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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.5.e9649" ID-PMC="PMC5345113" ID-Pensoft-Pub="1314-2828--9649" ID-PubMed="28325972" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1314-2828--e9649" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Nine new species of Uramya Robineau-Desvoidy (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica, with a key to their identification" checkinTime="1585357165802" checkinUser="pensoft" docAuthor="Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Hallwachs, Winnie, Janzen, Daniel &amp; Dapkey, Tanya" docDate="2017" docId="8420D05751AE5579D1BEC73D7D521054" docLanguage="en" docName="BiodivDatJour 5: e9649" docOrigin="Biodiversity Data Journal 5" docSource="http://dx.doi.org/10.3897/BDJ.5.e9649" docTitle="Uramya contraria Fleming &amp; Wood, sp. n." docType="treatment" docVersion="3" lastPageNumber="9649" masterDocId="1B26FFD87A2883622824B027C36CFFE1" masterDocTitle="Nine new species of Uramya Robineau-Desvoidy (Diptera: Tachinidae) from Area de Conservacion Guanacaste in northwestern Costa Rica, with a key to their identification" masterLastPageNumber="9649" masterPageNumber="9649" pageNumber="9649" updateTime="1668124532315" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Nine new species of Uramya Robineau-Desvoidy (Diptera: Tachinidae) from Area de Conservacion Guanacaste in northwestern Costa Rica, with a key to their identification</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fleming, AJ</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wood, D. Monty</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, M. Alex</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hallwachs, Winnie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Janzen, Daniel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dapkey, Tanya</mods:namePart>
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<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
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<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>5</mods:number>
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<mods:start>9649</mods:start>
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<mods:url>http://dx.doi.org/10.3897/BDJ.5.e9649</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.5.e9649</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--9649</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:8420D05751AE5579D1BEC73D7D521054" httpUri="http://treatment.plazi.org/id/8420D05751AE5579D1BEC73D7D521054" lastPageNumber="9649" pageId="0" pageNumber="9649">
<subSubSection pageId="0" pageNumber="9649" type="nomenclature">
<paragraph pageId="0" pageNumber="9649">
<taxonomicName LSID="urn:lsid:zoobank.org:act:A95CABAB-81A0-4C60-BA89-8A457B1E773E" authority="Fleming &amp; Wood" class="Insecta" family="Tachinidae" genus="Uramya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uramya contraria" order="Diptera" pageId="0" pageNumber="9649" phylum="Arthropoda" rank="species" species="contraria">Uramya contraria Fleming &amp; Wood</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="9649">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9649" type="materials_examined">
<paragraph pageId="0" pageNumber="9649">Materials</paragraph>
<paragraph pageId="0" pageNumber="9649">
<materialsCitation collectingDate="2007-09-01" collectingMethod="reared from caterpillar of CaviriareginaDHJ 04 (Erebidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen &amp; W. Hallwachs, Calixto Moraga, Petrona Rios, Manuel Rios" country="Costa Rica" latitude="11.014" location="Sector Pitilla" longitude="-85.424" pageId="0" pageNumber="9649" specimenCode="DHJPAR 0022003" specimenCount="1" typeStatus="Holotype">
Type status:
<typeStatus pageId="0" pageNumber="9649">Holotype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="9649">DHJPAR0022003</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="9649">D.H. Janzen &amp; W. Hallwachs, Calixto Moraga, Petrona Rios, Manuel Rios</collectorName>
; individualID: DHJPAR0022003; individualCount:
<specimenCount pageId="0" pageNumber="9649">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="9649">M</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="9649">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: ASTAT1141-07, 07-SRNP-32916; Taxon: scientificName: Uramyacontraria; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Uramya; specificEpithet: contraria; scientificNameAuthorship: Fleming &amp; Wood; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="9649">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Area de
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
Guanacaste; locality:
<location LSID="urn:lsid:plazi:treatment:8420D05751AE5579D1BEC73D7D521054:8A4E2654A41BE04E7E243AB608306AA4" country="Costa Rica" latitude="11.014" longitude="-85.424" name="Sector Pitilla" pageId="0" pageNumber="9649">Sector Pitilla</location>
; verbatimLocality: Sendero Rotulo; verbatimElevation:
<elevation pageId="0" pageNumber="9649">510</elevation>
; verbatimLatitude: 11.014; verbatimLongitude: -85.424; verbatimCoordinateSystem: Decimal; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="9649" value="11.014">11.014</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="9649" value="-85.424">-85.424</geoCoordinate>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="9649">AJ Fleming</determinerName>
; dateIdentified: 2015; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="9649">reared from caterpillar of CaviriareginaDHJ04 (Erebidae)</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="9649" value="2007-09-01">01-Sep-2007</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="9649">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="9649">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation collectingDate="2007-08-26" collectingMethod="reared from caterpillar of CaviriareginaDHJ 04 (Erebidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen &amp; W. Hallwachs, Calixto Moraga, Petrona Rios, Manuel Rios" country="Costa Rica" latitude="11.011" location="Sector Pitilla" longitude="-85.421" pageId="0" pageNumber="9649" specimenCode="DHJPAR 0021038" specimenCount="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="9649">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="9649">DHJPAR0021038</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="9649">D.H. Janzen &amp; W. Hallwachs, Calixto Moraga, Petrona Rios, Manuel Rios</collectorName>
; individualID: DHJPAR0021038; individualCount:
<specimenCount pageId="0" pageNumber="9649">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="9649">F</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="9649">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: ASTA1381-07, 07-SRNP-32877; Taxon: scientificName: Uramyacontraria; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Uramya; specificEpithet: contraria; scientificNameAuthorship: Fleming &amp; Wood; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="9649">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Area de
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
Guanacaste; locality:
<location LSID="urn:lsid:plazi:treatment:8420D05751AE5579D1BEC73D7D521054:586C90870006CF6B16E47BA8632BFA25" country="Costa Rica" latitude="11.011" longitude="-85.421" name="Sector Pitilla" pageId="0" pageNumber="9649">Sector Pitilla</location>
; verbatimLocality: Casa Roberto; verbatimElevation:
<elevation pageId="0" pageNumber="9649">520</elevation>
; verbatimLatitude: 11.011; verbatimLongitude: -85.421; verbatimCoordinateSystem: Decimal; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="9649" value="11.011">11.011</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="9649" value="-85.421">-85.421</geoCoordinate>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="9649">AJ Fleming</determinerName>
; dateIdentified: 2015; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="9649">reared from caterpillar of CaviriareginaDHJ04 (Erebidae)</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="9649" value="2007-08-26">26-Aug-2007</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="9649">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="9649">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation collectingDate="2007-08-31" collectingMethod="reared from caterpillar of CaviriareginaDHJ 04 (Erebidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen &amp; W. Hallwachs, Calixto Moraga, Petrona Rios, Manuel Rios" country="Costa Rica" latitude="11.011" location="Sector Pitilla" longitude="-85.421" pageId="0" pageNumber="9649" specimenCode="DHJPAR 0021039" specimenCount="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="9649">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="9649">DHJPAR0021039</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="9649">D.H. Janzen &amp; W. Hallwachs, Calixto Moraga, Petrona Rios, Manuel Rios</collectorName>
; individualID: DHJPAR0021039; individualCount:
<specimenCount pageId="0" pageNumber="9649">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="9649">F</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="9649">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: ASTA1382-07, 07-SRNP-32922; Taxon: scientificName: Uramyacontraria; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Uramya; specificEpithet: contraria; scientificNameAuthorship: Fleming &amp; Wood; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="9649">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Area de
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
Guanacaste; locality:
<location LSID="urn:lsid:plazi:treatment:8420D05751AE5579D1BEC73D7D521054:63556E81A51262CC2593D9359E588C26" country="Costa Rica" latitude="11.011" longitude="-85.421" name="Sector Pitilla" pageId="0" pageNumber="9649">Sector Pitilla</location>
; verbatimLocality: Casa Roberto; verbatimElevation:
<elevation pageId="0" pageNumber="9649">520</elevation>
; verbatimLatitude: 11.011; verbatimLongitude: -85.421; verbatimCoordinateSystem: Decimal; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="9649" value="11.011">11.011</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="9649" value="-85.421">-85.421</geoCoordinate>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="9649">AJ Fleming</determinerName>
; dateIdentified: 2015; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="9649">reared from caterpillar of CaviriareginaDHJ04 (Erebidae)</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="9649" value="2007-08-31">31-Aug-2007</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="9649">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="9649">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation collectingDate="2007-09-13" collectingMethod="reared from caterpillar of CaviriareginaDHJ 04 (Erebidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen &amp; W. Hallwachs, Calixto Moraga, Petrona Rios, Manuel Rios" country="Costa Rica" latitude="11.019" location="Sector Pitilla" longitude="-85.41" pageId="0" pageNumber="9649" specimenCode="DHJPAR 0022005" specimenCount="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="9649">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="9649">DHJPAR0022005</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="9649">D.H. Janzen &amp; W. Hallwachs, Calixto Moraga, Petrona Rios, Manuel Rios</collectorName>
; individualID: DHJPAR0022005; individualCount:
<specimenCount pageId="0" pageNumber="9649">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="9649">F</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="9649">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: ASTAT1143-07, 07-SRNP-33078; Taxon: scientificName: Uramyacontraria; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Uramya; specificEpithet: contraria; scientificNameAuthorship: Fleming &amp; Wood; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="9649">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Area de
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
Guanacaste; locality:
<location LSID="urn:lsid:plazi:treatment:8420D05751AE5579D1BEC73D7D521054:D87BD2E717500A01654BF68E1EDB064C" country="Costa Rica" latitude="11.019" longitude="-85.41" name="Sector Pitilla" pageId="0" pageNumber="9649">Sector Pitilla</location>
; verbatimLocality: Pasmompa; verbatimElevation:
<elevation pageId="0" pageNumber="9649">440</elevation>
; verbatimLatitude: 11.019; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="9649" value="11.019">11.019</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="9649" value="-85.41">-85.41</geoCoordinate>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="9649">AJ Fleming</determinerName>
; dateIdentified: 2015; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="9649">reared from caterpillar of CaviriareginaDHJ04 (Erebidae)</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="9649" value="2007-09-13">13-Sep-2007</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="9649">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="9649">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9649" type="description">
<paragraph pageId="0" pageNumber="9649">Description</paragraph>
<paragraph pageId="0" pageNumber="9649">
Male (Fig. 7). Length: 12
<normalizedToken originalValue="-">--</normalizedToken>
13 mm. Head (Fig. 7b): antenna black, with orange medially and at base, adjacent to pedicel; pedicel black; arista 1.5X as long as first flagellomere, dark brown and minutely pubescent; palpus dark yellow and haired; fronto-orbital plate, parafacial and gena silver pollinose; gena bearing fine hairs along lower margin. Thorax (Fig. 7b, c): entirely gray pollinose; dorsum of thorax and scutellum covered by conspicuous yellow-white hairs; sternopleura, hypopleura, pteropleura, and ventral surface of abdomen yellow-white pilose; 2 katepisternal bristles; 3 postsutural supra-alar bristles, 2nd postsutural supra-alar 4X as long as first; postpronotum bearing fine black hairs and anepisternum with fine yellow-white hairs; scutellum bearing 1 pair of discal bristles; underside of scutellum bearing a tuft of white hairs near basal marginal bristle. Legs: reddish-yellow in ground color; femora covered in long, yellow hairs interspersed among darker hairs and bristles; tarsi all black. Wing: smoky gray translucent; wing veins strongly infuscate; infuscation of wing veins becoming more generalized and blending together to a dark gray tone along basal portion of the wing. Abdomen (Fig. 7a): 1 pair of median marginal bristles on ST1+2 and T3; row of marginal bristles on T4 and T5; 2 pairs of median discal bristles on T3 and 1 pair on T4; abdomen dark brown dorsally, with silver pollen on anterior half of T3, T4 and T5; ST1+2 with 2 pollinose spots on either side of mid-dorsal depression; underside of abdomen entirely covered in silver pollinosity. Terminalia (Fig. 7d, e, f): sternite 5 with two small lobes; inner margin covered in dense pollinosity, appearing darker than surrounding cuticle; apical edges of lobes of sternite 5 bearing many long, stout, outwardly pointed bristles interspersed with longer bristles close to lobe margin; sternite 5 with wide V-shaped median cleft, 0.32X length of sternite from lobe apex to base; cercus sharply pointed and distinctly tapered; apical section 1.7X length of upper lobe; slightly curved when viewed laterally, with a slight upward hook at its tip; surstylus narrow, curved and scythe-like in lateral view; middle third of surstylus haired, otherwise almost bare; tip of surstylus not lobed when viewed dorsally and distinctly inwardly angled; surstylus 0.9X as long as cercus.
</paragraph>
<paragraph pageId="0" pageNumber="9649">Female (Fig. 8). Length: 8-11 mm. As male, except with a slight golden tinge around the thoracic suture, one pair of median discal bristles on T3, T4 and T5, wing smoky brown translucent, only slightly infuscate around major wing veins.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9649" type="diagnosis">
<paragraph pageId="0" pageNumber="9649">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="9649">
<taxonomicName class="Insecta" family="Tachinidae" genus="Uramya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uramya contraria" order="Diptera" pageId="0" pageNumber="9649" phylum="Arthropoda" rank="species" species="contraria">Uramya contraria</taxonomicName>
can be distinguished from all other Neotropical species of
<taxonomicName class="Insecta" family="Tachinidae" genus="Uramya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uramya" order="Diptera" pageId="0" pageNumber="9649" phylum="Arthropoda" rank="genus">Uramya</taxonomicName>
by the following combination of traits: dark brown to black antennae, 3 postsutural supra-alar bristles, 2 strong lateral scutellar bristles, no discal scutellar bristles, underside of scutellum with a tuft of white hairs near basal marginal bristle, abdomen flattened dorsoventrally, ST1+2 with lateral white pollinose spots on either side of mid-dorsal depression, T5 subtriangular, not strongly produced into a long, tail-like process, 1 pair of median discal bristles on T3, T4 and T5, and silver pollinosity on underside of abdomen.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9649" type="etymology">
<paragraph pageId="0" pageNumber="9649">Etymology</paragraph>
<paragraph pageId="0" pageNumber="9649">
The species epithet is derived from the latin adjective &quot; contrarius &quot; for contrary or opposed, referring to its overall similarity but differing terminalia to
<taxonomicName lsidName="U. halisidotae" pageId="0" pageNumber="9649" rank="species" species="halisidotae">U. halisidotae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9649" type="distribution">
<paragraph pageId="0" pageNumber="9649">Distribution</paragraph>
<paragraph pageId="0" pageNumber="9649">Costa Rica, ACG (Prov. Guanacaste), 440-520 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9649" type="ecology">
<paragraph pageId="0" pageNumber="9649">Ecology</paragraph>
<paragraph pageId="0" pageNumber="9649">
<taxonomicName class="Insecta" family="Tachinidae" genus="Uramya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uramya contraria" order="Diptera" pageId="0" pageNumber="9649" phylum="Arthropoda" rank="species" species="contraria">Uramya contraria</taxonomicName>
has been reared four times from a sample of 100 wild-caught, non-sibling
<taxonomicName class="Insecta" family="Lymantriidae" genus="Caviria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caviria regina" order="Lepidoptera" pageId="0" pageNumber="9649" phylum="Arthropoda" rank="species" species="regina">Caviria regina</taxonomicName>
Cramer (
<taxonomicName family="Erebidae" lsidName="" pageId="0" pageNumber="9649" rank="family">Erebidae</taxonomicName>
,
<taxonomicName lsidName="" pageId="0" pageNumber="9649" rank="subfamily" subfamily="Lymantriinae">Lymantriinae</taxonomicName>
) in ACG rain forest.
</paragraph>
</subSubSection>
</treatment>
</document>