265 lines
33 KiB
XML
265 lines
33 KiB
XML
<document id="D7542DA3310C6C4F5F7B96D95C7B123A" ID-DOI="10.11646/zootaxa.5033.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5499124" ID-ZooBank="4314F784-A510-4F36-9E11-ED1EAC83CEBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631246977811" checkinUser="plazi" docAuthor="Johnson, Robert A." docDate="2021" docId="5027B677FFFFA428FF25FCE1FC2C6C7D" docLanguage="en" docName="zootaxa.5033.1.1.pdf" docOrigin="Zootaxa 5033 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5033.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pogonomyrmex bolivianus Johnson 2021, NEW SPECIES" docType="treatment" docVersion="3" lastPageNumber="55" masterDocId="AC1ECE0FFFCAA41EFFB2FF86FFC76B0C" masterDocTitle="A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part II" masterLastPageNumber="230" masterPageNumber="1" pageNumber="54" updateTime="1699406547276" updateUser="plazi">
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<mods:titleInfo id="5EB39C3868A1B16FF6876EA74FBF2523">
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<mods:title id="63DCC7E1B1C7FE00861ECE47E4586717">A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part II</mods:title>
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<mods:name id="69AC00ADC5D3ADA11EFE6AB64F72EACF" type="personal">
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<mods:namePart id="AE19DECB0D194EDF651FB513A74B7B49">Johnson, Robert A.</mods:namePart>
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<mods:title id="8507F7061F41C7862DB3857B97638A63">Zootaxa</mods:title>
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<mods:part id="4DADC896C861BD4FF9F29686582D09A2">
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<mods:date id="1899FDD846721A109AFFA31F21B7CF98">2021</mods:date>
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<mods:number id="AA60483A73E589E54CD8B17C3D2B7708">2021-09-10</mods:number>
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<mods:detail id="933E645731B9AE8A3C9A9E82F40EAC7F" type="volume">
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<mods:number id="936DBC3E558266420C4C76718E0F3C4C">5033</mods:number>
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<mods:number id="418958A6D2AF844E467CDFCB9F2376B9">1</mods:number>
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<mods:start id="550E94DD67483F70FD61B9D92B68C21A">1</mods:start>
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<mods:url id="4D4A8C30E264D5C19F8FAAC2B6267623">http://dx.doi.org/10.11646/zootaxa.5033.1.1</mods:url>
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<mods:classification id="7B4D11BF729D3F307788364B5BBC6FB3">journal article</mods:classification>
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<mods:identifier id="9A6AC899F158D89B2E85A4CFCA3CB081" type="DOI">10.11646/zootaxa.5033.1.1</mods:identifier>
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<mods:identifier id="97C188CE66C07775651FF31C7F90367B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1C079EB1A7C527DC59BC1C95F7C1FB95" type="Zenodo-Dep">5499124</mods:identifier>
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<mods:identifier id="CEABA0ABCF6740034F520FA947D73AC3" type="ZooBank">4314F784-A510-4F36-9E11-ED1EAC83CEBF</mods:identifier>
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<treatment id="5027B677FFFFA428FF25FCE1FC2C6C7D" ID-DOI="http://doi.org/10.5281/zenodo.5499160" ID-Zenodo-Dep="5499160" LSID="urn:lsid:plazi:treatment:5027B677FFFFA428FF25FCE1FC2C6C7D" httpUri="http://treatment.plazi.org/id/5027B677FFFFA428FF25FCE1FC2C6C7D" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
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<subSubSection id="909454EAFFFFA42BFF25FCE1FD6E688C" box="[151,681,870,897]" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="D8310761FFFFA42BFF25FCE1FD6E688C" blockId="53.[151,681,870,969]" box="[151,681,870,897]" pageId="53" pageNumber="54">
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<heading id="8379B00DFFFFA42BFF25FCE1FD6E688C" bold="true" box="[151,681,870,897]" fontSize="11" level="2" pageId="53" pageNumber="54" reason="4">
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<emphasis id="EAFADB73FFFFA42BFF25FCE1FD6E688C" bold="true" box="[151,681,870,897]" pageId="53" pageNumber="54">
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<taxonomicName id="1F8E7CE2FFFFA42BFF25FCE1FE1C688D" authority="Johnson, 2021" authorityName="Johnson" authorityYear="2021" box="[151,475,870,897]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bolivianus" status="sp. nov.">
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<emphasis id="EAFADB73FFFFA42BFF25FCE1FE1C688D" bold="true" box="[151,475,870,897]" italics="true" pageId="53" pageNumber="54">Pogonomyrmex bolivianus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F1C96608FFFFA42BFE50FCE1FD6E688C" box="[482,681,870,897]" pageId="53" pageNumber="54" rank="species">NEW SPECIES</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="909454EAFFFFA428FF25FC0DFDB86FAD" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="description">
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<paragraph id="D8310761FFFFA42BFF25FC0DFED168A9" blockId="53.[151,681,870,969]" box="[151,278,907,933]" pageId="53" pageNumber="54">
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(
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<figureCitation id="40B51BE4FFFFA42BFF2DFC0DFEC968A9" box="[159,270,907,933]" captionStart="FIGURE 23" captionStartId="55.[151,250,1916,1940]" captionTargetBox="[418,1168,181,1892]" captionTargetId="figure-15@55.[418,1170,181,1892]" captionTargetPageId="55" captionText="FIGURE 23. Photographs of Pogonomyrmex bolivianus holotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0914125). Photographs by Michele Esposito from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.5499188" httpUri="https://zenodo.org/record/5499188/files/figure.png" pageId="53" pageNumber="54">Figure 23</figureCitation>
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)
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</paragraph>
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<paragraph id="D8310761FFFFA42BFF25FC29FEAD68C5" blockId="53.[151,681,870,969]" box="[151,362,943,969]" pageId="53" pageNumber="54">Distribution—22B</paragraph>
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<paragraph id="D8310761FFFFA42BFF25FC71FD936F56" blockId="53.[151,1437,1014,2013]" pageId="53" pageNumber="54">
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<materialsCitation id="68E60D3CFFFFA42BFF25FC71FEB06F3A" collectingDate="1951-02-25" collectionCode="CASC" collectorName="E. S. Ross & A. E. Michelbacher" country="Bolivia" location="La Paz" pageId="53" pageNumber="54" specimenCode="CASENT0914125" specimenCount="1" stateProvince="La Paz" typeStatus="holotype">
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<typeStatus id="0735B9C3FFFFA42BFF25FC71FF386F1D" box="[151,255,1015,1041]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
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worker [
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<collectionCode id="BE9F9FA4FFFFA42BFED1FC71FE746F1D" box="[355,435,1015,1041]" pageId="53" pageNumber="54">CASC</collectionCode>
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:
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<specimenCode id="8828AF1AFFFFA42BFE0CFC70FD536F1D" box="[446,660,1014,1041]" collectionCode="CASENT" pageId="53" pageNumber="54">CASENT0914125</specimenCode>
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]:
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<collectingCountry id="A09947F1FFFFA42BFD17FC71FCDC6F1D" box="[677,795,1015,1041]" name="Bolivia" pageId="53" pageNumber="54">BOLIVIA</collectingCountry>
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,
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<collectingRegion id="1A4AC983FFFFA42BFC94FC71FCBF6F1D" box="[806,888,1015,1041]" country="Bolivia" name="La Paz" pageId="53" pageNumber="54">La Paz</collectingRegion>
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:
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<quantity id="1F76AA84FFFFA42BFC30FC70FC046F1D" box="[898,963,1014,1041]" metricMagnitude="4" metricUnit="m" metricValue="7.242048" pageId="53" pageNumber="54" unit="mi" value="45.0">45 mi</quantity>
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S
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<collectingRegion id="1A4AC983FFFFA42BFC52FC71FBF76F1D" box="[992,1072,1015,1041]" country="Bolivia" name="La Paz" pageId="53" pageNumber="54">La Paz</collectingRegion>
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,
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<date id="AC3021A1FFFFA42BFB8EFC70FAC46F1D" box="[1084,1283,1014,1041]" pageId="53" pageNumber="54" value="1951-02-25">
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<collectingDate id="BC74D849FFFFA42BFB8EFC70FAC46F1D" box="[1084,1283,1014,1041]" pageId="53" pageNumber="54" value="1951-02-25">25 February 1951</collectingDate>
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</date>
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(
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<collectorName id="757B62B7FFFFA42BFAA1FC71FAB86F1D" box="[1299,1407,1015,1041]" pageId="53" pageNumber="54">E.S. Ross</collectorName>
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&
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<collectorName id="757B62B7FFFFA42BFF25FB9DFEB76F3A" box="[151,368,1051,1078]" pageId="53" pageNumber="54">A.E. Michelbacher</collectorName>
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)
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</materialsCitation>
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.
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<materialsCitation id="68E60D3CFFFFA42BFE32FB9DFC0A6F3A" box="[384,973,1051,1078]" collectingDate="1951-02-25" collectionCode="CASC" collectorName="E. S. Ross & A. E. Michelbacher" country="Bolivia" location="La Paz" pageId="53" pageNumber="54" specimenCount="7" specimenCount-worker="7" stateProvince="La Paz" typeStatus="paratype">
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<typeStatus id="0735B9C3FFFFA42BFE32FB9DFE376F39" box="[384,496,1051,1077]" pageId="53" pageNumber="54" type="paratype">Paratypes</typeStatus>
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, same data as holotype:
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<collectionCode id="BE9F9FA4FFFFA42BFD48FB9DFC846F39" box="[762,835,1051,1077]" pageId="53" pageNumber="54">CASC</collectionCode>
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||
(
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<specimenCount id="CE88CCE8FFFFA42BFCE0FB9DFC026F3A" box="[850,965,1051,1078]" pageId="53" pageNumber="54" type="worker">7 workers</specimenCount>
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)
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</materialsCitation>
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,
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<materialsCitation id="68E60D3CFFFFA42BFC65FB9DFB776F3A" box="[983,1200,1051,1078]" collectingDate="1951-02-25" collectionCode="MCZC" collectorName="E. S. Ross & A. E. Michelbacher" country="Bolivia" isTailEnumeration="true" location="La Paz" pageId="53" pageNumber="54" specimenCount="2" specimenCount-worker="2" stateProvince="La Paz" typeStatus="paratype">
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<collectionCode id="BE9F9FA4FFFFA42BFC65FB9DFBE16F39" box="[983,1062,1051,1077]" pageId="53" pageNumber="54">MCZC</collectionCode>
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(
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<specimenCount id="CE88CCE8FFFFA42BFB86FB9DFB6F6F3A" box="[1076,1192,1051,1078]" pageId="53" pageNumber="54" type="worker">2 workers</specimenCount>
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)
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</materialsCitation>
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,
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<materialsCitation id="68E60D3CFFFFA42BFB0BFB9DFA5F6F3A" box="[1209,1432,1051,1078]" collectionCode="MNHN" country="Bolivia" pageId="53" pageNumber="54" specimenCount="2" specimenCount-worker="2" stateProvince="La Paz" typeStatus="paratype">
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<collectionCode id="BE9F9FA4FFFFA42BFB0BFB9DFAC86F39" box="[1209,1295,1051,1077]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="53" pageNumber="54">MNHN</collectionCode>
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(
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<specimenCount id="CE88CCE8FFFFA42BFAAFFB9DFA566F3A" box="[1309,1425,1051,1078]" pageId="53" pageNumber="54" type="worker">2 workers</specimenCount>
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||
)
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</materialsCitation>
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||
,
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<materialsCitation id="68E60D3CFFFFA42BFF25FBB9FEAF6F56" box="[151,360,1087,1114]" collectionCode="RAJC" country="Bolivia" pageId="53" pageNumber="54" specimenCount="2" specimenCount-worker="2" stateProvince="La Paz" typeStatus="paratype">
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<collectionCode id="BE9F9FA4FFFFA42BFF25FBB9FF1C6F55" box="[151,219,1087,1113]" pageId="53" pageNumber="54">RAJC</collectionCode>
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(
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<specimenCount id="CE88CCE8FFFFA42BFF59FBB9FEA76F56" box="[235,352,1087,1114]" pageId="53" pageNumber="54" type="worker">2 workers</specimenCount>
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)
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</materialsCitation>
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,
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<materialsCitation id="68E60D3CFFFFA42BFEC0FBB9FD976F56" box="[370,592,1087,1114]" collectionCode="USNM" country="Bolivia" pageId="53" pageNumber="54" specimenCount="2" specimenCount-worker="2" stateProvince="La Paz" typeStatus="paratype">
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<collectionCode id="BE9F9FA4FFFFA42BFEC0FBB9FE046F55" box="[370,451,1087,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="53" pageNumber="54" type="Museum">USNM</collectionCode>
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(
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<specimenCount id="CE88CCE8FFFFA42BFE61FBB9FD8F6F56" box="[467,584,1087,1114]" pageId="53" pageNumber="54" type="worker">2 workers</specimenCount>
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)
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="D8310761FFFFA42BFF75FBE5FCC06E75" blockId="53.[151,1437,1014,2013]" pageId="53" pageNumber="54">
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<emphasis id="EAFADB73FFFFA42BFF75FBE5FE646F70" bold="true" box="[199,419,1122,1149]" pageId="53" pageNumber="54">Worker diagnosis.</emphasis>
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Workers of this species are uniquely characterized by the following combination of features: (1) first gastral tergum with striae along base to extending over anterior one-third to one-half of medial portion of segment, rest of tergum smooth and polished, strongly shining, (2) longitudinal rugae on dorsum of promesonotum irregular, continuing to anterior margin of pronotum, (3) mesosoma dark ferruginous orange to orangish-brown, (4) posterior corners of head rugose, (5) superior propodeal spines consist of short, triangular teeth with a length less than width at base, (6) height of inferior propodeal spines greater than length of superior propodeal spines, (7) mandibles with six teeth, (8) rugae on frontal triangle subparallel-parallel, continuing onto clypeus, and (9) rugae on dorsum of postpetiole mostly transverse (
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<figureCitation id="40B51BE4FFFFA42BFD3BFAD9FD3C6E75" box="[649,763,1375,1401]" captionStart="FIGURE 23" captionStartId="55.[151,250,1916,1940]" captionTargetBox="[418,1168,181,1892]" captionTargetId="figure-15@55.[418,1170,181,1892]" captionTargetPageId="55" captionText="FIGURE 23. Photographs of Pogonomyrmex bolivianus holotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0914125). Photographs by Michele Esposito from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.5499188" httpUri="https://zenodo.org/record/5499188/files/figure.png" pageId="53" pageNumber="54">Figure 23</figureCitation>
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).
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</paragraph>
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<paragraph id="D8310761FFFFA42BFF75FA05FD196D05" blockId="53.[151,1437,1014,2013]" pageId="53" pageNumber="54">
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<emphasis id="EAFADB73FFFFA42BFF75FA05FEB16E91" bold="true" box="[199,374,1411,1437]" pageId="53" pageNumber="54">Measurements</emphasis>
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—
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<typeStatus id="0735B9C3FFFFA42BFE21FA05FE336E91" box="[403,500,1411,1437]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
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(
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<emphasis id="EAFADB73FFFFA42BFDB7FA05FDD46E90" box="[517,531,1411,1436]" italics="true" pageId="53" pageNumber="54">n</emphasis>
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||
=
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<specimenCount id="CE88CCE8FFFFA42BFD81FA05FD0F6E91" box="[563,712,1411,1437]" pageId="53" pageNumber="54" type="generic" typeStatus="paratypes">12 paratypes</specimenCount>
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). HL 1.54 (1.43–1.61); HW 1.66 (1.53–1.69); MOD 0.32 (0.31– 0.35); OMD 0.42 (0.41–0.47); SL 1.16 (1.07–1.31); PNW 1.08 (1.06–1.18); HFL 1.57 (1.42–1.65); ML 1.87 (1.65– 1.95); PW 0.40 (0.37–0.43); PPW 0.55 (0.54–0.69). Indices: SI 69.88 (66.46–77.51); CI 107.79 (100.64–110.00); OI 19.28 (18.45–22.88); HFI 94.58 (89.38–101.90).
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</paragraph>
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<paragraph id="D8310761FFFFA42BFF75F994FE376CB5" blockId="53.[151,1437,1014,2013]" pageId="53" pageNumber="54">
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<emphasis id="EAFADB73FFFFA42BFF75F994FE966D20" bold="true" box="[199,337,1554,1580]" pageId="53" pageNumber="54">Description</emphasis>
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. Head quadrate to wider than long (CI = 100.64–110.00); posterior margin flat in full-face view. Longitudinal rugae on cephalic dorsum prominent; in full-face view, medial rugae diverging toward posterior corners of head. Rugae on frontal triangle subparallel-parallel, continuing onto clypeus. Interrugae on cephalic dorsum weakly to moderately granulate-punctate, weakly shining to shining; posterior corners rugose, interrugae mostly smooth and shining. Anterior margin of clypeus moderately concave, dorsum with numerous subparallel, longitudinal rugae. Numerous long, curved, bristle-like, yellowish macrochaetae project from anterior margin of clypeus and basolateral margin of mandibles. Mandibles with six teeth; mandibular dorsum coarsely rugose. MOD ranging from 0.19–0.25× HL. In profile, eyes situated slightly anterior to middle of head, OMD = 1.20–1.43× MOD. In fullface view, eyes protruding slightly beyond lateral margins of head. Antennal scapes (SI = 66.46–77.51) reaching posterior corners of head or failing to do so by less than length of basal funicular segment. Antennal scapes strongly striate, interstriae moderately granulate, dull to weakly shining; basal flange well developed with carinate margin. Psammophore well developed.
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</paragraph>
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<paragraph id="D8310761FFFFA428FF75F845FBD36AF9" blockId="53.[151,1437,1014,2013]" lastBlockId="54.[151,1437,151,1905]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">Mesosomal profile weakly to moderately convex; all mesosomal surfaces with prominent widely spaced, wavy to irregular rugae. In dorsal view, humeral shoulders of pronotum enlarged, dorsolateral margin weakly to moderately angulate. Promesonotal suture usually weakly to moderately impressed. Wavy to irregular, longitudinal rugae on mesonotum continue onto pronotum, medial rugae continue to anterior margin of pronotum, lateral rugae diverging to humeral shoulders of pronotum. Rugae on pronotal sides irregular, traversing ventrally or posteroventrally or weakly rugoreticulate. Irregular rugae on mesopleura angle posterodorsally; irregular transverse or oblique rugae on dorsum of propodeum traverse ventrally or anteroventrally on sides. Superior propodeal spines consist of short, triangular, acuminate teeth, spines connected by a weak to well defined keel. Inferior propodeal spines moderately well developed, wider than high, triangular, tips rounded to broadly rounded, height greater than length of superior spines. Propodeal spiracles narrowly ovate facing posterad. Interrugae on mesosoma weakly granulate-punctate, weakly shining to smooth and shining. Legs weakly coriarious, weakly shining.</paragraph>
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<paragraph id="D8310761FFFCA428FF75FE79FE066851" blockId="54.[151,1437,151,1905]" pageId="54" pageNumber="55">Peduncle of petiole about as long as petiolar node, anteroventral margin with poorly developed rounded process. In profile, posterior surface of petiolar node weakly convex; node asymmetrical with anterior surface longer than posterior surface, apex subangulate to weakly rounded. In dorsal view, petiolar node longer than wide, sides subparallel, narrowest posterad, anterior margin spatulate to broadly rounded. Sides and posterior surface of petiolar node with regular to wavy transverse to occasionally oblique rugae. Dorsum of postpetiole convex in profile; in dorsal view, widest near posterior margin, narrowing to anterior margin, width and length similar. Transverse rugae on dorsum of postpetiole finer, denser than those on posterior surface of petiolar node, interrugae on both surfaces weakly granulate, weakly shining to smooth and strongly shining. Striae extending over anterior one-third to onehalf of medial portion of first gastral tergum, weakly shining with a silky appearance, posterior portion smooth and polished, strongly shining.</paragraph>
|
||
<paragraph id="D8310761FFFCA428FF75FCE1FD786F39" blockId="54.[151,1437,151,1905]" pageId="54" pageNumber="55">
|
||
Erect, mostly short to medium length, whitish pilosity moderately abundant on head, few hairs> 0.4–0.5× MOD. Moderately abundant suberect to semidecumbent pilosity on scape; abundant decumbent hairs on funicular segments. Legs with moderately abundant suberect to decumbent whitish to pale yellow setae. Rest of body with moderately dense, erect, medium-length, whitish to pale yellow setae. Head and mesosoma dark ferruginous orange to orangish-brown, mesosoma sometimes slightly darker than head; mandibles, antennae, legs, petiolar node, postpetiole, gaster dark brownish-orange (
|
||
<figureCitation id="40B51BE4FFFCA428FDF2FB9DFD756F39" box="[576,690,1051,1077]" captionStart="FIGURE 23" captionStartId="55.[151,250,1916,1940]" captionTargetBox="[418,1168,181,1892]" captionTargetId="figure-15@55.[418,1170,181,1892]" captionTargetPageId="55" captionText="FIGURE 23. Photographs of Pogonomyrmex bolivianus holotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0914125). Photographs by Michele Esposito from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.5499188" httpUri="https://zenodo.org/record/5499188/files/figure.png" pageId="54" pageNumber="55">Figure 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D8310761FFFCA428FF75FBB8FE5E6F55" blockId="54.[151,1437,151,1905]" box="[199,409,1086,1113]" pageId="54" pageNumber="55">
|
||
<emphasis id="EAFADB73FFFCA428FF75FBB8FEDB6F54" bold="true" box="[199,284,1086,1112]" pageId="54" pageNumber="55">Queen.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="D8310761FFFCA428FF75FBE5FE4F6F71" blockId="54.[151,1437,151,1905]" box="[199,392,1123,1149]" pageId="54" pageNumber="55">
|
||
<emphasis id="EAFADB73FFFCA428FF75FBE5FECC6F71" bold="true" box="[199,267,1123,1149]" pageId="54" pageNumber="55">Male.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="D8310761FFFCA428FF75FB00FDB86FAD" blockId="54.[151,1437,151,1905]" box="[199,639,1158,1185]" pageId="54" pageNumber="55">
|
||
<emphasis id="EAFADB73FFFCA428FF75FB00FDF36FAC" bold="true" box="[199,564,1158,1184]" pageId="54" pageNumber="55">Additional material examined.</emphasis>
|
||
None.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="909454EAFFFCA428FF75FB2DFEC96FE5" pageId="54" pageNumber="55" type="etymology">
|
||
<paragraph id="D8310761FFFCA428FF75FB2DFEC96FE5" blockId="54.[151,1437,151,1905]" pageId="54" pageNumber="55">
|
||
<emphasis id="EAFADB73FFFCA428FF75FB2DFE886FC9" bold="true" box="[199,335,1195,1221]" pageId="54" pageNumber="55">Etymology.</emphasis>
|
||
The specific epithet,
|
||
<taxonomicName id="1F8E7CE2FFFCA428FD8EFB2DFD766FC9" authorityName="Johnson" authorityYear="2021" box="[572,689,1195,1221]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis id="EAFADB73FFFCA428FD8EFB2DFD766FC9" box="[572,689,1195,1221]" italics="true" pageId="54" pageNumber="55">bolivianus</emphasis>
|
||
</taxonomicName>
|
||
, is derived from Ross & Michelbacher collecting the
|
||
<typeStatus id="0735B9C3FFFCA428FAB2FB2DFA906FC9" box="[1280,1367,1195,1221]" pageId="54" pageNumber="55" type="syntype">syntype</typeStatus>
|
||
series in
|
||
<collectingCountry id="A09947F1FFFCA428FF06FB49FECD6FE5" box="[180,266,1231,1257]" name="Bolivia" pageId="54" pageNumber="55">Bolivia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="909454EAFFFCA428FF75FB74FDC46DB1" pageId="54" pageNumber="55" type="discussion">
|
||
<paragraph id="D8310761FFFCA428FF75FB74FDC46DB1" blockId="54.[151,1437,151,1905]" pageId="54" pageNumber="55">
|
||
<emphasis id="EAFADB73FFFCA428FF75FB74FE8B6E00" bold="true" box="[199,332,1266,1292]" pageId="54" pageNumber="55">Discussion.</emphasis>
|
||
<taxonomicName id="1F8E7CE2FFFCA428FEE7FB75FD446E01" authorityName="Johnson" authorityYear="2021" box="[341,643,1267,1293]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis id="EAFADB73FFFCA428FEE7FB75FD446E01" box="[341,643,1267,1293]" italics="true" pageId="54" pageNumber="55">Pogonomyrmex bolivianus</emphasis>
|
||
</taxonomicName>
|
||
is not known to co-occur with any congeners, but note that very few series of
|
||
<taxonomicName id="1F8E7CE2FFFCA428FF4FFA91FE6A6E3C" box="[253,429,1303,1328]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EAFADB73FFFCA428FF4FFA91FE6A6E3C" box="[253,429,1303,1328]" italics="true" pageId="54" pageNumber="55">Pogonomyrmex</emphasis>
|
||
</taxonomicName>
|
||
have been collected from the Bolivian altiplano. Regardless,
|
||
<taxonomicName id="1F8E7CE2FFFCA428FBD6FA91FB3F6E3D" authorityName="Johnson" authorityYear="2021" box="[1124,1272,1303,1329]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis id="EAFADB73FFFCA428FBD6FA91FBB26E3C" box="[1124,1141,1303,1328]" italics="true" pageId="54" pageNumber="55">P</emphasis>
|
||
.
|
||
<emphasis id="EAFADB73FFFCA428FB31FA91FB3F6E3D" box="[1155,1272,1303,1329]" italics="true" pageId="54" pageNumber="55">bolivianus</emphasis>
|
||
</taxonomicName>
|
||
is easily diagnosed by longitudinal striae that cover only the anterior one-third to one-half of the first gastral tergum with the rest of the tergum smooth and polished, strongly shining.
|
||
<taxonomicName id="1F8E7CE2FFFCA428FD57FAD9FC3D6E74" authorityName="Johnson" authorityYear="2021" box="[741,1018,1375,1400]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="atacama">
|
||
<emphasis id="EAFADB73FFFCA428FD57FAD9FC3D6E74" box="[741,1018,1375,1400]" italics="true" pageId="54" pageNumber="55">Pogonomyrmex atacama</emphasis>
|
||
</taxonomicName>
|
||
is the only species with this character that is likely to be confused with
|
||
<taxonomicName id="1F8E7CE2FFFCA428FDBFFA05FD5A6E91" authorityName="Johnson" authorityYear="2021" box="[525,669,1411,1437]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis id="EAFADB73FFFCA428FDBFFA05FD5A6E91" box="[525,669,1411,1437]" italics="true" pageId="54" pageNumber="55">P. bolivianus</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="1F8E7CE2FFFCA428FD78FA05FC9D6E91" authorityName="Johnson" authorityYear="2021" box="[714,858,1411,1437]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis id="EAFADB73FFFCA428FD78FA05FC9D6E91" box="[714,858,1411,1437]" italics="true" pageId="54" pageNumber="55">P. bolivianus</emphasis>
|
||
</taxonomicName>
|
||
: (1) the posterior corners of head rugose, (2) longitudinal rugae on dorsum of promesonotum irregular, continuing to anterior margin of pronotum, (3) mandibles with six teeth, (4) rugae on dorsum of postpetiole mostly transverse, and (5) rugae on frontal triangle subparallel-parallel, continuing onto clypeus. In
|
||
<taxonomicName id="1F8E7CE2FFFCA428FE6AFA69FD926D04" authorityName="Johnson" authorityYear="2021" box="[472,597,1519,1544]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="atacama">
|
||
<emphasis id="EAFADB73FFFCA428FE6AFA69FD926D04" box="[472,597,1519,1544]" italics="true" pageId="54" pageNumber="55">P. atacama</emphasis>
|
||
</taxonomicName>
|
||
: (1) the posterior corners of head smooth and shining to strongly shining for most to all workers in a series, (2) mandibles with seven teeth, (3) longitudinal rugae on dorsum of mesonotum regular; anterior margin of pronotum usually with one to few transverse rugae, (4) rugae on dorsum of postpetiole mostly longitudinal, and (5) one to few rugae on frontal triangle converging anterad forming a narrowly rounded V-shaped pattern.
|
||
<taxonomicName id="1F8E7CE2FFFCA428FED3F9F9FD466D94" authorityName="Johnson" authorityYear="2021" box="[353,641,1663,1688]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cusquena">
|
||
<emphasis id="EAFADB73FFFCA428FED3F9F9FD466D94" box="[353,641,1663,1688]" italics="true" pageId="54" pageNumber="55">Pogonomyrmex cusquena</emphasis>
|
||
</taxonomicName>
|
||
is another congener from high elevation habitats, but
|
||
<taxonomicName id="1F8E7CE2FFFCA428FB67F9F9FA9A6D94" authorityName="Johnson" authorityYear="2021" box="[1237,1373,1663,1689]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cusquena">
|
||
<emphasis id="EAFADB73FFFCA428FB67F9F9FB216D94" box="[1237,1254,1663,1688]" italics="true" pageId="54" pageNumber="55">P</emphasis>
|
||
.
|
||
<emphasis id="EAFADB73FFFCA428FB41F9F9FA9A6D94" box="[1267,1373,1663,1688]" italics="true" pageId="54" pageNumber="55">cusquena</emphasis>
|
||
</taxonomicName>
|
||
lacks striae on the first gastral tergum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="909454EAFFFCA428FF75F940FC2C6C7D" pageId="54" pageNumber="55" type="biology_ecology">
|
||
<paragraph id="D8310761FFFCA428FF75F940FC2C6C7D" blockId="54.[151,1437,151,1905]" pageId="54" pageNumber="55">
|
||
<emphasis id="EAFADB73FFFCA428FF75F940FEE06DEC" bold="true" box="[199,295,1734,1760]" pageId="54" pageNumber="55">Biology.</emphasis>
|
||
Nothing is known about the biology of
|
||
<taxonomicName id="1F8E7CE2FFFCA428FD5BF941FCBD6DED" authorityName="Johnson" authorityYear="2021" box="[745,890,1735,1761]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis id="EAFADB73FFFCA428FD5BF941FCBD6DED" box="[745,890,1735,1761]" italics="true" pageId="54" pageNumber="55">P. bolivianus</emphasis>
|
||
</taxonomicName>
|
||
. The collection locale for the
|
||
<typeStatus id="0735B9C3FFFCA428FB79F941FAE56DED" box="[1227,1314,1735,1761]" pageId="54" pageNumber="55" type="syntype">syntype</typeStatus>
|
||
series was given only as
|
||
<quantity id="1F76AA84FFFCA428FE84F96CFEB06C09" box="[310,375,1770,1797]" metricMagnitude="4" metricUnit="m" metricValue="7.242048" pageId="54" pageNumber="55" unit="mi" value="45.0">45 mi</quantity>
|
||
south of
|
||
<collectingRegion id="1A4AC983FFFCA428FE50F96DFDF56C09" box="[482,562,1771,1797]" country="Bolivia" name="La Paz" pageId="54" pageNumber="55">La Paz</collectingRegion>
|
||
, which is within the Bolivian altiplano at an elevation that likely ranges from
|
||
<quantity id="1F76AA84FFFCA428FF25F889FEF06C25" box="[151,311,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="4.09" metricValueMax="4.24" metricValueMin="3.94" pageId="54" pageNumber="55" unit="m" value="4090.0" valueMax="4240.0" valueMin="3940.0">3940–4240 m</quantity>
|
||
. It seems likely that
|
||
<taxonomicName id="1F8E7CE2FFFCA428FD94F889FD7E6C25" authorityName="Johnson" authorityYear="2021" box="[550,697,1807,1833]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis id="EAFADB73FFFCA428FD94F889FD7E6C25" box="[550,697,1807,1833]" italics="true" pageId="54" pageNumber="55">P. bolivianus</emphasis>
|
||
</taxonomicName>
|
||
is restricted to altiplano habitats of
|
||
<collectingCountry id="A09947F1FFFCA428FBE5F889FB6E6C25" box="[1111,1193,1807,1833]" name="Bolivia" pageId="54" pageNumber="55">Bolivia</collectingCountry>
|
||
and probably nearby areas of
|
||
<collectingCountry id="A09947F1FFFCA428FF4AF8B5FEE86C41" box="[248,303,1843,1869]" name="Peru" pageId="54" pageNumber="55">Peru</collectingCountry>
|
||
.
|
||
<taxonomicName id="1F8E7CE2FFFCA428FE88F8B5FDA06C41" authorityName="Johnson" authorityYear="2021" box="[314,615,1843,1869]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis id="EAFADB73FFFCA428FE88F8B5FDA06C41" box="[314,615,1843,1869]" italics="true" pageId="54" pageNumber="55">Pogonomyrmex bolivianus</emphasis>
|
||
</taxonomicName>
|
||
is only known from the type locality, which appears to be in the Central Andean dry puna ecoregion, as defined by
|
||
<bibRefCitation id="BC1F7A90FFFCA428FDCAF8D1FC8C6C7D" author="Olson, D. M. & Dinerstein, E. & Wikramanayake, E. D. & Burgess, N. D. & Powell, G. V. N. & Underwood, E. C. & D'Amico, J. A. & Itoua, I. & Strand, H. E. & Morrison, J. C. & Loucks, C. J. & Allnutt, T. F. & Ricketts, T. H. & Kura, Y. & Lamoreux, J. F. & Wettengel, W. W. & Hedao, P. & Kassem, K. R." box="[632,843,1879,1905]" pageId="54" pageNumber="55" pagination="933 - 938" refId="ref148173" refString="Olson, D. M., Dinerstein, E., Wikramanayake, E. D., Burgess, N. D., Powell, G. V. N., Underwood, E. C., D'Amico, J. A., Itoua, I., Strand, H. E., Morrison, J. C., Loucks, C. J., Allnutt, T. F., Ricketts, T. H., Kura, Y., Lamoreux, J. F., Wettengel, W. W., Hedao, P. & Kassem, K. R. (2001) Terrestrial ecoregions of the world: a new map of life on Earth. Bioscience, 51 (11), 933 - 938. https: // doi. org / 10.1641 / 0006 - 3568 (2001) 051 [0933: TEOTWA] 2.0. CO; 2" type="journal article" year="2001">
|
||
Olson
|
||
<emphasis id="EAFADB73FFFCA428FD70F8D1FD346C7D" box="[706,755,1879,1905]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
|
||
. (2001)
|
||
</bibRefCitation>
|
||
(
|
||
<figureCitation id="40B51BE4FFFCA428FCE8F8D1FC196C7D" box="[858,990,1879,1905]" captionStart="FIGURE 22" captionStartId="51.[151,250,1915,1939]" captionTargetBox="[423,1160,181,1891]" captionTargetId="figure-15@51.[423,1165,181,1891]" captionTargetPageId="51" captionText="FIGURE 22. Geographic distribution of: (A) Pogonomyrmex atratus, (B) P. bolivianus, and (C) P. carbonarius. The large black circle in each panel denotes the type locality. See text regarding the type locality for P. atratus." figureDoi="http://doi.org/10.5281/zenodo.5499186" httpUri="https://zenodo.org/record/5499186/files/figure.png" pageId="54" pageNumber="55">Figure 22B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |