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<document id="1073E6AEA874D8E87F0065D14A5B0CE1" ID-DOI="10.5281/zenodo.185067" ID-GBIF-Dataset="7efbee4b-7750-4ee7-a58b-5aac245f8daa" ID-ISSN="1175-5326" ID-Zenodo-Dep="185067" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459950457421" checkinUser="plazi" docAuthor="Soldán, Tomáš &amp; Godunko, Roman J." docDate="2009" docId="1E3887F1C07B4F30FF03FD7FFCCFFE49" docLanguage="en" docName="zt01972p019.pdf" docOrigin="Zootaxa 1972" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Baetis (Baetis) zdenkae Soldan and Godunko, sp. nov." docType="treatment" docVersion="10" lastPageNumber="13" masterDocId="E201FF89C07A4F3CFF94FFB8FFC0FF83" masterDocTitle="Baetis zdenkae sp. nov., a new representative of the Baetis buceratus species-group (Ephemeroptera: Baetidae) from Rhodos (Greece) with notes to species-grouping of the subgenus Baetis Leach, 1815 s. str." masterLastPageNumber="19" masterPageNumber="1" pageNumber="2" updateTime="1698230798979" updateUser="plazi">
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<mods:title id="046B91C4DEABDDEEFE3D2FFA777FC27B">Baetis zdenkae sp. nov., a new representative of the Baetis buceratus species-group (Ephemeroptera: Baetidae) from Rhodos (Greece) with notes to species-grouping of the subgenus Baetis Leach, 1815 s. str.</mods:title>
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<mods:role id="96D086F1E231CE90008A7577B3F55E9F">
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<mods:namePart id="51C723A37338CFB8C92EA761BB2D8F5E">Soldán, Tomáš</mods:namePart>
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<mods:namePart id="18490EFE05F5AFD888E2D26AA74EC7F2">Godunko, Roman J.</mods:namePart>
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<mods:title id="117906818DB5D168B9848DDFEDE8D572">Zootaxa</mods:title>
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<mods:date id="772883EFDC45592124CFE4DD6ADC3B4C">2009</mods:date>
<mods:detail id="417B059B3B45A8764357B3E4A1F389D4" type="volume">
<mods:number id="97045FE1C2883B29C9576315E10AE67C">1972</mods:number>
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<mods:identifier id="6D0E29A1F2BCC450A5384425985F99BE" type="DOI">10.5281/zenodo.185067</mods:identifier>
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<treatment id="1E3887F1C07B4F30FF03FD7FFCCFFE49" ID-DOI="http://doi.org/10.5281/zenodo.6223129" ID-GBIF-Taxon="119376452" ID-Zenodo-Dep="6223129" LSID="urn:lsid:plazi:treatment:1E3887F1C07B4F30FF03FD7FFCCFFE49" httpUri="http://treatment.plazi.org/id/1E3887F1C07B4F30FF03FD7FFCCFFE49" lastPageId="12" lastPageNumber="13" pageId="1" pageNumber="2">
<subSubSection id="DE8B656CC07B4F3DFF03FD7FFEDFFC87" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="962E36E7C07B4F3DFF03FD7FFCEDFD62" blockId="1.[151,813,711,772]" box="[151,813,711,737]" pageId="1" pageNumber="2">
<heading id="CD66818BC07B4F3DFF03FD7FFCEDFD62" bold="true" box="[151,813,711,737]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="A4E5EAF5C07B4F3DFF03FD7FFCEDFD62" bold="true" box="[151,813,711,737]" pageId="1" pageNumber="2">
<taxonomicName id="51914D64C07B4F3DFF03FD7FFD03FD62" ID-CoL="KD3S" authority="Soldan and Godunko" authorityName="Soldan and Godunko" box="[151,707,711,737]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov." subGenus="Baetis">
<emphasis id="A4E5EAF5C07B4F3DFF03FD7FFF22FD62" bold="true" box="[151,226,711,737]" italics="true" pageId="1" pageNumber="2">Baetis</emphasis>
(
<emphasis id="A4E5EAF5C07B4F3DFF67FD7FFEFEFD62" bold="true" box="[243,318,711,737]" italics="true" pageId="1" pageNumber="2">Baetis</emphasis>
)
<emphasis id="A4E5EAF5C07B4F3DFEDBFD7FFE71FD62" bold="true" box="[335,433,711,737]" italics="true" pageId="1" pageNumber="2">zdenkae</emphasis>
Soldán and Godunko
</taxonomicName>
,
<taxonomicNameLabel id="BFD6578EC07B4F3DFD44FD7FFCEDFD62" box="[720,813,711,737]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="962E36E7C07B4F3DFF03FD52FEDFFC87" blockId="1.[151,813,711,772]" box="[151,287,746,772]" pageId="1" pageNumber="2">
(
<figureCitation id="0EAA2A62C07B4F3DFF0BFD52FED5FC87" box="[159,277,746,772]" captionStart-0="FIGURES 1 4" captionStart-1="FIGURES 5 7" captionStart-2="FIGURES 8 12" captionStart-3="FIGURES 13 16" captionStart-4="FIGURE 17" captionStart-5="FIGURES 18 21" captionStart-6="FIGURES 22 25" captionStart-7="FIGURES 26 29" captionStartId-0="2.[151,269,1461,1485]" captionStartId-1="3.[151,269,1704,1728]" captionStartId-2="4.[151,269,1957,1981]" captionStartId-3="5.[151,269,1186,1210]" captionStartId-4="6.[151,255,1814,1838]" captionStartId-5="7.[151,269,1192,1216]" captionStartId-6="9.[151,270,1634,1658]" captionStartId-7="10.[151,269,1472,1496]" captionTargetBox-0="[178,1411,208,1428]" captionTargetBox-1="[173,1416,349,1667]" captionTargetBox-2="[197,1410,579,1912]" captionTargetBox-3="[151,1435,194,1162]" captionTargetBox-4="[630,979,231,1770]" captionTargetBox-5="[151,1435,197,1165]" captionTargetBox-6="[151,1435,640,1607]" captionTargetBox-7="[151,1435,477,1445]" captionTargetId-0="figure@2.[167,1420,197,1435]" captionTargetId-1="figure@3.[151,1436,342,1680]" captionTargetId-2="figure@4.[177,1410,574,1933]" captionTargetId-3="figure@5.[151,1436,194,1162]" captionTargetId-4="figure@6.[599,987,197,1787]" captionTargetId-5="figure@7.[151,1436,197,1165]" captionTargetId-6="figure@9.[151,1436,640,1608]" captionTargetId-7="figure@10.[151,1436,477,1445]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="FIGURES 1 4. Baetis zdenkae sp. nov., male imagoes, paratypes: 1, head (lateral view); 2, head (dorsal view); 3, hind right wing (dorsal view); 4, hind left wing (dorsal view)." captionText-1="FIGURES 5 7. Baetis zdenkae sp. nov., male imagoes, paratypes: 5 6, forceps (ventral view); 7, tip of abdomen (lateral view)." captionText-2="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." captionText-3="FIGURES 13 16. Baetis zdenkae sp. nov., mature larvae: 13, labrum (dorsal view); 14, apical portion of the last segment of maxillary palpus; 15, arrangement of spatulate setae on the surface of thorax; 16, outer margin of femur (hind leg). Scale bars: 13 = 500 nm; 14, 15 = 100 nm; 16 = 500 nm." captionText-4="FIGURE 17. Baetis zdenkae sp. nov., mature larva, general view of body (dorsal view)." captionText-5="FIGURES 18 21. Baetis zdenkae sp. nov., mature larvae, outer margin of femora (hind legs). Scale bars: 18 = 200 nm; 19 = 100 nm; 20, 21 = 100 nm." captionText-6="FIGURES 22 25. Baetis zdenkae sp. nov., mature larvae: 22, surface of femur; 23, outer margin of tibia; 24, inner margin of tibia; 25, tarsal claw (hind legs). Scale bars: 22 24 = 200 nm; 25 = 500 nm." captionText-7="FIGURES 26 29. Baetis zdenkae sp. nov., mature larvae: 26, surface of abdominal tergum IV; 27, surface of abdominal tergum VI; 28, posterior margin of abdominal tergum IV; 29, outer margin of paraproct plate. Scale bars: 26 29 = 20 mm." httpUri-0="https://zenodo.org/record/185068/files/figure.png" httpUri-1="https://zenodo.org/record/185069/files/figure.png" httpUri-2="https://zenodo.org/record/185070/files/figure.png" httpUri-3="https://zenodo.org/record/185071/files/figure.png" httpUri-4="https://zenodo.org/record/185072/files/figure.png" httpUri-5="https://zenodo.org/record/185073/files/figure.png" httpUri-6="https://zenodo.org/record/185074/files/figure.png" httpUri-7="https://zenodo.org/record/185075/files/figure.png" pageId="1" pageNumber="2">Figs 129</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DE8B656CC07B4F39FF03FC82FD4BF954" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2" type="description">
<paragraph id="962E36E7C07B4F3DFF03FC82FC95FC27" blockId="1.[151,1437,826,2012]" pageId="1" pageNumber="2">
<emphasis id="A4E5EAF5C07B4F3DFF03FC82FEEFFCD7" bold="true" box="[151,303,826,852]" pageId="1" pageNumber="2">Description.</emphasis>
<emphasis id="A4E5EAF5C07B4F3DFEA2FC82FE03FCD0" box="[310,451,826,851]" italics="true" pageId="1" pageNumber="2">Male imago</emphasis>
. Size: body length:
<quantity id="51699B02C07B4F3DFD39FC82FCFDFCD7" box="[685,829,826,852]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.3" metricValueMin="5.8" pageId="1" pageNumber="2" unit="mm" value="6.05" valueMax="6.3" valueMin="5.8">5.86.3 mm</quantity>
; fore wings length:
<quantity id="51699B02C07B4F3DFBB1FC82FB75FCD7" box="[1061,1205,826,852]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.2" metricValueMin="5.6" pageId="1" pageNumber="2" unit="mm" value="5.9" valueMax="6.2" valueMin="5.6">5.66.2 mm</quantity>
; cerci length:
<quantity id="51699B02C07B4F3DFACEFC82FEC5FCFF" metricMagnitude="-2" metricUnit="m" metricValue="1.22" metricValueMax="1.38" metricValueMin="1.06" pageId="1" pageNumber="2" unit="mm" value="12.2" valueMax="13.8" valueMin="10.6">10.6 13.8 mm</quantity>
; tarsal segments: T1 =
<quantity id="51699B02C07B4F3DFD8BFCDAFD0FFCFF" box="[543,719,866,892]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.199999999999999" metricValueMin="6.8" pageId="1" pageNumber="2" unit="mm" value="0.7" valueMax="0.72" valueMin="0.68">0.680.72 mm</quantity>
; T2 =
<quantity id="51699B02C07B4F3DFCB6FCDAFC12FCFF" box="[802,978,866,892]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.699999999999999" metricValueMin="5.4" pageId="1" pageNumber="2" unit="mm" value="0.5549999999999999" valueMax="0.57" valueMin="0.54">0.540.57 mm</quantity>
; T3 =
<quantity id="51699B02C07B4F3DFBB0FCDAFB14FCFF" box="[1060,1236,866,892]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="1" pageNumber="2" unit="mm" value="0.33" valueMax="0.34" valueMin="0.32">0.320.34 mm</quantity>
; T4 =
<quantity id="51699B02C07B4F3DFAB3FCDAFF08FC27" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="1" pageNumber="2" unit="mm" value="0.24" valueMax="0.25" valueMin="0.23">0.230.25 mm</quantity>
; general relative tarsal segments length ratio: 1&gt;2&gt;3&gt;4.
</paragraph>
<paragraph id="962E36E7C07B4F3DFF51FC0AFD8CFA07" blockId="1.[151,1437,826,2012]" pageId="1" pageNumber="2">
General body color light, yellowish-brown to brown. Head brown with darker smudges laterally. Antennae brown, flagellum slightly paler. Ocelli whitish-brown at the apex with brown bases. Eyes black, surrounded by a light ring, occasionally with some yellowish smudges basally. Turbinate eyes oval in dorsal view with convex external margins. Facetted surface yellowish-brown to light brown. Distal part of stalk with narrow yellow ring; central part slightly darker, dirty-yellow, basal brown ring well distinguished (
<figureCitation id="0EAA2A62C07B4F3DFA83FBEAFA4DFBEF" box="[1303,1421,1106,1132]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,1461,1485]" captionTargetBox="[178,1411,208,1428]" captionTargetId="figure@2.[167,1420,197,1435]" captionTargetPageId="2" captionText="FIGURES 1 4. Baetis zdenkae sp. nov., male imagoes, paratypes: 1, head (lateral view); 2, head (dorsal view); 3, hind right wing (dorsal view); 4, hind left wing (dorsal view)." httpUri="https://zenodo.org/record/185068/files/figure.png" pageId="1" pageNumber="2">Figs. 1, 2</figureCitation>
). Thorax darker than head, yellowish-brown to brown, with light to dark brown maculation along sutures. Pronotum and distal part of mesonotum yellowish-brown to brown, proximal part distinctly brown. Thoracic sterna light brown to brown. Wings hyaline, transparent. Fore wings with slightly opaque pterostigma.
<collectingCountry id="EE867677C07B4F3DFAF5FB72FA5CFB67" box="[1377,1436,1226,1252]" name="India" pageId="1" pageNumber="2">Hind</collectingCountry>
wings with costal process small, acute and rounded at apex with three simple longitudinal veins with two free veins in the third field, second vein occasionally with distal bifurcation, third vein relatively short, length about 0.380.40 of wing (
<figureCitation id="0EAA2A62C07B4F3DFE53FAFAFDF9FADF" box="[455,569,1346,1372]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,1461,1485]" captionTargetBox="[178,1411,208,1428]" captionTargetId="figure@2.[167,1420,197,1435]" captionTargetPageId="2" captionText="FIGURES 1 4. Baetis zdenkae sp. nov., male imagoes, paratypes: 1, head (lateral view); 2, head (dorsal view); 3, hind right wing (dorsal view); 4, hind left wing (dorsal view)." httpUri="https://zenodo.org/record/185068/files/figure.png" pageId="1" pageNumber="2">Figs. 3, 4</figureCitation>
). Fore legs light brown, femora with diffuse smudges distally, middle and hind legs yellow to yellowish-brown.
</paragraph>
<paragraph id="962E36E7C07B4F3DFF51FA2AFF11F9A7" blockId="1.[151,1437,826,2012]" pageId="1" pageNumber="2">Abdominal terga with small brownish lateral smudges. Tergum I brown, terga IIVI whitish-yellow with a pair of diffuse central brownish smudges, terga VIIX distinctly darker, yellowish-brown, with narrow elongate central spots. Sterna whitish-yellow to dirty yellow with elongate whitish central spot on segments VII VIII.</paragraph>
<paragraph id="962E36E7C07B4F3DFF51F98AFC24F96F" blockId="1.[151,1437,826,2012]" pageId="1" pageNumber="2">
Genitalia pale, yellowish-white to dirty yellow. Forceps distinctly bent up at approximately 45° angle in lateral view (
<figureCitation id="0EAA2A62C07B4F3DFEBBF9E2FEB6F9F7" box="[303,374,1626,1652]" captionStart="FIGURES 5 7" captionStartId="3.[151,269,1704,1728]" captionTargetBox="[173,1416,349,1667]" captionTargetId="figure@3.[151,1436,342,1680]" captionTargetPageId="3" captionText="FIGURES 5 7. Baetis zdenkae sp. nov., male imagoes, paratypes: 5 6, forceps (ventral view); 7, tip of abdomen (lateral view)." httpUri="https://zenodo.org/record/185069/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
). Basal segment about as long as wide without a posteromedial projection, segment 1 elongate conical with slightly convex external margin, segment 2 narrow at the base, slightly expanded distally (widened part as long as 2/33/4 of the segment length), segment 3 elongate, slightly longer than wide (
<figureCitation id="0EAA2A62C07B4F3DFAF5F912FF05F96F" captionStart="FIGURES 5 7" captionStartId="3.[151,269,1704,1728]" captionTargetBox="[173,1416,349,1667]" captionTargetId="figure@3.[151,1436,342,1680]" captionTargetPageId="3" captionText="FIGURES 5 7. Baetis zdenkae sp. nov., male imagoes, paratypes: 5 6, forceps (ventral view); 7, tip of abdomen (lateral view)." httpUri="https://zenodo.org/record/185069/files/figure.png" pageId="1" pageNumber="2">Figs. 5, 6</figureCitation>
). Cerci whitish-grey to dirty yellow, first 47 segments light brown.
</paragraph>
<paragraph id="962E36E7C07B4F3DFF51F943FA93F897" blockId="1.[151,1437,826,2012]" box="[197,1363,1786,1812]" pageId="1" pageNumber="2">
<emphasis id="A4E5EAF5C07B4F3DFF51F943FEADF897" box="[197,365,1787,1812]" italics="true" pageId="1" pageNumber="2">Female imago</emphasis>
. Size: body length:
<quantity id="51699B02C07B4F3DFDCDF942FD76F897" box="[601,694,1786,1812]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="2" unit="mm" value="6.5">6.5 mm</quantity>
; fore wings length:
<quantity id="51699B02C07B4F3DFC0AF942FC3CF897" box="[926,1020,1786,1812]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="2" unit="mm" value="6.2">6.2 mm</quantity>
; cerci length:
<quantity id="51699B02C07B4F3DFB34F942FA8FF897" box="[1184,1359,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" metricValueMax="1.34" metricValueMin="1.3199999999999998" pageId="1" pageNumber="2" unit="mm" value="13.3" valueMax="13.4" valueMin="13.2">13.213.4 mm</quantity>
.
</paragraph>
<paragraph id="962E36E7C07B4F3DFF51F89AFC8EF8E7" blockId="1.[151,1437,826,2012]" pageId="1" pageNumber="2">General body color yellowish-brown to brown. Head uniformly brown. Eyes black with yellowish maculations. Ocelli yellowish with black bases, antennae brown.</paragraph>
<paragraph id="962E36E7C07B4F3DFF51F8CAFB8FF85F" blockId="1.[151,1437,826,2012]" pageId="1" pageNumber="2">Thorax yellowish-brown to brown with dark brown smudges on mesonotum. Sterna whitish-yellow to light brown. Legs yellowish-brown to light brown, fore legs darker. Wings transparent, hyaline, fore wings with distinct yellowish-brown venation, hind wings similar to male, veins darker.</paragraph>
<caption id="C2EE666FC0784F3EFF03FA0DFD33FA6C" httpUri="https://zenodo.org/record/185068/files/figure.png" pageId="2" pageNumber="3" targetBox="[178,1411,208,1428]" targetPageId="2">
<paragraph id="962E36E7C0784F3EFF03FA0DFD33FA6C" blockId="2.[151,1435,1461,1519]" pageId="2" pageNumber="3">
<emphasis id="A4E5EAF5C0784F3EFF03FA0DFE83FA4E" bold="true" box="[151,323,1461,1485]" pageId="2" pageNumber="3">FIGURES 14.</emphasis>
<taxonomicName id="51914D64C0784F3EFEDFFA0EFE2BFA4E" box="[331,491,1462,1485]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0784F3EFEDFFA0EFE2BFA4E" box="[331,491,1462,1485]" italics="true" pageId="2" pageNumber="3">Baetis zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0784F3EFE66FA0DFD88FA4E" box="[498,584,1461,1485]" pageId="2" pageNumber="3" rank="species">
<emphasis id="A4E5EAF5C0784F3EFE66FA0DFD88FA4E" bold="true" box="[498,584,1461,1485]" pageId="2" pageNumber="3">sp. nov.</emphasis>
</taxonomicNameLabel>
, male imagoes, paratypes: 1, head (lateral view); 2, head (dorsal view); 3, hind right wing (dorsal view); 4, hind left wing (dorsal view).
</paragraph>
</caption>
<paragraph id="962E36E7C0784F3EFF51F9A6FE45F9E3" blockId="2.[151,1436,1566,2032]" pageId="2" pageNumber="3">Abdominal terga uniform, yellowish-brown to light brown, sterna paler. Cerci light brown to whitish-yellow, darker at bases.</paragraph>
<paragraph id="962E36E7C0784F3EFF51F9D6FAAFF90B" blockId="2.[151,1436,1566,2032]" box="[197,1391,1646,1672]" pageId="2" pageNumber="3">
<emphasis id="A4E5EAF5C0784F3EFF51F9D6FE9FF904" box="[197,351,1646,1671]" italics="true" pageId="2" pageNumber="3">Mature larva</emphasis>
. Size: body length:
<quantity id="51699B02C0784F3EFDDFF9D6FD18F90B" box="[587,728,1646,1672]" metricMagnitude="-3" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="2" pageNumber="3" unit="mm" value="6.45" valueMax="7.5" valueMin="5.4">5.47.5 mm</quantity>
(male),
<quantity id="51699B02C0784F3EFCAFF9D6FC08F90B" box="[827,968,1646,1672]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="8.5" metricValueMin="5.8" pageId="2" pageNumber="3" unit="mm" value="7.15" valueMax="8.5" valueMin="5.8">5.88.5 mm</quantity>
(female); cerci length: 2.0
<quantity id="51699B02C0784F3EFA9BF9D6FAABF90B" box="[1295,1387,1646,1672]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="3" unit="mm" value="3.3">3.3 mm</quantity>
.
</paragraph>
<paragraph id="962E36E7C0784F3FFF51F92EFF76FEA9" blockId="2.[151,1436,1566,2032]" lastBlockId="3.[151,1437,152,298]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
General body color dark, light brown to dark brown. Head yellowish-brown to brown, frons and clypeus paler. The whole head surface covered with short setae and solitary bases of setae. Surface of clypeus with numerous setae. Eyes black, developing turbinate eyes light brown to brown, surrounded by a narrow yellowish ring. Antennae whitish-yellow to light brown, flagellum paler than pedicel and scape, with numerous fine hairs. Pedicel surface with short setae only, scape surface with solitary setae concentrated distally. Labrum relatively wide (width/length ratio 1.521.55), with 1 + 47 long submarginal bristles and a few pointed lateral setae (
<figureCitation id="0EAA2A62C0784F3EFF70F83EFEFAF823" box="[228,314,1926,1952]" captionStart="FIGURES 8 12" captionStartId="4.[151,269,1957,1981]" captionTargetBox="[197,1410,579,1912]" captionTargetId="figure@4.[177,1410,574,1933]" captionTargetPageId="4" captionText="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="2" pageNumber="3">Figs. 9</figureCitation>
,
<figureCitation id="0EAA2A62C0784F3EFEDEF83EFEAAF823" box="[330,362,1926,1952]" captionStart="FIGURES 13 16" captionStartId="5.[151,269,1186,1210]" captionTargetBox="[151,1435,194,1162]" captionTargetId="figure@5.[151,1436,194,1162]" captionTargetPageId="5" captionText="FIGURES 13 16. Baetis zdenkae sp. nov., mature larvae: 13, labrum (dorsal view); 14, apical portion of the last segment of maxillary palpus; 15, arrangement of spatulate setae on the surface of thorax; 16, outer margin of femur (hind leg). Scale bars: 13 = 500 nm; 14, 15 = 100 nm; 16 = 500 nm." httpUri="https://zenodo.org/record/185071/files/figure.png" pageId="2" pageNumber="3">13</figureCitation>
). Mandible incisors as in
<figureCitation id="0EAA2A62C0784F3EFD32F83EFD32F823" box="[678,754,1926,1952]" captionStart="FIGURES 8 12" captionStartId="4.[151,269,1957,1981]" captionTargetBox="[197,1410,579,1912]" captionTargetId="figure@4.[177,1410,574,1933]" captionTargetPageId="4" captionText="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="2" pageNumber="3">Figs 8</figureCitation>
a, b. Tip of maxillary palpus conical with barely visible, small apical spine, surface with numerous fine hairs (
<figureCitation id="0EAA2A62C0784F3EFC96F816FC99F84B" box="[770,857,1966,1992]" captionStart="FIGURES 13 16" captionStartId="5.[151,269,1186,1210]" captionTargetBox="[151,1435,194,1162]" captionTargetId="figure@5.[151,1436,194,1162]" captionTargetPageId="5" captionText="FIGURES 13 16. Baetis zdenkae sp. nov., mature larvae: 13, labrum (dorsal view); 14, apical portion of the last segment of maxillary palpus; 15, arrangement of spatulate setae on the surface of thorax; 16, outer margin of femur (hind leg). Scale bars: 13 = 500 nm; 14, 15 = 100 nm; 16 = 500 nm." httpUri="https://zenodo.org/record/185071/files/figure.png" pageId="2" pageNumber="3">Fig. 14</figureCitation>
). Segment 2 of labial palpus relatively broad with slightly concave inner margin, about twice as broad at apex than at base, distomedial lobe as wide as about 1/ 3 of segment 3 at the base. Segment 3 slightly asymmetrical, slightly longer than 1/2 length of segment 2. The length/width ratio of segments 2 and 3 combined is 1.82.1 (
<figureCitation id="0EAA2A62C0794F3FFCFFFF78FC04FF59" box="[875,964,192,218]" captionStart="FIGURES 8 12" captionStartId="4.[151,269,1957,1981]" captionTargetBox="[197,1410,579,1912]" captionTargetId="figure@4.[177,1410,574,1933]" captionTargetPageId="4" captionText="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="3" pageNumber="4">Fig. 10</figureCitation>
). Glossae and paraglossae wide, robust. Paraglossae with 3 regular rows of long bristles apically, basal row consisting of at least 16 bristles (
<figureCitation id="0EAA2A62C0794F3FFAB7FF50FA41FE81" box="[1315,1409,232,258]" captionStart="FIGURES 8 12" captionStartId="4.[151,269,1957,1981]" captionTargetBox="[197,1410,579,1912]" captionTargetId="figure@4.[177,1410,574,1933]" captionTargetPageId="4" captionText="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="3" pageNumber="4">Figs. 11</figureCitation>
a, b).
</paragraph>
<caption id="C2EE666FC0794F3FFF03F910FF1BF961" httpUri="https://zenodo.org/record/185069/files/figure.png" pageId="3" pageNumber="4" targetBox="[173,1416,349,1667]" targetPageId="3">
<paragraph id="962E36E7C0794F3FFF03F910FF1BF961" blockId="3.[151,1436,1704,1762]" pageId="3" pageNumber="4">
<emphasis id="A4E5EAF5C0794F3FFF03F910FE82F943" bold="true" box="[151,322,1704,1728]" pageId="3" pageNumber="4">FIGURES 57.</emphasis>
<taxonomicName id="51914D64C0794F3FFEDCF911FE27F943" box="[328,487,1705,1728]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0794F3FFEDCF911FE27F943" box="[328,487,1705,1728]" italics="true" pageId="3" pageNumber="4">Baetis zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0794F3FFE79F910FD82F943" box="[493,578,1704,1728]" pageId="3" pageNumber="4" rank="species">
<emphasis id="A4E5EAF5C0794F3FFE79F910FD82F943" bold="true" box="[493,578,1704,1728]" pageId="3" pageNumber="4">sp. nov.</emphasis>
</taxonomicNameLabel>
, male imagoes, paratypes: 56, forceps (ventral view); 7, tip of abdomen (lateral view).
</paragraph>
</caption>
<paragraph id="962E36E7C0794F38FF51F8A9FC72FDC1" blockId="3.[151,1437,1809,2035]" lastBlockId="4.[151,1437,152,578]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Thorax dark, light to dark brown, pronotum with a pair of large pale subtriangular areas laterally, mesonotal color pattern as in
<figureCitation id="0EAA2A62C0794F3FFE00F881FE2AF8D0" box="[404,490,1849,1875]" captionStart="FIGURE 17" captionStartId="6.[151,255,1814,1838]" captionTargetBox="[630,979,231,1770]" captionTargetId="figure@6.[599,987,197,1787]" captionTargetPageId="6" captionText="FIGURE 17. Baetis zdenkae sp. nov., mature larva, general view of body (dorsal view)." httpUri="https://zenodo.org/record/185072/files/figure.png" pageId="3" pageNumber="4">Fig. 17</figureCitation>
. Metanotum brownish with central pale spot. Thorax surface with numerous fine hairs and slender spatulate setae rounded apically (
<figureCitation id="0EAA2A62C0794F3FFD61F8D9FC8DF8F8" box="[757,845,1889,1915]" captionStart="FIGURES 13 16" captionStartId="5.[151,269,1186,1210]" captionTargetBox="[151,1435,194,1162]" captionTargetId="figure@5.[151,1436,194,1162]" captionTargetPageId="5" captionText="FIGURES 13 16. Baetis zdenkae sp. nov., mature larvae: 13, labrum (dorsal view); 14, apical portion of the last segment of maxillary palpus; 15, arrangement of spatulate setae on the surface of thorax; 16, outer margin of femur (hind leg). Scale bars: 13 = 500 nm; 14, 15 = 100 nm; 16 = 500 nm." httpUri="https://zenodo.org/record/185071/files/figure.png" pageId="3" pageNumber="4">Fig. 15</figureCitation>
). Sterna pale, yellowish to light brown. Legs yellowish to light brown with distinct large central dark spot on all femora (occasionally with relatively small central spot). Tibiae with brown distal spots (occasionally only with diffuse brownish smudges), tarsi darker distally. Outer margin of femora with a single, sparse row of large spatulate bristles alternating with fine hairs and small submarginal spatulate setae (
<figureCitation id="0EAA2A62C07E4F38FDEBFF20FD1EFF31" box="[639,734,152,178]" captionStart="FIGURES 13 16" captionStartId="5.[151,269,1186,1210]" captionTargetBox="[151,1435,194,1162]" captionTargetId="figure@5.[151,1436,194,1162]" captionTargetPageId="5" captionText="FIGURES 13 16. Baetis zdenkae sp. nov., mature larvae: 13, labrum (dorsal view); 14, apical portion of the last segment of maxillary palpus; 15, arrangement of spatulate setae on the surface of thorax; 16, outer margin of femur (hind leg). Scale bars: 13 = 500 nm; 14, 15 = 100 nm; 16 = 500 nm." httpUri="https://zenodo.org/record/185071/files/figure.png" pageId="4" pageNumber="5">Fig. 16</figureCitation>
). Spatulate bristles relatively short and robust, distinctly rounded at apex, distal portion with shallowly splintered margins (
<figureCitation id="0EAA2A62C07E4F38FC3EFF78FBA9FF59" box="[938,1129,192,218]" captionStart="FIGURES 18 21" captionStartId="7.[151,269,1192,1216]" captionTargetBox="[151,1435,197,1165]" captionTargetId="figure@7.[151,1436,197,1165]" captionTargetPageId="7" captionText="FIGURES 18 21. Baetis zdenkae sp. nov., mature larvae, outer margin of femora (hind legs). Scale bars: 18 = 200 nm; 19 = 100 nm; 20, 21 = 100 nm." httpUri="https://zenodo.org/record/185073/files/figure.png" pageId="4" pageNumber="5">Figs. 18, 19, 21</figureCitation>
). Femora with (1) slender spatulate setae with convergent margins (
<figureCitation id="0EAA2A62C07E4F38FD14FF50FCCEFE81" box="[640,782,232,258]" captionStart="FIGURES 18 21" captionStartId="7.[151,269,1192,1216]" captionTargetBox="[151,1435,197,1165]" captionTargetId="figure@7.[151,1436,197,1165]" captionTargetPageId="7" captionText="FIGURES 18 21. Baetis zdenkae sp. nov., mature larvae, outer margin of femora (hind legs). Scale bars: 18 = 200 nm; 19 = 100 nm; 20, 21 = 100 nm." httpUri="https://zenodo.org/record/185073/files/figure.png" pageId="4" pageNumber="5">Figs. 1820</figureCitation>
) and (2) relatively wide and apically rounded spatulate setae with subparallel margins (
<figureCitation id="0EAA2A62C07E4F38FD86FEA8FDABFEA9" box="[530,619,272,298]" captionStart="FIGURES 18 21" captionStartId="7.[151,269,1192,1216]" captionTargetBox="[151,1435,197,1165]" captionTargetId="figure@7.[151,1436,197,1165]" captionTargetPageId="7" captionText="FIGURES 18 21. Baetis zdenkae sp. nov., mature larvae, outer margin of femora (hind legs). Scale bars: 18 = 200 nm; 19 = 100 nm; 20, 21 = 100 nm." httpUri="https://zenodo.org/record/185073/files/figure.png" pageId="4" pageNumber="5">Fig. 21</figureCitation>
). Submarginal spatulate setae of distal portion of femora with splintered margins. Inner margin of femora with uniform row of small pointed spines and fine hairs; all surface of femora with spatulate setae and fine hairs (
<figureCitation id="0EAA2A62C07E4F38FD08FED8FD37FEF9" box="[668,759,352,378]" captionStart="FIGURES 22 25" captionStartId="9.[151,270,1634,1658]" captionTargetBox="[151,1435,640,1607]" captionTargetId="figure@9.[151,1436,640,1608]" captionTargetPageId="9" captionText="FIGURES 22 25. Baetis zdenkae sp. nov., mature larvae: 22, surface of femur; 23, outer margin of tibia; 24, inner margin of tibia; 25, tarsal claw (hind legs). Scale bars: 22 24 = 200 nm; 25 = 500 nm." httpUri="https://zenodo.org/record/185074/files/figure.png" pageId="4" pageNumber="5">Fig. 22</figureCitation>
). Outer margins of tibiae with deeply splintered, slender spatulas alternating with fine hairs (
<figureCitation id="0EAA2A62C07E4F38FDD0FE30FD5DFE21" box="[580,669,392,418]" captionStart="FIGURES 22 25" captionStartId="9.[151,270,1634,1658]" captionTargetBox="[151,1435,640,1607]" captionTargetId="figure@9.[151,1436,640,1608]" captionTargetPageId="9" captionText="FIGURES 22 25. Baetis zdenkae sp. nov., mature larvae: 22, surface of femur; 23, outer margin of tibia; 24, inner margin of tibia; 25, tarsal claw (hind legs). Scale bars: 22 24 = 200 nm; 25 = 500 nm." httpUri="https://zenodo.org/record/185074/files/figure.png" pageId="4" pageNumber="5">Fig. 23</figureCitation>
). Inner margins with irregular row of spines and fine hairs (
<figureCitation id="0EAA2A62C07E4F38FAF9FE30FF78FE49" captionStart="FIGURES 22 25" captionStartId="9.[151,270,1634,1658]" captionTargetBox="[151,1435,640,1607]" captionTargetId="figure@9.[151,1436,640,1608]" captionTargetPageId="9" captionText="FIGURES 22 25. Baetis zdenkae sp. nov., mature larvae: 22, surface of femur; 23, outer margin of tibia; 24, inner margin of tibia; 25, tarsal claw (hind legs). Scale bars: 22 24 = 200 nm; 25 = 500 nm." httpUri="https://zenodo.org/record/185074/files/figure.png" pageId="4" pageNumber="5">Fig. 24</figureCitation>
). Outer margins of tarsi with small pointed spines and hairs; small submarginal spatulate setae, with distally splintered edges. Inner margins with large pointed spines; splintered submarginal spatulate setae, and fine hairs. Surface of tarsi with numerous hairs and splintered spatulate setae. Tarsal claws elongate, relatively slender with at least 10 teeth, without subapical setae (
<figureCitation id="0EAA2A62C07E4F38FC81FD90FCB7FDC1" box="[789,887,552,578]" captionStart="FIGURES 8 12" captionStartId="4.[151,269,1957,1981]" captionTargetBox="[197,1410,579,1912]" captionTargetId="figure@4.[177,1410,574,1933]" captionTargetPageId="4" captionText="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="4" pageNumber="5">Figs. 12</figureCitation>
,
<figureCitation id="0EAA2A62C07E4F38FC17FD90FC64FDC1" box="[899,932,552,578]" captionStart="FIGURES 22 25" captionStartId="9.[151,270,1634,1658]" captionTargetBox="[151,1435,640,1607]" captionTargetId="figure@9.[151,1436,640,1608]" captionTargetPageId="9" captionText="FIGURES 22 25. Baetis zdenkae sp. nov., mature larvae: 22, surface of femur; 23, outer margin of tibia; 24, inner margin of tibia; 25, tarsal claw (hind legs). Scale bars: 22 24 = 200 nm; 25 = 500 nm." httpUri="https://zenodo.org/record/185074/files/figure.png" pageId="4" pageNumber="5">25</figureCitation>
).
</paragraph>
<caption id="C2EE666FC07E4F38FF03F81DFB73F85C" httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="4" pageNumber="5" targetBox="[197,1410,579,1912]" targetPageId="4">
<paragraph id="962E36E7C07E4F38FF03F81DFB73F85C" blockId="4.[151,1436,1957,2015]" pageId="4" pageNumber="5">
<emphasis id="A4E5EAF5C07E4F38FF03F81DFE91F83E" bold="true" box="[151,337,1957,1981]" pageId="4" pageNumber="5">FIGURES 812.</emphasis>
<taxonomicName id="51914D64C07E4F38FECCF81EFE39F83E" box="[344,505,1958,1981]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C07E4F38FECCF81EFE39F83E" box="[344,505,1958,1981]" italics="true" pageId="4" pageNumber="5">Baetis zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC07E4F38FD94F81DFD96F83E" box="[512,598,1957,1981]" pageId="4" pageNumber="5" rank="species">
<emphasis id="A4E5EAF5C07E4F38FD94F81DFD96F83E" bold="true" box="[512,598,1957,1981]" pageId="4" pageNumber="5">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg).
</paragraph>
</caption>
<caption id="C2EE666FC07F4F39FF03FB1AFCEBFB7C" httpUri="https://zenodo.org/record/185071/files/figure.png" pageId="5" pageNumber="6" targetBox="[151,1435,194,1162]" targetPageId="5">
<paragraph id="962E36E7C07F4F39FF03FB1AFCEBFB7C" blockId="5.[151,1436,1186,1279]" pageId="5" pageNumber="6">
<emphasis id="A4E5EAF5C07F4F39FF03FB1AFE9FFB39" bold="true" box="[151,351,1186,1210]" pageId="5" pageNumber="6">FIGURES 1316.</emphasis>
<taxonomicName id="51914D64C07F4F39FEF3FB1BFDC8FB39" box="[359,520,1187,1210]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C07F4F39FEF3FB1BFDC8FB39" box="[359,520,1187,1210]" italics="true" pageId="5" pageNumber="6">Baetis zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC07F4F39FD84FB1AFDA7FB39" box="[528,615,1186,1210]" pageId="5" pageNumber="6" rank="species">
<emphasis id="A4E5EAF5C07F4F39FD84FB1AFDA7FB39" bold="true" box="[528,615,1186,1210]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larvae: 13, labrum (dorsal view); 14, apical portion of the last segment of maxillary palpus; 15, arrangement of spatulate setae on the surface of thorax; 16, outer margin of femur (hind leg). Scale bars: 13 = 500 nm; 14, 15 = 100 nm; 16 = 500 nm.
</paragraph>
</caption>
<paragraph id="962E36E7C07F4F39FF51FA95FBC6F92C" blockId="5.[151,1437,1325,1991]" pageId="5" pageNumber="6">
Abdominal terga dark, yellowish-brown to brown, color pattern as in
<figureCitation id="0EAA2A62C07F4F39FB94FA95FB97FAC4" box="[1024,1111,1325,1351]" captionStart="FIGURE 17" captionStartId="6.[151,255,1814,1838]" captionTargetBox="[630,979,231,1770]" captionTargetId="figure@6.[599,987,197,1787]" captionTargetPageId="6" captionText="FIGURE 17. Baetis zdenkae sp. nov., mature larva, general view of body (dorsal view)." httpUri="https://zenodo.org/record/185072/files/figure.png" pageId="5" pageNumber="6">Fig. 17</figureCitation>
, muscle insertions pale (cf.
<bibRefCitation id="F2004B16C07F4F39FF03FAEDFEDFFAEC" author="Jacob" box="[151,287,1365,1391]" pageId="5" pageNumber="17" refString="Jacob, U. (2003) Baetis Leach, 1815 sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimungslusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob 2003</bibRefCitation>
), posterior margins with broad triangular bluntly pointed teeth alternating with setae (
<figureCitation id="0EAA2A62C07F4F39FA97FAEDFA99FAEC" box="[1283,1369,1365,1391]" captionStart="FIGURES 26 29" captionStartId="10.[151,269,1472,1496]" captionTargetBox="[151,1435,477,1445]" captionTargetId="figure@10.[151,1436,477,1445]" captionTargetPageId="10" captionText="FIGURES 26 29. Baetis zdenkae sp. nov., mature larvae: 26, surface of abdominal tergum IV; 27, surface of abdominal tergum VI; 28, posterior margin of abdominal tergum IV; 29, outer margin of paraproct plate. Scale bars: 26 29 = 20 mm." httpUri="https://zenodo.org/record/185075/files/figure.png" pageId="5" pageNumber="6">Fig. 28</figureCitation>
), surface with numerous fine hairs, wide bases of hairs and small spatulate setae. Two distinct
<typeStatus id="492A8845C07F4F39FB58FAC5FACAFA14" box="[1228,1290,1405,1431]" pageId="5" pageNumber="6">types</typeStatus>
of spatulate setae discernible on abdominal tergal surface: (1) elongate narrow ones with lateral convergent margins, and (2) relatively rare ones with subparallel or divergent lateral margins, rounded at apex (
<figureCitation id="0EAA2A62C07F4F39FB33FA75FAFBFA64" box="[1191,1339,1485,1511]" captionStart="FIGURES 26 29" captionStartId="10.[151,269,1472,1496]" captionTargetBox="[151,1435,477,1445]" captionTargetId="figure@10.[151,1436,477,1445]" captionTargetPageId="10" captionText="FIGURES 26 29. Baetis zdenkae sp. nov., mature larvae: 26, surface of abdominal tergum IV; 27, surface of abdominal tergum VI; 28, posterior margin of abdominal tergum IV; 29, outer margin of paraproct plate. Scale bars: 26 29 = 20 mm." httpUri="https://zenodo.org/record/185075/files/figure.png" pageId="5" pageNumber="6">Figs. 26, 27</figureCitation>
). Sterna uniform, yellowish-brown to brown with a pair of short transversal divergent strokes near the anterior margins. Inner margin of paraprocts with a single row of at least 10 (usually 1215) distinct teeth (
<figureCitation id="0EAA2A62C07F4F39FA94F9A5FA98F9B4" box="[1280,1368,1565,1591]" captionStart="FIGURES 26 29" captionStartId="10.[151,269,1472,1496]" captionTargetBox="[151,1435,477,1445]" captionTargetId="figure@10.[151,1436,477,1445]" captionTargetPageId="10" captionText="FIGURES 26 29. Baetis zdenkae sp. nov., mature larvae: 26, surface of abdominal tergum IV; 27, surface of abdominal tergum VI; 28, posterior margin of abdominal tergum IV; 29, outer margin of paraproct plate. Scale bars: 26 29 = 20 mm." httpUri="https://zenodo.org/record/185075/files/figure.png" pageId="5" pageNumber="6">Fig. 29</figureCitation>
), surface with fine hairs and sparse spatulate setae. Gills 1 and 7 slightly asymmetric and subequal in size. Gills 2 6 asymmetric and distally rounded. Both outer and inner gill margins with small teeth alternating with fine hairs. Cerci uniform, yellowish-brown to brown, paracercus shorter by 1/2.
</paragraph>
<paragraph id="962E36E7C07F4F39FF51F905FD4BF954" blockId="5.[151,1437,1325,1991]" box="[197,651,1725,1751]" pageId="5" pageNumber="6">Male and female subimagos unknown.</paragraph>
</subSubSection>
<subSubSection id="DE8B656CC07F4F39FF51F95DFC3BF844" pageId="5" pageNumber="6" type="etymology">
<paragraph id="962E36E7C07F4F39FF51F95DFCE9F8A4" blockId="5.[151,1437,1325,1991]" pageId="5" pageNumber="6">
<emphasis id="A4E5EAF5C07F4F39FF51F95DFE8AF97C" bold="true" box="[197,330,1765,1791]" pageId="5" pageNumber="6">Etymology</emphasis>
. The species is named for Zdenka, wife of the senior author, to acknowledge her distinguished field assistance during sampling of mayflies on Rhodos.
</paragraph>
<paragraph id="962E36E7C07F4F39FF51F88DFC3BF844" blockId="5.[151,1437,1325,1991]" pageId="5" pageNumber="6">
<emphasis id="A4E5EAF5C07F4F39FF51F88DFED5F8CC" bold="true" box="[197,277,1845,1871]" pageId="5" pageNumber="6">
<typeStatus id="492A8845C07F4F39FF51F88DFECBF8CC" box="[197,267,1845,1871]" pageId="5" pageNumber="6">Types</typeStatus>
.
</emphasis>
<typeStatus id="492A8845C07F4F39FE8AF88DFE7DF8CC" box="[286,445,1845,1871]" pageId="5" pageNumber="6" type="holotype">HOLOTYPE</typeStatus>
: male imago,
<collectingCountry id="EE867677C07F4F39FDF2F88DFD1BF8CC" box="[614,731,1845,1871]" name="Greece" pageId="5" pageNumber="6">GREECE</collectingCountry>
, Rhodos (Rodos)
<collectingCountry id="EE867677C07F4F39FC26F88DFC3CF8CC" box="[946,1020,1845,1871]" name="Iceland" pageId="5" pageNumber="6">Island</collectingCountry>
, stream at the Kremastis Bridge in Kremastí, about
<quantity id="51699B02C07F4F39FEC8F8E5FE68F8F4" box="[348,424,1885,1911]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="6" unit="km" value="14.0">14 km</quantity>
SW of Rhodos City, about
<quantity id="51699B02C07F4F39FD60F8E5FCF1F8F4" box="[756,817,1885,1911]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="km" value="3.0">3 km</quantity>
NE of the Diagoras International Airport and about
<quantity id="51699B02C07F4F39FF03F83DFF21F81C" box="[151,225,1925,1951]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="m" value="500.0">500 m</quantity>
from its mouth to the Aegean Sea at about
<quantity id="51699B02C07F4F39FD74F83DFCDCF81C" box="[736,796,1925,1951]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="m" value="10.0">10 m</quantity>
a. s. l.,
<date id="E22F1027C07F4F39FCECF83DFBDFF81C" box="[888,1055,1925,1951]" pageId="5" pageNumber="6" value="2005-05-28">May 28, 2005</date>
.
<typeStatus id="492A8845C07F4F39FBBFF83DFB12F81C" box="[1067,1234,1925,1951]" pageId="5" pageNumber="6" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="8097FD6EC07F4F39FB48F83DFAEBF81C" box="[1244,1323,1925,1951]" pageId="5" pageNumber="6" type="male">6 male</specimenCount>
imagoes,
<specimenCount id="8097FD6EC07F4F39FF03F815FF3DF844" box="[151,253,1965,1991]" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
imago,
<specimenCount id="8097FD6EC07F4F39FEC8F815FE19F844" box="[348,473,1965,1991]" pageId="5" pageNumber="6" type="larva">275 larvae</specimenCount>
, same locality and collection date as
<typeStatus id="492A8845C07F4F39FC18F815FC37F844" box="[908,1015,1965,1991]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="C2EE666FC07C4F3AFF03F8AEFB86F8AE" box="[151,1094,1814,1838]" httpUri="https://zenodo.org/record/185072/files/figure.png" pageId="6" pageNumber="7" targetBox="[630,979,231,1770]" targetPageId="6">
<paragraph id="962E36E7C07C4F3AFF03F8AEFB86F8AE" blockId="6.[151,1094,1814,1838]" box="[151,1094,1814,1838]" pageId="6" pageNumber="7">
<emphasis id="A4E5EAF5C07C4F3AFF03F8AEFEE6F8AD" bold="true" box="[151,294,1814,1838]" pageId="6" pageNumber="7">FIGURE 17.</emphasis>
<taxonomicName id="51914D64C07C4F3AFEB9F8AEFE0CF8AE" box="[301,460,1814,1837]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C07C4F3AFEB9F8AEFE0CF8AE" box="[301,460,1814,1837]" italics="true" pageId="6" pageNumber="7">Baetis zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC07C4F3AFE47F8AEFDE8F8AD" box="[467,552,1814,1838]" pageId="6" pageNumber="7" rank="species">
<emphasis id="A4E5EAF5C07C4F3AFE47F8AEFDE8F8AD" bold="true" box="[467,552,1814,1838]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larva, general view of body (dorsal view).
</paragraph>
</caption>
<subSubSection id="DE8B656CC07C4F34FF51F8D8FCB1F949" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="962E36E7C07C4F3AFF51F8D8FE82F870" blockId="6.[151,1436,1888,2035]" pageId="6" pageNumber="7">
<emphasis id="A4E5EAF5C07C4F3AFF51F8D8FD62F8F8" bold="true" box="[197,674,1888,1915]" pageId="6" pageNumber="7">
Further material examined (no
<typeStatus id="492A8845C07C4F3AFDC4F8D8FD53F8F9" box="[592,659,1888,1914]" pageId="6" pageNumber="7">types</typeStatus>
).
</emphasis>
<specimenCount id="8097FD6EC07C4F3AFD3EF8D9FCE7F8F8" box="[682,807,1889,1915]" pageId="6" pageNumber="7" type="larva">256 larvae</specimenCount>
,
<collectingCountry id="EE867677C07C4F3AFCA6F8D9FC67F8F8" box="[818,935,1889,1915]" name="Greece" pageId="6" pageNumber="7">GREECE</collectingCountry>
, Rhodos (Rodos)
<collectingCountry id="EE867677C07C4F3AFBEDF8D9FB03F8F8" box="[1145,1219,1889,1915]" name="Iceland" pageId="6" pageNumber="7">Island</collectingCountry>
, same locality and collection date as
<typeStatus id="492A8845C07C4F3AFEF9F831FE17F820" box="[365,471,1929,1955]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
;
<specimenCount id="8097FD6EC07C4F3AFE71F831FD86F820" box="[485,582,1929,1955]" pageId="6" pageNumber="7" type="larva">8 larvae</specimenCount>
,
<collectingCountry id="EE867677C07C4F3AFDC7F831FD69F820" box="[595,681,1929,1955]" name="Greece" pageId="6" pageNumber="7">Greece</collectingCountry>
, Rhodos Island, Argiros Potamos (stream), about
<quantity id="51699B02C07C4F3AFB6FF831FA86F820" box="[1275,1350,1929,1955]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="km" value="30.0">30 km</quantity>
SW of Rhodos City,
<quantity id="51699B02C07C4F3AFEACF809FEB5F848" box="[312,373,1969,1995]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="km" value="1.0">1 km</quantity>
NE of Kalavárda and about
<quantity id="51699B02C07C4F3AFD5CF809FCC5F848" box="[712,773,1969,1995]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="km" value="1.0">1 km</quantity>
from its mouth to the Aegean Sea at about
<quantity id="51699B02C07C4F3AFA9EF809FA86F848" box="[1290,1350,1969,1995]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="m" value="10.0">10 m</quantity>
a. s. l.,
<date id="E22F1027C07C4F3AFF03F861FEFEF870" box="[151,318,2009,2035]" pageId="6" pageNumber="7" value="2005-05-26">May 26, 2005</date>
.
</paragraph>
<caption id="C2EE666FC07D4F3BFF03FB10FE24FB61" httpUri="https://zenodo.org/record/185073/files/figure.png" pageId="7" pageNumber="8" targetBox="[151,1435,197,1165]" targetPageId="7">
<paragraph id="962E36E7C07D4F3BFF03FB10FE24FB61" blockId="7.[151,1436,1192,1250]" pageId="7" pageNumber="8">
<emphasis id="A4E5EAF5C07D4F3BFF03FB10FE9EFB43" bold="true" box="[151,350,1192,1216]" pageId="7" pageNumber="8">FIGURES 1821.</emphasis>
<taxonomicName id="51914D64C07D4F3BFEF1FB10FDC4FB3C" box="[357,516,1192,1215]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C07D4F3BFEF1FB10FDC4FB3C" box="[357,516,1192,1215]" italics="true" pageId="7" pageNumber="8">Baetis zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC07D4F3BFD98FB10FDA1FB43" box="[524,609,1192,1216]" pageId="7" pageNumber="8" rank="species">
<emphasis id="A4E5EAF5C07D4F3BFD98FB10FDA1FB43" bold="true" box="[524,609,1192,1216]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larvae, outer margin of femora (hind legs). Scale bars: 18 = 200 nm; 19 = 100 nm; 20, 21 = 100 nm.
</paragraph>
</caption>
<paragraph id="962E36E7C07D4F3BFF51FAA8FCBBFA49" blockId="7.[151,1437,1296,2002]" pageId="7" pageNumber="8">
All material (preserved in 70% alcohol) was collected by Zdenka and Tomáš Soldán. Genitalia of two male imagoes on slides.
<typeStatus id="492A8845C07D4F3BFE2CFA80FDE8FAD1" box="[440,552,1336,1362]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
, most
<typeStatus id="492A8845C07D4F3BFDE2FA80FD26FAD1" box="[630,742,1336,1362]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
and other material examined deposited in collection of the Biological Centre, Academy of Sciences of the
<collectingCountry id="EE867677C07D4F3BFD58FAD8FC4BFAF9" box="[716,907,1376,1402]" name="Czech Republic" pageId="7" pageNumber="8">Czech Republic</collectingCountry>
, Institute of Entomology, Č eské Budĕjovice,
<collectingCountry id="EE867677C07D4F3BFF03FA30FE97FA21" box="[151,343,1416,1442]" name="Czech Republic" pageId="7" pageNumber="8">Czech Republic</collectingCountry>
;
<specimenCount id="8097FD6EC07D4F3BFEF0FA30FE13FA21" box="[356,467,1416,1442]" pageId="7" pageNumber="8" type="larva">50 larvae</specimenCount>
and
<specimenCount id="8097FD6EC07D4F3BFD9BFA30FD9FFA21" box="[527,607,1416,1442]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
imago (all
<typeStatus id="492A8845C07D4F3BFD7FFA30FC9DFA21" box="[747,861,1416,1442]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
) deposited in the State Museum of Natural History, National Academy of Sciences of
<collectingCountry id="EE867677C07D4F3BFDF0FA08FD06FA49" box="[612,710,1456,1482]" name="Ukraine" pageId="7" pageNumber="8">Ukraine</collectingCountry>
, Lviv,
<collectingCountry id="EE867677C07D4F3BFC81FA08FCB7FA49" box="[789,887,1456,1482]" name="Ukraine" pageId="7" pageNumber="8">Ukraine</collectingCountry>
.
</paragraph>
<paragraph id="962E36E7C07D4F3BFF51FA60FA43F939" blockId="7.[151,1437,1296,2002]" pageId="7" pageNumber="8">
The comparative material of
<taxonomicName id="51914D64C07D4F3BFD8AFA60FD72FA72" box="[542,690,1496,1521]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C07D4F3BFD8AFA60FD72FA72" box="[542,690,1496,1521]" italics="true" pageId="7" pageNumber="8">B. buceratus</emphasis>
</taxonomicName>
was collected in Central
<collectingCountry id="EE867677C07D4F3BFC75FA60FB8BFA71" box="[993,1099,1496,1522]" name="Czech Republic" pageId="7" pageNumber="8">Bohemia</collectingCountry>
(the Berounka River basin),
<collectingCountry id="EE867677C07D4F3BFF03F9B8FF3EF999" box="[151,254,1536,1562]" name="Slovakia" pageId="7" pageNumber="8">Slovakia</collectingCountry>
and
<collectingCountry id="EE867677C07D4F3BFEA1F9B8FE53F999" box="[309,403,1536,1562]" name="Ukraine" pageId="7" pageNumber="8">Ukraine</collectingCountry>
(the Latorica, Tisza and Dniester River basins) at numerous localities. Larvae and adults of
<taxonomicName id="51914D64C07D4F3BFF23F990FEDBF9C2" box="[183,283,1576,1601]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C07D4F3BFF23F990FEDBF9C2" box="[183,283,1576,1601]" italics="true" pageId="7" pageNumber="8">B. nexus</emphasis>
</taxonomicName>
were collected at several localities of the Dyje (Thaya) River basin in Moravia (
<collectingCountry id="EE867677C07D4F3BFB5AF990FA4FF9C1" box="[1230,1423,1576,1602]" name="Czech Republic" pageId="7" pageNumber="8">Czech Republic</collectingCountry>
),
<collectingCountry id="EE867677C07D4F3BFF03F9E8FF3EF9E9" box="[151,254,1616,1642]" name="Slovakia" pageId="7" pageNumber="8">Slovakia</collectingCountry>
and
<collectingCountry id="EE867677C07D4F3BFEA2F9E8FE54F9E9" box="[310,404,1616,1642]" name="Ukraine" pageId="7" pageNumber="8">Ukraine</collectingCountry>
(the Latorica and Tisza River basins). For descriptions of some localities see
<bibRefCitation id="F2004B16C07D4F3BFAB5F9E8FE81F911" author="Landa" pageId="7" pageNumber="18" refString="Landa, V. &amp; Soldan, T. (1989) Distribution of Mayflies (Ephemeroptera) and its changes in the Labe Basin of Czechoslovakia in Connection with Changes of Water Quality. Academia, Praha, Studie CSAV, 17, 171 pp. (in Czech, Engl. sum.)" type="book" year="1989">
<collectingCountry id="EE867677C07D4F3BFAB5F9E8FAAAF9E9" box="[1313,1386,1616,1642]" name="Netherlands" pageId="7" pageNumber="8">Landa</collectingCountry>
and Soldán (1989)
</bibRefCitation>
,
<bibRefCitation id="F2004B16C07D4F3BFEDAF9C0FD7EF911" author="Soldan" box="[334,702,1656,1682]" pageId="7" pageNumber="18" refString="Soldan, T. &amp; Zahradkova, S. (2000) Ephemeroptera of the Czech Republic: Atlas of Distribution. In: Helesic, J. &amp; Zahradkova, S. (Eds). Fauna Aquatica Europae Centralis I. Masaryk University, Brno (Czech Republic), 401 pp." type="book" year="2000">Soldán and Zahrádková (2000)</bibRefCitation>
, and
<bibRefCitation id="F2004B16C07D4F3BFD69F9C0FC6CF911" author="Horsak" box="[765,940,1656,1682]" pageId="7" pageNumber="17" refString="Horsak, M. (2001) Contribution to the knowledge of macroinvertebrate fauna of the Dyje River downstream of the Nove Mlyny Reservoirs (Czech Republic). Scripta Facultatis scientiarum naturalium Universitatis Masarykianae brunensis, Biologia, Supplementum, 27, 41 - 62." type="journal article" year="2001">Horsák (2001)</bibRefCitation>
. Material of
<taxonomicName id="51914D64C07D4F3BFBD7F9C0FB54F912" box="[1091,1172,1656,1681]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C07D4F3BFBD7F9C0FB54F912" box="[1091,1172,1656,1681]" italics="true" pageId="7" pageNumber="8">B. spei</emphasis>
</taxonomicName>
was not studied, morphological characters and other data are based on the original description by
<bibRefCitation id="F2004B16C07D4F3BFB8EF918FAEAF939" author="Thomas" box="[1050,1322,1696,1722]" pageId="7" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas and Dia (1985</bibRefCitation>
,
<bibRefCitation id="F2004B16C07D4F3BFAA3F918FAB6F939" author="Thomas" box="[1335,1398,1696,1722]" pageId="7" pageNumber="19" refString="Thomas, A. &amp; Dia, A. (2007) Complements et corrections a la faune des Ephemeropteres du Proche-Orient. 7. Description de l'imago male de Baetis spei Thomas, 1985 [Ephemeroptera, Baetidae]. Ephemera, 8 (2): 75 - 78." type="journal article" year="2007">2007</bibRefCitation>
).
</paragraph>
<paragraph id="962E36E7C07D4F34FF51F970FEB1FC29" blockId="7.[151,1437,1296,2002]" lastBlockId="8.[151,1437,152,2020]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="A4E5EAF5C07D4F3BFF51F970FD42F961" bold="true" box="[197,642,1736,1762]" pageId="7" pageNumber="8">Differential diagnosis and affinities.</emphasis>
<taxonomicName id="51914D64C07D4F3BFD1DF970FCC7F962" box="[649,775,1736,1761]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C07D4F3BFD1DF970FCC7F962" box="[649,775,1736,1761]" italics="true" pageId="7" pageNumber="8">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC07D4F3BFC9AF970FCACF961" box="[782,876,1736,1762]" pageId="7" pageNumber="8" rank="species">
<emphasis id="A4E5EAF5C07D4F3BFC9AF970FCACF961" bold="true" box="[782,876,1736,1762]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
can be easily distinguished from the other three representatives of the
<taxonomicName id="51914D64C07D4F3BFE08F948FDF1F88A" box="[412,561,1776,1801]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C07D4F3BFE08F948FDF1F88A" box="[412,561,1776,1801]" italics="true" pageId="7" pageNumber="8">B. buceratus</emphasis>
</taxonomicName>
species-group (
<taxonomicName id="51914D64C07D4F3BFD7AF948FB92F889" authority="Eaton, 1870" authorityName="Eaton" authorityYear="1870" box="[750,1106,1776,1802]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="buceratus" subGenus="Baetis">
<emphasis id="A4E5EAF5C07D4F3BFD7AF948FCC7F88A" box="[750,775,1776,1801]" italics="true" pageId="7" pageNumber="8">B.</emphasis>
(
<emphasis id="A4E5EAF5C07D4F3BFC8EF948FCF3F88A" box="[794,819,1776,1801]" italics="true" pageId="7" pageNumber="8">B.</emphasis>
)
<emphasis id="A4E5EAF5C07D4F3BFCD1F948FC79F88A" box="[837,953,1776,1801]" italics="true" pageId="7" pageNumber="8">buceratus</emphasis>
<bibRefCitation id="F2004B16C07D4F3BFC55F948FB92F889" author="Eaton" box="[961,1106,1776,1802]" pageId="7" pageNumber="17" refString="Eaton, A. E. (1870) On some new British species of Ephemeridae. Transactions of the Entomological Society of London, 1870 (1), 1 - 8." type="journal article" year="1870">Eaton, 1870</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="51914D64C07D4F3BFBCBF948FA57F889" authority="Navas, 1918" authorityName="Navas" authorityYear="1918" box="[1119,1431,1776,1802]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nexus" subGenus="Baetis">
<emphasis id="A4E5EAF5C07D4F3BFBCBF948FBB8F88A" box="[1119,1144,1776,1801]" italics="true" pageId="7" pageNumber="8">B.</emphasis>
(
<emphasis id="A4E5EAF5C07D4F3BFB1FF948FB64F88A" box="[1163,1188,1776,1801]" italics="true" pageId="7" pageNumber="8">B.</emphasis>
)
<emphasis id="A4E5EAF5C07D4F3BFB22F948FB39F88A" box="[1206,1273,1776,1801]" italics="true" pageId="7" pageNumber="8">nexus</emphasis>
<bibRefCitation id="F2004B16C07D4F3BFA95F948FA57F889" author="Navas" box="[1281,1431,1776,1802]" pageId="7" pageNumber="18" refString="Navas, L. (1918) Excursiones entomologicas por el Norte de la provincia de Lerida (6 - 24 de Julio de 1917). Butlleti de la Institucio catalana d'Historia Natural, 1918, 36 - 49." type="journal article" year="1918">Navás, 1918</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="51914D64C07D4F3BFF5FF8A0FD9FF8B1" authority="Thomas &amp; Dia, 1985" authorityName="Thomas &amp; Dia" authorityYear="1985" box="[203,607,1816,1842]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spei" subGenus="Baetis">
<emphasis id="A4E5EAF5C07D4F3BFF5FF8A0FF1DF8B2" box="[203,221,1816,1841]" italics="true" pageId="7" pageNumber="8">B</emphasis>
. (
<emphasis id="A4E5EAF5C07D4F3BFF6CF8A0FED1F8B2" box="[248,273,1816,1841]" italics="true" pageId="7" pageNumber="8">B.</emphasis>
)
<emphasis id="A4E5EAF5C07D4F3BFEB1F8A0FE94F8B2" box="[293,340,1816,1841]" italics="true" pageId="7" pageNumber="8">spei</emphasis>
<bibRefCitation id="F2004B16C07D4F3BFECAF8A0FD9FF8B1" author="Thomas" box="[350,607,1816,1842]" pageId="7" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas &amp; Dia, 1985</bibRefCitation>
</taxonomicName>
) by the following combination of morphological characters:
<emphasis id="A4E5EAF5C07D4F3BFAABF8A0FF22F8D9" bold="true" italics="true" pageId="7" pageNumber="8">in male imago</emphasis>
: (1) facetted surface of turbinate eyes yellowish-brown to light brown (yellowish brown to orange brown in
<taxonomicName id="51914D64C07D4F3BFE98F8D0FE61F802" box="[268,417,1896,1921]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C07D4F3BFE98F8D0FE61F802" box="[268,417,1896,1921]" italics="true" pageId="7" pageNumber="8">B. buceratus</emphasis>
</taxonomicName>
, dark brownish in
<taxonomicName id="51914D64C07D4F3BFD13F8D0FD2DF802" box="[647,749,1896,1921]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C07D4F3BFD13F8D0FD2DF802" box="[647,749,1896,1921]" italics="true" pageId="7" pageNumber="8">B. nexus</emphasis>
</taxonomicName>
and flesh-colored in
<taxonomicName id="51914D64C07D4F3BFC7BF8D0FB81F802" box="[1007,1089,1896,1921]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C07D4F3BFC7BF8D0FB81F802" box="[1007,1089,1896,1921]" italics="true" pageId="7" pageNumber="8">B. spei</emphasis>
</taxonomicName>
), (2) tricolor combination of stalk of turbinate eyes (stalk paler, light orange in
<taxonomicName id="51914D64C07D4F3BFD7CF828FCBCF82A" box="[744,892,1936,1961]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C07D4F3BFD7CF828FCBCF82A" box="[744,892,1936,1961]" italics="true" pageId="7" pageNumber="8">B. buceratus</emphasis>
</taxonomicName>
, eyes unicolorous in
<taxonomicName id="51914D64C07D4F3BFBEDF828FB1DF82A" box="[1145,1245,1936,1961]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C07D4F3BFBEDF828FB1DF82A" box="[1145,1245,1936,1961]" italics="true" pageId="7" pageNumber="8">B. nexus</emphasis>
</taxonomicName>
, stalk flesh-colored in
<taxonomicName id="51914D64C07D4F3BFF61F800FE87F852" box="[245,327,1976,2001]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C07D4F3BFF61F800FE87F852" box="[245,327,1976,2001]" italics="true" pageId="7" pageNumber="8">B. spei</emphasis>
</taxonomicName>
with diffuse and intermittent basal ring), (3) third vein of hind wings relatively short, about 0.380.40 wing length (about half of wing length in
<taxonomicName id="51914D64C0724F34FD6CFF20FC4BFF32" box="[760,907,152,177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FD6CFF20FC4BFF32" box="[760,907,152,177]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
, 1/2 length to longer in
<taxonomicName id="51914D64C0724F34FB36FF20FAC4FF32" box="[1186,1284,152,177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FB36FF20FAC4FF32" box="[1186,1284,152,177]" italics="true" pageId="8" pageNumber="9">B. nexus</emphasis>
</taxonomicName>
), (4) forceps conspicuously bent up in lateral view (as in
<taxonomicName id="51914D64C0724F34FD3FFF78FC81FF5A" box="[683,833,192,217]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FD3FFF78FC81FF5A" box="[683,833,192,217]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
, nearly straight in
<taxonomicName id="51914D64C0724F34FBB2FF78FB4BFF5A" box="[1062,1163,192,217]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FBB2FF78FB4BFF5A" box="[1062,1163,192,217]" italics="true" pageId="8" pageNumber="9">B. nexus</emphasis>
</taxonomicName>
and
<taxonomicName id="51914D64C0724F34FB5CFF78FADAFF5A" box="[1224,1306,192,217]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FB5CFF78FADAFF5A" box="[1224,1306,192,217]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
), (5) basal segment of forceps without posteromedial projection (posteromedially produced into lobe-like projection in
<emphasis id="A4E5EAF5C0724F34FF03FEA8FE5BFEAA" box="[151,411,272,297]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="51914D64C0724F34FF03FEA8FEEDFEAA" box="[151,301,272,297]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">B. buceratus</taxonomicName>
,
<taxonomicName id="51914D64C0724F34FEADFEA8FE5BFEAA" box="[313,411,272,297]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">B. nexus</taxonomicName>
</emphasis>
and
<taxonomicName id="51914D64C0724F34FE47FEA8FDE2FEAA" box="[467,546,272,297]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FE47FEA8FDE2FEAA" box="[467,546,272,297]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
), and (6) segment 1 of forceps elongate conical (conical in
<taxonomicName id="51914D64C0724F34FB43FEA8FAABFEAA" box="[1239,1387,272,297]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FB43FEA8FAABFEAA" box="[1239,1387,272,297]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
and
<taxonomicName id="51914D64C0724F34FF03FE80FF27FED2" box="[151,231,312,337]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FF03FE80FF27FED2" box="[151,231,312,337]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
, oval, bulbous with apparently convex inner and outer lateral margins in
<taxonomicName id="51914D64C0724F34FBD1FE80FB69FED2" box="[1093,1193,312,337]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FBD1FE80FB69FED2" box="[1093,1193,312,337]" italics="true" pageId="8" pageNumber="9">B. nexus</emphasis>
</taxonomicName>
);
<emphasis id="A4E5EAF5C0724F34FB56FE8FFAE0FED2" bold="true" box="[1218,1312,311,337]" italics="true" pageId="8" pageNumber="9">in larva</emphasis>
: (7) structure of segments 2 and 3 of labial palpi (segment 3 shorter than or equal to 1/2 of segment
<quantity id="51699B02C0724F34FB5CFED8FB35FEF9" box="[1224,1269,352,378]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="9" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="51914D64C0724F34FB69FED8FAA0FEFA" box="[1277,1376,352,377]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FB69FED8FAA0FEFA" box="[1277,1376,352,377]" italics="true" pageId="8" pageNumber="9">B. nexus</emphasis>
</taxonomicName>
), (8) structure of mandible incisors (cf. e.g., our
<figureCitation id="0EAA2A62C0724F34FD00FE30FD25FE21" box="[660,741,392,418]" captionStart="FIGURES 8 12" captionStartId="4.[151,269,1957,1981]" captionTargetBox="[197,1410,579,1912]" captionTargetId="figure@4.[177,1410,574,1933]" captionTargetPageId="4" captionText="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="8" pageNumber="9">Figs. 8</figureCitation>
a, b with
<figureCitation id="0EAA2A62C0724F34FCC6FE30FC57FE21" box="[850,919,392,418]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,1461,1485]" captionTargetBox="[178,1411,208,1428]" captionTargetId="figure@2.[167,1420,197,1435]" captionTargetPageId="2" captionText="FIGURES 1 4. Baetis zdenkae sp. nov., male imagoes, paratypes: 1, head (lateral view); 2, head (dorsal view); 3, hind right wing (dorsal view); 4, hind left wing (dorsal view)." httpUri="https://zenodo.org/record/185068/files/figure.png" pageId="8" pageNumber="9">
Fig.
<quantity id="51699B02C0724F34FC1CFE30FC57FE21" box="[904,919,392,418]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="8" pageNumber="9" unit="g" value="3.0">3</quantity>
</figureCitation>
g, d by
<bibRefCitation id="F2004B16C0724F34FC64FE30FB3AFE21" author="Thomas" box="[1008,1274,392,418]" pageId="8" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas and Dia 1985</bibRefCitation>
: 242 showing
<taxonomicName id="51914D64C0724F34FF03FE08FF26FE4A" box="[151,230,432,457]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FF03FE08FF26FE4A" box="[151,230,432,457]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
), (9) paraglossae with basal row of apical bristles consisting of at least 16 bristles (about 1012 bristles in
<taxonomicName id="51914D64C0724F34FF20FE60FEC3FE72" box="[180,259,472,497]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FF20FE60FEC3FE72" box="[180,259,472,497]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
), (10) outer margins of femora with single sparse row of spatulate bristles, alternating with fine hairs and submarginal small spatulate setae (as in
<taxonomicName id="51914D64C0724F34FD08FDB8FD2BFD9A" box="[668,747,512,537]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FD08FDB8FD2BFD9A" box="[668,747,512,537]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
), (11) structure of small spatulate setae of femora (some of them conspicuously splintered as in
<taxonomicName id="51914D64C0724F34FDDFFD90FD5DFDC2" box="[587,669,552,577]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FDDFFD90FD5DFDC2" box="[587,669,552,577]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
), (12) arrangement of small spatulate setae on dorsal surface of femora, (13) posterior margins of abdominal terga with broad bluntly pointed triangular teeth, alternating with setae (teeth rounded, semicircular in
<taxonomicName id="51914D64C0724F34FDD8FDC0FD5DFD12" box="[588,669,632,657]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FDD8FDC0FD5DFD12" box="[588,669,632,657]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
, narrow triangular blunt pointed teeth in
<taxonomicName id="51914D64C0724F34FB13FDC0FADBFD12" box="[1159,1307,632,657]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FB13FDC0FADBFD12" box="[1159,1307,632,657]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
), (14) surface of terga with numerous fine hairs, spatulate setae, and wide bases of hairs and spatulate setae (without spatulate setae in
<taxonomicName id="51914D64C0724F34FEFCFD70FE78FD62" box="[360,440,712,737]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FEFCFD70FE78FD62" box="[360,440,712,737]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
), (15) inner margin of paraproct with 1015 distinct teeth (about 2025 teeth of different shape in
<taxonomicName id="51914D64C0724F34FEDFFD48FE5BFC8A" box="[331,411,752,777]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FEDFFD48FE5BFC8A" box="[331,411,752,777]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
; about 20 teeth in
<taxonomicName id="51914D64C0724F34FDE3FD48FCCBFC8A" box="[631,779,752,777]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FDE3FD48FCCBFC8A" box="[631,779,752,777]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
and
<bibRefCitation id="F2004B16C0724F34FCD0FD48FB8DFC89" author="Thomas" box="[836,1101,752,778]" pageId="8" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas and Dia 1985</bibRefCitation>
: 243,
<figureCitation id="0EAA2A62C0724F34FB07FD48FB29FC89" box="[1171,1257,752,778]" captionStart="FIGURES 8 12" captionStartId="4.[151,269,1957,1981]" captionTargetBox="[197,1410,579,1912]" captionTargetId="figure@4.[177,1410,574,1933]" captionTargetPageId="4" captionText="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
; about 25 teeth in
<taxonomicName id="51914D64C0724F34FF21FCA0FDB1FCB1" authority="Muller-Liebenau 1969: 149" authorityName="Muller-Liebenau" authorityPageNumber="149" authorityYear="1969" box="[181,625,792,818]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FF21FCA0FEE0FCB2" box="[181,288,792,817]" italics="true" pageId="8" pageNumber="9">B. nexus,</emphasis>
<bibRefCitation id="F2004B16C0724F34FEBDFCA0FDF6FCB1" author="Muller-Liebenau" box="[297,566,792,818]" pageId="8" pageNumber="18" refString="Muller-Liebenau, I. (1969) Revision der europaischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera). Gewasser und Abwasser, 48 / 49, 1 - 214." type="journal article" year="1969">Müller-Liebenau 1969</bibRefCitation>
: 149
</taxonomicName>
, Abb. 109), (16) paracercus shorter than cerci by 1/2 (subequal in
<taxonomicName id="51914D64C0724F34FA17FCA0FECBFCDA" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FA17FCA0FECBFCDA" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
and
<taxonomicName id="51914D64C0724F34FED6FCF8FE65FCDA" box="[322,421,832,857]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FED6FCF8FE65FCDA" box="[322,421,832,857]" italics="true" pageId="8" pageNumber="9">B. nexus</emphasis>
</taxonomicName>
, about 2/3 of cerci in
<taxonomicName id="51914D64C0724F34FD30FCF8FD33FCDA" box="[676,755,832,857]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FD30FCF8FD33FCDA" box="[676,755,832,857]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
), and (17) specific color pattern of thorax and abdomen in dorsal view (e.g., a single, conspicuous pale band in the middle of pronotum in
<taxonomicName id="51914D64C0724F34FBC0FCD0FB66FC02" box="[1108,1190,872,897]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FBC0FCD0FB66FC02" box="[1108,1190,872,897]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
, in
<bibRefCitation id="F2004B16C0724F34FB43FCD0FF16FC29" author="Thomas" pageId="8" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas and Dia 1985</bibRefCitation>
: 243,
<figureCitation id="0EAA2A62C0724F34FE8FFC28FEA2FC29" box="[283,354,912,938]" captionStart="FIGURES 5 7" captionStartId="3.[151,269,1704,1728]" captionTargetBox="[173,1416,349,1667]" captionTargetId="figure@3.[151,1436,342,1680]" captionTargetPageId="3" captionText="FIGURES 5 7. Baetis zdenkae sp. nov., male imagoes, paratypes: 5 6, forceps (ventral view); 7, tip of abdomen (lateral view)." httpUri="https://zenodo.org/record/185069/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="962E36E7C0724F34FF51FC00FEFFFB69" blockId="8.[151,1437,152,2020]" pageId="8" pageNumber="9">
The male genitalia of
<taxonomicName id="51914D64C0724F34FE58FC00FD8CFC52" box="[460,588,952,977]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FE58FC00FD8CFC52" box="[460,588,952,977]" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FDC0FC0FFD73FC52" box="[596,691,951,977]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FDC0FC0FFD73FC52" bold="true" box="[596,691,951,977]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
are similar to
<taxonomicName id="51914D64C0724F34FCF5FC00FC37FC52" box="[865,1015,952,977]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FCF5FC00FC37FC52" box="[865,1015,952,977]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
, in the structure and proportions of segments 13, and conspicuously bent up forceps (see
<bibRefCitation id="F2004B16C0724F34FCA7FC58FB83FC79" author="Muller-Liebenau" box="[819,1091,992,1018]" pageId="8" pageNumber="18" refString="Muller-Liebenau, I. (1969) Revision der europaischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera). Gewasser und Abwasser, 48 / 49, 1 - 214." type="journal article" year="1969">Müller-Liebenau 1969</bibRefCitation>
: 140, Abb. 99100 or
<bibRefCitation id="F2004B16C0724F34FACDFC58FF16FBA1" author="Jacob" pageId="8" pageNumber="17" refString="Jacob, U. (2003) Baetis Leach, 1815 sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimungslusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob 2003</bibRefCitation>
: 96, Abb. 17ab). On the other hand, the male imago of the new species differs markedly from
<taxonomicName id="51914D64C0724F34FABBFBB0FF08FBCA" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FABBFBB0FF08FBCA" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
in color pattern of the facetted surface of the turbinate eyes and their stalks and in hind wing venation (cf. characters 13, see also
<bibRefCitation id="F2004B16C0724F34FE26FBE0FD7EFBF1" author="Muller-Liebenau" box="[434,702,1112,1138]" pageId="8" pageNumber="18" refString="Muller-Liebenau, I. (1969) Revision der europaischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera). Gewasser und Abwasser, 48 / 49, 1 - 214." type="journal article" year="1969">Müller-Liebenau 1969</bibRefCitation>
: 139140, Abb. 104j or
<bibRefCitation id="F2004B16C0724F34FC4DFBE0FBA2FBF1" author="Jacob" box="[985,1122,1112,1138]" pageId="8" pageNumber="17" refString="Jacob, U. (2003) Baetis Leach, 1815 sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimungslusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob 2003</bibRefCitation>
: 97). On the other hand,
<taxonomicName id="51914D64C0724F34FA17FBE0FF35FB1A" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FA17FBE0FF35FB1A" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FF68FBC7FE99FB1A" box="[252,345,1151,1177]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FF68FBC7FE99FB1A" bold="true" box="[252,345,1151,1177]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
can be easily distinguished from
<taxonomicName id="51914D64C0724F34FD4AFB38FC81FB1A" box="[734,833,1152,1177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FD4AFB38FC81FB1A" box="[734,833,1152,1177]" italics="true" pageId="8" pageNumber="9">B. nexus</emphasis>
</taxonomicName>
by the different arrangement of all critical imaginal characters (16, see above and cf.
<bibRefCitation id="F2004B16C0724F34FDA6FB10FC80FB41" author="Muller-Liebenau" box="[562,832,1192,1218]" pageId="8" pageNumber="18" refString="Muller-Liebenau, I. (1969) Revision der europaischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera). Gewasser und Abwasser, 48 / 49, 1 - 214." type="journal article" year="1969">Müller-Liebenau 1969</bibRefCitation>
: 146147, Abb. 101, 107, and 108, or
<bibRefCitation id="F2004B16C0724F34FA9AFB10FA58FB41" author="Jacob" box="[1294,1432,1192,1218]" pageId="8" pageNumber="17" refString="Jacob, U. (2003) Baetis Leach, 1815 sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimungslusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob 2003</bibRefCitation>
: 98, Abb. 17c).
</paragraph>
<paragraph id="962E36E7C0724F34FF51FB40FCB1F949" blockId="8.[151,1437,152,2020]" pageId="8" pageNumber="9">
Larvae of
<taxonomicName id="51914D64C0724F34FEAAFB40FE7DFA92" box="[318,445,1272,1297]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FEAAFB40FE7DFA92" box="[318,445,1272,1297]" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FE51FB4FFDE2FA92" box="[453,546,1271,1297]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FE51FB4FFDE2FA92" bold="true" box="[453,546,1271,1297]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from all remaining representatives of the
<taxonomicName id="51914D64C0724F34FA9CFB40FA5BFA92" box="[1288,1435,1272,1297]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FA9CFB40FA5BFA92" box="[1288,1435,1272,1297]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
species-group by the combination of characters (9 and 14), and, particularly really the critical characters (8 and 16) separate this species from
<taxonomicName id="51914D64C0724F34FDBDFAF0FD7DFAE2" box="[553,701,1352,1377]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FDBDFAF0FD7DFAE2" box="[553,701,1352,1377]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
and
<taxonomicName id="51914D64C0724F34FD61FAF0FC98FAE2" box="[757,856,1352,1377]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FD61FAF0FC98FAE2" box="[757,856,1352,1377]" italics="true" pageId="8" pageNumber="9">B. nexus</emphasis>
</taxonomicName>
. Additionally,
<taxonomicName id="51914D64C0724F34FB92FAF0FB45FAE2" box="[1030,1157,1352,1377]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FB92FAF0FB45FAE2" box="[1030,1157,1352,1377]" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FB18FAFFFB29FAE2" box="[1164,1257,1351,1377]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FB18FAFFFB29FAE2" bold="true" box="[1164,1257,1351,1377]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="51914D64C0724F34FA17FAF0FF1AFA0A" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0724F34FA17FAF0FF1AFA0A" italics="true" pageId="8" pageNumber="9">B. nexus</emphasis>
</taxonomicName>
by the structure of outer margin of femora and paraproct (characters 10 and 15). The latter two characters and the arrangement of triangular teeth on the posterior margin of abdominal terga (character 13), distinguish the new species from
<taxonomicName id="51914D64C0724F34FE74FA78FDF0FA5A" box="[480,560,1472,1497]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FE74FA78FDF0FA5A" box="[480,560,1472,1497]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
with small rounded, nearly semicircular teeth, cf.
<bibRefCitation id="F2004B16C0724F34FB10FA78FEF0F981" author="Thomas" pageId="8" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">
Thomas and Dia (1985: 243:
<figureCitation id="0EAA2A62C0724F34FF46FA50FEE6F981" box="[210,294,1512,1538]" captionStart="FIGURES 8 12" captionStartId="4.[151,269,1957,1981]" captionTargetBox="[197,1410,579,1912]" captionTargetId="figure@4.[177,1410,574,1933]" captionTargetPageId="4" captionText="FIGURES 8 12. Baetis zdenkae sp. nov., mature larvae: 8, left (a) and right (b) mandibular incisors; 9, labrum (dorsal view); 10, labial palpus (ventral view); 11, paraglossa (a) and glossa (b); 12, tarsal claw (hind leg)." httpUri="https://zenodo.org/record/185070/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
)
</bibRefCitation>
. Contrary to
<taxonomicName id="51914D64C0724F34FE53FA50FDD6F982" box="[455,534,1512,1537]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FE53FA50FDD6F982" box="[455,534,1512,1537]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
(muscle insertions on head well visible and contrasting), the head of
<taxonomicName id="51914D64C0724F34FAA8FA50FF00F9AA" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FAA8FA50FF00F9AA" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FF53F9B7FEE4F9AA" box="[199,292,1551,1577]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FF53F9B7FEE4F9AA" bold="true" box="[199,292,1551,1577]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
, is rather unicolorous. Labrum of
<taxonomicName id="51914D64C0724F34FD23F9A8FCF6F9AA" box="[695,822,1552,1577]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FD23F9A8FCF6F9AA" box="[695,822,1552,1577]" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FCA9F9B7FC5BF9AA" box="[829,923,1551,1577]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FCA9F9B7FC5BF9AA" bold="true" box="[829,923,1551,1577]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
, provided with 1 + 47 bristles is relatively wider (width/length ratio 1.521.55) vs. 1 + 68 bristles and width/length ratio
<quantity id="51699B02C0724F34FBDCF980FB1DF9D1" box="[1096,1245,1592,1618]" metricMagnitude="-2" metricUnit="m" metricValue="3.6195" metricValueMax="3.81" metricValueMin="3.429" pageId="8" pageNumber="9" unit="in" value="1.425" valueMax="1.5" valueMin="1.35">1.351.50 in</quantity>
<taxonomicName id="51914D64C0724F34FB71F980FAF6F9D2" box="[1253,1334,1592,1617]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FB71F980FAF6F9D2" box="[1253,1334,1592,1617]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
. Further differences between
<taxonomicName id="51914D64C0724F34FE1CF9D8FDC6F9FA" box="[392,518,1632,1657]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FE1CF9D8FDC6F9FA" box="[392,518,1632,1657]" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FD99F9E7FDAAF9FA" box="[525,618,1631,1657]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FD99F9E7FDAAF9FA" bold="true" box="[525,618,1631,1657]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="51914D64C0724F34FDECF9D8FCCBF9FA" box="[632,779,1632,1657]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0724F34FDECF9D8FCCBF9FA" box="[632,779,1632,1657]" italics="true" pageId="8" pageNumber="9">B. buceratus</emphasis>
</taxonomicName>
and
<taxonomicName id="51914D64C0724F34FCD7F9D8FC52F9FA" box="[835,914,1632,1657]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0724F34FCD7F9D8FC52F9FA" box="[835,914,1632,1657]" italics="true" pageId="8" pageNumber="9">B. spei</emphasis>
</taxonomicName>
can be found also in thoracic and abdominal terga color patterns (see
<bibRefCitation id="F2004B16C0724F34FE54F930FD0FF921" author="Muller-Liebenau" box="[448,719,1672,1698]" pageId="8" pageNumber="18" refString="Muller-Liebenau, I. (1969) Revision der europaischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera). Gewasser und Abwasser, 48 / 49, 1 - 214." type="journal article" year="1969">Müller-Liebenau 1969</bibRefCitation>
: 141145 and 147150, Abb. 102103, 109 or
<bibRefCitation id="F2004B16C0724F34FA99F930FA58F921" author="Jacob" box="[1293,1432,1672,1698]" pageId="8" pageNumber="17" refString="Jacob, U. (2003) Baetis Leach, 1815 sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimungslusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob 2003</bibRefCitation>
: 9698, Abb. 3l, 18, 19, and
<bibRefCitation id="F2004B16C0724F34FE4BF908FD23F949" author="Thomas" box="[479,739,1712,1738]" pageId="8" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas and Dia 1985</bibRefCitation>
for details).
</paragraph>
</subSubSection>
<subSubSection id="DE8B656CC0724F37FF51F96FFD6BFAE7" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="962E36E7C0724F35FF51F96FFC76FDC7" blockId="8.[151,1437,152,2020]" lastBlockId="9.[151,1437,152,580]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="A4E5EAF5C0724F34FF51F96FFEEAF972" bold="true" box="[197,298,1751,1777]" pageId="8" pageNumber="9">Biology.</emphasis>
Larvae of
<taxonomicName id="51914D64C0724F34FE23F960FDFBF972" box="[439,571,1752,1777]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FE23F960FDFBF972" box="[439,571,1752,1777]" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FDD2F96FFD68F972" box="[582,680,1751,1777]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FDD2F96FFD68F972" bold="true" box="[582,680,1751,1777]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
evidently prefer stream run places of the “lowland” biotopes within the island. These localities (mostly at elevations of
<quantity id="51699B02C0724F34FCD8F8B8FC12F899" box="[844,978,1792,1818]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="8" pageNumber="9" unit="m" value="175.0" valueMax="250.0" valueMin="100.0">100250 m</quantity>
a. s. l.) were inhabited by other baetid species, e.g., by apparently dominant
<taxonomicName id="51914D64C0724F34FDC2F890FB54F8C1" authority="Bengtsson, 1912" authorityName="Bengtsson" authorityYear="1912" box="[598,1172,1832,1858]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="digitatus" subGenus="Nigrobaetis">
<emphasis id="A4E5EAF5C0724F34FDC2F890FC74F8C2" box="[598,948,1832,1857]" italics="true" pageId="8" pageNumber="9">Baetis (Nigrobaetis) digitatus</emphasis>
(
<bibRefCitation id="F2004B16C0724F34FC50F890FB4CF8C1" author="Bengtsson" box="[964,1164,1832,1858]" pageId="8" pageNumber="17" refString="Bengtsson, S. (1912) Neue Ephemeriden aus Schweden. Entomologisk Tidskrift, 33, 107 - 117." type="journal article" year="1912">Bengtsson, 1912</bibRefCitation>
)
</taxonomicName>
which is the first faunistic record from Rhodos and
<emphasis id="A4E5EAF5C0724F34FD90F8E8FD7CF8EA" box="[516,700,1872,1897]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="51914D64C0724F34FD90F8E8FD78F8EA" box="[516,696,1872,1897]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fascicaudale">P. fascicaudale</taxonomicName>
,
</emphasis>
but no larvae of
<taxonomicName id="51914D64C0724F34FC12F8E8FBC5F8EA" box="[902,1029,1872,1897]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0724F34FC12F8E8FBC5F8EA" box="[902,1029,1872,1897]" italics="true" pageId="8" pageNumber="9">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0724F34FB99F8F7FBAAF8EA" box="[1037,1130,1871,1897]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A4E5EAF5C0724F34FB99F8F7FBAAF8EA" bold="true" box="[1037,1130,1871,1897]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
have been collected. Larvae of this new species were found in two streams with permanent water of about
<quantity id="51699B02C0724F34FB0FF8C0FB2AF811" box="[1179,1258,1912,1938]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="9" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">13 m</quantity>
across and an average depth of
<quantity id="51699B02C0724F34FEF3F818FE21F839" box="[359,481,1952,1978]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="9" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">1030 cm</quantity>
, moderate to fast current velocity and stony bottom overgrown, in some places, with submerged aquatic vegetation. The current velocity was about
<quantity id="51699B02C0724F34FC39F872FBE5F867" box="[941,1061,1994,2020]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="8" pageNumber="9" unit="cm" value="35.0" valueMax="50.0" valueMin="20.0">2050 cm</quantity>
.s -1 at the most densely inhabited places, the density of larvae, roughly estimated at about 100300 ind.m -1, decreased rapidly with decreasing current velocity, places with slower current were inhabited rather sparsely. Water temperature ranged from 17 ºC early in the morning to 2326.5 ºC in the afternoon at sun-exposed, densely inhabited places (
<typeStatus id="492A8845C0734F35FA97FF52FAF7FE87" box="[1283,1335,234,260]" pageId="9" pageNumber="10">type</typeStatus>
locality, up to 28.5 ºC in pools with very rare occurrence of larvae). As to substrate roughness, larvae preferred gravel places (lithal) covered with cobble and pebble. Their densities in places covered with granules (2.0 mm in diameter) and very coarse (1.0 mm) or coarse (
<quantity id="51699B02C0734F35FD56FEDAFCE0FEFF" box="[706,800,354,380]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="mm" value="0.5">0.5 mm</quantity>
) sand were very low showing rather incidental occurrence. Larvae were not found on fine substrates like medium to very fine sand or silt. If submerged aquatic plants occurred at the places preferred, larval densities among vegetation were comparable with those of stony habitats. However, substrate requirements of larvae do not seem to be pronounced. Stream banks are regulated at numerous places in both biotopes inhabited. For instance, the Argiros Potamos near Kalávarda is a very shallow stream with an artificial concrete streambed in some places.
</paragraph>
<caption id="C2EE666FC0734F35FF03F9DAFBC3F91F" httpUri="https://zenodo.org/record/185074/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1435,640,1607]" targetPageId="9">
<paragraph id="962E36E7C0734F35FF03F9DAFBC3F91F" blockId="9.[151,1436,1634,1692]" pageId="9" pageNumber="10">
<emphasis id="A4E5EAF5C0734F35FF03F9DAFE9EF9F9" bold="true" box="[151,350,1634,1658]" pageId="9" pageNumber="10">FIGURES 2225.</emphasis>
<taxonomicName id="51914D64C0734F35FEF0F9DAFDC3F9FA" box="[356,515,1634,1657]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0734F35FEF0F9DAFDC3F9FA" box="[356,515,1634,1657]" italics="true" pageId="9" pageNumber="10">Baetis zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0734F35FD9EF9DAFD9FF9F9" box="[522,607,1634,1658]" pageId="9" pageNumber="10" rank="species">
<emphasis id="A4E5EAF5C0734F35FD9EF9DAFD9FF9F9" bold="true" box="[522,607,1634,1658]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larvae: 22, surface of femur; 23, outer margin of tibia; 24, inner margin of tibia; 25, tarsal claw (hind legs). Scale bars: 2224 = 200 nm; 25 = 500 nm.
</paragraph>
</caption>
<paragraph id="962E36E7C0734F35FF51F972FCC0F807" blockId="9.[151,1437,1738,2004]" pageId="9" pageNumber="10">
At the
<typeStatus id="492A8845C0734F35FE81F972FE88F967" box="[277,328,1738,1764]" pageId="9" pageNumber="10">type</typeStatus>
locality, larvae of
<taxonomicName id="51914D64C0734F35FDB5F972FD5FF960" box="[545,671,1738,1763]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0734F35FDB5F972FD5FF960" box="[545,671,1738,1763]" italics="true" pageId="9" pageNumber="10">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0734F35FD33F972FCC4F967" box="[679,772,1738,1764]" pageId="9" pageNumber="10" rank="species">
<emphasis id="A4E5EAF5C0734F35FD33F972FCC4F967" bold="true" box="[679,772,1738,1764]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
are clearly dominant forming at least 75% of the mayfly standing crop, they seem to be very rare at the locality of the Argiros Potamos (at most 10% of the mayfly standing crop in May,
<taxonomicName id="51914D64C0734F35FE30F8A2FC89F8B7" authority="Burmeister, 1839" authorityName="Burmeister" authorityYear="1839" box="[420,841,1818,1844]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis id="A4E5EAF5C0734F35FE30F8A2FDA1F8B0" box="[420,609,1818,1843]" italics="true" pageId="9" pageNumber="10">Caenis luctuosa</emphasis>
(
<bibRefCitation id="F2004B16C0734F35FDE6F8A2FC81F8B7" author="Burmeister" box="[626,833,1818,1844]" pageId="9" pageNumber="17" refString="Burmeister, H. (1839) Ephemerina. Handbuch der Entomologie, 2 (2), 788 - 804." type="journal article" year="1839">Burmeister, 1839</bibRefCitation>
)
</taxonomicName>
and a new species of the
<taxonomicName id="51914D64C0734F35FB14F8A2FB30F8B0" box="[1152,1264,1818,1843]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lutheri">
<emphasis id="A4E5EAF5C0734F35FB14F8A2FB30F8B0" box="[1152,1264,1818,1843]" italics="true" pageId="9" pageNumber="10">B. lutheri</emphasis>
</taxonomicName>
species-group were dominant). At the
<typeStatus id="492A8845C0734F35FE3AF8FAFE20F8DF" box="[430,480,1858,1884]" pageId="9" pageNumber="10">type</typeStatus>
locality, the larvae were collected together with those of
<taxonomicName id="51914D64C0734F35FBEBF8FAFAC4F8D8" box="[1151,1284,1858,1883]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis id="A4E5EAF5C0734F35FBEBF8FAFAC4F8D8" box="[1151,1284,1858,1883]" italics="true" pageId="9" pageNumber="10">C. luctuosa</emphasis>
</taxonomicName>
,
<taxonomicName id="51914D64C0734F35FA80F8FAFF0AF800" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fascicaudale">
<emphasis id="A4E5EAF5C0734F35FA80F8FAFF0AF800" italics="true" pageId="9" pageNumber="10">P. fascicaudale</emphasis>
</taxonomicName>
and
<taxonomicName id="51914D64C0734F35FE96F8D2FD3CF807" authority="Pictet, 1843" authorityName="Pictet" authorityYear="1843" box="[258,764,1898,1924]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rhodani" subGenus="Rhodobaetis">
<emphasis id="A4E5EAF5C0734F35FE96F8D2FEDCF800" box="[258,284,1898,1923]" italics="true" pageId="9" pageNumber="10">B.</emphasis>
(
<emphasis id="A4E5EAF5C0734F35FEB8F8D2FE7FF800" box="[300,447,1898,1923]" italics="true" pageId="9" pageNumber="10">Rhodobaetis</emphasis>
) cf.
<emphasis id="A4E5EAF5C0734F35FE61F8D2FD91F800" box="[501,593,1898,1923]" italics="true" pageId="9" pageNumber="10">rhodani</emphasis>
(
<bibRefCitation id="F2004B16C0734F35FDF5F8D2FD33F807" author="Pictet" box="[609,755,1898,1924]" pageId="9" pageNumber="18" refString="Pictet, F. J. (1843 - 1845) Histoire naturelle generale et particuliere des insectes nevropteres. Famille des ephemerines. Chez Kessmann, J. &amp; Cherbuliz, Ab., Geneve, 1 - 300." type="book chapter" year="1843">Pictet, 1843</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="962E36E7C0734F36FF51F82AFBDFFE21" blockId="9.[151,1437,1738,2004]" lastBlockId="10.[151,1437,152,418]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
The life cycle of
<taxonomicName id="51914D64C0734F35FE19F82AFDCBF828" box="[397,523,1938,1963]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0734F35FE19F82AFDCBF828" box="[397,523,1938,1963]" italics="true" pageId="9" pageNumber="10">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0734F35FD86F82AFDAFF82F" box="[530,623,1938,1964]" pageId="9" pageNumber="10" rank="species">
<emphasis id="A4E5EAF5C0734F35FD86F82AFDAFF82F" bold="true" box="[530,623,1938,1964]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
remains unknown in detail. However, judging from the size of larvae in
<date id="E22F1027C0734F35FF21F802FEEEF857" box="[181,302,1978,2004]" pageId="9" pageNumber="10" value="2005-05">May 2005</date>
and timing of mating flight we might suppose a seasonal, bivoltine winter life cycle, the MBws
<typeStatus id="492A8845C0704F36FF03FF20FF09FF31" box="[151,201,152,178]" pageId="10" pageNumber="11">type</typeStatus>
according to the classification by
<bibRefCitation id="F2004B16C0704F36FDCDFF20FCD0FF31" author="Clifford" box="[601,784,152,178]" pageId="10" pageNumber="17" refString="Clifford, H. F. (1982) Life cycles of mayflies (Ephemeroptera), with special reference to voltinism. Quaestiones Entomologicae, 18, 15 - 90." type="journal article" year="1982">Clifford (1982)</bibRefCitation>
. Adult emergence by mid May indicates a possibility of two generations a year (second generation might fly from late August to early October). Owing to favorable conditions (generally mild climate, relatively high water temperatures) there is a chance to complete development of larvae originating from the eggs laid in May of the same year. Taking into account approximately linear growth of larvae during winter followed by its acceleration after spring warming of water, and the different size categories of larvae collected at the same time in May, several egg cohorts are likely, as in numerous other mild climate
<taxonomicName id="51914D64C0704F36FE64FE30FDF9FE22" box="[496,569,392,417]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="A4E5EAF5C0704F36FE64FE30FDF9FE22" box="[496,569,392,417]" italics="true" pageId="10" pageNumber="11">Baetis</emphasis>
</taxonomicName>
species (cf.
<bibRefCitation id="F2004B16C0704F36FD5FFE30FC8FFE21" author="Sowa" box="[715,847,392,418]" pageId="10" pageNumber="18" refString="Sowa, R. (1975) Ecology and biogeography of mayflies (Ephemeroptera) of running waters in the Polish part of the Carpathians. 2. Life cycles. Acta Hydrobiologica, 17, 319 - 353." type="journal article" year="1975">Sowa 1975</bibRefCitation>
or
<bibRefCitation id="F2004B16C0704F36FCE1FE30FBD7FE21" author="Clifford" box="[885,1047,392,418]" pageId="10" pageNumber="17" refString="Clifford, H. F. (1982) Life cycles of mayflies (Ephemeroptera), with special reference to voltinism. Quaestiones Entomologicae, 18, 15 - 90." type="journal article" year="1982">Clifford 1982</bibRefCitation>
)
</paragraph>
<caption id="C2EE666FC0704F36FF03FA78FA57FA79" httpUri="https://zenodo.org/record/185075/files/figure.png" pageId="10" pageNumber="11" targetBox="[151,1435,477,1445]" targetPageId="10">
<paragraph id="962E36E7C0704F36FF03FA78FA57FA79" blockId="10.[151,1437,1472,1530]" pageId="10" pageNumber="11">
<emphasis id="A4E5EAF5C0704F36FF03FA78FE9DFA5B" bold="true" box="[151,349,1472,1496]" pageId="10" pageNumber="11">FIGURES 2629.</emphasis>
<taxonomicName id="51914D64C0704F36FEF0FA78FDC3FA54" box="[356,515,1472,1495]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0704F36FEF0FA78FDC3FA54" box="[356,515,1472,1495]" italics="true" pageId="10" pageNumber="11">Baetis zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0704F36FD9DFA78FD9EFA5B" box="[521,606,1472,1496]" pageId="10" pageNumber="11" rank="species">
<emphasis id="A4E5EAF5C0704F36FD9DFA78FD9EFA5B" bold="true" box="[521,606,1472,1496]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larvae: 26, surface of abdominal tergum IV; 27, surface of abdominal tergum VI; 28, posterior margin of abdominal tergum IV; 29, outer margin of paraproct plate. Scale bars: 2629 = 20mm.
</paragraph>
</caption>
<paragraph id="962E36E7C0704F36FF51F990FCE4F8D9" blockId="10.[151,1436,1576,2002]" pageId="10" pageNumber="11">
Subimagines emerge from the water surface and places overgrown with submerged vegetation during daylight hours, from about 10.00 a.m16.00 p.m. local time.
<taxonomicName id="51914D64C0704F36FCAFF9E8FC2AF9EA" box="[827,1002,1616,1641]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0704F36FCAFF9E8FC2AF9EA" box="[827,1002,1616,1641]" italics="true" pageId="10" pageNumber="11">Baetis zdenkae</emphasis>
</taxonomicName>
<emphasis id="A4E5EAF5C0704F36FC66F9E8FB8FF9E9" bold="true" box="[1010,1103,1616,1642]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="BFD6578EC0704F36FC66F9E8FB8FF9E9" box="[1010,1103,1616,1642]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
showed early morning mating flight activity with the mating flight generally occurring from about 6.308.30 a.m. local time. First males started to appear approximately a half hour before sunrise, and showed a typical “pendular” flight pattern usually in groups of about 310 individuals, females were observed about 0.51 hour later. Although only a few females were observed flying upstream, their flight activity seemed to last at least one hour after males had disappeared. A single female was observed ovipositing on a submerged stone, hence the way of oviposition seems to be similar to other species of the genus
<taxonomicName id="51914D64C0704F36FD40F8F8FCDDF8DA" box="[724,797,1856,1881]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="A4E5EAF5C0704F36FD40F8F8FCDDF8DA" box="[724,797,1856,1881]" italics="true" pageId="10" pageNumber="11">Baetis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="962E36E7C0704F37FF51F8D0FD1AFE4F" blockId="10.[151,1436,1576,2002]" lastBlockId="11.[151,1437,151,1980]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Generally, the biology of
<taxonomicName id="51914D64C0704F36FD96F8D0FD45F802" box="[514,645,1896,1921]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0704F36FD96F8D0FD45F802" box="[514,645,1896,1921]" italics="true" pageId="10" pageNumber="11">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0704F36FD04F8D0FD31F801" box="[656,753,1896,1922]" pageId="10" pageNumber="11" rank="species">
<emphasis id="A4E5EAF5C0704F36FD04F8D0FD31F801" bold="true" box="[656,753,1896,1922]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
resembles that of
<taxonomicName id="51914D64C0704F36FC4DF8D0FBECF802" box="[985,1068,1896,1921]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0704F36FC4DF8D0FBECF802" box="[985,1068,1896,1921]" italics="true" pageId="10" pageNumber="11">B. spei</emphasis>
</taxonomicName>
which was characterized as a rhithro-potamal species by
<bibRefCitation id="F2004B16C0704F36FE4DF828FD35F829" author="Thomas" box="[473,757,1936,1962]" pageId="10" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas and Dia (1985)</bibRefCitation>
, while the remaining representatives of
<taxonomicName id="51914D64C0704F36FB5AF828FAA2F82A" box="[1230,1378,1936,1961]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0704F36FB5AF828FAA2F82A" box="[1230,1378,1936,1961]" italics="true" pageId="10" pageNumber="11">B. buceratus</emphasis>
</taxonomicName>
species-group occur at rather different (mostly potamal) habitats. However, larvae of
<taxonomicName id="51914D64C0704F36FBEBF800FB12F852" box="[1151,1234,1976,2001]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0704F36FBEBF800FB12F852" box="[1151,1234,1976,2001]" italics="true" pageId="10" pageNumber="11">B. spei</emphasis>
</taxonomicName>
showed a wider range of vertical distribution than those of
<taxonomicName id="51914D64C0714F37FD13FF20FCC5FF32" box="[647,773,152,177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0714F37FD13FF20FCC5FF32" box="[647,773,152,177]" italics="true" pageId="11" pageNumber="12">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0714F37FC98FF2FFCA9FF32" box="[780,873,151,177]" pageId="11" pageNumber="12" rank="species">
<emphasis id="A4E5EAF5C0714F37FC98FF2FFCA9FF32" bold="true" box="[780,873,151,177]" pageId="11" pageNumber="12">sp. nov.</emphasis>
</taxonomicNameLabel>
being collected at altitudes of
<quantity id="51699B02C0714F37FB5AFF20FAF5FF31" box="[1230,1333,152,178]" metricMagnitude="2" metricUnit="m" metricValue="4.775" metricValueMax="9.5" metricValueMin="0.05" pageId="11" pageNumber="12" unit="m" value="477.5" valueMax="950.0" valueMin="5.0">5950 m</quantity>
a. s. l. at places
<quantity id="51699B02C0714F37FF73FF78FE8DFF59" box="[231,333,192,218]" metricMagnitude="4" metricUnit="m" metricValue="2.5" metricValueMax="4.8" metricValueMin="0.2" pageId="11" pageNumber="12" unit="km" value="25.0" valueMax="48.0" valueMin="2.0">248 km</quantity>
from source in well oxygenated water. They were moderately abundant to abundant at streams about
<quantity id="51699B02C0714F37FF75FF50FEF3FE81" box="[225,307,232,258]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="11" pageNumber="12" unit="m" value="6.5" valueMax="10.0" valueMin="3.0">310m</quantity>
across, including also a temporary stream. The larvae were characterized as thermophilous (in contrast to those of
<taxonomicName id="51914D64C0714F37FE14FEA8FD16FEA9" authority="Thomas &amp; Dia, 1983" authorityName="Thomas &amp; Dia" authorityYear="1983" box="[384,726,272,298]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bisri">
<emphasis id="A4E5EAF5C0714F37FE14FEA8FE17FEAA" box="[384,471,272,297]" italics="true" pageId="11" pageNumber="12">B. bisri</emphasis>
<bibRefCitation id="F2004B16C0714F37FE4AFEA8FD16FEA9" author="Thomas" box="[478,726,272,298]" pageId="11" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1983) Baetis bisri n. sp., Ephemeroptere nouveau du Liban (Baetidae). Annales de Limnologie, 19 (3), 213 - 217." type="journal article" year="1983">Thomas &amp; Dia, 1983</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="51914D64C0714F37FC84FEA8FB04FEA9" authority="Thomas &amp; Dia, 1984" authorityName="Thomas &amp; Dia" authorityYear="1984" box="[784,1220,272,298]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="baroukianus">
<emphasis id="A4E5EAF5C0714F37FC84FEA8FC02FEAA" box="[784,962,272,297]" italics="true" pageId="11" pageNumber="12">B. baroukianus</emphasis>
<bibRefCitation id="F2004B16C0714F37FC5FFEA8FB04FEA9" author="Thomas" box="[971,1220,272,298]" pageId="11" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1984) Baetis baroukianus n. sp., Ephemeroptere nouveau du Liban (Baetidae). Bulletin de la Societe de Histoire Naturelle Toulouse, 120, 7 - 11." type="journal article" year="1984">Thomas &amp; Dia, 1984</bibRefCitation>
</taxonomicName>
living at the same area (
<bibRefCitation id="F2004B16C0714F37FF43FE80FE0EFED1" author="Thomas" box="[215,462,312,338]" pageId="11" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1983) Baetis bisri n. sp., Ephemeroptere nouveau du Liban (Baetidae). Annales de Limnologie, 19 (3), 213 - 217." type="journal article" year="1983">Thomas &amp; Dia 1983</bibRefCitation>
,
<bibRefCitation id="F2004B16C0714F37FE4FFE80FDDAFED1" author="Thomas" box="[475,538,312,338]" pageId="11" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1984) Baetis baroukianus n. sp., Ephemeroptere nouveau du Liban (Baetidae). Bulletin de la Societe de Histoire Naturelle Toulouse, 120, 7 - 11." type="journal article" year="1984">1984</bibRefCitation>
), maximal water temperature ranged from 1729 ºC (pH 7.48.25), conductivity 290440 µS.cm -1. The life cycle of
<taxonomicName id="51914D64C0714F37FDECFEDBFD07FEFF" box="[632,711,355,380]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0714F37FDECFEDBFD07FEFF" box="[632,711,355,380]" italics="true" pageId="11" pageNumber="12">B. spei</emphasis>
</taxonomicName>
is not fully known, mature larvae were collected from May to August (
<bibRefCitation id="F2004B16C0714F37FF69FE32FE39FE27" author="Thomas" box="[253,505,394,420]" pageId="11" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas &amp; Dia 1985</bibRefCitation>
), were collected from May to August (
<bibRefCitation id="F2004B16C0714F37FC59FE32FB09FE27" author="Thomas" box="[973,1225,394,420]" pageId="11" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas &amp; Dia 1985</bibRefCitation>
), and adults were collected at the same time (
<bibRefCitation id="F2004B16C0714F37FE42FE0AFD0DFE4F" author="Thomas" box="[470,717,434,460]" pageId="11" pageNumber="19" refString="Thomas, A. &amp; Dia, A. (2007) Complements et corrections a la faune des Ephemeropteres du Proche-Orient. 7. Description de l'imago male de Baetis spei Thomas, 1985 [Ephemeroptera, Baetidae]. Ephemera, 8 (2): 75 - 78." type="journal article" year="2007">Thomas &amp; Dia 2007</bibRefCitation>
).
</paragraph>
<paragraph id="962E36E7C0714F37FF51FE62FBEFFC57" blockId="11.[151,1437,151,1980]" pageId="11" pageNumber="12">
Larvae of
<taxonomicName id="51914D64C0714F37FEA9FE63FE10FE77" box="[317,464,475,500]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0714F37FEA9FE63FE10FE77" box="[317,464,475,500]" italics="true" pageId="11" pageNumber="12">B. buceratus</emphasis>
</taxonomicName>
have been found mostly at the localities of large lowland rivers (epipotamal). They apparently can tolerate a certain degree of pollution being sometimes dominant at these biotopes. Larvae prefer medium to high current speed and stony substrata (gravel and larger stones) but often occur also among submerged plants. Such habitats are described as typical for this species (
<bibRefCitation id="F2004B16C0714F37FB97FDEAFAE1FDEF" author="Buffagni" box="[1027,1313,594,620]" pageId="11" pageNumber="17" refString="Buffagni, A. &amp; Desio, F. (1994) Le specie del genere Baetis LEACH (Ephemeroptera: Baetidae) nell'Italia settentrionale: sintesi ecologia. In: Atti XVII Congresso Ital. entomologia, Undine, pp. 413 - 416." type="book chapter" year="1994">Buffagni &amp; Desio 1994</bibRefCitation>
; Gonzáles del
<bibRefCitation id="F2004B16C0714F37FF56FDC2FEA1FD17" author="Tanago" box="[194,353,634,660]" pageId="11" pageNumber="17" refString="Gonzales del Tanago, M. (1984) Distribucion y biologia de la familia Baetidae (Ephem.) en la Cuenca del Duero. Bolletino de la Asociation de espanol Entomologia, 8, 73 - 94." type="journal article" year="1984">Tánago 1984</bibRefCitation>
).
<taxonomicName id="51914D64C0714F37FEE1FDC3FDCAFD17" box="[373,522,635,660]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0714F37FEE1FDC3FDCAFD17" box="[373,522,635,660]" italics="true" pageId="11" pageNumber="12">B. buceratus</emphasis>
</taxonomicName>
shows a typical bivoltine winter life cycle (MBws) with overwintering larvae of the first generation. However, there are some data indicating plasticity of the cycle. In
<collectingCountry id="EE867677C0714F37FB2DFD1AFA88FD3F" box="[1209,1352,674,700]" name="Switzerland" pageId="11" pageNumber="12">Switzerland</collectingCountry>
, larvae live also in epirhithral stream segments although they prefer lower elevations and places with lower current velocity at this area (
<bibRefCitation id="F2004B16C0714F37FE1AFD4AFD69FC8F" author="Sartori" box="[398,681,754,780]" pageId="11" pageNumber="18" refString="Sartori, M. &amp; Landolt, P. (1999) Atlas de distribution des Ephemeres de Suisse (Insecta, Ephemeroptera). Fauna helvetica, 3, SEG - CSCF ed., Neuchatel, 214 pp." type="book" year="1999">Sartori &amp; Landolt 1999</bibRefCitation>
). According to
<bibRefCitation id="F2004B16C0714F37FCF5FD4AFBC1FC8F" author="Landa" box="[865,1025,754,780]" pageId="11" pageNumber="18" refString="Landa, V. (1968) Developmental cycles of Central European Ephemeroptera and their interrelations. Acta Entomologica Bohemoslovaca, 65, 276 - 284." type="journal article" year="1968">
<collectingCountry id="EE867677C0714F37FCF5FD4AFC6AFC8F" box="[865,938,754,780]" name="Netherlands" pageId="11" pageNumber="12">Landa</collectingCountry>
(1968)
</bibRefCitation>
older larvae of the first generation appear from March to May this may indicate ceasing of larval development during winter months rarely found in species with the MBws life cycle
<typeStatus id="492A8845C0714F37FD04FCFAFD06FCDF" box="[656,710,834,860]" pageId="11" pageNumber="12">type</typeStatus>
. According to
<bibRefCitation id="F2004B16C0714F37FCE0FCFAFBEAFCDF" author="Haybach" box="[884,1066,834,860]" pageId="11" pageNumber="17" refString="Haybach, A. (1998) Die Eintagsfliegen (Insecta: Ephemeroptera) von Rheinland - Pfalz - Zoogeographie, Faunistik, Okologie, Taxonomie und Nomenklatur - Unter besonderer Berucksichtigung der Familie Heptageniidae und unter Einbeziehung die ubrigen aus Deutschland bekannten Arten. Dissertation am Fachbereich Biologie der Johannes Gutenberg-Universitat Mainz, 417 pp." type="book" year="1998">Haybach (1998</bibRefCitation>
,
<bibRefCitation id="F2004B16C0714F37FBA3FCFAFBB3FCDF" author="Haybach" box="[1079,1139,834,860]" pageId="11" pageNumber="17" refString="Haybach, A. (2006) Die Eintagsfliegen von Reinland-Pfalz (Insecta Ephemeroptera). Mainzer Naturwissenschaftliches Archiv, Naturhistorisches Museum Mainz / Landessammlung fur Naturkunde Rheiland-Pfalz, 29, 121 pp." type="book" year="2006">2006</bibRefCitation>
) there is one overwintering generation with flight period in spring and two generations following late summer and autumn. Since the adults apparently fly from July to October (
<bibRefCitation id="F2004B16C0714F37FD02FC2AFCE7FC2F" author="Landa" box="[662,807,914,940]" pageId="11" pageNumber="18" refString="Landa, V. (1968) Developmental cycles of Central European Ephemeroptera and their interrelations. Acta Entomologica Bohemoslovaca, 65, 276 - 284." type="journal article" year="1968">
<collectingCountry id="EE867677C0714F37FD02FC2AFD21FC2F" box="[662,737,914,940]" name="Netherlands" pageId="11" pageNumber="12">Landa</collectingCountry>
1968
</bibRefCitation>
;
<bibRefCitation id="F2004B16C0714F37FCA0FC2AFC27FC2F" author="Langford" box="[820,999,914,940]" pageId="11" pageNumber="18" refString="Langford, T. E. (1971) The distribution, abundance and life histories of stoneflies (Plecoptera) and mayflies (Ephemeroptera) in a British river, warmed by cooling-water from a power station. Hydrobiologia, 38 (2), 339 - 377." type="journal article" year="1971">Langford 1971</bibRefCitation>
;
<bibRefCitation id="F2004B16C0714F37FC67FC2AFBBAFC2F" author="Sowa" box="[1011,1146,914,940]" pageId="11" pageNumber="18" refString="Sowa, R. (1975) Ecology and biogeography of mayflies (Ephemeroptera) of running waters in the Polish part of the Carpathians. 2. Life cycles. Acta Hydrobiologica, 17, 319 - 353." type="journal article" year="1975">Sowa 1975</bibRefCitation>
), two “summer” generations and, consequently, some
<typeStatus id="492A8845C0714F37FE68FC02FDEEFC57" box="[508,558,954,980]" pageId="11" pageNumber="12">type</typeStatus>
of polyvoltinism (MP) cannot be excluded.
</paragraph>
<paragraph id="962E36E7C0714F37FF51FC5AFD6BFAE7" blockId="11.[151,1437,151,1980]" pageId="11" pageNumber="12">
Larvae of
<taxonomicName id="51914D64C0714F37FEABFC5BFE62FC7F" box="[319,418,995,1020]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0714F37FEABFC5BFE62FC7F" box="[319,418,995,1020]" italics="true" pageId="11" pageNumber="12">B. nexus</emphasis>
</taxonomicName>
live in lowland waters of different size, being mostly found in rivers, their backwaters and artificial freshwater biotopes (e.g., in the “roach zone”), at places with very low current velocity. They evidently prefer places rich in submerged aquatic vegetation. Azonal species according to
<bibRefCitation id="F2004B16C0714F37FB4AFB8AFA57FBCF" author="Haybach" box="[1246,1431,1074,1100]" pageId="11" pageNumber="17" refString="Haybach, A. (1998) Die Eintagsfliegen (Insecta: Ephemeroptera) von Rheinland - Pfalz - Zoogeographie, Faunistik, Okologie, Taxonomie und Nomenklatur - Unter besonderer Berucksichtigung der Familie Heptageniidae und unter Einbeziehung die ubrigen aus Deutschland bekannten Arten. Dissertation am Fachbereich Biologie der Johannes Gutenberg-Universitat Mainz, 417 pp." type="book" year="1998">Haybach (1998</bibRefCitation>
,
<bibRefCitation id="F2004B16C0714F37FF03FBE2FF16FBF7" author="Haybach" box="[151,214,1114,1140]" pageId="11" pageNumber="17" refString="Haybach, A. (2006) Die Eintagsfliegen von Reinland-Pfalz (Insecta Ephemeroptera). Mainzer Naturwissenschaftliches Archiv, Naturhistorisches Museum Mainz / Landessammlung fur Naturkunde Rheiland-Pfalz, 29, 121 pp." type="book" year="2006">2006</bibRefCitation>
), habitat preference cannot be evaluated from the rhithron-potamon concept. However, in south Moravia (
<collectingCountry id="EE867677C0714F37FF0BFB3AFEA4FB1F" box="[159,356,1154,1180]" name="Czech Republic" pageId="11" pageNumber="12">Czech Republic</collectingCountry>
), the larvae are usually found in a typical epipotamal river, moreover influenced by the upstream impoundment (
<bibRefCitation id="F2004B16C0714F37FE28FB12FD97FB47" author="Horsak" box="[444,599,1194,1220]" pageId="11" pageNumber="17" refString="Horsak, M. (2001) Contribution to the knowledge of macroinvertebrate fauna of the Dyje River downstream of the Nove Mlyny Reservoirs (Czech Republic). Scripta Facultatis scientiarum naturalium Universitatis Masarykianae brunensis, Biologia, Supplementum, 27, 41 - 62." type="journal article" year="2001">Horsák 2001</bibRefCitation>
) like they do in southern
<collectingCountry id="EE867677C0714F37FC12FB12FC16FB47" box="[902,982,1194,1220]" name="France" pageId="11" pageNumber="12">France</collectingCountry>
(Cammouseight &amp;
<bibRefCitation id="F2004B16C0714F37FB2AFB12FAAAFB47" author="Fontaine" box="[1214,1386,1194,1220]" pageId="11" pageNumber="17" refString="Camousseight, A. &amp; Fontaine, J. (1990) The biological cycle of Baetis pentaphlebodes Ujhelyi 1966, in an old meander of the Rhone River, France (Ephemeroptera: Baetidae). In: Campbell, I. C. (Ed.), Mayflies and Stoneflies: Life Histories and Biology, Kluwer Academic Publishers, Dordrecht (Netherlands), pp. 27 - 34." type="book chapter" year="1990">Fontaine 1990</bibRefCitation>
, see this paper also for further details on ecological requirements of larvae and their quantitative presentation). Seasonal bivoltine winter life cycles (MBws) have been observed in
<collectingCountry id="EE867677C0714F37FC7BFB42FBFFFA97" box="[1007,1087,1274,1300]" name="France" pageId="11" pageNumber="12">France</collectingCountry>
(Cammouseight &amp;
<bibRefCitation id="F2004B16C0714F37FAA2FB42FF16FABF" author="Fontaine" pageId="11" pageNumber="17" refString="Camousseight, A. &amp; Fontaine, J. (1990) The biological cycle of Baetis pentaphlebodes Ujhelyi 1966, in an old meander of the Rhone River, France (Ephemeroptera: Baetidae). In: Campbell, I. C. (Ed.), Mayflies and Stoneflies: Life Histories and Biology, Kluwer Academic Publishers, Dordrecht (Netherlands), pp. 27 - 34." type="book chapter" year="1990">Fontaine 1990</bibRefCitation>
), with two flight periods in May and August, and also in
<collectingCountry id="EE867677C0714F37FC12FA9AFC2CFABF" box="[902,1004,1314,1340]" name="Hungary" pageId="11" pageNumber="12">Hungary</collectingCountry>
(
<bibRefCitation id="F2004B16C0714F37FC69FA9AFB5EFABF" author="Ujhelyi" box="[1021,1182,1314,1340]" pageId="11" pageNumber="19" refString="Ujhelyi, S. (1966) The mayflies of Hungary, with the description of a new species, Baetis pentaphlebodes sp. n. (Ephemeroptera). Acta Zoologica Academiae Scientiarum Hungaricae, 12 (1 - 2), 203 - 210." type="journal article" year="1966">Ujhelyi 1966</bibRefCitation>
),
<collectingCountry id="EE867677C0714F37FB20FA9AFAE1FABF" box="[1204,1313,1314,1340]" name="Germany" pageId="11" pageNumber="12">Germany</collectingCountry>
(
<bibRefCitation id="F2004B16C0714F37FAA6FA9AFF14FAE7" author="Haybach" pageId="11" pageNumber="17" refString="Haybach, A. (1998) Die Eintagsfliegen (Insecta: Ephemeroptera) von Rheinland - Pfalz - Zoogeographie, Faunistik, Okologie, Taxonomie und Nomenklatur - Unter besonderer Berucksichtigung der Familie Heptageniidae und unter Einbeziehung die ubrigen aus Deutschland bekannten Arten. Dissertation am Fachbereich Biologie der Johannes Gutenberg-Universitat Mainz, 417 pp." type="book" year="1998">Haybach 1998</bibRefCitation>
,
<bibRefCitation id="F2004B16C0714F37FF74FAF2FEDCFAE7" author="Haybach" box="[224,284,1354,1380]" pageId="11" pageNumber="17" refString="Haybach, A. (2006) Die Eintagsfliegen von Reinland-Pfalz (Insecta Ephemeroptera). Mainzer Naturwissenschaftliches Archiv, Naturhistorisches Museum Mainz / Landessammlung fur Naturkunde Rheiland-Pfalz, 29, 121 pp." type="book" year="2006">2006</bibRefCitation>
) and
<collectingCountry id="EE867677C0714F37FEC9FAF2FE6EFAE7" box="[349,430,1354,1380]" name="Poland" pageId="11" pageNumber="12">Poland</collectingCountry>
(
<bibRefCitation id="F2004B16C0714F37FE2AFAF2FD5FFAE7" author="Keffermuller" box="[446,671,1354,1380]" pageId="11" pageNumber="17" refString="Keffermuller, M. (1972) Investigations on the mayflies (Ephemeroptera) fauna in Great Poland. V. Polskie Pismo Entomologiczne, 43, 527 - 533. (in Polish, Engl. sum.)" type="journal article" year="1972">Keffermüller 1972</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DE8B656CC0714F30FF51FACAFA4AFF5A" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="distribution">
<paragraph id="962E36E7C0714F37FF51FACAFDD3F927" blockId="11.[151,1437,151,1980]" pageId="11" pageNumber="12">
<emphasis id="A4E5EAF5C0714F37FF51FACAFE9FFA0F" bold="true" box="[197,351,1394,1420]" italics="true" pageId="11" pageNumber="12">Distribution.</emphasis>
Since
<taxonomicName id="51914D64C0714F37FE26FACBFDF1FA0F" box="[434,561,1395,1420]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0714F37FE26FACBFDF1FA0F" box="[434,561,1395,1420]" italics="true" pageId="11" pageNumber="12">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0714F37FDAFFACAFD59FA0F" box="[571,665,1394,1420]" pageId="11" pageNumber="12" rank="species">
<emphasis id="A4E5EAF5C0714F37FDAFFACAFD59FA0F" bold="true" box="[571,665,1394,1420]" pageId="11" pageNumber="12">sp. nov.</emphasis>
</taxonomicNameLabel>
is so far known only from Rhodos Island it might be considered endemic to this island. However, the true distribution of
<taxonomicName id="51914D64C0714F37FCB3FA23FC65FA37" box="[807,933,1435,1460]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0714F37FCB3FA23FC65FA37" box="[807,933,1435,1460]" italics="true" pageId="11" pageNumber="12">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0714F37FC38FA22FBC9FA37" box="[940,1033,1434,1460]" pageId="11" pageNumber="12" rank="species">
<emphasis id="A4E5EAF5C0714F37FC38FA22FBC9FA37" bold="true" box="[940,1033,1434,1460]" pageId="11" pageNumber="12">sp. nov.</emphasis>
</taxonomicNameLabel>
still remains poorly known so that its classification is preliminary since these possible “endemics” might inhabit a much larger area as shown, for example, by our finding of presumably “endemic” species
<taxonomicName id="51914D64C0714F37FCC9FA53FC12F987" box="[861,978,1515,1540]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mirkae">
<emphasis id="A4E5EAF5C0714F37FCC9FA53FC12F987" box="[861,978,1515,1540]" italics="true" pageId="11" pageNumber="12">B. mirkae</emphasis>
</taxonomicName>
group described from
<collectingCountry id="EE867677C0714F37FB73FA52FAFCF987" box="[1255,1340,1514,1540]" name="Cyprus" pageId="11" pageNumber="12">Cyprus</collectingCountry>
also on Rhodos (
<bibRefCitation id="F2004B16C0714F37FE96F9AAFDFDF9AF" author="Soldan" box="[258,573,1554,1580]" pageId="11" pageNumber="18" refString="Soldan, T. &amp; Godunko, R. J. (2008) Two new species of the genus Baetis Laech, 1815 (Ephemeroptera: Baetidae) from Cyprus. Annales Zoologici, 58 (1): 79 - 104." type="journal article" year="2008">Soldán &amp; Godunko, 2008</bibRefCitation>
). That is why we prefer to wait until further collection records are published. The distribution of
<taxonomicName id="51914D64C0714F37FE58F983FD8AF9D7" box="[460,586,1595,1620]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0714F37FE58F983FD8AF9D7" box="[460,586,1595,1620]" italics="true" pageId="11" pageNumber="12">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0714F37FDC5F982FD6EF9D7" box="[593,686,1594,1620]" pageId="11" pageNumber="12" rank="species">
<emphasis id="A4E5EAF5C0714F37FDC5F982FD6EF9D7" bold="true" box="[593,686,1594,1620]" pageId="11" pageNumber="12">sp. nov.</emphasis>
</taxonomicNameLabel>
can be compared to that of
<taxonomicName id="51914D64C0714F37FC7BF983FBFEF9D7" box="[1007,1086,1595,1620]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="spei">
<emphasis id="A4E5EAF5C0714F37FC7BF983FBFEF9D7" box="[1007,1086,1595,1620]" italics="true" pageId="11" pageNumber="12">B. spei</emphasis>
</taxonomicName>
, the species known only from
<collectingCountry id="EE867677C0714F37FF03F9DAFEC1F9FF" box="[151,257,1634,1660]" name="Lebanon" pageId="11" pageNumber="12">Lebanon</collectingCountry>
, only from two river basins (Nahr el Aouali and Nahr ed Damour) as well, so far altogether from 9 localities (
<bibRefCitation id="F2004B16C0714F37FE84F932FDC6F927" author="Thomas" box="[272,518,1674,1700]" pageId="11" pageNumber="19" refString="Thomas, A. G. B. &amp; Dia, A. (1985) Baetis spei n. sp., Ephemeroptere nouveau du Liban: description et ecologie (Baetidae). Annales de Limnologie, 21 (3), 241 - 245." type="journal article" year="1985">Thomas &amp; Dia 1985</bibRefCitation>
).
</paragraph>
<paragraph id="962E36E7C0714F30FF51F90AFA4AFF5A" blockId="11.[151,1437,151,1980]" lastBlockId="12.[151,1436,152,458]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
On the other hand the distribution of
<taxonomicName id="51914D64C0714F37FDE3F90BFD36F94F" box="[631,758,1715,1740]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0714F37FDE3F90BFD36F94F" box="[631,758,1715,1740]" italics="true" pageId="11" pageNumber="12">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0714F37FD69F90AFC9AF94F" box="[765,858,1714,1740]" pageId="11" pageNumber="12" rank="species">
<emphasis id="A4E5EAF5C0714F37FD69F90AFC9AF94F" bold="true" box="[765,858,1714,1740]" pageId="11" pageNumber="12">sp. nov.</emphasis>
</taxonomicNameLabel>
seems to be quite different from the distributions of
<taxonomicName id="51914D64C0714F37FF2DF963FE90F977" box="[185,336,1755,1780]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0714F37FF2DF963FE90F977" box="[185,336,1755,1780]" italics="true" pageId="11" pageNumber="12">B. buceratus</emphasis>
</taxonomicName>
and
<taxonomicName id="51914D64C0714F37FE1AF963FE35F977" box="[398,501,1755,1780]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0714F37FE1AF963FE35F977" box="[398,501,1755,1780]" italics="true" pageId="11" pageNumber="12">B. nexus</emphasis>
</taxonomicName>
. The former species (
<typeStatus id="492A8845C0714F37FD6AF962FCF2F977" box="[766,818,1754,1780]" pageId="11" pageNumber="12">type</typeStatus>
locality the Kenneth and Holybrook near Reading, Berkshire,
<collectingCountry id="EE867677C0714F37FE83F8BAFEBBF89F" box="[279,379,1794,1820]" name="United Kingdom" pageId="11" pageNumber="12">England</collectingCountry>
) belongs to widely distributed Westpalaearctic species. It is known from Europe including the British Islands and some Mediterranean islands, e.g., Sardinia (
<bibRefCitation id="F2004B16C0714F37FC35F892FB72F8C7" author="Belfiore" box="[929,1202,1834,1860]" pageId="11" pageNumber="16" refString="Belfiore, C. &amp; Gaino, E. (1988) Il popolamento a Efemerotteri della Sardegna (Ephemeroptera). Bolletino della Societa Entomologica Italiana, 20 (2), 75 - 83." type="journal article" year="1988">Belfiore &amp; Gaino 1988</bibRefCitation>
) but probably missing in North Africa. Northern limits are situated in Scandinavia
<collectingCountry id="EE867677C0714F37FC2DF8EAFBD6F8EF" box="[953,1046,1874,1900]" name="Sweden" pageId="11" pageNumber="12">Sweden</collectingCountry>
and
<collectingCountry id="EE867677C0714F37FBC0F8EAFB01F8EF" box="[1108,1217,1874,1900]" name="Denmark" pageId="11" pageNumber="12">Denmark</collectingCountry>
(
<bibRefCitation id="F2004B16C0714F37FB47F8EAFA57F8EF" author="Bengtsson" box="[1235,1431,1874,1900]" pageId="11" pageNumber="17" refString="Bengtsson, S. (1917) Weitere Beitrage zur Kenntnis der nordischen Eintagsfliegen. Entomologisk Tidskrift, 38, 174 - 194." type="journal article" year="1917">Bengtsson 1917</bibRefCitation>
;
<bibRefCitation id="F2004B16C0714F37FF03F8C2FEEBF817" author="Jensen" box="[151,299,1914,1940]" pageId="11" pageNumber="17" refString="Jensen, C. F. (1969) Baetis buceratus EAT. og Metretopus borealis EAT. nye for Danmark Ephemeroptera-Dognfluer). Flora og Fauna, 75, 129 - 134." type="journal article" year="1969">Jensen 1969</bibRefCitation>
), southeastern ones in
<collectingCountry id="EE867677C0714F37FDA0F8C2FD7CF817" box="[564,700,1914,1940]" name="Turkey" pageId="11" pageNumber="12">Asia Minor</collectingCountry>
,
<collectingCountry id="EE867677C0714F37FD5CF8C2FCDBF817" box="[712,795,1914,1940]" name="Turkey" pageId="11" pageNumber="12">Turkey</collectingCountry>
and
<collectingCountry id="EE867677C0714F37FCC0F8C2FC43F817" box="[852,899,1914,1940]" name="Iraq" pageId="11" pageNumber="12">Iraq</collectingCountry>
(
<bibRefCitation id="F2004B16C0714F37FC07F8C2FBDAF817" author="Koch" box="[915,1050,1914,1940]" pageId="11" pageNumber="18" refString="Koch, S. (1988) Mayflies of the northern Levant (Insecta: Ephemeroptera). Zoology in the Middle East, 2, 89 - 112." type="journal article" year="1988">Koch 1988</bibRefCitation>
;
<bibRefCitation id="F2004B16C0714F37FBB2F8C2FB0BF817" author="Kazanci" box="[1062,1227,1914,1940]" pageId="11" pageNumber="17" refString="Kazanci, N. (1984) New Ephemeroptera (Insecta) from Turkey. Aquatic Insects, 6, 253 - 258." type="journal article" year="1984">Kazanci 1984</bibRefCitation>
;
<bibRefCitation id="F2004B16C0714F37FB43F8C2FF15F83F" author="Al-Zubaidi" pageId="11" pageNumber="16" refString="Al-Zubaidi, F., Braasch, D. &amp; Al-Kayatt, A. (1987) Mayflies from Iraq (Insecta, Ephemeroptera). Faunistische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 14, 179 - 184." type="journal article" year="1987">
Al-Zubaidi
<emphasis id="A4E5EAF5C0714F37FAF5F8C3FA5CF817" box="[1377,1436,1915,1940]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
1987
</bibRefCitation>
), eastern limits probably in southern Ural, Caucasus and Central Asia (cf.
<bibRefCitation id="F2004B16C0714F37FBA8F81AFB09F83F" author="Kluge" box="[1084,1225,1954,1980]" pageId="11" pageNumber="17" refString="Kluge, N. J. (1997). Order Ephemeroptera. In: Tsalolikhin, S. J. (Ed.). Key to Freshwater Invertebrates of Russia and adjacent lands, Vol. 3, Zoological Institute of the Russian Academy of Sciences, St. Petersburg, pp. 175 - 220, Tab. 8 - 20 (304 - 329). (in Russian)" type="book chapter" year="1997">Kluge 1997</bibRefCitation>
).
<taxonomicName id="51914D64C0714F37FB4AF81BFAB2F83F" box="[1246,1394,1955,1980]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="A4E5EAF5C0714F37FB4AF81BFAB2F83F" box="[1246,1394,1955,1980]" italics="true" pageId="11" pageNumber="12">B. buceratus</emphasis>
</taxonomicName>
is classified either as a Holomediterranean faunistic element of the expansive
<typeStatus id="492A8845C0764F30FB9FFF20FBFEFF31" box="[1035,1086,152,178]" pageId="12" pageNumber="13">type</typeStatus>
according to
<bibRefCitation id="F2004B16C0764F30FB49FF20FAB8FF31" author="Jacob" box="[1245,1400,152,178]" pageId="12" pageNumber="17" refString="Jacob, U. (1972) Beitrag zur autochthonen Ephemeropterenfauna in der Deutschen Demokratischen Republik. Dissertation Universitat Leipzig, Leipzig, 158 pp." type="book" year="1972">Jacob (1972)</bibRefCitation>
, or a Pontokaspian element according to
<bibRefCitation id="F2004B16C0764F30FDD9FF78FCCBFF59" author="Haybach" box="[589,779,192,218]" pageId="12" pageNumber="17" refString="Haybach, A. (1998) Die Eintagsfliegen (Insecta: Ephemeroptera) von Rheinland - Pfalz - Zoogeographie, Faunistik, Okologie, Taxonomie und Nomenklatur - Unter besonderer Berucksichtigung der Familie Heptageniidae und unter Einbeziehung die ubrigen aus Deutschland bekannten Arten. Dissertation am Fachbereich Biologie der Johannes Gutenberg-Universitat Mainz, 417 pp." type="book" year="1998">Haybach (1998)</bibRefCitation>
and its area is sympatric to that of
<taxonomicName id="51914D64C0764F30FB32FF78FAE5FF5A" box="[1190,1317,192,217]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="zdenkae" status="sp. nov.">
<emphasis id="A4E5EAF5C0764F30FB32FF78FAE5FF5A" box="[1190,1317,192,217]" italics="true" pageId="12" pageNumber="13">B. zdenkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFD6578EC0764F30FAB8FF07FA4AFF5A" box="[1324,1418,191,217]" pageId="12" pageNumber="13" rank="species">
<emphasis id="A4E5EAF5C0764F30FAB8FF07FA4AFF5A" bold="true" box="[1324,1418,191,217]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DE8B656CC0764F30FF51FF50FCCFFE49" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="962E36E7C0764F30FF51FF50FCCFFE49" blockId="12.[151,1436,152,458]" pageId="12" pageNumber="13">
<taxonomicName id="51914D64C0764F30FF51FF50FE98FE82" ID-CoL="KCZM" box="[197,344,232,257]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nexus">
<emphasis id="A4E5EAF5C0764F30FF51FF50FE98FE82" box="[197,344,232,257]" italics="true" pageId="12" pageNumber="13">Baetis nexus</emphasis>
</taxonomicName>
belongs to widely distributed Westpalaearctic species as well, however its area is restricted to about 50º N. Northern limits are situated in
<collectingCountry id="EE867677C0764F30FD09FEA8FCCAFEA9" box="[669,778,272,298]" name="Germany" pageId="12" pageNumber="13">Germany</collectingCountry>
and
<collectingCountry id="EE867677C0764F30FCD1FEA8FC5AFEA9" box="[837,922,272,298]" name="Poland" pageId="12" pageNumber="13">Poland</collectingCountry>
. This species is known from Europe except for the British Islands, Fennoscandia and most of the Mediterranean, and is missing in North Africa (see
<bibRefCitation id="F2004B16C0764F30FAFFFE80FE4FFEF9" author="Sartori" pageId="12" pageNumber="18" refString="Sartori, M. &amp; Soldan, T. (2006) Case 3322. Baetis pentaphlebodes Ujhelyi, 1966 (Insecta, Ephemeroptera) proposed precedence over Baetis nexus Navas, 1918. Bulletin of Zoological Nomenclature, 63 (1), 23 - 27." type="journal article" year="2006">Sartori &amp; Soldán (2006)</bibRefCitation>
for complete list of distributional records). Eastern limits are uncertain, the species has been found easternmost in the Asian part of
<collectingCountry id="EE867677C0764F30FD0EFE30FD2DFE21" box="[666,749,392,418]" name="Turkey" pageId="12" pageNumber="13">Turkey</collectingCountry>
(
<bibRefCitation id="F2004B16C0764F30FD68FE30FC63FE21" author="Kazanci" box="[764,931,392,418]" pageId="12" pageNumber="17" refString="Kazanci, N. (1984) New Ephemeroptera (Insecta) from Turkey. Aquatic Insects, 6, 253 - 258." type="journal article" year="1984">Kazanci 1984</bibRefCitation>
). The species is considered a typical Pontic faunistic element according to
<bibRefCitation id="F2004B16C0764F30FD94FE08FD76FE49" author="Haybach" box="[512,694,432,458]" pageId="12" pageNumber="17" refString="Haybach, A. (1998) Die Eintagsfliegen (Insecta: Ephemeroptera) von Rheinland - Pfalz - Zoogeographie, Faunistik, Okologie, Taxonomie und Nomenklatur - Unter besonderer Berucksichtigung der Familie Heptageniidae und unter Einbeziehung die ubrigen aus Deutschland bekannten Arten. Dissertation am Fachbereich Biologie der Johannes Gutenberg-Universitat Mainz, 417 pp." type="book" year="1998">Haybach (1998</bibRefCitation>
,
<bibRefCitation id="F2004B16C0764F30FD57FE08FCC2FE49" author="Haybach" box="[707,770,432,458]" pageId="12" pageNumber="17" refString="Haybach, A. (2003) Zoogeographische Aspekte der Eintagfliegenbesiedlung Deutschlands (Insecta, Ephemeroptera. Verhandlungen der westdeutschen Entomologentagung, 2002, 187 - 209." type="journal article" year="2003">2003</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>