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<document id="D699AAFB07C3FBCC1DDB94AE78C99ED1" ID-DOI="10.11646/zootaxa.5543.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14386297" ID-ZooBank="C90518B7-B880-4290-95FE-A0F51EDE1E98" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1733918563993" checkinUser="plazi" docAuthor="Kunte, Krushnamegh &amp; Basu, Dipendra Nath" docDate="2024" docId="61670B57DA1C783321B48657AD6BFEB1" docLanguage="en" docName="zootaxa.5543.3.3.pdf" docOrigin="Zootaxa 5543 (3)" docSource="https://doi.org/10.11646/zootaxa.5543.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athyma inara subsp. inara" docType="treatment" docVersion="3" lastPageNumber="345" masterDocId="9D5E732FDA1E783021238164AC3EFFD4" masterDocTitle="Two new taxa of brush-footed butterflies (Lepidoptera: Nymphalidae) from the Western Ghats biodiversity hotspot, southern India" masterLastPageNumber="367" masterPageNumber="343" pageNumber="345" updateTime="1734447740349" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="DE2BE2822E89AB1A73155EE6CAA08BD5">Two new taxa of brush-footed butterflies (Lepidoptera: Nymphalidae) from the Western Ghats biodiversity hotspot, southern India</mods:title>
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<mods:namePart id="0C679AD35CD42B0B320AFF16EDA9884F">Kunte, Krushnamegh</mods:namePart>
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<mods:affiliation id="4B34C61B051AC3824D586600326409A4">National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560 065, India</mods:affiliation>
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<mods:namePart id="4B7FB50A9911996235E7B54E9707324C">Basu, Dipendra Nath</mods:namePart>
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<mods:affiliation id="D0740F008A018E8745ECB4BD3A14B995">National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560 065, India &amp; Ashoka University, Rajiv Gandhi Education City, Sonipat, Haryana 131029, India</mods:affiliation>
<mods:nameIdentifier id="D4BFC5FD0A36E3E5E0B437703E4EE6E6" type="email">dipendra.basu@ashoka.edu.in</mods:nameIdentifier>
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<subSubSection id="A1D4E9CADA1C783221B48657ADFFF899" box="[151,449,1842,1869]" pageId="2" pageNumber="345" type="nomenclature">
<paragraph id="E971BA41DA1C783221B48657ADFFF899" blockId="2.[151,1437,150,2013]" box="[151,449,1842,1869]" pageId="2" pageNumber="345">
12.
<emphasis id="DBBA6653DA1C783221E48656ADFFF899" bold="true" box="[199,449,1842,1869]" pageId="2" pageNumber="345">
<taxonomicName id="2ECEC1C2DA1C783221E48656AD86F899" box="[199,440,1842,1869]" class="Insecta" family="Nymphalidae" genus="Athyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="345" phylum="Arthropoda" rank="subSpecies" species="inara" subSpecies="inara">
<emphasis id="DBBA6653DA1C783221E48656AD86F899" bold="true" box="[199,440,1842,1869]" italics="true" pageId="2" pageNumber="345">Athyma inara inara</emphasis>
</taxonomicName>
:
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A1D4E9CADA1C783220ED8657A8B7F8A5" pageId="2" pageNumber="345" type="reference_group">
<paragraph id="E971BA41DA1C783220ED8657A8B7F8A5" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="345">
This includes the following synonyms: (a)
<taxonomicName id="2ECEC1C2DA1C783222F08657A90AF899" authority="(Evans 1932)" baseAuthorityName="Evans" baseAuthorityYear="1932" box="[979,1332,1843,1869]" class="Insecta" family="Nymphalidae" genus="Athyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="345" phylum="Arthropoda" rank="species" species="inarina">
<emphasis id="DBBA6653DA1C783222F08657A8B6F898" box="[979,1160,1843,1869]" italics="true" pageId="2" pageNumber="345">Athyma inarina</emphasis>
(
<bibRefCitation id="8D5FC7B0DA1C783225BE8657A913F899" author="Evans, W. H." box="[1181,1325,1843,1869]" pageId="2" pageNumber="345" refId="ref13172" refString="Evans, W. H. (1932) The Identification of Indian Butterflies (2 nd edn.). Bombay Natural History Society, Mumbai, 454 pp." type="book" year="1932">Evans 1932</bibRefCitation>
)
</taxonomicName>
, and (b)
<taxonomicName id="2ECEC1C2DA1C783221E48633AE03F8A5" authority="Swinhoe, 1917" authorityName="Swinhoe" authorityYear="1917" box="[199,573,1879,1905]" class="Insecta" family="Nymphalidae" genus="Pantoporia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="345" phylum="Arthropoda" rank="species" species="mera">
<emphasis id="DBBA6653DA1C783221E48633ADB4F8A4" box="[199,394,1879,1904]" italics="true" pageId="2" pageNumber="345">Pantoporia mera</emphasis>
<bibRefCitation id="8D5FC7B0DA1C783220B08633AE03F8A5" author="Swinhoe, C." box="[403,573,1879,1905]" pageId="2" pageNumber="345" pagination="407 - 412" refId="ref15041" refString="Swinhoe, C. (1917) New Indo-Malayan Lepidoptera. Annals and Magazine of Natural History, 20, 407 - 412. https: // doi. org / 10.1080 / 00222931709487028" type="journal article" year="1917">Swinhoe, 1917</bibRefCitation>
</taxonomicName>
(see below for a note on synonymy of this taxon).
</paragraph>
</subSubSection>
<subSubSection id="A1D4E9CADA1C783325B2863CAD6BFEB1" lastPageId="3" lastPageNumber="346" pageId="2" pageNumber="345" type="materials_examined">
<paragraph id="E971BA41DA1C783325B2863CAD6BFEB1" blockId="2.[151,1437,150,2013]" lastBlockId="3.[151,1437,150,753]" lastPageId="3" lastPageNumber="346" pageId="2" pageNumber="345">
<materialsCitation id="59A6B01CDA1C783325B2863CAD6CFEB1" lastPageId="3" lastPageNumber="346" pageId="2" pageNumber="345">
<emphasis id="DBBA6653DA1C783225B2863CA88EF8A0" bold="true" box="[1169,1200,1880,1908]" pageId="2" pageNumber="345">
<specimenCount id="FFC871C8DA1C783225B2863CA896F8A0" box="[1169,1192,1880,1908]" count="1" pageId="2" pageNumber="345" type="male"></specimenCount>
:
</emphasis>
BMNH(E) #933444 (
<typeStatus id="367504E3DA1C783221EC861FAD07F841" box="[207,313,1915,1941]" pageId="2" pageNumber="345" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="2ECEC1C2DA1C78322046861FAD9CF840" authorityName="Kunte &amp; Basu" authorityYear="2024" box="[357,418,1915,1940]" class="Insecta" family="Nymphalidae" genus="Athyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="345" phylum="Arthropoda" rank="species" species="inara">
<emphasis id="DBBA6653DA1C78322046861FAD9CF840" box="[357,418,1915,1940]" italics="true" pageId="2" pageNumber="345">inara</emphasis>
</taxonomicName>
. Silhet,
<collectingCountry id="91D9FAD1DA1C78322325861FAEB2F841" box="[518,652,1915,1941]" name="Bangladesh" pageId="2" pageNumber="345">Bangladesh</collectingCountry>
.
<figureCitation id="71F5A6C4DA1C783223B8861FAF33F841" box="[667,781,1915,1941]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="16.[151,250,1981,2005]" captionStartId-1="17.[151,250,1191,1215]" captionTargetBox-0="[168,1415,786,1947]" captionTargetBox-1="[156,1434,187,1159]" captionTargetId-0="figure-368@16.[164,1423,777,1958]" captionTargetId-1="figure-19@17.[151,1436,181,1167]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 9. Comparison of Athyma inara sahyadriensis ssp. nov. with other similar Athyma (see Materials and Methods for specimen details). A millimetre scale is at the bottom. Photographs: Krushnamegh Kunte." captionText-1="FIGURE 10. Diagnostic characters of Athyma inara sahyadriensis ssp. nov., marked on digitally enhanced images from which wing-wear has been removed to illustrate the distinctive phenotype of the subspecies. Male of A. i. sahyadriensis ssp. nov. is compared with males of A. i. inara and two subspecies of A. nefte. Numbers point to distinctive characters: 1: FW subapical markings, 2: FW submarginal markings, 3: FW spot in interspace 3, 4: FW markings in the discal cell, 5: HW post-discal band, 6: UNF markings in the discal cell, 7: UN base colour of the wings, 8: UNF submarginal markings, 9: large white spot in interspace 2, 10: UNH post-discal band (further details in the text). Images are not to scale. Photographs: Krushnamegh Kunte." figureDoi-0="http://doi.org/10.5281/zenodo.14386322" figureDoi-1="http://doi.org/10.5281/zenodo.14386326" httpUri-0="https://zenodo.org/record/14386322/files/figure.png" httpUri-1="https://zenodo.org/record/14386326/files/figure.png" pageId="2" pageNumber="345">Fig. 910</figureCitation>
), BMNH(E) #933429 (
<typeStatus id="367504E3DA1C78322538861FA8BBF841" box="[1051,1157,1915,1941]" pageId="2" pageNumber="345" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="2ECEC1C2DA1C78322592861FA93DF840" baseAuthorityName="Evans" baseAuthorityYear="1932" box="[1201,1283,1915,1940]" class="Insecta" family="Nymphalidae" genus="Athyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="345" phylum="Arthropoda" rank="species" species="inarina">
<emphasis id="DBBA6653DA1C78322592861FA93DF840" box="[1201,1283,1915,1940]" italics="true" pageId="2" pageNumber="345">inarina</emphasis>
</taxonomicName>
. “
<collectingRegion id="2B0A74A3DA1C7832243C861FA94EF841" box="[1311,1392,1915,1941]" country="India" name="Assam" pageId="2" pageNumber="345">Assam</collectingRegion>
” as per a specimen label; border of
<collectingRegion id="2B0A74A3DA1C7832230E86FBAE47F86D" box="[557,633,1951,1977]" country="India" name="Assam" pageId="2" pageNumber="345">Assam</collectingRegion>
and
<collectingRegion id="2B0A74A3DA1C7832239186FBAF2AF86D" box="[690,788,1951,1977]" country="India" name="Manipur" pageId="2" pageNumber="345">Manipur</collectingRegion>
, NE
<collectingCountry id="91D9FAD1DA1C7832226E86FBAFB7F86D" box="[845,905,1951,1977]" name="India" pageId="2" pageNumber="345">India</collectingCountry>
, as per original description.
<figureCitation id="71F5A6C4DA1C783225E986FBA930F86D" box="[1226,1294,1951,1977]" captionStart="FIGURE 9" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[168,1415,786,1947]" captionTargetId="figure-368@16.[164,1423,777,1958]" captionTargetPageId="16" captionText="FIGURE 9. Comparison of Athyma inara sahyadriensis ssp. nov. with other similar Athyma (see Materials and Methods for specimen details). A millimetre scale is at the bottom. Photographs: Krushnamegh Kunte." figureDoi="http://doi.org/10.5281/zenodo.14386322" httpUri="https://zenodo.org/record/14386322/files/figure.png" pageId="2" pageNumber="345">Fig. 9</figureCitation>
), BMNH(E) #933424 (
<typeStatus id="367504E3DA1C7832201486A7AD9FF809" box="[311,417,1987,2013]" pageId="2" pageNumber="345" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="2ECEC1C2DA1C783220E686A7ADC1F808" authorityName="Swinhoe" authorityYear="1917" box="[453,511,1987,2012]" class="Insecta" family="Nymphalidae" genus="Pantoporia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="345" phylum="Arthropoda" rank="species" species="mera">
<emphasis id="DBBA6653DA1C783220E686A7ADC1F808" box="[453,511,1987,2012]" italics="true" pageId="2" pageNumber="345">mera</emphasis>
</taxonomicName>
. “Borneo”, in error, collected almost certainly from either NE
<collectingCountry id="91D9FAD1DA1C7832259486A7A8CFF809" box="[1207,1265,1987,2013]" name="India" pageId="2" pageNumber="345">India</collectingCountry>
or N
<collectingCountry id="91D9FAD1DA1C7832240C86A7A9A2F809" box="[1327,1436,1987,2013]" name="Myanmar" pageId="2" pageNumber="345">Myanmar</collectingCountry>
based on the phenotype.
<figureCitation id="71F5A6C4DA1D783320D081F3AE02FF65" box="[499,572,151,177]" captionStart="FIGURE 9" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[168,1415,786,1947]" captionTargetId="figure-368@16.[164,1423,777,1958]" captionTargetPageId="16" captionText="FIGURE 9. Comparison of Athyma inara sahyadriensis ssp. nov. with other similar Athyma (see Materials and Methods for specimen details). A millimetre scale is at the bottom. Photographs: Krushnamegh Kunte." figureDoi="http://doi.org/10.5281/zenodo.14386322" httpUri="https://zenodo.org/record/14386322/files/figure.png" pageId="3" pageNumber="346">Fig. 9</figureCitation>
), BMNH(E) #933502 (Darjeeling,
<collectingRegion id="2B0A74A3DA1D783322FF81F3A84CFF65" box="[988,1138,151,177]" country="India" name="West Bengal" pageId="3" pageNumber="346">West Bengal</collectingRegion>
,
<collectingCountry id="91D9FAD1DA1D783325A081F3A8FEFF65" box="[1155,1216,151,177]" name="India" pageId="3" pageNumber="346">India</collectingCountry>
. 1894), BMNH(E) #933423 (
<collectingRegion id="2B0A74A3DA1D7833201581DFADB2FF01" box="[310,396,187,213]" country="India" name="Sikkim" pageId="3" pageNumber="346">Sikkim</collectingRegion>
. 1893), BMNH(E) #933496 (
<collectingRegion id="2B0A74A3DA1D783323ED81DFAF1AFF01" box="[718,804,187,213]" country="India" name="Sikkim" pageId="3" pageNumber="346">Sikkim</collectingRegion>
. 1893), BMNH(E) #933472 (Kurseong, near Darjeeling), BMNH(E) #933481 (Darjeeling. 1894).
<emphasis id="DBBA6653DA1D7833238B8184AEF8FF28" bold="true" box="[680,710,224,252]" pageId="3" pageNumber="346">
<specimenCount id="FFC871C8DA1D7833238B8184AE81FF28" box="[680,703,224,252]" count="1" pageId="3" pageNumber="346" type="female"></specimenCount>
:
</emphasis>
BMNH(E) #933428 (Darjeeling. 1894), BMNH(E) #933461 (Darjeeling. 1909), BMNH(E) #933418 (
<collectingRegion id="2B0A74A3DA1D7833238E8067AF39FEC9" box="[685,775,259,285]" country="India" name="Sikkim" pageId="3" pageNumber="346">Sikkim</collectingRegion>
, Singlah, likely from Darjeeling/ Singalila area. 1894), BMNH(E) #933425 (
<collectingRegion id="2B0A74A3DA1D7833209D8043AE26FE95" box="[446,536,295,321]" country="India" name="Sikkim" pageId="3" pageNumber="346">Sikkim</collectingRegion>
, Teesta. 1927), BMNH(E) #933400 (Silhet,
<collectingCountry id="91D9FAD1DA1D783325398043A89EFE95" box="[1050,1184,295,321]" name="Bangladesh" pageId="3" pageNumber="346">Bangladesh</collectingCountry>
. 1909). All specimens in NHMUK
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>