treatments-xml/data/05/4F/DB/054FDB70FF2CFF35B49C1FCE7CCB25B4.xml
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<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FF2CFF35B49C1FCE7CCB25B4" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax bison Prebus 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="240" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="236" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PeerJ</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="series">
<mods:title>e 11514</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2021-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>462</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
<mods:identifier type="ISSN">2167-8359</mods:identifier>
<mods:identifier type="PMC">PMC8254503</mods:identifier>
<mods:identifier type="PubMed">34249486</mods:identifier>
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102555" ID-GBIF-Taxon="183542626" ID-Zenodo-Dep="5102555" LSID="urn:lsid:zoobank.org:act:F59A4882-C8C7-4C0C-A340-F2DC34D2EB0A" httpUri="http://treatment.plazi.org/id/054FDB70FF2CFF35B49C1FCE7CCB25B4" lastPageId="239" lastPageNumber="240" pageId="235" pageNumber="236">
<subSubSection box="[498,836,1491,1518]" pageId="235" pageNumber="236" type="nomenclature">
<paragraph blockId="235.[498,982,1491,1558]" box="[498,836,1491,1518]" pageId="235" pageNumber="236">
<heading box="[498,836,1491,1518]" level="2" pageId="235" pageNumber="236">
<emphasis bold="true" box="[498,836,1491,1518]" pageId="235" pageNumber="236">
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,732,1491,1518]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="235" pageNumber="236" phylum="Arthropoda" rank="species" species="bison" status="sp. nov.">
<emphasis bold="true" box="[498,732,1491,1518]" italics="true" pageId="235" pageNumber="236">Temnothorax bison</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[742,836,1492,1518]" pageId="235" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,982,1532,1558]" pageId="235" pageNumber="236" type="distribution">
<paragraph blockId="235.[498,982,1491,1558]" box="[498,982,1532,1558]" pageId="235" pageNumber="236">
Distribution:
<figureCitation box="[657,770,1532,1558]" captionStart="Figure 127" captionStartId="235.[524,589,731,753]" captionTargetBox="[498,1541,234,717]" captionTargetId="figure-274@235.[498,1534,234,702]" captionTargetPageId="235" captionText="Figure 127 Geographical and elevational distribution of the pergandei group. (A) Temnothorax bison sp. nov. (B) T. pergandei. The colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles; if present, types of synonyms are named; non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-127" figureDoi="http://doi.org/10.5281/zenodo.5102369" httpUri="https://zenodo.org/record/5102369/files/figure.png" pageId="235" pageNumber="236">Fig. 127A</figureCitation>
; worker:
<figureCitation box="[882,976,1532,1558]" captionStart="Figure 128" captionStartId="236.[524,589,1078,1100]" captionTargetBox="[498,1541,234,1065]" captionTargetId="figure-240@236.[498,1295,234,1050]" captionTargetPageId="236" captionText="Figure 128 Temnothorax bison sp. nov. holotype worker (CASENT0636947). (A) Profile view. (B) Full-face view. (C) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-128" figureDoi="http://doi.org/10.5281/zenodo.5102373" httpUri="https://zenodo.org/record/5102373/files/figure.png" pageId="235" pageNumber="236">Fig. 128</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[498,1084,1579,1606]" pageId="235" pageNumber="236" type="reference_group">
<paragraph blockId="235.[498,1084,1579,1606]" box="[498,1084,1579,1606]" pageId="235" pageNumber="236">
<treatmentCitationGroup box="[498,1084,1579,1606]" pageId="235" pageNumber="236">
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1580,1606]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="235" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis box="[498,655,1580,1606]" italics="true" pageId="235" pageNumber="236">Temnothorax</emphasis>
</taxonomicName>
mmp05
<treatmentCitation author="Prebus M." box="[763,1080,1580,1606]" page="8" pageId="235" pageNumber="236" year="2017">
<bibRefCitation author="Prebus M." box="[763,942,1580,1606]" journalOrPublisher="BMC Evolutionary Biology" pageId="235" pageNumber="236" pagination="250" part="17" refId="ref272195" refString="Prebus M. 2017. Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae). BMC Evolutionary Biology 17 (1): 250 DOI 10.1186 / s 12862 - 017 - 1095 - 8." title="Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae)" type="journal article" year="2017">
<emphasis box="[763,911,1580,1606]" italics="true" pageId="235" pageNumber="236">
<author box="[763,843,1580,1606]" fns="*" inm="true" npo="*" pageId="235" pageNumber="236">Prebus</author>
,
<year box="[856,911,1580,1606]" pageId="235" pageNumber="236">2017</year>
</emphasis>
: 8
</bibRefCitation>
. Phylogeny
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="236" lastPageNumber="237" pageId="235" pageNumber="236" type="materials_examined">
<paragraph blockId="235.[498,1542,1642,1908]" pageId="235" pageNumber="236">
<emphasis bold="true" box="[530,831,1642,1668]" pageId="235" pageNumber="236">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861379" collectingDate="2015-06-18" collectingDateMax="2015-06-21" collectingDateMin="2015-06-18" collectionCode="CASC" collectorName="Z. Falin" country="Guatemala" elevation="200" latitude="14.99336" location="finca near University Landivar campus" longLatPrecision="1" longitude="-89.54607" pageId="235" pageNumber="236" specimenCode="GUATIF15-188, CASENT0636947" specimenCount="1" stateProvince="Zacapa" typeStatus="holotype">
<emphasis box="[838,1035,1642,1668]" italics="true" pageId="235" pageNumber="236">
<typeStatus box="[838,941,1642,1668]" pageId="235" pageNumber="236">Holotype</typeStatus>
<specimenCount box="[948,1028,1642,1668]" pageId="235" pageNumber="236" type="worker">worker</specimenCount>
:
</emphasis>
<collectingCountry box="[1042,1223,1642,1668]" name="Guatemala" pageId="235" pageNumber="236">GUATEMALA</collectingCountry>
:
<collectingRegion box="[1233,1319,1643,1669]" country="Guatemala" name="Zacapa" pageId="235" pageNumber="236">Zacapa</collectingRegion>
: Zacapa,
<location LSID="urn:lsid:plazi:treatment:054FDB70FF2CFF35B49C1FCE7CCB25B4:88393CBDFF2CFF31B0FD1C777A0321FA" country="Guatemala" latitude="14.99336" longLatPrecision="1" longitude="-89.54607" name="finca near University Landivar campus" pageId="235" pageNumber="236" stateProvince="Zacapa">finca near University Landivar campus</location>
,
<geoCoordinate box="[829,961,1677,1709]" degrees="14.99336" direction="north" orientation="latitude" pageId="235" pageNumber="236" precision="1" value="14.99336">
14.99336
<emphasis box="[928,938,1677,1695]" italics="true" pageId="235" pageNumber="236">
<superScript attach="right" box="[928,938,1677,1695]" fontSize="8" pageId="235" pageNumber="236">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[969,1109,1677,1709]" degrees="89.54607" direction="west" orientation="longitude" pageId="235" pageNumber="236" precision="1" value="-89.54607">
89.54607
<emphasis box="[1068,1078,1677,1695]" italics="true" pageId="235" pageNumber="236">
<superScript attach="right" box="[1068,1078,1677,1695]" fontSize="8" pageId="235" pageNumber="236">°</superScript>
</emphasis>
W
</geoCoordinate>
,
<quantity box="[1122,1195,1682,1709]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="235" pageNumber="236" unit="m" value="200.0">
<elevation box="[1122,1195,1682,1709]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="235" pageNumber="236" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date box="[1208,1399,1682,1709]" pageId="235" pageNumber="236" value="2015-06-18" valueMax="2015-06-21" valueMin="2015-06-18">
<collectingDate box="[1208,1399,1682,1709]" pageId="235" pageNumber="236" value="2015-06-18" valueMax="2015-06-21" valueMin="2015-06-18">1821 June 2015</collectingDate>
</date>
,
<collectorName pageId="235" pageNumber="236">Z. Falin</collectorName>
#
<specimenCode box="[571,747,1722,1749]" pageId="235" pageNumber="236">GUATIF15-188</specimenCode>
, dry thorn scrub forest, flight intercept trap (
<specimenCode box="[1231,1434,1722,1749]" pageId="235" pageNumber="236">CASENT0636947</specimenCode>
) [
<collectionCode box="[1454,1532,1721,1748]" pageId="235" pageNumber="236">CASC</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="235.[498,1542,1642,1908]" box="[530,1469,1761,1789]" pageId="235" pageNumber="236">
<materialsCitation ID-GBIF-Occurrence="3321861652" box="[530,1465,1761,1789]" collectingDate="2015-06-18" collectingDateMax="2015-06-21" collectingDateMin="2015-06-18" collectionCode="UCDC" collectorName="Z. Falin" country="Guatemala" elevation="200" latitude="14.99336" location="University Landivar" longLatPrecision="1" longitude="-89.54607" pageId="235" pageNumber="236" specimenCount="1" specimenCount-worker="1" stateProvince="Zacapa" typeStatus="paratype">
<emphasis box="[530,728,1762,1788]" italics="true" pageId="235" pageNumber="236">
<typeStatus box="[530,632,1763,1788]" pageId="235" pageNumber="236">Paratype</typeStatus>
worker:
</emphasis>
same data as holotype,
<specimenCount box="[1012,1117,1762,1788]" pageId="235" pageNumber="236" type="worker">1 worker</specimenCount>
(CASENT0636946) [
<collectionCode box="[1368,1458,1761,1788]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="235" pageNumber="236" type="Museum">UCDC</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="235.[498,1542,1642,1908]" lastBlockId="236.[498,1542,1180,1928]" lastPageId="236" lastPageNumber="237" pageId="235" pageNumber="236">
<emphasis bold="true" box="[530,888,1801,1828]" pageId="235" pageNumber="236">Non-type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861384" collectingDate="2007-07-07" collectionCode="JTLC, UVGC" collectorName="J. Longino" country="Guatemala" elevation="530" latitude="14.907267" location="Morazan" longLatPrecision="5" longitude="-90.1548" pageId="235" pageNumber="236" specimenCode="JTLC000010341, JTLC000010340" specimenCount="2" specimenCount-worker="2" stateProvince="El Progreso">
<collectingCountry box="[897,1078,1802,1828]" name="Guatemala" pageId="235" pageNumber="236">GUATEMALA</collectingCountry>
:
<collectingRegion box="[1090,1228,1802,1828]" country="Guatemala" name="El Progreso" pageId="235" pageNumber="236">El Progreso</collectingRegion>
:
<quantity box="[1240,1322,1802,1828]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="235" pageNumber="236" unit="km" value="3.7">3.7 km</quantity>
SW
<location LSID="urn:lsid:plazi:treatment:054FDB70FF2CFF35B49C1FCE7CCB25B4:88393CBDFF2CFF31B00B1D177CE22073" box="[1381,1488,1802,1828]" country="Guatemala" latitude="14.907267" longLatPrecision="5" longitude="-90.1548" name="Morazan" pageId="235" pageNumber="236" stateProvince="El Progreso">Morazán</location>
,
<geoCoordinate box="[498,650,1837,1868]" degrees="14.907267" direction="north" orientation="latitude" pageId="235" pageNumber="236" precision="1" value="14.907267">
14.907267
<emphasis box="[617,627,1837,1855]" italics="true" pageId="235" pageNumber="236">
<superScript attach="right" box="[617,627,1837,1855]" fontSize="8" pageId="235" pageNumber="236">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[657,789,1837,1868]" degrees="90.1548" direction="west" orientation="longitude" pageId="235" pageNumber="236" precision="5" value="-90.1548">
90.1548
<emphasis box="[747,757,1837,1855]" italics="true" pageId="235" pageNumber="236">
<superScript attach="right" box="[747,757,1837,1855]" fontSize="8" pageId="235" pageNumber="236">°</superScript>
</emphasis>
W
</geoCoordinate>
,
<quantity box="[801,874,1841,1868]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="235" pageNumber="236" unit="m" value="530.0">
<elevation box="[801,874,1841,1868]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="235" pageNumber="236" unit="m" value="530.0">530 m</elevation>
</quantity>
,
<date box="[887,1018,1841,1868]" pageId="235" pageNumber="236" value="2007-07-07">
<collectingDate box="[887,1018,1841,1868]" pageId="235" pageNumber="236" value="2007-07-07">7 July 2007</collectingDate>
</date>
,
<collectorName box="[1030,1154,1841,1868]" pageId="235" pageNumber="236">J. Longino</collectorName>
#6021-s, scrubby xeric vegetation, on ground,
<specimenCount box="[633,736,1881,1907]" pageId="235" pageNumber="236" type="worker">1 worker</specimenCount>
(
<specimenCode box="[750,943,1881,1907]" collectionCode="JTLC" pageId="235" pageNumber="236">JTLC000010341</specimenCode>
) [
<collectionCode box="[963,1032,1881,1908]" pageId="235" pageNumber="236">JTLC</collectionCode>
]
<specimenCount box="[1045,1148,1881,1907]" pageId="235" pageNumber="236" type="worker">1 worker</specimenCount>
(
<specimenCode box="[1162,1355,1881,1907]" collectionCode="JTLC" pageId="235" pageNumber="236">JTLC000010340</specimenCode>
) [
<collectionCode box="[1376,1466,1881,1908]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="235" pageNumber="236">UVGC</collectionCode>
]
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3321861348" collectingDate="2003-11-17" collectionCode="USNM" collectorName="A. L. Wild" country="Guatemala" elevation="400" lastPageId="236" lastPageNumber="237" latitude="14.916667" location="El Rancho" longLatPrecision="1" longitude="-90.066666" pageId="235" pageNumber="236" specimenCount="1" specimenCount-worker="1" stateProvince="El Progreso">
<quantity box="[1483,1542,1881,1908]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="235" pageNumber="236" unit="km" value="5.0">5 km</quantity>
W
<location LSID="urn:lsid:plazi:treatment:054FDB70FF2CFF35B49C1FCE7CCB25B4:88393CBDFF2BFF36B7791EBD7BA123ED" box="[535,659,1184,1210]" country="Guatemala" latitude="14.916667" longLatPrecision="1" longitude="-90.066666" name="El Rancho" pageId="236" pageNumber="237" stateProvince="El Progreso">El Rancho</location>
,
<geoCoordinate box="[672,824,1180,1211]" degrees="14.916667" direction="north" orientation="latitude" pageId="236" pageNumber="237" precision="1" value="14.916667">
14.916667
<emphasis box="[790,800,1180,1198]" italics="true" pageId="236" pageNumber="237">
<superScript attach="right" box="[790,800,1180,1198]" fontSize="8" pageId="236" pageNumber="237">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[832,992,1180,1211]" degrees="90.066667" direction="west" orientation="longitude" pageId="236" pageNumber="237" precision="1" value="-90.066666">
90.066667
<emphasis box="[950,960,1180,1198]" italics="true" pageId="236" pageNumber="237">
<superScript attach="right" box="[950,960,1180,1198]" fontSize="8" pageId="236" pageNumber="237">°</superScript>
</emphasis>
W
</geoCoordinate>
,
<quantity box="[1005,1081,1185,1211]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="236" pageNumber="237" unit="m" value="400.0">
<elevation box="[1005,1081,1185,1211]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="236" pageNumber="237" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date box="[1095,1321,1184,1211]" pageId="236" pageNumber="237" value="2003-11-17">
<collectingDate box="[1095,1321,1184,1211]" pageId="236" pageNumber="237" value="2003-11-17">17 November 2003</collectingDate>
</date>
,
<collectorName box="[1334,1449,1184,1211]" pageId="236" pageNumber="237">A.L. Wild</collectorName>
#AW2082, tropical dry forest,
<specimenCount box="[858,963,1224,1251]" pageId="236" pageNumber="237" type="worker">1 worker</specimenCount>
(CASENT0915984) [
<collectionCode box="[1214,1306,1224,1251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="236" pageNumber="237" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102373" ID-Zenodo-Dep="5102373" httpUri="https://zenodo.org/record/5102373/files/figure.png" pageId="236" pageNumber="237" startId="236.[524,589,1078,1100]" targetBox="[498,1541,234,1065]" targetPageId="236">
<paragraph blockId="236.[524,1515,1078,1130]" pageId="236" pageNumber="237">
<emphasis bold="true" box="[524,1306,1078,1100]" pageId="236" pageNumber="237">
Figure 128
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[645,838,1078,1100]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="236" pageNumber="237" phylum="Arthropoda" rank="species" species="bison" status="sp. nov.">
<emphasis bold="true" box="[645,838,1078,1100]" italics="true" pageId="236" pageNumber="237">Temnothorax bison</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[847,926,1078,1099]" pageId="236" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
holotype worker (CASENT0636947).
</emphasis>
(A) Profile view. (B) Full-face view. (C) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-128
</paragraph>
</caption>
<paragraph blockId="236.[498,1542,1180,1928]" pageId="236" pageNumber="237">
<materialsCitation ID-GBIF-Occurrence="3321861688" collectingDate="1978-08-28" collectionCode="MCZC" collectorName="John E. Rawlins" country="Mexico" elevation="560" latitude="16.966667" location="Yaxoquintela" longLatPrecision="1" longitude="-91.78333" pageId="236" pageNumber="237" specimenCount="1" specimenCount-worker="1" stateProvince="Chiapas">
<collectingCountry box="[530,645,1264,1290]" name="Mexico" pageId="236" pageNumber="237">MEXICO</collectingCountry>
:
<collectingRegion box="[656,753,1264,1290]" country="Mexico" name="Chiapas" pageId="236" pageNumber="237">Chiapas</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:054FDB70FF2CFF35B49C1FCE7CCB25B4:88393CBDFF2BFF36B7921EED7AAA225D" box="[764,920,1264,1290]" country="Mexico" latitude="16.966667" longLatPrecision="1" longitude="-91.78333" name="Yaxoquintela" pageId="236" pageNumber="237" stateProvince="Chiapas">Yaxoquintela</location>
,
<geoCoordinate box="[932,1084,1260,1291]" degrees="16.966667" direction="north" orientation="latitude" pageId="236" pageNumber="237" precision="1" value="16.966667">
16.966667
<emphasis box="[1051,1061,1260,1278]" italics="true" pageId="236" pageNumber="237">
<superScript attach="right" box="[1051,1061,1260,1278]" fontSize="8" pageId="236" pageNumber="237">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[1092,1252,1260,1291]" degrees="91.783333" direction="west" orientation="longitude" pageId="236" pageNumber="237" precision="1" value="-91.78333">
91.783333
<emphasis box="[1210,1220,1260,1278]" italics="true" pageId="236" pageNumber="237">
<superScript attach="right" box="[1210,1220,1260,1278]" fontSize="8" pageId="236" pageNumber="237">°</superScript>
</emphasis>
W
</geoCoordinate>
,
<quantity box="[1264,1339,1264,1291]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="236" pageNumber="237" unit="m" value="560.0">
<elevation box="[1264,1339,1264,1291]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="236" pageNumber="237" unit="m" value="560.0">560 m</elevation>
</quantity>
,
<date box="[1352,1537,1264,1291]" pageId="236" pageNumber="237" value="1978-08-28">
<collectingDate box="[1352,1537,1264,1291]" pageId="236" pageNumber="237" value="1978-08-28">28 August 1978</collectingDate>
</date>
,
<collectorName box="[498,688,1304,1331]" pageId="236" pageNumber="237">John E. Rawlins</collectorName>
,
<specimenCount box="[701,806,1304,1330]" pageId="236" pageNumber="237" type="worker">1 worker</specimenCount>
(MCZ-ENT00541468) [
<collectionCode box="[1091,1182,1303,1330]" pageId="236" pageNumber="237">MCZC</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="236.[498,1542,1180,1928]" pageId="236" pageNumber="237">
<materialsCitation ID-GBIF-Occurrence="3321861407" collectingDate="2011-04-23" collectionCode="UCDC, LACM" collectorName="J. Longino" country="Nicaragua" elevation="0" latitude="13.40699" location="Somoto" longLatPrecision="1" longitude="-86.59106" pageId="236" pageNumber="237" specimenCount="2" specimenCount-worker="2" stateProvince="Madriz">
<collectingCountry box="[530,704,1344,1370]" name="Nicaragua" pageId="236" pageNumber="237">NICARAGUA</collectingCountry>
:
<collectingRegion box="[716,803,1344,1370]" country="Nicaragua" name="Madriz" pageId="236" pageNumber="237">Madriz</collectingRegion>
:
<quantity box="[815,876,1344,1370]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="236" pageNumber="237" unit="km" value="8.0">8 km</quantity>
S
<location LSID="urn:lsid:plazi:treatment:054FDB70FF2CFF35B49C1FCE7CCB25B4:88393CBDFF2BFF36B6E51F5D7AD5220D" box="[907,999,1344,1370]" country="Nicaragua" latitude="13.40699" longLatPrecision="1" longitude="-86.59106" name="Somoto" pageId="236" pageNumber="237" stateProvince="Madriz">Somoto</location>
,
<geoCoordinate box="[1012,1150,1339,1371]" degrees="13.40699" direction="north" orientation="latitude" pageId="236" pageNumber="237" precision="1" value="13.40699">
13.40699
<emphasis box="[1117,1127,1339,1357]" italics="true" pageId="236" pageNumber="237">
<superScript attach="right" box="[1117,1127,1339,1357]" fontSize="8" pageId="236" pageNumber="237">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[1159,1302,1339,1371]" degrees="86.59106" direction="west" orientation="longitude" pageId="236" pageNumber="237" precision="1" value="-86.59106">
86.59106
<emphasis box="[1263,1273,1339,1357]" italics="true" pageId="236" pageNumber="237">
<superScript attach="right" box="[1263,1273,1339,1357]" fontSize="8" pageId="236" pageNumber="237">°</superScript>
</emphasis>
W
</geoCoordinate>
±
<elevation box="[1327,1500,1344,1371]" isEnumeration="true" pageId="236" pageNumber="237">
<quantity box="[1327,1390,1344,1371]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="236" pageNumber="237" unit="m" value="50.0">50 m</quantity>
,
<quantity box="[1404,1500,1344,1371]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="236" pageNumber="237" unit="m" value="1260.0">1,260 m</quantity>
</elevation>
,
<date box="[498,657,1384,1410]" pageId="236" pageNumber="237" value="2011-04-23">
<collectingDate box="[498,657,1384,1410]" pageId="236" pageNumber="237" value="2011-04-23">23 April 2011</collectingDate>
</date>
,
<collectorName box="[667,783,1384,1410]" pageId="236" pageNumber="237">J. Longino</collectorName>
#JTL7396, dry oak woodland, at bait,
<specimenCount box="[1211,1313,1384,1410]" pageId="236" pageNumber="237" type="worker">1 worker</specimenCount>
(CASENT0619615) [
<collectionCode box="[505,591,1423,1450]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="236" pageNumber="237" type="Museum">UCDC</collectionCode>
]
<specimenCount box="[606,710,1424,1450]" pageId="236" pageNumber="237" type="worker">1 worker</specimenCount>
(CASENT0619345) [
<collectionCode box="[956,1045,1423,1450]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="236" pageNumber="237" type="Museum">LACM</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[530,1542,1463,1490]" pageId="236" pageNumber="237" type="distribution">
<paragraph blockId="236.[498,1542,1180,1928]" box="[530,1542,1463,1490]" pageId="236" pageNumber="237">
<emphasis bold="true" box="[530,754,1463,1490]" pageId="236" pageNumber="237">Geographic range:</emphasis>
Low-to-mid elevations, southern
<collectingCountry box="[1151,1237,1463,1489]" name="Mexico" pageId="236" pageNumber="237">Mexico</collectingCountry>
to
<collectingCountry box="[1276,1395,1463,1489]" name="Nicaragua" pageId="236" pageNumber="237">Nicaragua</collectingCountry>
(
<figureCitation box="[1413,1525,1463,1490]" captionStart="Figure 127" captionStartId="235.[524,589,731,753]" captionTargetBox="[498,1541,234,717]" captionTargetId="figure-274@235.[498,1534,234,702]" captionTargetPageId="235" captionText="Figure 127 Geographical and elevational distribution of the pergandei group. (A) Temnothorax bison sp. nov. (B) T. pergandei. The colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles; if present, types of synonyms are named; non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-127" figureDoi="http://doi.org/10.5281/zenodo.5102369" httpUri="https://zenodo.org/record/5102369/files/figure.png" pageId="236" pageNumber="237">Fig. 127A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="237" lastPageNumber="238" pageId="236" pageNumber="237" type="diagnosis">
<paragraph blockId="236.[498,1542,1180,1928]" pageId="236" pageNumber="237">
<emphasis bold="true" box="[530,754,1503,1529]" pageId="236" pageNumber="237">Worker diagnosis:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[763,990,1503,1529]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="236" pageNumber="237" phylum="Arthropoda" rank="species" species="bison" status="sp. nov.">
<emphasis box="[763,990,1503,1529]" italics="true" pageId="236" pageNumber="237">Temnothorax bison</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[998,1090,1504,1530]" pageId="236" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from all other species in the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[660,735,1543,1569]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="236" pageNumber="237" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[660,735,1543,1569]" italics="true" pageId="236" pageNumber="237">salvini</emphasis>
</taxonomicName>
clade by the following character combination: promesonotum evenly convex; metanotal groove deeply impressed and propodeum depressed below the level of the promesonotum; propodeal spines absent: dorsal face of propodeum transitioning evenly in to the propodeal declivity; in profile view, postpetiole bulging anterodorsally and flat posterodorsally; in dorsal view, petiolar node 1.3 to 1.6 times as broad as caudal cylinder; integument predominantly dark brown; setae on head, mesosoma, legs, waist segments and gaster erect, moderately long, abundant and tapering.
</paragraph>
<paragraph blockId="236.[498,1542,1180,1928]" lastBlockId="237.[498,1542,234,1934]" lastPageId="237" lastPageNumber="238" pageId="236" pageNumber="237">
<emphasis bold="true" box="[530,719,1822,1848]" pageId="236" pageNumber="237">Similar species:</emphasis>
this striking species is difficult to confuse with any other species in
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1862,1888]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="236" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis box="[498,655,1862,1888]" italics="true" pageId="236" pageNumber="237">Temnothorax</emphasis>
</taxonomicName>
, except the closely related
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1895" box="[968,1111,1862,1888]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="236" pageNumber="237" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis box="[968,1111,1862,1888]" italics="true" pageId="236" pageNumber="237">T. pergandei</emphasis>
</taxonomicName>
. The absence of propodeal teeth, the structure of the postpetiole, which is evenly convex dorsally in
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1895" box="[1243,1388,1902,1928]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="236" pageNumber="237" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis box="[1243,1388,1902,1928]" italics="true" pageId="236" pageNumber="237">T. pergandei</emphasis>
</taxonomicName>
, and the relative width of the petiolar node in dorsal view (1.1 to 1.3 times as broad as the caudal cylinder in
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1895" box="[634,778,274,300]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="237" pageNumber="238" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis box="[634,778,274,300]" italics="true" pageId="237" pageNumber="238">T. pergandei</emphasis>
</taxonomicName>
) will separate the two species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="239" lastPageNumber="240" pageId="237" pageNumber="238" type="description">
<paragraph blockId="237.[498,1542,234,1934]" pageId="237" pageNumber="238">
<emphasis bold="true" box="[530,1018,313,340]" pageId="237" pageNumber="238">
Worker measurements &amp; indices (
<emphasis bold="true" box="[941,957,314,340]" italics="true" pageId="237" pageNumber="238">n</emphasis>
= 8):
</emphasis>
SL =
<geoCoordinate box="[1084,1146,314,341]" degrees="0.535" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.535">0.535</geoCoordinate>
<geoCoordinate box="[1146,1223,314,341]" degrees="0.675" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.675">0.675</geoCoordinate>
(0.596); FRS =
<geoCoordinate box="[1402,1464,314,340]" degrees="0.171" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.171">0.171</geoCoordinate>
<geoCoordinate box="[1464,1541,314,340]" degrees="0.244" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.244">0.244</geoCoordinate>
(0.214); CW =
<geoCoordinate box="[678,740,354,380]" degrees="0.602" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.602">0.602</geoCoordinate>
<geoCoordinate box="[740,817,354,380]" degrees="0.748" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.748">0.748</geoCoordinate>
(0.678); CWb =
<geoCoordinate box="[1020,1082,354,381]" degrees="0.548" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.548">0.548</geoCoordinate>
<geoCoordinate box="[1082,1159,354,380]" degrees="0.686" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.686">0.686</geoCoordinate>
(0.627); PoOC =
<geoCoordinate box="[1372,1434,354,380]" degrees="0.229" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.229">0.229</geoCoordinate>
<geoCoordinate box="[1434,1511,354,380]" degrees="0.282" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.282">0.282</geoCoordinate>
(0.253); CL =
<geoCoordinate box="[665,727,393,420]" degrees="0.651" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.651">0.651</geoCoordinate>
<geoCoordinate box="[727,804,394,420]" degrees="0.783" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.783">0.783</geoCoordinate>
(0.710); EL =
<geoCoordinate box="[977,1039,393,420]" degrees="0.158" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.158">0.158</geoCoordinate>
<geoCoordinate box="[1039,1116,394,420]" degrees="0.207" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.207">0.207</geoCoordinate>
(0.184); EW =
<geoCoordinate box="[1302,1364,394,420]" degrees="0.114" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.114">0.114</geoCoordinate>
<geoCoordinate box="[1364,1441,394,420]" degrees="0.136" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.136">0.136</geoCoordinate>
(0.124); MD =
<geoCoordinate box="[580,642,434,460]" degrees="0.143" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.143">0.143</geoCoordinate>
<geoCoordinate box="[642,719,433,460]" degrees="0.178" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.178">0.178</geoCoordinate>
(0.158); WL =
<geoCoordinate box="[904,966,433,459]" degrees="0.816" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.816">0.816</geoCoordinate>
<geoCoordinate box="[966,1043,433,460]" degrees="0.990" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.99">0.990</geoCoordinate>
(0.901); SPST =
<geoCoordinate box="[1245,1307,434,460]" degrees="0.119" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.119">0.119</geoCoordinate>
<geoCoordinate box="[1308,1385,433,460]" degrees="0.158" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.158">0.158</geoCoordinate>
(0.141); MPST =
<geoCoordinate box="[607,669,473,499]" degrees="0.242" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.242">0.242</geoCoordinate>
<geoCoordinate box="[669,746,473,499]" degrees="0.336" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.336">0.336</geoCoordinate>
(0.300); PEL =
<geoCoordinate box="[935,997,473,499]" degrees="0.311" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.311">0.311</geoCoordinate>
<geoCoordinate box="[997,1074,473,499]" degrees="0.413" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.413">0.413</geoCoordinate>
(0.366); NOL =
<geoCoordinate box="[1274,1336,473,499]" degrees="0.196" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.196">0.196</geoCoordinate>
<geoCoordinate box="[1336,1413,473,499]" degrees="0.227" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.227">0.227</geoCoordinate>
(0.211); NOH =
<geoCoordinate box="[598,660,513,539]" degrees="0.094" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.094">0.094</geoCoordinate>
<geoCoordinate box="[660,737,513,539]" degrees="0.129" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.129">0.129</geoCoordinate>
(0.112); PEH =
<geoCoordinate box="[933,995,513,540]" degrees="0.175" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.175">0.175</geoCoordinate>
<geoCoordinate box="[995,1072,513,539]" degrees="0.239" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.239">0.239</geoCoordinate>
(0.209); PPL =
<geoCoordinate box="[1262,1324,513,540]" degrees="0.156" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.156">0.156</geoCoordinate>
<geoCoordinate box="[1324,1401,513,539]" degrees="0.193" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.193">0.193</geoCoordinate>
(0.173); PPH =
<geoCoordinate box="[587,649,553,579]" degrees="0.217" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.217">0.217</geoCoordinate>
<geoCoordinate box="[649,726,553,579]" degrees="0.278" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.278">0.278</geoCoordinate>
(0.248); PW =
<geoCoordinate box="[911,973,553,580]" degrees="0.395" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.395">0.395</geoCoordinate>
<geoCoordinate box="[973,1050,553,580]" degrees="0.513" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.513">0.513</geoCoordinate>
(0.450); SBPA = n/a; SPTI =
<geoCoordinate box="[1402,1464,553,579]" degrees="0.163" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.163">0.163</geoCoordinate>
<geoCoordinate box="[1464,1541,553,579]" degrees="0.211" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.211">0.211</geoCoordinate>
(0.186); PEW =
<geoCoordinate box="[691,753,593,619]" degrees="0.141" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.141">0.141</geoCoordinate>
<geoCoordinate box="[754,830,593,619]" degrees="0.176" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.176">0.176</geoCoordinate>
(0.161); PNW =
<geoCoordinate box="[1038,1100,593,619]" degrees="0.197" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.197">0.197</geoCoordinate>
<geoCoordinate box="[1100,1177,593,619]" degrees="0.282" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.282">0.282</geoCoordinate>
(0.238); PPW =
<geoCoordinate box="[1379,1441,593,619]" degrees="0.234" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.234">0.234</geoCoordinate>
<geoCoordinate box="[1441,1518,593,619]" degrees="0.314" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.314">0.314</geoCoordinate>
(0.270); HFL =
<geoCoordinate box="[683,745,632,659]" degrees="0.555" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.555">0.555</geoCoordinate>
<geoCoordinate box="[745,822,633,659]" degrees="0.732" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.732">0.732</geoCoordinate>
(0.648); HFWmax =
<geoCoordinate box="[1079,1141,633,659]" degrees="0.126" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.126">0.126</geoCoordinate>
<geoCoordinate box="[1141,1218,632,659]" degrees="0.157" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.157">0.157</geoCoordinate>
(0.141); HFWmin =
<geoCoordinate box="[645,707,672,698]" degrees="0.046" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.046">0.046</geoCoordinate>
<geoCoordinate box="[708,785,672,699]" degrees="0.060" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.06">0.060</geoCoordinate>
(0.052); CS =
<geoCoordinate box="[958,1020,673,699]" degrees="0.874" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.874">0.874</geoCoordinate>
<geoCoordinate box="[1021,1098,672,699]" degrees="1.069" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-1.069">1.069</geoCoordinate>
(0.982); ES =
<geoCoordinate box="[1269,1331,672,699]" degrees="0.215" direction="north" orientation="latitude" pageId="237" pageNumber="238" precision="55" value="0.215">0.215</geoCoordinate>
<geoCoordinate box="[1331,1408,673,699]" degrees="0.273" direction="west" orientation="longitude" pageId="237" pageNumber="238" precision="55" value="-0.273">0.273</geoCoordinate>
(0.246); SI = 8799 (95); OI = 2327 (25); CI = 8492 (88); WLI = 141150 (144); SBI = n/a; PSI = 1418 (16); PWI = 149184 (167); PLI = 194232 (212); NI = 167209 (189); PNWI = 125164 (147); NLI = 5363 (58); FI = 254292 (275).
</paragraph>
<paragraph blockId="237.[498,1542,234,1934]" pageId="237" pageNumber="238">
<emphasis bold="true" box="[530,775,832,858]" pageId="237" pageNumber="238">Worker description:</emphasis>
In full-face view, head subquadrate, longer than broad (CI 8492). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin weakly convex medially. Antennal scapes very long: when fully retracted, surpassing the posterior margin of the head capsule by about two times the maximum width of the scape (SI 8799). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment as long as the preceding two in combination. Frontal carinae short, extending past the antennal toruli by about one and a half times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin weakly convex, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="237.[498,1542,234,1934]" lastBlockId="238.[498,1542,234,1894]" lastPageId="238" lastPageNumber="239" pageId="237" pageNumber="238">
In profile view, compound eyes elongate ovular and moderately large (OI 2327), with 11 ommatidia in longest row. Pronotal declivity indistinct, neck and anterior face of pronotum forming a ~120
<emphasis box="[809,819,1425,1443]" italics="true" pageId="237" pageNumber="238">
<superScript attach="left" box="[809,819,1425,1443]" fontSize="8" pageId="237" pageNumber="238">°</superScript>
</emphasis>
angle. Mesosoma with promesonotum forming an even convexity; the propodeum depressed below the level of the promesonotum and dorsally concave. Promesonotal suture extending from the posterior margin of the procoxal insertion to the mesothoracic spiracle, which is moderately well developed, then continuing to the dorsal surface as a very weak sulcus. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture; continuing dorsally as a distinct sulcus. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about three spiracle diameters. Propodeal spines absent, but with a cuticular flange extending from the propodeal dorsum to the propodeal lobes in their place. Propodeal declivity flat, forming a rounded ~100
<emphasis box="[1150,1160,1863,1881]" italics="true" pageId="237" pageNumber="238">
<superScript attach="left" box="[1150,1160,1863,1881]" fontSize="8" pageId="237" pageNumber="238">°</superScript>
</emphasis>
angle with propodeal dorsum. Propodeal lobes rounded and weakly developed. Metapleural gland bulla moderately large, extending from the metacoxal insertion two thirds of the way to the propodeal spiracle. Petiole long (PLI 194232), without tubercles anterodorsally. Subpetiolar process in the form of a small, triangular, blunt tooth, which continues as a low carina to the caudal cylinder; ventral margin of petiole posterior to the subpetiolar process strongly concave. Petiolar peduncle moderately long: comprising about half the length of the petiole. Petiolar node cuneiform but rounded dorsally: transition between peduncle and node marked by a rounded angle of ~150
<emphasis box="[767,777,469,487]" italics="true" pageId="238" pageNumber="239">
<superScript attach="none" box="[767,777,469,487]" fontSize="8" pageId="238" pageNumber="239">°</superScript>
</emphasis>
, resulting in a weakly concave anterior node face; anterior face forming a broadly rounded ~60
<emphasis box="[863,873,509,527]" italics="true" pageId="238" pageNumber="239">
<superScript attach="left" box="[863,873,509,527]" fontSize="8" pageId="238" pageNumber="239">°</superScript>
</emphasis>
angle with the posterior face; caudal cylinder short, about half the maximum width of the antennal scape. Postpetiole weakly convex anteriorly, rounding evenly into the dorsal face, which bulges strongly before flattening posteriorly; lobed ventrally.
</paragraph>
<paragraph blockId="238.[498,1542,234,1894]" pageId="238" pageNumber="239">In dorsal view, humeri weakly developed: evenly rounded and wider than the rest of the mesosoma, but not distinct from it; mesothoracic spiracles weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Promesonotal suture visible as a weak sulcus. Metanotal strongly impressed, distinctly dividing the promesonotum from the propodeum. Propodeal spines absent, but flanges diverging at the apex of the propodeal dorsum. Petiolar peduncle with spiracles weakly protruding past the lateral margins, but not noticeably constricted anterior to them. Petiolar node, when viewed posterodorsally, trapezoidal and apically broadened, the apex weakly convex; caudal cylinder narrower than the apex of the node. Postpetiole narrow (PWI 149184) and trapezoidal: widest anteriorly. Anterior margin of the postpetiole weakly convex and meeting the lateral margins at a rounded angle; lateral margins converge moderately to the angulate posterior corners; posterior margin broadly concave. Metafemur moderately incrassate (FI 254292).</paragraph>
<paragraph blockId="238.[498,1542,234,1894]" pageId="238" pageNumber="239">Sculpture: median clypeal carina present, extending posteriorly to the level of the anterior margins of the antennal insertions. Lateral clypeal lobes with additional, weaker carinae; ground sculpture smooth and shining. Antennal scapes shining through weak coriarious ground sculpture. Cephalic dorsum predominantly coriarious, with costulae flanking the frontal carinae; concentric costulae surrounding the antennal insertions, extending from the apices of the frontal carinae, to the anterior margins of the lateral clypeal lobes. Lateral surfaces of head with weak coriarious sculpture posterior to the compound eye, denser coriarious sculpture surrounding the compound eye, and stronger costulae between the compound eye and the mandibular insertion. Ventral surface of head smooth and shining. Pronotal neck areolate. Lateral surface of the pronotum areolate-coriarious on the anterior third, otherwise smooth and shining. Lateral face of the propodeum, meso- and metapleurae predominantly costate, over areolate ground sculpture; area around propodeal spiracle finely areolate. Propodeal declivity smooth and shining. Dorsal surface of promesonotum coriarious. Dorsum of propodeum strongly rugose. Femora smooth and shining. Petiole smooth and shining ventrally; node weakly coriarious; dorsum of peduncle finely areolate. Postpetiole weakly coriarious. First gastral tergite weakly coriarious; first gastral sternite smooth and shining. Spectral iridescence absent.</paragraph>
<paragraph blockId="239.[498,1542,234,739]" pageId="239" pageNumber="240">Setae: antennal scapes and funiculi with moderately long, decumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, tapering setae, the longest of which are about the length of the compound eye. The head bears ~40, mesosoma ~38, petiole ~14, postpetiole ~20, and first gastral tergite ~60 setae. Pubescence is long, coarse, difficult to distinguish from the setae, and present over the entire body except for the ventral surface of the petiole.</paragraph>
<paragraph blockId="239.[498,1542,234,739]" box="[530,1542,473,500]" pageId="239" pageNumber="240">Color: predominantly dark brown, with mandibles, antennae, tibiae and tarsi testaceous.</paragraph>
<paragraph blockId="239.[498,1542,234,739]" box="[530,734,513,539]" pageId="239" pageNumber="240">
<emphasis bold="true" box="[530,601,513,539]" pageId="239" pageNumber="240">Gyne:</emphasis>
Unknown.
</paragraph>
<paragraph blockId="239.[498,1542,234,739]" box="[530,730,553,579]" pageId="239" pageNumber="240">
<emphasis bold="true" box="[530,597,553,579]" pageId="239" pageNumber="240">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection box="[530,1542,593,619]" pageId="239" pageNumber="240" type="etymology">
<paragraph blockId="239.[498,1542,234,739]" box="[530,1542,593,619]" pageId="239" pageNumber="240">
<emphasis bold="true" box="[530,668,593,619]" pageId="239" pageNumber="240">Etymology:</emphasis>
Morphological, in reference to the humpbacked appearance of the worker.
</paragraph>
</subSubSection>
<subSubSection pageId="239" pageNumber="240" type="discussion">
<paragraph blockId="239.[498,1542,234,739]" pageId="239" pageNumber="240">
<emphasis bold="true" box="[530,670,632,658]" pageId="239" pageNumber="240">Comments:</emphasis>
Despite multiple collections, very little is known about the biology of
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,725,673,699]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="239" pageNumber="240" phylum="Arthropoda" rank="species" species="bison" status="sp. nov.">
<emphasis box="[498,725,673,699]" italics="true" pageId="239" pageNumber="240">Temnothorax bison</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[732,824,673,699]" pageId="239" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
It appears to inhabit low-to-mid elevation tropical dry forest or xeric scrub habitats and forage terrestrially, much like its close relative,
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1895" box="[1378,1523,712,738]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="239" pageNumber="240" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis box="[1378,1523,712,738]" italics="true" pageId="239" pageNumber="240">T. pergandei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>