278 lines
34 KiB
XML
278 lines
34 KiB
XML
<document id="3BB903DFBFFB946B850889CE62FDC659" ID-DOI="10.5852/ejt.2020.626" ID-GBIF-Dataset="c906d9cd-cfcb-43db-a24c-b57b39d9184b" ID-Zenodo-Dep="3754989" ID-ZooBank="D91881D6-55D8-48FC-A383-069BC643A91E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587066773978" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2020" docId="5C11879EFFBBFFD5FD86E9D7FC7F1642" docLanguage="en" docName="ejt-626_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 626" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor longipala Enghoff, in Enghoff & Frederiksen 2015" docType="treatment" docVersion="6" lastPageNumber="16" masterDocId="A028FFE6FFB6FFDAFFC1ED40FFC7143C" masterDocTitle="A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="14" updateTime="1698738618004" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="77E6442CF340C5DCFF727E2353A5CB49">A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="B76475F3EA185B85387CDAC68EF759DD">Enghoff, Henrik</mods:namePart>
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<mods:title id="F06AFE916E16FE9D189D85753F06C4A9">European Journal of Taxonomy</mods:title>
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<mods:part id="1F2CC539AAF70459B86053F9E1C19522">
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<mods:date id="9B1DCC64705CE4F85116FD0E0783F4F2">2020</mods:date>
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<mods:number id="DE1555AA58F55A38704F037811A672E8">2020-04-14</mods:number>
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<mods:number id="B7CD96E59033830AC5DD44BED2E765FF">626</mods:number>
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<treatment id="5C11879EFFBBFFD5FD86E9D7FC7F1642" ID-DOI="http://doi.org/10.5281/zenodo.3808352" ID-GBIF-Taxon="163642161" ID-Zenodo-Dep="3808352" LSID="urn:lsid:plazi:treatment:5C11879EFFBBFFD5FD86E9D7FC7F1642" httpUri="http://treatment.plazi.org/id/5C11879EFFBBFFD5FD86E9D7FC7F1642" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
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<subSubSection id="9CA26503FFBBFFD7FD86E9D7FC2A108E" box="[583,1005,1175,1202]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="D4073688FFBBFFD7FD86E9D7FC2A108E" blockId="13.[583,1005,1175,1237]" box="[583,1005,1175,1202]" pageId="13" pageNumber="14">
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<heading id="8F4F81E4FFBBFFD7FD86E9D7FC2A108E" box="[583,1005,1175,1202]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
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<taxonomicName id="13B84D0BFFBBFFD7FD86E9D7FC2A108E" ID-CoL="FZL2" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[583,1005,1175,1202]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipala">
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<emphasis id="E6CCEA9AFFBBFFD7FD86E9D7FCFB108E" bold="true" box="[583,828,1175,1202]" italics="true" pageId="13" pageNumber="14">Aquattuor longipala</emphasis>
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<bibRefCitation id="B0294B79FFBBFFD7FC85E9D7FC2A108E" author="Enghoff H. & Frederiksen S. B." box="[836,1005,1175,1202]" pageId="13" pageNumber="14" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9CA26503FFBBFFD7FD50E9FAFC6510E8" box="[657,930,1210,1237]" pageId="13" pageNumber="14" type="description">
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<paragraph id="D4073688FFBBFFD7FD50E9FAFC6510E8" blockId="13.[583,1005,1175,1237]" box="[657,930,1210,1237]" pageId="13" pageNumber="14">
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<figureCitation id="4C832A0DFFBBFFD7FD50E9FAFD1D10E8" box="[657,730,1210,1236]" captionStart="Fig" captionStartId="2.[189,232,1948,1974]" captionTargetBox="[192,1385,1023,1872]" captionTargetId="figure@2.[186,1391,1009,1876]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites for Aquattuor spp. in the Udzungwa Mts. Based on Marshall et al. (2010: fig. 1); inset by permission of the Eastern Arc Mountains Conservation Endowment Fund." figureDoi="http://doi.org/10.5281/zenodo.3754991" httpUri="https://zenodo.org/record/3754991/files/figure.png" pageId="13" pageNumber="14">Figs 1</figureCitation>
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,
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<figureCitation id="4C832A0DFFBBFFD7FD29E9FAFD3210E9" box="[744,757,1210,1237]" captionStart="Fig" captionStartId="7.[189,232,1887,1913]" captionTargetBox="[234,1343,270,1842]" captionTargetId="figure@7.[229,1354,262,1848]" captionTargetPageId="7" captionText="Fig. 5. Aquattuor spp., first pair of male legs. A–C. A. major Enghoff, 2015, paratype, ♂ (NHMD 621644), first pair of legs in situ. D–F. A. longipala Enghoff, 2015, male from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642). A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754999" httpUri="https://zenodo.org/record/3754999/files/figure.png" pageId="13" pageNumber="14">5</figureCitation>
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D–F, 8B, 9, 11
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</paragraph>
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</subSubSection>
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<subSubSection id="9CA26503FFBBFFD7FF7CE9BCFC0B112A" box="[189,972,1275,1302]" pageId="13" pageNumber="14" type="reference_group">
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<paragraph id="D4073688FFBBFFD7FF7CE9BCFC0B112A" blockId="13.[189,972,1275,1302]" box="[189,972,1275,1302]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="F4A811A6FFBBFFD7FF7CE9BCFC0B112A" box="[189,972,1275,1302]" pageId="13" pageNumber="14">
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<treatmentCitation id="55191099FFBBFFD7FF7CE9BCFC0F112A" ID-DOI="http://doi.org/10.5281/zenodo.3793781" ID-Zenodo-Dep="3793781" author="Enghoff H. & Frederiksen S. B." box="[189,968,1275,1302]" httpUri="http://treatment.plazi.org/id/03A487F2FFF2FFB27564682A977CB6E3" page="12" pageId="13" pageNumber="14" year="2015">
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<taxonomicName id="13B84D0BFFBBFFD7FF7CE9BCFC0F112A" ID-CoL="FZL2" authority="Enghoff, in Enghoff & Frederiksen, 2015: 12" authorityName="Enghoff, in Enghoff & Frederiksen" authorityPageNumber="12" authorityYear="2015" box="[189,968,1275,1302]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipala">
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<emphasis id="E6CCEA9AFFBBFFD7FF7CE9BCFE6C112A" box="[189,427,1276,1302]" italics="true" pageId="13" pageNumber="14">Aquattuor longipala</emphasis>
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Enghoff,
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<emphasis id="E6CCEA9AFFBBFFD7FDE0E9BCFDFF1129" box="[545,568,1276,1301]" italics="true" pageId="13" pageNumber="14">in</emphasis>
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<bibRefCitation id="B0294B79FFBBFFD7FDFEE9BBFC0F112A" author="Enghoff H. & Frederiksen S. B." box="[575,968,1275,1302]" pageId="13" pageNumber="14" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff & Frederiksen, 2015: 12</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="9CA26503FFBBFFD5FF7CE87DFC7115D2" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="D4073688FFBBFFD7FF7CE87DFEF1116B" blockId="13.[189,310,1341,1367]" box="[189,310,1341,1367]" pageId="13" pageNumber="14">
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<heading id="8F4F81E4FFBBFFD7FF7CE87DFEF1116B" bold="true" box="[189,310,1341,1367]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
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<emphasis id="E6CCEA9AFFBBFFD7FF7CE87DFEF1116B" bold="true" box="[189,310,1341,1367]" pageId="13" pageNumber="14">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D4073688FFBBFFD7FF7CE82BFCF31209" blockId="13.[189,1399,1387,1590]" pageId="13" pageNumber="14">
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Diameter
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<quantity id="13409B6DFFBBFFD7FEF4E82CFE2011BA" box="[309,487,1388,1414]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.02" metricValueMin="1.78" pageId="13" pageNumber="14" unit="mm" value="1.9" valueMax="2.02" valueMin="1.78">1.78–2.02 mm</quantity>
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, 50–52 podous rings (
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<figureCitation id="4C832A0DFFBBFFD7FD38E82BFC8211BA" box="[761,837,1387,1414]" captionStart="Fig" captionStartId="11.[187,230,1893,1919]" captionTargetBox="[193,1391,1095,1856]" captionTargetId="figure@11.[192,1402,1089,1860]" captionTargetPageId="11" captionText="Fig. 9. Aquattuor spp. Body size (number of podous rings and midbody vertical diameter) in males of Aquattuor species from the Udzungwa Mts, and of A. claudiahempae Enghoff & Frederiksen, 2015 from Mt Kilimanjaro. In case of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3755007" httpUri="https://zenodo.org/record/3755007/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
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). Differs from congeners, except
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<taxonomicName id="13B84D0BFFBBFFD7FB23E82CFAB111BA" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[1250,1398,1388,1414]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipala">
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<emphasis id="E6CCEA9AFFBBFFD7FB23E82CFAB111BA" box="[1250,1398,1388,1414]" italics="true" pageId="13" pageNumber="14">A. longipala</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="13B84D0BFFBBFFD7FF31E8CFFE9F1195" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[240,344,1423,1449]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="major">
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<emphasis id="E6CCEA9AFFBBFFD7FF31E8CFFE9F1195" box="[240,344,1423,1449]" italics="true" pageId="13" pageNumber="14">A. major</emphasis>
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</taxonomicName>
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, by having the gonopodal palette considerably longer than broad, and by having a distal prefemoral lobe on the first pair of male legs, in addition to the normal prefemoral process. Differs from
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<taxonomicName id="13B84D0BFFBBFFD7FF7CE896FE9E11D3" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[189,345,1493,1520]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
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<emphasis id="E6CCEA9AFFBBFFD7FF7CE896FE9E11D3" box="[189,345,1493,1520]" italics="true" pageId="13" pageNumber="14">A. mollilobus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FDFF57E1FFBBFFD7FEA1E896FE7D11CC" box="[352,442,1494,1520]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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by having distinct lateral and mesal incisions on the gonopod coxa and by having the tarsal setae of the first pair of male legs of normal length; differs from
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<taxonomicName id="13B84D0BFFBBFFD7FBCCE8B9FBB4122E" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[1037,1139,1529,1555]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="major">
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<emphasis id="E6CCEA9AFFBBFFD7FBCCE8B9FBB4122E" box="[1037,1139,1529,1555]" italics="true" pageId="13" pageNumber="14">A. major</emphasis>
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</taxonomicName>
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in lacking a triangular tooth on the anterior margin of the gonopod telomere.
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</paragraph>
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<paragraph id="D4073688FFBBFFD7FF7CEB1DFD131244" blockId="13.[189,724,1629,1656]" box="[189,724,1629,1656]" pageId="13" pageNumber="14">
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||
<emphasis id="E6CCEA9AFFBBFFD7FF7CEB1DFE4B124B" bold="true" box="[189,396,1629,1655]" pageId="13" pageNumber="14">Material studied</emphasis>
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(total: 3 ♂♂,
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<specimenCount id="C2BEFD01FFBBFFD7FDF3EB1DFDB21244" box="[562,629,1629,1656]" pageId="13" pageNumber="14" type="female">3 ♀♀</specimenCount>
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, 3 juvs)
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</paragraph>
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<paragraph id="D4073688FFBBFFD7FF7CEBCCFD3F130E" blockId="13.[189,1399,1675,1843]" pageId="13" pageNumber="14">
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||
<materialsCitation id="64D03CD5FFBBFFD7FF7CEBCCFCB212D0" ID-GBIF-Occurrence="2598688700" collectingDate="2014-10-27" collectingMethod="pitfall trapping (summed catch from two traps" collectionCode="NHMD" collectorName="J. Malumbres-Olarte" country="Tanzania" county="Udzungwa Mts National Park" elevation="1552" latitude="-7.827722" location="Mito Mitatu" longLatPrecision="1" longitude="36.840553" municipality="Mito Mitatu" pageId="13" pageNumber="14" specimenCode="NHMD 621642" specimenCount="8" specimenCount-female="3" stateProvince="Morogoro Region">
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||
<collectingCountry id="ACAF7618FFBBFFD7FF7CEBCCFE96129A" box="[189,337,1676,1702]" name="Tanzania" pageId="13" pageNumber="14">TANZANIA</collectingCountry>
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||
•
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||
<specimenCount id="C2BEFD01FFBBFFD7FEB1EBCCFE70129A" box="[368,439,1676,1702]" pageId="13" pageNumber="14">2 ♂♂</specimenCount>
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||
,
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||
<specimenCount id="C2BEFD01FFBBFFD7FE06EBCCFDC9129A" box="[455,526,1676,1702]" pageId="13" pageNumber="14" type="female">3 ♀♀</specimenCount>
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||
,
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||
<specimenCount id="C2BEFD01FFBBFFD7FDDEEBCCFDAA1299" box="[543,621,1675,1702]" pageId="13" pageNumber="14">3 juvs</specimenCount>
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||
;
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||
<collectingRegion id="167CF86AFFBBFFD7FDBDEBCCFC921299" box="[636,853,1675,1702]" country="Tanzania" name="Morogoro" pageId="13" pageNumber="14">Morogoro Region</collectingRegion>
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||
,
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||
<collectingCounty id="3D664E04FFBBFFD7FCA5EBCBFB091299" box="[868,1230,1675,1702]" pageId="13" pageNumber="14">Udzungwa Mts National Park</collectingCounty>
|
||
,
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||
<collectingMunicipality id="3463ACF2FFBBFFD7FB1CEBCBFAB51299" box="[1245,1394,1675,1701]" pageId="13" pageNumber="14">Mito Mitatu</collectingMunicipality>
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, Plot 15;
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||
<geoCoordinate id="B18C504FFFBBFFD7FEE4EBEFFE0912F5" box="[293,462,1709,1737]" degrees="07" direction="south" minutes="49" orientation="latitude" pageId="13" pageNumber="14" precision="1" seconds="39.8" value="-7.827722">07°49′39.8″ S</geoCoordinate>
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||
,
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||
<geoCoordinate id="B18C504FFFBBFFD7FE1CEBEFFD4F12F5" box="[477,648,1709,1738]" degrees="36" direction="east" minutes="50" orientation="longitude" pageId="13" pageNumber="14" precision="1" seconds="26.0" value="36.840553">36°50′26.0″ E</geoCoordinate>
|
||
;
|
||
<elevation id="5F95D1BBFFBBFFD7FD56EBEFFCF212F5" box="[663,821,1711,1738]" metricMagnitude="3" metricUnit="m" metricValue="1.552" pageId="13" pageNumber="14" unit="m" value="1552.0">
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<quantity id="13409B6DFFBBFFD7FD56EBEFFD3412F5" box="[663,755,1711,1738]" metricMagnitude="3" metricUnit="m" metricValue="1.552" pageId="13" pageNumber="14" unit="m" value="1552.0">1552 m</quantity>
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||
a.s.l.
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||
</elevation>
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||
;
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||
<date id="A0061048FFBBFFD7FC84EBEFFC2112F5" box="[837,998,1711,1737]" pageId="13" pageNumber="14" value="2014-10-27">
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<collectingDate id="B042E9A0FFBBFFD7FC84EBEFFC2112F5" box="[837,998,1711,1737]" pageId="13" pageNumber="14" value="2014-10-27">27 Oct. 2014</collectingDate>
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</date>
|
||
;
|
||
<collectorName id="794D535EFFBBFFD7FC35EBEFFB2D12F5" box="[1012,1258,1711,1737]" pageId="13" pageNumber="14">J. Malumbres-Olarte</collectorName>
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||
leg.;
|
||
<collectingMethod id="0DF94E9FFFBBFFD7FAF1EBEFFD6512D0" pageId="13" pageNumber="14">pitfall trapping (summed catch from two traps</collectingMethod>
|
||
);
|
||
<specimenCode id="841E9EF3FFBBFFD7FD78EB92FCB212D0" box="[697,885,1746,1772]" collectionCode="NHMD" pageId="13" pageNumber="14">NHMD 621642</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="64D03CD5FFBBFFD7FC54EB92FD33130E" ID-GBIF-Occurrence="2598688707" collectingDate="2013-11-25" collectingMethod="pitfall trapping" collectionCode="NHMD" collectorName="T. Pape & N. Scharff" country="Tanzania" elevation="1207" latitude="-7.8373055" location="Mito Mitatu" longLatPrecision="1" longitude="36.846333" municipality="Udzungwa Mts National Park" pageId="13" pageNumber="14" specimenCode="NHMD 621643" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region">
|
||
<specimenCount id="C2BEFD01FFBBFFD7FC54EB92FC0112D0" box="[917,966,1746,1772]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="167CF86AFFBBFFD7FC16EB92FB7712D0" box="[983,1200,1746,1772]" country="Tanzania" name="Morogoro" pageId="13" pageNumber="14">Morogoro Region</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="3463ACF2FFBBFFD7FB7EEB92FEA21333" pageId="13" pageNumber="14">Udzungwa Mts National Park</collectingMunicipality>
|
||
,
|
||
<location id="D1676053FFBBFFD7FEB3EBB5FDC01333" LSID="urn:lsid:plazi:treatment:5C11879EFFBBFFD5FD86E9D7FC7F1642:D1676053FFBBFFD7FEB3EBB5FDC01333" box="[370,519,1781,1807]" country="Tanzania" latitude="-7.8373055" longLatPrecision="1" longitude="36.846333" municipality="Udzungwa Mts National Park" name="Mito Mitatu" pageId="13" pageNumber="14" stateProvince="Morogoro Region">Mito Mitatu</location>
|
||
;
|
||
<geoCoordinate id="B18C504FFFBBFFD7FDD4EBB5FD7A1333" box="[533,701,1779,1808]" degrees="07" direction="south" minutes="50" orientation="latitude" pageId="13" pageNumber="14" precision="1" seconds="14.3" value="-7.8373055">07°50′14.3″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B18C504FFFBBFFD7FD0CEBB5FCBF132C" box="[717,888,1779,1808]" degrees="36" direction="east" minutes="50" orientation="longitude" pageId="13" pageNumber="14" precision="1" seconds="46.8" value="36.846333">36°50′46.8″ E</geoCoordinate>
|
||
;
|
||
<elevation id="5F95D1BBFFBBFFD7FC46EBB5FBE31333" box="[903,1060,1781,1808]" metricMagnitude="3" metricUnit="m" metricValue="1.207" pageId="13" pageNumber="14" unit="m" value="1207.0">
|
||
<quantity id="13409B6DFFBBFFD7FC46EBB5FC25132C" box="[903,994,1781,1808]" metricMagnitude="3" metricUnit="m" metricValue="1.207" pageId="13" pageNumber="14" unit="m" value="1207.0">1207 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="A0061048FFBBFFD7FBF5EBB5FB1C1333" box="[1076,1243,1781,1808]" pageId="13" pageNumber="14" value="2013-11-25">
|
||
<collectingDate id="B042E9A0FFBBFFD7FBF5EBB5FB1C1333" box="[1076,1243,1781,1808]" pageId="13" pageNumber="14" value="2013-11-25">25 Nov. 2013</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="794D535EFFBBFFD7FB28EBB6FA85132C" box="[1257,1346,1782,1808]" pageId="13" pageNumber="14">T. Pape</collectorName>
|
||
and
|
||
<collectorName id="794D535EFFBBFFD7FF7CEA59FEF0130E" box="[189,311,1816,1843]" pageId="13" pageNumber="14">N. Scharff</collectorName>
|
||
leg.;
|
||
<collectingMethod id="0DF94E9FFFBBFFD7FEB8EA58FDEB130E" box="[377,556,1816,1842]" pageId="13" pageNumber="14">pitfall trapping</collectingMethod>
|
||
;
|
||
<specimenCode id="841E9EF3FFBBFFD7FDF6EA59FD33130E" box="[567,756,1816,1843]" collectionCode="NHMD" pageId="13" pageNumber="14">NHMD 621643</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D4073688FFBBFFD7FF7CEA1AFE541348" blockId="13.[189,403,1882,1908]" box="[189,403,1882,1908]" pageId="13" pageNumber="14">
|
||
<heading id="8F4F81E4FFBBFFD7FF7CEA1AFE541348" bold="true" box="[189,403,1882,1908]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="E6CCEA9AFFBBFFD7FF7CEA1AFE541348" bold="true" box="[189,403,1882,1908]" pageId="13" pageNumber="14">Descriptive notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFBBFFD7FF7CEAC8FDCC13D5" blockId="13.[189,1399,1928,2026]" pageId="13" pageNumber="14">
|
||
The new specimens perfectly fit the original description of
|
||
<taxonomicName id="13B84D0BFFBBFFD7FCAFEAC9FC38139E" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[878,1023,1928,1955]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipala">
|
||
<emphasis id="E6CCEA9AFFBBFFD7FCAFEAC9FC38139E" box="[878,1023,1928,1955]" italics="true" pageId="13" pageNumber="14">A. longipala</emphasis>
|
||
</taxonomicName>
|
||
, except that they are marginally larger (51–52 podous rings, diameter
|
||
<quantity id="13409B6DFFBBFFD7FDBEEAECFCE813FA" box="[639,815,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" metricValueMax="2.02" metricValueMin="1.84" pageId="13" pageNumber="14" unit="mm" value="1.93" valueMax="2.02" valueMin="1.84">1.84–2.02 mm</quantity>
|
||
, vs 50–52 podous rings, diameter
|
||
<quantity id="13409B6DFFBBFFD7FB0BEAECFAB013FA" box="[1226,1399,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.9" metricValueMin="1.78" pageId="13" pageNumber="14" unit="mm" value="1.8399999999999999" valueMax="1.9" valueMin="1.78">1.78–1.90 mm</quantity>
|
||
for
|
||
<typeStatus id="0B03882AFFBBFFD7FF26EA8FFEDD13D5" box="[231,282,1999,2025]" pageId="13" pageNumber="14">type</typeStatus>
|
||
specimens) (
|
||
<figureCitation id="4C832A0DFFBBFFD7FE75EA8FFE3B13D5" box="[436,508,1999,2025]" captionStart="Fig" captionStartId="11.[187,230,1893,1919]" captionTargetBox="[193,1391,1095,1856]" captionTargetId="figure@11.[192,1402,1089,1860]" captionTargetPageId="11" captionText="Fig. 9. Aquattuor spp. Body size (number of podous rings and midbody vertical diameter) in males of Aquattuor species from the Udzungwa Mts, and of A. claudiahempae Enghoff & Frederiksen, 2015 from Mt Kilimanjaro. In case of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3755007" httpUri="https://zenodo.org/record/3755007/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="80C76600FFB8FFD4FF7CEA5AFEA613D9" ID-DOI="http://doi.org/10.5281/zenodo.3755013" ID-Zenodo-Dep="3755013" httpUri="https://zenodo.org/record/3755013/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1818,1844]" targetBox="[254,1335,280,1761]" targetPageId="14">
|
||
<paragraph id="D4073688FFB8FFD4FF7CEA5AFEA613D9" blockId="14.[189,1399,1818,2021]" pageId="14" pageNumber="15">
|
||
<emphasis id="E6CCEA9AFFB8FFD4FF7CEA5AFEDF1308" bold="true" box="[189,280,1818,1844]" pageId="14" pageNumber="15">Fig. 11.</emphasis>
|
||
<taxonomicName id="13B84D0BFFB8FFD4FEE1EA5BFD031309" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[288,708,1818,1845]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="longipala">
|
||
<emphasis id="E6CCEA9AFFB8FFD4FEE1EA5BFDC81308" box="[288,527,1818,1845]" italics="true" pageId="14" pageNumber="15">Aquattuor longipala</emphasis>
|
||
<bibRefCitation id="B0294B79FFB8FFD4FDD6EA5AFD031309" author="Enghoff H. & Frederiksen S. B." box="[535,708,1818,1845]" pageId="14" pageNumber="15" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
, male, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642), left gonopod.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FD4CEA7DFD01136B" bold="true" box="[653,710,1853,1880]" pageId="14" pageNumber="15">A–C</emphasis>
|
||
. Coxa.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FCE1EA7DFCF2136B" bold="true" box="[800,821,1853,1879]" pageId="14" pageNumber="15">A</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FC31EA7EFBC31364" bold="true" box="[1008,1028,1854,1880]" pageId="14" pageNumber="15">B</emphasis>
|
||
. Mesal view.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FB64EA7DFB7D136B" bold="true" box="[1189,1210,1853,1879]" pageId="14" pageNumber="15">C</emphasis>
|
||
. Posterior view.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FF7CEA21FF3F1347" bold="true" box="[189,248,1889,1915]" pageId="14" pageNumber="15">D–H</emphasis>
|
||
. Telopodite.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FE56EA21FE6B1347" bold="true" box="[407,428,1889,1915]" pageId="14" pageNumber="15">D</emphasis>
|
||
. Tip of telomere.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FD48EA21FD5A1347" bold="true" box="[649,669,1889,1915]" pageId="14" pageNumber="15">E</emphasis>
|
||
. Posterior view.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FCABEA21FCBB1347" bold="true" box="[874,892,1889,1915]" pageId="14" pageNumber="15">F</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FB80EA21FB9F1347" bold="true" box="[1089,1112,1889,1915]" pageId="14" pageNumber="15">G</emphasis>
|
||
. Posterior-ventral view.
|
||
<emphasis id="E6CCEA9AFFB8FFD4FF7CEAC4FF1313A2" bold="true" box="[189,212,1924,1950]" pageId="14" pageNumber="15">H</emphasis>
|
||
. Distal (ventral) view. Abbreviations:
|
||
<emphasis id="E6CCEA9AFFB8FFD4FD54EAC4FD6213A2" box="[661,677,1924,1950]" italics="true" pageId="14" pageNumber="15">li</emphasis>
|
||
= lateral incision;
|
||
<emphasis id="E6CCEA9AFFB8FFD4FC40EAC5FC5913A2" box="[897,926,1925,1950]" italics="true" pageId="14" pageNumber="15">mi</emphasis>
|
||
= mesal incision;
|
||
<emphasis id="E6CCEA9AFFB8FFD4FBB4EAC4FB6613A2" box="[1141,1185,1924,1950]" italics="true" pageId="14" pageNumber="15">mpl</emphasis>
|
||
= mesal-posterior telomeral lamella;
|
||
<emphasis id="E6CCEA9AFFB8FFD4FE5EEAE8FE7B13FD" box="[415,444,1960,1985]" italics="true" pageId="14" pageNumber="15">pa</emphasis>
|
||
= palette;
|
||
<emphasis id="E6CCEA9AFFB8FFD4FD80EAE7FDAD13FD" box="[577,618,1959,1985]" italics="true" pageId="14" pageNumber="15">slm</emphasis>
|
||
= solenomere;
|
||
<emphasis id="E6CCEA9AFFB8FFD4FCE8EAE8FC8113FD" box="[809,838,1960,1985]" italics="true" pageId="14" pageNumber="15">tm</emphasis>
|
||
= telomere. Scale bars: A–C, E–H = 0.1 mm; D = 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D4073688FFB9FFD5FF7CEC4BFBA21550" blockId="15.[189,1399,267,364]" pageId="15" pageNumber="16">
|
||
First pair of male legs (
|
||
<figureCitation id="4C832A0DFFB9FFD5FE0AEC4BFDC9151A" box="[459,526,267,294]" captionStart="Fig" captionStartId="7.[189,232,1887,1913]" captionTargetBox="[234,1343,270,1842]" captionTargetId="figure@7.[229,1354,262,1848]" captionTargetPageId="7" captionText="Fig. 5. Aquattuor spp., first pair of male legs. A–C. A. major Enghoff, 2015, paratype, ♂ (NHMD 621644), first pair of legs in situ. D–F. A. longipala Enghoff, 2015, male from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642). A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754999" httpUri="https://zenodo.org/record/3754999/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
|
||
D–F) each with a single slender, finger-shaped prefemoral process (
|
||
<emphasis id="E6CCEA9AFFB9FFD5FAE3EC4CFA80151A" box="[1314,1351,268,294]" italics="true" pageId="15" pageNumber="16">pfp</emphasis>
|
||
); in addition, each prefemur has a smaller, irregular lobe (
|
||
<emphasis id="E6CCEA9AFFB9FFD5FCF5EC6FFC9E1575" box="[820,857,303,329]" italics="true" pageId="15" pageNumber="16">dpl</emphasis>
|
||
) on the oral side of the distal margin; setae of femur much shorter than corresponding setae on normal walking legs (
|
||
<figureCitation id="4C832A0DFFB9FFD5FC3AEC12FB901550" box="[1019,1111,338,364]" captionStart="Fig" captionStartId="10.[189,232,1878,1904]" captionTargetBox="[205,1373,1111,1837]" captionTargetId="figure@10.[189,1399,1092,1845]" captionTargetPageId="10" captionText="Fig. 8. Aquattuor spp., males, left leg of first pair. A. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650): setae same length as in normal walking legs. B. A. longipala Enghoff, 2015, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642): setae of femur shortened. C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621646): all setae shortened. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755005" httpUri="https://zenodo.org/record/3755005/files/figure.png" pageId="15" pageNumber="16">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB9FFD5FF7CECD3FC0F1591" blockId="15.[189,968,403,430]" box="[189,968,403,430]" pageId="15" pageNumber="16">
|
||
<figureCitation id="4C832A0DFFB9FFD5FF7CECD3FEC81591" box="[189,271,403,429]" captionStart="Fig" captionStartId="14.[189,232,1818,1844]" captionTargetBox="[254,1335,280,1761]" captionTargetId="figure@14.[231,1356,265,1775]" captionTargetPageId="14" captionText="Fig. 11. Aquattuor longipala Enghoff, 2015, male, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642), left gonopod. A–C. Coxa. A. Anterior view. B. Mesal view. C. Posterior view. D–H. Telopodite. D. Tip of telomere. E. Posterior view. F. Anterior view. G. Posterior-ventral view. H. Distal (ventral) view. Abbreviations: li = lateral incision; mi = mesal incision; mpl = mesal-posterior telomeral lamella; pa = palette; slm = solenomere; tm = telomere. Scale bars: A–C, E–H = 0.1 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3755013" httpUri="https://zenodo.org/record/3755013/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
|
||
shows a gonopod of one of the newly collected specimens.
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB9FFD5FF7CEC95FC7115D2" blockId="15.[189,950,469,496]" box="[189,950,469,496]" pageId="15" pageNumber="16">
|
||
<taxonomicName id="13B84D0BFFB9FFD5FF7CEC95FE6C15D3" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[189,427,469,495]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="longipala">
|
||
<emphasis id="E6CCEA9AFFB9FFD5FF7CEC95FE6C15D3" box="[189,427,469,495]" italics="true" pageId="15" pageNumber="16">Aquattuor longipala</emphasis>
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="13B84D0BFFB9FFD5FD41EC95FD2015D2" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[640,743,469,495]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="major">
|
||
<emphasis id="E6CCEA9AFFB9FFD5FD41EC95FD2015D2" box="[640,743,469,495]" italics="true" pageId="15" pageNumber="16">A. major</emphasis>
|
||
</taxonomicName>
|
||
(see Discussion).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA26503FFB9FFD5FF7CEF52FC7F1642" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="D4073688FFB9FFD5FF7CEF52FE911610" blockId="15.[189,342,530,556]" box="[189,342,530,556]" pageId="15" pageNumber="16">
|
||
<heading id="8F4F81E4FFB9FFD5FF7CEF52FE911610" bold="true" box="[189,342,530,556]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="E6CCEA9AFFB9FFD5FF7CEF52FE911610" bold="true" box="[189,342,530,556]" pageId="15" pageNumber="16">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB9FFD5FF7CEF01FC7F1642" blockId="15.[189,1398,577,639]" pageId="15" pageNumber="16">
|
||
Known from two sites in the Udzungwa Mts, viz., West Kilombero FR and Mito Mitatu in Udzungwa Mts National Park (
|
||
<figureCitation id="4C832A0DFFB9FFD5FE65EF24FE2E1642" box="[420,489,612,638]" captionStart="Fig" captionStartId="2.[189,232,1948,1974]" captionTargetBox="[192,1385,1023,1872]" captionTargetId="figure@2.[186,1391,1009,1876]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites for Aquattuor spp. in the Udzungwa Mts. Based on Marshall et al. (2010: fig. 1); inset by permission of the Eastern Arc Mountains Conservation Endowment Fund." figureDoi="http://doi.org/10.5281/zenodo.3754991" httpUri="https://zenodo.org/record/3754991/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
|
||
). Altitudinal range:
|
||
<quantity id="13409B6DFFB9FFD5FD19EF24FCBD1643" box="[728,890,612,639]" metricMagnitude="3" metricUnit="m" metricValue="1.3794999999999997" metricValueMax="1.552" metricValueMin="1.207" pageId="15" pageNumber="16" unit="m" value="1379.5" valueMax="1552.0" valueMin="1207.0">1207–1552 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |