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<mods:title id="F449EF2949767373161730327E1712B2">Description of two new Australian species of Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) and females of two further species with notes on their biogeography</mods:title>
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<mods:namePart id="2610BF70975CEB01B6CC97B45B4F7B3F">Moore, Michael D.</mods:namePart>
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<mods:namePart id="530FB9DC055EF1DCAFC0E9D1B5AB357E">Beaver, Ethan P.</mods:namePart>
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<mods:affiliation id="3D2198EE2BDA49494BC0C96CBE3DCD46">Biological and Earth Sciences, South Australian Museum, SA 5000, Australia &amp; ethan. beaver @ live. com. au; https: // orcid. org / 0000 - 0002 - 0613 - 7046</mods:affiliation>
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<mods:namePart id="0200BFD826918DE48DBBDE9A6F783773">Velasco-Castrillón, Alejandro</mods:namePart>
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<mods:affiliation id="BCC5DC449DEDCBB7E9A76484FAF324BD">Biological and Earth Sciences, South Australian Museum, SA 5000, Australia &amp; a. velascocastrillon @ gmail. com; https: // orcid. org / 0000 - 0002 - 3516 - 6655</mods:affiliation>
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<mods:namePart id="84A5DE712E266E3505DAA824D63B4E09">Stevens, Mark I.</mods:namePart>
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<treatment id="216587E00D21FF9AEFC4F1A231BEFBEF" ID-DOI="http://doi.org/10.5281/zenodo.4401264" ID-GBIF-Taxon="177010365" ID-Zenodo-Dep="4401264" LSID="urn:lsid:plazi:treatment:216587E00D21FF9AEFC4F1A231BEFBEF" httpUri="http://treatment.plazi.org/id/216587E00D21FF9AEFC4F1A231BEFBEF" lastPageId="15" lastPageNumber="86" pageId="13" pageNumber="84" scope_family="Hepialidae" scope_order="Lepidoptera">
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<heading id="F23B819A0D21FF98EFC4F1A2332DFF07" bold="true" box="[151,846,223,250]" fontSize="11" level="1" pageId="13" pageNumber="84" reason="1">
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<taxonomicName id="6ECC4D750D21FF98EFC4F1A23282FF07" ID-CoL="99VVG" authority="Simonsen, 2018" authorityName="Simonsen" authorityYear="2018" box="[151,737,223,250]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D21FF98EFC4F1A23274FF04" bold="true" box="[151,535,223,250]" italics="true" pageId="13" pageNumber="84">Abantiades macropusinsulariae</emphasis>
<bibRefCitation id="CD5D4B070D21FF98ED4DF1A23282FF07" author="Simonsen, T. J." box="[542,737,223,249]" pageId="13" pageNumber="84" pagination="1 - 300" refId="ref12387" refString="Simonsen, T. J. (2018) Splendid Ghost Moths and Their Allies. Monographs on Australian Lepidoptera, 12, i-xii + 1 - 300. https: // doi. org / 10.1071 / 9781486307487" type="journal article" year="2018">Simonsen, 2018</bibRefCitation>
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(female)
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</paragraph>
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<paragraph id="A97336F60D21FF98EFC4F07E3132FEE3" blockId="13.[151,846,223,285]" box="[151,337,259,285]" pageId="13" pageNumber="84">
<figureCitation id="31F72A730D21FF98EFC4F07E30BFFEE3" box="[151,220,259,285]" captionStart="FIGURE 6" captionStartId="11.[151,260,1810,1836]" captionTargetBox="[173,1412,602,1776]" captionTargetId="figure-196@11.[151,1436,594,1785]" captionTargetPageId="11" captionText="FIGURE 6. Abantiades macropusinsulariae. (a) Adult male, SAMA. SA, Kangaroo Island, North Cape Road, Bay of Shoals, M+M Moore, 35o3547.7”S 137o3540.7”E, 2 May 2019. Specimen No 19173. Leg removed for DNA analysis; Sample designation G105. SAMA No. 31-021004. (b) Adult female, SAMA. SA, Kangaroo Island, North Cape Road, Bay of Shoals, M+M Moore, 35o3547.7”S 137o3540.7”E, 2 May 2019. Specimen No 19169. Leg removed for DNA analysis; Sample designation G116. Dissected M.D. Moore 12th Feb. 2020. SAMA No. 31-021005. Arrow: white spot half way along dorsum." figureDoi="http://doi.org/10.5281/zenodo.4401282" httpUri="https://zenodo.org/record/4401282/files/figure.png" pageId="13" pageNumber="84">Figs 6</figureCitation>
ab, 7ad.
</paragraph>
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<subSubSection id="E1D6657D0D21FF98EFC4F036327EFE2C" pageId="13" pageNumber="84" type="materials_examined">
<paragraph id="A97336F60D21FF98EFC4F0363114FE9B" blockId="13.[151,1437,331,466]" box="[151,375,331,357]" pageId="13" pageNumber="84">Material Examined:</paragraph>
<paragraph id="A97336F60D21FF98EF94F012327EFE2C" blockId="13.[151,1437,331,466]" pageId="13" pageNumber="84">
<materialsCitation id="19A43CAB0D21FF98EF94F0123124FE77" ID-GBIF-Occurrence="3019881308" box="[199,327,367,393]" collectionCode="SAMA" pageId="13" pageNumber="84" specimenCount="1" specimenCount-female="1">
<collectionCode id="CFDDAE330D21FF98EF94F012317FFE77" box="[199,284,367,393]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="13" pageNumber="84" type="Museum">SAMA</collectionCode>
.
<specimenCount id="BFCAFD7F0D21FF98EE77F0123124FE77" box="[292,327,367,393]" pageId="13" pageNumber="84" type="female">1♀</specimenCount>
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,
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<collectionCode id="CFDDAE330D21FF98EE00F012311AFE77" box="[339,377,367,393]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="13" pageNumber="84">SA</collectionCode>
,
<location id="AC13602D0D21FF98EED0F00D325EFE77" LSID="urn:lsid:plazi:treatment:216587E00D21FF9AEFC4F1A231BEFBEF:AC13602D0D21FF98EED0F00D325EFE77" box="[387,573,367,394]" county="Bay of Shoals" latitude="-35.59658" longLatPrecision="1" longitude="137.59464" municipality="North Cape Road" name="Kangaroo Island" pageId="13" pageNumber="84">Kangaroo Island</location>
,
<collectingMunicipality id="4917AC8C0D21FF98ED15F012336AFE77" box="[582,777,367,393]" pageId="13" pageNumber="84">North Cape Road</collectingMunicipality>
,
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Bay of
<collectorName id="043953200D21FF98EC33F01233CEFE77" box="[864,941,367,393]" pageId="13" pageNumber="84">Shoals</collectorName>
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, M+
<collectorName id="043953200D21FF98EC8CF00D3428FE74" box="[991,1099,368,394]" pageId="13" pageNumber="84">M Moore</collectorName>
,
<geoCoordinate id="CCF850310D21FF98EB07F0123488FE74" box="[1108,1259,367,394]" degrees="35" direction="south" minutes="35" orientation="latitude" pageId="13" pageNumber="84" precision="1" seconds="47.7" value="-35.59658">
35
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3547.7”S
</geoCoordinate>
<geoCoordinate id="CCF850310D21FF98EBBCF01235FBFE74" box="[1263,1432,367,394]" degrees="137" direction="east" minutes="35" orientation="longitude" pageId="13" pageNumber="84" precision="1" seconds="40.7" value="137.59464">
137
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3540.7”E
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,
<date id="DD7210360D21FF98EFC4F0EE3140FE53" box="[151,291,403,430]" pageId="13" pageNumber="84" value="2019-05-02">
<collectingDate id="CD36E9DE0D21FF98EFC4F0EE3140FE53" box="[151,291,403,430]" pageId="13" pageNumber="84" value="2019-05-02">2 May 2019</collectingDate>
</date>
.
<collectorName id="043953200D21FF98EE7EF0EE31A6FE50" box="[301,453,403,430]" pageId="13" pageNumber="84">Specimen No</collectorName>
19169.
<collectorName id="043953200D21FF98ED72F0E9322EFE50" box="[545,589,404,430]" pageId="13" pageNumber="84">Leg</collectorName>
removed for DNA analysis;
<collectorName id="043953200D21FF98ECC2F0EE3456FE53" box="[913,1077,403,429]" pageId="13" pageNumber="84">Designation G</collectorName>
116.
<collectorName id="043953200D21FF98EB38F0EE350EFE50" box="[1131,1389,403,430]" pageId="13" pageNumber="84">Dissected M.D. Moore</collectorName>
<date id="DD7210360D21FF98EA20F0EE3168FE2F" pageId="13" pageNumber="84" value="2020-02-12">
<collectingDate id="CD36E9DE0D21FF98EA20F0EE3168FE2F" pageId="13" pageNumber="84" value="2020-02-12">
12
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Feb. 2020
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</date>
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. SAMA No. 31-021005.
</paragraph>
</subSubSection>
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<paragraph id="A97336F60D21FF98EFC4F35E317AFDC3" blockId="13.[151,281,547,573]" box="[151,281,547,573]" pageId="13" pageNumber="84">
<heading id="F23B819A0D21FF98EFC4F35E317AFDC3" bold="true" box="[151,281,547,573]" fontSize="11" level="1" pageId="13" pageNumber="84" reason="1">
<emphasis id="9BB8EAE40D21FF98EFC4F35E317AFDC3" bold="true" box="[151,281,547,573]" pageId="13" pageNumber="84">Diagnosis:</emphasis>
</heading>
</paragraph>
<paragraph id="A97336F60D21FF98EFC4F316325FFC13" blockId="13.[151,1437,619,1005]" pageId="13" pageNumber="84">
Endemic to Kangaroo Island, the males of
<taxonomicName id="6ECC4D750D21FF98ED22F316330CFD7B" authorityName="Simonsen" authorityYear="2018" box="[625,879,619,645]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D21FF98ED22F316330CFD7B" box="[625,879,619,645]" italics="true" pageId="13" pageNumber="84">A. macropusinsulariae</emphasis>
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have patterned forewings with many scattered, irregular shaped, white coloured marks, and lighter coloured hindwings giving a two-tone effect to the specimens and they also have a red-brown antennal flagellum. Specifically,
<bibRefCitation id="CD5D4B070D21FF98EC6DF3CE339DFD33" author="Simonsen, T. J." box="[830,1022,691,717]" pageId="13" pageNumber="84" pagination="1 - 300" refId="ref12387" refString="Simonsen, T. J. (2018) Splendid Ghost Moths and Their Allies. Monographs on Australian Lepidoptera, 12, i-xii + 1 - 300. https: // doi. org / 10.1071 / 9781486307487" type="journal article" year="2018">Simonsen (2018)</bibRefCitation>
states, “second segment of the labial palp very long, more than 3x the length of the terminal segment, S8 short and pentagonal with a medially slightly projecting posterior margin, distal pseudoteguminal lobes smoothly triangularly rounded without a concave curvature on the dorso-posterior margin, disto-posterior arm of vinculum with a small, lateral ridge; saccus in posteroventral view narrowly V-shaped”. The female of this species is black/grey and white, has patterned forewings and a red-brown antennal flagellum with distinctly triforked pectinations. Specifically, it varies from all other triforked
<taxonomicName id="6ECC4D750D21FF98EFC4F2F63176FC5B" authorityName="Herrich-Schaffer" authorityYear="1855" box="[151,277,907,933]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="9BB8EAE40D21FF98EFC4F2F63176FC5B" box="[151,277,907,933]" italics="true" pageId="13" pageNumber="84">Abantiades</emphasis>
</taxonomicName>
females by; i) the heavily sclerotized bands bordering the medial lobe of the antevaginal lamella, and ii) the long, conical end projection on the corpus bursae. The wings in both sexes of this species are shorter and broader than in the other species in this clade.
</paragraph>
</subSubSection>
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<paragraph id="A97336F60D21FF98EFC4F5423151FBA7" blockId="13.[151,306,1087,1113]" box="[151,306,1087,1113]" pageId="13" pageNumber="84">
<heading id="F23B819A0D21FF98EFC4F5423151FBA7" bold="true" box="[151,306,1087,1113]" fontSize="11" level="1" pageId="13" pageNumber="84" reason="1">
<emphasis id="9BB8EAE40D21FF98EFC4F5423151FBA7" bold="true" box="[151,306,1087,1113]" pageId="13" pageNumber="84">Description:</emphasis>
</heading>
</paragraph>
<paragraph id="A97336F60D21FF98EFC4F5FA308AFB5F" blockId="13.[151,1437,1159,2014]" box="[151,233,1159,1185]" pageId="13" pageNumber="84">Female</paragraph>
<paragraph id="A97336F60D21FF98EF94F5D635E2FA87" blockId="13.[151,1437,1159,2014]" pageId="13" pageNumber="84">
<emphasis id="9BB8EAE40D21FF98EF94F5D6316EFB3B" box="[199,269,1195,1221]" italics="true" pageId="13" pageNumber="84">Head:</emphasis>
Head, thickly covered in medium grey piliform scales, fronto-clypeal scales darker. Two large compound eyes, each only slightly smaller than head capsule. Labial palps approximately straight, three segmented, second palpomere longest, third smallest, sub-spherical, all densely covered in brown coloured scales. Antennae: (
<figureCitation id="31F72A730D21FF98EA39F58E30C6FACC" captionStart="FIGURE 7" captionStartId="14.[151,260,1433,1459]" captionTargetBox="[173,1410,616,1401]" captionTargetId="figure-146@14.[151,1436,594,1409]" captionTargetPageId="14" captionText="FIGURE 7. Abantiades macropusinsulariae. (SA, Kangaroo Island, North Cape Road, Bay of Shoals, M+M Moore, 35o3547.7”S 137o3540.7”E, 2 May 2019. Specimen No 19169. Leg removed for DNA analysis; Sample designation G116. Dissected M.D. Moore 12th Feb. 2020. SAMA No. 31-021005.) (a) posterior view female genitalia, (b) corpus bursae and ductus bursae, (c) cruciform pectination (lateral view), (d) cruciform pectinations (frontal view). Morphological structures: CB: corpus bursae; Db: ductus bursae; Dp: dorsal plate; Lal: lateral lobe of the antevaginal lamella; Mal: medial lobe of the antevaginal lamella; Sub: sub-anal plate. Arrow: triangular projection on dorsal plate." figureDoi="http://doi.org/10.5281/zenodo.4401284" httpUri="https://zenodo.org/record/4401284/files/figure.png" pageId="13" pageNumber="84">Figs 7</figureCitation>
cd). total 48 segments, length
<quantity id="6E349B130D21FF98ED5CF46A32E8FACC" box="[527,651,1303,1330]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="13" pageNumber="84" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
, 20% of forewing length; flagellum dark red-brown, monopectinate, basal pectinations tear-drop shaped, then transitional to triforked about 1/5 from base. Lateral projections red-brown approximately parallel sides with blunt, rounded end, central fork white, long triangular with narrowly acute tip.
</paragraph>
<paragraph id="A97336F60D21FF98EF94F4FE3523F8D4" blockId="13.[151,1437,1159,2014]" pageId="13" pageNumber="84">
<emphasis id="9BB8EAE40D21FF98EF94F4FE3142FA63" box="[199,289,1411,1437]" italics="true" pageId="13" pageNumber="84">Thorax:</emphasis>
Densely clothed in medium-grey piliform scales. Forewing: length
<quantity id="6E349B130D21FF98EB47F4FE3407FA60" box="[1044,1124,1411,1438]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="13" pageNumber="84" unit="mm" value="48.0">48 mm</quantity>
, membrane whitish opaque, veins white, thinly covered with short, dark grey scales. Dorsal surface: space between costa and Sc vein thickly covered in small piliform grey scales particularly near base. Two large white areas; discoidal, filled with dull white scales, and in this specimen divided medially into two irregular patches, patch of dark brown/grey scales along anterior edge in basal half. Sub-terminal band dull white, sublinear between M3 and near apex. Also, a series of intervein, irregular white shapes along termen. White, strongly marked scrolling lines in apical area, less apparent elsewhere. Circular white mark present on posterior wing half way along the dorsum (
<figureCitation id="31F72A730D21FF98EBC3F7263482F98B" box="[1168,1249,1627,1653]" captionStart="FIGURE 6" captionStartId="11.[151,260,1810,1836]" captionTargetBox="[173,1412,602,1776]" captionTargetId="figure-196@11.[151,1436,594,1785]" captionTargetPageId="11" captionText="FIGURE 6. Abantiades macropusinsulariae. (a) Adult male, SAMA. SA, Kangaroo Island, North Cape Road, Bay of Shoals, M+M Moore, 35o3547.7”S 137o3540.7”E, 2 May 2019. Specimen No 19173. Leg removed for DNA analysis; Sample designation G105. SAMA No. 31-021004. (b) Adult female, SAMA. SA, Kangaroo Island, North Cape Road, Bay of Shoals, M+M Moore, 35o3547.7”S 137o3540.7”E, 2 May 2019. Specimen No 19169. Leg removed for DNA analysis; Sample designation G116. Dissected M.D. Moore 12th Feb. 2020. SAMA No. 31-021005. Arrow: white spot half way along dorsum." figureDoi="http://doi.org/10.5281/zenodo.4401282" httpUri="https://zenodo.org/record/4401282/files/figure.png" pageId="13" pageNumber="84">Fig. 6b</figureCitation>
arrow). White markings edged in blackish-brown scales with lighter, grey scales filling the inter-scrolled spaces. Ventral surface unmarked but with dorsal markings faintly visible, sparsely covered by reddish-brown digitiform scales that are longer near dorsum particularly near the base. Hindwing:
<quantity id="6E349B130D21FF98EC7AF7BA3318F91C" box="[809,891,1735,1762]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="84" unit="mm" value="39.0">39 mm</quantity>
, dorsal surface with whitish veins, and elongate reddish-brown piliform scales. Ventral surface, as for forewing. Legs: Dorsal and lateral surfaces densely covered in grey piliform scales, longer laterally, ventrally covered in sandy red scales. Epiphysis and arolium present.
</paragraph>
<paragraph id="A97336F60D21FF98EF94F64E3378F8B3" blockId="13.[151,1437,1159,2014]" box="[199,795,1843,1870]" pageId="13" pageNumber="84">
<emphasis id="9BB8EAE40D21FF98EF94F64E3159F8B3" box="[199,314,1843,1869]" italics="true" pageId="13" pageNumber="84">Abdomen:</emphasis>
Covered in long dark grey piliform scales.
</paragraph>
<paragraph id="A97336F60D21FF9BEF94F62A3438FF07" blockId="13.[151,1437,1159,2014]" lastBlockId="14.[151,1436,151,249]" lastPageId="14" lastPageNumber="85" pageId="13" pageNumber="84">
<emphasis id="9BB8EAE40D21FF98EF94F62A3159F88F" box="[199,314,1879,1905]" italics="true" pageId="13" pageNumber="84">Genitalia:</emphasis>
(
<figureCitation id="31F72A730D21FF98EE19F62A31F2F88C" box="[330,401,1879,1906]" captionStart="FIGURE 7" captionStartId="14.[151,260,1433,1459]" captionTargetBox="[173,1410,616,1401]" captionTargetId="figure-146@14.[151,1436,594,1409]" captionTargetPageId="14" captionText="FIGURE 7. Abantiades macropusinsulariae. (SA, Kangaroo Island, North Cape Road, Bay of Shoals, M+M Moore, 35o3547.7”S 137o3540.7”E, 2 May 2019. Specimen No 19169. Leg removed for DNA analysis; Sample designation G116. Dissected M.D. Moore 12th Feb. 2020. SAMA No. 31-021005.) (a) posterior view female genitalia, (b) corpus bursae and ductus bursae, (c) cruciform pectination (lateral view), (d) cruciform pectinations (frontal view). Morphological structures: CB: corpus bursae; Db: ductus bursae; Dp: dorsal plate; Lal: lateral lobe of the antevaginal lamella; Mal: medial lobe of the antevaginal lamella; Sub: sub-anal plate. Arrow: triangular projection on dorsal plate." figureDoi="http://doi.org/10.5281/zenodo.4401284" httpUri="https://zenodo.org/record/4401284/files/figure.png" pageId="13" pageNumber="84">Figs 7</figureCitation>
ab). Dorsal plate lightly sclerotized, bilobed, with deep “V” shaped central invagination of plate. Outer margin curved with triangular prominences in central region of each lobe (
<figureCitation id="31F72A730D21FF98EB05F60634C7F868" box="[1110,1188,1915,1942]" captionStart="FIGURE 7" captionStartId="14.[151,260,1433,1459]" captionTargetBox="[173,1410,616,1401]" captionTargetId="figure-146@14.[151,1436,594,1409]" captionTargetPageId="14" captionText="FIGURE 7. Abantiades macropusinsulariae. (SA, Kangaroo Island, North Cape Road, Bay of Shoals, M+M Moore, 35o3547.7”S 137o3540.7”E, 2 May 2019. Specimen No 19169. Leg removed for DNA analysis; Sample designation G116. Dissected M.D. Moore 12th Feb. 2020. SAMA No. 31-021005.) (a) posterior view female genitalia, (b) corpus bursae and ductus bursae, (c) cruciform pectination (lateral view), (d) cruciform pectinations (frontal view). Morphological structures: CB: corpus bursae; Db: ductus bursae; Dp: dorsal plate; Lal: lateral lobe of the antevaginal lamella; Mal: medial lobe of the antevaginal lamella; Sub: sub-anal plate. Arrow: triangular projection on dorsal plate." figureDoi="http://doi.org/10.5281/zenodo.4401284" httpUri="https://zenodo.org/record/4401284/files/figure.png" pageId="13" pageNumber="84">Fig. 7a</figureCitation>
arrow).Antevaginal lamella trilobate with broad areas of connection between lateral and medial plates. Medial plate more heavily sclerotized, dorsal margin centrally raised with minute bilobed prominence and heavily sclerotized convex sides. Long, thick setae on prominence and generally over central surface of medial plate. Also, on inner margins of lateral lobes and on lobes of dorsal plate. Sub-anal plates sclerotized, pentagonal. Corpus bursae broad at junction with narrow ductus bursae and then tapering as a long conical terminal. No diverticulum observed.
</paragraph>
</subSubSection>
<subSubSection id="E1D6657D0D22FF9BEFC4F0373190FDC3" pageId="14" pageNumber="85" type="distribution">
<paragraph id="A97336F60D22FF9BEFC4F03731B5FE9A" blockId="14.[151,470,330,357]" box="[151,470,330,357]" pageId="14" pageNumber="85">
<heading id="F23B819A0D22FF9BEFC4F03731B5FE9A" bold="true" box="[151,470,330,357]" fontSize="11" level="1" pageId="14" pageNumber="85" reason="1">
<emphasis id="9BB8EAE40D22FF9BEFC4F03731B5FE9A" bold="true" box="[151,470,330,357]" pageId="14" pageNumber="85">Distribution and Biology:</emphasis>
</heading>
</paragraph>
<paragraph id="A97336F60D22FF9BEFC4F0EE3243FE2C" blockId="14.[151,1436,403,573]" pageId="14" pageNumber="85">
<emphasis id="9BB8EAE40D22FF9BEFC4F0EE311CFE53" box="[151,383,403,429]" italics="true" pageId="14" pageNumber="85">Recorded Localities:</emphasis>
(
<figureCitation id="31F72A730D22FF9BEEDEF0EE3180FE53" box="[397,483,403,429]" captionStart="FIGURE 10" captionStartId="18.[151,260,1371,1397]" captionTargetBox="[172,1427,185,1340]" captionTargetId="figure-40@18.[151,1436,179,1347]" captionTargetPageId="18" captionText="FIGURE 10. Distribution Map of Australia for Abantiades kristenseni, A. inexpecta, A. macropusinsulariae, and A. cephalocorvus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4401290" httpUri="https://zenodo.org/record/4401290/files/figure.png" pageId="14" pageNumber="85">Figs 10</figureCitation>
,
<figureCitation id="31F72A730D22FF9BEEBCF0EE326AFE50" box="[495,521,403,430]" captionStart="FIGURE 12" captionStartId="20.[151,260,1736,1762]" captionTargetBox="[151,1436,270,1712]" captionTargetId="figure-84@20.[151,1436,270,1712]" captionTargetPageId="20" captionText="FIGURE 12. Habitat images. (a)+(b) Googs Lake, site of Abantiades cephalocorvus sp. nov. collection. (c)+(d) Capture site of A. tembyi sp. nov., 30 km ENE of Swan Reach. (e) Scrub on North Cape. A site of A. macropusinsulariae collection. (f) Scrub at Vivonne Bay. A site of A. macropusinsulariae collection." figureDoi="http://doi.org/10.5281/zenodo.4401294" httpUri="https://zenodo.org/record/4401294/files/figure.png" pageId="14" pageNumber="85">12</figureCitation>
ef) Coastal, previously only from Vivonne Bay (south-central of
<collectingCountry id="D1DB76660D22FF9BEBB4F0EE3553FE50" box="[1255,1328,403,430]" name="Iceland" pageId="14" pageNumber="85">Island</collectingCountry>
), but now also on North Cape (NE of
<collectingCountry id="D1DB76660D22FF9BEE98F0CA3277FE2C" box="[459,532,439,466]" name="Iceland" pageId="14" pageNumber="85">Island</collectingCountry>
).
</paragraph>
<paragraph id="A97336F60D22FF9BEF94F0A63177FDE7" blockId="14.[151,1436,403,573]" pageId="14" pageNumber="85">
<emphasis id="9BB8EAE40D22FF9BEF94F0A6313BFE0A" box="[199,344,475,501]" italics="true" pageId="14" pageNumber="85">Flight times:</emphasis>
April 19
<superScript id="5EB99BBE0D22FF9BEEEDF0A731A8FE16" attach="left" box="[446,459,474,488]" fontSize="6" pageId="14" pageNumber="85">th</superScript>
to May 3
<superScript id="5EB99BBE0D22FF9BED6BF0A73225FE16" attach="left" box="[568,582,474,488]" fontSize="6" pageId="14" pageNumber="85">rd</superScript>
(
<bibRefCitation id="CD5D4B070D22FF9BED07F0A6336AFE0B" author="Simonsen, T. J." box="[596,777,475,501]" pageId="14" pageNumber="85" pagination="1 - 300" refId="ref12387" refString="Simonsen, T. J. (2018) Splendid Ghost Moths and Their Allies. Monographs on Australian Lepidoptera, 12, i-xii + 1 - 300. https: // doi. org / 10.1071 / 9781486307487" type="journal article" year="2018">Simonsen 2018</bibRefCitation>
). The female and an associated male were collected on the 2
<superScript id="5EB99BBE0D22FF9BEFF6F08330D6FDF2" attach="left" box="[165,181,510,524]" fontSize="6" pageId="14" pageNumber="85">nd</superScript>
of May.
</paragraph>
<paragraph id="A97336F60D22FF9BEF94F35E3190FDC3" blockId="14.[151,1436,403,573]" box="[199,499,547,573]" pageId="14" pageNumber="85">
<emphasis id="9BB8EAE40D22FF9BEF94F35E3115FDC3" box="[199,374,547,573]" italics="true" pageId="14" pageNumber="85">Larval biology:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="FDB3667E0D22FF9BEFC4F4E43142F94E" ID-DOI="http://doi.org/10.5281/zenodo.4401284" ID-Zenodo-Dep="4401284" httpUri="https://zenodo.org/record/4401284/files/figure.png" pageId="14" pageNumber="85" startId="14.[151,260,1433,1459]" targetBox="[173,1410,616,1401]" targetPageId="14">
<paragraph id="A97336F60D22FF9BEFC4F4E430BDF9BB" blockId="14.[151,1437,1433,1712]" pageId="14" pageNumber="85">
<emphasis id="9BB8EAE40D22FF9BEFC4F4E4317CFA4A" bold="true" box="[151,287,1433,1460]" pageId="14" pageNumber="85">FIGURE 7.</emphasis>
<taxonomicName id="6ECC4D750D22FF9BEE77F4E732E5FA4A" authorityName="Simonsen" authorityYear="2018" box="[292,646,1434,1460]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D22FF9BEE77F4E732E5FA4A" box="[292,646,1434,1460]" italics="true" pageId="14" pageNumber="85">Abantiades macropusinsulariae</emphasis>
</taxonomicName>
. (SA, Kangaroo Island, North Cape Road, Bay of Shoals, M+M Moore, 35
<superScript id="5EB99BBE0D22FF9BEFE0F4C330D8FA32" attach="none" box="[179,187,1470,1484]" fontSize="6" pageId="14" pageNumber="85">o</superScript>
3547.7”S 137
<superScript id="5EB99BBE0D22FF9BEE32F4C3310AFA32" attach="none" box="[353,361,1470,1484]" fontSize="6" pageId="14" pageNumber="85">o</superScript>
3540.7”E, 2 May 2019. Specimen No 19169. Leg removed for DNA analysis; Sample designation G116. Dissected M.D. Moore 12
<superScript id="5EB99BBE0D22FF9BED09F49C3204FA11" attach="left" box="[602,615,1505,1519]" fontSize="6" pageId="14" pageNumber="85">th</superScript>
Feb. 2020. SAMA No. 31-021005.)
<emphasis id="9BB8EAE40D22FF9BEB5FF49C344EFA05" bold="true" box="[1036,1069,1505,1531]" pageId="14" pageNumber="85">(a)</emphasis>
posterior view female genitalia,
<emphasis id="9BB8EAE40D22FF9BEFC4F77830D9F9E1" bold="true" box="[151,186,1541,1567]" pageId="14" pageNumber="85">(b)</emphasis>
corpus bursae and ductus bursae,
<emphasis id="9BB8EAE40D22FF9BED15F7783205F9E1" bold="true" box="[582,614,1541,1567]" pageId="14" pageNumber="85">(c)</emphasis>
cruciform pectination (lateral view),
<emphasis id="9BB8EAE40D22FF9BEB46F7783454F9E1" bold="true" box="[1045,1079,1541,1567]" pageId="14" pageNumber="85">(d)</emphasis>
cruciform pectinations (frontal view).
</paragraph>
<paragraph id="A97336F60D22FF9BEFC4F7333142F94E" blockId="14.[151,1437,1433,1712]" pageId="14" pageNumber="85">
Morphological structures:
<emphasis id="9BB8EAE40D22FF9BEE97F7303196F999" bold="true" box="[452,501,1613,1639]" pageId="14" pageNumber="85">CB:</emphasis>
corpus bursae;
<emphasis id="9BB8EAE40D22FF9BEDF9F73332B4F996" bold="true" box="[682,727,1614,1640]" pageId="14" pageNumber="85">Db:</emphasis>
ductus bursae;
<emphasis id="9BB8EAE40D22FF9BECDFF73333DAF996" bold="true" box="[908,953,1614,1640]" pageId="14" pageNumber="85">Dp:</emphasis>
dorsal plate;
<emphasis id="9BB8EAE40D22FF9BEB05F73334EBF996" bold="true" box="[1110,1160,1614,1640]" pageId="14" pageNumber="85">Lal:</emphasis>
lateral lobe of the antevaginal lamella;
<emphasis id="9BB8EAE40D22FF9BEE01F70F31E8F972" bold="true" box="[338,395,1650,1676]" pageId="14" pageNumber="85">Mal:</emphasis>
medial lobe of the antevaginal lamella;
<emphasis id="9BB8EAE40D22FF9BEC1FF70C33E7F975" bold="true" box="[844,900,1649,1675]" pageId="14" pageNumber="85">Sub:</emphasis>
sub-anal plate.
<emphasis id="9BB8EAE40D22FF9BEB66F70C34EEF975" bold="true" box="[1077,1165,1649,1675]" pageId="14" pageNumber="85">Arrow:</emphasis>
triangular projection on dorsal plate.
</paragraph>
</caption>
<subSubSection id="E1D6657D0D22FF9AEFC4F67F31BEFBEF" lastPageId="15" lastPageNumber="86" pageId="14" pageNumber="85" type="discussion">
<paragraph id="A97336F60D22FF9BEFC4F67F3170F8E2" blockId="14.[151,275,1794,1820]" box="[151,275,1794,1820]" pageId="14" pageNumber="85">
<heading id="F23B819A0D22FF9BEFC4F67F3170F8E2" bold="true" box="[151,275,1794,1820]" fontSize="11" level="1" pageId="14" pageNumber="85" reason="1">
<emphasis id="9BB8EAE40D22FF9BEFC4F67F3170F8E2" bold="true" box="[151,275,1794,1820]" pageId="14" pageNumber="85">Remarks:</emphasis>
</heading>
</paragraph>
<paragraph id="A97336F60D22FF9BEFC4F63734BCF852" blockId="14.[151,1437,1865,2036]" pageId="14" pageNumber="85">
The female specimen (
<accessionNumber id="B69FAB150D22FF9BEEC5F634327BF89A" box="[406,536,1865,1892]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT052730" pageId="14" pageNumber="85">MT052730</accessionNumber>
) in
<figureCitation id="31F72A730D22FF9BED10F63732E6F89A" box="[579,645,1866,1892]" captionStart="FIGURE 1" captionStartId="3.[151,260,1815,1841]" captionTargetBox="[151,1427,283,1786]" captionTargetId="figure-67@3.[151,1436,270,1791]" captionTargetPageId="3" captionText="FIGURE 1. Maximum credibility tree from Bayesian phylogenetic analysis of the mitochondrial DNA COI gene of all 42 Abantiades sequences used and the ♀ specimens for A. macropusinsulariae and A. pallida. Posterior probabilities greater than 0.7 are shown at each node. Details for all sequences including GenBank accessions are available in Table S1." figureDoi="http://doi.org/10.5281/zenodo.4401270" httpUri="https://zenodo.org/record/4401270/files/figure.png" pageId="14" pageNumber="85">Fig. 1</figureCitation>
was initially identified as
<taxonomicName id="6ECC4D750D22FF9BECFDF63734CFF89A" authorityName="Simonsen" authorityYear="2018" box="[942,1196,1866,1892]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D22FF9BECFDF63733DCF89A" box="[942,959,1866,1892]" italics="true" pageId="14" pageNumber="85">A</emphasis>
.
<emphasis id="9BB8EAE40D22FF9BEC9EF63734CFF89A" box="[973,1196,1866,1892]" italics="true" pageId="14" pageNumber="85">macropusinsulariae</emphasis>
</taxonomicName>
by sequencing of the mtDNA (COI) gene, and groups with the four other male
<taxonomicName id="6ECC4D750D22FF9BEC40F6133473F876" authorityName="Simonsen" authorityYear="2018" box="[787,1040,1902,1928]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D22FF9BEC40F6133473F876" box="[787,1040,1902,1928]" italics="true" pageId="14" pageNumber="85">A. macropusinsulariae</emphasis>
</taxonomicName>
specimens. All five
<taxonomicName id="6ECC4D750D22FF9BEBA1F6133090F852" authorityName="Simonsen" authorityYear="2018" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D22FF9BEBA1F6133090F852" italics="true" pageId="14" pageNumber="85">A. macropusinsulariae</emphasis>
</taxonomicName>
specimens are closely related and form a sister group to
<taxonomicName id="6ECC4D750D22FF9BEC23F6EF33BDF852" authorityName="Simonsen" authorityYear="2018" box="[880,990,1938,1964]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D22FF9BEC23F6EF33BDF852" box="[880,990,1938,1964]" italics="true" pageId="14" pageNumber="85">A. pallida</emphasis>
</taxonomicName>
and
<taxonomicName id="6ECC4D750D22FF9BEB46F6EF341CF852" authority="Moore &amp; Beaver &amp; Velasco-Castrillón &amp; Stevens, 2020" authorityName="Moore &amp; Beaver &amp; Velasco-Castrillón &amp; Stevens" authorityYear="2020" box="[1045,1151,1938,1964]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="tembyi" status="sp. nov.">
<emphasis id="9BB8EAE40D22FF9BEB46F6EF341CF852" box="[1045,1151,1938,1964]" italics="true" pageId="14" pageNumber="85">A. tembyi</emphasis>
</taxonomicName>
<emphasis id="9BB8EAE40D22FF9BEBD6F6EF34BCF852" bold="true" box="[1157,1247,1938,1964]" pageId="14" pageNumber="85">
<taxonomicNameLabel id="808B579F0D22FF9BEBD6F6EF34BCF852" box="[1157,1247,1938,1964]" pageId="14" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="A97336F60D22FF9AEF94F6CB3391FE77" blockId="14.[151,1437,1865,2036]" lastBlockId="15.[151,1437,151,1041]" lastPageId="15" lastPageNumber="86" pageId="14" pageNumber="85">
Comparison with the females of
<taxonomicName id="6ECC4D750D22FF9BED66F6CB32DDF831" authorityName="Simonsen" authorityYear="2018" box="[565,702,1974,2000]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="inexpecta">
<emphasis id="9BB8EAE40D22FF9BED66F6CB32DDF831" box="[565,702,1974,2000]" italics="true" pageId="14" pageNumber="85">A. inexpecta</emphasis>
</taxonomicName>
,
<taxonomicName id="6ECC4D750D22FF9BED9EF6CB33EDF82E" authority="Moore &amp; Beaver &amp; Velasco-Castrillón &amp; Stevens, 2020" authorityName="Moore &amp; Beaver &amp; Velasco-Castrillón &amp; Stevens" authorityYear="2020" box="[717,910,1974,2000]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="cephalocorvus" status="sp. nov.">
<emphasis id="9BB8EAE40D22FF9BED9EF6CB33EDF82E" box="[717,910,1974,2000]" italics="true" pageId="14" pageNumber="85">A. cephalocorvus</emphasis>
</taxonomicName>
<emphasis id="9BB8EAE40D22FF9BECC5F6CB338CF82E" bold="true" box="[918,1007,1974,2000]" pageId="14" pageNumber="85">
<taxonomicNameLabel id="808B579F0D22FF9BECC5F6CB338CF82E" box="[918,1007,1974,2000]" pageId="14" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="6ECC4D750D22FF9BEB7FF6CB34F5F82E" authority="Moore &amp; Beaver &amp; Velasco-Castrillón &amp; Stevens, 2020" authorityName="Moore &amp; Beaver &amp; Velasco-Castrillón &amp; Stevens" authorityYear="2020" box="[1068,1174,1974,2000]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="tembyi" status="sp. nov.">
<emphasis id="9BB8EAE40D22FF9BEB7FF6CB34F5F82E" box="[1068,1174,1974,2000]" italics="true" pageId="14" pageNumber="85">A. tembyi</emphasis>
</taxonomicName>
<emphasis id="9BB8EAE40D22FF9BEBCFF6CB3495F82E" bold="true" box="[1180,1270,1974,2000]" pageId="14" pageNumber="85">
<taxonomicNameLabel id="808B579F0D22FF9BEBCFF6CB3495F82E" box="[1180,1270,1974,2000]" pageId="14" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is not possible because none have been located in collections. We can compare the females of the three remaining species if we use
<bibRefCitation id="CD5D4B070D23FF9AEFC4F1EA3134FF4F" author="Simonsen, T. J." box="[151,343,151,177]" pageId="15" pageNumber="86" pagination="1 - 300" refId="ref12387" refString="Simonsen, T. J. (2018) Splendid Ghost Moths and Their Allies. Monographs on Australian Lepidoptera, 12, i-xii + 1 - 300. https: // doi. org / 10.1071 / 9781486307487" type="journal article" year="2018">Simonsen (2018)</bibRefCitation>
for the features of
<taxonomicName id="6ECC4D750D23FF9AED7AF1EA32A3FF4F" authorityName="Simonsen" authorityYear="2018" box="[553,704,151,177]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="9BB8EAE40D23FF9AED7AF1EA32A3FF4F" box="[553,704,151,177]" italics="true" pageId="15" pageNumber="86">A. kristenseni</emphasis>
</taxonomicName>
. In
<taxonomicName id="6ECC4D750D23FF9AEDB8F1EA338AFF4F" authorityName="Simonsen" authorityYear="2018" box="[747,1001,151,177]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D23FF9AEDB8F1EA338AFF4F" box="[747,1001,151,177]" italics="true" pageId="15" pageNumber="86">A. macropusinsulariae</emphasis>
</taxonomicName>
the structure of the genitalia is particularly interesting as the heavily sclerotized bands bordering the medial lobe of antevaginal lamella are not present in any other as yet dissected
<taxonomicName id="6ECC4D750D23FF9AEEB3F1A2323DFF07" authorityName="Herrich-Schaffer" authorityYear="1855" box="[480,606,223,249]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="9BB8EAE40D23FF9AEEB3F1A2323DFF07" box="[480,606,223,249]" italics="true" pageId="15" pageNumber="86">Abantiades</emphasis>
</taxonomicName>
female, triforked or otherwise. These bands presumably provide rigidity. In our opinion this provides a three dimensionality to the structure not seen in any other triforked
<taxonomicName id="6ECC4D750D23FF9AEBB6F07E3500FEE3" authorityName="Herrich-Schaffer" authorityYear="1855" box="[1253,1379,259,285]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="9BB8EAE40D23FF9AEBB6F07E3500FEE3" box="[1253,1379,259,285]" italics="true" pageId="15" pageNumber="86">Abantiades</emphasis>
</taxonomicName>
. The only other female
<taxonomicName id="6ECC4D750D23FF9AEE35F05A3187FEBF" authorityName="Herrich-Schaffer" authorityYear="1855" box="[358,484,295,321]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="9BB8EAE40D23FF9AEE35F05A3187FEBF" box="[358,484,295,321]" italics="true" pageId="15" pageNumber="86">Abantiades</emphasis>
</taxonomicName>
that shows any marked three dimensionality in its structure is
<taxonomicName id="6ECC4D750D23FF9AEBFBF05A30B0FE9B" authority="Moore, 2014" authorityName="Moore" authorityYear="2014" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="equipalpus">
<emphasis id="9BB8EAE40D23FF9AEBFBF05A3520FEBF" box="[1192,1347,295,321]" italics="true" pageId="15" pageNumber="86">A. equipalpus</emphasis>
<bibRefCitation id="CD5D4B070D23FF9AEA18F05A30B0FE9B" author="Moore M. D." pageId="15" pageNumber="86" pagination="217 - 231" refId="ref12224" refString="Moore M. D. (2014) Two further new species of Abantiades Herrich-Schaffer (Lepidoptera: Hepialidae) from Western Australia. Australian Entomologist, 41 (4), 217 - 231." type="journal article" year="2014">Moore, 2014</bibRefCitation>
</taxonomicName>
. The corpus bursae is also unique in having a long taper at its distal end, that was particularly notable upon first opening the abdomen, the taper being bent forward over the whole structure.
</paragraph>
<paragraph id="A97336F60D23FF9AEF94F0EE34D9FCC7" blockId="15.[151,1437,151,1041]" pageId="15" pageNumber="86">
<taxonomicName id="6ECC4D750D23FF9AEF94F0EE324DFE53" authorityName="Simonsen" authorityYear="2018" box="[199,558,403,429]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D23FF9AEF94F0EE324DFE53" box="[199,558,403,429]" italics="true" pageId="15" pageNumber="86">Abantiades macropusinsulariae</emphasis>
</taxonomicName>
is a dark grey/black coloured moth with distinct discoidal and submarginal white marks whereas both
<taxonomicName id="6ECC4D750D23FF9AEE90F0CA3238FE2F" authorityName="Simonsen" authorityYear="2018" box="[451,603,439,465]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="9BB8EAE40D23FF9AEE90F0CA3238FE2F" box="[451,603,439,465]" italics="true" pageId="15" pageNumber="86">A. kristenseni</emphasis>
</taxonomicName>
and
<taxonomicName id="6ECC4D750D23FF9AEDC2F0CA329CFE2F" authorityName="Simonsen" authorityYear="2018" box="[657,767,439,465]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D23FF9AEDC2F0CA329CFE2F" box="[657,767,439,465]" italics="true" pageId="15" pageNumber="86">A. pallida</emphasis>
</taxonomicName>
are light grey in colour. In the specimens available it seems that
<taxonomicName id="6ECC4D750D23FF9AEF9AF0A63102FE0B" authorityName="Simonsen" authorityYear="2018" box="[201,353,475,501]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="9BB8EAE40D23FF9AEF9AF0A63102FE0B" box="[201,353,475,501]" italics="true" pageId="15" pageNumber="86">A. kristenseni</emphasis>
</taxonomicName>
will usually have two large marks on the forewings, discoidal and submarginal, whereas
<taxonomicName id="6ECC4D750D23FF9AEA03F0A630A0FDE7" authorityName="Simonsen" authorityYear="2018" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D23FF9AEA03F0A630A0FDE7" italics="true" pageId="15" pageNumber="86">A. pallida</emphasis>
</taxonomicName>
will only have a submarginal mark. Upon dissection
<taxonomicName id="6ECC4D750D23FF9AEC44F0823476FDE7" authorityName="Simonsen" authorityYear="2018" box="[791,1045,511,537]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D23FF9AEC44F0823476FDE7" box="[791,1045,511,537]" italics="true" pageId="15" pageNumber="86">A. macropusinsulariae</emphasis>
</taxonomicName>
stands out having a narrow medial lobe to the antevaginal lamella edged in thick sclerotization, more regularly shaped, more rounded and more widely separated lobes on the dorsal plate and sub-anal plates that are double the area of the other two species.
<taxonomicName id="6ECC4D750D23FF9AEA4DF33A3085FD7B" authorityName="Simonsen" authorityYear="2018" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D23FF9AEA4DF33A3085FD7B" italics="true" pageId="15" pageNumber="86">Abantiades pallida</emphasis>
</taxonomicName>
and
<taxonomicName id="6ECC4D750D23FF9AEE73F31631D9FD7B" authorityName="Simonsen" authorityYear="2018" box="[288,442,619,645]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="9BB8EAE40D23FF9AEE73F31631D9FD7B" box="[288,442,619,645]" italics="true" pageId="15" pageNumber="86">A. kristenseni</emphasis>
</taxonomicName>
both have shallow rounded lobes on the dorsal plate and no thick sclerotization on the medial lobe. Also, the medial lobe of the antevaginal lamella is over triple the width in
<taxonomicName id="6ECC4D750D23FF9AEB33F3F2349BFD57" authorityName="Simonsen" authorityYear="2018" box="[1120,1272,655,681]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="9BB8EAE40D23FF9AEB33F3F2349BFD57" box="[1120,1272,655,681]" italics="true" pageId="15" pageNumber="86">A. kristenseni</emphasis>
</taxonomicName>
and the lateral processes on the inner margin are triangular and pointed whereas in
<taxonomicName id="6ECC4D750D23FF9AECC0F3CE3461FD33" authorityName="Simonsen" authorityYear="2018" box="[915,1026,691,717]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D23FF9AECC0F3CE3461FD33" box="[915,1026,691,717]" italics="true" pageId="15" pageNumber="86">A. pallida</emphasis>
</taxonomicName>
where the medial lobe is a little narrower than
<taxonomicName id="6ECC4D750D23FF9AEE4AF3AA31D1FD0F" authorityName="Simonsen" authorityYear="2018" box="[281,434,727,753]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="9BB8EAE40D23FF9AEE4AF3AA31D1FD0F" box="[281,434,727,753]" italics="true" pageId="15" pageNumber="86">A. kristenseni</emphasis>
</taxonomicName>
(but still triple the width of
<taxonomicName id="6ECC4D750D23FF9AEDABF3AA3394FD0F" authorityName="Simonsen" authorityYear="2018" box="[760,1015,727,753]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D23FF9AEDABF3AA336AFD0F" box="[760,777,727,753]" italics="true" pageId="15" pageNumber="86">A</emphasis>
.
<emphasis id="9BB8EAE40D23FF9AEC4BF3AA3394FD0F" box="[792,1015,727,753]" italics="true" pageId="15" pageNumber="86">macropusinsulariae</emphasis>
</taxonomicName>
) the lateral processes are ten percent longer, trapezoidal in form and blunt ended. The setose nipple like eruptions (
<figureCitation id="31F72A730D23FF9AECA6F3863454FCEB" box="[1013,1079,763,789]" captionStart="FIGURE 9" captionStartId="17.[151,260,1702,1728]" captionTargetBox="[182,1411,1189,1660]" captionTargetId="figure-385@17.[151,1437,1170,1678]" captionTargetPageId="17" captionText="FIGURE 9. Abantiades pallida. (West.Aust. Nullarbor, Quarantine Station, Border Village, 31o3819”S, 129o07”E, R. Penning 7 May 18. Spec. No. 18122, leg removed, for tissue, storage. M.D. Moore. Sample designation No. H 122. SAMA No. 31-020414. Dissected E.P. Beaver.) (left) posterior view female genitalia, (right) lateral view female genitalia. Morphological structures: Dp: dorsal plate; Lal: lateral lobe of the antevaginal lamella; Mal: medial lobe of the antevaginal lamella; Sub: sub-anal plate. Arrow: Setose nipple." figureDoi="http://doi.org/10.5281/zenodo.4401288" httpUri="https://zenodo.org/record/4401288/files/figure.png" pageId="15" pageNumber="86">Fig. 9</figureCitation>
arrow)) on the lateral lobes of the antevaginal lamella seen in
<taxonomicName id="6ECC4D750D23FF9AEEA4F2623206FCC7" authorityName="Simonsen" authorityYear="2018" box="[503,613,799,825]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D23FF9AEEA4F2623206FCC7" box="[503,613,799,825]" italics="true" pageId="15" pageNumber="86">A. pallida</emphasis>
</taxonomicName>
has not been seen on either of the two other females.
</paragraph>
<paragraph id="A97336F60D23FF9AEF94F23E31BEFBEF" blockId="15.[151,1437,151,1041]" pageId="15" pageNumber="86">
Endemic to Kangaroo Island,
<taxonomicName id="6ECC4D750D23FF9AED41F23E336CFCA3" authorityName="Simonsen" authorityYear="2018" box="[530,783,835,861]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D23FF9AED41F23E336CFCA3" box="[530,783,835,861]" italics="true" pageId="15" pageNumber="86">A. macropusinsulariae</emphasis>
</taxonomicName>
can be separated from the other two triforked species present on the island by its antennae.
<taxonomicName id="6ECC4D750D23FF9AED4AF21A331DFC7F" authorityName="Simonsen" authorityYear="2018" box="[537,894,871,897]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D23FF9AED4AF21A331DFC7F" box="[537,894,871,897]" italics="true" pageId="15" pageNumber="86">Abantiades macropusinsulariae</emphasis>
</taxonomicName>
has a dark red flagellum, distinctly three lobed pectinations with a long, pointed pale central fork.
<taxonomicName id="6ECC4D750D23FF9AED85F2F6341BFC5B" authority="(Walker, 1856)" baseAuthorityName="Walker" baseAuthorityYear="1856" box="[726,1144,906,933]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="atripalpis">
<emphasis id="9BB8EAE40D23FF9AED85F2F633ABFC5B" box="[726,968,907,933]" italics="true" pageId="15" pageNumber="86">Abantiades atripalpis</emphasis>
(
<bibRefCitation id="CD5D4B070D23FF9AEC84F2F6340CFC5B" author="Walker, F." box="[983,1135,906,933]" pageId="15" pageNumber="86" pagination="1508 - 1808" refId="ref12657" refString="Walker, F. (1856) List of the specimens of Lepidopterous Insects in the collection of the British Museum. Lepidoptera Heterocera, 7, 1508 - 1808. [incl. Index to parts 1 - 7]" type="journal article" year="1856">Walker, 1856</bibRefCitation>
)
</taxonomicName>
antennae are tan/russet in colour, pectinations are more rhomboidal and with a shorter central fork, and scrolling lines that are fine and indistinct.
<taxonomicName id="6ECC4D750D23FF9AEF84F2AE31ABFC12" baseAuthorityName="Tindale" baseAuthorityYear="1932" box="[215,456,979,1005]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="argentata">
<emphasis id="9BB8EAE40D23FF9AEF84F2AE31ABFC12" box="[215,456,979,1005]" italics="true" pageId="15" pageNumber="86">Abantiades argentata</emphasis>
</taxonomicName>
has dark brown, black rami similar in shape to
<taxonomicName id="6ECC4D750D23FF9AEC8FF2AE340BFC13" baseAuthorityName="Walker" baseAuthorityYear="1856" box="[988,1128,979,1005]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="atripalpis">
<emphasis id="9BB8EAE40D23FF9AEC8FF2AE340BFC13" box="[988,1128,979,1005]" italics="true" pageId="15" pageNumber="86">A. atripalpis</emphasis>
</taxonomicName>
, and brown forewings with more obvious scrolling lines.
</paragraph>
</subSubSection>
</treatment>
</document>