treatments-xml/data/A2/09/49/A209493BFD7BFFA4AFA619C1FB95BB11.xml

<document id="A5B2557E2590BF209501B71C1D03BEEB" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. &amp; Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD7BFFA4AFA619C1FB95BB11" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ligyrus (Ligyrus) allonasutus Lopez-Garcia &amp; Deloya 2022, new species" docType="treatment" docVersion="7" lastPageNumber="38" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="37" updateTime="1699396337306" updateUser="plazi">
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<mods:title id="84A8FA3145C877AF7074CBB7044812DD">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
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<mods:namePart id="4C77FA935092F83A59B2E393AC6B90DF">Deloya, Cuauhtémoc</mods:namePart>
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<taxonomicName id="EDA083AEFD7BFFA5AFA619C1FC8EBCD0" authority="Lopez-Garcia &amp; Deloya" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2022" box="[151,816,150,178]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="allonasutus" status="sp. nov." subGenus="Ligyrus">
<emphasis id="18D4243FFD7BFFA5AFA619C1FF4ABCD0" bold="true" box="[151,244,151,177]" italics="true" pageId="36" pageNumber="37">Ligyrus</emphasis>
(
<emphasis id="18D4243FFD7BFFA5AE3419C1FEDCBCD0" bold="true" box="[261,354,151,177]" italics="true" pageId="36" pageNumber="37">Ligyrus</emphasis>
) 
<emphasis id="18D4243FFD7BFFA5AE4219C1FE41BCD3" bold="true" box="[371,511,151,178]" italics="true" pageId="36" pageNumber="37">allonasutus</emphasis>
López-García &amp; Deloya
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<taxonomicNameLabel id="03E79944FD7BFFA5AC0919C1FC76BCD0" box="[824,968,151,177]" pageId="36" pageNumber="37" rank="species">new species</taxonomicNameLabel>
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<paragraph id="2A1FF82DFD7BFFA5AFA619EDFD64BCB4" blockId="36.[151,968,150,214]" box="[151,730,187,214]" pageId="36" pageNumber="37">
(
<figureCitation id="B29BE4A8FD7BFFA5AFAE19EDFEBCBCB4" box="[159,258,187,213]" captionStart="FIGURE 2" captionStartId="82.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1548]" captionTargetId="figure-24@82.[151,1436,181,1560]" captionTargetPageId="82" captionText="FIGURE 2. Labium. A) Ligyrus (Anagrylius) moroni, B) Ligyrus (Ligyrus) allonasutus, C) L. (Ligyrus) gibbosus, D) L. (Ligyrus) villosus, E) L. (Ligyrodes) relictus, F) Proculigyrus cicatricosus, G) Euligyrus ebenus, H) E. similis, I) Tomarus gyas, J) Tomarus selanderi, K) Tomarus subtropicus. Abbreviations: LL: Ligular lobes, LP: Labial palpus, M: Mentum, S: Suture." figureDoi="http://doi.org/10.5281/zenodo.7345015" httpUri="https://zenodo.org/record/7345015/files/figure.png" pageId="36" pageNumber="37">Figs. 2B</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AE3C19EDFE8CBCB4" box="[269,306,187,213]" captionStart="FIGURE 3" captionStartId="83.[151,250,998,1022]" captionTargetBox="[156,1436,185,967]" captionTargetId="figure-17@83.[151,1436,181,974]" captionTargetPageId="83" captionText="FIGURE 3. Maxilla. A) Ligyrus (Anagrylius) moroni, B) L. (Ligyrodes) sallaei, C) Ligyrus (Ligyrus) allonasutus. Numeration of teeth of galea: D) Euligyrus similis, E) L. (Ligyrodes) sallaei, F) Tomarus gyas, G) L. (Ligyrus) fossor, H) L. (Ligyrus) allonasutus." figureDoi="http://doi.org/10.5281/zenodo.7345017" httpUri="https://zenodo.org/record/7345017/files/figure.png" pageId="36" pageNumber="37">3H</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AE0F19EDFED6BCB4" box="[318,360,187,213]" captionStart="FIGURE 11" captionStartId="88.[151,250,931,955]" captionTargetBox="[234,1354,186,899]" captionTargetId="figure-24@88.[231,1357,181,907]" captionTargetPageId="88" captionText="FIGURE 11. Protibia in frontal view.A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) relictus, E) L. (Ligyrodes) sallaei, F) L. (Ligyrus) allonasutus, G) L. (Ligyrus) villosus, H) Tomarus bituberculatus, I) T. colombianus, J) T. pullus." figureDoi="http://doi.org/10.5281/zenodo.7345051" httpUri="https://zenodo.org/record/7345051/files/figure.png" pageId="36" pageNumber="37">11F</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AE4519EDFE78BCB7" box="[372,454,187,214]" captionStart="FIGURE 12" captionStartId="88.[151,250,1921,1945]" captionTargetBox="[237,1351,1055,1897]" captionTargetId="figure-89@88.[233,1355,1052,1897]" captionTargetPageId="88" captionText="FIGURE 12. Mesotibia (A–F). A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei; E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) villosus. Metatibia (G–L). G) Euligyrus similis, H) Proculigyrus cicatricosus, I) Ligyrus (Anagrylius) moroni, J) L. (Ligyrodes) sallaei, K) L. (Ligyrus) allonasutus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345053" httpUri="https://zenodo.org/record/7345053/files/figure.png" pageId="36" pageNumber="37">12E, K</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AEE019EDFDBFBCB4" box="[465,513,187,213]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="36" pageNumber="37">13E</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AD3D19EDFD8DBCB7" box="[524,563,187,214]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="36" pageNumber="37">15I</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AD7119EDFDD1BCB4" box="[576,623,187,213]" captionStart="FIGURE 21" captionStartId="96.[151,250,1739,1763]" captionTargetBox="[161,1418,194,1706]" captionTargetId="figure-24@96.[151,1435,181,1715]" captionTargetPageId="96" captionText="FIGURE 21. Internal sac. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus. Abbreviations:AL:Accessory lamellae, Br: Brush of setae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula." figureDoi="http://doi.org/10.5281/zenodo.7345085" httpUri="https://zenodo.org/record/7345085/files/figure.png" pageId="36" pageNumber="37">21E</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AD4A19EDFD17BCB7" box="[635,681,187,214]" captionStart="FIGURE 25" captionStartId="100.[151,250,1841,1865]" captionTargetBox="[163,1432,189,1806]" captionTargetId="figure-24@100.[151,1436,181,1817]" captionTargetPageId="100" captionText="FIGURE 25. Habitus of the species of Ligyrus. Subgenus Anagrylius: A) Ligyrus cuniculus, B) L. moroni. Subgenus Ligyrodes: C) L. peruvianus, D) L. relictus, E) L. sallaei. Subgenus Ligyrus: F) L. allonasutus, G) L. bidentulus, H) L. burmeisteri. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345097" httpUri="https://zenodo.org/record/7345097/files/figure.png" pageId="36" pageNumber="37">25F</figureCitation>
; 
<figureCitation id="B29BE4A8FD7BFFA5AD8419EDFD6CBCB4" box="[693,722,187,213]" captionStart="FIGURE 34" captionStartId="107.[151,250,2007,2031]" captionTargetBox="[246,1346,1181,1977]" captionTargetId="figure-32@107.[242,1346,1181,1983]" captionTargetPageId="107" captionText="FIGURE 34. Distributional records of Ligyrus (Ligyrus) allonasutus and L. nasutus." figureDoi="http://doi.org/10.5281/zenodo.7345125" httpUri="https://zenodo.org/record/7345125/files/figure.png" pageId="36" pageNumber="37">34</figureCitation>
)
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<emphasis id="18D4243FFD7BFFA5AFA61855FEFBBD7C" bold="true" box="[151,325,259,285]" pageId="36" pageNumber="37">Type material.</emphasis>
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(
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) “ 
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, 
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, 
<collectingMunicipality id="CA7B6257FD7BFFA5AC091852FC32BD7F" box="[824,908,260,286]" pageId="36" pageNumber="37">Juan R.</collectingMunicipality>
/ 
<location id="2F7FAEF6FD7BFFA5AC921855FB7ABD7C" LSID="urn:lsid:plazi:treatment:A209493BFD7BFFA4AFA619C1FB95BB11:2F7FAEF6FD7BFFA5AC921855FB7ABD7C" box="[931,1220,259,286]" country="Mexico" municipality="Juan R." name="Escudero, El Terrero, 360" pageId="36" pageNumber="37" stateProvince="Guerrero">Escudero, El Terrero, 360</location>
/ msnm, 24-vii-12, / Alumbrado público / Héctor Calvo Gatica // 
<taxonomicName id="EDA083AEFD7BFFA5AD941871FC3DBD20" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2022" box="[677,899,295,321]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="allonasutus">Ligyrus allonasutus</taxonomicName>
/ López-García &amp; Deloya / HOLOTYPE”.
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<materialsCitation id="9AC8F270FD7BFFA5AA46187EFE86BDE8" ID-GBIF-Occurrence="3986696503" collectionCode="IEXA" country="Mexico" location="Escudero, El Terrero, 360" municipality="Juan R." pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero" typeStatus="holotype">
Female allotype (IEXA) labeled as holotype and with yellow label “ 
<taxonomicName id="EDA083AEFD7BFFA5ACB3181DFBDCBD04" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2022" box="[898,1122,331,357]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="allonasutus">Ligyrus allonasutus</taxonomicName>
/ López-García &amp; Deloya / ALLOTYPE”
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. 
<typeStatus id="F51B468FFD7BFFA5AE721826FE0EBDEB" box="[323,432,368,394]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
(
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, 
<specimenCount id="3CA633A4FD7BFFA5AEC21839FDA9BDEB" box="[499,535,367,394]" pageId="36" pageNumber="37" type="female">5♀</specimenCount>
): One female labeled as 
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. One male and 
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) “ 
<collectingCountry id="52B7B8BDFD7BFFA5AA0C1839FA26BDE8" box="[1341,1432,367,393]" name="Mexico" pageId="36" pageNumber="37">México</collectingCountry>
, 
<collectingRegion id="E86436CFFD7BFFA5AFA618C5FF41BDCC" box="[151,255,403,429]" country="Mexico" name="Guerrero" pageId="36" pageNumber="37">Guerrero</collectingRegion>
, Juan R. / Escudero, El Terrero, 360 / msnm, 01-31-VIII-12, / Alumbrado público / Héctor Calvo Gatica” Two males and 
<specimenCount id="3CA633A4FD7BFFA5AE7D18EEFE76BDB0" box="[332,456,439,466]" pageId="36" pageNumber="37" type="female">one female</specimenCount>
(
<collectionCode id="4CB160E8FD7BFFA5AEE818E1FDA1BDB0" box="[473,543,439,465]" pageId="36" pageNumber="37">IEXA</collectionCode>
) “ 
<collectingCountry id="52B7B8BDFD7BFFA5AD0818E1FD2ABDB0" box="[569,660,439,465]" name="Mexico" pageId="36" pageNumber="37">México</collectingCountry>
, 
<collectingRegion id="E86436CFFD7BFFA5AD9118E1FCB6BDB0" box="[672,776,439,465]" country="Mexico" name="Guerrero" pageId="36" pageNumber="37">Guerrero</collectingRegion>
, Juan R. / Escudero, El Terrero, 360 / msnm, 25-VII-12, / Alumbrado público / Héctor Calvo Gatica” One male (
<collectionCode id="4CB160E8FD7BFFA5AC35188DFCF4BD94" box="[772,842,475,501]" pageId="36" pageNumber="37">IEXA</collectionCode>
) “ 
<collectingCountry id="52B7B8BDFD7BFFA5AC55188DFC01BD94" box="[868,959,475,501]" name="Mexico" pageId="36" pageNumber="37">México</collectingCountry>
, 
<collectingRegion id="E86436CFFD7BFFA5ACFA188DFB8DBD94" box="[971,1075,475,501]" country="Mexico" name="Guerrero" pageId="36" pageNumber="37">Guerrero</collectingRegion>
, Juan R. / Escudero, El Terrero, 360 / msnm, 08-IX-12, / Alumbrado público / Héctor Calvo Gatica” One male (
<collectionCode id="4CB160E8FD7BFFA5AB2018A9FBE9BE78" box="[1041,1111,511,537]" pageId="36" pageNumber="37">IEXA</collectionCode>
) “ 
<collectingCountry id="52B7B8BDFD7BFFA5AB4118A9FB75BE78" box="[1136,1227,511,537]" name="Mexico" pageId="36" pageNumber="37">México</collectingCountry>
, 
<collectingRegion id="E86436CFFD7BFFA5ABE418A9FA83BE78" box="[1237,1341,511,537]" country="Mexico" name="Guerrero" pageId="36" pageNumber="37">Guerrero</collectingRegion>
, Juan R. / Escudero, El Terrero, Luz / mercurial, 
<specimenCount id="3CA633A4FD7BFFA5AD691B75FD19BE5F" box="[600,679,547,574]" pageId="36" pageNumber="37" type="generic">
<quantity id="ED5855C8FD7BFFA5AD691B75FD1DBE5F" box="[600,675,547,574]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="36" pageNumber="37" unit="m" value="350.0">350 m</quantity>
.
</specimenCount>
/ 01-30-VI-12, / Héctor Calvo Gatica” One male (
<collectionCode id="4CB160E8FD7BFFA5ABEF1B75FA9ABE5C" box="[1246,1316,547,573]" pageId="36" pageNumber="37">IEXA</collectionCode>
) “ 
<collectingCountry id="52B7B8BDFD7BFFA5AA0C1B75FA26BE5C" box="[1341,1432,547,573]" name="Mexico" pageId="36" pageNumber="37">México</collectingCountry>
, 
<collectingRegion id="E86436CFFD7BFFA5AFA61B11FF41BE00" box="[151,255,583,609]" country="Mexico" name="Guerrero" pageId="36" pageNumber="37">Guerrero</collectingRegion>
, Juan R. / Escudero, El Terrero, 360 / msnm, 25-VIII-12, / Excremento bovino / Héctor Calvo Gatica” One female (
<collectionCode id="4CB160E8FD7BFFA5AFC01B3DFE89BEE4" box="[241,311,619,645]" pageId="36" pageNumber="37">IEXA</collectionCode>
) “ 
<collectingCountry id="52B7B8BDFD7BFFA5AE7F1B3DFE17BEE4" box="[334,425,619,645]" name="Mexico" pageId="36" pageNumber="37">México</collectingCountry>
, 
<collectingRegion id="E86436CFFD7BFFA5AE821B3DFDA5BEE4" box="[435,539,619,645]" country="Mexico" name="Guerrero" pageId="36" pageNumber="37">Guerrero</collectingRegion>
, Juan R. / Escudero, El Terrero, Luz / mercurial, 
<specimenCount id="3CA633A4FD7BFFA5AB031B3DFBC1BEE7" box="[1074,1151,619,646]" pageId="36" pageNumber="37" type="generic">
<quantity id="ED5855C8FD7BFFA5AB031B3DFBC4BEE7" box="[1074,1146,619,646]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="36" pageNumber="37" unit="m" value="350.0">350 m</quantity>
.
</specimenCount>
/ 25-V-12, / Héctor Calvo Gatica” All 
<typeStatus id="F51B468FFD7BFFA5AE2C1BC6FE36BECB" box="[285,392,656,682]" pageId="36" pageNumber="37" type="paratype">paratypes</typeStatus>
with yellow label “ 
<taxonomicName id="EDA083AEFD7BFFA5AD521BD9FCFFBEC8" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2022" box="[611,833,655,681]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="allonasutus">Ligyrus allonasutus</taxonomicName>
/ López-García &amp; Deloya / 
<typeStatus id="F51B468FFD7BFFA5AB4A1BD9FAB3BEC8" box="[1147,1293,655,681]" pageId="36" pageNumber="37" type="paratype">PARATYPE</typeStatus>
”.
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD7BFFA5AFF61BE5FE3EB9FC" pageId="36" pageNumber="37" type="description">
<paragraph id="2A1FF82DFD7BFFA5AFF61BE5FD7EB888" blockId="36.[151,1437,259,2013]" pageId="36" pageNumber="37">
<emphasis id="18D4243FFD7BFFA5AFF61BE5FE69BEAC" bold="true" box="[199,471,691,717]" pageId="36" pageNumber="37">
Description. 
<typeStatus id="F51B468FFD7BFFA5AE521BE5FE6CBEAC" box="[355,466,691,717]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
.
</emphasis>
Habitus as in 
<figureCitation id="B29BE4A8FD7BFFA5ADB71BE5FD55BEAF" box="[646,747,691,718]" captionStart="FIGURE 25" captionStartId="100.[151,250,1841,1865]" captionTargetBox="[163,1432,189,1806]" captionTargetId="figure-24@100.[151,1436,181,1817]" captionTargetPageId="100" captionText="FIGURE 25. Habitus of the species of Ligyrus. Subgenus Anagrylius: A) Ligyrus cuniculus, B) L. moroni. Subgenus Ligyrodes: C) L. peruvianus, D) L. relictus, E) L. sallaei. Subgenus Ligyrus: F) L. allonasutus, G) L. bidentulus, H) L. burmeisteri. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345097" httpUri="https://zenodo.org/record/7345097/files/figure.png" pageId="36" pageNumber="37">
Fig. 
<specimenCount id="3CA633A4FD7BFFA5AD8C1BE5FD55BEAF" box="[701,747,691,718]" pageId="36" pageNumber="37" type="generic">25F</specimenCount>
</figureCitation>
. Length 
<quantity id="ED5855C8FD7BFFA5AC631BE5FC03BEAF" box="[850,957,691,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="36" pageNumber="37" unit="mm" value="16.1">16.1 mm</quantity>
; humeral width 
<quantity id="ED5855C8FD7BFFA5AB4C1BE5FB64BEAF" box="[1149,1242,691,718]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="36" pageNumber="37" unit="mm" value="8.2">8.2 mm</quantity>
. Color dark reddish brown. 
<emphasis id="18D4243FFD7BFFA5AE151B81FED3BE90" bold="true" box="[292,365,727,753]" pageId="36" pageNumber="37">Head:</emphasis>
Frons deeply rugopunctate, with large and sparse punctures, nearly smooth on vertex and with only sparse, small punctures. Frontoclypeal carina straight, complete. Ocular canthus acute, with 10 ventral setae. Clypeal surface rugopunctate to transversely rugose, striae denser than those on frons. Clypeus triangular, base 3.5 times wider than apex. Clypeal teeth triangular, separated by 1 tooth diameter. Mandible with 2 apical and 1 lateral, acute tooth, tooth 1 longer and narrower than tooth 2. Maxilla rectangular, 3.5 times longer than wide; galea with 3 dorsal teeth. Apex of labrum evenly rounded. Interocular distance 3.5 times an eye width. Antennal club very long. 
<emphasis id="18D4243FFD7BFFA5AFA61AF9FEA7BFA8" bold="true" box="[151,281,943,969]" pageId="36" pageNumber="37">Pronotum:</emphasis>
Surface with large, dense punctures. Apex without tubercle or fovea. 
<emphasis id="18D4243FFD7BFFA5AB0C1AF9FB2CBFA8" bold="true" box="[1085,1170,943,969]" pageId="36" pageNumber="37">Elytra:</emphasis>
First interval punctate, punctures similar to the other intervals. Inner surface of apex with small transverse tubercles forming about 70 parallel lines. 
<emphasis id="18D4243FFD7BFFA5AE061AA1FE0BB870" bold="true" box="[311,437,1015,1041]" pageId="36" pageNumber="37">Abdomen:</emphasis>
Apex of tergite IV with a diagonal line of 11 transverse, parallel, short carinae. Pygidial surface with small punctures, denser on base and anterior corners. Surface in lateral view strongly convex. 
<emphasis id="18D4243FFD7BFFA5AA6A1D4DFA22B854" bold="true" box="[1371,1436,1051,1077]" pageId="36" pageNumber="37">Legs:</emphasis>
Protibia tridentate; basal tooth slightly distant from others (
<figureCitation id="B29BE4A8FD7BFFA5AC1E1D69FC2FB838" box="[815,913,1087,1113]" captionStart="FIGURE 11" captionStartId="88.[151,250,931,955]" captionTargetBox="[234,1354,186,899]" captionTargetId="figure-24@88.[231,1357,181,907]" captionTargetPageId="88" captionText="FIGURE 11. Protibia in frontal view.A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) relictus, E) L. (Ligyrodes) sallaei, F) L. (Ligyrus) allonasutus, G) L. (Ligyrus) villosus, H) Tomarus bituberculatus, I) T. colombianus, J) T. pullus." figureDoi="http://doi.org/10.5281/zenodo.7345051" httpUri="https://zenodo.org/record/7345051/files/figure.png" pageId="36" pageNumber="37">Fig. 11F</figureCitation>
). Protibial surface sparsely punctate. Protarsus of male simple, inner claw simple. Metatibia strongly contracted at apex (
<figureCitation id="B29BE4A8FD7BFFA5ACE01D35FB87B81C" box="[977,1081,1123,1149]" captionStart="FIGURE 12" captionStartId="88.[151,250,1921,1945]" captionTargetBox="[237,1351,1055,1897]" captionTargetId="figure-89@88.[233,1355,1052,1897]" captionTargetPageId="88" captionText="FIGURE 12. Mesotibia (A–F). A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei; E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) villosus. Metatibia (G–L). G) Euligyrus similis, H) Proculigyrus cicatricosus, I) Ligyrus (Anagrylius) moroni, J) L. (Ligyrodes) sallaei, K) L. (Ligyrus) allonasutus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345053" httpUri="https://zenodo.org/record/7345053/files/figure.png" pageId="36" pageNumber="37">Fig. 12K</figureCitation>
). Apex of metatibia entire, with 13 spinules. 
<emphasis id="18D4243FFD7BFFA5AE131DD1FE6AB8C0" bold="true" box="[290,468,1159,1185]" pageId="36" pageNumber="37">Male genitalia:</emphasis>
Spiculum gastrale thin, apex truncate. Parameres with sides nearly parallel, lateral ventral teeth short (
<figureCitation id="B29BE4A8FD7BFFA5AE771DFDFE15B8A7" box="[326,427,1195,1222]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="36" pageNumber="37">Figs. 15I</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AE861DFDFE55B8A4" box="[439,491,1195,1221]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="36" pageNumber="37">18A</figureCitation>
). Internal sac with lamellar spiny belt long (5 times longer than copulatory lamellae); accessory lamella thickened, elongate (
<figureCitation id="B29BE4A8FD7BFFA5AD7C1D99FD0DB888" box="[589,691,1231,1257]" captionStart="FIGURE 21" captionStartId="96.[151,250,1739,1763]" captionTargetBox="[161,1418,194,1706]" captionTargetId="figure-24@96.[151,1435,181,1715]" captionTargetPageId="96" captionText="FIGURE 21. Internal sac. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus. Abbreviations:AL:Accessory lamellae, Br: Brush of setae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula." figureDoi="http://doi.org/10.5281/zenodo.7345085" httpUri="https://zenodo.org/record/7345085/files/figure.png" pageId="36" pageNumber="37">Fig. 21E</figureCitation>
).
</paragraph>
<paragraph id="2A1FF82DFD7BFFA5AFF61DA5FAE6B934" blockId="36.[151,1437,259,2013]" pageId="36" pageNumber="37">
<emphasis id="18D4243FFD7BFFA5AFF61DA5FE8DB96C" bold="true" box="[199,307,1267,1293]" pageId="36" pageNumber="37">
<typeStatus id="F51B468FFD7BFFA5AFF61DA5FE90B96C" box="[199,302,1267,1293]" pageId="36" pageNumber="37" type="allotype">Allotype</typeStatus>
.
</emphasis>
Length 
<quantity id="ED5855C8FD7BFFA5AEBE1DA5FE4BB96F" box="[399,501,1267,1294]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="36" pageNumber="37" unit="mm" value="16.8">16.8 mm</quantity>
; humeral width 
<quantity id="ED5855C8FD7BFFA5AD961DA5FD41B96F" box="[679,767,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="37" unit="mm" value="8.9">8.9 mm</quantity>
. 
<emphasis id="18D4243FFD7BFFA5AC391DA5FC68B96C" bold="true" box="[776,982,1267,1293]" pageId="36" pageNumber="37">Female genitalia:</emphasis>
Subcoxite subrectangular (2 times wider than long). Coxite subrectangular (wider than long); surface strongly concave. Subcoxite wider and longer than coxite (
<figureCitation id="B29BE4A8FD7BFFA5AFDA1C6DFEEEB934" box="[235,336,1339,1365]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="36" pageNumber="37">Fig. 13E</figureCitation>
). 
<emphasis id="18D4243FFD7BFFA5AE551C6DFE5EB934" bold="true" box="[356,480,1339,1365]" pageId="36" pageNumber="37">Pygidium:</emphasis>
Surface in lateral view strongly flat. 
<emphasis id="18D4243FFD7BFFA5ACB01C6DFC7CB934" bold="true" box="[897,962,1339,1365]" pageId="36" pageNumber="37">Legs:</emphasis>
Apex of metatibia with 19 spinules.
</paragraph>
<paragraph id="2A1FF82DFD7BFFA5AFF61C09FE3EB9FC" blockId="36.[151,1437,259,2013]" pageId="36" pageNumber="37">
<emphasis id="18D4243FFD7BFFA5AFF61C09FE81B918" bold="true" box="[199,319,1375,1401]" pageId="36" pageNumber="37">Variation.</emphasis>
Eleven 
<typeStatus id="F51B468FFD7BFFA5AEAF1C36FDB3B91B" box="[414,525,1376,1402]" pageId="36" pageNumber="37" type="paratype">paratypes</typeStatus>
. Length 
<quantity id="ED5855C8FD7BFFA5AD401C09FCA6B91B" box="[625,792,1375,1402]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" metricValueMax="1.7399999999999998" metricValueMin="1.48" pageId="36" pageNumber="37" unit="mm" value="16.1" valueMax="17.4" valueMin="14.8">14.8–17.4 mm</quantity>
; humeral width 
<quantity id="ED5855C8FD7BFFA5ACE31C09FBC4B91B" box="[978,1146,1375,1402]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" metricValueMax="8.98" metricValueMin="7.82" pageId="36" pageNumber="37" unit="mm" value="8.4" valueMax="8.98" valueMin="7.82">7.82–8.98 mm</quantity>
. 
<emphasis id="18D4243FFD7BFFA5ABB71C09FB79B918" bold="true" box="[1158,1223,1375,1401]" pageId="36" pageNumber="37">Legs:</emphasis>
Apex of metatibia with 13–20 spinules.
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD7BFFA5AFF61CF1FDF6BA30" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="2A1FF82DFD7BFFA5AFF61CF1FDF6BA30" blockId="36.[151,1437,259,2013]" pageId="36" pageNumber="37">
<emphasis id="18D4243FFD7BFFA5AFF61CF1FEFCB9A0" bold="true" box="[199,322,1447,1473]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
<taxonomicName id="EDA083AEFD7BFFA5AE7B1CFEFD97B9A0" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2022" box="[330,553,1447,1473]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="allonasutus">
<emphasis id="18D4243FFD7BFFA5AE7B1CFEFD97B9A0" box="[330,553,1447,1473]" italics="true" pageId="36" pageNumber="37">Ligyrus allonasutus</emphasis>
</taxonomicName>
can be recognized by the triangular clypeus; frontal carina straight, not interrupted; mandible with outer margin acute; pronotum without tubercle or fovea, without an apical swelling; pronotal punctation deep and large; metatibia elongate with apex strongly narrowed; parameres with sides nearly parallel; lateral ventral teeth short (
<figureCitation id="B29BE4A8FD7BFFA5AE8C1F45FD9DBA4F" box="[445,547,1555,1582]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="36" pageNumber="37">Figs. 15I</figureCitation>
, 
<figureCitation id="B29BE4A8FD7BFFA5AD1E1F45FDDCBA4C" box="[559,610,1555,1581]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="36" pageNumber="37">18A</figureCitation>
); internal sac with accessory lamella thick and elongate (
<figureCitation id="B29BE4A8FD7BFFA5ABEE1F45FAFABA4C" box="[1247,1348,1555,1581]" captionStart="FIGURE 21" captionStartId="96.[151,250,1739,1763]" captionTargetBox="[161,1418,194,1706]" captionTargetId="figure-24@96.[151,1435,181,1715]" captionTargetPageId="96" captionText="FIGURE 21. Internal sac. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus. Abbreviations:AL:Accessory lamellae, Br: Brush of setae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula." figureDoi="http://doi.org/10.5281/zenodo.7345085" httpUri="https://zenodo.org/record/7345085/files/figure.png" pageId="36" pageNumber="37">Fig. 21E</figureCitation>
); coxite of female strongly concave (
<figureCitation id="B29BE4A8FD7BFFA5AEE61F61FD85BA30" box="[471,571,1591,1617]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="36" pageNumber="37">Fig. 13E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD7BFFA4AFF61F0DFB95BB11" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="etymology">
<paragraph id="2A1FF82DFD7BFFA5AFF61F0DFB23BAF8" blockId="36.[151,1437,259,2013]" pageId="36" pageNumber="37">
<emphasis id="18D4243FFD7BFFA5AFF61F0DFEF0BA14" bold="true" box="[199,334,1627,1653]" pageId="36" pageNumber="37">Etymology.</emphasis>
The name “ 
<taxonomicName id="EDA083AEFD7BFFA5AEEE1F0DFDDEBA14" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2022" box="[479,608,1627,1653]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="allonasutus">
<emphasis id="18D4243FFD7BFFA5AEEE1F0DFDDEBA14" box="[479,608,1627,1653]" italics="true" pageId="36" pageNumber="37">allonasutus</emphasis>
</taxonomicName>
” comes from the Greek 
<emphasis id="18D4243FFD7BFFA5ACB31F0DFC06BA14" box="[898,952,1627,1653]" italics="true" pageId="36" pageNumber="37">allós</emphasis>
(other or different) and the name 
<taxonomicName id="EDA083AEFD7BFFA5AA761F0AFA23BA14" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1351,1437,1628,1653]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="18D4243FFD7BFFA5AA761F0AFA23BA14" box="[1351,1437,1628,1653]" italics="true" pageId="36" pageNumber="37">nasutus</emphasis>
</taxonomicName>
referring to the species from which the new species was separated (see taxonomic remarks).
</paragraph>
<paragraph id="2A1FF82DFD7BFFA5AFF61FF5FCFEBBBC" blockId="36.[151,1437,259,2013]" pageId="36" pageNumber="37">
<emphasis id="18D4243FFD7BFFA5AFF61FF5FE03BADC" bold="true" box="[199,445,1699,1725]" pageId="36" pageNumber="37">Taxonomic remarks.</emphasis>
The new species is purposed here to place populations from western 
<collectingCountry id="52B7B8BDFD7BFFA5ABF51FF5FAA4BADC" box="[1220,1306,1699,1725]" name="Mexico" pageId="36" pageNumber="37">Mexico</collectingCountry>
to 
<collectingCountry id="52B7B8BDFD7BFFA5AA0D1FF2FA26BADF" box="[1340,1432,1700,1726]" name="Panama" pageId="36" pageNumber="37">Panama</collectingCountry>
, which have been traditionally known as 
<taxonomicName id="EDA083AEFD7BFFA5AD691F9EFCB4BA80" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[600,778,1736,1761]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="18D4243FFD7BFFA5AD691F9EFCB4BA80" box="[600,778,1736,1761]" italics="true" pageId="36" pageNumber="37">Ligyrus nasutus</emphasis>
</taxonomicName>
. The species was described by 
<bibRefCitation id="4E3185DCFD7BFFA5AB571F91FA8DBA80" author="Burmeister, H." box="[1126,1331,1735,1761]" pageId="36" pageNumber="37" refId="ref88626" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, viii + 584 pp." type="book" year="1847">Burmeister (1847)</bibRefCitation>
with 
<specimenCount id="3CA633A4FD7BFFA5AA431F9EFF50BB64" pageId="36" pageNumber="37" type="female">two females</specimenCount>
from Yucatán and 
<collectingCountry id="52B7B8BDFD7BFFA5AEF91FBDFDFBBB64" box="[456,581,1771,1797]" name="Guatemala" pageId="36" pageNumber="37">Guatemala</collectingCountry>
. 
<bibRefCitation id="4E3185DCFD7BFFA5AD611FBDFD40BB64" author="Endrodi, S." box="[592,766,1771,1797]" pageId="36" pageNumber="37" pagination="1 - 145" refId="ref89768" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen, 37, 1 - 145." type="journal article" year="1969">Endrödi (1969)</bibRefCitation>
choose the female from 
<collectingRegion id="E86436CFFD7BFFA5AB281FBDFBCDBB64" box="[1049,1139,1771,1797]" country="Mexico" name="Yucatan" pageId="36" pageNumber="37">Yucatán</collectingRegion>
(in bad conditions) as the 
<typeStatus id="F51B468FFD7BFFA5AFA61E59FF40BB48" box="[151,254,1807,1833]" pageId="36" pageNumber="37" type="lectotype">lectotype</typeStatus>
for 
<taxonomicName id="EDA083AEFD7BFFA5AE1F1E46FE1CBB48" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[302,418,1808,1833]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="18D4243FFD7BFFA5AE1F1E46FE1CBB48" box="[302,418,1808,1833]" italics="true" pageId="36" pageNumber="37">L. nasutus</emphasis>
</taxonomicName>
, but his descriptions and illustrations are based only on populations from western 
<collectingCountry id="52B7B8BDFD7BFFA5AA761E59FA23BB48" box="[1351,1437,1807,1833]" name="Mexico" pageId="36" pageNumber="37">Mexico</collectingCountry>
and 
<collectingCountry id="52B7B8BDFD7BFFA5AFF91E65FEFFBB2C" box="[200,321,1843,1869]" name="Guatemala" pageId="36" pageNumber="37">Guatemala</collectingCountry>
to 
<collectingCountry id="52B7B8BDFD7BFFA5AE561E62FE7DBB2F" box="[359,451,1844,1870]" name="Panama" pageId="36" pageNumber="37">Panama</collectingCountry>
. After his work, the name 
<taxonomicName id="EDA083AEFD7BFFA5ADC31E62FCD9BB2C" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[754,871,1844,1869]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="18D4243FFD7BFFA5ADC31E62FCD9BB2C" box="[754,871,1844,1869]" italics="true" pageId="36" pageNumber="37">L. nasutus</emphasis>
</taxonomicName>
was widely used for supposedly a unique species with wide distribution, but there was confusion since the original description as 
<bibRefCitation id="4E3185DCFD7BFFA5AB011E01FABEBB10" author="Burmeister, H." box="[1072,1280,1879,1905]" pageId="36" pageNumber="37" refId="ref88626" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, viii + 584 pp." type="book" year="1847">Burmeister (1847)</bibRefCitation>
included 
<specimenCount id="3CA633A4FD7BFFA5AA431E0EFF5DBBF4" pageId="36" pageNumber="37" type="female">two female</specimenCount>
specimens from different species as types for 
<taxonomicName id="EDA083AEFD7BFFA5ADC91E2AFCD0BBF4" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[760,878,1916,1941]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="18D4243FFD7BFFA5ADC91E2AFCD0BBF4" box="[760,878,1916,1941]" italics="true" pageId="36" pageNumber="37">L. nasutus</emphasis>
</taxonomicName>
. However, as the 
<typeStatus id="F51B468FFD7BFFA5AB0C1E2DFB1ABBF4" box="[1085,1188,1915,1941]" pageId="36" pageNumber="37" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="EDA083AEFD7BFFA5ABFC1E2AFAFCBBF4" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1229,1346,1916,1941]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="18D4243FFD7BFFA5ABFC1E2AFAFCBBF4" box="[1229,1346,1916,1941]" italics="true" pageId="36" pageNumber="37">L. nasutus</emphasis>
</taxonomicName>
is from 
<collectingRegion id="E86436CFFD7BFFA5AFA61EC9FF4ABBD8" box="[151,244,1951,1977]" country="Mexico" name="Yucatan" pageId="36" pageNumber="37">Yucatán</collectingRegion>
, only populations of this region correspond to 
<taxonomicName id="EDA083AEFD7BFFA5AC3B1EF6FCC0BBD8" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[778,894,1952,1977]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="18D4243FFD7BFFA5AC3B1EF6FCC0BBD8" box="[778,894,1952,1977]" italics="true" pageId="36" pageNumber="37">L. nasutus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="EDA083AEFD7BFFA5AC8F1EF6FBE0BBD8" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2022" box="[958,1118,1951,1977]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="allonasutus">
<emphasis id="18D4243FFD7BFFA5AC8F1EF6FBE0BBD8" box="[958,1118,1951,1977]" italics="true" pageId="36" pageNumber="37">L. allonasutus</emphasis>
</taxonomicName>
is proposed here for the remaining populations (western 
<collectingCountry id="52B7B8BDFD7BFFA5AEDA1E95FDFFBBBC" box="[491,577,1987,2013]" name="Mexico" pageId="36" pageNumber="37">Mexico</collectingCountry>
and Central America).
</paragraph>
<paragraph id="2A1FF82DFD7AFFA4AFF619C0FE2CBCB4" blockId="37.[151,1437,150,1905]" pageId="37" pageNumber="38">
<emphasis id="18D4243FFD7AFFA4AFF619C0FE51BCD1" bold="true" box="[199,495,150,176]" pageId="37" pageNumber="38">Geographic distribution.</emphasis>
<taxonomicName id="EDA083AEFD7AFFA4AEC419C1FD6DBCD1" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2022" box="[501,723,150,176]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="allonasutus">
<emphasis id="18D4243FFD7AFFA4AEC419C1FD6DBCD1" box="[501,723,150,176]" italics="true" pageId="37" pageNumber="38">Ligyrus allonasutus</emphasis>
</taxonomicName>
is widely distributed from western 
<collectingCountry id="52B7B8BDFD7AFFA4AB5119C0FB08BCD1" box="[1120,1206,150,176]" name="Mexico" pageId="37" pageNumber="38">Mexico</collectingCountry>
to 
<collectingCountry id="52B7B8BDFD7AFFA4ABE819C1FA8BBCD0" box="[1241,1333,151,177]" name="Panama" pageId="37" pageNumber="38">Panama</collectingCountry>
, between sea level and 
<quantity id="ED5855C8FD7AFFA4AE1E19ECFE30BCB4" box="[303,398,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="37" pageNumber="38" unit="m" value="1850.0">1,850 m</quantity>
.
</paragraph>
<paragraph id="2A1FF82DFD7AFFA4AFF61988FDF4BA81" blockId="37.[151,1437,150,1905]" pageId="37" pageNumber="38">
<emphasis id="18D4243FFD7AFFA4AFF61988FDBBBC99" bold="true" box="[199,517,222,248]" pageId="37" pageNumber="38">
Locality records (
<figureCitation id="B29BE4A8FD7AFFA4AEAE1988FE46BC99" box="[415,504,222,248]" captionStart="FIGURE 34" captionStartId="107.[151,250,2007,2031]" captionTargetBox="[246,1346,1181,1977]" captionTargetId="figure-32@107.[242,1346,1181,1983]" captionTargetPageId="107" captionText="FIGURE 34. Distributional records of Ligyrus (Ligyrus) allonasutus and L. nasutus." figureDoi="http://doi.org/10.5281/zenodo.7345125" httpUri="https://zenodo.org/record/7345125/files/figure.png" pageId="37" pageNumber="38">Fig. 34</figureCitation>
).
</emphasis>
383 examined specimens from CAFESI, CIIDIR, CZUG, EMAC, IEXA, MN/ UFRJ, UNSM, USNM. Some data from 
<bibRefCitation id="4E3185DCFD7AFFA4AD431854FC95BD7D" author="Ratcliffe, B. C." box="[626,811,258,284]" pageId="37" pageNumber="38" pagination="1 - 506" refId="ref92614" refString="Ratcliffe, B. C. (2003) The dynastine scarab beetles of Costa Rica and Panama. Bulletin of the University of Nebraska State Museum, 16, 1 - 506." type="journal article" year="2003">Ratcliffe (2003)</bibRefCitation>
, 
<bibRefCitation id="4E3185DCFD7AFFA4AC0B1854FBE9BD7D" author="Ratcliffe, B. C. &amp; Cave, R. D." box="[826,1111,258,284]" pageId="37" pageNumber="38" pagination="1 - 424" refId="ref92768" refString="Ratcliffe, B. C. &amp; Cave, R. D. (2006) The dynastine scarab beetles of Honduras, Nicaragua, and El Salvador. Bulletin of the University of Nebraska State Museum, 21, 1 - 424." type="journal article" year="2006">Ratcliffe &amp; Cave (2006)</bibRefCitation>
, and 
<bibRefCitation id="4E3185DCFD7AFFA4ABA81854FA26BD7D" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Cano, E. B." box="[1177,1432,258,285]" pageId="37" pageNumber="38" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. &amp; Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
Ratcliffe 
<emphasis id="18D4243FFD7AFFA4AA341855FA87BD7D" box="[1285,1337,258,284]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. (2013)
</bibRefCitation>
. 
<emphasis id="18D4243FFD7AFFA4AFA61870FDA4BD21" bold="true" box="[151,538,294,320]" pageId="37" pageNumber="38">
<collectingCountry id="52B7B8BDFD7AFFA4AFA61870FEFFBD21" box="[151,321,294,320]" name="Costa Rica" pageId="37" pageNumber="38">COSTA RICA</collectingCountry>
(20). 
<collectingRegion id="E86436CFFD7AFFA4AEB51870FE57BD21" box="[388,489,294,320]" country="Costa Rica" name="Alajuela" pageId="37" pageNumber="38">Alajuela</collectingRegion>
(1):
</emphasis>
Caño Negro. 
<emphasis id="18D4243FFD7AFFA4AD861870FC3CBD21" bold="true" box="[695,898,294,320]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AD861870FCFDBD21" box="[695,835,294,320]" country="Costa Rica" name="Guanacaste" pageId="37" pageNumber="38">Guanacaste</collectingRegion>
(16):
</emphasis>
Cañas; Barra Honda; Estación Enrique Jiménez Nuñes; Estación Almendros; Estación Murciélagos; Estación Palo Verde; Finca Jenny; Finca La Pacífica; Lomas Barbudal; Playa Naranjo; Reserva Carara; Reserva Monteverde; Río San Lorenzo; 
<collectingRegion id="E86436CFFD7AFFA4AB7C1838FB6EBDE8" box="[1101,1232,366,393]" inferredBySuffix="true" pageId="37" pageNumber="38">Santa Rosa</collectingRegion>
; Tierras Morenas. 
<emphasis id="18D4243FFD7AFFA4AFA618C4FDD2BDCD" bold="true" box="[151,620,401,428]" pageId="37" pageNumber="38">
<collectingCountry id="52B7B8BDFD7AFFA4AFA618C4FEE5BDCD" box="[151,347,402,428]" name="El Salvador" pageId="37" pageNumber="38">EL SALVADOR</collectingCountry>
(68). 
<collectingRegion id="E86436CFFD7AFFA4AE9018C7FD87BDCD" box="[417,569,401,428]" country="El Salvador" name="Ahuachapan" pageId="37" pageNumber="38">Ahuachapán</collectingRegion>
(6):
</emphasis>
Guaymango; Izcaquillo; Parque Nacional El Imposible. 
<emphasis id="18D4243FFD7AFFA4ABC218C4FA22BDCD" bold="true" box="[1267,1436,402,428]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ABC218C4FAE5BDCD" box="[1267,1371,402,428]" country="El Salvador" name="Cabanas" pageId="37" pageNumber="38">Cabañas</collectingRegion>
(33):
</emphasis>
Cinquera; Ilobasco. 
<emphasis id="18D4243FFD7AFFA4AE4F18E3FD97BDB1" bold="true" box="[382,553,437,464]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AE4F18E3FE48BDB1" box="[382,502,437,464]" country="El Salvador" name="Cuscatlan" pageId="37" pageNumber="38">Cuscatlán</collectingRegion>
(2):
</emphasis>
El Playón (
<quantity id="ED5855C8FD7AFFA4AD8418E0FD4FBDB1" box="[693,753,438,464]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="38" unit="km" value="7.0">7 km</quantity>
NE Coctepeque); San Pedro Perulapán; 
<emphasis id="18D4243FFD7AFFA4ABF618E0FA22BDB1" bold="true" box="[1223,1436,438,464]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ABF618E0FAE5BDB1" box="[1223,1371,438,464]" country="El Salvador" name="La Libertad" pageId="37" pageNumber="38">La Libertad</collectingRegion>
(14):
</emphasis>
Canton El Barillo; 
<collectingRegion id="E86436CFFD7AFFA4AE5D188DFE4DBD95" box="[364,499,474,501]" inferredBySuffix="true" pageId="37" pageNumber="38">La Libertad</collectingRegion>
; Parque Natural Walter Thilo Deininger; San Diego; San Andrés; Santa Tecla; Valle Zapotitán; Zaragoza. 
<emphasis id="18D4243FFD7AFFA4AEB418A8FDB8BE79" bold="true" box="[389,518,510,536]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AEB418A8FE69BE79" box="[389,471,510,536]" country="El Salvador" name="La Paz" pageId="37" pageNumber="38">La Paz</collectingRegion>
(5):
</emphasis>
Campo San Luís Talpa; Caprex; Zacatecoluca. 
<emphasis id="18D4243FFD7AFFA4AB2518A8FB0ABE79" bold="true" box="[1044,1204,509,536]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AB2518A8FB3ABE79" box="[1044,1156,509,536]" country="El Salvador" name="La Union" pageId="37" pageNumber="38">La Unión</collectingRegion>
(1):
</emphasis>
<collectingRegion id="E86436CFFD7AFFA4AB8818A9FA99BE79" box="[1209,1319,510,537]" inferredBySuffix="true" pageId="37" pageNumber="38">La Unión</collectingRegion>
. 
<emphasis id="18D4243FFD7AFFA4AA1E18ABFF7FBE5D" bold="true" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AA1E18ABFA22BE79" box="[1327,1436,509,536]" country="El Salvador" name="Morazan" pageId="37" pageNumber="38">Morazán</collectingRegion>
(1):
</emphasis>
Arambala; Río Sapo. 
<emphasis id="18D4243FFD7AFFA4AE8B1B74FDCDBE5D" bold="true" box="[442,627,546,572]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AE8B1B74FDFCBE5D" box="[442,578,546,572]" country="El Salvador" name="San Miguel" pageId="37" pageNumber="38">San Miguel</collectingRegion>
(1):
</emphasis>
Laguna El Jocotal. 
<emphasis id="18D4243FFD7AFFA4AC631B74FB9CBE5D" bold="true" box="[850,1058,546,572]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AC631B74FC4CBE5D" box="[850,1010,546,572]" country="El Salvador" name="San Salvador" pageId="37" pageNumber="38">San Salvador</collectingRegion>
(3):
</emphasis>
Parque Saburo Hirao; 
<collectingRegion id="E86436CFFD7AFFA4AA151B74FF00BE01" inferredBySuffix="true" pageId="37" pageNumber="38">San Salvador</collectingRegion>
. 
<emphasis id="18D4243FFD7AFFA4AFF91B10FECCBE01" bold="true" box="[200,370,582,608]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AFF91B10FEFDBE01" box="[200,323,582,608]" country="El Salvador" name="Santa Ana" pageId="37" pageNumber="38">Santa Ana</collectingRegion>
(3):
</emphasis>
Los Planes; San José Ingenio. 
<emphasis id="18D4243FFD7AFFA4ADF91B10FCCCBE01" bold="true" box="[712,882,582,608]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ADF91B10FCFFBE01" box="[712,833,582,608]" country="El Salvador" name="Sonsonate" pageId="37" pageNumber="38">Sonsonate</collectingRegion>
(1):
</emphasis>
Sonsonate. 
<emphasis id="18D4243FFD7AFFA4ACC61B10FF7FBEE5" bold="true" pageId="37" pageNumber="38">
<collectingCountry id="52B7B8BDFD7AFFA4ACC61B10FB12BE01" box="[1015,1196,582,608]" name="Guatemala" pageId="37" pageNumber="38">GUATEMALA</collectingCountry>
(104). 
<collectingRegion id="E86436CFFD7AFFA4ABCD1B10FA23BE01" box="[1276,1437,582,608]" country="Guatemala" name="Baja Verapaz" pageId="37" pageNumber="38">Baja Verapaz</collectingRegion>
(5):
</emphasis>
Niño Perdido; Rabinal; Salama; San Jerónimo. 
<emphasis id="18D4243FFD7AFFA4ADEB1B3CFC24BEE5" bold="true" box="[730,922,618,644]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ADEB1B3CFCD7BEE5" box="[730,873,618,644]" country="Guatemala" name="Chiquimula" pageId="37" pageNumber="38">Chiquimula</collectingRegion>
(1):
</emphasis>
Quetzaltepeque. 
<emphasis id="18D4243FFD7AFFA4AB6D1B3CFAA6BEE5" bold="true" box="[1116,1304,618,644]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AB6D1B3CFB59BEE5" box="[1116,1255,618,644]" country="Guatemala" name="El Progreso" pageId="37" pageNumber="38">El Progreso</collectingRegion>
(3):
</emphasis>
El Rancho; San Agustín Acasaguastlán. 
<emphasis id="18D4243FFD7AFFA4AEE51BD8FD3CBEC9" bold="true" box="[468,642,654,680]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AEE51BD8FDFABEC9" box="[468,580,654,680]" country="Guatemala" name="Escuintla" pageId="37" pageNumber="38">Escuintla</collectingRegion>
(17):
</emphasis>
Chulamar; Likin; Peñón de Siquinalá; Puerto de San José; Santa Lucía Cotzumalguapa; 
<collectingRegion id="E86436CFFD7AFFA4AE691BE4FE62BEAC" box="[344,476,690,717]" inferredBySuffix="true" pageId="37" pageNumber="38">Santa Rosa</collectingRegion>
; Taxisco (
<quantity id="ED5855C8FD7AFFA4AD631BE4FD23BEAD" box="[594,669,690,716]" metricMagnitude="4" metricUnit="m" metricValue="8.6" pageId="37" pageNumber="38" unit="km" value="86.0">86 km</quantity>
N); Tiquisate. 
<emphasis id="18D4243FFD7AFFA4AC7C1BE4FBADBEAD" bold="true" box="[845,1043,690,716]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AC7C1BE4FC6FBEAD" box="[845,977,690,716]" country="Guatemala" name="Guatemala" pageId="37" pageNumber="38">Guatemala</collectingRegion>
(10):
</emphasis>
Amatitlán; Ciudad de Guatemala; Puerto Parada; San Jerónimo; Villa Canales. 
<emphasis id="18D4243FFD7AFFA4ADA71B80FC3BBE91" bold="true" box="[662,901,726,752]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ADA71B80FCECBE91" box="[662,850,726,752]" country="Guatemala" name="Huehuetenango" pageId="37" pageNumber="38">Huehuetenango</collectingRegion>
(4):
</emphasis>
Jacaltenango; La Laguna; Nenton. 
<emphasis id="18D4243FFD7AFFA4AA2B1B80FA23BE91" bold="true" box="[1306,1437,726,752]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AA2B1B80FAD7BE91" box="[1306,1385,726,752]" country="Guatemala" name="Jalapa" pageId="37" pageNumber="38">Jalapa</collectingRegion>
(1):
</emphasis>
Jalapa. 
<emphasis id="18D4243FFD7AFFA4AFDF1BACFEC5BF75" bold="true" box="[238,379,762,788]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AFDF1BACFEF6BF75" box="[238,328,762,788]" country="Guatemala" name="Jutiapa" pageId="37" pageNumber="38">Jutiapa</collectingRegion>
(6):
</emphasis>
Cañón ed Monjoy; Comapa; Jutiapa (
<quantity id="ED5855C8FD7AFFA4AC051BACFCCFBF75" box="[820,881,762,788]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="37" pageNumber="38" unit="mi" value="4.0">4 mi.</quantity>
E); Quezada. 
<emphasis id="18D4243FFD7AFFA4AB291BACFAB6BF75" bold="true" box="[1048,1288,762,788]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AB291BACFB6ABF75" box="[1048,1236,762,788]" country="Guatemala" name="Quetzaltenango" pageId="37" pageNumber="38">Quetzaltenango</collectingRegion>
(1):
</emphasis>
Coatepeque. 
<emphasis id="18D4243FFD7AFFA4AFA61A48FEF0BF59" bold="true" box="[151,334,798,824]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AFA61A48FEA4BF59" box="[151,282,798,824]" country="Guatemala" name="Retalhuleu" pageId="37" pageNumber="38">Retalhuleu</collectingRegion>
(8):
</emphasis>
Retalhuleu. 
<emphasis id="18D4243FFD7AFFA4AED31A4BFD06BF59" bold="true" box="[482,696,797,824]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AED31A4BFD3ABF59" box="[482,644,797,824]" country="Guatemala" name="Sacatepequez" pageId="37" pageNumber="38">Sacatepéquez</collectingRegion>
(2):
</emphasis>
San Lucas de Sacatepéquez; Santo Domingo. 
<emphasis id="18D4243FFD7AFFA4ABE61A48FA22BF59" bold="true" box="[1239,1436,798,824]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ABE61A48FAD6BF59" box="[1239,1384,798,824]" country="Guatemala" name="San Marcos" pageId="37" pageNumber="38">San Marcos</collectingRegion>
(4):
</emphasis>
Ayutla; Ixbén. 
<emphasis id="18D4243FFD7AFFA4AE711A14FE44BF3D" bold="true" box="[320,506,834,860]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AE711A14FE76BF3D" box="[320,456,834,860]" country="Guatemala" name="Santa Rosa" pageId="37" pageNumber="38">Santa Rosa</collectingRegion>
(8):
</emphasis>
Chiquimulilla; Cuilapa; Monterrico; Taxisco. 
<emphasis id="18D4243FFD7AFFA4AB3B1A17FB49BF3D" bold="true" box="[1034,1271,833,860]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AB3B1A17FB09BF3D" box="[1034,1207,833,860]" country="Guatemala" name="Suchitepequez" pageId="37" pageNumber="38">Suchitepéquez</collectingRegion>
(13):
</emphasis>
Cuyotenango; Finca Reforma; Ixtacapa; Mazatenango; Reserva Los Tarrales; San Juan Bautista; Santa Bárbara. 
<emphasis id="18D4243FFD7AFFA4ABE71A30FAD3BFE1" bold="true" box="[1238,1389,870,896]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ABE71A30FA91BFE1" box="[1238,1327,870,896]" country="Guatemala" name="Zacapa" pageId="37" pageNumber="38">Zacapa</collectingRegion>
(21):
</emphasis>
Río Hondo; Río Teculután; San José (
<quantity id="ED5855C8FD7AFFA4AD3D1ADCFDFBBFC5" box="[524,581,906,932]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="37" pageNumber="38" unit="km" value="3.0">3 km</quantity>
SE); San Lorenzo; Santa Cruz; Tres Marías. 
<emphasis id="18D4243FFD7AFFA4AB0B1ADCFF63BFA9" bold="true" pageId="37" pageNumber="38">
<collectingCountry id="52B7B8BDFD7AFFA4AB0B1ADCFB64BFC5" box="[1082,1242,906,932]" name="Honduras" pageId="37" pageNumber="38">HONDURAS</collectingCountry>
(555). 
<collectingRegion id="E86436CFFD7AFFA4AA1A1ADFFA23BFC5" box="[1323,1437,905,932]" country="Honduras" name="Atlantida" pageId="37" pageNumber="38">Atlántida</collectingRegion>
(302):
</emphasis>
Carretera a Cortés; Curla (
<quantity id="ED5855C8FD7AFFA4AD241AF8FDEEBFA9" box="[533,592,942,968]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="38" unit="km" value="8.0">8 km</quantity>
NW La Ceiba); Islita (
<quantity id="ED5855C8FD7AFFA4AC691AF8FC1CBFA9" box="[856,930,942,968]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="38" unit="km" value="60.0">60 km</quantity>
La Ceiba); La Ceiba; Tela. 
<emphasis id="18D4243FFD7AFFA4ABD31AF8FA22BFA9" bold="true" box="[1250,1436,942,968]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ABD31AF8FAE5BFA9" box="[1250,1371,942,968]" country="Honduras" name="Choluteca" pageId="37" pageNumber="38">Choluteca</collectingRegion>
(33):
</emphasis>
Choluteca; Duyure; Pespire; Tierra Blanca. 
<emphasis id="18D4243FFD7AFFA4ADBC1A84FCE0BF8D" bold="true" box="[653,862,978,1004]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ADBC1A84FCA2BF8D" box="[653,796,978,1004]" country="Honduras" name="Comayagua" pageId="37" pageNumber="38">Comayagua</collectingRegion>
(49):
</emphasis>
Comayagua; Siguatepeque; Taladro. 
<emphasis id="18D4243FFD7AFFA4AA3A1A87FA22BF8D" bold="true" box="[1291,1436,977,1004]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AA3A1A87FAE4BF8D" box="[1291,1370,977,1004]" country="Honduras" name="Cortes" pageId="37" pageNumber="38">Cortés</collectingRegion>
(39):
</emphasis>
San Pedro Sula. 
<emphasis id="18D4243FFD7AFFA4AE651AA0FDA1B871" bold="true" box="[340,543,1014,1040]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AE651AA0FE6FB871" box="[340,465,1014,1040]" country="Honduras" name="El Paraiso" pageId="37" pageNumber="38">El Paraiso</collectingRegion>
(101):
</emphasis>
Barrio Las Flores; Capire (
<quantity id="ED5855C8FD7AFFA4AC681AA0FC17B871" box="[857,937,1014,1040]" metricMagnitude="3" metricUnit="m" metricValue="8.3" pageId="37" pageNumber="38" unit="km" value="8.3">8.3 km</quantity>
SE); Danlí, El Barro; Morocelí (
<quantity id="ED5855C8FD7AFFA4AA101AA0FAD4B871" box="[1313,1386,1014,1040]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="km" value="10.0">10 km</quantity>
W). 
<emphasis id="18D4243FFD7AFFA4AFA61D4CFE7CB855" bold="true" box="[151,450,1049,1076]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AFA61D4CFE3FB855" box="[151,385,1049,1076]" country="Honduras" name="Francisco Morazan" pageId="37" pageNumber="38">Francisco Morazán</collectingRegion>
(25):
</emphasis>
Casa Blanca; El Zamorano; La Lima; Santa Lucía; Tatumbla; Villa San Francisco. 
<emphasis id="18D4243FFD7AFFA4AA491D4CFEA9B839" bold="true" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AA491D4CFF5CB839" country="Honduras" name="Intibuca" pageId="37" pageNumber="38">Intibucá</collectingRegion>
(1):
</emphasis>
Jesús de Otoro. 
<emphasis id="18D4243FFD7AFFA4AEEC1D68FD21B839" bold="true" box="[477,671,1085,1112]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AEEC1D68FDD4B839" box="[477,618,1085,1112]" country="Honduras" name="Isla Roatan" pageId="37" pageNumber="38">Isla Roatán</collectingRegion>
(1):
</emphasis>
Jicaro Galán. 
<emphasis id="18D4243FFD7AFFA4AC7A1D68FC59B839" bold="true" box="[843,999,1086,1112]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AC7A1D68FC0DB839" box="[843,947,1086,1112]" country="Honduras" name="Lempira" pageId="37" pageNumber="38">Lempira</collectingRegion>
(1):
</emphasis>
Erandique. 
<emphasis id="18D4243FFD7AFFA4AB471D68FA86B839" bold="true" box="[1142,1336,1086,1112]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AB471D68FABAB839" box="[1142,1284,1086,1112]" country="Honduras" name="Ocotepeque" pageId="37" pageNumber="38">Ocotepeque</collectingRegion>
(1):
</emphasis>
Reserva Güisayote. 
<emphasis id="18D4243FFD7AFFA4AE261D34FE3BB81D" bold="true" box="[279,389,1122,1148]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AE261D34FEEDB81D" box="[279,339,1122,1148]" country="Honduras" name="Valle" pageId="37" pageNumber="38">Valle</collectingRegion>
(1):
</emphasis>
Nacaome. 
<emphasis id="18D4243FFD7AFFA4AD351D34FDD1B81D" bold="true" box="[516,623,1122,1148]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AD351D34FD83B81D" box="[516,573,1122,1148]" country="Honduras" name="Yoro" pageId="37" pageNumber="38">Yoro</collectingRegion>
(1):
</emphasis>
Parque Nacional Pico Pijol. 
<emphasis id="18D4243FFD7AFFA4AC841D34FA9EB81D" bold="true" box="[949,1312,1122,1148]" pageId="37" pageNumber="38">
<collectingCountry id="52B7B8BDFD7AFFA4AC841D34FB95B81D" box="[949,1067,1122,1148]" name="Mexico" pageId="37" pageNumber="38">MEXICO</collectingCountry>
(327). 
<collectingRegion id="E86436CFFD7AFFA4AB4F1D34FB5EB81D" box="[1150,1248,1122,1148]" country="Mexico" name="Chiapas" pageId="37" pageNumber="38">Chiapas</collectingRegion>
(96):
</emphasis>
Aguacero; Albino Corzo; Berriozabal; Cahuare; Chiapa de Corzo; Cinco Cerros; Cintalapa; Comitán de Domínguez; El Chorreadero; El Jardín; Laguna Bélgica; Musté; Ocosingo; Ocozocoautla; Pijijiapan; Rizo de Oro; Suchiapa; Tapachula; Tuxtla Gutiérrez; Villaflores. 
<emphasis id="18D4243FFD7AFFA4AEDB1D98FDCBB889" bold="true" box="[490,629,1230,1256]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AEDB1D98FDFFB889" box="[490,577,1230,1256]" country="Mexico" name="Colima" pageId="37" pageNumber="38">Colima</collectingRegion>
(5):
</emphasis>
Colima; Comala. 
<emphasis id="18D4243FFD7AFFA4AC7B1D98FC42B889" bold="true" box="[842,1020,1230,1256]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AC7B1D98FC04B889" box="[842,954,1230,1256]" country="Mexico" name="Guerrero" pageId="37" pageNumber="38">Guerrero</collectingRegion>
(86):
</emphasis>
Acapulco; Acahuizotla; Aserradero; Caletilla; Chilpancingo; Juan R. Escudero, El Terrero; Mochitlán; Quechultenango; Rodecia; San Luís Acatlán, Tlaxcalixtlahuaca; Tecpán; Zihuatajeno. 
<emphasis id="18D4243FFD7AFFA4AD5A1C40FD41B951" bold="true" box="[619,767,1302,1328]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AD5A1C40FD74B951" box="[619,714,1302,1328]" country="Mexico" name="Hidalgo" pageId="37" pageNumber="38">Hidalgo</collectingRegion>
(2):
</emphasis>
Mixquiahuala. 
<emphasis id="18D4243FFD7AFFA4AC861C40FBF5B951" bold="true" box="[951,1099,1302,1328]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AC861C40FBB6B951" box="[951,1032,1302,1328]" country="Mexico" name="Jalisco" pageId="37" pageNumber="38">Jalisco</collectingRegion>
(54):
</emphasis>
Casimiro Castillo; Chamela; Guadalajara; La Cañada; Melaque; Nevado del Colima; Puerto Vallarta; Tomatlán. 
<emphasis id="18D4243FFD7AFFA4AB771C6FFB71B935" bold="true" box="[1094,1231,1337,1364]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AB771C6FFB23B935" box="[1094,1181,1337,1364]" country="Mexico" name="Mexico" pageId="37" pageNumber="38">México</collectingRegion>
(4):
</emphasis>
Soyaniquilpan de Juárez; Valle de Bravo. 
<emphasis id="18D4243FFD7AFFA4AE931C0BFDD8B919" bold="true" box="[418,614,1373,1400]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AE931C0BFD98B919" box="[418,550,1373,1400]" country="Mexico" name="Michoacan de Ocampo" pageId="37" pageNumber="38">Michoacán</collectingRegion>
(12):
</emphasis>
Apatzingan; Chinicuila; Coahuayana; San Telmo, Ojo de Agua. 
<emphasis id="18D4243FFD7AFFA4AA0A1C08FF71B9FD" bold="true" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AA0A1C08FA22B919" box="[1339,1436,1374,1400]" country="Mexico" name="Morelos" pageId="37" pageNumber="38">Morelos</collectingRegion>
(16):
</emphasis>
Acamilpa; Jantetelco; Jojutla de Juárez; Tlaltizapan. 
<emphasis id="18D4243FFD7AFFA4AC021CD4FC7CB9FD" bold="true" box="[819,962,1410,1436]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AC021CD4FC31B9FD" box="[819,911,1410,1436]" country="Mexico" name="Nayarit" pageId="37" pageNumber="38">Nayarit</collectingRegion>
(4):
</emphasis>
San Blas (
<quantity id="ED5855C8FD7AFFA4AB701CD4FBC0B9FD" box="[1089,1150,1410,1436]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="37" pageNumber="38" unit="mi" value="9.0">9 mi.</quantity>
E); Sayulita; Volcán Ceboruco. 
<emphasis id="18D4243FFD7AFFA4AFC51CF0FE30B9A1" bold="true" box="[244,398,1446,1472]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AFC51CF0FEF0B9A1" box="[244,334,1446,1472]" country="Mexico" name="Oaxaca" pageId="37" pageNumber="38">Oaxaca</collectingRegion>
(41):
</emphasis>
Almoloya; Juchitán de Zaragoza; Matías Romero (
<quantity id="ED5855C8FD7AFFA4ACE31CF0FBA5B9A1" box="[978,1051,1446,1472]" metricMagnitude="4" metricUnit="m" metricValue="3.7014912" pageId="37" pageNumber="38" unit="mi" value="23.0">23 mi.</quantity>
S); Oaxaca; Puerto Ángel; Puerto Escondido; Salina Cruz; Tapanatepec (
<quantity id="ED5855C8FD7AFFA4AD681C9CFD29B985" box="[601,663,1482,1508]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="37" pageNumber="38" unit="mi" value="8.0">8 mi.</quantity>
W); Tehuantepec. 
<emphasis id="18D4243FFD7AFFA4AC471C9CFC43B985" bold="true" box="[886,1021,1482,1508]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AC471C9CFC76B985" box="[886,968,1482,1508]" country="Mexico" name="Puebla" pageId="37" pageNumber="38">Puebla</collectingRegion>
(1):
</emphasis>
Tehuixtla. 
<emphasis id="18D4243FFD7AFFA4ABB21C9CFAC7B985" bold="true" box="[1155,1401,1481,1508]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ABB21C9CFAFAB985" box="[1155,1348,1481,1508]" country="Mexico" name="San Luis Potosi" pageId="37" pageNumber="38">San Luís Potosí</collectingRegion>
(1):
</emphasis>
El Salto. 
<emphasis id="18D4243FFD7AFFA4AFD11CB8FE20BA69" bold="true" box="[224,414,1518,1544]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AFD11CB8FED5BA69" box="[224,363,1518,1544]" country="Mexico" name="Tamaulipas" pageId="37" pageNumber="38">Tamaulipas</collectingRegion>
(2):
</emphasis>
Ciudad Victoria. 
<emphasis id="18D4243FFD7AFFA4AD581CB8FCB7BA69" bold="true" box="[617,777,1518,1544]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AD581CB8FD68BA69" box="[617,726,1518,1544]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="37" pageNumber="38">Veracruz</collectingRegion>
(3):
</emphasis>
Córdoba; Santiago Tuxtla. 
<emphasis id="18D4243FFD7AFFA4AB751CB8FF7FBA4D" bold="true" pageId="37" pageNumber="38">
<collectingCountry id="52B7B8BDFD7AFFA4AB751CB8FB4FBA69" box="[1092,1265,1518,1544]" name="Nicaragua" pageId="37" pageNumber="38">NICARAGUA</collectingCountry>
(618). 
<collectingRegion id="E86436CFFD7AFFA4AA741CB8FA22BA69" box="[1349,1436,1518,1544]" country="Nicaragua" name="Carazo" pageId="37" pageNumber="38">Carazo</collectingRegion>
(4):
</emphasis>
San Francisco; 
<collectingRegion id="E86436CFFD7AFFA4AE441F44FE40BA4C" box="[373,510,1554,1581]" inferredBySuffix="true" pageId="37" pageNumber="38">San Marcos</collectingRegion>
. 
<emphasis id="18D4243FFD7AFFA4AD381F44FD74BA4D" bold="true" box="[521,714,1554,1580]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AD381F44FD24BA4D" box="[521,666,1554,1580]" country="Nicaragua" name="Chinandega" pageId="37" pageNumber="38">Chinandega</collectingRegion>
(8):
</emphasis>
Chichigalpa; Chinandega; San Benito; El Viejo. 
<emphasis id="18D4243FFD7AFFA4ABC31F44FA22BA4D" bold="true" box="[1266,1436,1554,1580]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ABC31F44FAE3BA4D" box="[1266,1373,1554,1580]" country="Nicaragua" name="Granada" pageId="37" pageNumber="38">Granada</collectingRegion>
(46):
</emphasis>
Nandaime; Volcán Mombacho. 
<emphasis id="18D4243FFD7AFFA4AECB1F63FD3EBA31" bold="true" box="[506,640,1589,1616]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AECB1F63FD89BA31" box="[506,567,1589,1616]" country="Nicaragua" name="Leon" pageId="37" pageNumber="38">León</collectingRegion>
(111):
</emphasis>
<collectingRegion id="E86436CFFD7AFFA4ADB91F61FD66BA30" box="[648,728,1591,1617]" inferredBySuffix="true" pageId="37" pageNumber="38">La Paz</collectingRegion>
; Laguna de Asososca; Las Marías; Lechecuago; León; 
<taxonomicName id="EDA083AEFD7AFFA4AA701F60FA22BA31" authority="El" authorityName="El" box="[1345,1436,1590,1616]" class="Insecta" family="Staphylinidae" genus="Mina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">Mina El</taxonomicName>
Limón; Posoltega; Santa Clara; Valle de las Zapatas. 
<emphasis id="18D4243FFD7AFFA4ADC41F0CFC19BA15" bold="true" box="[757,935,1626,1652]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4ADC41F0CFCD9BA15" box="[757,871,1626,1652]" country="Nicaragua" name="Managua" pageId="37" pageNumber="38">Managua</collectingRegion>
(34):
</emphasis>
La Calera; La Chinampa; Laguna de Xiloa; Managua; Matano; Quinta Salvatierra; Tipitapa. 
<emphasis id="18D4243FFD7AFFA4AD8C1F28FCD7BAF9" bold="true" box="[701,873,1662,1688]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AD8C1F28FCA4BAF9" box="[701,794,1662,1688]" country="Nicaragua" name="Masaya" pageId="37" pageNumber="38">Masaya</collectingRegion>
(412):
</emphasis>
Las Flores; Masatepe. 
<emphasis id="18D4243FFD7AFFA4AB431F28FA9BBAF9" bold="true" box="[1138,1317,1662,1688]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AB431F28FB4DBAF9" box="[1138,1267,1662,1688]" country="Nicaragua" name="Matagalpa" pageId="37" pageNumber="38">Matagalpa</collectingRegion>
(3):
</emphasis>
Las Pilas; Matagalpa; San Isidro. 
<emphasis id="18D4243FFD7AFFA4AEAE1FF4FD76BADD" bold="true" box="[415,712,1697,1724]" pageId="37" pageNumber="38">
<collectingCountry id="52B7B8BDFD7AFFA4AEAE1FF4FDA6BADD" box="[415,536,1698,1724]" name="Panama" pageId="37" pageNumber="38">PANAMA</collectingCountry>
(3). 
<collectingRegion id="E86436CFFD7AFFA4AD7E1FF7FD29BADD" box="[591,663,1697,1724]" country="Panama" name="Colon" pageId="37" pageNumber="38">Colón</collectingRegion>
(2):
</emphasis>
Madden Dam; Pipeline Road (
<quantity id="ED5855C8FD7AFFA4AB181FF4FBDDBADD" box="[1065,1123,1698,1724]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="km" value="2.0">2 km</quantity>
W Gamboa); 
<emphasis id="18D4243FFD7AFFA4AA351FF7FA23BADD" bold="true" box="[1284,1437,1697,1724]" pageId="37" pageNumber="38">
<collectingRegion id="E86436CFFD7AFFA4AA351FF7FAD5BADD" box="[1284,1387,1697,1724]" country="Panama" name="Chiriqui" pageId="37" pageNumber="38">Chiriquí</collectingRegion>
(1):
</emphasis>
Finca La Suiza (
<quantity id="ED5855C8FD7AFFA4AE7F1F90FE23BA81" box="[334,413,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="5.3" pageId="37" pageNumber="38" unit="km" value="5.3">5.3 km</quantity>
N Los Planes).
</paragraph>
<paragraph id="2A1FF82DFD7AFFA4AFF61FBCFB95BB11" blockId="37.[151,1437,150,1905]" pageId="37" pageNumber="38">
<emphasis id="18D4243FFD7AFFA4AFF61FBCFE39BB65" bold="true" box="[199,391,1770,1796]" pageId="37" pageNumber="38">Natural history.</emphasis>
One examined specimen was collected in detritus of 
<taxonomicName id="EDA083AEFD7AFFA4ACDB1FBDFB55BB65" authority="(Smith)" baseAuthorityName="Smith" box="[1002,1259,1770,1797]" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="18D4243FFD7AFFA4ACDB1FBDFB32BB65" box="[1002,1164,1771,1797]" italics="true" pageId="37" pageNumber="38">Atta mexicana</emphasis>
(Smith)
</taxonomicName>
(
<taxonomicName id="EDA083AEFD7AFFA4ABCB1FBCFA27BB65" box="[1274,1433,1770,1796]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
: 
<taxonomicName id="EDA083AEFD7AFFA4AFA61E58FEA2BB49" box="[151,284,1806,1832]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
). The larval stages were described by 
<bibRefCitation id="4E3185DCFD7AFFA4ADFD1E58FBA7BB49" author="Ramirez-Salinas, C. &amp; Pacheco-Flores, C. &amp; Castro-Ramirez, A. E. &amp; Moron, M. A." box="[716,1049,1806,1832]" pageId="37" pageNumber="38" pagination="617 - 625" refId="ref92549" refString="Ramirez-Salinas, C., Pacheco-Flores, C., Castro-Ramirez, A. E. &amp; Moron, M. A. (2010) Descripcion de los estados inmaduros de Ligyrus nasutus (Burmeister, 1847) (Coleoptera: Melolonthidae: Dynastinae) con informacion sobre su biologia. Acta Zoologica Mexicana, New Series, 26, 617 - 625." type="journal article" year="2010">
Ramírez-Salinas 
<emphasis id="18D4243FFD7AFFA4ACBC1E59FC79BB49" box="[909,967,1806,1832]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
(2010)
</bibRefCitation>
(as 
<taxonomicName id="EDA083AEFD7AFFA4AB7B1E59FB00BB49" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1098,1214,1807,1832]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="18D4243FFD7AFFA4AB7B1E59FB00BB49" box="[1098,1214,1807,1832]" italics="true" pageId="37" pageNumber="38">L. nasutus</emphasis>
</taxonomicName>
), who found larvae in agricultural soils where cattle grazed. It is a common species in tropical deciduous forests between sea level and 
<quantity id="ED5855C8FD7AFFA4AFA61E00FF50BB10" box="[151,238,1878,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="37" pageNumber="38" unit="m" value="1800.0">1800 m</quantity>
, but mainly below 
<quantity id="ED5855C8FD7AFFA4AEF41E00FDB2BB10" box="[453,524,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="38" unit="m" value="500.0">500 m</quantity>
(
<bibRefCitation id="4E3185DCFD7AFFA4AD2B1E00FC91BB11" author="Ratcliffe, B. C. &amp; Moron, M. A." box="[538,815,1878,1904]" pageId="37" pageNumber="38" pagination="53 - 98" refId="ref92970" refString="Ratcliffe, B. C. &amp; Moron, M. A. (1997) Dynastinae. In: Moron, M. A., Ratcliffe, B. C. &amp; Deloya, C. (Eds.), Atlas de los escarabajos de Mexico. Coleoptera: Lamellicornia. Vol. I. Familia Melolonthidae. Subfamilias Rutelinae, Dynastinae, Cetoniinae, Trichinae, Valginae y Melolonthinae. Comision Nacional para el Conocimiento y Uso de la Biodiversidad (CONABIO), Mexico City, pp. 53 - 98." type="book chapter" year="1997">Ratcliffe &amp; Morón 1997</bibRefCitation>
; 
<bibRefCitation id="4E3185DCFD7AFFA4AC0A1E00FB9EBB11" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Cano, E. B." box="[827,1056,1878,1904]" pageId="37" pageNumber="38" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. &amp; Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
Ratcliffe 
<emphasis id="18D4243FFD7AFFA4AC951E01FC62BB11" box="[932,988,1878,1904]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>