207 lines
24 KiB
XML
207 lines
24 KiB
XML
<document id="764AADD28E171A6948CF6D5C468F6D68" ID-CLB-Dataset="28679" ID-DOI="http://dx.doi.org/10.3897/zookeys.810.29824" ID-GBIF-Dataset="f67f7966-5859-4a73-8e51-18b05d559ba1" ID-PMC="PMC6308220" ID-Pensoft-Pub="1313-2970-810-19" ID-PubMed="30613172" ID-ZooBank="F67F5B77293D49D997D93E147A5B80C0" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-810-19" ModsDocOrigin="ZooKeys 810" ModsDocTitle="Enantiomorphs and taxonomy of three conchological species in flat-shelled snails Trichocathaica (Pulmonata, Camaenidae)" checkinTime="1545315431060" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, unyadi, Andras & Asami, Takahiro" docDate="2018" docId="2F2663F6B68ECD1B119F1E9EAF91CA1D" docLanguage="en" docName="ZooKeys 810: 19-44" docOrigin="ZooKeys 810" docSource="http://dx.doi.org/10.3897/zookeys.810.29824" docTitle="Trichocathaica vestita Pilsbry 1934, comb. n." docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="FFD9FF9B4C0AF709FFC2FF86FFD8FFD3" masterDocTitle="Enantiomorphs and taxonomy of three conchological species in flat-shelled snails Trichocathaica (Pulmonata, Camaenidae)" masterLastPageNumber="44" masterPageNumber="19" pageNumber="25" updateTime="1701384931438" updateUser="ExternalLinkService">
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<mods:title id="620BA90A551DBDBAF3DE11BDDC9FB4B7">Enantiomorphs and taxonomy of three conchological species in flat-shelled snails Trichocathaica (Pulmonata, Camaenidae)</mods:title>
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<mods:namePart id="5143C60930387283CC78B24D5CF4B451">Pall-Gergely, Barna</mods:namePart>
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<mods:namePart id="30EBB54BCB49AEE394148240DEB302BF">unyadi, Andras</mods:namePart>
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<mods:namePart id="1F9D3FFDAD35B0E1751F9EE7C110A02C">Asami, Takahiro</mods:namePart>
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<mods:date id="F6E70EF6FAA1013CCC95EE8907CFD5B4">2018</mods:date>
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<mods:number id="2A8FA31B3D77910B2991B6EB615292B5">810</mods:number>
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<mods:classification id="77DF9E6FB58EE58C55C8E8C03AE1B28A">journal article</mods:classification>
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<mods:identifier id="526CA5560375553FFD6DE10D58D0BFC0" type="DOI">http://dx.doi.org/10.3897/zookeys.810.29824</mods:identifier>
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<treatment id="2F2663F6B68ECD1B119F1E9EAF91CA1D" ID-GBIF-Taxon="154127333" LSID="urn:lsid:plazi:treatment:2F2663F6B68ECD1B119F1E9EAF91CA1D" httpUri="http://treatment.plazi.org/id/2F2663F6B68ECD1B119F1E9EAF91CA1D" lastPageId="8" lastPageNumber="27" pageId="6" pageNumber="25">
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<subSubSection id="4AA89BE1F319E86B8D1E1F6F7B20FC1A" pageId="6" pageNumber="25" type="nomenclature">
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<paragraph id="131F9C8CBDB0E08C505F3712EC21BC36" pageId="6" pageNumber="25">
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<taxonomicName id="E99579AA221E01A3F3D6E37196F80F71" ID-CoL="7D2F7" authority="Pilsbry, 1934" authorityName="Pilsbry" authorityYear="1934" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica vestita" order="Stylommatophora" pageId="6" pageNumber="25" phylum="Mollusca" rank="species" species="vestita">Trichocathaica vestita (Pilsbry, 1934)</taxonomicName>
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<taxonomicNameLabel id="0B824BB6742084FFE3CC180B903EED00" pageId="6" pageNumber="25">comb. n.</taxonomicNameLabel>
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Figures 4A, 5, 6, 7, 8
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</paragraph>
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</subSubSection>
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<subSubSection id="03A7709BAAAE6D0AF7EF0457D0A12F0F" pageId="6" pageNumber="25" type="reference_group">
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<paragraph id="033726953283A6733035501DF8DEAA61" pageId="6" pageNumber="25">
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<taxonomicName id="D431F65AEE124057AAE7D92F12EF850D" class="Gastropoda" family="Camaenidae" genus="Cathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cathaica constantinae" order="Stylommatophora" pageId="6" pageNumber="25" phylum="Mollusca" rank="subSpecies" species="constantinae" subSpecies="vestita">Cathaica constantinae vestita</taxonomicName>
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Pilsbry, 1934: 15, plate 3, figs 5-7.
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</paragraph>
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</subSubSection>
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<subSubSection id="7C7EDC6710E63C416EA8641EB0867234" pageId="6" pageNumber="25" type="types examined">
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<paragraph id="74E991DB710D170A76CD135912735B38" pageId="6" pageNumber="25">Types examined.</paragraph>
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<paragraph id="4EAC3DB6B422B686BC51AEF59A2797E3" pageId="6" pageNumber="25">Between Wenchwan and Weichow, June, 1931, ANSP 159708 (photos of the holotype were examined).</paragraph>
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</subSubSection>
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<subSubSection id="1326BA13BF82949732EB5FA58929BE83" pageId="6" pageNumber="25" type="materials_examined">
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<paragraph id="8D141A44147D622E543BB3306B3FEED2" pageId="6" pageNumber="25">Additional material examined.</paragraph>
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<paragraph id="764DA8F55E5733B4AD636EFEE7E46F67" pageId="6" pageNumber="25">
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2015/64 Sichuan, Ganzi Zhou, Kangding Xian, Wasihekou, southern side of the river, along the highway, 1420 m a.s.l., 30°04.564'N 102°09.865'E, leg. A. Hunyadi, 14.06.2015, HNHM 103470 (dextral shell, Fig. 5
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<normalizedToken id="EF48995F4B520AB32990E36970E3DE79" originalValue="D–F">D-F</normalizedToken>
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+ ethanol-preserved body: Figs 6A, 7A, 8A), HA/15 dextral shells, PGB/3 dextral shells; 2015/65 China, Sichuan, Ganzi Zhou, Kangding Xian, Wasihekou 200 m towards Guzan Zhen, around the stupa, 1420 m a.s.l., 30°04.565'N, 102°10.085'E, leg. Hunyadi, A. & Szekeres, M., 14.06.2015, HNHM 103471 (1 sinistral shell, Fig. 5
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<normalizedToken id="BCEA667889421584424514BF6969BC55" originalValue="A–C">A-C</normalizedToken>
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+ ethanol-preserved body: Figs 6
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<normalizedToken id="2895B8ABC3E71F9C4F8DD5F0573C8B1E" originalValue="B–C">B-C</normalizedToken>
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, 7B, 8B), HA/6 sinistral shells + 2 dextral shells, PGB/3 sinistral shells; W-Sytschuan, Wa-sy-Kou am Tung, coll.
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<normalizedToken id="E11A5D477E80F2263C72D8D5FBD04D1A" originalValue="Möllendorff">Moellendorff</normalizedToken>
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ex coll. Potanin, SMF 8940 (1 dextral shell, paralectotype of
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<taxonomicName id="C66CF25FBC78E6D1380D5D3B2272DB0C" lsidName="T amphidroma" pageId="6" pageNumber="25" rank="species" species="amphidroma">amphidroma</taxonomicName>
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); China, W-Sytschuan: Wa-sy-Kou am Tung, coll. O.
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<normalizedToken id="78F9BC7D7B5B9106E24FF2939D03273D" originalValue="Möllendorff">Moellendorff</normalizedToken>
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ex coll. Potanin, SMF 349506 (1 dextral shell, paralectotype of
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<taxonomicName id="D55E67E8427F7284F586623DDA15AA28" lsidName="T amphidroma" pageId="6" pageNumber="25" rank="species" species="amphidroma">amphidroma</taxonomicName>
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, ex SMF 8941); Sy-tschuan, rechter Ufer des Flusses Tun bei dem Torfe (?) Wa-sy-ku, coll. O.
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<normalizedToken id="D24292A049F6B46C7885ED4ABAB3CFB8" originalValue="Möllendorff">Moellendorff</normalizedToken>
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ex coll. Potanin 3898a, 1903, SMF 95002 (1 dextral shell, paralectotype of
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<taxonomicName id="73F1D8CD68F441F27D21EFD21F7E44A0" lsidName="T amphidroma" pageId="6" pageNumber="25" rank="species" species="amphidroma">amphidroma</taxonomicName>
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); W-Sy-tschuan, coll.
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<normalizedToken id="735BBD7AD95103B8979E5C569B0E1471" originalValue="Möllendorff">Moellendorff</normalizedToken>
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, SMF 349505 (1 dextral shell, paralectotype of
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<taxonomicName id="CBF3BE850B353D7995C3C0C3B74A9432" lsidName="T amphidroma" pageId="6" pageNumber="25" rank="species" species="amphidroma">amphidroma</taxonomicName>
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, ex SMF 9171); W-Sytschuan, Liu-Ting am Tung, coll.
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<normalizedToken id="E9C94AB783710EAC94F0401FC28C1639" originalValue="Möllendorff">Moellendorff</normalizedToken>
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ex coll. Potanin, SMF 349503 (2 dextral shells, paralectotypes of
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<taxonomicName id="8460528957A82B406D6716C24EA81F30" lsidName="T amphidroma" pageId="6" pageNumber="25" rank="species" species="amphidroma">amphidroma</taxonomicName>
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, ex SMF 8938); W-Ufer d. Lu Ho (Ta Tu Ho) zw. Tapien und Ja
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<normalizedToken id="9FDD3461471F41FE3BA38682ACAD15EA" originalValue="sz’kou">sz'kou</normalizedToken>
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, ex Krejci-Graf, 05.08.1930, SMF 349507 (2 dextral shells, ex SMF 24667).
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</paragraph>
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<caption id="CBD3F7CDD6C4F656C6C1192499A658E5" pageId="6" pageNumber="25">
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<paragraph id="FA05EF61AA63DE742E0EC0F76F3F473F" pageId="6" pageNumber="25">
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Figure 4. Shells of
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<taxonomicName id="DFE8E93CBF1F8AC543EF18995E7526E5" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica vestita" order="Stylommatophora" pageId="6" pageNumber="25" phylum="Mollusca" rank="species" species="vestita">Trichocathaica vestita</taxonomicName>
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(Pilsbry, 1934) (holotype, ANSP 159708) (A) and
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<taxonomicName id="504CB721D2193CBB7A6263B1DBD7AA91" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Camaena) constantinae" order="Stylommatophora" pageId="6" pageNumber="25" phylum="Mollusca" rank="species" species="constantinae" subGenus="Camaena">Helix (Camaena) constantinae</taxonomicName>
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Adams, 1870 (B) (illustration from the original description). Scale bars: Scale refers only to A.
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</paragraph>
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</caption>
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<caption id="79C3EE79895AD4B18E78308097EB2117" pageId="6" pageNumber="25">
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<paragraph id="AFB598323494578AAD7FE34C675049E0" pageId="6" pageNumber="25">
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Figure 5. Shells (A, D), dorsal (B, E), and ventral (C, F) sculpture of
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<taxonomicName id="6413E02297206EE9638028FE935C085B" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica vestita" order="Stylommatophora" pageId="6" pageNumber="25" phylum="Mollusca" rank="species" species="vestita">Trichocathaica vestita</taxonomicName>
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<normalizedToken id="CA66787624D2542F283AB4D05A57F049" originalValue="A–C">A-C</normalizedToken>
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HNHM 103471 (D = 22.7 mm)
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<normalizedToken id="53467888D3D6E7C34B8B485A8AF9A5C6" originalValue="D–F">D-F</normalizedToken>
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HNHM 103470 (D = 22.8 mm). Upper scale (A, D), lower scale (B, C, E, F). All photos: B.
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<normalizedToken id="E60ACE7B449D5FC95A259A96B55A8B0D" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="E2D145962569399263E7CB1C166B8AB7" pageId="6" pageNumber="25" type="diagnosis">
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<paragraph id="8C2DE5ED53130AD9E8A9567E559A6B7D" pageId="6" pageNumber="25">Diagnosis.</paragraph>
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<paragraph id="E50F4086EE0FE7D47A61F04BCAD80E7C" pageId="6" pageNumber="25">Shell sinistral or dextral, body whorl rounded to keeled, teleoconch finely wrinkled with small scale-like periostracal folds; fold scars (if visible) represented as short curved lines.</paragraph>
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</subSubSection>
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<subSubSection id="507E0DA3A324C7517482498C19EA37CD" pageId="6" pageNumber="25" type="description">
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<paragraph id="593378B2F73521E091CC5A3E84C53A56" pageId="6" pageNumber="25">Description.</paragraph>
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<paragraph id="9AEAFFC5B25ABD1E1074E8362523371C" pageId="6" pageNumber="25">Shell sinistral or dextral, spire slightly elevated; body whorl rounded (with a very slight indication of a keel) to keeled, protoconch consists of 1.25-1.5 whorls, finely, irregularly wrinkled; entire shell with 5.25-5.75 whorls; teleoconch finely, irregularly wrinkled, with small, low, dense periostracal folds having curved (C-shaped) base; scales not visible to the naked eye; in specimens/shell parts without periostracum the bases of folds or sometimes not visible as small curved lines; aperture subcircular, peristome slightly expanded, thin, sharp; inner, white thickening parallel to the peristome prominent, situated in some distance from peristome edge.</paragraph>
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</subSubSection>
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<subSubSection id="E2A8651D7D2A164D77C7CE98EAC21267" pageId="6" pageNumber="25" type="description">
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<paragraph id="2AF35FD1A537E944443D52E5F3603BBA" pageId="6" pageNumber="25">Measurements (in mm).</paragraph>
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<paragraph id="5901F87F3DC290B3FEFE14991167A5A3" pageId="6" pageNumber="25">D = 21.2-23.6, H = 11.1-13.6 (n = 12).</paragraph>
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</subSubSection>
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<subSubSection id="DFD7BF1C11A03CB785DF6461964B8A56" lastPageId="8" lastPageNumber="27" pageId="6" pageNumber="25" type="anatomy">
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<paragraph id="AE6246C91B573CC888D10FDFB5A2DA29" pageId="6" pageNumber="25">Anatomy</paragraph>
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<paragraph id="0D7D94CC786AD9F681C5EF0A15E970D7" pageId="6" pageNumber="25">
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(Figs 6, 7). Genital morphology of two specimens (HNHM 103470, dextral specimen and HNHM 103471, sinistral specimen) showed that the left ommatophoral retractor crosses between penis and vagina in the sinistral specimen, and that the right retractor in the dextral specimen. Atrium short, penis with a slimmer, shorter distal, and a thicker, longer proximal portion, distal portion covered by a weak penial sheath; epiphallus much more slender than penis, approximately as long as penis; retractor muscle shorter in sinistral and longer in dextral specimen, inserts on epiphallus, close to its meeting point with penis; proximal part of penis internally with reticulated zigzag sculpture caused by the perpendicular projections of longitudinal folds (Fig. 8); dart sac well developed, with thickened, larger basal part and smaller head part; dart was only found in the dextral specimen (Fig. 7); long glandulae insert on 4-6 points on the
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<normalizedToken id="DB1BE85AE256DB4974331934047AC61D" originalValue="“neck”">"neck"</normalizedToken>
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of the dart sac (at the meeting point of the body and head of the dart sac) (Fig. 6C); vagina short in dextral and longer in sinistral specimen, stalk of bursa copulatrix long, relatively slender, bursa ovoid, diverticulum absent; spermoviduct slender, no embryos found; albumen gland crescent shaped, talon relatively large.
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</paragraph>
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<caption id="52EE6F950CDA7E7AAC00C01A64006E9F" pageId="6" pageNumber="25">
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<paragraph id="8E5BBAD34BE227A2BFEC1F0FFE5F5236" pageId="6" pageNumber="25">
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Figure 6. Genital anatomy of
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<taxonomicName id="952D4DA7DAD95F077853A5C90CF5B089" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica vestita" order="Stylommatophora" pageId="6" pageNumber="25" phylum="Mollusca" rank="species" species="vestita">Trichocathaica vestita</taxonomicName>
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A Dextral specimen (HNHM 103470) B, C Sinistral specimen (HNHM 103471).
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</paragraph>
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</caption>
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<caption id="2AF9DE77BBA90620D6E031675CE274EC" pageId="6" pageNumber="25">
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<paragraph id="B263BC534FCE7DDC89B5B6E178BE7644" pageId="6" pageNumber="25">
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Figure 7. Opened dart sac of sculpture of
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<taxonomicName id="93BA28B6E4CA0654EE09A73AF208B63E" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica vestita" order="Stylommatophora" pageId="6" pageNumber="25" phylum="Mollusca" rank="species" species="vestita">Trichocathaica vestita</taxonomicName>
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A Dextral specimen (HNHM 103470) B, C Sinistral specimen (HNHM 103471). Arrow shows dart. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption id="6D0C8F23A1792A6A5E3125DB20C85D54" pageId="6" pageNumber="25">
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<paragraph id="787A9A6954FC8E520FA40C0BABE48691" pageId="6" pageNumber="25">
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Figure 8. Penial sculpture of
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<taxonomicName id="A8D250DFBD7F7514B7BD7B4FA08B0873" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica vestita" order="Stylommatophora" pageId="6" pageNumber="25" phylum="Mollusca" rank="species" species="vestita">Trichocathaica vestita</taxonomicName>
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A Dextral specimen (HNHM 103470) B, C Sinistral specimen (HNHM 103471). Scale bars: 1 mm.
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</paragraph>
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</caption>
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<paragraph id="C1E5B2F546D684332F4C83B381DD5EA2" lastPageId="8" lastPageNumber="27" pageId="6" pageNumber="25">
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We found no discrete differences between the dextral and sinistral individuals in gross anatomy of the genital system or in the internal structure of the dart sac (Figs 6, 7).
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<pageBreakToken id="04D53D01705838ABCED7ECD1F8B87BEF" pageId="7" pageNumber="26" start="start">The</pageBreakToken>
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internal surface of the penial tube exhibited complex patterns of microsculpture typical to camaenid snails (Fig. 8). In this structure, several slight differences were noticeable between these specimens of enantiomorphs. Around the area leading to the epiphallus (upper edge in Fig. 8), thin longitudinal pilasters are more tightly gathered with narrower furrows in the dextral than in the sinistral. In both of them, toward the middle of the penial tube, these pilasters become thick and sparse with wider furrows and form a reticulate pattern. Longitudinally (in the direction towards the genital orifice, the bottom in Fig. 8) under the reticulated range, zigzag crenulated pilasters are present in parallel. In these portions, structural change from the reticulate pattern to the parallel pilasters is more distinct in the dextral than in the sinistral. Smooth-bottomed furrows separate the zigzag pilasters in the dextral, whereas those furrows are not obviously present between the irregularly zigzag-shaped pilasters in the sinistral. This structure of the dextral is present in a longitudinally wider range than the sinistral. In the sinistral, instead, the irregular zigzag structure becomes weak or disappears from the longitudinal pilasters, which become thick and pronounced near the genital orifice. This longitudinal structural change is not present in the dextral. The dextral
|
||
<pageBreakToken id="795EE2DD6935A7952CFE465855A221C1" pageId="8" pageNumber="27" start="start">instead</pageBreakToken>
|
||
exhibits a different pattern such that the longitudinally parallel zigzag pilasters are continuously present and merge with one another without forming major pilasters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="80AAEEC47F087654CA0040747B2910EE" pageId="8" pageNumber="27" type="differential diagnosis">
|
||
<paragraph id="58B6A25CC93069B49507603BA00F6D4B" pageId="8" pageNumber="27">Differential diagnosis.</paragraph>
|
||
<paragraph id="4C5E71530A63A20AF28DA929D2DA2D59" pageId="8" pageNumber="27">
|
||
<taxonomicName id="415D4D045B7365B16B38E633B9130EC8" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica vestita" order="Stylommatophora" pageId="8" pageNumber="27" phylum="Mollusca" rank="species" species="vestita">Trichocathaica vestita</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="95A09E5D69A4EF0BA5D67AA25CCE798B" lsidName="T. macrosquamata" pageId="8" pageNumber="27" rank="species" species="macrosquamata">T. macrosquamata</taxonomicName>
|
||
sp. n. by having the smaller periostracal folds (scales) on the entire shell surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="26576B924DF322298488D3D41A087803" pageId="8" pageNumber="27" type="distribution">
|
||
<paragraph id="AE1F930184E1FC75091B404FFDD85135" pageId="8" pageNumber="27">Distribution.</paragraph>
|
||
<paragraph id="51BEB1F31039051D988E33F791F9C93B" pageId="8" pageNumber="27">This species is known from the valley of the Dadu River at Luding and Wasigou.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="384F36A3BE2E1E93FB6552F53F6288A1" pageId="8" pageNumber="27" type="remarks">
|
||
<paragraph id="6B4318E9E4F8F9FC2FCE187E1E3CC1B5" pageId="8" pageNumber="27">Remarks.</paragraph>
|
||
<paragraph id="77E6FBC15CDDB9439F7855462FE0D423" pageId="8" pageNumber="27">
|
||
This species was described as a subspecies of
|
||
<taxonomicName id="AAC9FBE13F5E7B7CB9DC698050825164" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Camaena) constantinae" order="Stylommatophora" pageId="8" pageNumber="27" phylum="Mollusca" rank="species" species="constantinae" subGenus="Camaena">Helix (Camaena) constantinae</taxonomicName>
|
||
Adams, 1870. We had no possibility to examine that species; however, it has remarkably different shell traits, such as the strongly sculptured shell surface and a white band (see Fig. 4B). That species is probably not a
|
||
<taxonomicName id="33D658857E851E989DAADCDAC75A35C6" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica" order="Stylommatophora" pageId="8" pageNumber="27" phylum="Mollusca" rank="genus">Trichocathaica</taxonomicName>
|
||
, but something entirely different, as
|
||
<bibRefCitation id="765847ECD4F63FDBFA0DD4A05D35F085" author="Pilsbry, HA" journalOrPublisher="Proceedings of the Academy of Natural Science of Philadelphia" pageId="19" pageNumber="38" pagination="5 - 28" title="Zoological results of the Dolan West China expedition of 1931, Part II, Mollusks." volume="86" year="1934">Pilsbry (1934)</bibRefCitation>
|
||
already suspected. Moreover,
|
||
<taxonomicName id="FDB86B27DF9DCB6ABEEC710A635BA9F3" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica" order="Stylommatophora" pageId="8" pageNumber="27" phylum="Mollusca" rank="genus">Trichocathaica</taxonomicName>
|
||
seems to inhabit only the mountains in Sichuan and southern Gansu, and
|
||
<taxonomicName id="BA55A001A27B1C2D35780ACF95D4B3B6" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Camaena) constantinae" order="Stylommatophora" pageId="8" pageNumber="27" phylum="Mollusca" rank="species" species="constantinae" subGenus="Camaena">Helix (Camaena) constantinae</taxonomicName>
|
||
was collected in the "Ichang gorge" on the Yangtze River in Hubei Province (ca 30°55'N, 110°50'E,
|
||
<bibRefCitation id="CE52774C8AA3C492BD1128C308E0B9FA" author="Adams, H" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="18" pageNumber="37" pagination="377 - 380" title="Descriptions of ten new species of land and freshwater shells collected by Robert Swinhoe, Esq., in China and Formosa." volume="1870" year="1870">Adams 1870</bibRefCitation>
|
||
). Therefore, we handle
|
||
<taxonomicName id="991DE3DF726E4325FF4C8CAB04BD3A8F" class="Gastropoda" family="Camaenidae" genus="Trichocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocathaica vestita" order="Stylommatophora" pageId="8" pageNumber="27" phylum="Mollusca" rank="species" species="vestita">Trichocathaica vestita</taxonomicName>
|
||
(Pilsbry, 1934) as a species of its own right.
|
||
</paragraph>
|
||
<paragraph id="2ACF10D406B6F2863F70E68D8CE3F38F" pageId="8" pageNumber="27">
|
||
<bibRefCitation id="6B84296503DC382C0FC6D8C1BC2D159D" author="Pilsbry, HA" journalOrPublisher="Proceedings of the Academy of Natural Science of Philadelphia" pageId="19" pageNumber="38" pagination="5 - 28" title="Zoological results of the Dolan West China expedition of 1931, Part II, Mollusks." volume="86" year="1934">Pilsbry (1934)</bibRefCitation>
|
||
already noted in the original description that the species show an extreme variability in terms of the development of the keel. Our data also indicates that the keel morphology is variable within and between populations. In the sample of 2015/64, 15 shells had nearly rounded body whorls, two were keeled, and two were intermediate between those rounded and keeled forms. Although every shell in the sample of 2015/65 had a keel, two of them were similar to the intermediate form of the 2015/64 sample.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |