291 lines
32 KiB
XML
291 lines
32 KiB
XML
<document id="B929344A755D0CD26E032B5425943B42" ID-DOI="10.5252/adansonia2018v40a12" ID-ISSN="1639-4798" ID-Zenodo-Dep="4602024" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615574952122" checkinUser="carolina" docAuthor="Castillon, Jean-Philippe" docDate="2018" docId="614E97046767D02FFC32FDE6FB553EDC" docLanguage="fr" docName="Adansonia.40.12.163-170.pdf" docOrigin="Adansonia (3) 40 (12)" docStyle="DocumentStyle:2FE19AC740987067EC2A6C89A85F9648.8:Adansonia.2018-.journal_article" docStyleId="2FE19AC740987067EC2A6C89A85F9648" docStyleName="Adansonia.2018-.journal_article" docStyleVersion="8" docTitle="Euphorbia valeryae Castillon, 2018, sp. nov." docType="treatment" docVersion="3" lastPageNumber="168" masterDocId="9D77EF7C6762D028FFB1FFB3FF9F3B31" masterDocTitle="Deux nouvelles espèces d’euphorbes (Euphorbia L. section Goniostema, Euphorbiaceae) de la région de Fort-Dauphin, Madagascar" masterLastPageNumber="170" masterPageNumber="163" pageNumber="166" updateTime="1698885597061" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="829AD06D4B1C3D0EAB0DFC3723ABC695">Deux nouvelles espèces d’euphorbes (Euphorbia L. section Goniostema, Euphorbiaceae) de la région de Fort-Dauphin, Madagascar</mods:title>
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<mods:namePart id="A07FE4CF70A9A7FB27E3A38E770FC902">Castillon, Jean-Philippe</mods:namePart>
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<mods:title id="73CFE36DFBC251B34699CDCD50D9491D">Adansonia</mods:title>
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<mods:part id="A1D72DD370A7673083C37D7A137BDD7C">
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<mods:date id="04752DD6FE4FA810174C198C53C94AB5">2018</mods:date>
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<mods:title id="8D1F224B8E92C0B549F347B34941F79B">3</mods:title>
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<mods:number id="B990E6E3F136E08FFD4FB167863E8969">2018-10-22</mods:number>
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<mods:number id="05996DBF3FB66335B801C426E7FC43E4">12</mods:number>
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<mods:identifier id="39206CA60D8C8BF7B4A1F6DE3227BC87" type="DOI">10.5252/adansonia2018v40a12</mods:identifier>
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<mods:identifier id="BA4DF84377AFCD331DC9CB9BD66D56E3" type="ISSN">1639-4798</mods:identifier>
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<treatment id="614E97046767D02FFC32FDE6FB553EDC" LSID="urn:lsid:plazi:treatment:614E97046767D02FFC32FDE6FB553EDC" httpUri="http://treatment.plazi.org/id/614E97046767D02FFC32FDE6FB553EDC" lastPageId="7" lastPageNumber="168" pageId="5" pageNumber="166">
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<subSubSection id="A1FD75996767D02DFC32FDE6FAC63941" box="[899,1369,597,624]" pageId="5" pageNumber="166" type="nomenclature">
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<paragraph id="E95826126767D02DFC32FDE6FAC63941" blockId="5.[899,1369,597,656]" box="[899,1369,597,624]" pageId="5" pageNumber="166">
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<heading id="B210917E6767D02DFC32FDE6FAC63941" box="[899,1369,597,624]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="166" reason="6">
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<taxonomicName id="2EE75D916767D02DFC32FDE6FBC93941" ID-CoL="8Y3Q6" authorityName="J.-P.Castillon" authorityYear="2018" box="[899,1110,597,624]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="166" phylum="Tracheophyta" rank="species" species="valeryae">
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<emphasis id="DB93FA006767D02DFC32FDE6FBC93941" bold="true" box="[899,1110,597,624]" italics="true" pageId="5" pageNumber="166">Euphorbia valeryae</emphasis>
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</taxonomicName>
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J.-P. Castillon,
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<taxonomicNameLabel id="C0A0477B6767D02DFAB5FDE5FAC63941" box="[1284,1369,598,624]" pageId="5" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A1FD75996767D02FFBF7FDC5FA313A00" lastPageId="7" lastPageNumber="168" pageId="5" pageNumber="166" type="description">
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<paragraph id="E95826126767D02DFBF7FDC5FB0939A1" blockId="5.[899,1369,597,656]" box="[1094,1174,630,656]" pageId="5" pageNumber="166">
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(
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<figureCitation id="71DC3A976767D02DFBFFFDC5FB1239A1" box="[1102,1165,630,656]" captionStart="FIG" captionStartId="8.[132,143,1985,2003]" captionTargetBox="[196,1391,215,1948]" captionTargetId="figure-31@8.[747,1300,763,1314]" captionTargetPageId="8" captionText="FIG. 3. — Euphorbia valeryae J.-P. Castillon, sp. nov.: A, port général; B, tige, feuilles et cyathes; C, cyathe à fleur femelle réceptive; D, cyathe à fleurs mâles mûres; E, fruit." figureDoi="http://doi.org/10.5281/zenodo.4602030" httpUri="https://zenodo.org/record/4602030/files/figure.png" pageId="5" pageNumber="166">Fig. 3</figureCitation>
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)
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</paragraph>
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<paragraph id="E95826126767D02DFC9CFD1CFC46387A" blockId="5.[811,1457,687,1044]" pageId="5" pageNumber="166">
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<emphasis id="DB93FA006767D02DFC9CFD1CFCC239F7" box="[813,861,687,710]" italics="true" pageId="5" pageNumber="166">Sicut</emphasis>
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<taxonomicName id="2EE75D916767D02DFCD4FD1CFC5039F6" authorityName="Rauh" authorityYear="1994" box="[869,975,687,711]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="166" phylum="Tracheophyta" rank="species" species="geroldii">E. geroldii</taxonomicName>
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E. tardieuanaque,
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<emphasis id="DB93FA006767D02DFB23FD1CFB2339F6" box="[1170,1212,687,711]" italics="true" pageId="5" pageNumber="166">haec</emphasis>
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<taxonomicName id="2EE75D916767D02DFB77FD1CFAAF39F6" box="[1222,1328,687,711]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="166" phylum="Tracheophyta" rank="genus">Euphorbia</taxonomicName>
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<emphasis id="DB93FA006767D02DFA87FD1CFC223801" italics="true" pageId="5" pageNumber="166">
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non spinosus dumus est sed sequentibus characteribus praecipue differt: 1) caulibus minus latis intermiscentibusque, saepe recumbentibus et radices agentibus, foliorum basi 3-5 filamenta ferentibus super
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<quantity id="2E1F8BF76767D02DFAF1FD4DFA0D3827" box="[1344,1426,766,790]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="166" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">5-10 cm</quantity>
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ad caulium apices.
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</emphasis>
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2
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<emphasis id="DB93FA006767D02DFC7AFCAAFC46387A" italics="true" pageId="5" pageNumber="166">) Incyathescentiis 1-3 terminalibus, quisque cum uno albo-viride cyatho.</emphasis>
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</paragraph>
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<paragraph id="E95826126767D02DFC9CFCD0FA1B3881" blockId="5.[811,1457,687,1044]" pageId="5" pageNumber="166">
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<materialsCitation id="598F2C4F6767D02DFC9CFCD0FBA23881" collectingDate="2017-08" collectionCode="TYPUS" collectorName="de Fort-Dauphin" country="Madagascar" elevation="300" latitude="-25.066668" location="District de Fort-Dauphin" longLatPrecision="1248" longitude="46.6" pageId="5" pageNumber="166" specimenCount="1" typeStatus="holotype">
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<typeStatus id="365C98B06767D02DFC9CFCD0FCF63848" box="[813,873,867,890]" pageId="5" pageNumber="166">TYPUS</typeStatus>
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. —
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<emphasis id="DB93FA006767D02DFC20FCD0FB99384A" bold="true" box="[913,1030,867,891]" pageId="5" pageNumber="166">
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<collectingCountry id="91F066826767D02DFC20FCD0FB99384A" box="[913,1030,867,891]" name="Madagascar" pageId="5" pageNumber="166">Madagascar</collectingCountry>
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</emphasis>
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.
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<location id="EC3870C96767D02DFBA3FCD0FA98384A" LSID="urn:lsid:plazi:treatment:614E97046767D02FFC32FDE6FB553EDC:EC3870C96767D02DFBA3FCD0FA98384A" box="[1042,1287,867,891]" country="Madagascar" latitude="-25.066668" longLatPrecision="1248" longitude="46.6" name="District de Fort-Dauphin" pageId="5" pageNumber="166">
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District
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<collectorName id="441243C46767D02DFBD0FCD0FA98384A" box="[1121,1287,867,891]" pageId="5" pageNumber="166">de Fort-Dauphin</collectorName>
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</location>
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, montagne granitique vers Ranomainty, dans des lambeaux de forêt,
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<quantity id="2E1F8BF76767D02DFAA5FCCEFACB38A4" box="[1300,1364,893,917]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="166" unit="m" value="300.0">
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<elevation id="62CAC1216767D02DFAA5FCCEFACB38A4" box="[1300,1364,893,917]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="166" unit="m" value="300.0">300 m</elevation>
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</quantity>
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,
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<geoCoordinate id="8CD340D56767D02DFAECFCCEFA3438A4" box="[1373,1451,893,917]" degrees="25" direction="south" minutes="04" orientation="latitude" pageId="5" pageNumber="166" precision="925" value="-25.066668">25°04’S</geoCoordinate>
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,
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<geoCoordinate id="8CD340D56767D02DFC9AFC24FCE5389E" box="[811,890,919,943]" degrees="46" direction="east" minutes="36" orientation="longitude" pageId="5" pageNumber="166" precision="925" value="46.6">46°36’E</geoCoordinate>
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environ,
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<date id="9D5900D26767D02DFC66FC2BFBA23881" box="[983,1085,920,944]" pageId="5" pageNumber="166" value="2017-08">
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<collectingDate id="8D1DF93A6767D02DFC66FC2BFBA23881" box="[983,1085,920,944]" pageId="5" pageNumber="166" value="2017-08">VIII.2017</collectingDate>
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</date>
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,
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<emphasis id="DB93FA006767D02DFBF8FC2BFB70389E" box="[1097,1263,919,944]" italics="true" pageId="5" pageNumber="166">J.-B. Castillon 65</emphasis>
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||
(holo-, TAN).
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||
</paragraph>
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||
<paragraph id="E95826126767D02DFC9CFC7AFA853F25" blockId="5.[811,1457,687,1044]" pageId="5" pageNumber="166">ÉTYMOLOGIE. — Cette espèce est dédiée à Mme Valery pour ses contributions, discrètes mais nombreuses, à la connaissance de la flore malgache, dont la découverte de cette espèce.</paragraph>
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||
<paragraph id="E95826126767D02DFC9CFB86FC223F7C" blockId="5.[813,1455,1077,1199]" box="[813,957,1077,1103]" pageId="5" pageNumber="166">DESCRIPTION</paragraph>
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<paragraph id="E95826126767D02DFC9CFBE6FC583F5F" blockId="5.[813,1455,1077,1199]" box="[813,967,1108,1134]" pageId="5" pageNumber="166">
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||
<emphasis id="DB93FA006767D02DFC9CFBE6FC583F5F" box="[813,967,1108,1134]" italics="true" pageId="5" pageNumber="166">Forme générale</emphasis>
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</paragraph>
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||
<paragraph id="E95826126767D02DFC9CFBC7FC683F9F" blockId="5.[813,1455,1077,1199]" pageId="5" pageNumber="166">
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||
Arbrisseau ou sous-arbrisseau non épineux, très ramifié, de
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||
<quantity id="2E1F8BF76767D02DFC9FFB27FC083F9F" box="[814,919,1172,1199]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="5" pageNumber="166" unit="cm" value="65.0" valueMax="80.0" valueMin="50.0">50-80 cm</quantity>
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||
de haut.
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||
</paragraph>
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||
<paragraph id="E95826126767D02DFC9CFB67FCC53FDC" blockId="5.[811,1457,1236,1613]" box="[813,858,1236,1261]" pageId="5" pageNumber="166">
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<emphasis id="DB93FA006767D02DFC9CFB67FCC53FDC" box="[813,858,1236,1261]" italics="true" pageId="5" pageNumber="166">Tige</emphasis>
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||
</paragraph>
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<paragraph id="E95826126767D02DFC9CFB40FB563D7D" blockId="5.[811,1457,1236,1613]" pageId="5" pageNumber="166">
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Tige principale de
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<quantity id="2E1F8BF76767D02DFC42FB47FBB73E3F" box="[1011,1064,1268,1294]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="166" unit="cm" value="1.0">1 cm</quantity>
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||
de diamètre à la base, à écorce relativement lisse, vert-marron tachetée de blanc.Tiges secondaires 10-50 ×
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||
<quantity id="2E1F8BF76767D02DFC21FA80FB903E7F" box="[912,1039,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="166" unit="cm" value="0.4" valueMax="0.5" valueMin="0.3">0,3-0,5 cm</quantity>
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||
, tortueuses, parfois entremêlées, sans direction particulière, parfois couchées ou retombant sur le sol où elles s’enracinent à nouveau, se terminant généralement en 3-4 ramifications ultimes de
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||
<quantity id="2E1F8BF76767D02DFBD0FA20FB5F3E9F" box="[1121,1216,1427,1454]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="5" pageNumber="166" unit="cm" value="8.0" valueMax="10.0" valueMin="6.0">6-10 cm</quantity>
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||
. Les 5-10 derniers cm des tiges présentent des boursouflures formées des cicatrices rondes ou réniformes 3 ×
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||
<quantity id="2E1F8BF76767D02DFB8CFA60FBE23EDC" box="[1085,1149,1491,1517]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="166" unit="mm" value="2.0">2 mm</quantity>
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||
des feuilles récemment tombées, entourées par des groupes de 5-10 stipules. L’extrémité des tiges porte une rosette de 2-5 feuilles, de nombreuses stipules ainsi que 2-3 incyathescences.
|
||
</paragraph>
|
||
<paragraph id="E95826126767D02DFC9CF9C1FCE63DBD" blockId="5.[812,1457,1650,2028]" box="[813,889,1650,1676]" pageId="5" pageNumber="166">
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<emphasis id="DB93FA006767D02DFC9CF9C1FCE63DBD" box="[813,889,1650,1676]" italics="true" pageId="5" pageNumber="166">Feuilles</emphasis>
|
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</paragraph>
|
||
<paragraph id="E95826126767D02DFC9CF921FB523CDA" blockId="5.[812,1457,1650,2028]" pageId="5" pageNumber="166">
|
||
Soit caulinaires et rapidement caduques, soit persistantes et groupées par 2-5 en rosettes terminales, de forme ovale à rhomboïdale arrondie, 90 ×
|
||
<quantity id="2E1F8BF76767D02DFBCFF961FB503DDD" box="[1150,1231,1746,1773]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="166" unit="mm" value="45.0">45 mm</quantity>
|
||
au maximum, 50 ×
|
||
<quantity id="2E1F8BF76767D02DFC9CF941FC1F3C3D" box="[813,896,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="166" unit="mm" value="25.0">25 mm</quantity>
|
||
en moyenne, à extrémité généralement arrondie, avec un mucron de
|
||
<quantity id="2E1F8BF76767D02DFBBFF8A1FBCE3C1D" box="[1038,1105,1810,1836]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="166" unit="mm" value="1.0">1 mm</quantity>
|
||
; limbe lisse, vert foncé brillant sur le dessus, plus clair en dessous; marge continue, parfois légèrement ondulée; pétiole 5-10 ×
|
||
<quantity id="2E1F8BF76767D02DFB08F8E1FB603C5D" box="[1209,1279,1874,1900]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="166" unit="mm" value="2.0">2 mm</quantity>
|
||
, canaliculé, vert foncé au-dessus, blanchâtre au-dessous; nervation pennée, une nervure centrale blanchâtre en relief sous la feuille, 8-10 nervures secondaires généralement peu distinctes, parfois blanchâtres et alors nettement visibles.
|
||
</paragraph>
|
||
<caption id="BD98769A6764D02EFED9F864FB523CD8" ID-DOI="http://doi.org/10.5281/zenodo.4602028" ID-Zenodo-Dep="4602028" box="[360,1229,2007,2025]" httpUri="https://zenodo.org/record/4602028/files/figure.png" pageId="6" pageNumber="167" startId="6.[360,371,2007,2025]" targetBox="[235,1352,215,1954]" targetPageId="6">
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<paragraph id="E95826126764D02EFED9F864FB523CD8" blockId="6.[360,1229,2007,2025]" box="[360,1229,2007,2025]" pageId="6" pageNumber="167">
|
||
FIG. 2. — Paratype de
|
||
<taxonomicName id="2EE75D916764D02EFD93F864FD5C3CD8" authorityName="J.-P.Castillon" authorityYear="2018" box="[546,707,2007,2025]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="167" phylum="Tracheophyta" rank="species" species="gigantea">
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||
<emphasis id="DB93FA006764D02EFD93F864FD5C3CD8" box="[546,707,2007,2025]" italics="true" pageId="6" pageNumber="167">Euphorbia gigantea</emphasis>
|
||
</taxonomicName>
|
||
J.-P. Castillon,
|
||
<taxonomicNameLabel id="C0A0477B6764D02EFCF4F864FC153CD8" box="[837,906,2007,2025]" pageId="6" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, échantillon
|
||
<emphasis id="DB93FA006764D02EFC5EF864FBC73CD8" box="[1007,1112,2007,2025]" italics="true" pageId="6" pageNumber="167">Decary 9432</emphasis>
|
||
(P00217865).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E95826126765D02FFF35FF64FF4C3BC0" blockId="7.[132,777,215,369]" box="[132,211,215,241]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FF64FF4C3BC0" box="[132,211,215,241]" italics="true" pageId="7" pageNumber="168">Stipules</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FF44FDBD3A40" blockId="7.[132,777,215,369]" pageId="7" pageNumber="168">
|
||
Piliformes souples, présentes uniquement sur les derniers cm des tiges, par groupes de 5-10 autour des cicatrices des feuilles, plus nombreuses à l’extrémité des tiges, rapidement caduques, 1-3 ×
|
||
<quantity id="2E1F8BF76765D02FFE86FEE4FE0F3A40" box="[311,400,343,369]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="168" unit="mm" value="0.1">0,1 mm</quantity>
|
||
, gris-marron.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FE24FEE83A81" blockId="7.[132,595,406,465]" box="[132,375,406,433]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FE24FEE83A81" box="[132,375,406,433]" italics="true" pageId="7" pageNumber="168">Système incyathescentiel</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FE05FDCC3AE1" blockId="7.[132,595,406,465]" box="[132,595,438,465]" pageId="7" pageNumber="168">Subterminal formé de 1-3 incyathescences.</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FE45FE893921" blockId="7.[130,775,502,593]" box="[132,278,502,528]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FE45FE893921" box="[132,278,502,528]" italics="true" pageId="7" pageNumber="168">Incyathescence</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FDA4FC993960" blockId="7.[130,775,502,593]" pageId="7" pageNumber="168">Portant un unique cyathium bisexué (très exceptionnellement les boutons axillaires de ce cyathium peuvent se développer).</paragraph>
|
||
<paragraph id="E95826126765D02FFF32FDC5FF7539A1" blockId="7.[131,775,630,720]" box="[131,234,630,656]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF32FDC5FF7539A1" box="[131,234,630,656]" italics="true" pageId="7" pageNumber="168">Pédoncule</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FD25FEAB39FE" blockId="7.[131,775,630,720]" pageId="7" pageNumber="168">
|
||
Collant, vert, 12-18 ×
|
||
<quantity id="2E1F8BF76765D02FFEC9FD26FE4E3981" box="[376,465,661,688]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="168" unit="mm" value="0.5">0,5 mm</quantity>
|
||
, recourbé, se prolongeant en 2 cyathophylles.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FD46FE8F383E" blockId="7.[131,777,757,880]" box="[132,272,757,783]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FD46FE8F383E" box="[132,272,757,783]" italics="true" pageId="7" pageNumber="168">Cyathophylles</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FCA4FDC2385E" blockId="7.[131,777,757,880]" pageId="7" pageNumber="168">
|
||
Étalés, environ 7 ×
|
||
<quantity id="2E1F8BF76765D02FFED0FCA5FE363801" box="[353,425,790,816]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="168" unit="mm" value="7.0">7 mm</quantity>
|
||
, verts au niveau du pédoncule, blancs avec des traces jaunes ou vertes sur le dessus, rosissant en séchant, à extrémité légèrement apiculée.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FC26FF7C389E" blockId="7.[132,348,917,975]" box="[132,227,917,943]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FC26FF7C389E" box="[132,227,917,943]" italics="true" pageId="7" pageNumber="168">Involucre</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FC06FEC338FE" blockId="7.[132,348,917,975]" box="[132,348,949,975]" pageId="7" pageNumber="168">
|
||
Conique 3 ×
|
||
<quantity id="2E1F8BF76765D02FFEA5FC06FEC638FE" box="[276,345,949,975]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="168" unit="mm" value="3.0">3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FC47FF483F3F" blockId="7.[132,775,1012,1134]" box="[132,215,1012,1038]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FC47FF483F3F" box="[132,215,1012,1038]" italics="true" pageId="7" pageNumber="168">Glandes</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FBA7FE223F5F" blockId="7.[132,775,1012,1134]" pageId="7" pageNumber="168">
|
||
Cinq glandes vertes puis jaunes à l’anthèse, elliptiques-réniformes, 2 ×
|
||
<quantity id="2E1F8BF76765D02FFEB4FB87FED63F7E" box="[261,329,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="168" unit="mm" value="1.0">1 mm</quantity>
|
||
, contiguës, étalées mais avec la lèvre extérieure légèrement plus haute.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FB27FE153F9F" blockId="7.[132,775,1172,1262]" box="[132,394,1172,1198]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FB27FE153F9F" box="[132,394,1172,1198]" italics="true" pageId="7" pageNumber="168">Bractées interglandulaires</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FB07FE343FDF" blockId="7.[132,775,1172,1262]" pageId="7" pageNumber="168">Vertes, à extrémités divisées en 5-7 languettes blanches translucides, recouvrant l’ovaire.</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FAA0FE883E1C" blockId="7.[130,776,1299,1454]" box="[132,279,1299,1325]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FAA0FE883E1C" box="[132,279,1299,1325]" italics="true" pageId="7" pageNumber="168">Fleurs femelles</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FA80FC993E9C" blockId="7.[130,776,1299,1454]" pageId="7" pageNumber="168">
|
||
Ovaire à 3loges, blanc, 2 ×
|
||
<quantity id="2E1F8BF76765D02FFE2DFA80FE6E3E7F" box="[412,497,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="168" unit="mm" value="1.5">1,5 mm</quantity>
|
||
, glabre; 3 styles blanc-vert,
|
||
<quantity id="2E1F8BF76765D02FFF33FAE0FF5B3E5F" box="[130,196,1363,1390]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="168" unit="mm" value="2.0">2 mm</quantity>
|
||
, arqués-ascendants, non soudés, divisés à l’extrémité en 2 lobes recourbés. Stigmates sphériques, blancs, translucides. Chez les cyathes âgés, les 3 styles se ressoudent en une colonne.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FA60FE9D3EDC" blockId="7.[132,775,1491,1581]" box="[132,258,1491,1517]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35FA60FE9D3EDC" box="[132,258,1491,1517]" italics="true" pageId="7" pageNumber="168">Fleurs mâles</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35FA40FE543D1D" blockId="7.[132,775,1491,1581]" pageId="7" pageNumber="168">
|
||
Environ 10-12;filet et pédicelle de
|
||
<quantity id="2E1F8BF76765D02FFDB1FA41FDD03D3C" box="[512,591,1522,1549]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="168" unit="mm" value="1.5">1,5mm</quantity>
|
||
environ, blancs; 2 anthères jaunes par étamine.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35F9E0FF5E3D5D" blockId="7.[132,661,1619,1677]" box="[132,193,1619,1644]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35F9E0FF5E3D5D" box="[132,193,1619,1644]" italics="true" pageId="7" pageNumber="168">Fruits</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35F9C1FD0A3DBD" blockId="7.[132,661,1619,1677]" box="[132,661,1650,1677]" pageId="7" pageNumber="168">
|
||
Blancs, triloculaires, 4 ×
|
||
<quantity id="2E1F8BF76765D02FFE3FF9C1FE4D3DBC" box="[398,466,1650,1677]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="168" unit="mm" value="3.0">3 mm</quantity>
|
||
, à style persistant.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35F901FF4C3DFD" blockId="7.[132,297,1714,1772]" box="[132,211,1714,1740]" pageId="7" pageNumber="168">
|
||
<emphasis id="DB93FA006765D02FFF35F901FF4C3DFD" box="[132,211,1714,1740]" italics="true" pageId="7" pageNumber="168">Graines</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFF35F961FEB63DDD" blockId="7.[132,297,1714,1772]" box="[132,297,1746,1772]" pageId="7" pageNumber="168">Non observées.</paragraph>
|
||
<paragraph id="E95826126765D02FFF35F8A1FEC93C1D" blockId="7.[131,776,1810,2028]" box="[132,342,1810,1836]" pageId="7" pageNumber="168">NOTE ÉCOLOGIQUE</paragraph>
|
||
<paragraph id="E95826126765D02FFF35F881FA313A00" blockId="7.[131,776,1810,2028]" lastBlockId="7.[811,1455,215,306]" pageId="7" pageNumber="168">
|
||
Cette plante pousse sur le flanc d’un inselberg granitique, dans une forêt semi-humide formée d’arbres feuillus, et non dans le bush à
|
||
<taxonomicName id="2EE75D916765D02FFED2F8C2FE523CBA" box="[355,461,1905,1931]" class="Insecta" family="Cerambycidae" genus="Alluaudia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DB93FA006765D02FFED2F8C2FE523CBA" box="[355,461,1905,1931]" italics="true" pageId="7" pageNumber="168">Alluaudia</emphasis>
|
||
</taxonomicName>
|
||
, sous une canopée de
|
||
<quantity id="2E1F8BF76765D02FFD7DF8C2FC983CBD" box="[716,775,1905,1932]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="168" unit="m" value="10.0">10 m</quantity>
|
||
environ, dans des sols latéritiques recouverts d’humus. Elle ne voit quasiment jamais le soleil. Elle doit évidemment se multiplier par graines, mais son mode de reproduction prin- cipal semble être plutôt végétatif: ses tiges minces et souples courbent sous leur propre poids et touchent le sol où elles s’enracinent pour donner naissance à de nouveaux plants.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1FD75996765D02FFC9CFEE4FB553EDC" pageId="7" pageNumber="168" type="discussion">
|
||
<paragraph id="E95826126765D02FFC9CFEE4FC2C3A5E" blockId="7.[811,1457,343,816]" box="[813,947,343,369]" pageId="7" pageNumber="168">DISCUSSION</paragraph>
|
||
<paragraph id="E95826126765D02FFC9CFEC4FAAD3801" blockId="7.[811,1457,343,816]" pageId="7" pageNumber="168">
|
||
Les affinités de cette espèce sont peu évidentes: son absence d’épines la rapproche des autres euphorbes non épineuses de la section
|
||
<emphasis id="DB93FA006765D02FFC0DFE04FBA63AE0" box="[956,1081,439,465]" italics="true" pageId="7" pageNumber="168">Goniostema</emphasis>
|
||
en provenance de la côte est malgache (
|
||
<taxonomicName id="2EE75D916765D02FFCCEFE64FBA53AC1" authority="Rauh" authorityName="Rauh" authorityYear="1994" box="[895,1082,470,496]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="168" phylum="Tracheophyta" rank="species" species="geroldii">
|
||
<emphasis id="DB93FA006765D02FFCCEFE64FC6E3AC1" box="[895,1009,470,496]" italics="true" pageId="7" pageNumber="168">E. geroldii</emphasis>
|
||
Rauh
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2EE75D916765D02FFBFAFE64FAAC3AC1" authority="Baill." authorityName="Baill." box="[1099,1331,470,496]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="168" phylum="Tracheophyta" rank="species" species="thouarsiana">
|
||
<emphasis id="DB93FA006765D02FFBFAFE64FB753AC1" box="[1099,1258,470,496]" italics="true" pageId="7" pageNumber="168">E. thouarsiana</emphasis>
|
||
Baill.
|
||
</taxonomicName>
|
||
, etc.) mais la simple absence d’épines, qui semble fortement corrélée à l’habitat en zones humides, ne me semble pas être un caractère de forte importance. Son port en petits buissons à tiges retombantes la rapproche de certaines euphorbes épineuses du sud malgache (
|
||
<taxonomicName id="2EE75D916765D02FFBD1FDC4FA3539A1" authority="Denis ex Leandri" authorityName="Leandri" box="[1120,1450,630,656]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="168" phylum="Tracheophyta" rank="species" species="beharensis">
|
||
<emphasis id="DB93FA006765D02FFBD1FDC4FB7739A1" box="[1120,1256,630,656]" italics="true" pageId="7" pageNumber="168">E. beharensis</emphasis>
|
||
Denis ex Leandri
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2EE75D916765D02FFC9AFD25FB843980" authority="Denis" authorityName="Denis" box="[811,1051,662,689]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="168" phylum="Tracheophyta" rank="species" species="mahafalensis">
|
||
<emphasis id="DB93FA006765D02FFC9AFD25FC513981" box="[811,974,662,688]" italics="true" pageId="7" pageNumber="168">E. mahafalensis</emphasis>
|
||
Denis
|
||
</taxonomicName>
|
||
), ses pédoncules collants ainsi que ses larges cyathophylles la rapprochent du groupe de
|
||
<taxonomicName id="2EE75D916765D02FFAEDFD05FC3A39DE" authority="Des Moul." authorityName="Des Moul." class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="168" phylum="Tracheophyta" rank="species" species="milii">
|
||
<emphasis id="DB93FA006765D02FFAEDFD05FA2F39FE" box="[1372,1456,693,719]" italics="true" pageId="7" pageNumber="168">E. milii</emphasis>
|
||
Des Moul.
|
||
</taxonomicName>
|
||
, tandis que ses incyathescences toujours simples sont un caractère propre à l’espèce et qu’on ne retrouve chez aucun autre membre de la section
|
||
<emphasis id="DB93FA006765D02FFB1CFCA5FAB53801" box="[1197,1322,790,816]" italics="true" pageId="7" pageNumber="168">Goniostema</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E95826126765D02FFC9CFCE5FBCD385F" blockId="7.[811,1457,854,1517]" box="[813,1106,854,880]" pageId="7" pageNumber="168">STATUT DE CONSERVATION</paragraph>
|
||
<paragraph id="E95826126765D02FFC9CFCC6FB553EDC" blockId="7.[811,1457,854,1517]" pageId="7" pageNumber="168">
|
||
Cette plante a été trouvée dans un lambeau de forêt semihumide composée d’arbres feuillus, dans une zone dominée par le bush à
|
||
<taxonomicName id="2EE75D916765D02FFC08FC06FBBC38FE" box="[953,1059,949,975]" class="Insecta" family="Cerambycidae" genus="Alluaudia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DB93FA006765D02FFC08FC06FBBC38FE" box="[953,1059,949,975]" italics="true" pageId="7" pageNumber="168">Alluaudia</emphasis>
|
||
</taxonomicName>
|
||
et où ces forêts sont rares. Ses caractéristiques morphologiques, intermédiaires entre celles des euphorbes de zones sèches et celles des euphorbes de forêts humides, en font une plante adaptée aux zones de transition, comme les forêts de l’extrémité sud de l’Andohahelo, et il est probable qu’on ne trouvera cette plante ni dans les forêts humides de l’est de l’Andohahelo, ni dans le bush de l’Androy. Or les zones de transition n’occupent qu’une surface très limitée; la zone d’occurrence de cette espèce, ainsi que sa zone d’occupation, sont donc probablement très petites, ce qui est confirmé par le fait qu’aucune autre collecte n’en ait jamais été effectuée à ce jour, malgré les nombreuses recherches entreprises dans la région. Les données actuellement disponibles sur cette espèce, seulement quelques dizaines de plants dans une forêt dégradée, sans aucune protection, et sur une surface de quelques centaines de m² m’incitent à considérer cette plante comme en danger critique d’extinction (CR B2a,b; D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |