date id="C0CB01EB27C37A3D540B97DEAC2D96CD" value="2021-06-09">
<collectingDate id="6005F1F4AAC35B5D675E0F9BB4B996EC" value="2021-06-09">9 Jun. 2021</collectingDate>
</date>
; C 57-3
</materialsCitation>
<materialsCitation id="EB95CCBFF014A7AB85A37B880B6E0AC4" ID-GBIF-Occurrence="4904612368" collectingDate="2020-06-26" collectorName="Wei Jiang &amp; Geng Qin" country="China" elevation="20" latitude="15.88625" location="Zhongsha Islands" longLatPrecision="1" longitude="114.7916" specimenCount="1" specimenCount-male="1">
<specimenCount id="2443D28EB1B86E53CE24E0C71334B1FC" count="1" type="male">1 male</specimenCount>
;
<abbrev id="ABBRID0EBWCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="92CA1F3DA4027401559729AE127E3B13" metricMagnitude="-3" metricUnit="m" metricValue="5.4" unit="mm" value="5.4">5.4 mm</quantity>
,
<abbrev id="ABBRID0EFWCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="34A76285353605C37B57A974EE595EDF" metricMagnitude="-3" metricUnit="m" metricValue="3.6" unit="mm" value="3.6">3.6 mm</quantity>
;
<location id="651F47D68128CCD14EFDA0245D120FBC" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:651F47D68128CCD14EFDA0245D120FBC" country="China" latitude="15.88625" longLatPrecision="1" longitude="114.7916" name="Zhongsha Islands">Zhongsha Islands</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMWCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[114.791600,15.886250]}">
<geoCoordinate id="85651C4326AD88ED975F0B38E1EB1354" degrees="15" direction="north" minutes="53" orientation="latitude" precision="1" seconds="10.5" value="15.88625">15 ° 53 ' 10.5 &quot; N</geoCoordinate>
,
<geoCoordinate id="CDCF40955576743B7D1137CBD668BC8C" degrees="114" direction="east" minutes="47" orientation="longitude" precision="1" seconds="29.76" value="114.7916">114 ° 47 ' 29.76 &quot; E</geoCoordinate>
</named-content>
,
<quantity id="7B1E37FD6697CC4B4BECED351ACB214F" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
<elevation id="8FC84D36296BBB450D97CCE90D94943A" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</elevation>
</quantity>
;
<date id="2F2CC0AFAF3538D7AF5485275D8CC7D9" value="2020-06-26">
<collectingDate id="3A8EEEB1F5F215ADAD129A7A6C386B53" value="2020-06-26">26 Jun. 2020</collectingDate>
</date>
;
<collectorName id="15D2CB0DFD7F9C08988ADD6D22727448">Wei Jiang</collectorName>
,
<collectorName id="B169AC22BAC0F8ED5B77E92B0556C2FB">Geng Qin</collectorName>
coll.; ZS 233 C 07
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERWCI" type="material">
<paragraph id="99AB7DB7CE7FA3E0399B1084A17821CB">
<heading id="077A33E0FE742393D6A04281CDC17E3B" reason="title">Comparative material.</heading>
</paragraph>
<paragraph id="9350DC998FECB7CD9F63873177D4E6A4">
<taxonomicName id="3CA2F9017876D21E869F29A8F8C48A29" authority="Guinot, 1976" authorityName="Guinot" authorityYear="1976" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="mutatus">
<emphasis id="C0197A5FB8FBB9040B14D74D70AA54AA" bold="true">
<emphasis id="831C49EE74D8CD214EE6E3F0DDDDD9C9" bold="true" italics="true">Actaeodes mutatus</emphasis>
<bibRefCitation id="DAFAF82A13045B9B1FE1219F432CB0DF" author="Guinot D" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustaces Decapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Memoires du Museum National d'Histoire Naturelle, Paris: A, 97, 1 - 308. [figs 1 - 47, pls 1 - 19]" year="1976">Guinot, 1976</bibRefCitation>
</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="D339FB342A68525AC9C5DDF2FE5589C4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza &amp; PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron &amp; Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 F</figureCitation>
).
<materialsCitation id="8E214D3E876E7583C32B1E3706595854" ID-GBIF-Occurrence="4904612350" collectingDate="2007-11-25" collectorName="Xan" country="China" location="Hainan" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
<collectingCountry id="AA9723FBD2763D1B8E25E0F5EA711254" name="China">China</collectingCountry>
<specimenCount id="04B5D9941957A6634D213CC46F71C0AB" count="1" type="male">1 male</specimenCount>
; Lingao,
<collectingRegion id="0C166C8369194EF2AB7AF305B2A35C11" country="China" name="Hainan">Hainan</collectingRegion>
Island;
<date id="A646FC0A6ED088ED27294DFB64EF86DC" value="2007-11-25">
<collectingDate id="3785B460A319DAA2C8182E74E1836798" value="2007-11-25">25 Nov. 2007</collectingDate>
</date>
; Xan 074
</materialsCitation>
<materialsCitation id="2215CA4DB1B071D56782872A99DF8B9C" ID-GBIF-Occurrence="4904612314" collectorName="Island &amp; Xu Zhang" country="China" location="Hainan" specimenCount="1" specimenCount-female="1" stateProvince="Hainan">
<specimenCount id="54F198BC00E225D5E6F811A68F8CD40F" count="1" type="female">1 female</specimenCount>
;
<collectingRegion id="DD01F4D3DE9C374AFEF85909533D66B9" country="China" name="Hainan">Hainan</collectingRegion>
<collectorName id="4FBEE8032701AA2398D21469C565D1C7">Island</collectorName>
; 2022;
<collectorName id="53B5172628E3C2EF7A22374832B3D844">Xu Zhang</collectorName>
coll.; BF 01
</materialsCitation>
<materialsCitation id="DEA05F9D06159617CE93712C66EE9995" ID-GBIF-Occurrence="4904612318" collectorName="Xu Zhang" country="China" location="Phoenix Island" municipality="Sanya" specimenCount="1" specimenCount-female="1" stateProvince="Hainan Island">
<specimenCount id="B0E21B859DD4B372DF9F1A1A9C6F2F53" count="1" type="female">1 female</specimenCount>
;
<location id="6AD5A3FDFD3B9E46780ADDDED4032ABF" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:6AD5A3FDFD3B9E46780ADDDED4032ABF" country="China" municipality="Sanya" name="Phoenix Island" stateProvince="Hainan Island">Phoenix Island</location>
,
<collectingMunicipality id="45F628841599E66AD9484146F3E5239B">Sanya</collectingMunicipality>
,
<collectingRegion id="74604C7B6A6605EAB7CA54C675E8FA87">Hainan Island</collectingRegion>
; 2021;
<collectorName id="2C52CB9510BB010676C15B2A07D81FF1">Xu Zhang</collectorName>
coll.; BF 02.
<abbrev id="ABBRID0EJXCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="1B93A49BE8ED0014872267343ABEBF32" metricMagnitude="-2" metricUnit="m" metricValue="1.78" metricValueMax="2.08" metricValueMin="1.48" unit="mm" value="17.8" valueMax="20.8" valueMin="14.8">14.820.8 mm</quantity>
,
<abbrev id="ABBRID0ENXCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="B612CFBA9EC858A6C106252B40D9FCCC" metricMagnitude="-2" metricUnit="m" metricValue="1.055" metricValueMax="1.25" metricValueMin="0.86" unit="mm" value="10.55" valueMax="12.5" valueMin="8.6">8.612.5 mm</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="FB52D686E7020C9133649E96298360F9">
<taxonomicName id="4F8BEA1991BF8DCCFAE2CCE1236E3533" authority="(Ruppell, 1830)" baseAuthorityName="Ruppell" baseAuthorityYear="1830" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="hirsutissimus">
<emphasis id="7018656E70CA2447A4D47968315A9EA5" bold="true">
<emphasis id="9172DBFF667DA43F1EEB9C248CE91A62" bold="true" italics="true">Actaeodes hirsutissimus</emphasis>
(
<bibRefCitation id="731A59748E11711DC071151923FBD090" author="Rüppell E" journalOrPublisher="Brönner, Frankfurt am Main" refId="B39" refString="Ruppell E (1830) Beschreibung und abbildung von 24 arten kurzschwanzigen krabben, als beitrag zur naturgeschichte des rothen meeres. H. L. Bronner, Frankfurt am Main, 28 pp. [6 pls]" title="Beschreibung und abbildung von 24 arten kurzschwänzigen krabben, als beitrag zur naturgeschichte des rothen meeres. H. L." year="1830">Rüppell, 1830</bibRefCitation>
)
</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="6C46B5C5442252A4CA737622CC5A7B18" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza &amp; PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron &amp; Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 G</figureCitation>
).
<materialsCitation id="493D02F9B1EFC892BB826E23B7FE3CDA" ID-GBIF-Occurrence="4904612348" collectingDate="1976-04-15" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Zhao Shu Island" specimenCode="MBM 164262" specimenCount="3" specimenCount-male="3">
<collectingCountry id="9B6B14695C4A3A70D2376D0689E12C57" name="China">China</collectingCountry>
<specimenCount id="EE87252F71D476713FBF7C62DB0C209B" count="3" type="male">3 males</specimenCount>
;
<collectingMunicipality id="8A817C4ECCC6A1BC7948BDAD4AE52B59">Zhao Shu Island</collectingMunicipality>
,
<location id="D39694FD087BF056FF1B7F5DEA2DC82C" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:D39694FD087BF056FF1B7F5DEA2DC82C" country="China" municipality="Zhao Shu Island" name="Xisha Islands">Xisha Islands</location>
;
<date id="90647EAB5EF8BCA47A373F49799632FC" value="1976-04-15">
<collectingDate id="F3C7B28F7D338B4A53A0779CF01A7855" value="1976-04-15">15 Apr. 1976</collectingDate>
</date>
;
<specimenCode id="D1FF2F5939659565F69C91DD0B18DCB6" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164262</specimenCode>
</materialsCitation>
<materialsCitation id="9DA1DA62823DE80B91953E424AEBA599" ID-GBIF-Occurrence="4904612321" collectingDate="1975-06-09" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Dong Island" specimenCode="MBM 164166" specimenCount="1" specimenCount-female="1">
<specimenCount id="59278366063B5FB5728D576B4CBDCB71" count="1" type="female">1 female</specimenCount>
; north of
<collectingMunicipality id="3953B45EAF97568040F815373CF2E580">Dong Island</collectingMunicipality>
,
<location id="DCD88389E30C8F95AE81EBBFB649B0D5" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:DCD88389E30C8F95AE81EBBFB649B0D5" country="China" municipality="Dong Island" name="Xisha Islands">Xisha Islands</location>
;
<date id="9D03DCE417E9CC03587E4788B1A904FD" value="1975-06-09">
<collectingDate id="25823D0B27EC47BC8C443524B1AE8E90" value="1975-06-09">9 Jun. 1975</collectingDate>
</date>
;
<specimenCode id="2DCB08F7444032814BCECBE4C7F5D076" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164166</specimenCode>
</materialsCitation>
<materialsCitation id="28FE064BE8EC1A11B5062174205544E8" ID-GBIF-Occurrence="4904612345" collectingDate="1975-06-09" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Dong Island" specimenCode="MBM 164155" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<specimenCount id="B81A7AF555C6E206A561FD8DBB9E44C0" count="2" type="male">2 males</specimenCount>
,
<specimenCount id="F62312F327A894A1DDCF160FF038256C" count="3" type="female">3 ovigerous females</specimenCount>
;
<collectingMunicipality id="744853272C57AAC908E6764FC00C12B7">Dong Island</collectingMunicipality>
,
<location id="FBF3827BC8A9534FD52907423BDF21F8" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:FBF3827BC8A9534FD52907423BDF21F8" country="China" municipality="Dong Island" name="Xisha Islands">Xisha Islands</location>
;
<date id="897E553A0415549A2C5AF11E8924C51F" value="1975-06-09">
<collectingDate id="8F2A362708FE554D4E4F2CEBD92C65B7" value="1975-06-09">9 Jun. 1975</collectingDate>
</date>
;
<specimenCode id="ED2C8FDB7F94CA89977AE064C49C5CAE" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164155</specimenCode>
</materialsCitation>
<materialsCitation id="E3809E7A29438A7DE4F17E45625E9DFA" ID-GBIF-Occurrence="4904612369" collectingDate="1975-06-10" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Northeast of Dong Island" specimenCode="MBM 164148" specimenCount="5" specimenCount-female="1" specimenCount-juvenile="3" specimenCount-male="1">
<specimenCount id="23E246E87B9D26E5C9338C97A0AA5867" count="1" type="male">1 male</specimenCount>
,
<specimenCount id="59293304C07924899B47B18E43DDDF0B" count="1" type="female">1 female</specimenCount>
,
<specimenCount id="FE9706BE4F808762717AB726FF5760C3" count="3" type="juvenile">3 juveniles</specimenCount>
;
<collectingMunicipality id="098D77CDECD7FC2548DB52FC33F41FA9">Northeast of Dong Island</collectingMunicipality>
,
<location id="AAC717CE359B32872D755077F65BE5F2" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:AAC717CE359B32872D755077F65BE5F2" country="China" municipality="Northeast of Dong Island" name="Xisha Islands">Xisha Islands</location>
;
<date id="203EC38FFA1095952C76BAD1CD0D76B6" value="1975-06-10">
<collectingDate id="44DCEA022C14979630EB9828DE04D107" value="1975-06-10">10 Jun. 1975</collectingDate>
</date>
;
<specimenCode id="A82A41539D048FE4EE8A60BA85991E72" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164148</specimenCode>
</materialsCitation>
<materialsCitation id="45991D06D461C5FAE8D97A13B7884030" ID-GBIF-Occurrence="4904612301" collectingDate="1981-06-02" collectingDateMax="1981-06-04" collectingDateMin="1981-06-02" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Rocky Island" specimenCode="MBM 164151" specimenCount="6" specimenCount-female="2" specimenCount-male="4">
<specimenCount id="84DF1D22E3BF9ED613FE2BEAF5DE716D" count="4" type="male">4 males</specimenCount>
,
<specimenCount id="E3D4B901F658B8B727E8A1B21438F928" count="2" type="female">2 females</specimenCount>
;
<collectingMunicipality id="E3536DBDE3F2A4BD7CDF98108B2C15D4">Rocky Island</collectingMunicipality>
,
<location id="69F4B890C323002ECE8B52C3C973A5AD" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:69F4B890C323002ECE8B52C3C973A5AD" country="China" municipality="Rocky Island" name="Xisha Islands">Xisha Islands</location>
;
<date id="212B44FD2C593CE75AAE8E4863A495AF" value="1981-06-02" valueMax="1981-06-04" valueMin="1981-06-02">
<collectingDate id="5F9D11AA161DD3172DAEB56884435260" value="1981-06-02" valueMax="1981-06-04" valueMin="1981-06-02">24 Jun. 1981</collectingDate>
</date>
;
<specimenCode id="A984FDBB2DD27AC65C6A632FBCAB9C57" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164151</specimenCode>
</materialsCitation>
<materialsCitation id="87B5EEB0F32DC18A6A6752394700BEA2" ID-GBIF-Occurrence="4904612359" collectingDate="1980-05-28" collectingDateMax="1980-05-30" collectingDateMin="1980-05-28" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Dong Island" specimenCode="MBM 164140" specimenCount="1" specimenCount-female="1">
<specimenCount id="8424BBA981BCBCB78794DB51698ECC00" count="1" type="female">1 female</specimenCount>
;
<collectingMunicipality id="1703F556F1FB71C837F4C1AE53FA9B82">Dong Island</collectingMunicipality>
,
<location id="055E679687352A89C4A1046690185834" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:055E679687352A89C4A1046690185834" country="China" municipality="Dong Island" name="Xisha Islands">Xisha Islands</location>
;
<date id="EFA167059B69E0751CC89989E53ED91A" value="1980-05-28" valueMax="1980-05-30" valueMin="1980-05-28">
<collectingDate id="F81D4AB475F486D117F55E03A749F7EB" value="1980-05-28" valueMax="1980-05-30" valueMin="1980-05-28">2830 May 1980</collectingDate>
</date>
;
<specimenCode id="15695493373C11E722A3A3DC746571A6" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164140</specimenCode>
</materialsCitation>
<materialsCitation id="CDA48AA296C78D428DF4B32B2B133FC6" ID-GBIF-Occurrence="4904612324" collectingDate="2019-07-10" country="China" location="Xisha Islands" municipality="Jinqing Island" specimenCount="2" specimenCount-juvenile="2">
<specimenCount id="D58EE38AA1E130214980F87006ACC769" count="2" type="juvenile">2 juveniles</specimenCount>
;
<collectingMunicipality id="D5246FB529122A3B4A99DC0CE87BC51F">Jinqing Island</collectingMunicipality>
,
<location id="7C51C20DF634B8796677CD836E401CA3" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:7C51C20DF634B8796677CD836E401CA3" country="China" municipality="Jinqing Island" name="Xisha Islands">Xisha Islands</location>
;
<date id="8BBB5A68869DE2C42F1547C43784E9B9" value="2019-07-10">
<collectingDate id="5A8D0D10E0E3BF4106AA1AC2AD747323" value="2019-07-10">10 Jul. 2019</collectingDate>
</date>
; MF 01
</materialsCitation>
<materialsCitation id="0084FD856E1905B3D7B0013936675E85" ID-GBIF-Occurrence="4904612310" collectingDate="2021-11-07" collectorName="E Xuan Port &amp; Xan" country="China" location="Island" municipality="Danzhou" specimenCount="2" specimenCount-male="2" stateProvince="Hainan">
<specimenCount id="87D86F2CDA3C4A7173D75EBB9608C2B4" count="2" type="male">2 males</specimenCount>
;
<collectorName id="AD20BC4CBB7FEDEC5B7BA2011A9E172F">E Xuan Port</collectorName>
,
<collectingMunicipality id="5A5BD1FFA73CB12AE6C25EE2C9FAF102">Danzhou</collectingMunicipality>
,
<collectingRegion id="7B082EF91D0785B2E5592801A58CAD06" country="China" name="Hainan">Hainan</collectingRegion>
<location id="3936999BA31FB1EC2CA8493BBC77922B" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:3936999BA31FB1EC2CA8493BBC77922B" country="China" municipality="Danzhou" name="Island" stateProvince="Hainan">Island</location>
;
<date id="99794D29D32737E05BFF7C0CBB0A2205" value="2021-11-07">
<collectingDate id="A754A4C69B4E628FBBFAB6B9A81B41DC" value="2021-11-07">7 Nov. 2021</collectingDate>
</date>
; Xan 179
</materialsCitation>
<materialsCitation id="B8D39249B2A51B4A75E874DE9F25D812" ID-GBIF-Occurrence="4904612335" collectingDate="2019-07-09" country="China" location="Xisha Islands" municipality="Yuzhuo Reef" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
<specimenCount id="E1EDE0D15F3D0AE6BBE561CEBE6EBFB3" count="1" type="male">1 male</specimenCount>
;
<collectingMunicipality id="C69D84FAD924D92B99BBF1BB11CEAB05">Yuzhuo Reef</collectingMunicipality>
,
<location id="D929361955936BF676C84821C13ABBA0" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:D929361955936BF676C84821C13ABBA0" country="China" municipality="Yuzhuo Reef" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
;
<date id="2E729737CF838A26EFA8B55B5DAA1572" value="2019-07-09">
<collectingDate id="BA23C0AB8AC6C0550234AB9FF25CEDC8" value="2019-07-09">9 Jul. 2019</collectingDate>
</date>
; MF 02
</materialsCitation>
<materialsCitation id="B28F2DB57C166AB0F658FE44E5233602" ID-GBIF-Occurrence="4904612338" collectingDate="1975-05-09" collectionCode="MBM" collectorName="Xianqiu Ren" country="China" location="Xisha Islands" municipality="Rocky Island" specimenCode="MBM 164298" specimenCount="18" specimenCount-female="9" specimenCount-male="9" stateProvince="Hainan">
<specimenCount id="33AE796956333B9121662E8B35E6817C" count="9" type="male">9 males</specimenCount>
,
<specimenCount id="60FB743C9D977C2593C388328DC6B77B" count="9" type="female">9 females</specimenCount>
;
<collectingMunicipality id="C68D59F314CB05E46BD21648255174E1">Rocky Island</collectingMunicipality>
,
<location id="0B853FE064711FF89A0F0B90F6B1AE83" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:0B853FE064711FF89A0F0B90F6B1AE83" country="China" municipality="Rocky Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
;
<date id="4CC966DA83004FF2E31E035F70D8C1A5" value="1975-05-09">
<collectingDate id="71F50748184413EA00C969D053AADF8C" value="1975-05-09">9 May 1975</collectingDate>
</date>
;
<collectorName id="2F40F84FAE9C6A0B90BEE2A2D2450CEB">Xianqiu Ren</collectorName>
coll.;
<specimenCode id="C0C22871BC2596CD426C45AA88D41C69" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164298</specimenCode>
</materialsCitation>
<materialsCitation id="3252FF1C96D9C45918E7F0E99A202955" ID-GBIF-Occurrence="4904612357" collectingDate="1980-05-19" collectingDateMax="1980-05-23" collectingDateMin="1980-05-19" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Coral Island" specimenCode="MBM 164189" specimenCount="15" specimenCount-female="5" specimenCount-juvenile="2" specimenCount-male="8" stateProvince="Hainan">
<specimenCount id="A7FC3BBCE3E43A56E6AAEDBD0432BC98" count="8" type="male">8 males</specimenCount>
,
<specimenCount id="BA1B10A9242C24624335007A364746AF" count="5" type="female">5 females</specimenCount>
,
<specimenCount id="4750077320ED44398E1B5198518C4E40" count="2" type="juvenile">2 juveniles</specimenCount>
;
<collectingMunicipality id="937F1C0EA8D924E85DF88D6EB2DFF923">Coral Island</collectingMunicipality>
,
<location id="CF878DC3119FEA377A79DA252A4BC737" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:CF878DC3119FEA377A79DA252A4BC737" country="China" municipality="Coral Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
;
<date id="730C05707DBBA5C7499E39676EBDF9EA" value="1980-05-19" valueMax="1980-05-23" valueMin="1980-05-19">
<collectingDate id="7658B197A50B34C7C6912406B18BB44C" value="1980-05-19" valueMax="1980-05-23" valueMin="1980-05-19">1923 May 1980</collectingDate>
</date>
;
<specimenCode id="799D7EC097F26EA8ADF316D27A038449" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164189</specimenCode>
</materialsCitation>
<materialsCitation id="C591D7647D15FDCDE5B94062D169D5BE" ID-GBIF-Occurrence="4904612312" collectingDate="1980-05-09" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Jinqing Island" specimenCode="MBM 164190" specimenCount="7" specimenCount-female="4" specimenCount-juvenile="1" specimenCount-male="2" stateProvince="Hainan">
<specimenCount id="5617F05DF87BF928CDCE6DF9C3D40CEF" count="2" type="male">2 males</specimenCount>
,
<specimenCount id="1B0D3D99990B28C7C19B9CE42FF16182" count="4" type="female">4 females</specimenCount>
,
<specimenCount id="A38BA6A7B4774427578D1C7BF72951C7" count="1" type="juvenile">1 juvenile</specimenCount>
;
<collectingMunicipality id="FE7D7F1CE8196E62F3BD44A3B6D78853">Jinqing Island</collectingMunicipality>
,
<location id="5C6AFCED88E635FA8E454FD78F3D3CCC" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:5C6AFCED88E635FA8E454FD78F3D3CCC" country="China" municipality="Jinqing Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
;
<date id="67FF72925FAC12F43FCD7C6155E34BDC" value="1980-05-09">
<collectingDate id="3EB4620D75A26FBEE25862E80502C23F" value="1980-05-09">9 May 1980</collectingDate>
</date>
;
<specimenCode id="232C8F1EF5E1F31EEA3187954F4AD7A2" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164190</specimenCode>
</materialsCitation>
<materialsCitation id="2852740F43316A89D439A5105FB1ADE5" ID-GBIF-Occurrence="4904612341" collectingDate="1989-05-12" collectionCode="MBM" country="China" location="Nansha Islands" municipality="Xian'e Reef" specimenCode="MBM 164196" specimenCount="1" specimenCount-female="1" stateProvince="Hainan">
<specimenCount id="A62584AFBE00C875FD5E999C72E2B3C3" count="1" type="female">1 female</specimenCount>
;
<collectingMunicipality id="EADC5E4D47476EF968307B768E3B3A4E">Xiane Reef</collectingMunicipality>
,
<location id="DE389CB27BAFBD77C14E7C37916E75AA" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:DE389CB27BAFBD77C14E7C37916E75AA" country="China" municipality="Xian'e Reef" name="Nansha Islands" stateProvince="Hainan">Nansha Islands</location>
;
<date id="F1420C2E39799801450FE80EEFE79224" value="1989-05-12">
<collectingDate id="12E3DBEA09381B457CDE26DF49008C6E" value="1989-05-12">12 May 1989</collectingDate>
</date>
;
<specimenCode id="9C317CAEFF0356204C846B3239D56E37" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164196</specimenCode>
</materialsCitation>
<materialsCitation id="673DB6D129F1F9DC9D94764A169B2F51" ID-GBIF-Occurrence="4904612316" collectingDate="1980-05-14" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Jinyin Island" specimenCode="MBM 164185" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Hainan">
<specimenCount id="085839170C70E6340B671A884F77A543" count="2" type="male">2 males</specimenCount>
,
<specimenCount id="0FDA38B00A829A054812C6EB8A9B42DC" count="2" type="female">2 females</specimenCount>
;
<collectingMunicipality id="2EA1C064118D1DC7FBC970BE8ED8FB89">Jinyin Island</collectingMunicipality>
,
<location id="3794F5307B5D1D80AC01E8E33336E8C2" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:3794F5307B5D1D80AC01E8E33336E8C2" country="China" municipality="Jinyin Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
;
<date id="ACB4869EADC71B21CF2F38517FAECCB8" value="1980-05-14">
<collectingDate id="C78A93D5AF382570D5C100DBD8F46D3E" value="1980-05-14">14 May 1980</collectingDate>
</date>
;
<specimenCode id="313BA574F959A255A9552DAC3014EFF7" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164185</specimenCode>
</materialsCitation>
<materialsCitation id="91C6F82518AFC2B5BEE7D59F17FFEE41" ID-GBIF-Occurrence="4904612322" collectingDate="1975-06-10" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Northeast of Dong Island" specimenCode="MBM 164261" specimenCount="1" stateProvince="Hainan">
2 ovigerous females;
<collectingMunicipality id="54F3E4677341347E109E4D893C568EF6">Northeast of Dong Island</collectingMunicipality>
,
<location id="5752625241D85BA36EB6E45C502C75D9" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:5752625241D85BA36EB6E45C502C75D9" country="China" municipality="Northeast of Dong Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
;
<date id="19772A696501E0BD5E269981856FD4CF" value="1975-06-10">
<collectingDate id="55AED2E03F7C27B8FEF85B2DEB2482F6" value="1975-06-10">10 Jun. 1975</collectingDate>
</date>
;
<specimenCode id="019C872783E3D6180DD44CB1B280DB63" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164261</specimenCode>
</materialsCitation>
<materialsCitation id="A6041D2C72FFF75555510A43370FC3E5" ID-GBIF-Occurrence="4904612319" collectingDate="1981-05-19" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Jinqing Island" specimenCode="MBM 164225" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Hainan">
<specimenCount id="A84C854064D3B8091881A40D56FC00EA" count="5" type="male">5 males</specimenCount>
,
<specimenCount id="43B01B49A2600DC4CF0B99030D18AD78" count="4" type="female">4 females</specimenCount>
;
<collectingMunicipality id="BCC0F534F53F9C093D41C27B5C3770B6">Jinqing Island</collectingMunicipality>
,
<location id="77E24AFE2405379E8C68C2F6014028DE" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:77E24AFE2405379E8C68C2F6014028DE" country="China" municipality="Jinqing Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
;
<date id="26DBF8BF7AC294D9A8D895DDBAFD8D18" value="1981-05-19">
<collectingDate id="83AA549679A3340DCDFF5C01666A9C4E" value="1981-05-19">19 May 1981</collectingDate>
</date>
;
<specimenCode id="5FF198DF7AEAC3E9F28ADB53964AA84E" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164225</specimenCode>
.
<abbrev id="ABBRID0EFYCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="074DDD68355231654B0E940742F8DFDD" metricMagnitude="-2" metricUnit="m" metricValue="2.095" metricValueMax="3.42" metricValueMin="0.77" unit="mm" value="20.95" valueMax="34.2" valueMin="7.7">7.734.2 mm</quantity>
,
<abbrev id="ABBRID0EJYCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="061F1B5AAA0321A9C00D7C98BFDB4976" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" metricValueMax="2.2" metricValueMin="0.52" unit="mm" value="13.6" valueMax="22.0" valueMin="5.2">5.222 mm</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="103DDF2BDA55CF3BC93A5285A19DD8B9">
<emphasis id="94A1AC94CD427EBFE0B86476BAF10966" bold="true">
<taxonomicName id="9FE4F138D68880C8B77F35FB410D49A2" baseAuthorityName="H. Milne-Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="tomentosus">
<emphasis id="9C63DBFE59125F6A43193516E0CEE330" bold="true" italics="true">Actaeodes tomentosus</emphasis>
</taxonomicName>
(H. Milne Edwards, 1834)
</emphasis>
(Fig.
<figureCitation id="DE6A0023304F13674AB79824035BE5E1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza &amp; PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron &amp; Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 H</figureCitation>
).
<materialsCitation id="779553F4A1D8C1F10F458296DEB65728" ID-GBIF-Occurrence="4904612327" collectingDate="2021-07-24" country="Russia" location="Wenchang" municipality="Island" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
<collectingCountry id="C50DDDE27B6B4AC7C203D0D1E208D9F1" name="China">China</collectingCountry>
<specimenCount id="A14315E6A35D5E024441D15C3E66AB1B" count="1" type="male">1 male</specimenCount>
;
<location id="948D1AE4F8682F505A944665DDBAB5D2" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:948D1AE4F8682F505A944665DDBAB5D2" country="Russia" municipality="Island" name="Wenchang" stateProvince="Hainan">Wenchang</location>
,
<collectingRegion id="A2544C7B7DCA78A0A2497A909CAE280D" country="China" name="Hainan">Hainan</collectingRegion>
<collectingMunicipality id="6EEC80E7E1402D5B64CA3369F3975ED9">Island</collectingMunicipality>
;
<date id="8789AC16F06EEBBE01F95548ABC2BCB3" value="2021-07-24">
<collectingDate id="9FCC0071DC4838FEAFFAE21D0BAA9336" value="2021-07-24">24 Jul. 2021</collectingDate>
</date>
; RF 01
</materialsCitation>
<materialsCitation id="B19244383FD4E783B189D692245B4DBC" ID-GBIF-Occurrence="4904612329" collectingDate="2008-03-29" collectionCode="MBM" collectorName="Ping Lan &amp; Yongqiang Wang" country="Russia" location="Xincun" municipality="Hainan Island" specimenCode="MBM 282509" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
<specimenCount id="535B359175D8E6D9BBC469E34BA47F75" count="1" type="male">1 male</specimenCount>
;
<location id="6E1D42190AED0B15CD4F68D4B47B85B4" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:6E1D42190AED0B15CD4F68D4B47B85B4" country="Russia" municipality="Hainan Island" name="Xincun" stateProvince="Hainan">Xincun</location>
,
<collectingMunicipality id="7F9C58BD859243D557C82637F9D55AE4">Hainan Island</collectingMunicipality>
;
<date id="07351E9A60D2F01AB7C3EE009BBA38D5" value="2008-03-29">
<collectingDate id="FF5EC17AEDB8E5F3E39D8060A3329BA5" value="2008-03-29">29 Mar. 2008</collectingDate>
</date>
;
<collectorName id="E5A4085731641C85227CFEE0B0CCD0B6">Ping Lan</collectorName>
,
<collectorName id="09F681D9E016368251582DDBD45A152B">Yongqiang Wang</collectorName>
coll.;
<specimenCode id="CC42CFBE47D440C09D29B856C51BAF99" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 282509</specimenCode>
</materialsCitation>
<materialsCitation id="029E0B37ED8BF61BB6E63C2ADABFFCFE" ID-GBIF-Occurrence="4904612339" collectingDate="2008-03-23" collectionCode="MBM" country="Russia" location="Xiaodonghai" municipality="Hainan Island" specimenCode="MBM 282414" specimenCount="3" specimenCount-male="3" stateProvince="Hainan">
<specimenCount id="2CE597F2078C55EB18F2CF7053604501" count="3" type="male">3 males</specimenCount>
;
<location id="D14ED0AAF16E5E9232DBCB1522B271C9" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:D14ED0AAF16E5E9232DBCB1522B271C9" country="Russia" municipality="Hainan Island" name="Xiaodonghai" stateProvince="Hainan">Xiaodonghai</location>
,
<collectingMunicipality id="33272AC97197FEBDCC96CB8A8A886EDC">Hainan Island</collectingMunicipality>
;
<date id="FA66BD90E77FAFBE9EAF7756FBBCDA0B" value="2008-03-23">
<collectingDate id="41BC84DDE1F3C7EFC1FF545ABC389426" value="2008-03-23">23 Mar. 2008</collectingDate>
</date>
;
<specimenCode id="13733459F849A7E49161CFD2498C2553" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 282414</specimenCode>
</materialsCitation>
<materialsCitation id="26961EF07346D3D16D64D3CDEBA7EC08" ID-GBIF-Occurrence="4904612366" collectingDate="2007-12-25" collectionCode="MBM" country="Russia" location="Xiaodonghai" municipality="Hainan Island" specimenCode="MBM 283216" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Hainan">
<specimenCount id="6BC6522DA8F3273A3E849DD5E450C917" count="4" type="male">4 males</specimenCount>
,
<specimenCount id="567773B4EACEA7ACDFAB0BF072A4A910" count="4" type="female">4 females</specimenCount>
; same collection data as for preceding;
<date id="F4DA36D546C96B4358707DACE00474F5" value="2007-12-25">
<collectingDate id="4D0FB6E843CFBDC7561A62D6F4BC9688" value="2007-12-25">25 Dec. 2007</collectingDate>
</date>
;
<specimenCode id="6A1E7ADE5BD44DDC62DCA2A92DB14480" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 283216</specimenCode>
</materialsCitation>
<materialsCitation id="52EBEEAF4003A4D6625F3357FFBF4D29" ID-GBIF-Occurrence="4904612367" collectingDate="2007-12-24" collectionCode="MBM" collectorName="Xan" country="Russia" location="Xiaodonghai" municipality="Hainan Island" specimenCode="MBM 283218" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Hainan">
<specimenCount id="1795927C8E9172C4D965D25914A0848E" count="9" type="male">9 males</specimenCount>
,
<specimenCount id="C5CC50DE464E04BEE4F50EF5395B1294" count="6" type="female">6 females</specimenCount>
; same collection data as for preceding;
<date id="416A8A981E7D7076539652EABBC9DE82" value="2007-12-24">
<collectingDate id="F5FEC54D8C4769D71A24A3DE823A8907" value="2007-12-24">24 Dec. 2007</collectingDate>
</date>
; Xan 045,
<specimenCode id="5C06EB74243FA52F84AA608A603C8C78" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 283218</specimenCode>
</materialsCitation>
<materialsCitation id="EE58E7A659035646A9E494960968C4DC" ID-GBIF-Occurrence="4904612323" collectingDate="2008-03-23" collectionCode="MBM" collectorName="Wei Jiang &amp; Xan" country="Russia" location="Xiaodonghai" municipality="Hainan Island" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Hainan">
<specimenCount id="B501F37C9A73BDDB0BE47D3C5953E6A8" count="2" type="male">2 males</specimenCount>
,
<specimenCount id="18F57DAACAAFD12615D026F20DEA9DB1" count="1" type="female">1 female</specimenCount>
; same collection data as for preceding;
<date id="CCAE1A9D2A63467CE0A3ACBF71B2DE24" value="2008-03-23">
<collectingDate id="4497FBD87114EBA4E4112A8C28146313" value="2008-03-23">23 Mar. 2008</collectingDate>
</date>
;
<collectorName id="3EB9EC0FE7F73DADAC0B7CF24BBB472A">Wei Jiang</collectorName>
coll.; Xan 041
</materialsCitation>
<materialsCitation id="B546CFFA35C8100BC9E73B2F6C67B569" ID-GBIF-Occurrence="4904612306" collectingDate="2015-05-11" collectorName="Xan" country="Russia" location="Langhua Reef" municipality="Xisha Islands" specimenCount="1" specimenCount-female="1" stateProvince="Hainan">
<specimenCount id="5C26B45C8F38B620CF3DDAF4EF3D477D" count="1" type="female">1 female</specimenCount>
;
<location id="7DD85FDEC6A8C9BEDEE5FC47A3A34840" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:7DD85FDEC6A8C9BEDEE5FC47A3A34840" country="Russia" municipality="Xisha Islands" name="Langhua Reef" stateProvince="Hainan">Langhua Reef</location>
,
<collectingMunicipality id="D7ED64B6856BBDA87866B3DBB344A5F6">Xisha Islands</collectingMunicipality>
;
<date id="411A07ABAB72DCC1008F2A79982CFC46" value="2015-05-11">
<collectingDate id="02AA22B031D4EB178B15B6CEB749CB1A" value="2015-05-11">11 May 2015</collectingDate>
</date>
; Xan 126
</materialsCitation>
<materialsCitation id="C41AA3F7636028DC5C7AAEB49C4E2314" ID-GBIF-Occurrence="4904612320" collectingDate="2018-03-22" collectorName="Xan" country="Russia" county="Hainan Island" location="Houhai" municipality="Sanya" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
<specimenCount id="42062EC0698FFCD9909851A8F9249AF9" count="1" type="male">1 male</specimenCount>
;
<location id="499C0D19DF86A7A5991A2D97E4CB866E" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:499C0D19DF86A7A5991A2D97E4CB866E" country="Russia" county="Hainan Island" municipality="Sanya" name="Houhai" stateProvince="Hainan">Houhai</location>
,
<collectingMunicipality id="B5D5DA7D5E36053A60DF934C6E56CC00">Sanya</collectingMunicipality>
,
<collectingCounty id="ED79979F0E1A81D378EF2E03A9922269">Hainan Island</collectingCounty>
;
<date id="3D8516447178917320AC0E0CC113CC69" value="2018-03-22">
<collectingDate id="1F52ABC110D2829051BEC9C3F3ECF8EE" value="2018-03-22">22 Mar. 2018</collectingDate>
</date>
; Xan 048
</materialsCitation>
<materialsCitation id="7C474067564EA29144FE80EC5EC7F138" ID-GBIF-Occurrence="4904612343" collectorName="Xu Zhang" country="Russia" location="Dadonghai" municipality="Hainan Island" specimenCount="1" specimenCount-female="1">
<specimenCount id="07D222ABB216D0D0A829D8C828209F36" count="1" type="female">1 female</specimenCount>
;
<location id="595F342951FD463269EA1DD1BA2BF5EF" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:595F342951FD463269EA1DD1BA2BF5EF" country="Russia" municipality="Hainan Island" name="Dadonghai">Dadonghai</location>
,
<collectingMunicipality id="7F6E848B09DDD71E1ECA95F845666B70">Hainan Island</collectingMunicipality>
; 2021;
<collectorName id="5663EE7A1192AFAFDE5A728DA80CB0D1">Xu Zhang</collectorName>
coll.; RF 03
</materialsCitation>
<materialsCitation id="C8BAFF2BD17FDBAD9AF25B2A97DF19CF" ID-GBIF-Occurrence="4904612305" collectorName="Xu Zhang" country="Russia" location="Sanya" municipality="Hainan Island" specimenCount="1" specimenCount-male="1">
<specimenCount id="6EAF4F927DCEB91E022B96554AC85F12" count="1" type="male">1 male</specimenCount>
;
<location id="DBCFF3DB52002FF91266895C088B6C06" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:DBCFF3DB52002FF91266895C088B6C06" country="Russia" municipality="Hainan Island" name="Sanya">Sanya</location>
,
<collectingMunicipality id="CBCAEFC588B7644CD1C569616538D3C1">Hainan Island</collectingMunicipality>
; 2022;
<collectorName id="E4AC69BBDA79121E90ADE91402760CB9">Xu Zhang</collectorName>
coll.; RF 04
</materialsCitation>
<materialsCitation id="3AAC1CF0C9BBE4D41B648EF6F95C1118" ID-GBIF-Occurrence="4904612309" collectingDate="2019-08-20" collectorName="Yunhao Pan" country="Russia" location="Sanya" municipality="Hainan Island" specimenCount="1" specimenCount-female="1">
<specimenCount id="C19A64DDC4462579C83BD7E2D5F2DA18" count="1" type="female">1 female</specimenCount>
;
<location id="0B509C7980D44913978D90328AD1329F" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:0B509C7980D44913978D90328AD1329F" country="Russia" municipality="Hainan Island" name="Sanya">Sanya</location>
,
<collectingMunicipality id="40E3A9953B549F965B2C79D38DD13D7D">Hainan Island</collectingMunicipality>
;
<date id="27C341B9D8C5E81FDA53988782734154" value="2019-08-20">
<collectingDate id="5979EF6A2BE451C7B7A4AD44D9D8819B" value="2019-08-20">20 Aug. 2019</collectingDate>
</date>
;
<collectorName id="764BCAFE48C0DAFAB2EF77A833C2CEBB">Yunhao Pan</collectorName>
coll.; RF 05
</materialsCitation>
<materialsCitation id="FD700CF04E9E9B00EA8B433ED58D385A" ID-GBIF-Occurrence="4904612346" collectingDate="2019-07-09" country="Russia" location="Yuzhuo Reef" municipality="Xisha Islands" specimenCount="1" specimenCount-male="1">
<specimenCount id="0AD3F77312AD968D90630573B93B87ED" count="1" type="male">1 male</specimenCount>
;
<location id="579BFF26DA0A64BDA10F3C47206EAF9A" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:579BFF26DA0A64BDA10F3C47206EAF9A" country="Russia" municipality="Xisha Islands" name="Yuzhuo Reef">Yuzhuo Reef</location>
,
<collectingMunicipality id="89D1AE1D719727C43315E5E6FE171010">Xisha Islands</collectingMunicipality>
;
<date id="90AA2E51B8E2D7B75B2426ABB34EBE4D" value="2019-07-09">
<collectingDate id="2946E607B281B94900F42E1335EF178B" value="2019-07-09">9 Jul. 2019</collectingDate>
</date>
; RF 06
</materialsCitation>
<materialsCitation id="A2730B8F6B34CDCDCD6D6E514E5F5A99" ID-GBIF-Occurrence="4904612358" collectingDate="2019-07-11" country="Russia" location="Lingyang Reef" municipality="Xisha Islands" specimenCount="1" specimenCount-male="1">
<specimenCount id="F7C3ED71387A71C52544CA9DF451D3FD" count="1" type="male">1 male</specimenCount>
;
<location id="65227F5DC3732D485DBBEF59A1D363EF" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:65227F5DC3732D485DBBEF59A1D363EF" country="Russia" municipality="Xisha Islands" name="Lingyang Reef">Lingyang Reef</location>
,
<collectingMunicipality id="93E6844874F625A116AC64A9BB2CD42F">Xisha Islands</collectingMunicipality>
;
<date id="31B4E6FA6C16C091EC4DC2F1949DEEB0" value="2019-07-11">
<collectingDate id="115F84EEB6CF44803E1804C061BDCDCA" value="2019-07-11">11 Jul. 2019</collectingDate>
</date>
; RF 07
</materialsCitation>
<materialsCitation id="8BCDDDD7A986CB78FA1C24194DD3CD73" ID-GBIF-Occurrence="4904612351" collectingDate="1980-03-27" collectionCode="MBM" country="Russia" location="Yongxing Island" municipality="Xisha Islands" specimenCode="MBM 164194" specimenCount="1" specimenCount-male="1">
<specimenCount id="4C8800AE3ED17A0C8B13368CA139340A" count="1" type="male">1 male</specimenCount>
;
<location id="228DA2285ABCB85E1CF3BDC53664D6AC" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:228DA2285ABCB85E1CF3BDC53664D6AC" country="Russia" municipality="Xisha Islands" name="Yongxing Island">Yongxing Island</location>
,
<collectingMunicipality id="0C3D86456CAC040069082586A737505F">Xisha Islands</collectingMunicipality>
;
<date id="34DE282A4E95F20D19369D95F1A48216" value="1980-03-27">
<collectingDate id="78B10583DC63884578E8AB04C7446110" value="1980-03-27">27 Mar. 1980</collectingDate>
</date>
;
<specimenCode id="D718928FE4843A5B7A9F95648B14629C" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164194</specimenCode>
</materialsCitation>
<materialsCitation id="9D4B8A8F5FEBD712CEFAA8D855D5536B" ID-GBIF-Occurrence="4904612334" collectingDate="1990-11-22" collectionCode="MBM" country="Russia" location="Sanya Bay" municipality="Hainan Island" specimenCode="MBM 164492" specimenCount="1" specimenCount-female="1">
<specimenCount id="BEE8A701115951ED4F48AE9E41195C19" count="1" type="female">1 female</specimenCount>
;
<location id="E36884A0F3D13259E086C478919EFC6D" LSID="urn:lsid:plazi:treatment:9682DAD4848258339F5C7FD265643EF7:E36884A0F3D13259E086C478919EFC6D" country="Russia" municipality="Hainan Island" name="Sanya Bay">Sanya Bay</location>
,
<collectingMunicipality id="718EF4AA9C1D9F55953ABDCC0D15218B">Hainan Island</collectingMunicipality>
;
<date id="1926D43AD5F3644E0B3EE987F3019573" value="1990-11-22">
<collectingDate id="F16E41A16B331B1398F7AA6F28BEF7F5" value="1990-11-22">22 Nov. 1990</collectingDate>
</date>
;
<specimenCode id="660F221A1F5DA97C3276F0E93E469C19" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 164492</specimenCode>
.
<abbrev id="ABBRID0EBZCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="380BBFF5A3C29838B0E49330B2D6AC32" metricMagnitude="-2" metricUnit="m" metricValue="2.4299999999999997" metricValueMax="3.7399999999999998" metricValueMin="1.1199999999999999" unit="mm" value="24.299999999999997" valueMax="37.4" valueMin="11.2">11.237.4 mm</quantity>
,
<abbrev id="ABBRID0EFZCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="8F49401C55D4551C592C573ACD0CF988" metricMagnitude="-2" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="2.4899999999999998" metricValueMin="0.74" unit="mm" value="16.15" valueMax="24.9" valueMin="7.4">7.424.9 mm</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJZCI" type="description">
<paragraph id="C50BF5BD5A51DF4D35F47671F7D83D52">
<heading id="E4C49911696EB1D93C810B6773B1E016" reason="title">Description.</heading>
</paragraph>
<paragraph id="948D2B66F3699B9DF065DDAB5ACD98A7">
Carapace (Figs
<figureCitation id="1C2623FE6675B38F4D1F67FEC6D08A8C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A, B</figureCitation>
,
<figureCitation id="D94E658C5C2DD527196759EC54FA4802" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 B</figureCitation>
,
<figureCitation id="8F572B9184B975F2C0B8F3A95513A228" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A, D, G</figureCitation>
) broader than long,
<abbrev id="ABBRID0E2ZCI" xlink_title="maximum carapace width">CW</abbrev>
about 1.5 times the
<abbrev id="ABBRID0E6ZCI" xlink_title="median carapace length">CL</abbrev>
, dorsal surface bearing round granules, granules interspersed with short pubescence; regions well defined, grooves wide and deep, 1 M separated from 2 M; 2 M completely divided; 3 M distinct, divided into three lobes; 4 M distinct; 1 L, 4 L indistinct; 2 L, 3 L, 5 l, 6 L distinct, 5 L, 6 L partially divided; 1 P, 2 P distinct; front broad, about 0.3 times
<abbrev id="ABBRID0ED1CI" xlink_title="maximum carapace width">CW</abbrev>
, not protruding, divided into two slightly triangular lobes by a wide and deep V-shaped notch, frontal lobes and dorsal inner orbital angle separated by shallow depression; eyestalks densely granulated; Anterolateral margin divided into four teeth by narrow but sometimes opened fissures; first tooth extremely flattened, second tooth broader, sometimes obtuse, third tooth prominent, fourth tooth slightly smaller than third; carapace widest at third tooth; posterolateral margin almost straight; subhepatic region densely granulated; Posterior margin nearly straight.
</paragraph>
<caption id="C1CD8F1DC18DE312BAADBB603CEDE7AD" ID-DOI="10.3897/zse.100.117859.figure7" ID-arpha="6C2A31F1-CA19-51B7-84BA-ECE755CFCB11" httpUri="https://binary.pensoft.net/fig/1088100" startId="F7">
<paragraph id="E7DDFB87BD0DB60739BC34A2707A46EB">
<label id="1C6F87BCFD3FD9EC9DE72DA5DD248843">Figure 7.</label>
</paragraph>
<paragraph id="65829DFF04D86551E2FD216EA6EA1BD0">
<taxonomicName id="8BD72294E0C449FC408A17F75C64C674" authority="(A. Milne-Edwards, 1873)" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="53212766DDA3E8EC2E05A120BAE5CCE6" italics="true">Gothus consobrinus</emphasis>
(A.
<bibRefCitation id="E9FECC84CD1142AF9ADAD6786067544A" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" journalOrPublisher="Journal des Museum Godeffroy" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustaces nouveaux ou peu connus provenant du musee de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253 - 264. [pls. 12 - 13] https: // doi. org / 10.5962 / bhl. title. 10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards, 1873</bibRefCitation>
)
</taxonomicName>
, male,
<abbrev id="ABBRID0E11CI" xlink_title="maximum carapace width">CW</abbrev>
6.7 mm,
<abbrev id="ABBRID0E51CI" xlink_title="median carapace length">CL</abbrev>
4.4 mm,
<specimenCode id="85E8D658C253630D255E09B5B5EC125D">NS-YS- 2022-1336</specimenCode>
.
<emphasis id="A7B8E4B2F87071685A2B7143E7413ACD" bold="true">A.</emphasis>
Dorsal habitus;
<emphasis id="E4F0D5354DFFAC02DB2C8D0E94EAAB66" bold="true">B.</emphasis>
Carapace;
<emphasis id="7AF80A0BA3022745FD1716AF94D067B5" bold="true">C.</emphasis>
Frontal view;
<emphasis id="90581261461F5C53D863541D5C158F05" bold="true">D.</emphasis>
Thoracic sternites and pleon;
<emphasis id="84A391AF68B27E77AA12E9F93B530206" bold="true">E.</emphasis>
Chelipeds. Scale bar: 2 mm.
</paragraph>
</caption>
<caption id="807CBFF02A5036CB71407844D1C871C8" ID-DOI="10.3897/zse.100.117859.figure8" ID-arpha="2E9C389E-B3F4-500A-A62F-477E632A74A5" httpUri="https://binary.pensoft.net/fig/1088101" startId="F8">
<paragraph id="83B01DE670A4C357AD46D605DA3FE52A">
<label id="18A85C4D703E14A577A6DEA8889CACE8">Figure 8.</label>
</paragraph>
<paragraph id="F43CFA2D00527913B3ECCC65F2B2AA45">
<taxonomicName id="614462F99CB2798455478E9981EE2FAA" authority="(A. Milne-Edwards, 1873)" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="E64A6E8F43FF726EC10C0223960B8D85" italics="true">Gothus consobrinus</emphasis>
(A.
<bibRefCitation id="B73C4DCFCAD954CC80C659E533D90BD6" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" journalOrPublisher="Journal des Museum Godeffroy" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustaces nouveaux ou peu connus provenant du musee de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253 - 264. [pls. 12 - 13] https: // doi. org / 10.5962 / bhl. title. 10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards, 1873</bibRefCitation>
)
</taxonomicName>
, male,
<abbrev id="ABBRID0EE3CI" xlink_title="maximum carapace width">CW</abbrev>
6.7 mm,
<abbrev id="ABBRID0EI3CI" xlink_title="median carapace length">CL</abbrev>
4.4 mm,
<specimenCode id="72D01809AC2FA036944AA68258E19780">NS-YS- 2022-1336</specimenCode>
.
<emphasis id="6AFB34B36AD1A7B682A2E606539552D1" bold="true">A.</emphasis>
Carapace;
<emphasis id="B416D67F9E74E1829F5473112E3ADC46" bold="true">B.</emphasis>
Maxilliped 3;
<emphasis id="91C5DF32814BA406F8A281D63DF5E7F5" bold="true">C.</emphasis>
Cheliped;
<emphasis id="7BB858CE8ABF9E7A3926CDACDF2B91BA" bold="true">D.</emphasis>
Pereopod 5;
<emphasis id="DD6A55C947D414F8BD36ABE08D7354D0" bold="true">E.</emphasis>
Pleon;
<emphasis id="599B7849E920027C172344FFA0952A41" bold="true">F.</emphasis>
Left
<abbrev id="ABBRID0EY3CI" xlink_title="first gonopod of male">G 1</abbrev>
, ventral view;
<emphasis id="A4D3EE6191E3A7FDA4C406FB2C6BCAE3" bold="true">G.</emphasis>
Left
<abbrev id="ABBRID0E53CI" xlink_title="first gonopod of male">G 1</abbrev>
;
<emphasis id="BEBA5584599334221E94DEFC103F3CD6" bold="true">H.</emphasis>
Left
<abbrev id="ABBRID0EE4CI" xlink_title="second gonopod of male">G 2</abbrev>
, ventral view. Scale bar: 2 mm (
<emphasis id="F809671930867E19E6EEB13034397730" bold="true">A</emphasis>
); 0.5 mm (
<emphasis id="27569B97B4A857F429FA63B26B86C8C6" bold="true">B, F, H</emphasis>
); 1 mm (
<emphasis id="B1A313339D445C1F547F6D06EEC9E7A9" bold="true">C, D, E</emphasis>
); 0.1 mm (
<emphasis id="1F9175D70DF42464F0C83C83FC53C9FD" bold="true">G</emphasis>
).
</paragraph>
</caption>
<caption id="5D80E164E51E87A88F362E2A35FD2930" ID-DOI="10.3897/zse.100.117859.figure9" ID-arpha="8A8080B2-BC39-550C-B8B7-3E4AFE725B0A" httpUri="https://binary.pensoft.net/fig/1088102" startId="F9">
<paragraph id="25B359A8BB34C859C9EE41FD3CC67B0A">
<label id="AE93A8AA39DA8D70361CEA2F7232F2FB">Figure 9.</label>
</paragraph>
<paragraph id="E7BC921BB065DBF94B7AFEA67E8CBFAC">
<taxonomicName id="E0B093D2120B4B77DB51C5D73DB28D98" authority="(A. Milne-Edwards, 1873)" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="70C43A2BD21E28A20B5A76B88AC1BAE3" italics="true">Gothus consobrinus</emphasis>
(A.
<bibRefCitation id="6054F6EA64E92345838FBC4DE0773036" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" journalOrPublisher="Journal des Museum Godeffroy" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustaces nouveaux ou peu connus provenant du musee de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253 - 264. [pls. 12 - 13] https: // doi. org / 10.5962 / bhl. title. 10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards, 1873</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="98CA80D16A7CBF20187056B1DA09F6B4" bold="true">A C.</emphasis>
Female,
<abbrev id="ABBRID0EK5CI" xlink_title="maximum carapace width">CW</abbrev>
10.5 mm,
<abbrev id="ABBRID0EO5CI" xlink_title="median carapace length">CL</abbrev>
6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S);
<emphasis id="620E7AECE2797C5FFF61DC38582EB9D0" bold="true">D F.</emphasis>
Female,
<abbrev id="ABBRID0EU5CI" xlink_title="maximum carapace width">CW</abbrev>
8.8 mm,
<abbrev id="ABBRID0EY5CI" xlink_title="median carapace length">CL</abbrev>
5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne;
<emphasis id="FAEC22892B6C9C02CE8295D37BE8AAA8" bold="true">G I.</emphasis>
Female,
<abbrev id="ABBRID0E55CI" xlink_title="maximum carapace width">CW</abbrev>
4.5 mm,
<abbrev id="ABBRID0EC6CI" xlink_title="median carapace length">CL</abbrev>
2.9 mm,
<specimenCode id="BA3FFEA6F96D0A478A74790C5E9ADB0D">XS-QL- 2022-1014</specimenCode>
. Scale bar: 2 mm (
<emphasis id="47C517C255A0818FDEEBE2347B9DD57E" bold="true">A F</emphasis>
); 1 mm (
<emphasis id="E83A0D66A2C39E8F4672188BF9465E8E" bold="true">G I</emphasis>
).
</paragraph>
</caption>
<paragraph id="DD465E47ECEFF8877ABE470C32FE12BB">
Epistome (Figs
<figureCitation id="F3CBD7C862350EEA9B553F5034ACD2DD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 C</figureCitation>
,
<figureCitation id="4BCA8BB5180D3B7CB8896D88B0741B97" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 B, E, H</figureCitation>
) Central region with median projection on posterior margin; lateral regions with undulating posterior margins. Maxilliped 3 (Figs
<figureCitation id="5CE1032C0867AD9C7EEF422AA1207CE2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 C</figureCitation>
,
<figureCitation id="2C00C0DCEAACA9CA649A3DA79AE4563C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 B</figureCitation>
) granulated, ischium subrectangular, presenting submedian groove; merus subquadrate; anterior margin indented; anteroexternal angle square-shaped expanded. Antennule (Fig.
<figureCitation id="03BA82569F24C44B405ADB4F54DF6C60" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 C</figureCitation>
,
<figureCitation id="62413EC8AD74047AFA676E81741A87DF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 B, E, H</figureCitation>
) folding transversely; basal segment of antenna subrectangular; contacting ventral external frontal margin and ventral internal orbital angle; antennal flagellum filling orbital hiatus.
</paragraph>
<paragraph id="8D4EAE06C781109BC4CEFF1AAEE1F83E">
Chelipeds (Figs
<figureCitation id="8870962DD6E2529E7E104885A4B07461" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 E</figureCitation>
,
<figureCitation id="47884CF24883C9512239880962CEF16C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 C</figureCitation>
) symmetrical, meri short; carpi robust, nearly spherical, surface granulated, aggregated into nodules; outer and dorsal surfaces of palms densely granulated; fingers elongated, with triangular teeth, tips sharp, crossing at extremities when closed; dorsal surface of movable finger with three granular ridges; outer surface of immovable finger with two ridges. Fingers brownish-black, coloration of immovable finger extending onto palm along both inner and outer surfaces in male.
</paragraph>
<paragraph id="1669AEE94C8999B35E06B801D83691AC">
Ambulatory legs (Figs
<figureCitation id="0C328EA1EEE24EE69BDD96475B2D415E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A</figureCitation>
,
<figureCitation id="240020AC789502A3F9A56B21F116EEF8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 D</figureCitation>
) with meri flattened, P 5 merus length about 3 times as wide as distal end, granulated dorsally and along anterior and posterior edges; carpi granulated dorsally and along anterior edge; dorsal surface with a grooved indentation near anterior edge; propodi granulated dorsally and along edges; dactyli elongated, margins armed with granules and setae, terminal end chitinous, sharp, slightly recurved, dactylo-propodal lock present.
</paragraph>
<paragraph id="9FCD2789B9A422363105D5F7BBE1A0F0">
Male thoracic sternum (Fig.
<figureCitation id="1CD93F2CF578812FB22994E0552D764C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 D</figureCitation>
) with sternites 1 and 2 completely fused, suture between sternites 2 and 3 straight, complete, sternites 3 and 4 mostly fused, suture between them visible only at margins, sternites 3 short, sternite 4 with central longitudinal groove, tubercle of sterno-pleonal lock (press-button mechanism) located on posterior of sternite 5. Male pleon (Figs
<figureCitation id="D557A4BDCD3D8AC65174C129E6A4216D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 D</figureCitation>
,
<figureCitation id="BB10522B26FF9C4F2A3AE46D96B3A9FC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 E</figureCitation>
) very narrow; pleonites 3 to 5 completely fused; lateral margins of pleonite 6 slightly concave; telson long, broad, longer than wide; truncated oval; basal margin wider than terminal margin of pleonite 6.
</paragraph>
<paragraph id="58D14D2800AA897F4F1DBD1D555A0429">
<abbrev id="ABBRID0ENBDI" xlink_title="first gonopod of male">G 1</abbrev>
(Fig.
<figureCitation id="972DF2E5883135AD1EDA2720DDD77CF0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 F G</figureCitation>
) slender, distal lobe prominent, elongated, curved upwards, long setae on inner subdistal side, small spines on outer side.
<abbrev id="ABBRID0EVBDI" xlink_title="second gonopod of male">G 2</abbrev>
(Fig.
<figureCitation id="03CECDD10963DD0959C5C5037D31CF3C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 H</figureCitation>
) short, distal lobe elongated, slightly curved upwards.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4BDI" type="Live coloration">
<paragraph id="DDB3C70C9AF40C6DAF317399282A5CAD">
<heading id="A585D46D19D8CE53362A642B86C9DEFA" reason="title">Live coloration.</heading>
</paragraph>
<paragraph id="7525FEFE6570E1D404F1153694CA423C">
Overall white to ivory-colored, carapace adorned with symmetrical black to brown stripes and orange spots, ambulatory legs and chelipeds bearing black to brown stripes, cheliped palm dorsal surface and anterior part of carpus sometimes coral pink (Fig.
<figureCitation id="FFF832831429CF6290740DF16BA30AEF" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Gothus consobrinus (A. Milne-Edwards, 1873). A. Male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336; B. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014, displaying live coloration. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure10" httpUri="https://binary.pensoft.net/fig/1088103">10</figureCitation>
). Fingers brownish-black, coloration of immovable finger extending onto inner and outer surfaces of palm in male (Fig.
<figureCitation id="A4F312C7323B90691A794EF2A28890C4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 E</figureCitation>
).
</paragraph>
<caption id="1E87DBBAAD4F8B4111094C211A137BA1" ID-DOI="10.3897/zse.100.117859.figure10" ID-arpha="1618F2E5-67B4-55B7-BAA0-980F5DCE6548" httpUri="https://binary.pensoft.net/fig/1088103" startId="F10">
<paragraph id="EE33B2E394BDB4D1962BF04BEC7D8929">
<label id="96AA36A71ADB85B46EDEB1F2291BCB98">Figure 10.</label>
</paragraph>
<paragraph id="EEB71B31DC2F93CBECF8BA8C25E488E5">
<taxonomicName id="AE3B4A548DD31967802D5CDAAA36DC3F" authority="(A. Milne-Edwards, 1873)" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="E963D235B64DD0AA6AD4C2E772FA3920" italics="true">Gothus consobrinus</emphasis>
(A.
<bibRefCitation id="C46616AB31845A799F3F92875E64B326" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" journalOrPublisher="Journal des Museum Godeffroy" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustaces nouveaux ou peu connus provenant du musee de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253 - 264. [pls. 12 - 13] https: // doi. org / 10.5962 / bhl. title. 10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards, 1873</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B04178BE802D086F5D23460B6F06620A" bold="true">A.</emphasis>
Male,
<abbrev id="ABBRID0EADDI" xlink_title="maximum carapace width">CW</abbrev>
6.7 mm,
<abbrev id="ABBRID0EEDDI" xlink_title="median carapace length">CL</abbrev>
4.4 mm,
<specimenCode id="1E72CCD213747606B2382D24175B5198">NS-YS- 2022-1336</specimenCode>
;
<emphasis id="0785649EA59BCB81F793E11013EE91EA" bold="true">B.</emphasis>
Female,
<abbrev id="ABBRID0EKDDI" xlink_title="maximum carapace width">CW</abbrev>
4.5 mm,
<abbrev id="ABBRID0EODDI" xlink_title="median carapace length">CL</abbrev>
2.9 mm,
<specimenCode id="674DD1A85A05D7656BF82F6074FB40C1">XS-QL- 2022-1014</specimenCode>
, displaying live coloration. Scale bar: 2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EXDDI" type="distribution">
<paragraph id="C7AF17BCDAA3487319D82A88677EEAEA">
<heading id="329433626DF2A499A1E4DF833EDCA752" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="94B37844961099D3DA7F2B612F0E0ED9">
Distributed in the Zhongsha (= Macclesfield Bank), Xisha (= Paracel Islands), and Nansha Islands (=
<collectingCountry id="E8255C942B6817BF95B8FE4E20DEBE5D" name="Spratly Islands">Spratly Islands</collectingCountry>
) of the
<collectingCountry id="B6411829CF6C111BD98B366703937D22" name="China">China</collectingCountry>
Sea; widely found in the Indo-West Pacific, with the type locality at Upolu Island, inhabiting crevices in shallow coral reefs.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3DDI" type="remarks">
<paragraph id="69F696F5DE5C0A0C57DE7F2DBCE57803">
<heading id="81EE82B43A5E7861AB76E1BC625C26D8" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="FCF0B44B2161A77FAA902B60AE62FA69">
This report constitutes the first record of this species in the Chinese sea. Alphonse
<bibRefCitation id="80A8EF7E59B6A44E30509F070E0E4910" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" firstAuthor="Milne-Edwards" journalOrPublisher="Journal des Museum Godeffroy" page="255" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253264. [pls. 1213] https://doi.org/10.5962/bhl.title.10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards (1873: 255</bibRefCitation>
[79]) briefly described “
<taxonomicName id="36467A0C599CE795945DE8D4802A5BF8" authorityName="Nobili" authorityYear="1907" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrina">
<emphasis id="148AF0A997DF3360FB3411E4B792A5E5" italics="true">Actaea consobrina</emphasis>
</taxonomicName>
” from Upolu in present-day
<collectingCountry id="C3A0F7E36B58A2A6DE3C07685FE0DEC4" name="Samoa">Samoa</collectingCountry>
and provided the carapace measurements of
<specimenCount id="8DE7351F1458BBF75D591D2FFE0A0693" count="1" type="generic">one specimen</specimenCount>
(
<abbrev id="ABBRID0EREDI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="7F85089976C840AE372DACAF5E980A08" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
,
<abbrev id="ABBRID0EVEDI" xlink_title="median carapace length">CL</abbrev>
<quantity id="B56E51F58DA3120F26722C5D3944F7AD" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
), though they did not indicate the sex of the specimen nor how many other specimens they examined. This species remained in the genus
<taxonomicName id="BBEBFE2D62EB14CB6ABD31EBE09AEFE4" authorityName="De Haan" authorityYear="1833" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="6EE91D0974E9BDF0428235157075EFB3" italics="true">Actaea</emphasis>
</taxonomicName>
until
<bibRefCitation id="F1200A3062373DF33AD818AB361F04BF" author="Guinot D" refId="B13" refString="Guinot D (1967 b) A propos des affinités des Genres Dairoides Stebbing et Daira de Haan. In: Recherches Préliminaires sur les Groupements Naturels chez les Crustacés, Décapodes, Brachyoures, III. Bulletin du Muséum National dHistoire Naturelle, Paris 2 39 (3): 540563. [figs 136]">Guinot (1967 b</bibRefCitation>
) transferred it to
<taxonomicName id="C2C78E00C279365395426651B6B769B0" authorityName=", Guinot" authorityYear="1976" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="FF199C1A63CE6A4FF88C77267B2811C6" italics="true">Actaeodes</emphasis>
</taxonomicName>
. As part of her revision of some
<taxonomicName id="E5FFA299CCABF5615267F8387D4FC598" authorityName="Alcock" authorityYear="1898" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="subFamily" subFamily="Actaeinae">Actaeinae</taxonomicName>
genera, including
<taxonomicName id="F61F2988AF0C5AAAC82BC6F76A293A26" authority=", Guinot (1976: 246)" authorityName=", Guinot" authorityPageNumber="246" authorityYear="1976" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="CE6931644D5410E04A8BBD0004A08786" italics="true">Actaeodes</emphasis>
,
<bibRefCitation id="EEACFD9CB4D0C19B8906BCA3E8418DEE" author="Guinot D" firstAuthor="Guinot" page="246" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">Guinot (1976: 246)</bibRefCitation>
</taxonomicName>
examined
<specimenCount id="6639A048C15C5D111CE660DCB3F745FE" count="2" type="female">two female</specimenCount>
specimens collected by A. Milne-Edwards and deposited in the
<collectionCode id="ED3A96CB291E963FC112D1A17F25ED93" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
(MP-B 3885 S from Upolu and a specimen without a collection number from “
<collectingCountry id="0EE44396735E0E2A6BF7F85845D0D178" name="Samoa">Samoa</collectingCountry>
? ”). She pointed out that based on the measurements (
<abbrev id="ABBRID0E2FDI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="6588234A3DDDBD3D09BA52F78E1BFAF7" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
,
<abbrev id="ABBRID0E6FDI" xlink_title="median carapace length">CL</abbrev>
<quantity id="340EA19BBE1C0703DA4D9E74F0339C1C" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
), this specimen MP-B 3885 S could be the
<typeStatus id="0248B4FFEEE7297B153DB4EA7681B7B0" type="holotype">holotype</typeStatus>
of
<taxonomicName id="51C660C93722A499079B230E2F5496CD" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="BDE55E53FE1B16D0D369C5932549C4B8" italics="true">Actaeodes consobrinus</emphasis>
</taxonomicName>
. For the purposes of this study, we consider this specimen to be typical of
<taxonomicName id="B2FD25E4207B525F943CB4AAB8D4FD33" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="D3B16E830C46A390B8D08443CCB41B1A" italics="true">A. consobrinus</emphasis>
</taxonomicName>
and use it for our comparisons (Fig.
<figureCitation id="2CA9322D6287A5F63DDF6550CB490BD5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A C</figureCitation>
). It exhibits a nearly truncated second anterolateral tooth with open fissures between the anterolateral teeth, which is also observed in some of the current specimens (Fig.
<figureCitation id="A5C23D79E7506CC361816089976DC7CB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 G I</figureCitation>
). Compared to the specimens from the
<collectingCountry id="C2600D0F4D561E711B2DBFB495D711EC" name="China">China</collectingCountry>
Sea, it has deeper grooves on the carapace, which may be attributed to growth-related differences considering its larger size. The other female individual (MNHN-IU- 2024-3483) found together with this specimen is likely the one collected by A. Milne-Edwards that lacks a collection number (
<bibRefCitation id="0D048E7E0AE341746697C9036E17B595" author="Guinot D" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustaces Decapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Memoires du Museum National d'Histoire Naturelle, Paris: A, 97, 1 - 308. [figs 1 - 47, pls 1 - 19]" year="1976">Guinot, 1976: 246</bibRefCitation>
;
<abbrev id="ABBRID0EBHDI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="84DB8FF335BE42B0D10B0539B33DEC08" metricMagnitude="-3" metricUnit="m" metricValue="8.5" unit="mm" value="8.5">8.5 mm</quantity>
,
<abbrev id="ABBRID0EFHDI" xlink_title="median carapace length">CL</abbrev>
<quantity id="26A4144BA21A2145934F9C29829C5221" metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
), based on its measurements (
<abbrev id="ABBRID0EJHDI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="178D8331BFD198D6540737383CED91B2" metricMagnitude="-3" metricUnit="m" metricValue="8.8" unit="mm" value="8.8">8.8 mm</quantity>
,
<abbrev id="ABBRID0ENHDI" xlink_title="median carapace length">CL</abbrev>
<quantity id="467CEB96E9CAAA02C7307926E197EA36" metricMagnitude="-3" metricUnit="m" metricValue="5.7" unit="mm" value="5.7">5.7 mm</quantity>
; Fig.
<figureCitation id="F4474BC3252DBCF7511667430EA924AF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 D F</figureCitation>
). This specimen possesses triangular anterior lateral teeth and shallower grooves on the carapace (Fig.
<figureCitation id="E331DA239E282B5C825D60CE756A26C4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 D F</figureCitation>
).
</paragraph>
<paragraph id="69F0FEB43E8DF80A343493DA8ADA0CBD">
There are some issues regarding the classification of this species within its genus: Alphonse
<bibRefCitation id="767D794CAE789E634453A96C4AD87BAE" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" firstAuthor="Milne-Edwards" journalOrPublisher="Journal des Museum Godeffroy" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253264. [pls. 1213] https://doi.org/10.5962/bhl.title.10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards (1873)</bibRefCitation>
put the species in
<taxonomicName id="28F0C681FB09EF201484576B8BFDBDA8" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="0AF2108ACE37809D8573EBD7E5D67730" italics="true">Actaea</emphasis>
De Haan, 1833
</taxonomicName>
, and initially compared it with
<taxonomicName id="6B92595029FF7B213633B2FF30481DF1" authority="(Ruppell, 1830)" baseAuthorityName="Ruppell" baseAuthorityYear="1830" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="hirsutissimus">
<emphasis id="ABB397244592FEF9902FE36317DAF548" italics="true">Actaea hirsutissimus</emphasis>
(
<bibRefCitation id="C8CACFBFE5538F54CCD51517AF6AD891" author="Rüppell E" journalOrPublisher="Brönner, Frankfurt am Main" refId="B39" refString="Ruppell E (1830) Beschreibung und abbildung von 24 arten kurzschwanzigen krabben, als beitrag zur naturgeschichte des rothen meeres. H. L. Bronner, Frankfurt am Main, 28 pp. [6 pls]" title="Beschreibung und abbildung von 24 arten kurzschwänzigen krabben, als beitrag zur naturgeschichte des rothen meeres. H. L." year="1830">Rüppell, 1830</bibRefCitation>
)
</taxonomicName>
(presently known as
<taxonomicName id="68859EDB9B09E42115C79BE7BE58C3CA" authorityName=", Guinot" authorityYear="1976" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="D617C3CEC3F74E8BD5F75AB058077BF7" italics="true">Actaeodes</emphasis>
</taxonomicName>
) and
<taxonomicName id="00AED65770619BB33010D10168303BD4" authority="(Heller, 1860)" baseAuthorityName="Heller" baseAuthorityYear="1860" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="kraussi">
<emphasis id="26F322145A90A00DC1B28B5753BD1DB8" italics="true">Actaea kraussi</emphasis>
(Heller, 1860)
</taxonomicName>
(presently known as
<taxonomicName id="82A52D6AF4E19D9A234920D72B7121F3" authorityName="A. Milne-Edwards" authorityYear="1869" class="Malacostraca" family="Xanthidae" genus="Banareia" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="353F18247D511C38BA4B4FBE7952D736" italics="true">Banareia</emphasis>
</taxonomicName>
) and primarily considered it similar to the former.
<bibRefCitation id="DBA04637B6564936EBD109450B99B53D" author="Sakai T" firstAuthor="Sakai" refId="B40" refString="Sakai T (1939) Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo Co., Tokyo, 365741. [figs 1129, pls 42111, table 1]" year="1939">Sakai (1939)</bibRefCitation>
considered the species to be close to
<taxonomicName id="37BDF401E9C6C7EDE13A0C9FDB2F2E53" authority="(Odhner, 1925)" baseAuthorityName="Odhner" baseAuthorityYear="1925" class="Malacostraca" family="Xanthidae" genus="Paractaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="tumulosa">
<emphasis id="7B756EF9A6C5DCC40978B0B51E759784" italics="true">Paractaea tumulosa</emphasis>
(
<bibRefCitation id="EEFCF015DAF74FF38DDEB3F13CDA7808" author="Odhner T" refId="B31" refString="Odhner T (1925) Monographierte gattungen der krabben-familie Xanthidae. I. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar (4) 29 (1): 1 - 92. [figs 1 - 7, pls 1 - 5]" year="1925">Odhner, 1925</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="712DFB1DB2E9D0CDD973EE9A12CE259B" author="Guinot D" refId="B13" refString="Guinot D (1967 b) A propos des affinités des Genres Dairoides Stebbing et Daira de Haan. In: Recherches Préliminaires sur les Groupements Naturels chez les Crustacés, Décapodes, Brachyoures, III. Bulletin du Muséum National dHistoire Naturelle, Paris 2 39 (3): 540563. [figs 136]">Guinot (1967 b</bibRefCitation>
,
<bibRefCitation id="EFED52ED50CF50EA0341BB4A437AA37D" author="Guinot" firstAuthor="Guinot" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">1976</bibRefCitation>
) opposed the similarity to
<taxonomicName id="14C834307D21EA03C6EC1DF91044B405" baseAuthorityName="Odhner" baseAuthorityYear="1925" class="Malacostraca" family="Xanthidae" genus="Paractaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="tumulosa">
<emphasis id="9985CA3A0CDBD8CDE5AD4764A983F342" italics="true">P. tumulosa</emphasis>
</taxonomicName>
but acknowledged its relationship with
<taxonomicName id="B526E44BDC7CFAC3E44D39EB01C1CC7D" authorityName=", Guinot" authorityYear="1976" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="1847D2142A4EA5285E6606FA2B3B0A1C" italics="true">Actaeodes</emphasis>
</taxonomicName>
, and upon reviewing
<taxonomicName id="B806FC45CDD54744AF33C0875A2444D4" authorityName=", Guinot" authorityYear="1976" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="D6407A7625C7C4CB5190E68D20C8988B" italics="true">Actaeodes</emphasis>
</taxonomicName>
,
<taxonomicName id="EB69F32FD925C9B8E96488C804CE1004" authorityName="Nobili" authorityYear="1907" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrina">
<emphasis id="AC5FAA467FE4A30AF072587C93A2C53F" italics="true">Actaea consobrina</emphasis>
</taxonomicName>
was classified into the genus
<taxonomicName id="15158B2AFD1EA68AC917B59A77361A21" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="B1EEBA58CE784BE8BF0F0C4D41D12C25" italics="true">Actaeodes</emphasis>
<bibRefCitation id="679DCE0DC0095FF145319897EF192CF5" author="Dana JD" refId="B4" refString="Dana JD (1851) On the Classification of the Cancroidea. The American Journal of Science and Arts 2 12 (34): 121 - 131." year="1851">Dana, 1851</bibRefCitation>
</taxonomicName>
, and supplemented its morphological characteristics.
</paragraph>
<paragraph id="87ADA51B159F46A137EA222526655631">
In Guinots review (1976), the definition of
<taxonomicName id="2DDE70311ECB3A348288047460BB761F" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="A99FB81EE0EF2F0F3F92DFAA843CADE5" italics="true">Actaeodes</emphasis>
</taxonomicName>
includes the following characteristics: 1) carapace wide to very wide; 2) long anterolateral margins curving back over branchial regions, divided into lobes by fissures that extend as grooves into the subhepatic region; 3) very short posterolateral margins with a strong concavity that coapted against the last three pairs of ambulatory legs; 4) developed areolation of the dorsal surface with granular and pilosity lobules; 5). The frontal edge slopes downward with a central notch leading to the anterior tip of the epistome; the frontal lobes barely form a canopy above the antennules; 6) orbits round and relatively small, with specific fissures on supraorbital and exorbital edges; no infraorbital fissure 7) Equal and short chelipeds, with fingers either ending in a spoon-shaped tip or crossing at tips; 8) antenna fitting between front and orbit or with a closed orbit in
<taxonomicName id="917E8C452F824E8DCB23FA91A923CFE4" authority="(Ortmann, 1894)" baseAuthorityName="Ortmann" baseAuthorityYear="1894" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="semoni">
<emphasis id="61BD396F279DC2CBE4C9330AF3B675F7" italics="true">Actaeodes semoni</emphasis>
(Ortmann, 1894)
</taxonomicName>
; 9) small and slightly depressed epistome, with the anterior tip projecting forward to join the anterior median groove of the dorsal face; 10) short ambulatory legs; 11) sub-hepatic region grooved; 12) thoracic sternite 4 traversed by two transverse grooves and two oblique grooves, and with a very clear longitudinal groove, hidden by telson; a median line present at levels of sternites 6, 7, and 8; 13) male pleon with fused pleonites 35, elongated and projecting forward, featuring a median longitudinal swelling from 3 to 6 pleonites; 14)
<abbrev id="ABBRID0EAMDI" xlink_title="first gonopod of male">G 1</abbrev>
with a tapered distal lobe adorned with relatively short bristles.
</paragraph>
<paragraph id="D7DF83C62DC6BF40BFABFBA8FC1A4282">
<taxonomicName id="D8D3FD748FD0DF32269C205841A98C89" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="269BE5FEE747DB6669DF96C408BD0AFB" italics="true">Actaeodes</emphasis>
</taxonomicName>
currently comprises six species, among which the type species
<taxonomicName id="E04037B33AF5F0E7C9DAA5011364798F" baseAuthorityName="H. Milne-Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="tomentosus">
<emphasis id="2B23FFB971D305EF4C041CE53DDA5512" italics="true">A. tomentosus</emphasis>
</taxonomicName>
,
<taxonomicName id="2A86B3A7E8459C6F0EF640771C87AE81" authority="(Ortmann, 1894)" baseAuthorityName="Ortmann" baseAuthorityYear="1894" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="semoni">
<emphasis id="10BAA2A1705B4C10827A572E189014E1" italics="true">A. semoni</emphasis>
(Ortmann, 1894)
</taxonomicName>
,
<taxonomicName id="8C469E46369D52467AF095EA75785B1E" authority="(Ruppell, 1830)" baseAuthorityName="Ruppell" baseAuthorityYear="1830" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="hirsutissimus">
<emphasis id="6C7FA826BA5644C18D0226694105A31D" italics="true">A. hirsutissimus</emphasis>
(
<bibRefCitation id="E06AEF81A3BE1579C713C45DB30CAD6F" author="Rüppell E" journalOrPublisher="Brönner, Frankfurt am Main" refId="B39" refString="Ruppell E (1830) Beschreibung und abbildung von 24 arten kurzschwanzigen krabben, als beitrag zur naturgeschichte des rothen meeres. H. L. Bronner, Frankfurt am Main, 28 pp. [6 pls]" title="Beschreibung und abbildung von 24 arten kurzschwänzigen krabben, als beitrag zur naturgeschichte des rothen meeres. H. L." year="1830">Rüppell, 1830</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4E27AD30AB127113B2C11528096555E4" authority="Guinot, 1976" authorityName="Guinot" authorityYear="1976" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="mutatus">
<emphasis id="1FC7011B7B2E8DF375E03966A0A014BC" italics="true">A. mutatus</emphasis>
<bibRefCitation id="10AFC4ABFD165604A434A5838AEECBBB" author="Guinot D" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustaces Decapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Memoires du Museum National d'Histoire Naturelle, Paris: A, 97, 1 - 308. [figs 1 - 47, pls 1 - 19]" year="1976">Guinot, 1976</bibRefCitation>
</taxonomicName>
share a relatively similar appearance and match the description above. The most direct similarity is likely due to the very short posterolateral margins with a strong concavity that coapted against the last three pairs of ambulatory legs. However,
<taxonomicName id="E82F6B05D76DCA4ADE469D783C3E1178" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="5D86E3300F1E75A66AB437B329DDF3E1" italics="true">A. consobrinus</emphasis>
</taxonomicName>
and
<taxonomicName id="2D9D2FCAADCAB57B16AC57F1A3BFE976" authority="Garth &amp; Kim, 1983" authorityName="Garth &amp; Kim" authorityYear="1983" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="quinquelobatus">
<emphasis id="5D24E576E3CE8F6D10EC720813D926D5" italics="true">A. quinquelobatus</emphasis>
<bibRefCitation id="7843A3C3D66024E46EACD91FADB30C9E" DOI="10.1080/00222938300770561" author="Garth JS &amp; Kim HS" issue="5" journalOrPublisher="Journal of Natural History" pagination="663-729" refId="B10" refString="Garth JS, Kim HS (1983) Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908 - 1909. Journal of Natural History 17 (5): 663 - 729. https: // doi. org / 10.1080 / 00222938300770561" title="Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908-1909." volume="17" year="1983">Garth &amp; Kim, 1983</bibRefCitation>
</taxonomicName>
exhibit morphologies that differ significantly and may not fit well within the genus
<taxonomicName id="CD373FABE07F4FB9C1101348CA530649" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="39306540220E2C77CB16FEECC48B3DB8" italics="true">Actaeodes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="94BA952E0D84B5894AB2D84743897D51">
For the current species
<taxonomicName id="4863B2067C8025CE846DA01298565300" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="BC03E346E9FBFCA683DAFBA2F5147C31" italics="true">A. consobrinus</emphasis>
</taxonomicName>
, it indeed exhibits several features similar to typical
<taxonomicName id="9763F2BFD2160ABD1F46F79EBA0B3864">
<emphasis id="209060D1238AE6CA8F4DDB1CEFA614FD" italics="true">Actaeodes</emphasis>
species
</taxonomicName>
, which mainly include a carapace with developed dorsal areolation with granular and setae, longer anterolateral margins and shorter posterolateral margins, symmetrical chelipeds with sharply crossing tips, an elongated male pleon, and a
<abbrev id="ABBRID0EKPDI" xlink_title="first gonopod of male">G 1</abbrev>
that is overall similar in morphology. However, this similarity is superficial, and there are some undeniable differences between
<taxonomicName id="B1F92A7E8C416EF7146328CBB973BD00" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="FBBBADC4149E5E3B63DF831F611D6C00" italics="true">A. consobrinus</emphasis>
</taxonomicName>
and typical
<taxonomicName id="650581DD594B60FDC2DE96D346EC8EE6" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="7D3E85274A75972745470584AEBA9F21" italics="true">Actaeodes</emphasis>
species. A
</taxonomicName>
. Milne-Edwards, in the original description, compared
<taxonomicName id="BB881E6CD75ACB5BDE45BDA28436F9BB" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="D54D035A108D62A84D199B5FBF2CB17E" italics="true">A. consobrinus</emphasis>
</taxonomicName>
with
<taxonomicName id="142C6D0F1DF96444451A19CD113F177B" baseAuthorityName="Ruppell" baseAuthorityYear="1830" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="hirsutissimus">
<emphasis id="2F4B50632167C20D6B1C59E4BE6F5D14" italics="true">A. hirsutissimus</emphasis>
</taxonomicName>
, noting the absence of a pronounced concavity in its posterior margin. In current observations, the posterior margin of this species is almost straight (Figs
<figureCitation id="0D873E3F59CDC01145BBE606985A410A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A, B</figureCitation>
,
<figureCitation id="707B31229B8451F1B602E449CD8C4C44" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 A</figureCitation>
,
<figureCitation id="B9A901F68F859288F8C840EA73825755" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A, D, F</figureCitation>
), which significantly deviates from
<taxonomicName id="C2B9B42FC92EBA990C6B429F85A07872" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="5071EF657816F1AE09F03EF5D824C10C" italics="true">Actaeodes</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="4F857221EDBF4E1C3B2C675ADAFC1E9D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza &amp; PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron &amp; Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 F H</figureCitation>
; cf.
<bibRefCitation id="7B9D2440914C78344BA96C925CBE8C61" author="Guinot" firstAuthor="Guinot" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">Guinot 1976</bibRefCitation>
: pl. XV, figs 14). Furthermore, the morphology of the thoracic sternum in the current species markedly differs from
<taxonomicName id="92490B149E24EEC1D4D662CC2068BA1F" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="C3F23AD03055D751FA8CF5D300DCEEB7" italics="true">Actaeodes</emphasis>
</taxonomicName>
, with its third sternite being very short (Fig.
<figureCitation id="EDF6C7605F8D33BE9994C2EE45CAAC4E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 D</figureCitation>
) (vs. elongated third sternite in
<taxonomicName id="A1D4E9D8F6C9135E5650FF2106A295E5" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="0DEA1CFD291BD0B3B2605F5C32B4A198" italics="true">Actaeodes</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="28E5494D035AE882B0684A797DDE20BC" author="Guinot" firstAuthor="Guinot" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">Guinot 1976</bibRefCitation>
: fig. 41 C) and the fourth sternite lacking oblique grooves (Fig.
<figureCitation id="796ECD135917F80666D473CC4D74A103" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 D</figureCitation>
) (fourth thoracic sternite of
<taxonomicName id="A89518BF029C6077805BE1B1731ED979" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="48ED90A33771DC18721D759DBAB58F29" italics="true">Actaeodes</emphasis>
</taxonomicName>
traversed by two transverse grooves and two oblique grooves; cf.
<bibRefCitation id="F4FBB278544877A4B565DF1A5AB0D687" author="Guinot" firstAuthor="Guinot" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">Guinot 1976</bibRefCitation>
: fig. 41 C). Other differences include
<taxonomicName id="34B34355BE963EAB2736A1DA700C9B54" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="2182D46F094CAC5DCA9331CE9B9FA491" italics="true">A. consobrinus</emphasis>
</taxonomicName>
having an elongated pleon that barely extends beyond the coxo-sternal condyles of pereiopod 1 (Fig.
<figureCitation id="8F6FDA0ED8EFD23018948C016E7529E4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 D</figureCitation>
) (vs. pleon being significantly elongated, clearly surpassing the coxo-sternal condyles of pereiopod
<quantity id="6604D5B2845CD60E6D80857EBFAC3B39" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="FAFA4921EE78821F6BEE375BB9589117" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="6662CD1654ED1EAAD5EE43D56871AD16" italics="true">Actaeodes</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="8BCC83064D1F58B27926F3DDFCE8045E" author="Guinot" firstAuthor="Guinot" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">Guinot 1976</bibRefCitation>
: fig. 41 C); like other species in
<taxonomicName id="D89C5A771756C321FFE4D7145BEF1AD9" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="FA1DDEB5DA64482EDEB91791D7461C0C" italics="true">Actaeodes</emphasis>
</taxonomicName>
,
<taxonomicName id="77433A55849C6A601AE3184DB6A41791" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="A1AA1EF399A1FA334344D24B6DDDBBE1" italics="true">A. consobrinus</emphasis>
</taxonomicName>
has an elongated telson, but its overall shape is truncate-oval, with relatively arcuate lateral edges (Fig.
<figureCitation id="DB36083E42B201A07D98C1954670229C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 D</figureCitation>
) (vs. triangular telson with converging lateral edges in
<taxonomicName id="397B204A04F3BB7984C2EEB48F391B99" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="2D753DEB9D4575886878126431EED6A9" italics="true">Actaeodes</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="E2ACF8D3952C5F1661A68A95855FFFF1" author="Guinot" firstAuthor="Guinot" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">Guinot 1976</bibRefCitation>
: fig. 41 C);
<taxonomicName id="D70EDE61BDFFBB00209B1A420BFC608A" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="07288F0E36204770D906AF623B27FF63" italics="true">A. consobrinus</emphasis>
</taxonomicName>
has the first tooth flattened and the subsequent three teeth prominent (Figs
<figureCitation id="0AF599C5170F9628A91EFEEDB6985F27" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A, B</figureCitation>
,
<figureCitation id="8BE79437717010F14B250E60A8100E4B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 A</figureCitation>
,
<figureCitation id="61C20BBCA8633775E67F5687AE4DF50B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A, D, F</figureCitation>
) (vs. anterolateral margin divided into four distinct but not very prominent lobes in
<taxonomicName id="2C7D1C0D451781102C5B33D5BECEF13E" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="FB99926E38E2CE831935632C1ABC25BF" italics="true">Actaeodes</emphasis>
</taxonomicName>
; Fig.
<figureCitation id="5F89FA3E1ECF70DE8DCBB30B6EFCE9D5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza &amp; PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron &amp; Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 F H</figureCitation>
; cf.
<bibRefCitation id="4E03D91F4770A22BAC5F68FC1F900BC6" author="Guinot" firstAuthor="Guinot" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">Guinot 1976</bibRefCitation>
: pl. XV, figs 14); the cheliped carpus is more robust in
<taxonomicName id="2E4AD9AB946F132EC23BCC3F54DCFB6F" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="69EAE24C38670BAC8195F2CD6416ABAA" italics="true">A. consobrinus</emphasis>
</taxonomicName>
(Figs
<figureCitation id="9C1DAC5164135478F70B0CBD81CDB3EA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A</figureCitation>
,
<figureCitation id="F6A0051B11450D98EF5B9569270B01CE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A, D, F</figureCitation>
) (vs. proportionally more slender carpus in
<taxonomicName id="0D5860694E7EAB832D43526CBBF6E314" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="45557DA254BBEA2CFE398A3024CAEE33" italics="true">Actaeodes</emphasis>
</taxonomicName>
; Fig.
<figureCitation id="FC08599362C91639C41951449BBF0488" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza &amp; PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron &amp; Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 F H</figureCitation>
; cf.
<bibRefCitation id="666884AFEEB7D9C60958EFF68A2B13F0" author="Guinot" firstAuthor="Guinot" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum National dHistoire Naturelle, Paris: A, 97, 1308. [figs 147, pls 119]" year="1976">Guinot 1976</bibRefCitation>
: pl. XV, figs 14). It is worth mentioning that although body color is generally not used as a basis for defining genera within the family
<taxonomicName id="FD85129998EDB033431C17533D237BD0" authorityName="MacLeay" authorityYear="1838" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="family">Xanthidae</taxonomicName>
, the vibrant and high-contrast living coloration of
<taxonomicName id="065FF404E9FD49312A9FCB92C46AA6B5" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="515CBE5CE9B84D6A174ED96DA2B7B5EB" italics="true">A. consobrinus</emphasis>
</taxonomicName>
is also quite unique in
<taxonomicName id="A6F40AFC16329701652A7B53EB9E283A" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="D82A185D3A3DCD38E72A9EEB46D7A675" italics="true">Actaeodes</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="4CE06E0C42F00043CC6124456BE428EE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Gothus consobrinus (A. Milne-Edwards, 1873). A. Male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336; B. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014, displaying live coloration. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure10" httpUri="https://binary.pensoft.net/fig/1088103">10</figureCitation>
). Based on the aforementioned reasons,
<taxonomicName id="670476E9398D2DF9227C8488D33F20B0" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="65CF023CC0586353539F7C620850DC87" italics="true">A. consobrinus</emphasis>
</taxonomicName>
is not suitable for placement within the genus
<taxonomicName id="EC4264E5C2B63DC08B1B7C6CAADD877B" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="311BF093AEF2F3C64F1B6D42C190C5D4" italics="true">Actaeodes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F517E71AAA6DF5D62332D2E4B4B90EAD">
Another genus worth considering is
<taxonomicName id="E0062602360234EBFBA57EC1500230CF" authority="Serene, 1984" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Meractaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="311776790167AD74871A9F0499D0E54C" italics="true">Meractaea</emphasis>
<bibRefCitation id="D527890F340D1FE6BBCE04CADB9B9A03" author="Serène R" refId="B42" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1 - 349. [figs A - C + 1 - 243, pls 1 - 48]" year="1984">Serène, 1984</bibRefCitation>
</taxonomicName>
, characterized by almost straight posterolateral margins, developed areolation on the dorsal surface of the carapace, and four underdeveloped small teeth on the anterolateral margins, all of which are similar to the current species. However, there are also differences between this genus and
<taxonomicName id="28A9481A18588230A2D34FA52460F958" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="1950D1D22E53E17CC7159AAC55C0E9FE" italics="true">A. consobrinus</emphasis>
</taxonomicName>
, including an almost straight, quadrilobate frontal margin with a rounded central notch (cf.
<bibRefCitation id="639A4086BDAE2BA3F3677443D7DB5C24" author="Serene" firstAuthor="Serene" refId="B42" refString="Serène R (1984) Crustacés Décapodes Brachyoures de lOcéan Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1349. [figs A C + 1243, pls 148]" year="1984">Serène 1984</bibRefCitation>
: pl. XIX, fig. C) (vs. front not very prominent but not straight, divided by a V-shaped notch into two inclined rounded lobes in
<taxonomicName id="BA1C6472E0E8C8502A4D3220933F2E10" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="1193378197E6D3FB1160B5466646476D" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="DBE03F198FE3D9FA0B6B81B47941142E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A, B</figureCitation>
,
<figureCitation id="691B276B07509DE4382EAA331CBB9DA2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 A</figureCitation>
,
<figureCitation id="EFB10A72007CA3BC92426F8114848217" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A, D, F</figureCitation>
); markedly slender ambulatory legs (cf.
<bibRefCitation id="E77A1242752F62A8B79EACDE32E7466E" author="Serene" firstAuthor="Serene" refId="B42" refString="Serène R (1984) Crustacés Décapodes Brachyoures de lOcéan Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1349. [figs A C + 1243, pls 148]" year="1984">Serène 1984</bibRefCitation>
: pl. XIX, fig. C) (vs. flat and robust ambulatory legs in
<taxonomicName id="3EA5C6996CBA645A469CCB3952B04C02" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="99F7EF1B0F6053A04B9BFA0B64DFD094" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="1D16CC0735436F86893AB2B9EF4BAB5B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A</figureCitation>
,
<figureCitation id="E93BEBBB88C25576F61761877119B6E2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 D</figureCitation>
); a completely smooth dorsal surface of the carapace with irregularly sized granules, sometimes connected (cf.
<bibRefCitation id="1B9017B48B37BFA9B82630399D056A94" author="Serene" firstAuthor="Serene" refId="B42" refString="Serène R (1984) Crustacés Décapodes Brachyoures de lOcéan Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1349. [figs A C + 1243, pls 148]" year="1984">Serène 1984</bibRefCitation>
: pl. XIX, fig. C) (vs. carapace dorsal surface with setae, regularly sized granules, never connected in
<taxonomicName id="9FC179C1AA51FFB59325B175DAB9D80E" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="0D493C999E73466A3EDE0F2D4126DF5C" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="58FFAB49F7C878A2FF40E12DEA13BC1C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A, B</figureCitation>
,
<figureCitation id="0C3327F5CA25EFF5D1544068FD7E927A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 A</figureCitation>
,
<figureCitation id="F7B853B887C0585B751DCBB40850C74B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A, D, F</figureCitation>
);
<abbrev id="ABBRID0EH2DI" xlink_title="first gonopod of male">G 1</abbrev>
distal lobe slightly curved outward (cf.
<bibRefCitation id="E888811AA8C48D2DE5352AF8B7AA4EE2" author="Serene" firstAuthor="Serene" refId="B42" refString="Serène R (1984) Crustacés Décapodes Brachyoures de lOcéan Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1349. [figs A C + 1243, pls 148]" year="1984">Serène 1984</bibRefCitation>
: fig. 63) (vs.
<abbrev id="ABBRID0EP2DI" xlink_title="first gonopod of male">G 1</abbrev>
distal lobe curved inward in
<taxonomicName id="D4112C62B043F32067A9B79DE2BEE1A2" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="AD0D52CEE8BE4D9350047F551DC7AE89" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Fig.
<figureCitation id="C7C7649675CA0E9588BD046D29F0E08E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 F G</figureCitation>
). Considering these significant differences,
<taxonomicName id="9B2EA5BBD2DA370E83F7EA6FB7CA5A9B" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="061C1AD2C8E849406B1609EECFE91B9D" italics="true">A. consobrinus</emphasis>
</taxonomicName>
also cannot be placed within this genus.
</paragraph>
<paragraph id="8C34245B8FC497B8B92A64B927D29564">
Compared with the species of
<taxonomicName id="B2623035323BAB86075A2BBAAAA6F624" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="1D318B6EC731842FEDF28E1D6CC2998C" italics="true">Actaeodes</emphasis>
</taxonomicName>
and
<taxonomicName id="9205B003CCF9B4B280FBFD250E970A12" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Meractaea" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="FC708AE5335705658077364081E9C87B" italics="true">Meractaea</emphasis>
</taxonomicName>
,
<taxonomicName id="9ACF31028ADD4DEFA35720551674F403" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="DCCD8E97138F6AF6A71957D503E5B1AA" italics="true">A. consobrinus</emphasis>
</taxonomicName>
is actually more closely related to
<taxonomicName id="2C2142D533B819385D236C125E50ED50" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo">
<emphasis id="C21AE0338105C053F6AE5A569B97EF3F" italics="true">G. teemo</emphasis>
</taxonomicName>
. Beyond the most noticeable similarity in vibrant living coloration, both share similar carapace contours, flattened first anterolateral teeth, robust cheliped carpus, similar states of thoracic sternum, and special male abdominal morphology, particularly the truncate-oval telson (see the comparison in the remarks of
<taxonomicName id="8ACEE69668364056ECC5047A3CBD90FC" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo">
<emphasis id="3E7F60DBBF9493A4A482B35D160FEC6A" italics="true">G. teemo</emphasis>
</taxonomicName>
). We believe that placing this species into the current new genus and new combination is more appropriate.
</paragraph>
<paragraph id="8B75D216CF9EE5B0F1E6BE9A443F02EC">
Regarding the status of
<taxonomicName id="AA352CD1D125EE6A01651AC54895B298" authorityName="Garth &amp; Kim" authorityYear="1983" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="quinquelobatus">
<emphasis id="CA7CE2B6F41B7155E70F19D1FA8C9202" italics="true">A. quinquelobatus</emphasis>
</taxonomicName>
, in the absence of specimens, we hereby present some limited queries. Similar to the new combination
<taxonomicName id="93EEA009C8C79B35DEF42A5D99C361F1" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="5F703301C49391F7BA2D20E670F7AE88" italics="true">G. consobrinus</emphasis>
</taxonomicName>
, the morphology of
<taxonomicName id="63AA729742894C23967C2A519B821D3D" authorityName="Garth &amp; Kim" authorityYear="1983" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="quinquelobatus">
<emphasis id="5C651C42B3ED82526FFE166BB2F63212" italics="true">A. quinquelobatus</emphasis>
</taxonomicName>
also appears to deviate from the definition of
<taxonomicName id="E0871DBB91BD0E00E105BF063F94125C" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="86EB35F80E88AED09D9FA4E21753C164" italics="true">Actaeodes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DA22FC95BF419D48F85B2E3FB9F7DBF1" sensu="stricto">sensu stricto</taxonomicNameLabel>
, featuring 5 instead of 4 anterolateral teeth and possessing non-concave posterior margins (cf.
<bibRefCitation id="10D772FFB31EC68771F868F047042376" DOI="10.1080/00222938300770561" author="Garth and Kim" firstAuthor="Garth" issue="5" journalOrPublisher="Journal of Natural History" pagination="663-729" refId="B10" refString="Garth JS, Kim HS (1983) Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 19081909. Journal of Natural History 17 (5): 663729. https://doi.org/10.1080/00222938300770561" title="Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908-1909." volume="17" year="1983">Garth and Kim 1983</bibRefCitation>
: fig. 5 A). As
<bibRefCitation id="0026E90C2FCF91988279BC284AF67D7C" DOI="10.1080/00222938300770561" author="Garth JS &amp; Kim HS" firstAuthor="Garth" issue="5" journalOrPublisher="Journal of Natural History" pagination="663-729" refId="B10" refString="Garth JS, Kim HS (1983) Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 19081909. Journal of Natural History 17 (5): 663729. https://doi.org/10.1080/00222938300770561" title="Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908-1909." volume="17" year="1983">Garth and Kim (1983)</bibRefCitation>
noted,
<taxonomicName id="3D7BAAE3D773699F21B2743984F64A80" authorityName="Garth &amp; Kim" authorityYear="1983" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="quinquelobatus">
<emphasis id="7CF99750AB0CFD68A2C7F5DAAA81BE7B" italics="true">A. quinquelobatus</emphasis>
</taxonomicName>
has carapace partitioning similar to
<taxonomicName id="756058F2A06CD341A0531AE05C49C55F" authorityName="Guinot" authorityYear="1967" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="25E60C39D8C378E3724E735C07E27030" italics="true">G. consobrinus</emphasis>
</taxonomicName>
. However, current evidence does not affirm its placement within the genus
<taxonomicName id="A88FD4043448D5EF059F69F5B673C582" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="330B8296CD5B6764035A46A18B9CE951" italics="true">Gothus</emphasis>
</taxonomicName>
, given it has 5 anterolateral teeth (cf.
<bibRefCitation id="AAB22F74518AAB0D36DEE6C1B3FC645A" DOI="10.1080/00222938300770561" author="Garth and Kim" firstAuthor="Garth" issue="5" journalOrPublisher="Journal of Natural History" pagination="663-729" refId="B10" refString="Garth JS, Kim HS (1983) Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 19081909. Journal of Natural History 17 (5): 663729. https://doi.org/10.1080/00222938300770561" title="Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908-1909." volume="17" year="1983">Garth and Kim 1983</bibRefCitation>
: fig. 5 A) (vs. 3 or
<quantity id="3DCC4460C4EF4A737EE273115131954D" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="DA61445929F714FA08C36B694586D32B" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="723F613929645DF4B72F786301FB457C" italics="true">Gothus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="E9B8E7D490D0247672DA126FBCB14915" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 C</figureCitation>
,
<figureCitation id="231A6B8BD5E04E43B174EECA955B36E5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A</figureCitation>
), and the carapace and chelae exhibit a multitude of developed nodules (cf.
<bibRefCitation id="A3436237C4036ECF092FB01F9979D372" DOI="10.1080/00222938300770561" author="Garth and Kim" firstAuthor="Garth" issue="5" journalOrPublisher="Journal of Natural History" pagination="663-729" refId="B10" refString="Garth JS, Kim HS (1983) Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 19081909. Journal of Natural History 17 (5): 663729. https://doi.org/10.1080/00222938300770561" title="Crabs of the family Xanthidae (Crustacea, Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908-1909." volume="17" year="1983">Garth and Kim 1983</bibRefCitation>
: fig. 5 A, B) (vs. surfaces have granules but lack nodules in
<taxonomicName id="5908F8263962571A83E116CC124FB5F0" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="61A8AE8163AB344DFD8F5CA00B2A0720" italics="true">Gothus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="52D161F4E0B9D560C0381B5E92C48270" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
,
<figureCitation id="BC7BCCF984E5D24FC1D79B00EA9F53A5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A</figureCitation>
). Further examination is necessary to confirm its taxonomic status.
</paragraph>
</subSubSection>
</treatment>
</document>