213 lines
20 KiB
XML
213 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/fr.25.83034" ID-Pensoft-Pub="2193-0074-1-187" ID-Pensoft-UUID="86764B0ACD9453CD965B288014B441CD" ID-ZooBank="6402F8F152294153823FCAEA106F90A1" ModsDocID="2193-0074-25-1-187" checkinTime="1667366246929" checkinUser="pensoft" docAuthor="Spasojevic, Tamara, Broad, Gavin R. & Klopfstein, Seraina" docDate="2022" docId="5A877DF635D25ECC8E3CB5E303CEF8E3" docLanguage="en" docName="FossilRecord 25(1): 187-212" docOrigin="Fossil Record 25 (1)" docPubDate="2022-06-07" docSource="http://dx.doi.org/10.3897/fr.25.83034" docTitle="Hallocinetus arvernus Spasojevic & Broad & Klopfstein 2022, comb. nov." docType="treatment" docVersion="1" id="86764B0ACD9453CD965B288014B441CD" lastPageNumber="187" masterDocId="86764B0ACD9453CD965B288014B441CD" masterDocTitle="Revision of 18 ichneumonid fossil species (Hymenoptera, Ichneumonidae) highlights the need for open nomenclature in palaeontology" masterLastPageNumber="212" masterPageNumber="187" pageNumber="187" updateTime="1667366246929" updateUser="pensoft">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Revision of 18 ichneumonid fossil species (Hymenoptera, Ichneumonidae) highlights the need for open nomenclature in palaeontology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Spasojevic, Tamara</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5301-5722</mods:nameIdentifier>
|
||
<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, Basel, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, Bern, Switzerland</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Broad, Gavin R.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7223-5333</mods:nameIdentifier>
|
||
<mods:affiliation>Natural History Museum, Cromwell Road, London SW 7 6 BD, UK</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Klopfstein, Seraina</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4025-975X</mods:nameIdentifier>
|
||
<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, Basel, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, Bern, Switzerland</mods:affiliation>
|
||
<mods:nameIdentifier type="email">seraina.klopfstein@bs.ch</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Fossil Record</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2022</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2022-06-07</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>25</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>187</mods:start>
|
||
<mods:end>212</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/fr.25.83034</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/fr.25.83034</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">2193-0074-1-187</mods:identifier>
|
||
<mods:identifier type="ZooBank">6402F8F152294153823FCAEA106F90A1</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">86764B0ACD9453CD965B288014B441CD</mods:identifier>
|
||
</mods:mods>
|
||
<treatment LSID="urn:lsid:plazi:treatment:5A877DF635D25ECC8E3CB5E303CEF8E3" httpUri="http://treatment.plazi.org/id/5A877DF635D25ECC8E3CB5E303CEF8E3" lastPageNumber="187" pageId="0" pageNumber="187">
|
||
<subSubSection pageId="0" pageNumber="187" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="187">
|
||
<taxonomicName LSID="5A877DF6-35D2-5ECC-8E3C-B5E303CEF8E3" authority="(Piton, 1940)" authorityName="Spasojevic & Broad & Klopfstein" authorityYear="2022" baseAuthorityName="Piton" baseAuthorityYear="1940" class="Insecta" family="Ichneumonidae" genus="Hallocinetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hallocinetus arvernus" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="arvernus" status="comb. nov.">Hallocinetus? arvernus (Piton, 1940)</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="187">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="187" type="description">
|
||
<paragraph pageId="0" pageNumber="187">
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hallocinetus? arvernus (Piton, 1940) comb. nov. A. Photograph of the holotype obtained from MNHN. F; B. Our interpretative drawing of the fossil. Thicker lines indicate outlines of body structures, thinner lines show characters inside these outlines, and dotted lines represent uncertain interpretations." figureDoi="10.3897/fr.25.83034.figure2" httpUri="https://binary.pensoft.net/fig/693061" pageId="0" pageNumber="187">Fig. 2</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="187" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="187">
|
||
<taxonomicName authorityName="Spasojevic & Broad & Klopfstein" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Phaenolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaenolobus arvernus" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="arvernus" status="comb. nov.">* Phaenolobus arvernus</taxonomicName>
|
||
Piton, 1940
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="187" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="187">Material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="187">
|
||
<materialsCitation accessionNumber="A57301" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" specimenCount="1" typeStatus="Holotype">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
(part only, MNHN.F.
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A57301">A57301</accessionNumber>
|
||
) examined at MNHN.F.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="187" type="stratum">
|
||
<paragraph pageId="0" pageNumber="187">Stratum.</paragraph>
|
||
<paragraph pageId="0" pageNumber="187">
|
||
Menat Formation,
|
||
<normalizedToken originalValue="Puy-de-Dôme">Puy-de-Dome</normalizedToken>
|
||
, France. Late Palaeocene (Thanetian), 58.7-55.8 Ma.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="187" type="description">
|
||
<paragraph pageId="0" pageNumber="187">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="187">Sex unknown. Holotype in lateral view, rather poorly preserved, with head, outline of mesosoma and partial metasoma, partial fore wing and outline of hind legs. Body length ~10 mm.</paragraph>
|
||
<paragraph pageId="0" pageNumber="187">Dark brown including wing venation, legs and posterior part of metasoma lighter brown or reddish.</paragraph>
|
||
<paragraph pageId="0" pageNumber="187">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Head</emphasis>
|
||
rather large and high, compound eye about 0.7-0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
head height in lateral view. Antennae incomplete, scape rather short.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Mesosoma</emphasis>
|
||
rather short, without any details preserved.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Fore wing</emphasis>
|
||
7.2 mm; areolet obliquely quadrate and strongly petiolate, 2 + 3M longer than 4M; 1cu-a meeting M + Cu nearly opposite of 1M & 1Rs; 3Cu a bit longer than 2cu-a; cell 2R1 3.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than wide; 2m-cu without bullae in posterior half, potentially with one or two bullae in anterior half.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Hind legs</emphasis>
|
||
rather stout; femur 3.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
, tibia 5.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than wide; hind tarsi with first segment much longer than second and third, which are about as long as wide.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Metasoma</emphasis>
|
||
with T1 elongate, probably with a dorso-lateral carina; sternites rather well sclerotized; tip not well preserved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="187" type="interpretation">
|
||
<paragraph pageId="0" pageNumber="187">Interpretation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="187">
|
||
This is a rather poorly preserved fossil, from which not many characters can be discerned. Comparing the holotype and its high-resolution photograph with the drawing in the original description (p. 228, fig. 94,
|
||
<bibRefCitation author="Piton, LEA" journalOrPublisher="Vallier, Clermont-Ferrand" pageId="0" pageNumber="187" refId="B32" refString="Piton, LEA, 1940. Paleontologie du gisement eocene de Menat (Puy-de-Dome): flore et faune. Imprimeries P. Vallier, Clermont-Ferrand" title="Paleontologie du gisement eocene de Menat (Puy-de-Dome): flore et faune. Imprimeries P." year="1940">Piton 1940</bibRefCitation>
|
||
), we found that the latter shows a considerable amount of artistic freedom. The antennae are drawn as very short and apically expanded, while they are incomplete and parallel-sided in the fossil. The head shows a beak-like structure not discernible clearly in the fossil and unknown in extant representatives of the subfamily, and the small eyes appear around 0.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as the head length in profile, compared to 0.7-0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
in the fossil. The areolet is depicted as being open, while we found it to be closed and strongly petiolate in the fossil. The tarsi of the fore and hind legs appear complete with claws present in the drawing, while at most the first tarsal segments are visible in the fossil. The femora of all legs are much thinner in the drawing than in the fossil, and the drawing of the posterior half of the fore wing, which largely overlaps the metasoma, contains veins not found in any ichneumonid wasp. Piton also drew the elongate-triangular hypopygium typical for the subfamily
|
||
<taxonomicName authorityName="Foerster" authorityYear="1869" lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Acaenitinae">Acaenitinae</taxonomicName>
|
||
; we are more cautious, noting that the apical part of the metasoma is too poorly preserved to draw conclusions about the shape of the hypopygium. Additional characters that could help subfamily placement, such as the shape of T1 and position of its spiracle, the propodeal carination and length of the ovipositor are indiscernible. The general habitus of the fossil, especially the shape of the metasoma in profile and stout legs, does resemble
|
||
<taxonomicName authorityName="Foerster" authorityYear="1869" lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Acaenitinae">Acaenitinae</taxonomicName>
|
||
, but in a rather vague fashion. Other characters are somewhat at odds with this placement, although not completely so. For instance, most
|
||
<taxonomicName authorityName="Foerster" authorityYear="1869" lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Acaenitinae">Acaenitinae</taxonomicName>
|
||
have 2m-cu with two bullae, one in the anterior and one in the posterior half, which would disagree with the fossil; however, some genera in former
|
||
<taxonomicName authorityName="Forster" authorityYear="1869" lsidName="" pageId="0" pageNumber="187" rank="tribe" tribe="Acaenitini">Acaenitini</taxonomicName>
|
||
, such as
|
||
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Hieroceryx" kingdom="Animalia" lsidName="Hieroceryx" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="187">Hieroceryx</emphasis>
|
||
</taxonomicName>
|
||
Tosquinet, 1896 and
|
||
<taxonomicName authorityName="Townes" authorityYear="1971" class="Insecta" family="Ichneumonidae" genus="Prosacron" kingdom="Animalia" lsidName="Prosacron" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="187">Prosacron</emphasis>
|
||
</taxonomicName>
|
||
Townes, 1971, have both bullae in the anterior half and separated by a very short tubular part of the vein. Moreover,
|
||
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Phaenolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaenolobus" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="187">Phaenolobus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Phorotrophus" kingdom="Animalia" lsidName="Phorotrophus" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="187">Phorotrophus</emphasis>
|
||
</taxonomicName>
|
||
Saussure, 1892 can in fact have a single bulla, in the former located in the dorsal half of 2m-cu. However, all former
|
||
<taxonomicName authorityName="Forster" authorityYear="1869" lsidName="" pageId="0" pageNumber="187" rank="tribe" tribe="Acaenitini">Acaenitini</taxonomicName>
|
||
have an open areolet, while a closed and petiolate areolet occurs only in the former tribe
|
||
<taxonomicName authorityName="Clement" authorityYear="1938" lsidName="" pageId="0" pageNumber="187" rank="tribe" tribe="Coleocentrini">Coleocentrini</taxonomicName>
|
||
, which has turned out paraphyletic with respect to the former
|
||
<taxonomicName authorityName="Forster" authorityYear="1869" lsidName="" pageId="0" pageNumber="187" rank="tribe" tribe="Acaenitini">Acaenitini</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.1046/j.1365-3113.1998.00057.x" author="Wahl, D" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="187" pagination="265 - 298" refId="B52" refString="Wahl, D, Gauld, I, 1998. The cladistics and higher classification of the Pimpliformes (Hymenoptera: Ichneumonidae). Systematic Entomology 23: 265 - 298, DOI: https://doi.org/10.1046/j.1365-3113.1998.00057.x" title="The cladistics and higher classification of the Pimpliformes (Hymenoptera: Ichneumonidae)." url="https://doi.org/10.1046/j.1365-3113.1998.00057.x" volume="23" year="1998">Wahl and Gauld 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1080/00222930902807783" author="Quicke, DLJ" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="187" pagination="1305 - 1421" refId="B34" refString="Quicke, DLJ, Laurenne, NM, Fitton, MG, Broad, GR, 2009. A thousand and one wasps: a 28S rDNA and morphological phylogeny of the Ichneumonidae (Insecta: Hymenoptera) with an investigation into alignment parameter space and elision. Journal of Natural History 43: 1305 - 1421, DOI: https://doi.org/10.1080/00222930902807783" title="A thousand and one wasps: a 28 S rDNA and morphological phylogeny of the Ichneumonidae (Insecta: Hymenoptera) with an investigation into alignment parameter space and elision." url="https://doi.org/10.1080/00222930902807783" volume="43" year="2009">Quicke et al. 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1111/syen.12333" author="Klopfstein, S" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="187" pagination="361 - 383" refId="B20" refString="Klopfstein, S, Langille, B, Spasojevic, T, Broad, GR, Cooper, SJB, Austin, AD, Niehuis, O, 2019. Hybrid capture data unravel a rapid radiation of pimpliform parasitoid wasps (Hymenoptera: Ichneumonidae: Pimpliformes). Systematic Entomology 44: 361 - 383, DOI: https://doi.org/10.1111/syen.12333" title="Hybrid capture data unravel a rapid radiation of pimpliform parasitoid wasps (Hymenoptera: Ichneumonidae: Pimpliformes)." url="https://doi.org/10.1111/syen.12333" volume="44" year="2019">Klopfstein et al. 2019</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1093/sysbio/syaa079" author="Spasojevic, T" journalOrPublisher="Systematic Biology" pageId="0" pageNumber="187" pagination="322 - 339" refId="B45" refString="Spasojevic, T, Broad, GR, Saeaeksjaervi, IE, Schwarz, M, Ito, M, Korenko, S, Klopfstein, S, 2021. Mind the Outgroup and Bare Branches in Total-Evidence Dating: a Case Study of Pimpliform Darwin Wasps (Hymenoptera, Ichneumonidae). Systematic Biology 70: 322 - 339, DOI: https://doi.org/10.1093/sysbio/syaa079" title="Mind the Outgroup and Bare Branches in Total-Evidence Dating: a Case Study of Pimpliform Darwin Wasps (Hymenoptera, Ichneumonidae)." url="https://doi.org/10.1093/sysbio/syaa079" volume="70" year="2021">Spasojevic et al. 2021</bibRefCitation>
|
||
). The characters for generic placement within the former
|
||
<taxonomicName authorityName="Clement" authorityYear="1938" lsidName="" pageId="0" pageNumber="187" rank="tribe" tribe="Coleocentrini">Coleocentrini</taxonomicName>
|
||
, such as the shape of the clypeus, size and shape of the hypopygium and modifications on tarsal claws (
|
||
<bibRefCitation author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="187" refId="B50" refString="Townes, HK, 1971. The genera of Ichneumonidae, Part 4. Memoirs of the American Entomological Institute: 1-372." title="The genera of Ichneumonidae, Part 4. Memoirs of the American Entomological Institute: 1 - 372." year="1971">Townes 1971</bibRefCitation>
|
||
) are not visible in the fossil. Since we also have no character evidence for placing the fossil in a different subfamily, we tentatively place it in
|
||
<taxonomicName authorityName="Viktorov" authorityYear="1962" class="Insecta" family="Ichneumonidae" genus="Hallocinetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hallocinetus" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="187">Hallocinetus</emphasis>
|
||
</taxonomicName>
|
||
Viktorov, 1962, to which it superficially resembles in the shape of the anterior margin of the mesoscutum and stout and large hind legs. However, we acknowledge the uncertainty in its placement by placing a question mark after the genus and subfamily names.
|
||
</paragraph>
|
||
<caption doi="10.3897/fr.25.83034.figure2" httpUri="https://binary.pensoft.net/fig/693061" pageId="0" pageNumber="187" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="187">
|
||
<emphasis bold="true" pageId="0" pageNumber="187">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Spasojevic & Broad & Klopfstein" authorityYear="2022" baseAuthorityName="Piton" baseAuthorityYear="1940" class="Insecta" family="Ichneumonidae" genus="Hallocinetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hallocinetus arvernus" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="arvernus">Hallocinetus? arvernus</taxonomicName>
|
||
(Piton, 1940) comb. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="187">A.</emphasis>
|
||
Photograph of the holotype obtained from MNHN.F;
|
||
<emphasis bold="true" pageId="0" pageNumber="187">B.</emphasis>
|
||
Our interpretative drawing of the fossil. Thicker lines indicate outlines of body structures, thinner lines show characters inside these outlines, and dotted lines represent uncertain interpretations.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="187">
|
||
<emphasis bold="true" pageId="0" pageNumber="187">
|
||
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1869" genus="Cremastinae" lsidName="Cremastinae" pageId="0" pageNumber="187" rank="genus">Cremastinae</taxonomicName>
|
||
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
|
||
, 1869
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="187">
|
||
<emphasis bold="true" pageId="0" pageNumber="187">
|
||
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Dimophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dimophora" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="187">Dimophora</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
|
||
, 1869
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |