255 lines
36 KiB
XML
255 lines
36 KiB
XML
<document id="091A761AC0F877C9B53023CB32CB861C" ID-CLB-Dataset="229105" ID-DOI="10.11646/zootaxa.5230.2.3" ID-GBIF-Dataset="997e16f6-a408-4411-93cc-53fa1b085c92" ID-ISSN="1175-5326" ID-Zenodo-Dep="7553485" ID-ZooBank="7D2F9E0B-9E32-40E4-8DED-9E6227BAEFD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1674200759302" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2023" docId="1F070A1EC6743C58A1DCFC17920EFD89" docLanguage="en" docName="zootaxa.5230.2.3.pdf" docOrigin="Zootaxa 5230 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyops confusus Sokolov 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="185" masterDocId="E33E7266C6713C5EA14BFFB9901CFF90" masterDocTitle="Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island" masterLastPageNumber="201" masterPageNumber="179" pageNumber="184" updateTime="1699474157326" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="404A6AF44E795CA6C506278A903F8104" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="82D07D7F52B023D2E8DD1F87C14EA22E">
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<mods:title id="773ACD7C764563ED53A0AAC5EE6E93F0">Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island</mods:title>
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</mods:titleInfo>
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<mods:name id="723158114040B2AC189C4B6D89D70904" type="personal">
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<mods:role id="6232AF72F89445777FE46B8485AE3F43">
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<mods:roleTerm id="55BABCB0EF12DD311299AE48BF2F480E">Author</mods:roleTerm>
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<mods:namePart id="958793FF9101EE052EDB82201C3CEBF9">Sokolov, Igor M.</mods:namePart>
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<mods:typeOfResource id="3A8B5509EFF7677C8533AC6A55116676">text</mods:typeOfResource>
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<mods:title id="3D400DDAF60FBCFDA2BCDBB77784BBBD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AAF46BA5A2A6E9C84423E9938FF86734">
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<mods:date id="58F5C61A6430F81CDA11B7B8B72EB650">2023</mods:date>
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<mods:number id="630B3DD09979B046F5F024921D83917B">2023-01-20</mods:number>
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<mods:number id="5691B3B277EA7D58FCB897CFC5A8B64D">5230</mods:number>
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<mods:detail id="8FB5DEA0B1B23E4D83406548B4688CC2" type="issue">
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<mods:number id="93F95E93959F64BC09CD49F48235650E">2</mods:number>
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<mods:start id="2FC243083A04D55520FA918631030653">179</mods:start>
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<mods:classification id="8448F75E4CCAABD6D32C3D97D8FE17B8">journal article</mods:classification>
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<mods:identifier id="17B692E42FA34495D59AFCF41AF00956" type="DOI">10.11646/zootaxa.5230.2.3</mods:identifier>
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<mods:identifier id="CA75BAB1608E1407F6FCA78E3185B546" type="GBIF-Dataset">997e16f6-a408-4411-93cc-53fa1b085c92</mods:identifier>
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<treatment id="1F070A1EC6743C58A1DCFC17920EFD89" ID-DOI="http://doi.org/10.5281/zenodo.7553493" ID-GBIF-Taxon="209011116" ID-Zenodo-Dep="7553493" LSID="urn:lsid:plazi:treatment:1F070A1EC6743C58A1DCFC17920EFD89" httpUri="http://treatment.plazi.org/id/1F070A1EC6743C58A1DCFC17920EFD89" lastPageId="6" lastPageNumber="185" pageId="5" pageNumber="184">
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<subSubSection id="DFB4E883C6743C5BA1DCFC179204FC59" box="[151,536,942,969]" pageId="5" pageNumber="184" type="nomenclature">
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<paragraph id="9711BB08C6743C5BA1DCFC179204FC59" blockId="5.[151,536,942,1005]" box="[151,536,942,969]" pageId="5" pageNumber="184">
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<heading id="CC590C64C6743C5BA1DCFC179204FC59" bold="true" box="[151,536,942,969]" fontSize="11" level="1" pageId="5" pageNumber="184" reason="1">
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<emphasis id="A5DA671AC6743C5BA1DCFC179204FC59" bold="true" box="[151,536,942,969]" pageId="5" pageNumber="184">
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<taxonomicName id="50AEC08BC6743C5BA1DCFC1791B7FC58" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" box="[151,427,942,969]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="confusus" status="sp. nov.">
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<emphasis id="A5DA671AC6743C5BA1DCFC1791B7FC58" bold="true" box="[151,427,942,969]" italics="true" pageId="5" pageNumber="184">Nesamblyops confusus</emphasis>
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="BEE9DA61C6743C5BA0F1FC169204FC59" box="[442,536,943,969]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="DFB4E883C6743C5BA1DCFC6A918AFC7D" box="[151,406,978,1005]" pageId="5" pageNumber="184" type="description">
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<paragraph id="9711BB08C6743C5BA1DCFC6A918AFC7D" blockId="5.[151,536,942,1005]" box="[151,406,978,1005]" pageId="5" pageNumber="184">
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<figureCitation id="0F95A78DC6743C5BA1DCFC6A90E4FC7D" box="[151,248,979,1005]" captionStart="FIGURE 2" captionStartId="7.[151,250,1692,1716]" captionTargetBox="[151,1423,154,1661]" captionTargetId="figure-172@7.[151,1436,151,1669]" captionTargetPageId="7" captionText="FIGURE 2. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO). Scale bar 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7553491" httpUri="https://zenodo.org/record/7553491/files/figure.png" pageId="5" pageNumber="184">Figs. 2C</figureCitation>
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,
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<figureCitation id="0F95A78DC6743C5BA04FFC6A913BFC7D" box="[260,295,979,1005]" captionStart="FIGURE 4" captionStartId="10.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1197]" captionTargetId="figure-28@10.[151,1436,181,1198]" captionTargetPageId="10" captionText="FIGURE 4. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO), G—N. parvulus (Mount Stokes, Marlborough Sounds, SO), H—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), I—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale bar 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7553497" httpUri="https://zenodo.org/record/7553497/files/figure.png" pageId="5" pageNumber="184">4C</figureCitation>
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,
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<figureCitation id="0F95A78DC6743C5BA078FC6B9172FC7D" box="[307,366,978,1005]" captionStart="FIGURE 5" captionStartId="11.[151,250,1558,1582]" captionTargetBox="[161,1423,196,1523]" captionTargetId="figure-16@11.[150,1437,181,1535]" captionTargetPageId="11" captionText="FIGURE 5. Line drawings of male genitalia of New Zealand Nesamblyops species. N. brouni (Lewis Pass, North Canterbury, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. carltoni (Dun Mountain, Nelson, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. confusus (Mount Stokes, Marlborough Sounds, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. distinctus (Fabians Valley, Marlborough, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. oreobius (Mount Pirongia, Waikato, NO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7553501" httpUri="https://zenodo.org/record/7553501/files/figure.png" pageId="5" pageNumber="184">5G–I</figureCitation>
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,
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<figureCitation id="0F95A78DC6743C5BA031FC6A918AFC7D" box="[378,406,979,1005]" captionStart="FIGURE 10" captionStartId="18.[151,250,1698,1722]" captionTargetBox="[195,1404,151,1693]" captionTargetId="figure-120@18.[195,1404,151,1695]" captionTargetPageId="18" captionText="FIGURE 10. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Black circles—N. carltoni; yellow circles—N. parvulus; red circles—N. lescheni; green circles—N. confusus. Scale bar= 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7553515" httpUri="https://zenodo.org/record/7553515/files/figure.png" pageId="5" pageNumber="184">10</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="DFB4E883C6743C5BA1DCFBA29136FB31" pageId="5" pageNumber="184" type="materials_examined">
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<paragraph id="9711BB08C6743C5BA1DCFBA29136FB31" blockId="5.[151,1436,1050,2013]" pageId="5" pageNumber="184">
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||
<emphasis id="A5DA671AC6743C5BA1DCFBA29154FBA4" bold="true" box="[151,328,1050,1077]" pageId="5" pageNumber="184">Type material.</emphasis>
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||
<materialsCitation id="27C6B155C6743C5BA01AFBA292C5FBC9" ID-GBIF-Occurrence="4016386383" collectionCode="NZAC, DSIR" collectorName="J. I. Townsend" country="New Zealand" elevation="1160" location="Mt Stokes" pageId="5" pageNumber="184" specimenCount="1" stateProvince="Marlborough" typeStatus="holotype">
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<emphasis id="A5DA671AC6743C5BA01AFBA291FAFBA5" bold="true" box="[337,486,1051,1077]" italics="true" pageId="5" pageNumber="184">
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||
<typeStatus id="481505AAC6743C5BA01AFBA291FAFBA5" box="[337,486,1051,1077]" pageId="5" pageNumber="184" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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, male, in
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<collectionCode id="F1BF23CDC6743C5BA316FBA292B0FBA5" box="[605,684,1051,1077]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="184" type="Other institutional type">NZAC</collectionCode>
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||
, dissected, labeled: \
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||
<location id="9271EDD3C6743C5BA2EEFBA29406FBA5" LSID="urn:lsid:plazi:treatment:1F070A1EC6743C58A1DCFC17920EFD89:9271EDD3C6743C5BA2EEFBA29406FBA5" box="[933,1050,1051,1077]" country="New Zealand" name="Mt Stokes" pageId="5" pageNumber="184" stateProvince="Marlborough">Mt Stokes</location>
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||
<quantity id="505616EDC6743C5BA56FFBA2946CFBA5" box="[1060,1136,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="5" pageNumber="184" unit="m" value="1160.0">
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<elevation id="1C835C3BC6743C5BA56FFBA2946CFBA5" box="[1060,1136,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="5" pageNumber="184" unit="m" value="1160.0">1160m</elevation>
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||
</quantity>
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||
<collectingRegion id="556A75EAC6743C5BA531FBA29512FBA5" box="[1146,1294,1051,1077]" country="New Zealand" name="Marlborough" pageId="5" pageNumber="184">Marlborough</collectingRegion>
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||
12
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<collectorName id="3A5BDEDEC6743C5BA476FBA29579FBA5" box="[1341,1381,1051,1077]" pageId="5" pageNumber="184">Oct</collectorName>
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67 \
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<collectorName id="3A5BDEDEC6743C5BA1DCFB869130FBC9" box="[151,300,1087,1113]" pageId="5" pageNumber="184">J.I.Townsend</collectorName>
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\
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<collectorName id="3A5BDEDEC6743C5BA00CFB869199FBC9" box="[327,389,1087,1113]" pageId="5" pageNumber="184">Litter</collectorName>
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67/244 \
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<collectionCode id="F1BF23CDC6743C5BA0B3FB86922BFBC9" box="[504,567,1087,1113]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="5" pageNumber="184">DSIR</collectionCode>
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\
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<collectorName id="3A5BDEDEC6743C5BA302FB869277FBC9" box="[585,619,1087,1113]" pageId="5" pageNumber="184">No</collectorName>
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\
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<collectingCountry id="EFB9FB98C6743C5BA3CDFB8692B7FBC9" box="[646,683,1087,1113]" name="New Zealand" pageId="5" pageNumber="184">NZ</collectingCountry>
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PB
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</materialsCitation>
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\.
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<materialsCitation id="27C6B155C6743C5BA3A4FB86959CFBC9" ID-GBIF-Occurrence="4016386387" box="[751,1408,1087,1113]" collectionCode="NZAC" country="New Zealand" pageId="5" pageNumber="184" specimenCount="3" specimenCount-male="1" stateProvince="Marlborough" typeStatus="paratype">
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<emphasis id="A5DA671AC6743C5BA3A4FB869396FBC9" bold="true" box="[751,906,1087,1113]" italics="true" pageId="5" pageNumber="184">
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<typeStatus id="481505AAC6743C5BA3A4FB869396FBC9" box="[751,906,1087,1113]" pageId="5" pageNumber="184" type="paratype">PARATYPES</typeStatus>
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||
</emphasis>
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||
(
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<specimenCount id="81A87081C6743C5BA2D7FB869431FBC9" box="[924,1069,1087,1113]" pageId="5" pageNumber="184" type="generic">2 specimens</specimenCount>
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||
, in
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<collectionCode id="F1BF23CDC6743C5BA512FB8694B4FBC9" box="[1113,1192,1087,1113]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="184" type="Other institutional type">NZAC</collectionCode>
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||
, dissected
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||
<specimenCount id="81A87081C6743C5BA46DFB869565FBC9" box="[1318,1401,1087,1113]" pageId="5" pageNumber="184" type="male">1 male</specimenCount>
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||
)
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||
</materialsCitation>
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||
,
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<specimenCount id="81A87081C6743C5BA4C5FB8690FFFBED" pageId="5" pageNumber="184" type="female">1 female</specimenCount>
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||
labeled same as
|
||
<typeStatus id="481505AAC6743C5BA0EFFBDA9216FBED" box="[420,522,1123,1149]" pageId="5" pageNumber="184" type="holotype">holotype</typeStatus>
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||
;
|
||
<specimenCount id="81A87081C6743C5BA35DFBDA927DFBED" box="[534,609,1123,1149]" pageId="5" pageNumber="184" type="male">1 male</specimenCount>
|
||
<materialsCitation id="27C6B155C6743C5BA322FBDA9100FB31" ID-GBIF-Occurrence="4016386354" collectionCode="DSIR" collectorName="J. I. Townsend & Ns" country="New Zealand" location="Mt Stokes Marlb." pageId="5" pageNumber="184" specimenCount="1" typeStatus="paratype">
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||
labeled: \
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||
<location id="9271EDD3C6743C5BA392FBDA93BCFBED" LSID="urn:lsid:plazi:treatment:1F070A1EC6743C58A1DCFC17920EFD89:9271EDD3C6743C5BA392FBDA93BCFBED" box="[729,928,1123,1149]" country="New Zealand" name="Mt Stokes Marlb." pageId="5" pageNumber="184">Mt Stokes Marlb.</location>
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||
11.x.67
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||
<collectorName id="3A5BDEDEC6743C5BA549FBDA948BFBED" box="[1026,1175,1123,1149]" pageId="5" pageNumber="184">J.I.Townsend</collectorName>
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\ moss 67/247 \
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||
<collectionCode id="F1BF23CDC6743C5BA41EFBDA9588FBED" box="[1365,1428,1123,1149]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="5" pageNumber="184">DSIR</collectionCode>
|
||
\
|
||
<collectorName id="3A5BDEDEC6743C5BA1DCFB3E90AAFB31" box="[151,182,1159,1185]" pageId="5" pageNumber="184">Ns</collectorName>
|
||
\
|
||
<collectingCountry id="EFB9FB98C6743C5BA187FB3E90EDFB31" box="[204,241,1159,1185]" name="New Zealand" pageId="5" pageNumber="184">NZ</collectingCountry>
|
||
PB
|
||
</materialsCitation>
|
||
\.
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="DFB4E883C6743C5BA18CFB129565FB79" pageId="5" pageNumber="184" type="etymology">
|
||
<paragraph id="9711BB08C6743C5BA18CFB129565FB79" blockId="5.[151,1436,1050,2013]" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA18CFB129152FB55" bold="true" box="[199,334,1195,1221]" pageId="5" pageNumber="184">Etymology.</emphasis>
|
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The specific epithet is a Latin adjective,
|
||
<taxonomicName id="50AEC08BC6743C5BA26FFB12939AFB55" authorityName="Sokolov" authorityYear="2023" box="[804,902,1195,1221]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="A5DA671AC6743C5BA26FFB12939AFB55" box="[804,902,1195,1221]" italics="true" pageId="5" pageNumber="184">confusus</emphasis>
|
||
</taxonomicName>
|
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, in the masculine form, meaning “having been mixed”, and refers to the similarity between the external appearance of the new species and its closest relatives.
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DFB4E883C6743C5BA18CFB4A91F1FAE9" pageId="5" pageNumber="184" type="materials_examined">
|
||
<paragraph id="9711BB08C6743C5BA18CFB4A9456FA9D" blockId="5.[151,1436,1050,2013]" box="[199,1098,1266,1293]" pageId="5" pageNumber="184">
|
||
<materialsCitation id="27C6B155C6743C5BA18CFB4A945AFA9D" ID-GBIF-Occurrence="4016386318" box="[199,1094,1266,1293]" country="New Zealand" county="South Island" location="Mount Stokes" municipality="Sounds" pageId="5" pageNumber="184" specimenCount="1" stateProvince="Marlborough" typeStatus="holotype">
|
||
<emphasis id="A5DA671AC6743C5BA18CFB4A917AFA9C" bold="true" box="[199,358,1266,1293]" pageId="5" pageNumber="184">
|
||
<typeStatus id="481505AAC6743C5BA18CFB4A911EFA9D" box="[199,258,1267,1293]" pageId="5" pageNumber="184">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="EFB9FB98C6743C5BA027FB4A9219FA9D" box="[364,517,1267,1293]" name="New Zealand" pageId="5" pageNumber="184">New Zealand</collectingCountry>
|
||
,
|
||
<collectingCounty id="7E70C384C6743C5BA35BFB4A92BCFA9D" box="[528,672,1267,1293]" pageId="5" pageNumber="184">South Island</collectingCounty>
|
||
,
|
||
<collectingRegion id="556A75EAC6743C5BA3E0FB4A9323FA9D" box="[683,831,1267,1293]" country="New Zealand" name="Marlborough" pageId="5" pageNumber="184">Marlborough</collectingRegion>
|
||
<collectingMunicipality id="77752172C6743C5BA20DFB4A9387FA9D" box="[838,923,1267,1293]" pageId="5" pageNumber="184">Sounds</collectingMunicipality>
|
||
,
|
||
<location id="9271EDD3C6743C5BA2EDFB4A945AFA9D" LSID="urn:lsid:plazi:treatment:1F070A1EC6743C58A1DCFC17920EFD89:9271EDD3C6743C5BA2EDFB4A945AFA9D" box="[934,1094,1267,1293]" country="New Zealand" county="South Island" municipality="Sounds" name="Mount Stokes" pageId="5" pageNumber="184" stateProvince="Marlborough">Mount Stokes</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6743C5BA18CFAAF91F1FAE9" blockId="5.[151,1436,1050,2013]" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA18CFAAF9141FAA0" bold="true" box="[199,349,1302,1328]" pageId="5" pageNumber="184">Recognition.</emphasis>
|
||
Adults of this species (
|
||
<figureCitation id="0F95A78DC6743C5BA321FAAE92DCFAA1" box="[618,704,1303,1329]" captionStart="FIGURE 2" captionStartId="7.[151,250,1692,1716]" captionTargetBox="[151,1423,154,1661]" captionTargetId="figure-172@7.[151,1436,151,1669]" captionTargetPageId="7" captionText="FIGURE 2. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO). Scale bar 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7553491" httpUri="https://zenodo.org/record/7553491/files/figure.png" pageId="5" pageNumber="184">Fig. 2C</figureCitation>
|
||
) are practically indistinguishable from the adults of many
|
||
<taxonomicName id="50AEC08BC6743C5BA410FAAE9153FAC5" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA410FAAE90E8FAC5" italics="true" pageId="5" pageNumber="184">Nesamblyops</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
based on external characters (cf. with
|
||
<figureCitation id="0F95A78DC6743C5BA25AFA8293D9FAC5" box="[785,965,1339,1365]" captionStart="FIGURE 2" captionStartId="7.[151,250,1692,1716]" captionTargetBox="[151,1423,154,1661]" captionTargetId="figure-172@7.[151,1436,151,1669]" captionTargetPageId="7" captionText="FIGURE 2. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO). Scale bar 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7553491" httpUri="https://zenodo.org/record/7553491/files/figure.png" pageId="5" pageNumber="184">Figs. 2A–B, 2F</figureCitation>
|
||
) and are distinguished from them by the structure of the male genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DFB4E883C6743C58A18CFA3B920EFD89" lastPageId="6" lastPageNumber="185" pageId="5" pageNumber="184" type="description">
|
||
<paragraph id="9711BB08C6743C5BA18CFA3B94C2FA0D" blockId="5.[151,1436,1050,2013]" box="[199,1246,1410,1437]" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA18CFA3B9145FA0C" bold="true" box="[199,345,1410,1436]" pageId="5" pageNumber="184">Description.</emphasis>
|
||
Average size for genus (SBL range
|
||
<quantity id="505616EDC6743C5BA3A5FA3A938FFA0D" box="[750,915,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="1.6549999999999998" metricValueMax="1.67" metricValueMin="1.64" pageId="5" pageNumber="184" unit="mm" value="1.6549999999999998" valueMax="1.67" valueMin="1.64">1.64–1.67 mm</quantity>
|
||
, mean 1.66±
|
||
<quantity id="505616EDC6743C5BA56AFA3A948AFA0D" box="[1057,1174,1411,1437]" metricMagnitude="-5" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="184" unit="mm" value="0.017">0.017 mm</quantity>
|
||
, n=3).
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6743C5BA18CFA1F92A9F999" blockId="5.[151,1436,1050,2013]" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA18CFA1F9136FA51" bold="true" box="[199,298,1446,1473]" italics="true" pageId="5" pageNumber="184">Habitus.</emphasis>
|
||
Body form (
|
||
<figureCitation id="0F95A78DC6743C5BA0FBFA1E921FFA51" box="[432,515,1447,1473]" captionStart="FIGURE 2" captionStartId="7.[151,250,1692,1716]" captionTargetBox="[151,1423,154,1661]" captionTargetId="figure-172@7.[151,1436,151,1669]" captionTargetPageId="7" captionText="FIGURE 2. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO). Scale bar 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7553491" httpUri="https://zenodo.org/record/7553491/files/figure.png" pageId="5" pageNumber="184">Fig.2C</figureCitation>
|
||
) markedly convex, elongate ovoid, general proportions wide (WE/SBL 0.42±0.012), head narrow relative to pronotum (WH/WPm 0.68±0.038), in comparison with other species pronotum less bulky in comparison to elytra (WPm/WE 0.72±0.027).
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6743C5BA18CF9AB9345F9BD" blockId="5.[151,1436,1050,2013]" box="[199,857,1554,1581]" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA18CF9AB9112F9BD" bold="true" box="[199,270,1554,1581]" italics="true" pageId="5" pageNumber="184">Color.</emphasis>
|
||
Body color dark brunneous, appendages testaceous.
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6743C5BA18CF98F90ECF971" blockId="5.[151,1436,1050,2013]" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA18CF98F9158F9C1" bold="true" box="[199,324,1590,1617]" italics="true" pageId="5" pageNumber="184">Prothorax.</emphasis>
|
||
Pronotum (
|
||
<figureCitation id="0F95A78DC6743C5BA080F98E923EF9C1" box="[459,546,1591,1617]" captionStart="FIGURE 4" captionStartId="10.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1197]" captionTargetId="figure-28@10.[151,1436,181,1198]" captionTargetPageId="10" captionText="FIGURE 4. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO), G—N. parvulus (Mount Stokes, Marlborough Sounds, SO), H—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), I—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale bar 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7553497" httpUri="https://zenodo.org/record/7553497/files/figure.png" pageId="5" pageNumber="184">Fig. 4C</figureCitation>
|
||
) moderately long in comparison to elytra (LP/LE 0.42±0.024) and moderately transverse (WPm/LP 1.24±0.013), with lateral margins arcuately and strongly constricted posteriorly (WPm/WPp 1.35±0.026). Anterior angles indistinct, posterior angles strongly obtuse (123–127°), widely rounded. Width between posterior angles slightly greater than between anterior angles (WPa/WPp 0.93±0.054). Basal margin slightly convex.
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6743C5BA18CF9539316F8DD" blockId="5.[151,1436,1050,2013]" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA18CF9539109F895" bold="true" box="[199,277,1770,1797]" italics="true" pageId="5" pageNumber="184">Elytra.</emphasis>
|
||
Ovoid, narrowly depressed along suture, comparatively long (LE/SBL 0.59±0.015) and moderately wide (WE/LE 0.71±0.034). Humeri completely rounded. Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third.
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6743C58A18CF8EF9325FF69" blockId="5.[151,1436,1050,2013]" lastBlockId="6.[151,1437,150,537]" lastPageId="6" lastPageNumber="185" pageId="5" pageNumber="184">
|
||
<emphasis id="A5DA671AC6743C5BA18CF8EF916BF8E1" bold="true" box="[199,375,1878,1905]" italics="true" pageId="5" pageNumber="184">Male genitalia.</emphasis>
|
||
Median lobe of aedeagus (
|
||
<figureCitation id="0F95A78DC6743C5BA3E5F8EE92E7F8E1" box="[686,763,1878,1905]" captionStart="FIGURE 5" captionStartId="11.[151,250,1558,1582]" captionTargetBox="[161,1423,196,1523]" captionTargetId="figure-16@11.[150,1437,181,1535]" captionTargetPageId="11" captionText="FIGURE 5. Line drawings of male genitalia of New Zealand Nesamblyops species. N. brouni (Lewis Pass, North Canterbury, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. carltoni (Dun Mountain, Nelson, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. confusus (Mount Stokes, Marlborough Sounds, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. distinctus (Fabians Valley, Marlborough, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. oreobius (Mount Pirongia, Waikato, NO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7553501" httpUri="https://zenodo.org/record/7553501/files/figure.png" pageId="5" pageNumber="184">Fig. 5I</figureCitation>
|
||
) moderately slightly arcuate and moderately twisted. Shaft arcuate, tapering in apical half. Apex slightly enlarged and straight with rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe slightly concave in apical part. Walls of shaft with few poriferous canals apically. Dorsal copulatory sclerites with distinct V-shaped structure, directed with its sharp end apically. Scaled membraneous fields lacking. Spines of internal sac absent. Left paramere (
|
||
<figureCitation id="0F95A78DC6773C58A5DEFF2E94F0FF21" box="[1173,1260,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1558,1582]" captionTargetBox="[161,1423,196,1523]" captionTargetId="figure-16@11.[150,1437,181,1535]" captionTargetPageId="11" captionText="FIGURE 5. Line drawings of male genitalia of New Zealand Nesamblyops species. N. brouni (Lewis Pass, North Canterbury, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. carltoni (Dun Mountain, Nelson, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. confusus (Mount Stokes, Marlborough Sounds, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. distinctus (Fabians Valley, Marlborough, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. oreobius (Mount Pirongia, Waikato, NO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7553501" httpUri="https://zenodo.org/record/7553501/files/figure.png" pageId="6" pageNumber="185">Fig. 5G</figureCitation>
|
||
) comparatively wide, with shortly attenuate apex, bearing three long setae. Right paramere (
|
||
<figureCitation id="0F95A78DC6773C58A2A4FF02945AFF45" box="[1007,1094,186,213]" captionStart="FIGURE 5" captionStartId="11.[151,250,1558,1582]" captionTargetBox="[161,1423,196,1523]" captionTargetId="figure-16@11.[150,1437,181,1535]" captionTargetPageId="11" captionText="FIGURE 5. Line drawings of male genitalia of New Zealand Nesamblyops species. N. brouni (Lewis Pass, North Canterbury, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. carltoni (Dun Mountain, Nelson, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. confusus (Mount Stokes, Marlborough Sounds, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. distinctus (Fabians Valley, Marlborough, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. lescheni (D’Urville Island, Cook Strait, Marlborough Sounds, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. oreobius (Mount Pirongia, Waikato, NO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7553501" httpUri="https://zenodo.org/record/7553501/files/figure.png" pageId="6" pageNumber="185">Fig. 5H</figureCitation>
|
||
) short and wide, bearing three long setae, which are slightly longer the length of paramere.
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6773C58A18CFEBB92BDFE8D" blockId="6.[151,1437,150,537]" box="[199,673,258,285]" pageId="6" pageNumber="185">
|
||
<emphasis id="A5DA671AC6773C58A18CFEBB91E8FE8D" bold="true" box="[199,500,258,285]" italics="true" pageId="6" pageNumber="185">Female internal genitalia.</emphasis>
|
||
Not examined.
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6773C58A18CFE9F9267FEF5" blockId="6.[151,1437,150,537]" pageId="6" pageNumber="185">
|
||
<emphasis id="A5DA671AC6773C58A18CFE9F921AFED0" bold="true" box="[199,518,294,320]" pageId="6" pageNumber="185">Geographical distribution.</emphasis>
|
||
This species is known only from Mount Stokes in the
|
||
<collectingRegion id="556A75EAC6773C58A53CFE9E9517FED1" box="[1143,1291,295,321]" country="New Zealand" name="Marlborough" pageId="6" pageNumber="185">Marlborough</collectingRegion>
|
||
Sounds area (
|
||
<bibRefCitation id="F33FC6F9C6773C58A1D4FEF2916CFEF5" author="Crosby, T. K. & Dugdale, J. S. & Watt, J. C." box="[159,368,331,357]" pageId="6" pageNumber="185" pagination="175 - 183" refId="ref15708" refString="Crosby, T. K., Dugdale, J. S. & Watt, J. C. (1998) Area codes for recording specimen localities in the New Zealand subregion. New Zealand Journal of Zoology, 25, 175 - 183. https: // doi. org / 10.1080 / 03014223.1998.9518148" type="journal article" year="1998">
|
||
Crosby
|
||
<emphasis id="A5DA671AC6773C58A1B3FEF2912DFEF5" box="[248,305,331,357]" italics="true" pageId="6" pageNumber="185">et al.</emphasis>
|
||
1998
|
||
</bibRefCitation>
|
||
) (
|
||
<figureCitation id="0F95A78DC6773C58A0C3FEF291C6FEF5" box="[392,474,331,357]" captionStart="FIGURE 10" captionStartId="18.[151,250,1698,1722]" captionTargetBox="[195,1404,151,1693]" captionTargetId="figure-120@18.[195,1404,151,1695]" captionTargetPageId="18" captionText="FIGURE 10. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Black circles—N. carltoni; yellow circles—N. parvulus; red circles—N. lescheni; green circles—N. confusus. Scale bar= 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7553515" httpUri="https://zenodo.org/record/7553515/files/figure.png" pageId="6" pageNumber="185">Fig. 10</figureCitation>
|
||
, green circle).
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6773C58A18CFED7912CFE3D" blockId="6.[151,1437,150,537]" pageId="6" pageNumber="185">
|
||
<emphasis id="A5DA671AC6773C58A18CFED79136FE18" bold="true" box="[199,298,366,392]" pageId="6" pageNumber="185">Habitat.</emphasis>
|
||
According to label data
|
||
<emphasis id="A5DA671AC6773C58A30EFED6924CFE19" bold="true" box="[581,592,367,393]" pageId="6" pageNumber="185">s</emphasis>
|
||
pecimens were collected from moss without more precise data about plant communities.
|
||
</paragraph>
|
||
<paragraph id="9711BB08C6773C58A18CFE0F920EFD89" blockId="6.[151,1437,150,537]" pageId="6" pageNumber="185">
|
||
<emphasis id="A5DA671AC6773C58A18CFE0F916CFE40" bold="true" box="[199,368,438,464]" pageId="6" pageNumber="185">Relationships.</emphasis>
|
||
The structure of the male genitalia suggests the relationship of
|
||
<taxonomicName id="50AEC08BC6773C58A513FE0E94C2FE41" authorityName="Sokolov" authorityYear="2023" box="[1112,1246,439,465]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="185" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="A5DA671AC6773C58A513FE0E94C2FE41" box="[1112,1246,439,465]" italics="true" pageId="6" pageNumber="185">N. confusus</emphasis>
|
||
</taxonomicName>
|
||
to other species having a similar V-shaped structure of dorsal copulatory sclerites in the internal sac of the median lobe, such as
|
||
<taxonomicName id="50AEC08BC6773C58A4C8FE629144FD89" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="185" phylum="Arthropoda" rank="species" species="oreobius">
|
||
<emphasis id="A5DA671AC6773C58A4C8FE6290E4FD89" italics="true" pageId="6" pageNumber="185">N. oreobius</emphasis>
|
||
(Broun)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="50AEC08BC6773C58A0C5FE469217FD89" authorityName="Sokolov" authorityYear="2023" box="[398,523,511,537]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="185" phylum="Arthropoda" rank="species" species="lescheni">
|
||
<emphasis id="A5DA671AC6773C58A0C5FE469217FD89" box="[398,523,511,537]" italics="true" pageId="6" pageNumber="185">N. lescheni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |