237 lines
29 KiB
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237 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.287.4617" ID-PMC="PMC3677354" ID-Pensoft-Pub="1313-2970-287-19" ID-Pensoft-UUID="FF94FFD2281CFF8FFF9BFFD43002FFCA" ID-PubMed="23794846" ID-Zenodo-Dep="578343" ModsDocID="1313-2970-287-19" checkinTime="1451247441750" checkinUser="pensoft" docAuthor="Kongim, Bangon, Sutcharit, Chirasak, Naggs, Fred & Panha, Somsak" docDate="2013" docId="0AA945F68CF743DE6273D5C962D2EB85" docLanguage="en" docName="ZooKeys 287: 19-40" docOrigin="ZooKeys 287" docPubDate="2013-04-11" docSource="http://dx.doi.org/10.3897/zookeys.287.4617" docTitle="Pollicaria elephas" docType="treatment" docVersion="5" id="FF94FFD2281CFF8FFF9BFFD43002FFCA" lastPageNumber="34" masterDocId="FF94FFD2281CFF8FFF9BFFD43002FFCA" masterDocTitle="Taxonomic revision of the Elephant Pupinid snail genus Pollicaria Gould, 1856 (Prosobranchia, Pupinidae)" masterLastPageNumber="40" masterPageNumber="19" pageNumber="32" updateTime="1668155620179" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Elephant Pupinid snail genus Pollicaria Gould, 1856 (Prosobranchia, Pupinidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kongim, Bangon</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Naggs, Fred</mods:namePart>
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<mods:affiliation>Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, United Kingdom</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-04-11</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>287</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>40</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.287.4617</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.287.4617</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-287-19</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FF94FFD2281CFF8FFF9BFFD43002FFCA</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578343</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152042440" LSID="urn:lsid:plazi:treatment:0AA945F68CF743DE6273D5C962D2EB85" httpUri="http://treatment.plazi.org/id/0AA945F68CF743DE6273D5C962D2EB85" lastPageId="15" lastPageNumber="34" pageId="13" pageNumber="32">
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<subSubSection pageId="13" pageNumber="32" type="nomenclature">
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<paragraph pageId="13" pageNumber="32">
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<taxonomicName LSID="0AA945F6-8CF7-43DE-6273-D5C962D2EB85" authority="(Morgan, 1885)" baseAuthorityName="Morgan" baseAuthorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Pollicaria elephas" order="Architaenioglossa" pageId="13" pageNumber="32" phylum="Mollusca" rank="species" species="elephas">Pollicaria elephas (Morgan, 1885)</taxonomicName>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Living snails. A Pollicaria myersii from Pahom, Vang Vieng, Laos (CUMZ 1572; shell height about 40 mm) B Pollicaria mouhoti mouhoti from Tam Wungdang, Phitsanulok (CUMZ 1533; shell height about 35 mm) C Pollicaria mouhoti monochroma ssp. n. from the type locality (paratype CUMZ 1548; shell height about 30 mm) D Pollicaria elephas from Ipoh, Perak, Malaysia (CUMZ 1536; shell height about 45 mm)." httpUri="https://binary.pensoft.net/fig/29853" pageId="13" pageNumber="32">Figs 2D</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Shell morphology of Pollicaria spp. A-C Pollicaria elephas A lectotype of Pollicaria elephas (MNHN 21309) B lectotype of Hybocystis jousseaumei Morgan, 1885 (MNHN 21308), and C specimen from Gunung Kenting, Ipoh, Perak, Malaysia (CUMZ 1536) D, E Pollicaria rochebruni D lectotype (MNHN 21305), and E specimen from Phuong Nga National Park, Vietnam (CUMZ 1568) F, G Pollicaria crossei, F lectotype (MNHN 21304), and G specimen from Cuc Phuong National Park, Vietnam (CUMZ 1588)." httpUri="https://binary.pensoft.net/fig/29859" pageId="13" pageNumber="32">5A-C</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Radular morphology of Pollicaria spp. A Pollicaria myersii from Pahom, Vang Vieng, Laos (CUMZ 1572) B Pollicaria mouhoti mouhoti from Tam Wungdang, Phitsanulok (CUMZ 1533) C Pollicaria mouhoti monochroma ssp. n. from the type locality (paratype CUMZ 1548) D Pollicaria elephas from Gunung Kenting, Ipoh, Perak, Malaysia (CUMZ 1536) E Pollicaria rochebruni from Phuong Nga National Park, Vietnam (CUMZ 1568) F Pollicaria crossei from Cuc Phuong National Park, Ninh Binh Province, Vietnam (CUMZ 1588). Numbers indicated order of lateral and marginal teeth. Central tooth indicated by ' C '." httpUri="https://binary.pensoft.net/fig/29862" pageId="13" pageNumber="32">6D</figureCitation>
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparative morphological characters and karyotype among Pollicaria species recognized in this study." httpUri="http://table.plazi.org/id/11DE3FA6CF5E2EAC69BF073D25EC6E3C" pageId="13" pageNumber="32" tableUuid="11DE3FA6CF5E2EAC69BF073D25EC6E3C">Tables 1</tableCitation>
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<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Shell size variation among Pollicaria species recognized in this study." httpUri="http://table.plazi.org/id/F662766526473E97F5D9D660DF7073C0" pageId="13" pageNumber="32" tableUuid="F662766526473E97F5D9D660DF7073C0">2</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="32" type="reference_group">
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<paragraph pageId="13" pageNumber="32">
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<taxonomicName authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Hybocystis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="13" pageNumber="32" phylum="Mollusca" rank="species" species="elephas">Hybocystis elephas</taxonomicName>
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Morgan, 1885b: 70. Type locality: Perak.
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<bibRefCitation author="Morgan, J de" journalOrPublisher="Bulletin de la Scoiete Zoologique de France" pageId="18" pageNumber="37" pagination="353 - 428" refId="B31" refString="Morgan, J de, 1885a. Mollusques terrestres et fluviatiles du royaume de Perak. Bulletin de la Scoiete Zoologique de France 10: 353 - 428" title="Mollusques terrestres et fluviatiles du royaume de Perak." volume="10" year="1885 a">Morgan 1885a</bibRefCitation>
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: 404, 405, pl. 7, fig. 1.
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<bibRefCitation author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="18" pageNumber="37" pagination="180 - 193" refId="B9" refString="Crosse, H, 1885. Etude monographique sur les especes du genre Hybocystis de Benson. Journal de Conchyliologie 33: 180 - 193" title="Etude monographique sur les especes du genre Hybocystis de Benson." volume="33" year="1885">Crosse 1885</bibRefCitation>
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: 183-186, pl. 11, fig. 1.
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<bibRefCitation author="Fischer, P" journalOrPublisher="Journal de Conchyliologie" pageId="18" pageNumber="37" pagination="174 - 179" refId="B11" refString="Fischer, P, 1885. Note sur ľanimal de Hybocystis elephas de Morgan. Journal de Conchyliologie 33: 174 - 179" title="Note sur ľanimal de Hybocystis elephas de Morgan." volume="33" year="1885">Fischer 1885</bibRefCitation>
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: 174.
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<bibRefCitation author="Moellendorff, OF" journalOrPublisher="Zoologischer Theil, zweiter Band. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei" pageId="18" pageNumber="37" refId="B29" refString="Moellendorff, OF, 1886. The land shells of Perak. Journal of the Asiatic Society of Bengal 55: 299-316." title="The land shells of Perak. Journal of the Asiatic Society of Bengal 55: 299 - 316." year="1886">
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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1886
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</bibRefCitation>
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: 314.
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<bibRefCitation author="Moellendorff, OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="18" pageNumber="37" pagination="330 - 348" refId="B30" refString="Moellendorff, OF, 1891. On the land and freshwater shells of Perak. Proceedings of the Zoological Society of London 1891: 330 - 348" title="On the land and freshwater shells of Perak." volume="1891" year="1891">
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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1891
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</bibRefCitation>
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: 346.
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="18" pageNumber="37" pagination="129 - 151" refId="B21" refString="Kobelt, W, Moellendorff, OF, 1899. Katalog der gegenwaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 31: 129 - 151" title="Katalog der gegenwaertig lebend bekannten Pneumonopomen." volume="31" year="1899">
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Kobelt and
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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1899
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</bibRefCitation>
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: 137.
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</paragraph>
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<paragraph pageId="13" pageNumber="32">
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<taxonomicName authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Hybocystis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="13" pageNumber="32" phylum="Mollusca" rank="species" species="jousseaumei">Hybocystis jousseaumei</taxonomicName>
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Morgan, 1885b: 70. Type locality: Kinta, Perak.
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<bibRefCitation author="Morgan, J de" journalOrPublisher="Bulletin de la Scoiete Zoologique de France" pageId="18" pageNumber="37" pagination="353 - 428" refId="B31" refString="Morgan, J de, 1885a. Mollusques terrestres et fluviatiles du royaume de Perak. Bulletin de la Scoiete Zoologique de France 10: 353 - 428" title="Mollusques terrestres et fluviatiles du royaume de Perak." volume="10" year="1885 a">Morgan 1885a</bibRefCitation>
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: 405, 406, pl. 7, fig. 2.
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<bibRefCitation author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="18" pageNumber="37" pagination="180 - 193" refId="B9" refString="Crosse, H, 1885. Etude monographique sur les especes du genre Hybocystis de Benson. Journal de Conchyliologie 33: 180 - 193" title="Etude monographique sur les especes du genre Hybocystis de Benson." volume="33" year="1885">Crosse 1885</bibRefCitation>
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: 184.
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</paragraph>
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<paragraph pageId="13" pageNumber="32">
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<taxonomicName baseAuthorityName="Morgan" baseAuthorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="13" pageNumber="32" phylum="Mollusca" rank="species" species="elephas">Pollicaria elephas</taxonomicName>
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-Kobelt, 1902: 289.
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<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal Malayan Branch Royal Asiatic Society" pageId="18" pageNumber="37" pagination="25 - 37" refId="B24" refString="Laidlaw, FF, 1928. A list of land and freshwater Mollusca of the Malay Peninsular with notes. Journal Malayan Branch Royal Asiatic Society 6: 25 - 37" title="A list of land and freshwater Mollusca of the Malay Peninsular with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
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: 33.
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<bibRefCitation author="van Benthem, Jutting WSS" journalOrPublisher="Basteria" pageId="18" pageNumber="37" pagination="10 - 20" refId="B5" refString="van Benthem, Jutting WSS, 1960. Some notes on land and freshwater Mollusca of Malaya. Basteria 24: 10 - 20" title="Some notes on land and freshwater Mollusca of Malaya." volume="24" year="1960">van Benthem Jutting 1960</bibRefCitation>
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: 12.
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<bibRefCitation author="Pain, T" journalOrPublisher="Journal of Conchology" pageId="18" pageNumber="37" pagination="173 - 178" refId="B34" refString="Pain, T, 1974. The land operculate genus Pollicaria Gould (Gastropoda), a systematic revision. Journal of Conchology 28: 173 - 178" title="The land operculate genus Pollicaria Gould (Gastropoda), a systematic revision." volume="28" year="1974">Pain 1974</bibRefCitation>
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: 176, pl. 6, fig. 1, 3.
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<bibRefCitation author="Abbott, RT" journalOrPublisher="American Malacologists, Inc., MA" pageId="18" pageNumber="37" refId="B1" refString="Abbott, RT, 1989. Compendium of Landshells. American Malacologists, Inc., MA, 240 pp." title="Compendium of Landshells." year="1989">Abbott 1989</bibRefCitation>
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: 46, 1 figure.
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<bibRefCitation author="Chan, SY" journalOrPublisher="Basteria" pageId="18" pageNumber="37" refId="B7" refString="Chan, SY, 1997. On Pollicaria elephas (de Morgan, 1885) West Malaysia. The Papustyla 11: 11-12." title="On Pollicaria elephas (de Morgan, 1885) West Malaysia. The Papustyla 11: 11 - 12." year="1997">Chan 1997</bibRefCitation>
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: 11, fig. 1-2.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="33" pageId="13" pageNumber="32" type="material examined">
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<paragraph pageId="13" pageNumber="32">Material examined.</paragraph>
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<paragraph lastPageId="14" lastPageNumber="33" pageId="13" pageNumber="32">
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Five lots with 13 specimens of syntype deposited in MNHN, the specimen figured in the original publication is designated as the lectotype of
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<taxonomicName authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Hybocystis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="13" pageNumber="32" phylum="Mollusca" rank="species" species="elephas">
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<emphasis italics="true" pageId="13" pageNumber="32">Hybocystis elephas</emphasis>
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</taxonomicName>
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Morgan, 1885 MNHN 21309 (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Shell morphology of Pollicaria spp. A-C Pollicaria elephas A lectotype of Pollicaria elephas (MNHN 21309) B lectotype of Hybocystis jousseaumei Morgan, 1885 (MNHN 21308), and C specimen from Gunung Kenting, Ipoh, Perak, Malaysia (CUMZ 1536) D, E Pollicaria rochebruni D lectotype (MNHN 21305), and E specimen from Phuong Nga National Park, Vietnam (CUMZ 1568) F, G Pollicaria crossei, F lectotype (MNHN 21304), and G specimen from Cuc Phuong National Park, Vietnam (CUMZ 1588)." httpUri="https://binary.pensoft.net/fig/29859" pageId="13" pageNumber="32">Fig. 5A</figureCitation>
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), paralectotype MNHN 21310 (5 shells), 21311 (2 shells), 21312 (3 shells), 21313 (2 shells), RBINS 525391 (1 shell).
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<pageBreakToken pageId="14" pageNumber="33" start="start">Single</pageBreakToken>
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syntype specimen is designated as the lectotype of
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<taxonomicName authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Hybocystis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="14" pageNumber="33" phylum="Mollusca" rank="species" species="jousseaumei">
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<emphasis italics="true" pageId="14" pageNumber="33">Hybocystis jousseaumei</emphasis>
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</taxonomicName>
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Morgan, 1885 MNHN 21308 (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Shell morphology of Pollicaria spp. A-C Pollicaria elephas A lectotype of Pollicaria elephas (MNHN 21309) B lectotype of Hybocystis jousseaumei Morgan, 1885 (MNHN 21308), and C specimen from Gunung Kenting, Ipoh, Perak, Malaysia (CUMZ 1536) D, E Pollicaria rochebruni D lectotype (MNHN 21305), and E specimen from Phuong Nga National Park, Vietnam (CUMZ 1568) F, G Pollicaria crossei, F lectotype (MNHN 21304), and G specimen from Cuc Phuong National Park, Vietnam (CUMZ 1588)." httpUri="https://binary.pensoft.net/fig/29859" pageId="14" pageNumber="33">Fig. 5B</figureCitation>
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). Ipoh, Perak, Malaysia: ZMA E.A. Meene colln. Acc. no. 1982 (1 shell). Near bridge over river, road Ipoh to Tanjong Rambutan, Perak, Malaysia: ZMA J. Drijver colln. (5 shells). Perak, Malaysia: ZMB 75821 (2 shells), 38044 (1 shell), M. Schulz colln. 1216 (3 shells, smallest shell excluded). Bukit Chintamani, Selangor, Malaysia: CUMZ 1534. Gunung Kenting, Ampang Baru, Ipoh, Perak, Malaysia: CUMZ 1535, 1536 (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Shell morphology of Pollicaria spp. A-C Pollicaria elephas A lectotype of Pollicaria elephas (MNHN 21309) B lectotype of Hybocystis jousseaumei Morgan, 1885 (MNHN 21308), and C specimen from Gunung Kenting, Ipoh, Perak, Malaysia (CUMZ 1536) D, E Pollicaria rochebruni D lectotype (MNHN 21305), and E specimen from Phuong Nga National Park, Vietnam (CUMZ 1568) F, G Pollicaria crossei, F lectotype (MNHN 21304), and G specimen from Cuc Phuong National Park, Vietnam (CUMZ 1588)." httpUri="https://binary.pensoft.net/fig/29859" pageId="14" pageNumber="33">Fig. 5C</figureCitation>
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), 1566, 1570.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="34" type="description">
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<paragraph pageId="15" pageNumber="34">
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<pageBreakToken pageId="15" pageNumber="34" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="15" pageNumber="34">
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<emphasis bold="true" pageId="15" pageNumber="34">Shell:</emphasis>
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Shell large, elongate pupoid uniform yellow to orange. Periostracum thin, corneous; shell surface with fine growth lines and last whorl with distinctly strong pitting dorsally. Whorls 6-7 whorls; sutures impressed; apex obtuse. Last whorl large about two-third of shell height, flattened in front. Aperture rounded, with shallow to deep posterior angle groove. Peristome continuous, little elevated, yellow to orange, internal parietal declining shoulder absent. Lip thickened, duplicated, and with distinct growth ridges; umbilicus narrow. Operculum thick, calcareous, concentric.
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</paragraph>
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<paragraph pageId="15" pageNumber="34">
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<emphasis bold="true" pageId="15" pageNumber="34">Radula:</emphasis>
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Taenioglossate radula, teeth arrangement with central, lateral and marginal teeth shape similar to
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<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="myersii">
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<emphasis italics="true" pageId="15" pageNumber="34">Pollicaria myersii</emphasis>
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</taxonomicName>
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. Minor differences are the well-developed central cusp with one to three small lateral cusps of the central tooth, and the slightly elongate and slender central cusp of the inner marginal teeth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="34" type="distribution">
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<paragraph pageId="15" pageNumber="34">Distribution.</paragraph>
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<paragraph pageId="15" pageNumber="34">
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This species has a restricted distribution and is known only from limestone outcrops in Perak, Peninsular Malaysia (
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<bibRefCitation author="Morgan, J de" journalOrPublisher="Bulletin de la Scoiete Zoologique de France" pageId="18" pageNumber="37" pagination="353 - 428" refId="B31" refString="Morgan, J de, 1885a. Mollusques terrestres et fluviatiles du royaume de Perak. Bulletin de la Scoiete Zoologique de France 10: 353 - 428" title="Mollusques terrestres et fluviatiles du royaume de Perak." volume="10" year="1885 a">Morgan 1885a</bibRefCitation>
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,
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<bibRefCitation author="Morgan, J de" journalOrPublisher="Le Naturaliste" pageId="18" pageNumber="37" pagination="68 - 70" refId="B32" refString="Morgan, J de, 1885b. Note sur quelques especes nouvelles de mollusques terrestres recruillis dans la Peninsula Malaise. Le Naturaliste 7: 68 - 70" title="Note sur quelques especes nouvelles de mollusques terrestres recruillis dans la Peninsula Malaise." volume="7" year="1885 b">b</bibRefCitation>
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||
). Material collected for this study was from Kinta valley, Perak, and the southern part of the
|
||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||
historical range in Bukit Chintamani, Selangor, Peninsular Malaysia is considered to be this locally endemic species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="15" pageNumber="34" type="remarks">
|
||
<paragraph pageId="15" pageNumber="34">Remarks.</paragraph>
|
||
<paragraph pageId="15" pageNumber="34">
|
||
The locally endemic
|
||
<taxonomicName baseAuthorityName="Morgan" baseAuthorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="elephas">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Pollicaria elephas</emphasis>
|
||
</taxonomicName>
|
||
is confined to a few limestone outcrops in Peninsular Malaysia and shows several unique shell characters that separate it from its congeners. The major distinguishing characters of this species are the very large, monochrome yellowish to pale orange shell with the last whorl distorted ventrally and sculptured with scattered, deep pits dorsally; and rounded and thickened aperture. (
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Shell size variation among Pollicaria species recognized in this study." httpUri="http://table.plazi.org/id/F662766526473E97F5D9D660DF7073C0" pageId="15" pageNumber="34" tableUuid="F662766526473E97F5D9D660DF7073C0">Table 2</tableCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Shell morphology of Pollicaria spp. A-C Pollicaria elephas A lectotype of Pollicaria elephas (MNHN 21309) B lectotype of Hybocystis jousseaumei Morgan, 1885 (MNHN 21308), and C specimen from Gunung Kenting, Ipoh, Perak, Malaysia (CUMZ 1536) D, E Pollicaria rochebruni D lectotype (MNHN 21305), and E specimen from Phuong Nga National Park, Vietnam (CUMZ 1568) F, G Pollicaria crossei, F lectotype (MNHN 21304), and G specimen from Cuc Phuong National Park, Vietnam (CUMZ 1588)." httpUri="https://binary.pensoft.net/fig/29859" pageId="15" pageNumber="34">Fig. 5A-C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="15" pageNumber="34">
|
||
<bibRefCitation author="Morgan, J de" journalOrPublisher="Bulletin de la Scoiete Zoologique de France" pageId="18" pageNumber="37" pagination="353 - 428" refId="B31" refString="Morgan, J de, 1885a. Mollusques terrestres et fluviatiles du royaume de Perak. Bulletin de la Scoiete Zoologique de France 10: 353 - 428" title="Mollusques terrestres et fluviatiles du royaume de Perak." volume="10" year="1885 a">Morgan (1885a</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Morgan, J de" journalOrPublisher="Le Naturaliste" pageId="18" pageNumber="37" pagination="68 - 70" refId="B32" refString="Morgan, J de, 1885b. Note sur quelques especes nouvelles de mollusques terrestres recruillis dans la Peninsula Malaise. Le Naturaliste 7: 68 - 70" title="Note sur quelques especes nouvelles de mollusques terrestres recruillis dans la Peninsula Malaise." volume="7" year="1885 b">b</bibRefCitation>
|
||
) proposed two nominal species of
|
||
<taxonomicName authorityName="Gould" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Pollicaria</emphasis>
|
||
</taxonomicName>
|
||
from Perak, which differed mainly by the shell size (larger shell
|
||
<taxonomicName authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Hybocystis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="elephas">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Hybocystis elephas</emphasis>
|
||
</taxonomicName>
|
||
and smaller shell
|
||
<taxonomicName authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Hybocystis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="jousseaumei">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Hybocystis jousseaumei</emphasis>
|
||
</taxonomicName>
|
||
). In the first revision of this genus,
|
||
<bibRefCitation author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="18" pageNumber="37" pagination="180 - 193" refId="B9" refString="Crosse, H, 1885. Etude monographique sur les especes du genre Hybocystis de Benson. Journal de Conchyliologie 33: 180 - 193" title="Etude monographique sur les especes du genre Hybocystis de Benson." volume="33" year="1885">Crosse (1885)</bibRefCitation>
|
||
assumed that they were the same species and recognized only
|
||
<taxonomicName baseAuthorityName="Morgan" baseAuthorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="elephas">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Pollicaria elephas</emphasis>
|
||
</taxonomicName>
|
||
. Thereafter
|
||
<taxonomicName class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="jousseaumei">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Pollicaria jousseaumei</emphasis>
|
||
</taxonomicName>
|
||
was recognized as a synonym of
|
||
<taxonomicName baseAuthorityName="Morgan" baseAuthorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="elephas">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Pollicaria elephas</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Kobelt, W" journalOrPublisher="Verlag von R. Friedlaender und Sohn, Berlin" pageId="18" pageNumber="37" refId="B20" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich, 16 Lieferung. Verlag von R. Friedlaender und Sohn, Berlin, 663 pp." title="Cyclophoridae. Das Tierreich, 16 Lieferung." year="1902">Kobelt 1902</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Pain, T" journalOrPublisher="Journal of Conchology" pageId="18" pageNumber="37" pagination="173 - 178" refId="B34" refString="Pain, T, 1974. The land operculate genus Pollicaria Gould (Gastropoda), a systematic revision. Journal of Conchology 28: 173 - 178" title="The land operculate genus Pollicaria Gould (Gastropoda), a systematic revision." volume="28" year="1974">Pain 1974</bibRefCitation>
|
||
). Examination of the type specimens (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Shell morphology of Pollicaria spp. A-C Pollicaria elephas A lectotype of Pollicaria elephas (MNHN 21309) B lectotype of Hybocystis jousseaumei Morgan, 1885 (MNHN 21308), and C specimen from Gunung Kenting, Ipoh, Perak, Malaysia (CUMZ 1536) D, E Pollicaria rochebruni D lectotype (MNHN 21305), and E specimen from Phuong Nga National Park, Vietnam (CUMZ 1568) F, G Pollicaria crossei, F lectotype (MNHN 21304), and G specimen from Cuc Phuong National Park, Vietnam (CUMZ 1588)." httpUri="https://binary.pensoft.net/fig/29859" pageId="15" pageNumber="34">Fig. 5A, B</figureCitation>
|
||
) confirmed
|
||
<taxonomicName class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="jousseaumei">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Pollicaria jousseaumei</emphasis>
|
||
</taxonomicName>
|
||
as junior synonym of
|
||
<taxonomicName baseAuthorityName="Morgan" baseAuthorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Pollicaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="34" phylum="Mollusca" rank="species" species="elephas">
|
||
<emphasis italics="true" pageId="15" pageNumber="34">Pollicaria elephas</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the recent land snail survey in Perak, Peninsular Malaysia recorded both large and small shell forms of the species from the same localities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |