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<document id="F1AD9330E4DFEC18E499BB6B047E7E61" ID-CLB-Dataset="55637" ID-DOI="10.5281/zenodo.5312184" ID-GBIF-Dataset="5ff783bc-d046-4426-8e03-e131d0e4dabe" ID-ISSN="0253-116X" ID-Zenodo-Dep="5312184" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630118963700" checkinUser="marcus" docAuthor="De, G." docDate="2014" docId="5C3487D2FFBDFFB341B3FCB2BA56499D" docLanguage="en" docName="LinzerbiolBeitr.2014.46.2.1737-1765.pdf" docOrigin="Linzer biologische Beiträge 46 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Naddia asetosa De 2014, nov.sp." docType="treatment" docVersion="9" lastPageNumber="1743" masterDocId="A00DFFAAFFB8FFB54101FFD9BB074A2F" masterDocTitle="The genus Naddia in Borneo (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="1765" masterPageNumber="1737" pageNumber="1742" updateTime="1699158966059" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9313FD3DCF3BE377D762CE862869A7D9">The genus Naddia in Borneo (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="CDCF5B652F5D4B9E6C141ABD8E69FC51" type="personal">
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<mods:namePart id="3FF539B72100F78C6D916427FBC64020">De, G.</mods:namePart>
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<mods:title id="80D93B078FBCC067C2B031489EBC8AAC">Linzer biologische Beiträge</mods:title>
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<mods:part id="0D944F6BF605DD8262C94C5B967863D1">
<mods:date id="4CB81B0C18959183C0F067A659F9A510">2014</mods:date>
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<mods:number id="D31FDC084C521FB3655ECE0EC3953172">2014-12-19</mods:number>
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<mods:number id="CFEBE39DC59F11F84F7C2BE23EC8F28A">46</mods:number>
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<treatment id="5C3487D2FFBDFFB341B3FCB2BA56499D" ID-DOI="http://doi.org/10.5281/zenodo.6532574" ID-GBIF-Taxon="195277723" ID-Zenodo-Dep="6532574" LSID="urn:lsid:plazi:treatment:5C3487D2FFBDFFB341B3FCB2BA56499D" httpUri="http://treatment.plazi.org/id/5C3487D2FFBDFFB341B3FCB2BA56499D" lastPageId="6" lastPageNumber="1743" pageId="5" pageNumber="1742">
<subSubSection id="9C87654FFFBDFFB041B3FCB2BAA449AF" box="[178,419,875,897]" pageId="5" pageNumber="1742" type="nomenclature">
<paragraph id="D42236C4FFBDFFB041B3FCB2BAA449AF" blockId="5.[178,503,875,897]" box="[178,419,875,897]" pageId="5" pageNumber="1742">
<heading id="8F6A81A8FFBDFFB041B3FCB2BAA449AF" box="[178,419,875,897]" fontSize="8" level="2" pageId="5" pageNumber="1742" reason="2">
<emphasis id="E6E9EAD6FFBDFFB041B3FCB2BAA449AF" bold="true" box="[178,419,875,897]" pageId="5" pageNumber="1742">
<taxonomicName id="139D4D47FFBDFFB041B3FCB2BA5749AF" ID-CoL="45JVQ" authority="De, 2014" authorityName="De" authorityYear="2014" box="[178,336,875,897]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1742" phylum="Arthropoda" rank="species" species="asetosa" status="sp. nov.">
<emphasis id="E6E9EAD6FFBDFFB041B3FCB2BA5749AF" bold="true" box="[178,336,875,897]" italics="true" pageId="5" pageNumber="1742">Naddia asetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDDA57ADFFBDFFB04057FCB2BAA449AF" box="[342,419,875,896]" pageId="5" pageNumber="1742" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9C87654FFFBDFFB040A8FCB2BAF049AE" box="[425,503,875,897]" pageId="5" pageNumber="1742" type="description">
<paragraph id="D42236C4FFBDFFB040A8FCB2BAF049AE" blockId="5.[178,503,875,897]" box="[425,503,875,897]" pageId="5" pageNumber="1742">
<heading id="8F6A81A8FFBDFFB040A8FCB2BAF049AE" box="[425,503,875,897]" fontSize="8" level="2" pageId="5" pageNumber="1742" reason="2">
<emphasis id="E6E9EAD6FFBDFFB040A8FCB2BAF049AE" bold="true" box="[425,503,875,897]" pageId="5" pageNumber="1742">(Fig. 1)</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9C87654FFFBDFFB041CAFC40BA2949F1" pageId="5" pageNumber="1742" type="materials_examined">
<paragraph id="D42236C4FFBDFFB041CAFC40BA2949F1" blockId="5.[203,1086,920,990]" pageId="5" pageNumber="1742">
<materialsCitation id="64F53C99FFBDFFB041CAFC40BA2949F1" ID-GBIF-Occurrence="3763851345" collectingDate="2001-08" collectionCode="NHML" country="Indonesia" location="Rekut" municipality="Busang" pageId="5" pageNumber="1742" specimenCode="BMNH(E)-2001-191" specimenCount="1" stateProvince="Borneo" typeStatus="holotype">
<typeStatus id="0B268866FFBDFFB041CAFC40BA184980" box="[203,287,921,943]" pageId="5" pageNumber="1742" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="AC8A7654FFBDFFB04029FC40BAA54981" box="[296,418,921,942]" name="Indonesia" pageId="5" pageNumber="1742">INDONESIA</collectingCountry>
,
<collectingCounty id="3D434E48FFBDFFB040ADFC40BAE94981" box="[428,494,921,942]" pageId="5" pageNumber="1742">Borneo</collectingCounty>
,
<collectingRegion id="1659F826FFBDFFB040F9FC40B9AF4980" box="[504,680,921,943]" country="Indonesia" name="Kalimantan Tengah" pageId="5" pageNumber="1742">Kalimantan Tengah</collectingRegion>
,
<collectingMunicipality id="3446ACBEFFBDFFB043B0FC40B9F64980" box="[689,753,921,943]" pageId="5" pageNumber="1742">Busang</collectingMunicipality>
/
<location id="D142601FFFBDFFB043F6FC40B82B4980" LSID="urn:lsid:plazi:treatment:5C3487D2FFBDFFB341B3FCB2BA56499D:D142601FFFBDFFB043F6FC40B82B4980" box="[759,812,921,943]" country="Indonesia" county="Borneo" municipality="Busang" name="Rekut" pageId="5" pageNumber="1742" stateProvince="Kalimantan Tengah">Rekut</location>
confl. 0˚03S 113˚59E,
<location id="D142601FFFBDFFB0450BFC40BF394980" LSID="urn:lsid:plazi:treatment:5C3487D2FFBDFFB341B3FCB2BA56499D:D142601FFFBDFFB0450BFC40BF394980" box="[1034,1086,921,943]" country="Indonesia" county="Borneo" municipality="Busang" name="Flight" pageId="5" pageNumber="1742" stateProvince="Kalimantan Tengah">Flight</location>
<collectingMethod id="0DDC4ED3FFBDFFB041DEFC68BA2B49E9" box="[223,300,945,966]" pageId="5" pageNumber="1742">Intercept</collectingMethod>
FIT 6,
<location id="D142601FFFBDFFB0407FFC68BACE49E8" LSID="urn:lsid:plazi:treatment:5C3487D2FFBDFFB341B3FCB2BA56499D:D142601FFFBDFFB0407FFC68BACE49E8" box="[382,457,945,967]" country="Indonesia" county="Borneo" municipality="Busang" name="Brendell" pageId="5" pageNumber="1742" stateProvince="Kalimantan Tengah">Brendell</location>
/
<location id="D142601FFFBDFFB040D1FC68B91449E8" LSID="urn:lsid:plazi:treatment:5C3487D2FFBDFFB341B3FCB2BA56499D:D142601FFFBDFFB040D1FC68B91449E8" box="[464,531,945,967]" country="Indonesia" county="Borneo" municipality="Busang" name="Mendel" pageId="5" pageNumber="1742" stateProvince="Kalimantan Tengah">Mendel</location>
,
<date id="A0231004FFBDFFB04320FC68B99E49E9" box="[545,665,945,966]" pageId="5" pageNumber="1742" value="2001-08">
<collectingDate id="B067E9ECFFBDFFB04320FC68B99E49E9" box="[545,665,945,966]" pageId="5" pageNumber="1742" value="2001-08">August 2001</collectingDate>
</date>
,
<location id="D142601FFFBDFFB043ABFC68B81749E8" LSID="urn:lsid:plazi:treatment:5C3487D2FFBDFFB341B3FCB2BA56499D:D142601FFFBDFFB043ABFC68B81749E8" box="[682,784,945,967]" country="Indonesia" county="Borneo" municipality="Busang" name="Barito Ulu" pageId="5" pageNumber="1742" stateProvince="Kalimantan Tengah">Barito Ulu</location>
2001, BMNH(E) 2001-191 (in
<collectionCode id="B28CAE01FFBDFFB041DEFC10BA2349F1" box="[223,292,969,990]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="5" pageNumber="1742">NHML</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9C87654FFFBDFFB041B3FC34B8464C14" pageId="5" pageNumber="1742" type="diagnosis">
<paragraph id="D42236C4FFBDFFB041B3FC34B8AC4E0F" blockId="5.[178,1085,1005,1056]" pageId="5" pageNumber="1742">
This species has the deep occipital emargination and salient postero-lateral lobes of the head characteristic of
<taxonomicName id="139D4D47FFBDFFB04090FBD3BADC4E30" box="[401,475,1034,1055]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1742" phylum="Arthropoda" rank="genus">
<emphasis id="E6E9EAD6FFBDFFB04090FBD3BADC4E30" box="[401,475,1034,1055]" italics="true" pageId="5" pageNumber="1742">Naddia</emphasis>
</taxonomicName>
, but differs notably in the following respects:
</paragraph>
<paragraph id="D42236C4FFBDFFB041E3FBF4BADE4EB6" blockId="5.[226,1086,1069,1419]" pageId="5" pageNumber="1742">
 the sides of the head, instead of being parallel or expanded posteriad as in
<taxonomicName id="139D4D47FFBDFFB04013FB92BA5B4E4F" box="[274,348,1099,1120]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1742" phylum="Arthropoda" rank="genus">
<emphasis id="E6E9EAD6FFBDFFB04013FB92BA5B4E4F" box="[274,348,1099,1120]" italics="true" pageId="5" pageNumber="1742">Naddia</emphasis>
</taxonomicName>
, are retracted between the posterior margins of eyes and the posterolateral angles, giving the insect a vague resemblance to some members of the
<emphasis id="E6E9EAD6FFBDFFB04013FB5ABA854EB7" box="[274,386,1155,1176]" italics="true" pageId="5" pageNumber="1742">Eucibdelus</emphasis>
lineage;
</paragraph>
<paragraph id="D42236C4FFBDFFB041E3FB7CB88E4EDB" blockId="5.[226,1086,1069,1419]" pageId="5" pageNumber="1742">
 the antennae are long, with all segments symmetrical, and except for the second, at least as long as broad, the terminal antennomeres not transverse, asymmetrical, flattened or sub-serrate as in most other
<taxonomicName id="139D4D47FFBDFFB04236FB06B8864EDB" box="[823,897,1247,1268]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1742" phylum="Arthropoda" rank="genus">
<emphasis id="E6E9EAD6FFBDFFB04236FB06B8864EDB" box="[823,897,1247,1268]" italics="true" pageId="5" pageNumber="1742">Naddia</emphasis>
</taxonomicName>
;
</paragraph>
<paragraph id="D42236C4FFBDFFB041E3FB27BF104F39" blockId="5.[226,1086,1069,1419]" box="[226,1047,1278,1302]" pageId="5" pageNumber="1742">
 the puncturation of head and pronotum is finer than in other
<taxonomicName id="139D4D47FFBDFFB04277FAD9BF144F39" box="[886,1043,1280,1302]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1742" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E6E9EAD6FFBDFFB04277FAD9B8C74F3A" box="[886,960,1280,1301]" italics="true" pageId="5" pageNumber="1742">Naddia</emphasis>
species
</taxonomicName>
;
</paragraph>
<paragraph id="D42236C4FFBDFFB041E3FAC6BA8C4FA4" blockId="5.[226,1086,1069,1419]" pageId="5" pageNumber="1742">
 the sides of head, elytra and pleurites ot the genital segment are totally devoid of long setae, which are always present in other
<taxonomicName id="139D4D47FFBDFFB04232FAE4B87A4F7D" box="[819,893,1341,1362]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1742" phylum="Arthropoda" rank="genus">
<emphasis id="E6E9EAD6FFBDFFB04232FAE4B87A4F7D" box="[819,893,1341,1362]" italics="true" pageId="5" pageNumber="1742">Naddia</emphasis>
</taxonomicName>
, (among the fine puncturation there are no larger punctures present from which setae might have broken off).
</paragraph>
<paragraph id="D42236C4FFBDFFB041B3FA40B8464C14" blockId="5.[178,1085,1433,1595]" pageId="5" pageNumber="1742">
These differences suggest that a new genus should be described to accommodate this species; however given the present uncertain knowledge of phylogenetic lineages in the
<taxonomicName id="139D4D47FFBDFFB041B3FA08BA374FC8" authorityName="Latreille" authorityYear="1802" box="[178,304,1489,1511]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1742" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
, and the absence of males, it seems more prudent to describe it, at least provisionally, as belonging to
<taxonomicName id="139D4D47FFBDFFB040F1FA34B93D4C2D" box="[496,570,1517,1538]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1742" phylum="Arthropoda" rank="genus">
<emphasis id="E6E9EAD6FFBDFFB040F1FA34B93D4C2D" box="[496,570,1517,1538]" italics="true" pageId="5" pageNumber="1742">Naddia</emphasis>
</taxonomicName>
. As the specimen is glued to the card with a cement that does not seem to be soluble in water or ethyl-acetate, I am unable to examine the ventral structures for fear of damaging this unique specimen.
</paragraph>
</subSubSection>
<subSubSection id="9C87654FFFBEFFB341B3FF02BA56499D" pageId="6" pageNumber="1743" type="description">
<paragraph id="D42236C4FFBEFFB341B3FF02BAB94B4A" blockId="6.[178,1085,219,358]" pageId="6" pageNumber="1743">
Description: Proportions of
<typeStatus id="0B268866FFBEFFB3432DFF02B98E4ADE" box="[556,649,219,241]" pageId="6" pageNumber="1743" type="holotype">holotype</typeStatus>
: length:
<quantity id="13659B21FFBEFFB343E2FF02B8284ADF" box="[739,815,219,240]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="1743" unit="mm" value="13.0">13 mm</quantity>
; length of head: 1.7; total length of head: 2.0; maximum breadth of head: 2.0; breadth of head in front of posterolateral angles: 1.8; length of eye: 0.8; length of temples: 1.00; length of antenna: 3.5; length of pronotum: 2.2; breadth of pronotum: 2.2; length of elytron: 2.9; breadth of elytra: 2.8. Habitus: fig. 1.
</paragraph>
<paragraph id="D42236C4FFBEFFB341B3FEACBF184832" blockId="6.[178,1085,373,541]" pageId="6" pageNumber="1743">
Temples retracted towards postero-lateral angles. Labrum divided into two separate lobes, each lobe bearing six terminal black setae; both mandibles with a broad and long single molar, the right one slightly but distinctly bicuspid; palpi as in most
<taxonomicName id="139D4D47FFBEFFB342B9FE77BF3E4BEA" box="[952,1081,430,454]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1743" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E6E9EAD6FFBEFFB342B9FE77BF004BE9" box="[952,1031,430,454]" italics="true" pageId="6" pageNumber="1743">Naddia</emphasis>
spp.
</taxonomicName>
, the terminal segments shorter than in
<taxonomicName id="139D4D47FFBEFFB3433EFE12B9C74BCC" box="[575,704,459,483]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1743" phylum="Arthropoda" rank="genus">
<emphasis id="E6E9EAD6FFBEFFB3433EFE12B9C74BCC" box="[575,704,459,483]" italics="true" pageId="6" pageNumber="1743">Platydracus</emphasis>
</taxonomicName>
and more inflated. Antennae almost filiform, all antennomeres elongate, antennomeres 7-10 less so, almost quadrate, all antennomeres symmetrical. Protarsi dilated, though much less so than in other species.
</paragraph>
<paragraph id="D42236C4FFBEFFB341B3FDF4BA9D48DD" blockId="6.[178,1085,557,754]" pageId="6" pageNumber="1743">Integument of entire body black except posterior 2/3rd of elytra brassy metallic. Legs black. Head with a few short brassy setae, more conspicuous on temples; sides of pronotum with some sparse pale brassy pubescence; pubescence of scutellum black; elytra entirely covered with moderately dense long golden pubescence, except narrowly along anterior margin; abdominal tergites V-VII with a pair of narrow antero-lateral fasciae of short silvery pubescence; on tergite VII the two fasciae meeting in the centre to form a single transverse band.</paragraph>
<paragraph id="D42236C4FFBEFFB341B3FCDBB88149A2" blockId="6.[178,1085,770,909]" pageId="6" pageNumber="1743">Puncturation of head close, sub-rugose, composed of umbilicate punctures on posterior 1/3rd, the interstices coalescing to form long longitudinal rugae in anterior 2/3rd; puncturation of pronotum similar to that of posterior 1/3rd of head; the mid-longitudinal band interrupted near anterior and posterior margins and in the middle, leaving only two narrow elongate shiny calluses; puncturation of elytra very fine and dense.</paragraph>
<paragraph id="D42236C4FFBEFFB341B3FC45BA56499D" blockId="6.[178,337,924,946]" box="[178,337,924,946]" pageId="6" pageNumber="1743">Male: unknown</paragraph>
</subSubSection>
</treatment>
</document>