481 lines
50 KiB
XML
481 lines
50 KiB
XML
<document ID-CLB-Dataset="8199" ID-DOI="http://dx.doi.org/10.3897/phytokeys.172.62511" ID-GBIF-Dataset="e3a1f979-8f8b-4095-8027-1959bcd2096e" ID-PMC="PMC7895818" ID-Pensoft-Pub="1314-2003-172-75" ID-Pensoft-UUID="FDAE8D4147695752AAF24B45EA7067B0" ID-PubMed="33633487" ModsDocID="1314-2003-172-75" checkinTime="1613177897904" checkinUser="pensoft" docAuthor="Xu, Ke-Wang, Wang, Lu-Lu & Zhang, Li-Bing" docDate="2021" docId="2F8AFE4EFBFB5A1087EFEB580AFE2515" docLanguage="en" docName="PhytoKeys 172: 75-91" docOrigin="PhytoKeys 172" docPubDate="2021-02-12" docSource="http://dx.doi.org/10.3897/phytokeys.172.62511" docTitle="Asplenium wrightii Eaton ex Hooker, Sp. Fil. 3: 113. pl. 182. 1860." docType="treatment" docVersion="6" id="FDAE8D4147695752AAF24B45EA7067B0" lastPageNumber="75" masterDocId="FDAE8D4147695752AAF24B45EA7067B0" masterDocTitle="Taxonomic revision of the Asplenium wrightii complex (Aspleniaceae) with reinstatement of A. alatulum and A. subcrenatum" masterLastPageNumber="91" masterPageNumber="75" pageNumber="75" updateTime="1732828702414" updateUser="ExternalLinkService">
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<mods:mods id="A3B3CA2F64A5FB84338780A4AB95E401" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5ED6B19BECBF5DC796EECEB74F829869">
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<mods:title id="EC62779618BE7E4FFE3561B529D7FF97">Taxonomic revision of the Asplenium wrightii complex (Aspleniaceae) with reinstatement of A. alatulum and A. subcrenatum</mods:title>
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<mods:name id="C8B0FD730A59020CD1F23F841185B9AC" type="personal">
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<mods:roleTerm id="8926EBC146A9ECF9C3A412E062E953A4">Author</mods:roleTerm>
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<mods:namePart id="6024589A047B7CDDE047F37CCE8A13BD">Xu, Ke-Wang</mods:namePart>
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<mods:nameIdentifier id="943D56B84A82C99B01918A3F90025456" type="ORCID">https://orcid.org/0000-0003-2239-0487</mods:nameIdentifier>
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<mods:affiliation id="189CDAC77C8EDD9D97011D1A2DBE7634">Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University, Nanjing; 510275, China</mods:affiliation>
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<mods:nameIdentifier id="A9E0A119F8C5A871EBB6666B74619E95" type="email">xukw10@njfu.edu.cn</mods:nameIdentifier>
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<mods:name id="7B94BD8DD4B1BDCB6E297FBBF5C92A69" type="personal">
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<mods:roleTerm id="E878BC1DA242043E9D3EBD21D1A67DC9">Author</mods:roleTerm>
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<mods:namePart id="D19831A104F691531374F98358E0A84F">Wang, Lu-Lu</mods:namePart>
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<mods:affiliation id="C1337341EE08088317B12E0031B4826D">Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University, Nanjing; 510275, China</mods:affiliation>
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<mods:roleTerm id="F45F7B0FF687FD910A72473BA5ADCBEE">Author</mods:roleTerm>
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<mods:namePart id="2B643E0A08C0564B3E28ACAD39F9961B">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="2D657A1D3AA4BD75836CA3C7CC77720B">Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166 - 0299, USA</mods:affiliation>
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<mods:nameIdentifier id="8E59CA072E2E10F3180FE6EA2FB28AB0" type="email">libing.zhang@mobot.org</mods:nameIdentifier>
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<mods:typeOfResource id="13AE20DF8A67880002A594345AC02853">text</mods:typeOfResource>
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<mods:titleInfo id="8FEF623F846323494AAE833EF65E73CD">
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<mods:title id="910EBDF42D0A7994FCDDEEB0FF00921A">PhytoKeys</mods:title>
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<mods:part id="495DA159A06F157B186D0A4766229CDE">
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<mods:date id="F4816284458226873B472F833EEA861D">2021</mods:date>
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<mods:number id="C650A3C3DCDB89E1A999797521E0EAC7">2021-02-12</mods:number>
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<mods:detail id="3393B79FF5A36EF44BA7F1338053AB26" type="volume">
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<mods:number id="8C45350E29B581FB42B41CE43D75D664">172</mods:number>
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<mods:start id="50619ECF64DAB415E4DEA6F35B04244B">75</mods:start>
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<mods:end id="41B88AB32773F09469C68441E1AABDB6">91</mods:end>
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<mods:url id="DD4D005EC43CE525D85D49459BBBC396">http://dx.doi.org/10.3897/phytokeys.172.62511</mods:url>
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<mods:classification id="6D24E925F40627BAE591E2B7C066B782">journal article</mods:classification>
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<mods:identifier id="626729858000CDD468DEE4F450B753C3" type="DOI">http://dx.doi.org/10.3897/phytokeys.172.62511</mods:identifier>
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<mods:identifier id="6568608109409E00924637FA1A9BECF8" type="Pensoft-Pub">1314-2003-172-75</mods:identifier>
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<mods:identifier id="86CB0C5F981A1A7385827B29B540D03C" type="Pensoft-UUID">FDAE8D4147695752AAF24B45EA7067B0</mods:identifier>
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</mods:mods>
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<subSection id="D8E0AD099C954E1BDA153953AA87F331" pageId="0" pageNumber="75" type="results">
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<treatment id="2F8AFE4EFBFB5A1087EFEB580AFE2515" ID-GBIF-Taxon="177383158" LSID="urn:lsid:plazi:treatment:2F8AFE4EFBFB5A1087EFEB580AFE2515" httpUri="http://treatment.plazi.org/id/2F8AFE4EFBFB5A1087EFEB580AFE2515" lastPageNumber="75" pageId="0" pageNumber="75" scope_family="Aspleniaceae">
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<subSubSection id="D38CB0D5C8C62D91B86025D1898B7046" pageId="0" pageNumber="75" type="nomenclature">
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<paragraph id="BD3B4622C60406820417289DA5DC08E8" pageId="0" pageNumber="75">
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<taxonomicName id="5A5040BB7836BD920AE98A513E458D81" ID-CoL="HLTM" LSID="2F8AFE4E-FBFB-5A10-87EF-EB580AFE2515" authority="Eaton ex Hooker, Sp. Fil. 3: 113. pl. 182. 1860." authorityName="Eaton ex Hooker, Sp. Fil. 3: 113. pl. 182. 1860." class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="wrightii">Asplenium wrightii Eaton ex Hooker, Sp. Fil. 3: 113. pl. 182. 1860.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9DAE5A22A399CD5433D713510142EC0" pageId="0" pageNumber="75" type="reference_group">
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<paragraph id="686C9FF347C1E5EFA22F2DCEC184A756" pageId="0" pageNumber="75">
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<taxonomicName id="63DBE98A9917378BE2397CA83DCFC1D5" authorityName="Ching ex S.H.Wu" authorityYear="1989" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium duplicatoserratum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="duplicatoserratum">Asplenium duplicatoserratum</taxonomicName>
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Ching ex S.H. Wu, Bull. Bot. Res., Harbin 9(2): 86, f. 4. 1989.
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</paragraph>
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<paragraph id="8267AE76A25307ABFE8A931597A88FFF" pageId="0" pageNumber="75">
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<taxonomicName id="6AE27FEAE664217619181040B957B0A8" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium fujianense" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="fujianense">Asplenium fujianense</taxonomicName>
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Ching ex S.H. Wu, Bull. Bot. Res., Harbin 9(2): 21, 88-89, f. 7. 1989.
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</paragraph>
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<paragraph id="934985A1E713057E9F31A7A6E0637389" pageId="0" pageNumber="75">
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<taxonomicName id="04F333ED82B2C8977DDA29CA3C72FAC9" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium laui" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="laui">Asplenium laui</taxonomicName>
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Ching, Acta Phytotax. Sin. 9(4): 360-361. 1964.
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</paragraph>
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<paragraph id="F793C13589689A014D2ADFED72349C4F" pageId="0" pageNumber="75">
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<taxonomicName id="E3BB7387DE1415B3A2BFC2B2CB1D4FA6" authorityName="Ching ex S.H.Wu" authorityYear="1989" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium neomultijugum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="neomultijugum">Asplenium neomultijugum</taxonomicName>
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Ching ex S. H. Wu, Bull. Bot. Res. 9 (2): 21, f. 6. 1989.
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</paragraph>
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<paragraph id="07FD445912D7689693D9806312C1A8EA" pageId="0" pageNumber="75">
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<taxonomicName id="93B0245296A5EB54BFC18DD336644550" authorityName="Ching" authorityYear="1964" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium pseudowrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="pseudowrightii">Asplenium pseudowrightii</taxonomicName>
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Ching, Acta Phytotax. Sin. 9(4): 360. 1964.
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</paragraph>
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<paragraph id="F0278F6B2604113F10D545F867ECC3AA" pageId="0" pageNumber="75">
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<taxonomicName id="B707289319C91AE96AA983204FCB4535" authorityName="Ching ex S.H.Wu" authorityYear="1989" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium serratissimum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="serratissimum">Asplenium serratissimum</taxonomicName>
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Ching, Bull. Bot. Res., Harbin 9(2): 87. 1989.
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</paragraph>
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<paragraph id="C64CC2C42AB8397D14B68B7C22011F0F" pageId="0" pageNumber="75">
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<taxonomicName id="A1A3314A0E6FEED7C020885175A65DDD" authorityName="Ching ex S.H.Wu" authorityYear="1989" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium taiwanense" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="taiwanense">Asplenium taiwanense</taxonomicName>
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Ching ex S.H. Wu, Bull. Bot. Res., Harbin 9(2): 85-86, f. 3. 1989.
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</paragraph>
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<paragraph id="63F4CE8522DF0A2714E96C0ADED03749" pageId="0" pageNumber="75">
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<taxonomicName id="D6095D380851C99C7DBFB68AF88379A1" authorityName="H.Christ" authorityYear="1902" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium wrightioides" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="wrightioides">Asplenium wrightioides</taxonomicName>
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Christ, Bull. Acad. Int.
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<normalizedToken id="7C7ED03B4D15922F62B2E48F3ABCC036" originalValue="Géogr">Geogr</normalizedToken>
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. Bot. 11(153-154): 238-239. 1902.
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</paragraph>
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</subSubSection>
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<subSubSection id="7FA64F60288BA2446E83627345D258AB" pageId="0" pageNumber="75" type="materials_examined">
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<paragraph id="E35224E8043B59E38D23BD07D98A7A68" pageId="0" pageNumber="75">Type.</paragraph>
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<paragraph id="0807BD8D61A56C79108D70A3D4B88548" pageId="0" pageNumber="75">
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Japan.
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<emphasis id="8B03C7E82BE8FE405011530EFE40A8C0" bold="true" pageId="0" pageNumber="75">Kagoshima</emphasis>
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: Ryukyu Islands, 1853-1856,
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<emphasis id="8799986514F7FEE6FA13E5A4DD4BEB28" italics="true" pageId="0" pageNumber="75">C. Wright, #s.n.</emphasis>
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(holotype: NY (NY-00128031 [image!], isotype: GH (GH00020612 [image!])). Fig.
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<figureCitation id="241A13BD2D48E3A08000E0C461DBBA5E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Type of Asplenium alatulum Ching (A barcode PE 00059412), A. subcrenatum Ching ex S. H. Wu (B barcode PE 00059494), A. wrightii Eaton ex Hooker (C barcode NY 00128031) and isotype of A. wrightii GH (D barcode GH 00020612)." figureDoi="10.3897/phytokeys.172.62511.figure3" httpUri="https://binary.pensoft.net/fig/509068" pageId="0" pageNumber="75">3C, D</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="AA02FEE82E450B38059C0A04A9148E6E" pageId="0" pageNumber="75" type="description">
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<paragraph id="E99575E33FF1F76424EFA1026BCEE522" pageId="0" pageNumber="75">Description.</paragraph>
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<paragraph id="BD6769C7D088D803F0708215186B02CD" pageId="0" pageNumber="75">
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Plants 35-70(-100) cm tall. Rhizome erect, short, scaly; scales brown to dark brown, lanceolate-triangular, 5-12
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<normalizedToken id="401689C7FE9D7C4B4A8D5C657D172050" originalValue="×">x</normalizedToken>
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0.6-1.2 mm, denticulate glandular margin or long fibrillose. Fronds tufted; stipe greyish-green to brown, (18-)23-29(-31) cm, base densely scaly, scales brown, lanceolate to linear-lanceolate, (4.6-)6.1-7.8(-8.8) mm long, with multicellular hairs, subglabrous towards apex; lamina ovate-lanceolate to lanceolate, (19-)39-70(-88)
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<normalizedToken id="6951A5BC7FF9D760860AF132BBB301FD" originalValue="×">x</normalizedToken>
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(9-)17-27(-35) cm, base truncate, apex acute, 1-pinnate; pinnae (12-)17-25(-34) pairs, basal pinnae subopposite, others alternate, at an angle of ca. 50°-60° to rachis, with stalks (2-)4-8 mm, lower pinnae slightly reduced, suprabasal pinnae narrowly oval-lanceolate and often falcate, (6-)9.1-13(-20)
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<normalizedToken id="3B8020A9188A1C82C80ED599397374B0" originalValue="×">x</normalizedToken>
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(0.9-)1.2-1.8(-2.5) cm, base asymmetrical, acroscopic side truncate at an angle of (40°-)55°-75°(-85°) to costa and often auriculate, basiscopic side cuneate, at an angle of (20°-)30°-40°(-60°), becoming decurrent on rachis in central part of lamina, margin serrate to coarsely dentate, apex acuminate. Veins (1 or)2-forked, with terminal hydathode (Fig.
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<figureCitation id="DE76CDFA6B251E2C7D2F192A12F671B0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Asplenium wrightii A Habitat and habit B Abaxial view of upper portion of lamina C Abaxial view of pinna showing the almost entire to crenate-sinuate margin D Adaxial view of upper portion of lamina E Upper portion of rachis showing the lateral wings." figureDoi="10.3897/phytokeys.172.62511.figure6" httpUri="https://binary.pensoft.net/fig/509071" pageId="0" pageNumber="75">6B, C, D</figureCitation>
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). Fronds papery, green to brownish-green when dry, subglabrous; rachis dull green to reddish-brown, terete abaxially, winged towards apex. Sori linear, (3-)6-10(-12) mm, on acroscopic veinlets, medial (Fig.
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<figureCitation id="DDD78B9558A4EE06E729629219082ED0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Asplenium wrightii A Habitat and habit B Abaxial view of upper portion of lamina C Abaxial view of pinna showing the almost entire to crenate-sinuate margin D Adaxial view of upper portion of lamina E Upper portion of rachis showing the lateral wings." figureDoi="10.3897/phytokeys.172.62511.figure6" httpUri="https://binary.pensoft.net/fig/509071" pageId="0" pageNumber="75">6B, C</figureCitation>
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); indusia brown, linear, papery, opening towards costa, persistent. Spores with average exospore length 32-45
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<normalizedToken id="A124EA887C67E9D04BFC91E6377FA9D1" originalValue="μm">μm</normalizedToken>
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, perispore cristato-alate.
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</paragraph>
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</subSubSection>
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<subSubSection id="0514692D3C434FED31F8829E50FCD01E" pageId="0" pageNumber="75" type="distribution">
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<paragraph id="12BED4A8CBD0F252A090F0301E71B53B" pageId="0" pageNumber="75">Distribution and habitat.</paragraph>
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<paragraph id="44E5E3CC3956275B1EC5C9B570198144" pageId="0" pageNumber="75">
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<taxonomicName id="A7DCE61B60D461A8E4736719943E218F" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium wrightii" order="Filicales" pageId="0" pageNumber="75" rank="species" species="wrightii">
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<emphasis id="E754EFAAE2998A391FB7AF29DD57B177" italics="true" pageId="0" pageNumber="75">Asplenium wrightii</emphasis>
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</taxonomicName>
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is commonly distributed in China and Japan and it might occur in Korea. This species is found in damp valleys under evergreen broad-leaved forests where it grows as a lithophyte by streams (Fig.
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<figureCitation id="A7430A3CB08DD00D2081049620682C98" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Asplenium wrightii A Habitat and habit B Abaxial view of upper portion of lamina C Abaxial view of pinna showing the almost entire to crenate-sinuate margin D Adaxial view of upper portion of lamina E Upper portion of rachis showing the lateral wings." figureDoi="10.3897/phytokeys.172.62511.figure6" httpUri="https://binary.pensoft.net/fig/509071" pageId="0" pageNumber="75">6A</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="0801B1D9A53458E865DABE9EDAB7FF7F" pageId="0" pageNumber="75" type="materials_examined">
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<paragraph id="C65BE1DEE1B55B2AD27070446642ACB5" pageId="0" pageNumber="75">Additional specimens examined.</paragraph>
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<paragraph id="AAAA8BAC62756303C2FFE295EB75FBD1" pageId="0" pageNumber="75">
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<emphasis id="8AA2D1A998FA90FCD1CBD681C3E2B389" bold="true" pageId="0" pageNumber="75">China. Anhui</emphasis>
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: Qimen County, Xifengsi, Nov. 1957,
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<emphasis id="AAB955FB764A4C1A0471AC782020B565" italics="true" pageId="0" pageNumber="75">M.B.Deng 5308</emphasis>
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(HZ-028581); Jing County, Dingxi Village, Suhong, 22 Sep 1959,
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<emphasis id="C8A7B3410565B981330D6E899906EC31" italics="true" pageId="0" pageNumber="75">0431</emphasis>
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(NA-S00153661); Tongling County, Xiaokeng, Jun 1985,
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<emphasis id="038D7AFF26E28381368C4B4A0B591AC5" italics="true" pageId="0" pageNumber="75">X.L.Liu 85053</emphasis>
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(HUST-00008926);
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<emphasis id="FAD204DF5CEB1B1D943833581F530DC3" bold="true" pageId="0" pageNumber="75">Chongqing</emphasis>
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: Pengshui County, Hanjia Village, elev. 370 m, 29°25'137.98"N,
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<geoCoordinate id="0C48FD58EC4C40EC6132753805BD44D7" degrees="108" direction="east" minutes="18" orientation="longitude" precision="1" seconds="19.1" value="108.305305">108°18'19.1"E</geoCoordinate>
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, 16 Oct 2012,
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<emphasis id="32F07ED28BF64F572DCF2EE215E7C949" italics="true" pageId="0" pageNumber="75">Pengshui expedition 500243-003-167</emphasis>
|
||
(IMC-0003680); Yangshan County, Nanling Nature Reserve, Xinerkeng, 29 Sep 2007,
|
||
<emphasis id="E47EC184FC29C7E18375EB345C5B5267" italics="true" pageId="0" pageNumber="75">X.L.Zhou & H.F.Chen 785</emphasis>
|
||
(HUST-00008767);
|
||
<emphasis id="CECB49AD0984897653153D39B7D072D5" bold="true" pageId="0" pageNumber="75">Fujian</emphasis>
|
||
: 22 Jul 2015,
|
||
<emphasis id="942E13D5EE84F34854858FE685D56062" italics="true" pageId="0" pageNumber="75">ZXL05538</emphasis>
|
||
(CSH-0101629); 10 Apr 2014,
|
||
<emphasis id="B8BC6A3740F6EBE1BA8A2CAD7E6B52A6" italics="true" pageId="0" pageNumber="75">H.Shang & Y.F.Gu SG134</emphasis>
|
||
(CSH-0034578); Wuyishan, elev. 750 m, 15 Aug 2011,
|
||
<emphasis id="EF38EF8ED8D16BEE79FD9A32E2357F28" italics="true" pageId="0" pageNumber="75">X.F.Zeng 11215</emphasis>
|
||
(CZH-0006554); Zhaoan County, Wushan, 12 Apr 2015,
|
||
<emphasis id="3D4B3804427D495F27B35FDB709ACB62" italics="true" pageId="0" pageNumber="75">X.F.Zeng ZXF20041</emphasis>
|
||
(CZH-0012412); Nanping City, Mangdangshan, 17 Feb 1999,
|
||
<emphasis id="3E1C2C79E40CE122B588A51E036A1E8A" italics="true" pageId="0" pageNumber="75">G.S.He 9643</emphasis>
|
||
(PE-00913318); Jianning County, 20 Nov 1977,
|
||
<emphasis id="BECEF3C3AB2BEF2B4F1D56B6FFF3C263" italics="true" pageId="0" pageNumber="75">Z.Y.Li 10619</emphasis>
|
||
(PE-0913321); Jiangle County, Longxishan, 27 May 1991,
|
||
<emphasis id="2C8A44994FC77D8E570AD73127A82F8E" italics="true" pageId="0" pageNumber="75">Longxishan expedition 0359</emphasis>
|
||
(PE-01555190);
|
||
<emphasis id="D00BD247F5AA54D50DB1C37BB4BBF719" bold="true" pageId="0" pageNumber="75">Guangxi</emphasis>
|
||
: Lingyun County, Yuhong Village, Donglan, elev. 1303 m,
|
||
<geoCoordinate id="16E620B4514095617293DDDD170F193C" degrees="24" direction="north" minutes="24" orientation="latitude" precision="1" seconds="13.19" value="24.403664">24°24'13.19"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2970A87E562F792676984D80E5986451" degrees="106" direction="east" minutes="29" orientation="longitude" precision="1" seconds="03.17" value="106.48421">106°29'03.17"E</geoCoordinate>
|
||
, 16 Aug 2013,
|
||
<emphasis id="7EE0113CA495CEFB4C4010A229DC3683" italics="true" pageId="0" pageNumber="75">Lingyun expedition 451027130816103</emphasis>
|
||
(GXMG-0117634); Longshen County, Heping Village, elev. 615 m,
|
||
<geoCoordinate id="CB0A55C1F8E7E34437D73BE18EC1988D" degrees="25" direction="north" minutes="41" orientation="latitude" precision="1" seconds="12.10" value="25.686693">25°41'12.10"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="76421387EB9944973681E1F672BD1FF5" degrees="110" direction="east" minutes="03" orientation="longitude" precision="1" seconds="22.40" value="110.05623">110°03'22.40"E</geoCoordinate>
|
||
, 6 Mar 2013,
|
||
<emphasis id="D7F0EA95F778A1D728567CBF19AC97B2" italics="true" pageId="0" pageNumber="75">Longshen expedition 450328130306038LY</emphasis>
|
||
(IBK-00362003); Xingan County, Maoershan, 5 Oct 2007,
|
||
<emphasis id="9E2AA23C8C263C913E67E8BCC53BFF6B" italics="true" pageId="0" pageNumber="75">L.Wu & X.X.Xu 1042</emphasis>
|
||
(HUST-00019436); Sanjiangdongzu County, Sanxingpo, Dudong Village, elev. 700 m, 18 Dec 2007,
|
||
<emphasis id="EC4F7BAA1B7110A03B5831587CA42E6E" italics="true" pageId="0" pageNumber="75">X.X.Xu & L.C.Qin 255</emphasis>
|
||
(HUST-00011550);
|
||
<emphasis id="364A2E92B4B1753586F7F341EAEC8735" bold="true" pageId="0" pageNumber="75">Guangdong</emphasis>
|
||
: Heping County, Daba Village, Dafukeng, 2 Jan 2007,
|
||
<emphasis id="E23BF50F087446C018DDF9ABFAF5E0E4" italics="true" pageId="0" pageNumber="75">C.M.Tan et al. Y06630</emphasis>
|
||
(HUST-00004651); Lechang County, 23 Nov 1931,
|
||
<emphasis id="7C65A12685AD6EF8B6D7ACEBF80EBFD7" italics="true" pageId="0" pageNumber="75">Z.Huang 31490</emphasis>
|
||
(IBK-00035278); the same locality, 21 Jun 1942,
|
||
<emphasis id="D9CEA81C7C3A37D75D4C7E6D32064D6A" italics="true" pageId="0" pageNumber="75">S.Q.Chen1611</emphasis>
|
||
(IBK-00035299); Boluo County, Luofushan, 18 Aug 1930,
|
||
<emphasis id="64430D8B4BB2316DC581CE7D59EC3A7C" italics="true" pageId="0" pageNumber="75">N.Q.Chen 41631</emphasis>
|
||
(IBK-00035279); Xinyi, 13 Aug 1931,
|
||
<emphasis id="AEBEC727C3C9D3A0E5C72B7F8ED0ED60" italics="true" pageId="0" pageNumber="75">Z.Huang 31164</emphasis>
|
||
(IBK-00035285); Dawuling, 4 Aug 2003, Y.H.Yue et al. 1568 (PE-01784537); 21 Mar 1931,
|
||
<emphasis id="9D53898581BF4E2EF7E55589DC3AF93C" italics="true" pageId="0" pageNumber="75">Z.Huang 31756</emphasis>
|
||
(PE-00913341); Ruyuan, Daxiagu, 2 Aug 2005,
|
||
<emphasis id="E01AED7644AB2A9B0A50248CBE23E214" italics="true" pageId="0" pageNumber="75">B.R.Liu 05100</emphasis>
|
||
(PE-01785865); Yingde, Shimentai Nature Reserve, Oct 2001,
|
||
<emphasis id="3CCF84B1E89A7CD1610C4A6918A89DED" italics="true" pageId="0" pageNumber="75">Y.H.Yue & F.W.Xing 13265</emphasis>
|
||
(PE-01784538); Mei County, Jiaying, 4 Aug 1932, Tsang,
|
||
<emphasis id="270DC3E4F6C26097F334EDA9F3CB18CD" italics="true" pageId="0" pageNumber="75">W.T. 21466</emphasis>
|
||
(PE-00913338);
|
||
<emphasis id="73639F697F5B6B3C229044BB35E03E94" bold="true" pageId="0" pageNumber="75">Guizhou</emphasis>
|
||
: Yongshun County, Xiaoxi Village, elev. 621 m, 28°82'21.10"N,
|
||
<geoCoordinate id="2E92FC4DB115423F2EFFD6F7A090F0CB" degrees="110" direction="east" minutes="25" orientation="longitude" precision="1" seconds="11.00" value="110.419716">110°25'11.00"E</geoCoordinate>
|
||
, 14 Jan 2014,
|
||
<emphasis id="375535D2EE157F48E5C7333F96DA6C63" italics="true" pageId="0" pageNumber="75">D.G.Zhang zdg9949</emphasis>
|
||
(JIU-04159); Liping County, Pingjia, elev. 670 m, 16 Dec 2007,
|
||
<emphasis id="D6B0F533E151D12F8C2B29AFB768DEC2" italics="true" pageId="0" pageNumber="75">X.X.Xu & L.C.Qin 349</emphasis>
|
||
(HUST-00011558); Libo County, Maolan Nature Reserve, 25 Apr 2015;
|
||
<emphasis id="0C32142233AB86B346BF2C3B5BA83A4D" italics="true" pageId="0" pageNumber="75">X.C.Zhang 7259</emphasis>
|
||
(PE-02051530); Shiqian County, Qiangyang Village, Longdong, 30 Jul 1988,
|
||
<emphasis id="2762A54DEAC48029D0E4EF8064C18F7A" italics="true" pageId="0" pageNumber="75">Wulingshan expedition 2355</emphasis>
|
||
(PE-01557785); Jiangkou County, 30 Aug 1986,
|
||
<emphasis id="50FF091CA7BF927029F54B46D491502E" italics="true" pageId="0" pageNumber="75">B.Bartholomew et al. 595</emphasis>
|
||
(PE-00913309); Jianhe County, Nanjiaqu, 30 Apr 1992,
|
||
<emphasis id="6454922CDF8ACCB57CF5381255C2BEBD" italics="true" pageId="0" pageNumber="75">F.Wang 651</emphasis>
|
||
(HGAS-055247); Jinping County, Gaodengpo, 27 Nov 1991,
|
||
<emphasis id="BDF24BC2A1CA7A4335D4E18948A25786" italics="true" pageId="0" pageNumber="75">fern expedition 91474</emphasis>
|
||
(HGAS-055248); Rongjiang County, Shuiwei Shuizu Village, 15 Oct 2014,
|
||
<emphasis id="2245F8CC833682E4DBD682D633A12771" italics="true" pageId="0" pageNumber="75">Wei et al. WYG036</emphasis>
|
||
(CSH-0043860); Shibing County, Maohao Village, 30 May 2016,
|
||
<emphasis id="FAA9E92D565EC5B6E8FA7D7CB0964EC9" italics="true" pageId="0" pageNumber="75">D.Y.Zhou 522623160530476LY</emphasis>
|
||
(GZTM-0066044); Danzhai County, 18 Oct 2012,
|
||
<emphasis id="8BFD725CFC2AEB933E68FE40322FA520" italics="true" pageId="0" pageNumber="75">Hou GZDZ201210180003</emphasis>
|
||
(GYBG-0009834);
|
||
<emphasis id="75E6AFAFBB80F75C063C8F0E47299045" bold="true" pageId="0" pageNumber="75">Hunan</emphasis>
|
||
: Sangzhi County, Bamaoxi Village, 21 Oct 2014,
|
||
<emphasis id="BE27ADCC3DEB8BC1CB42280D533BA582" italics="true" pageId="0" pageNumber="75">X.L.Zhou et al. ZXL09673</emphasis>
|
||
(CSH-0045103); Baojing County, Baiyunshan Nature Reserve, elev. 397 m,
|
||
<geoCoordinate id="9B5AAB4480043A51969C788232BC8CDD" degrees="28" direction="north" minutes="37" orientation="latitude" precision="1" seconds="51.90" value="28.631083">28°37'51.90"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="19131FFEE2C7F718169C872CBD325476" degrees="109" direction="east" minutes="17" orientation="longitude" precision="1" seconds="11.34" value="109.28648">109°17'11.34"E</geoCoordinate>
|
||
, 11 Aug 2012,
|
||
<emphasis id="1A3BEE14EC1699AC323D497A88DD5928" italics="true" pageId="0" pageNumber="75">X.J.Su. & H.B.Liu 433125D00030810086</emphasis>
|
||
(JIU-06613); Yongshun County, Xiaoxi, Daping, 12 Sep. 2009,
|
||
<emphasis id="27CB8E2B8B079AC1C804C24BB4C3E0B5" italics="true" pageId="0" pageNumber="75">L.Xu 090912005</emphasis>
|
||
(JIU-02238); Dongzu County, Pingyang Village, Yangdongtan, elev. 300 m, 10-15 Jan 2008,
|
||
<emphasis id="595EE22826FD44D19C42B477379B9027" italics="true" pageId="0" pageNumber="75">L.C.Qing & H.B.Ouyang 1080</emphasis>
|
||
(HUST-00012147); Shangyan, Sanxingpo, elev. 750 m, 14 Dec 2004,
|
||
<emphasis id="3E990968281982F7F840A3055D974396" italics="true" pageId="0" pageNumber="75">L.C.Qing. & X.X.Xu 691</emphasis>
|
||
(HUST-00011548); Suining County, Huangsang Nature Reserve, elev. 450 m, 10 Jan 2008,
|
||
<emphasis id="6857ABABF2E12DB47062E6BFD97C6FEB" italics="true" pageId="0" pageNumber="75">J.M.Xi & Y.B.Qin 07919</emphasis>
|
||
(HUST-00012150); Yanling County, Taoyuandong Nature Reserve, Jul 2008,
|
||
<emphasis id="DE5186DA65CED527EA6AD6FF5F03DF5D" italics="true" pageId="0" pageNumber="75">X.L.Zhou &Z.L.Zhu 2154</emphasis>
|
||
(PE-01964123); Rucheng County, Donggangling, Shuinishan, elev. 110 m, 31 Jul 2007,
|
||
<emphasis id="557529492AD6AA83C72CF7F8C2F96F0E" italics="true" pageId="0" pageNumber="75">L.Wu & H.B.Ouyang 88</emphasis>
|
||
(HUST-00008785); Pingjiang County, Sicun Village, Wudeng, elev. 500 m, 15 Dec 2007,
|
||
<emphasis id="A89BFD3F030283C0DA9F95E3E5345C26" italics="true" pageId="0" pageNumber="75">L.Wu & S.X.Qi W467</emphasis>
|
||
(HUST-00011555); Huaihua County, Bamianshan, 15 Dec 2007,
|
||
<emphasis id="D98BD9D03049B58AA54AF7B54AD7444E" italics="true" pageId="0" pageNumber="75">X.L.Zhou & Y.Q.Yao 1744</emphasis>
|
||
(HUST-00011562);
|
||
<emphasis id="EC8A0452A83A28CBD26C3F57F970FA5E" bold="true" pageId="0" pageNumber="75">Hubei</emphasis>
|
||
: Laifeng County, Dahe Village, Lianghekou, elev. 590 m, 25 Jul 2013,
|
||
<emphasis id="44DFECF4C82BA27C56642074D5DFB592" italics="true" pageId="0" pageNumber="75">W.Z.Zhu BLF077</emphasis>
|
||
(CCAU-0002976);
|
||
<emphasis id="C4F7F84CDA01AD7AA937D25B8426EEB7" bold="true" pageId="0" pageNumber="75">Jiangxi</emphasis>
|
||
: Xingzi County, Wenquan Village, Xiufeng, 26 Jul 2013,
|
||
<emphasis id="D92D56D0D6BDF79281400493877F7403" italics="true" pageId="0" pageNumber="75">A.M.Dong 2082</emphasis>
|
||
(JJF00010847); Wanzai County, Jiulong Forest Park, elev. 190 m,
|
||
<geoCoordinate id="AAD430AF32C8CCF69F3103B4638A7430" degrees="28" direction="north" minutes="21" orientation="latitude" precision="15" seconds="38" value="28.360556">28°21'38"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C7860AC6239139FD75F00AE3F6AB244A" degrees="114" direction="east" minutes="30" orientation="longitude" precision="15" seconds="47" value="114.513054">114°30'47"E</geoCoordinate>
|
||
, 15 Aug 2013,
|
||
<emphasis id="D9A5FB10EB030777CE301A32984B7239" italics="true" pageId="0" pageNumber="75">H.G.Ye & F.Y.Zeng LXP10-2161</emphasis>
|
||
(IBSC-0773610); Wuning County, Yuangkou Village, Dongkeng, elev. 400 m, 16 Oct 2012,
|
||
<emphasis id="7B377CDBEAA6B922222DFDF835926787" italics="true" pageId="0" pageNumber="75">J.H.Zhang 2045</emphasis>
|
||
(JJF-00010846); Chongyi County, Fengzhou Village, Changlongao, elev. 1000 m, 8 Jul 2007,
|
||
<emphasis id="D9E944DECADB291238BA4B0E9365617F" italics="true" pageId="0" pageNumber="75">L.Y.Wang W.181</emphasis>
|
||
(HUST-00008778); Jianggangshan, elev. 800 m, 10 Apr 2004,
|
||
<emphasis id="D5F162D11DEA411A327EFFA9BCDC6635" italics="true" pageId="0" pageNumber="75">Y.H.Yan & J.S.Zhou 3415</emphasis>
|
||
(HUST-00001166);
|
||
<emphasis id="151DC07170B7AA1A658B6D4DFBCC4328" bold="true" pageId="0" pageNumber="75">Jiangsu</emphasis>
|
||
: Yixing, Longchi, Shuizhushan, 1956,
|
||
<emphasis id="AF46C8E22E3E6722D03677F5F6A0DD5D" italics="true" pageId="0" pageNumber="75">Liu & Huang 2874</emphasis>
|
||
(NAS-00091223);
|
||
<emphasis id="D5EC8B612BF7D98895A9CA3F1A833AF7" bold="true" pageId="0" pageNumber="75">Sichuan</emphasis>
|
||
: Emeishan, 15 Sep 1963,
|
||
<emphasis id="4C300B874497325C3A7B4B864D87F648" italics="true" pageId="0" pageNumber="75">G.X.Xing & K.Y.Lang 1788</emphasis>
|
||
(PE-13307); the same locality, 21 Sep 1979,
|
||
<emphasis id="389AB0A4D4F4C6D20FF01817F6AC0FA0" italics="true" pageId="0" pageNumber="75">Z.R.Wang C122</emphasis>
|
||
(PE-96543); Zhulian County, 1 Jun 1978,
|
||
<emphasis id="D65EAD09E5111668BCA856EB83E3C044" italics="true" pageId="0" pageNumber="75">X.R.Kong 5204</emphasis>
|
||
(CDBI-003584);
|
||
<emphasis id="D018C53F459A930519CC8B8FB8780657" bold="true" pageId="0" pageNumber="75">Taiwan</emphasis>
|
||
: Wulai, Fushan-Hapen, 13 Oct 1984,
|
||
<emphasis id="9A261674F03E6444AC146E1E387A1EA1" italics="true" pageId="0" pageNumber="75">Y.Tateishi et al. 20492</emphasis>
|
||
(PE-01451621); Takao, 27 Jan 1939,
|
||
<emphasis id="F71B5AB587BE70ABD837E689F80A0679" italics="true" pageId="0" pageNumber="75">M.Tagawa 2049</emphasis>
|
||
(PE-00913326);
|
||
<emphasis id="ECFEB0B3EAA8F082928D837B6FB36D41" italics="true" pageId="0" pageNumber="75">W.Hancock 141</emphasis>
|
||
(PE-00913329); Taito, 4 Mar 1940,
|
||
<emphasis id="D6391C206A03AF88B202A8A0B92E3240" italics="true" pageId="0" pageNumber="75">M.Tagawa 3168</emphasis>
|
||
(PE-00913330); Huanlian County, Changliang, 8 Jun 2001,
|
||
<emphasis id="E57ECAED5B6EEBC83EA3BB270228A8BB" italics="true" pageId="0" pageNumber="75">T.C.Chen 11303</emphasis>
|
||
(PE-01451613); Taibei County, Lujiaokengxi, 28 Jan 2000,
|
||
<emphasis id="6804AAFF6C71712D0C5CC6AB15016199" italics="true" pageId="0" pageNumber="75">T.C.Chen 10393</emphasis>
|
||
(PE-01451614);
|
||
<emphasis id="4DDB030A530A2AB5B2B36FF611972711" bold="true" pageId="0" pageNumber="75">Zhejiang</emphasis>
|
||
: Tonglu County, Luci, Changzhou, 8 Nov 1989,
|
||
<emphasis id="043490BE35ABE3597A9D8E3AD2792BFF" italics="true" pageId="0" pageNumber="75">L.Hong s.n.</emphasis>
|
||
(HZ-028576); Shouchang Zhen, 19 Jun 1959,
|
||
<emphasis id="79F125736B395A052BB26EB9E5402E01" italics="true" pageId="0" pageNumber="75">Zhejiang expedition 27803</emphasis>
|
||
(HZ028578); Taishun County, Liguang, Huangshikeng, 27 Nov 1958,
|
||
<emphasis id="3AC9D9EF01C51D36BA5999FE6ECB6E73" italics="true" pageId="0" pageNumber="75">Anonymous 23818</emphasis>
|
||
(HZ-028579); Lishui, Suichang Nature Reserve, elev. 780 m,
|
||
<geoCoordinate id="E4091802AA248732A46E18E034A39303" degrees="25" direction="north" minutes="59" orientation="latitude" precision="15" seconds="11" value="25.98639">25°59'11"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="81ED5018F8D6995B2883FF55D30B2F88" degrees="116" direction="east" minutes="25" orientation="longitude" precision="15" seconds="11" value="116.419716">116°25'11"E</geoCoordinate>
|
||
, 5 Jun 2012,
|
||
<emphasis id="083A3030BBC6FC6014B2FC3056BD7A87" italics="true" pageId="0" pageNumber="75">Q. Tian TQ01773</emphasis>
|
||
(CSH-0002430); 25 Nov 2015,
|
||
<emphasis id="E1B9AC3B29F44CD81C88007FF790D80F" italics="true" pageId="0" pageNumber="75">H.J.Wei JSL3466</emphasis>
|
||
(CSH-0110219); 29 Jul 1958,
|
||
<emphasis id="ABA287D3B2CB9ECFFDF403E351DC5B7B" italics="true" pageId="0" pageNumber="75">Shan 5578</emphasis>
|
||
(PE-00913289);
|
||
</paragraph>
|
||
<caption id="A7F8B62E1BC856810C6D424E608DDE6D" doi="10.3897/phytokeys.172.62511.figure6" httpUri="https://binary.pensoft.net/fig/509071" pageId="0" pageNumber="75" start="Figure 6" startId="F6">
|
||
<paragraph id="60CCD87D6565E8BDF1499B663ACFE004" pageId="0" pageNumber="75">
|
||
<emphasis id="1F0250B9CAFC547748C353DE67E6C5E6" bold="true" pageId="0" pageNumber="75">Figure 6.</emphasis>
|
||
<taxonomicName id="7BD5B023D74681A740D47A6F937825BA" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium wrightii" order="Filicales" pageId="0" pageNumber="75" rank="species" species="wrightii">
|
||
<emphasis id="B94185A3D0F6340D9F92B50A8E0D4AA3" italics="true" pageId="0" pageNumber="75">Asplenium wrightii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B7CCACAB717E2E5840AB3DA87B628D7" bold="true" pageId="0" pageNumber="75">A</emphasis>
|
||
Habitat and habit
|
||
<emphasis id="87E5FCA07546BA07FBFD13DE12A0B9DA" bold="true" pageId="0" pageNumber="75">B</emphasis>
|
||
Abaxial view of upper portion of lamina
|
||
<emphasis id="6CFA96A68483B0A7B9DB7E852CD2EBA6" bold="true" pageId="0" pageNumber="75">C</emphasis>
|
||
Abaxial view of pinna showing the almost entire to crenate-sinuate margin
|
||
<emphasis id="BA2A751D240A535607DA1394BD279658" bold="true" pageId="0" pageNumber="75">D</emphasis>
|
||
Adaxial view of upper portion of lamina
|
||
<emphasis id="7B0917EF9562220B6D7421D981D6CA72" bold="true" pageId="0" pageNumber="75">E</emphasis>
|
||
Upper portion of rachis showing the lateral wings.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="11E95A8F9FD0B149307CC4709B61BD97" pageId="0" pageNumber="75">
|
||
<emphasis id="4FD17C6408E52B88A6355E47340D9C10" bold="true" pageId="0" pageNumber="75">Japan. Kyushu</emphasis>
|
||
: Chikushi-yabakei, Nakagawa-cho, Chikushi-gun, Fukuoka Pref., 27 Mar 1975,
|
||
<emphasis id="471E9400AFB6DF3092AE28D107A01706" italics="true" pageId="0" pageNumber="75">T.Nakaike et al. 105</emphasis>
|
||
(PE-01708808); Is. Yakushima (Kagoshima Pref.), from Kurio to Segiri through Ookonotaki, elev. 30-100 m, 16 Jul 1979,
|
||
<emphasis id="6A28C6C36670F6C78CFFFE19A132CCBD" italics="true" pageId="0" pageNumber="75">T.Yamazaki et al. 2327</emphasis>
|
||
(PE-01708809); Kagoshima Pref., Okuchi-shi, Jusso, 6 Sep 1959,
|
||
<emphasis id="83AD85919C5CD22ADF3CA1913F4E5535" italics="true" pageId="0" pageNumber="75">Tetsuji Yamanaka s.n.</emphasis>
|
||
(PE-01964369); Prefecture Kagoshina, Yakushina Island, Yaku-cho, Tainoko-gawa River, 10-11 Oct 1977,
|
||
<emphasis id="5AEC9EC76883B8EFA1270E5570A2D2FA" italics="true" pageId="0" pageNumber="75">Boufford, David Edward 20118</emphasis>
|
||
(MO-3136075); Along trail in gorge, foot of Mt. Awa. Motobu Peninsula, Okinawa Island, 10 Dec 1953,
|
||
<emphasis id="619F2CA78B7FC740DCF5BCC7A5FF6515" italics="true" pageId="0" pageNumber="75">Walker, EH 7614</emphasis>
|
||
(L-3508507);
|
||
<emphasis id="9E5746C78D4007D465BF64A02879BC66" bold="true" pageId="0" pageNumber="75">Honshu</emphasis>
|
||
: Owase City, Kuki, Namera Valley, 12 Nov 1991,
|
||
<emphasis id="21D3E45031B2A6D9A76A7C011CD363C5" italics="true" pageId="0" pageNumber="75">K.H.Shing 31</emphasis>
|
||
(PE-01708810); Pref. Mie,Kuki, Omse-shi, 5 Oct 1971,
|
||
<emphasis id="BA5C8C171D9F0AEF201797097F8FFFC3" italics="true" pageId="0" pageNumber="75">Kato, Masahiro-259</emphasis>
|
||
(MO-3136077); Pref. Schizouka: Aono, Minamiizu-cho, Kamo-gun, 30 Dec 1972,
|
||
<emphasis id="444346D7AB3E6A846376D6F14F50E7F3" italics="true" pageId="0" pageNumber="75">T.Nakaike 5</emphasis>
|
||
(L-3508506);
|
||
<emphasis id="EB06C7179C3925358084CAD46D6100FB" bold="true" pageId="0" pageNumber="75">Shikoku</emphasis>
|
||
: Nakaohsaka Ohsakadani, Nakatosa Town, Takaoka Distr. Kouchi Pref., elev. 60-80 m,
|
||
<geoCoordinate id="5CF58A2FC23D3EA2B17ECC4BEC8ADE5F" degrees="33" direction="north" minutes="18" orientation="latitude" precision="15" seconds="48" value="33.31333">33°18'48"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A5C3C0C203E3D7BC0DE6F2E61C5D0599" degrees="133" direction="east" minutes="23" orientation="longitude" precision="15" seconds="48" value="133.39667">133°23'48"E</geoCoordinate>
|
||
, 6 Jan 2012,
|
||
<emphasis id="E5DD3FF9CCC5F4B54E6FFF2E38CFF447" italics="true" pageId="0" pageNumber="75">Taku Miyazaki 1201164</emphasis>
|
||
(PE-02002704); Yatabe, Nakatosa Town, Takaoka District, Kouchi Pref., elev. 40-60 m, 33°15'70"N,
|
||
<geoCoordinate id="8235739EAA03DC24841E9B51018A1D73" degrees="133" direction="east" minutes="15" orientation="longitude" precision="15" seconds="37" value="133.26028">133°15'37"E</geoCoordinate>
|
||
, 3 Jan 2007,
|
||
<emphasis id="7D498D36C52DC51570A5B8E106D00D63" italics="true" pageId="0" pageNumber="75">T.Miyazaki 0701137</emphasis>
|
||
(PE-01963957); Takanoyama-rindou Yamauchi, Nakatosa Town, Takaoka Distr. Kochi Pref., elev. 60-90 m,
|
||
<geoCoordinate id="E481309535D1462EC843DD00AAD01E20" degrees="33" direction="north" minutes="17" orientation="latitude" precision="15" seconds="04" value="33.284443">33°17'04"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FA66B60C1C2AB33D844B3AD66D1CC6FC" degrees="133" direction="east" minutes="12" orientation="longitude" precision="15" seconds="18" value="133.205">133°12'18"E</geoCoordinate>
|
||
, 27 Mar 2009,
|
||
<emphasis id="2D04C0BDB94ED1DDEAED33EFA9FA27D7" italics="true" pageId="0" pageNumber="75">Taku Miyazaki 0903138</emphasis>
|
||
(PE-01963829); Ohkawauchi Kaminokae, Nakatosa Town, Takaoka District, Kouchi Pref., elev. 20 m, 33°16'67"N,
|
||
<geoCoordinate id="76AE3910EF4F7E647A91D94AD2E39395" degrees="133" direction="east" minutes="14" orientation="longitude" precision="15" seconds="15" value="133.2375">133°14'15"E</geoCoordinate>
|
||
, 29 Dec 2006,
|
||
<emphasis id="D88678D4725AF59AA9DB8029B3FA5559" italics="true" pageId="0" pageNumber="75">Taku Miyazaki 0701025</emphasis>
|
||
(PE-01962310).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F80F2831AB384B2C7AC180D9F9DBD7FC" pageId="0" pageNumber="75" type="notes">
|
||
<paragraph id="447829A9A2DFA2D035F42DBC09E7FF39" pageId="0" pageNumber="75">Note.</paragraph>
|
||
<paragraph id="9EC3F6B88EB5988F93AA033F5532CC6F" pageId="0" pageNumber="75">
|
||
<taxonomicName id="FE98B2E1D6A7611B592ED02ECE464B5B" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium wrightii" order="Filicales" pageId="0" pageNumber="75" rank="species" species="wrightii">
|
||
<emphasis id="142C349E886D43C5A7096EA97C8287E8" italics="true" pageId="0" pageNumber="75">Asplenium wrightii</emphasis>
|
||
</taxonomicName>
|
||
is described based on the type material from Japan and it is widely distributed in China. The morphology of this species is variable, which resulted in the taxonomic chaos amongst
|
||
<taxonomicName id="991CABB8636672D0FC5875F7F2801B65" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="DA619401D1821749E216FF6489A08EE1" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
and its closely-related species. Several ploidy levels (e.g. tetraploid, octoploid, decaploid and dodecaploid) have been reported for the
|
||
<taxonomicName id="3A09D079E981C533797D36F588122237" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="A7B79DF9112903651CBC87CF383FD355" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
complex, but no correlation was found between ploidy levels and morphological characters within the complex (
|
||
<bibRefCitation id="CBDDC4D941658CBE7A58E238E4214715" author="Mitui, K" journalOrPublisher="Shokubutsu Kenkyu Zasshi" pageId="0" pageNumber="75" pagination="105 - 110" refId="B14" refString="Mitui, K, 1967. Chromosome studies on Japanese ferns (3). Shokubutsu Kenkyu Zasshi 42: 105 - 110" title="Chromosome studies on Japanese ferns (3)." volume="42" year="1967">Mitui 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BC40CA3CC41ED5558D9BD1E38E1226CF" author="Wang, ZR" editor="Shing, KH" journalOrPublisher="China Science and Technology Press, Beijing" pageId="0" pageNumber="75" pagination="133 - 134" refId="B21" refString="Wang, ZR, 1988. A preliminary report on the cytology of some species of Asplenium from China. In: Shing, KH, Kramer, KU, Eds., Proceedings of International Symposium on Systematic Pteridology. China Science and Technology Press, Beijing: 133 - 134" title="A preliminary report on the cytology of some species of Asplenium from China." volumeTitle="Proceedings of International Symposium on Systematic Pteridology." year="1988">Wang 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E9E4E1E60B53943DAAD4D24CC9307B3B" author="Lin, YX" editor="Wu, ZY" journalOrPublisher="2 - 3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="75" pagination="267 - 316" refId="B12" refString="Lin, YX, Viane, R, 2013. Aspleniaceae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China (Vol. 2-3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis: 267 - 316" title="Aspleniaceae." volumeTitle="Flora of China (Vol." year="2013">Lin and Viane 2013</bibRefCitation>
|
||
).
|
||
<taxonomicName id="3175F65843701A69D666B7E1A4120F83" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium shikokianum" order="Filicales" pageId="0" pageNumber="75" rank="species" species="shikokianum">
|
||
Asplenium
|
||
<normalizedToken id="2A16FB2270AE0E44AEA5505B0E86EB7B" originalValue="×">x</normalizedToken>
|
||
shikokianum
|
||
</taxonomicName>
|
||
is a natural hexaploid hybrid between octoploid
|
||
<taxonomicName id="C20ED513F8FE16DF6613F8086EB5CB39" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="4DD43B4ED09A20EA81AB93352BCCA339" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
and tetraploid
|
||
<taxonomicName id="E74727E3E7BB329E7989D40C414E8630" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. ritoense" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="ritoense">
|
||
<emphasis id="05D50906C6DE030D3E10C6DDAC3B3F12" italics="true" pageId="0" pageNumber="75">A. ritoense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F845DA536BDEEC7E167621D7351E1E50" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium wangii" order="Filicales" pageId="0" pageNumber="75" rank="species" species="wangii">
|
||
Asplenium
|
||
<normalizedToken id="30D89D73BC9D6B65064B1813D09ADA93" originalValue="×">x</normalizedToken>
|
||
wangii
|
||
</taxonomicName>
|
||
is another hybrid between
|
||
<taxonomicName id="DF19939906A4F2250B398B0B0D7405EC" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="5B0395E832667CE69531497C15566D39" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="17530552A920684AEECFB5F30586EA89" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. bullatum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="bullatum">
|
||
<emphasis id="E575CCE25727D282A39A16A641C42356" italics="true" pageId="0" pageNumber="75">A. bullatum</emphasis>
|
||
</taxonomicName>
|
||
. These hybrids are usually uncommon in terms of their distributions where their parents grow together (
|
||
<bibRefCitation id="DBF30650067CE9BB9A9FCFE0D52F82AA" author="Kuo, CM" journalOrPublisher="Botanical Bulletin of Academia Sinica" pageId="0" pageNumber="75" pagination="109 - 111" refId="B11" refString="Kuo, CM, 1988. A new Asplenium hybrid from Taiwan. Botanical Bulletin of Academia Sinica 29: 109 - 111" title="A new Asplenium hybrid from Taiwan." volume="29" year="1988">Kuo 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="77AFAA12ADAE68F785E4A7B8C5CC45AC" author="Lin, YX" editor="Wu, ZY" journalOrPublisher="2 - 3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="75" pagination="267 - 316" refId="B12" refString="Lin, YX, Viane, R, 2013. Aspleniaceae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China (Vol. 2-3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis: 267 - 316" title="Aspleniaceae." volumeTitle="Flora of China (Vol." year="2013">Lin and Viane 2013</bibRefCitation>
|
||
). Due to hybridisation and polyploidisation, there are a large number of recognised species complexes with ambiguous boundaries between species in
|
||
<taxonomicName id="36B2CF7EA047FB99565C610751EBE5A0" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="family">Aspleniaceae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="BC4297BE8B2ED1C50458C08D2924BBEE" author="Reichstein, T" journalOrPublisher="Botanica Helvetica" pageId="0" pageNumber="75" pagination="89 - 139" refId="B19" refString="Reichstein, T, 1981. Hybrids in European Aspleniaceae (Pteridophyta). Botanica Helvetica 91: 89 - 139" title="Hybrids in European Aspleniaceae (Pteridophyta)." volume="91" year="1981">Reichstein 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="79CA3F538E3DAEAE9A86F068E985FC2A" DOI="https://doi.org/10.1093/aob/mcs202" author="Dyer, RJ" journalOrPublisher="Annals of Botany" pageId="0" pageNumber="75" pagination="1515 - 1529" refId="B5" refString="Dyer, RJ, Savolainen, V, Schneider, H, 2012. Apomixis and reticulate evolution in the Asplenium monanthes fern complex. Annals of Botany 110 (8): 1515 - 1529, DOI: https://doi.org/10.1093/aob/mcs202" title="Apomixis and reticulate evolution in the Asplenium monanthes fern complex." url="https://doi.org/10.1093/aob/mcs202" volume="110" year="2012">Dyer et al. 2012</bibRefCitation>
|
||
). More comprehensive taxonomic studies are still needed to elucidate the species delimitation in this species complex.
|
||
</paragraph>
|
||
<paragraph id="37A911D06113C0C91BF1A1CEA9ABB21D" pageId="0" pageNumber="75">
|
||
<taxonomicName id="0AFE8AC0063136338E97AD1007D737CB" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium wrightii" order="Filicales" pageId="0" pageNumber="75" rank="species" species="wrightii">
|
||
<emphasis id="FE5558FE875BF3FAEEE0249AC75322F3" italics="true" pageId="0" pageNumber="75">Asplenium wrightii</emphasis>
|
||
</taxonomicName>
|
||
was also documented in Vietnam (
|
||
<bibRefCitation id="77095E072BBDDD1CA3466871F59BB228" author="Pham, HH" journalOrPublisher="Nha Xuat Ban Tre, Ho Chi Minh" pageId="0" pageNumber="75" refId="B16" refString="Pham, HH, 1999. An illustrated flora of Vietnam (2nd ed., Vol. 1). Nha Xuat Ban Tre, Ho Chi Minh" title="An illustrated flora of Vietnam (2 nd ed., Vol. 1)." year="1999">Pham 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C2100F782F6BBD97AE7BA3FA77060513" author="Lin, YX" editor="Wu, ZY" journalOrPublisher="2 - 3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="75" pagination="267 - 316" refId="B12" refString="Lin, YX, Viane, R, 2013. Aspleniaceae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China (Vol. 2-3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis: 267 - 316" title="Aspleniaceae." volumeTitle="Flora of China (Vol." year="2013">Lin and Viane 2013</bibRefCitation>
|
||
). However, we checked specimens identified as
|
||
<taxonomicName id="DEF511DF1CA15DF82822A66F8BA12518" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="B43B9F3074BB9E71CBA33D0B159B4527" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
from Vietnam and found none of these specimens is true
|
||
<taxonomicName id="7CB6953CFB43886BC01BF0E3BD64D00F" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="AB6B6D1BC630E53D7FA679A86A2ACF56" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
. It is possible that the name
|
||
<taxonomicName id="4DD420DCE5E1FA9CDE88BFFAE7FDB32B" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="1A75ACB3B3B5B947DAEA8E55EFD46A34" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
has been used erroneously for species
|
||
<taxonomicName id="03F60C8D8F9084BB18A1AABB7B34AC09" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. subcrenatum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="subcrenatum">
|
||
<emphasis id="4FC1138B89ED28E99AD5532BF7567A79" italics="true" pageId="0" pageNumber="75">A. subcrenatum</emphasis>
|
||
</taxonomicName>
|
||
in Vietnam.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |