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<document id="A9CFD31B521759D814779FBB4E754B86" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. &amp; Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD1AFFC7AFA61F8DFE16B9EA" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomarus selanderi" docType="treatment" docVersion="8" lastPageNumber="71" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="70" updateTime="1699396337306" updateUser="plazi">
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<mods:titleInfo id="DA3A7A636F7B72E3CC2788AE18C569BD">
<mods:title id="998B68E727EFDB471538F2C8C04AB5DC">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
</mods:titleInfo>
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<mods:namePart id="0CEB4888E29F881C60049A9EE782FEC8">López-García, Margarita M.</mods:namePart>
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<mods:nameIdentifier id="9EA1C1774CB07A867F04EF6A43EB2F78" type="email">margaralopezg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6C59F080FD064C643734CE6E0622D15F">Deloya, Cuauhtémoc</mods:namePart>
<mods:nameIdentifier id="49EA47341CB4EB8C6E10A1CAFE27A15E" type="ORCID">0000-0002-4774-140X</mods:nameIdentifier>
<mods:nameIdentifier id="82449042BBA884F2962FD86EA74F8EC8" type="email">cuauhtemoc.deloya@inecol.mx</mods:nameIdentifier>
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<mods:title id="D16A31A1AC317BEA6433BF0E6739A951">Zootaxa</mods:title>
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<mods:part id="B79D8E4658CA283C4A03E959F3C76A6D">
<mods:date id="BDEC4CE543FF0E88F772E8B6A257D712">2022</mods:date>
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<mods:number id="1392DA15AC94980FE2CF5DB5F8165E74">2022-11-22</mods:number>
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<mods:classification id="5BE5A80D8ED75AE35D0A963E8FAB1FC0">journal article</mods:classification>
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<treatment id="A209493BFD1AFFC7AFA61F8DFE16B9EA" ID-DOI="http://doi.org/10.5281/zenodo.7345129" ID-GBIF-Taxon="205794529" ID-Zenodo-Dep="7345129" LSID="urn:lsid:plazi:treatment:A209493BFD1AFFC7AFA61F8DFE16B9EA" httpUri="http://treatment.plazi.org/id/A209493BFD1AFFC7AFA61F8DFE16B9EA" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
<subSubSection id="62BAABA6FD1AFFC4AFA61F8DFDD4BA95" box="[151,618,1754,1781]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="2A1FF82DFD1AFFC4AFA61F8DFDD4BA95" blockId="69.[151,638,1754,1817]" box="[151,618,1754,1781]" pageId="69" pageNumber="70">
<heading id="71574F41FD1AFFC4AFA61F8DFDD4BA95" bold="true" box="[151,618,1754,1781]" fontSize="11" level="2" pageId="69" pageNumber="70" reason="4">
<taxonomicName id="EDA083AEFD1AFFC4AFA61F8DFDD4BA95" authority="(Cartwright, 1959)" baseAuthorityName="Cartwright" baseAuthorityYear="1959" box="[151,618,1754,1781]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="selanderi">
<emphasis id="18D4243FFD1AFFC4AFA61F8DFDD4BA95" bold="true" box="[151,618,1754,1781]" pageId="69" pageNumber="70">
<emphasis id="18D4243FFD1AFFC4AFA61F8DFEC6BA94" bold="true" box="[151,376,1754,1781]" italics="true" pageId="69" pageNumber="70">Tomarus selanderi</emphasis>
(
<bibRefCitation id="4E3185DCFD1AFFC4AEB61F8CFDDFBA95" author="Cartwright, O. L." box="[391,609,1754,1780]" pageId="69" pageNumber="70" pagination="515 - 541" refId="ref88844" refString="Cartwright, O. L. (1959) Scarab beetles of the genus Bothynus in the United States (Coleoptera: Sacarabaeidae). Proceedings of the United States National Museum, 108, 515 - 541." type="journal article" year="1959">Cartwright, 1959</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1AFFC4AFA61FA9FDC0BB78" box="[151,638,1790,1817]" pageId="69" pageNumber="70" type="description">
<paragraph id="2A1FF82DFD1AFFC4AFA61FA9FDC0BB78" blockId="69.[151,638,1754,1817]" box="[151,638,1790,1817]" pageId="69" pageNumber="70">
(
<figureCitation id="B29BE4A8FD1AFFC4AFAE1FA9FF44BB78" box="[159,250,1791,1817]" captionStart="FIGURE 2" captionStartId="82.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1548]" captionTargetId="figure-24@82.[151,1436,181,1560]" captionTargetPageId="82" captionText="FIGURE 2. Labium. A) Ligyrus (Anagrylius) moroni, B) Ligyrus (Ligyrus) allonasutus, C) L. (Ligyrus) gibbosus, D) L. (Ligyrus) villosus, E) L. (Ligyrodes) relictus, F) Proculigyrus cicatricosus, G) Euligyrus ebenus, H) E. similis, I) Tomarus gyas, J) Tomarus selanderi, K) Tomarus subtropicus. Abbreviations: LL: Ligular lobes, LP: Labial palpus, M: Mentum, S: Suture." figureDoi="http://doi.org/10.5281/zenodo.7345015" httpUri="https://zenodo.org/record/7345015/files/figure.png" pageId="69" pageNumber="70">Figs. 2J</figureCitation>
,
<figureCitation id="B29BE4A8FD1AFFC4AE371FA9FE94BB78" box="[262,298,1791,1817]" captionStart="FIGURE 4" captionStartId="83.[151,250,1872,1896]" captionTargetBox="[151,1437,1118,1848]" captionTargetId="figure-81@83.[151,1437,1118,1848]" captionTargetPageId="83" captionText="FIGURE 4. Maxilla. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus gyas, G) Tomarus maternus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345023" httpUri="https://zenodo.org/record/7345023/files/figure.png" pageId="69" pageNumber="70">4H</figureCitation>
,
<figureCitation id="B29BE4A8FD1AFFC4AE071FA9FEEABB78" box="[310,340,1791,1817]" captionStart="FIGURE 6" captionStartId="84.[151,250,1963,1987]" captionTargetBox="[185,1409,1024,1940]" captionTargetId="figure-74@84.[178,1409,1015,1940]" captionTargetPageId="84" captionText="FIGURE 6. Labrum. A) Ligyrus (Ligyrus) burmeisteri, B) L. (Ligyrus) villosus, C) Ligyrus (Ligyrodes) relictus, D) Tomarus gyas, E) T. maternus, F) T. selanderi, G) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345029" httpUri="https://zenodo.org/record/7345029/files/figure.png" pageId="69" pageNumber="70">6F</figureCitation>
,
<figureCitation id="B29BE4A8FD1AFFC4AE6E1FA9FE2CBB78" box="[351,402,1791,1817]" captionStart="FIGURE 14" captionStartId="89.[151,250,1677,1701]" captionTargetBox="[152,1432,1161,1649]" captionTargetId="figure-90@89.[152,1436,1161,1653]" captionTargetPageId="89" captionText="FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345059" httpUri="https://zenodo.org/record/7345059/files/figure.png" pageId="69" pageNumber="70">14G</figureCitation>
,
<figureCitation id="B29BE4A8FD1AFFC4AEAC1FA9FE6EBB78" box="[413,464,1791,1817]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="69" pageNumber="70">16Q</figureCitation>
,
<figureCitation id="B29BE4A8FD1AFFC4AEED1FA9FDB0BB78" box="[476,526,1791,1817]" captionStart="FIGURE 20" captionStartId="95.[151,250,1225,1249]" captionTargetBox="[159,1435,191,1201]" captionTargetId="figure-17@95.[151,1436,181,1202]" captionTargetPageId="95" captionText="FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345083" httpUri="https://zenodo.org/record/7345083/files/figure.png" pageId="69" pageNumber="70">20H</figureCitation>
,
<figureCitation id="B29BE4A8FD1AFFC4AD2B1FA9FDF3BB78" box="[538,589,1791,1817]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="69" pageNumber="70">28H</figureCitation>
;
<figureCitation id="B29BE4A8FD1AFFC4AD681FA8FDC8BB78" box="[601,630,1790,1817]" captionStart="FIGURE 45" captionStartId="113.[151,250,1077,1101]" captionTargetBox="[207,1391,181,1048]" captionTargetId="figure-17@113.[197,1391,181,1053]" captionTargetPageId="113" captionText="FIGURE 45. Distributional records of Tomarus discrepans and T. selanderi." figureDoi="http://doi.org/10.5281/zenodo.7345168" httpUri="https://zenodo.org/record/7345168/files/figure.png" pageId="69" pageNumber="70">45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1AFFC4AFA61E15FC90BB3A" box="[151,814,1859,1883]" pageId="69" pageNumber="70" type="reference_group">
<paragraph id="2A1FF82DFD1AFFC4AFA61E15FC90BB3A" blockId="69.[151,1437,1859,2037]" box="[151,814,1859,1883]" pageId="69" pageNumber="70">
<treatmentCitationGroup id="0AB0DF03FD1AFFC4AFA61E15FC90BB3A" box="[151,814,1859,1883]" pageId="69" pageNumber="70">
<treatmentCitation id="AB01DE3CFD1AFFC4AFA61E15FDFABB3A" author="Cartwright, O. L." box="[151,580,1859,1883]" page="537" pageId="69" pageNumber="70" year="1959">
<taxonomicName id="EDA083AEFD1AFFC4AFA61E15FDFABB3A" authority="Cartwright, 1959: 537" authorityName="Cartwright" authorityPageNumber="537" authorityYear="1959" box="[151,580,1859,1883]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="selanderi">
<emphasis id="18D4243FFD1AFFC4AFA61E15FEE4BB3A" box="[151,346,1859,1883]" italics="true" pageId="69" pageNumber="70">Bothynus selanderi</emphasis>
<bibRefCitation id="4E3185DCFD1AFFC4AE511E15FDFABB3A" author="Cartwright, O. L." box="[352,580,1859,1883]" pageId="69" pageNumber="70" pagination="515 - 541" refId="ref88844" refString="Cartwright, O. L. (1959) Scarab beetles of the genus Bothynus in the United States (Coleoptera: Sacarabaeidae). Proceedings of the United States National Museum, 108, 515 - 541." type="journal article" year="1959">Cartwright, 1959: 537</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<collectingMunicipality id="CA7B6257FD1AFFC4AD7C1E15FD1FBB3A" box="[589,673,1859,1883]" pageId="69" pageNumber="70">Original</collectingMunicipality>
combination.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1AFFC7AC021E15FC4EBD6A" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="2A1FF82DFD1AFFC7AC021E15FC4EBD6A" blockId="69.[151,1437,1859,2037]" lastBlockId="70.[199,1436,150,267]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
<location id="2F7FAEF6FD1AFFC4AC021E15FCD9BB3A" LSID="urn:lsid:plazi:treatment:A209493BFD1AFFC7AFA61F8DFE16B9EA:2F7FAEF6FD1AFFC4AC021E15FCD9BB3A" box="[819,871,1859,1883]" country="Mexico" municipality="Original" name="Male" pageId="69" pageNumber="70" stateProvince="Sinaloa">Male</location>
<typeStatus id="F51B468FFD1AFFC4AC5F1E15FC79BB3A" box="[878,967,1859,1883]" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
(USNM) “
<collectingCountry id="52B7B8BDFD1AFFC4AB071E15FB22BB3A" box="[1078,1180,1859,1883]" name="Mexico" pageId="69" pageNumber="70">MEXICO</collectingCountry>
/ Los Mochis /
<collectingRegion id="E86436CFFD1AFFC4AA0D1E15FA36BB3A" box="[1340,1416,1859,1883]" country="Mexico" name="Sinaloa" pageId="69" pageNumber="70">Sinaloa</collectingRegion>
// at light /
<date id="5E1EDEEDFD1AFFC4AE111E34FE1FBB1B" box="[288,417,1890,1914]" pageId="69" pageNumber="70" value="1955-07-22">
<collectingDate id="4E5A2705FD1AFFC4AE111E34FE1FBB1B" box="[288,417,1890,1914]" pageId="69" pageNumber="70" value="1955-07-22">22 July 1955</collectingDate>
</date>
// RB&amp;JM Selander // Type No / 63912 / USNM //
<typeStatus id="F51B468FFD1AFFC4AC951E34FB94BB1B" box="[932,1066,1890,1914]" pageId="69" pageNumber="70" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="EDA083AEFD1AFFC4AB0D1E34FB23BB1B" authorityName="Hope" authorityYear="1837" box="[1084,1181,1890,1914]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Bothynus</taxonomicName>
/ selanderi / Cartwright”.
<collectorName id="87559DFBFD1AFFC4AFF61ED6FEACBBF9" box="[199,274,1920,1944]" pageId="69" pageNumber="70">Female</collectorName>
<typeStatus id="F51B468FFD1AFFC4AE261ED6FED7BBF9" box="[279,361,1920,1944]" pageId="69" pageNumber="70" type="allotype">allotype</typeStatus>
(USNM) “
<collectingCountry id="52B7B8BDFD1AFFC4AEE71ED7FD82BBF8" box="[470,572,1921,1945]" name="Mexico" pageId="69" pageNumber="70">MEXICO</collectingCountry>
/ Los Mochis /
<collectingRegion id="E86436CFFD1AFFC4ADE71ED6FC9CBBF9" box="[726,802,1920,1944]" country="Mexico" name="Sinaloa" pageId="69" pageNumber="70">Sinaloa</collectingRegion>
// at light /
<date id="5E1EDEEDFD1AFFC4ACA01ED7FBACBBF9" box="[913,1042,1920,1945]" pageId="69" pageNumber="70" value="1955-07-22">
<collectingDate id="4E5A2705FD1AFFC4ACA01ED7FBACBBF9" box="[913,1042,1920,1945]" pageId="69" pageNumber="70" value="1955-07-22">22 July 1955</collectingDate>
</date>
// RB&amp;JM Selander // Type No / 63912 / USNM //
<typeStatus id="F51B468FFD1AFFC4AE091EC9FE05BBD6" box="[312,443,1951,1975]" pageId="69" pageNumber="70" type="allotype">ALLOTYPE</typeStatus>
/
<taxonomicName id="EDA083AEFD1AFFC4AEFE1EC9FD8EBBD6" authority="Cartwright, 1959: 537" authorityName="Cartwright" authorityPageNumber="537" authorityYear="1959" box="[463,560,1951,1975]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="selanderi">Bothynus</taxonomicName>
/ selanderi / Cartwright”.
<collectorName id="87559DFBFD1AFFC4AC081EC9FC3DBBD6" box="[825,899,1951,1975]" pageId="69" pageNumber="70">Twelve</collectorName>
male and
<specimenCount id="3CA633A4FD1AFFC4ACDD1EC9FBEFBBD6" box="[1004,1105,1951,1975]" pageId="69" pageNumber="70" type="female">21 female</specimenCount>
<typeStatus id="F51B468FFD1AFFC4AB661EF6FB06BBD9" box="[1111,1208,1952,1976]" pageId="69" pageNumber="70" type="paratype">paratypes</typeStatus>
(USNM) “
<collectingCountry id="52B7B8BDFD1AFFC4AA191EC9FA30BBD6" box="[1320,1422,1951,1975]" name="Mexico" pageId="69" pageNumber="70">MEXICO</collectingCountry>
/ Los Mochis /
<collectingRegion id="E86436CFFD1AFFC4AE651EE8FE1EBBB7" box="[340,416,1982,2006]" country="Mexico" name="Sinaloa" pageId="69" pageNumber="70">Sinaloa</collectingRegion>
// at light /
<date id="5E1EDEEDFD1AFFC4AD261EE8FD24BBB7" box="[535,666,1982,2006]" pageId="69" pageNumber="70" value="1955-07-22">
<collectingDate id="4E5A2705FD1AFFC4AD261EE8FD24BBB7" box="[535,666,1982,2006]" pageId="69" pageNumber="70" value="1955-07-22">22 July 1955</collectingDate>
</date>
// RB&amp;JM Selander // Type No / 63912 / USNM //
<typeStatus id="F51B468FFD1AFFC4AB981EE8FA99BBB7" box="[1193,1319,1982,2006]" pageId="69" pageNumber="70" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="EDA083AEFD1AFFC4AA0A1EE8FA22BBB7" authorityName="Hope" authorityYear="1837" box="[1339,1436,1982,2006]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Bothynus</taxonomicName>
/ selanderi / Cartwright”. “
<collectingCountry id="52B7B8BDFD1AFFC4AEE21E8BFD87BB94" box="[467,569,2013,2037]" name="Mexico" pageId="69" pageNumber="70">MEXICO</collectingCountry>
/ Mazatlán /
<collectingRegion id="E86436CFFD1AFFC4AD8E1E8AFCB5BB95" box="[703,779,2012,2036]" country="Mexico" name="Sinaloa" pageId="69" pageNumber="70">Sinaloa</collectingRegion>
//
<date id="5E1EDEEDFD1AFFC4AC171E8BFC14BB95" box="[806,938,2012,2037]" pageId="69" pageNumber="70" value="1955-07-21">
<collectingDate id="4E5A2705FD1AFFC4AC171E8BFC14BB95" box="[806,938,2012,2037]" pageId="69" pageNumber="70" value="1955-07-21">21 July 1955</collectingDate>
</date>
// RB&amp;JM Selander // Type No / 63912 / USNM //
<typeStatus id="F51B468FFD19FFC7AFEC19C1FEE5BCCE" box="[221,347,151,175]" pageId="70" pageNumber="71" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="EDA083AEFD19FFC7AE4019C1FE6CBCCE" authorityName="Hope" authorityYear="1837" box="[369,466,151,175]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">Bothynus</taxonomicName>
/ selanderi / Cartwright”.
<collectorName id="87559DFBFD19FFC7ADD019C1FCABBCCE" box="[737,789,151,175]" pageId="70" pageNumber="71">Male</collectorName>
<typeStatus id="F51B468FFD19FFC7AC2C19C1FCCABCCE" box="[797,884,151,175]" pageId="70" pageNumber="71" type="paratype">paratype</typeStatus>
(MNHN) “Los Mochis /
<collectingRegion id="E86436CFFD19FFC7AB4C19C1FB72BCCE" box="[1149,1228,151,175]" country="Mexico" name="Sinaloa" pageId="70" pageNumber="71">Sinaloa</collectingRegion>
, Mex /
<date id="5E1EDEEDFD19FFC7AA2D19C1FA22BCCF" box="[1308,1436,150,175]" pageId="70" pageNumber="71" value="1955-07-22">
<collectingDate id="4E5A2705FD19FFC7AA2D19C1FA22BCCF" box="[1308,1436,150,175]" pageId="70" pageNumber="71" value="1955-07-22">22-VII-1955</collectingDate>
</date>
// R&amp;J Selander //
<typeStatus id="F51B468FFD19FFC7AEBD19E3FDB4BCAC" box="[396,522,181,205]" pageId="70" pageNumber="71" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="EDA083AEFD19FFC7AD1019E3FD3CBCAC" authorityName="Hope" authorityYear="1837" box="[545,642,181,205]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">Bothynus</taxonomicName>
/ selanderi / Cartwright //
<typeStatus id="F51B468FFD19FFC7ACAB19E3FBA6BCAC" box="[922,1048,181,205]" pageId="70" pageNumber="71" type="paratype">PARATYPE</typeStatus>
// MNHN /
<specimenCode id="7A065056FD19FFC7ABAD19E3FB4ABCAC" box="[1180,1268,181,205]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="70" pageNumber="71">EC6732</specimenCode>
”.
<collectorName id="87559DFBFD19FFC7AA3919E3FA82BCAC" box="[1288,1340,181,205]" pageId="70" pageNumber="71">Male</collectorName>
<typeStatus id="F51B468FFD19FFC7AA7419E0FA22BCAF" box="[1349,1436,182,206]" pageId="70" pageNumber="71" type="paratype">paratype</typeStatus>
(MNHN) “Los Mochis /
<collectingRegion id="E86436CFFD19FFC7AEF51982FDADBC8D" box="[452,531,212,236]" country="Mexico" name="Sinaloa" pageId="70" pageNumber="71">Sinaloa</collectingRegion>
, Mex /
<date id="5E1EDEEDFD19FFC7AD6E1982FD61BC8D" box="[607,735,212,236]" pageId="70" pageNumber="71" value="1955-07-22">
<collectingDate id="4E5A2705FD19FFC7AD6E1982FD61BC8D" box="[607,735,212,236]" pageId="70" pageNumber="71" value="1955-07-22">22-VII-1955</collectingDate>
</date>
// R&amp;J Selander //
<typeStatus id="F51B468FFD19FFC7AC921982FB9FBC8D" box="[931,1057,212,236]" pageId="70" pageNumber="71" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="EDA083AEFD19FFC7AB041982FB28BC8D" authorityName="Hope" authorityYear="1837" box="[1077,1174,212,236]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">Bothynus</taxonomicName>
/ selanderi / Cartwright //
<typeStatus id="F51B468FFD19FFC7AFF619A5FEFBBD6A" box="[199,325,243,267]" pageId="70" pageNumber="71" type="paratype">PARATYPE</typeStatus>
// MNHN /
<specimenCode id="7A065056FD19FFC7AEF019A4FDA7BD6B" box="[449,537,242,266]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="70" pageNumber="71">EC6733</specimenCode>
”. Type locality: Los Mochis,
<collectingRegion id="E86436CFFD19FFC7AC7319A5FC2FBD6A" box="[834,913,243,267]" country="Mexico" name="Sinaloa" pageId="70" pageNumber="71">Sinaloa</collectingRegion>
, Mexico.
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD19FFC7AFA6186EFD80BFF2" pageId="70" pageNumber="71" type="description">
<paragraph id="2A1FF82DFD19FFC7AFA6186EFD80BFF2" blockId="70.[151,1437,312,1419]" pageId="70" pageNumber="71">
<emphasis id="18D4243FFD19FFC7AFA6186EFE97BD33" bold="true" box="[151,297,312,338]" pageId="70" pageNumber="71">Description.</emphasis>
Habitus as in
<figureCitation id="B29BE4A8FD19FFC7AEE4186FFD81BD32" box="[469,575,313,339]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="70" pageNumber="71">Fig. 28H</figureCitation>
. Length 21.0–30.0 mm; humeral width
<quantity id="ED5855C8FD19FFC7AB3C186FFB08BD32" box="[1037,1206,313,340]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" metricValueMax="1.45" metricValueMin="1.01" pageId="70" pageNumber="71" unit="mm" value="12.3" valueMax="14.5" valueMin="10.1">10.1–14.5 mm</quantity>
. Color dark reddish brown to black.
<emphasis id="18D4243FFD19FFC7AE7E180BFE26BD16" bold="true" box="[335,408,349,375]" pageId="70" pageNumber="71">Head:</emphasis>
Frons and clypeus coarsely and densely rugose. Frontoclypeal region with 2 conical tubercles separated by about 4.5–5.0 tubercle diameters. Clypeus narrowed towards apex, base 2.5 times wider than apex. Clypeal teeth triangular, separated by a tooth diameter. Mandible with 2 apical teeth and a lateral, narrowly rounded tooth. Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed (
<figureCitation id="B29BE4A8FD19FFC7AA02189FFA2EBD82" box="[1331,1424,457,483]" captionStart="FIGURE 4" captionStartId="83.[151,250,1872,1896]" captionTargetBox="[151,1437,1118,1848]" captionTargetId="figure-81@83.[151,1437,1118,1848]" captionTargetPageId="83" captionText="FIGURE 4. Maxilla. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus gyas, G) Tomarus maternus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345023" httpUri="https://zenodo.org/record/7345023/files/figure.png" pageId="70" pageNumber="71">Fig. 4H</figureCitation>
). Interocular distance equal to 3.5 times an eye width.
<emphasis id="18D4243FFD19FFC7ADC618BBFCC4BE66" bold="true" box="[759,890,493,519]" pageId="70" pageNumber="71">Pronotum:</emphasis>
Surface with small punctures, punctures denser and deeper on anterior and posterior angles. Apical tubercle prominent, acute, slightly recurved. Subapical fovea deep, rounded, as wide as interocular distance or slightly narrower, equal in both sexes, surface transversely rugose.
<emphasis id="18D4243FFD19FFC7AFA61B0EFEA6BE13" bold="true" box="[151,280,600,626]" pageId="70" pageNumber="71">Scutellum:</emphasis>
Surface with minute, sparse punctures.
<emphasis id="18D4243FFD19FFC7ADD71B0FFC85BE12" bold="true" box="[742,827,601,627]" pageId="70" pageNumber="71">Elytra:</emphasis>
Punctures on first interval as large as those on other intervals. Inner surface of apex with rounded, large tubercles forming 17 parallel lines.
<emphasis id="18D4243FFD19FFC7AB5F1B2AFB52BEF7" bold="true" box="[1134,1260,636,662]" pageId="70" pageNumber="71">Abdomen:</emphasis>
Apex of tergite IV with a triangular wide area with large, rounded tubercles forming 11–12 diagonal, parallel lines. Pygidium with dense rugosity on basal 3rd. Apex rounded.
<emphasis id="18D4243FFD19FFC7ADB61B93FD5EBEBE" bold="true" box="[647,736,709,735]" pageId="70" pageNumber="71">Venter:</emphasis>
Apex of prosternal process flat, longitudinally oval to subrectangular. Metasternum with minute setae.
<emphasis id="18D4243FFD19FFC7AD5F1BBFFD11BF62" bold="true" box="[622,687,745,771]" pageId="70" pageNumber="71">Legs:</emphasis>
Protibia tridentate, sometimes with a basal small denticle. Apex of metatibia crenulate, with 13–15 spinules.
<emphasis id="18D4243FFD19FFC7AD5A1A5BFCA1BF47" bold="true" box="[619,799,780,807]" pageId="70" pageNumber="71">Male genitalia:</emphasis>
Spiculum gastrale without basal plates (
<figureCitation id="B29BE4A8FD19FFC7ABD41A5BFAF0BF46" box="[1253,1358,781,807]" captionStart="FIGURE 14" captionStartId="89.[151,250,1677,1701]" captionTargetBox="[152,1432,1161,1649]" captionTargetId="figure-90@89.[152,1436,1161,1653]" captionTargetPageId="89" captionText="FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345059" httpUri="https://zenodo.org/record/7345059/files/figure.png" pageId="70" pageNumber="71">Fig. 14G</figureCitation>
). Phallobase 1.1 times longer than parameres. Parameres without dorsal teeth, apex wide, triangularly dilated (
<figureCitation id="B29BE4A8FD19FFC7AA171A67FA29BF2A" box="[1318,1431,817,843]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="70" pageNumber="71">Figs. 16Q</figureCitation>
,
<figureCitation id="B29BE4A8FD19FFC7AFA61A03FF72BF0E" box="[151,204,853,879]" captionStart="FIGURE 20" captionStartId="95.[151,250,1225,1249]" captionTargetBox="[159,1435,191,1201]" captionTargetId="figure-17@95.[151,1436,181,1202]" captionTargetPageId="95" captionText="FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345083" httpUri="https://zenodo.org/record/7345083/files/figure.png" pageId="70" pageNumber="71">20H</figureCitation>
). Internal sac with copulatory lamella and short lamellar spiny belt; with a complex of 5 spine-like, accessory lamellae, with dense granules at base.
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD19FFC7AFF61ACAFB33B842" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="2A1FF82DFD19FFC7AFF61ACAFB33B842" blockId="70.[151,1437,312,1419]" pageId="70" pageNumber="71">
<emphasis id="18D4243FFD19FFC7AFF61ACAFEFCBFD7" bold="true" box="[199,322,924,950]" pageId="70" pageNumber="71">Diagnosis.</emphasis>
<taxonomicName id="EDA083AEFD19FFC7AE781AC8FDA4BFD6" baseAuthorityName="Cartwright" baseAuthorityYear="1959" box="[329,538,925,951]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="selanderi">
<emphasis id="18D4243FFD19FFC7AE781AC8FDA4BFD6" box="[329,538,925,951]" italics="true" pageId="70" pageNumber="71">Tomarus selanderi</emphasis>
</taxonomicName>
can be differentiated by the following character combination: frontoclypeal tubercles conical; pronotal tubercle acute and prominent (
<figureCitation id="B29BE4A8FD19FFC7AC3C1A97FCC8BFBA" box="[781,886,961,987]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="70" pageNumber="71">Fig. 28H</figureCitation>
); pronotal fovea shallow, as wide as or narrower than interocular distance; protibia tridentate, rarely with an additional basal denticle; apex of metatibia with 14–15 spinules; parameres without dorsal teeth; apex wide, triangularly dilated (
<figureCitation id="B29BE4A8FD19FFC7ACFF1D5FFBFEB842" box="[974,1088,1033,1059]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="70" pageNumber="71">Figs. 16Q</figureCitation>
,
<figureCitation id="B29BE4A8FD19FFC7AB7D1D5FFB3FB842" box="[1100,1153,1033,1059]" captionStart="FIGURE 20" captionStartId="95.[151,250,1225,1249]" captionTargetBox="[159,1435,191,1201]" captionTargetId="figure-17@95.[151,1436,181,1202]" captionTargetPageId="95" captionText="FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345083" httpUri="https://zenodo.org/record/7345083/files/figure.png" pageId="70" pageNumber="71">20H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD19FFC7AFF61D7AFE16B9EA" pageId="70" pageNumber="71" type="distribution">
<paragraph id="2A1FF82DFD19FFC7AFF61D7AFDC8B80A" blockId="70.[151,1437,312,1419]" pageId="70" pageNumber="71">
<emphasis id="18D4243FFD19FFC7AFF61D7AFEDFB827" bold="true" box="[199,353,1068,1094]" pageId="70" pageNumber="71">Distribution.</emphasis>
<taxonomicName id="EDA083AEFD19FFC7AE581D78FD85B826" baseAuthorityName="Cartwright" baseAuthorityYear="1959" box="[361,571,1069,1095]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="selanderi">
<emphasis id="18D4243FFD19FFC7AE581D78FD85B826" box="[361,571,1069,1095]" italics="true" pageId="70" pageNumber="71">Tomarus selanderi</emphasis>
</taxonomicName>
is an endemic species found along the Pacific coast of
<collectingCountry id="52B7B8BDFD19FFC7AB8A1D7BFAAFB826" box="[1211,1297,1069,1095]" name="Mexico" pageId="70" pageNumber="71">Mexico</collectingCountry>
(
<bibRefCitation id="4E3185DCFD19FFC7AA101D7BFF6CB80D" author="Cartwright, O. L." pageId="70" pageNumber="71" pagination="515 - 541" refId="ref88844" refString="Cartwright, O. L. (1959) Scarab beetles of the genus Bothynus in the United States (Coleoptera: Sacarabaeidae). Proceedings of the United States National Museum, 108, 515 - 541." type="journal article" year="1959">Cartwright 1959</bibRefCitation>
;
<bibRefCitation id="4E3185DCFD19FFC7AFEF1D07FEC4B80D" author="Endrodi, S." box="[222,378,1105,1132]" pageId="70" pageNumber="71" refId="ref89854" refString="Endrodi, S. (1985) The Dynastinae of the World. W. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrödi 1985</bibRefCitation>
;
<bibRefCitation id="4E3185DCFD19FFC7AEB71D07FDD4B80A" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Cano, E. B." box="[390,618,1105,1131]" pageId="70" pageNumber="71" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. &amp; Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
Ratcliffe
<emphasis id="18D4243FFD19FFC7AEDF1D04FD99B80A" box="[494,551,1105,1131]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
<paragraph id="2A1FF82DFD19FFC7AFF61D22FCF0B922" blockId="70.[151,1437,312,1419]" pageId="70" pageNumber="71">
<emphasis id="18D4243FFD19FFC7AFF61D22FDBFB8EF" bold="true" box="[199,513,1140,1167]" pageId="70" pageNumber="71">
Locality records (
<figureCitation id="B29BE4A8FD19FFC7AEAC1D22FE4AB8EF" box="[413,500,1140,1166]" captionStart="FIGURE 45" captionStartId="113.[151,250,1077,1101]" captionTargetBox="[207,1391,181,1048]" captionTargetId="figure-17@113.[197,1391,181,1053]" captionTargetPageId="113" captionText="FIGURE 45. Distributional records of Tomarus discrepans and T. selanderi." figureDoi="http://doi.org/10.5281/zenodo.7345168" httpUri="https://zenodo.org/record/7345168/files/figure.png" pageId="70" pageNumber="71">Fig. 45</figureCitation>
).
</emphasis>
<materialsCitation id="9AC8F270FD19FFC7AD3A1D23FCF4B922" ID-GBIF-Occurrence="3986696799" collectionCode="CAFESI, EMAC, FSCA, CNIN, IEXA, MNHN, UNSM, USNM" collectorName="El Venado &amp; El Marmol" country="Mexico" location="San Blas" pageId="70" pageNumber="71" specimenCount="1" stateProvince="Colima">
205 examined specimens from
<collectionCode id="4CB160E8FD19FFC7AC401D23FC6AB8EE" box="[881,980,1141,1167]" pageId="70" pageNumber="71">CAFESI</collectionCode>
,
<collectionCode id="4CB160E8FD19FFC7ACD01D23FB8BB8EE" box="[993,1077,1141,1167]" pageId="70" pageNumber="71">EMAC</collectionCode>
,
<collectionCode id="4CB160E8FD19FFC7AB731D23FB35B8EE" box="[1090,1163,1141,1167]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="70" pageNumber="71" type="Museum">FSCA</collectionCode>
,
<collectionCode id="4CB160E8FD19FFC7ABA91D23FB61B8EE" box="[1176,1247,1141,1167]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="70" pageNumber="71">CNIN</collectionCode>
,
<collectionCode id="4CB160E8FD19FFC7ABDC1D23FA8DB8EE" box="[1261,1331,1141,1167]" pageId="70" pageNumber="71">IEXA</collectionCode>
,
<collectionCode id="4CB160E8FD19FFC7AA711D23FA26B8EE" box="[1344,1432,1141,1167]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="70" pageNumber="71">MNHN</collectionCode>
,
<collectionCode id="4CB160E8FD19FFC7AFA61DCFFF55B8D2" box="[151,235,1177,1203]" country="USA" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="70" pageNumber="71">UNSM</collectionCode>
,
<collectionCode id="4CB160E8FD19FFC7AFC61DCFFEF2B8D2" box="[247,332,1177,1203]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="70" pageNumber="71" type="Museum">USNM</collectionCode>
. Some data from
<bibRefCitation id="4E3185DCFD19FFC7AD201DCFFCB6B8D2" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Cano, E. B." box="[529,776,1177,1203]" pageId="70" pageNumber="71" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. &amp; Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
Ratcliffe
<emphasis id="18D4243FFD19FFC7AD4B1DCCFD0DB8D2" box="[634,691,1177,1203]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
(2013)
</bibRefCitation>
.
<emphasis id="18D4243FFD19FFC7AC221DCEFBDBB8D3" bold="true" box="[787,1125,1176,1202]" pageId="70" pageNumber="71">
<collectingCountry id="52B7B8BDFD19FFC7AC221DCEFC37B8D3" box="[787,905,1176,1202]" name="Mexico" pageId="70" pageNumber="71">MEXICO</collectingCountry>
(214).
<collectingRegion id="E86436CFFD19FFC7ACED1DCEFB8DB8D3" box="[988,1075,1176,1202]" country="Mexico" name="Colima" pageId="70" pageNumber="71">Colima</collectingRegion>
(2):
</emphasis>
Manzanillo.
<emphasis id="18D4243FFD19FFC7ABCB1DCEFA22B8D3" bold="true" box="[1274,1436,1176,1202]" pageId="70" pageNumber="71">
<collectingRegion id="E86436CFFD19FFC7ABCB1DCEFAD4B8D3" box="[1274,1386,1176,1202]" country="Mexico" name="Guerrero" pageId="70" pageNumber="71">Guerrero</collectingRegion>
(2):
</emphasis>
Acapulco.
<emphasis id="18D4243FFD19FFC7AE211DEAFE1EB8B7" bold="true" box="[272,416,1212,1238]" pageId="70" pageNumber="71">
<collectingRegion id="E86436CFFD19FFC7AE211DEAFEDFB8B7" box="[272,353,1212,1238]" country="Mexico" name="Jalisco" pageId="70" pageNumber="71">Jalisco</collectingRegion>
(17):
</emphasis>
Careyes; Chamela; Guadalajara; Melaque; Puerto Vallarta.
<emphasis id="18D4243FFD19FFC7AB0C1DEAFB4CB8B7" bold="true" box="[1085,1266,1212,1239]" pageId="70" pageNumber="71">
<collectingRegion id="E86436CFFD19FFC7AB0C1DEAFB7FB8B6" box="[1085,1217,1212,1239]" country="Mexico" name="Michoacan de Ocampo" pageId="70" pageNumber="71">Michoacán</collectingRegion>
(1):
</emphasis>
Jungapeo.
<emphasis id="18D4243FFD19FFC7AA401DEBFEAFB89B" bold="true" pageId="70" pageNumber="71">
<collectingRegion id="E86436CFFD19FFC7AA401DEBFF6FB89B" country="Mexico" name="Nayarit" pageId="70" pageNumber="71">Nayarit</collectingRegion>
(85):
</emphasis>
Acaponeta;
<collectorName id="87559DFBFD19FFC7AEAE1DB7FDA9B89A" box="[415,535,1249,1275]" pageId="70" pageNumber="71">El Venado</collectorName>
; Ruiz;
<location id="2F7FAEF6FD19FFC7AD591DB7FD71B89A" LSID="urn:lsid:plazi:treatment:A209493BFD1AFFC7AFA61F8DFE16B9EA:2F7FAEF6FD19FFC7AD591DB7FD71B89A" box="[616,719,1249,1275]" country="Mexico" name="San Blas" pageId="70" pageNumber="71" stateProvince="Colima">San Blas</location>
;
<locationDeviation id="3BADE10FFD19FFC7ADED1DB7FC14B89A" box="[732,938,1249,1276]" location="San Blas" pageId="70" pageNumber="71">
<quantity id="ED5855C8FD19FFC7ADED1DB7FC9AB89A" box="[732,804,1249,1276]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="km" value="15.0">15 km</quantity>
S San Blas
</locationDeviation>
; Sayulita; Singayta; Tecuala; Yago.
<emphasis id="18D4243FFD19FFC7AA731DB6FF7FB97F" bold="true" pageId="70" pageNumber="71">
<collectingRegion id="E86436CFFD19FFC7AA731DB6FA22B89B" box="[1346,1436,1248,1274]" country="Mexico" name="Oaxaca" pageId="70" pageNumber="71">Oaxaca</collectingRegion>
(2):
</emphasis>
Pinotepa Nacional; Puerto Escondido.
<emphasis id="18D4243FFD19FFC7AD481C52FCAFB97F" bold="true" box="[633,785,1284,1310]" pageId="70" pageNumber="71">
<collectingRegion id="E86436CFFD19FFC7AD481C52FD6CB97F" box="[633,722,1284,1310]" country="Mexico" name="Sinaloa" pageId="70" pageNumber="71">Sinaloa</collectingRegion>
(98):
</emphasis>
<collectorName id="87559DFBFD19FFC7AC291C53FC28B97E" box="[792,918,1285,1311]" pageId="70" pageNumber="71">El Mármol</collectorName>
; Escuinapa; Los Mochis; Mazatlan; Novalato; Piedra Blanca.
<emphasis id="18D4243FFD19FFC7AE701C7EFE76B923" bold="true" box="[321,456,1320,1346]" pageId="70" pageNumber="71">
<collectingRegion id="E86436CFFD19FFC7AE701C7EFE29B923" box="[321,407,1320,1346]" country="Mexico" name="Sonora" pageId="70" pageNumber="71">Sonora</collectingRegion>
(6):
</emphasis>
Esperanza, Guaymas, Hermosillo
</materialsCitation>
.
</paragraph>
<paragraph id="2A1FF82DFD19FFC7AFF61C1BFE16B9EA" blockId="70.[151,1437,312,1419]" pageId="70" pageNumber="71">
<emphasis id="18D4243FFD19FFC7AFF61C1BFE39B907" bold="true" box="[199,391,1356,1383]" pageId="70" pageNumber="71">Natural history.</emphasis>
Specimens have been collected between
<quantity id="ED5855C8FD19FFC7AC6F1C1BFC61B906" box="[862,991,1357,1384]" metricMagnitude="2" metricUnit="m" metricValue="7.65" metricValueMax="15.0" metricValueMin="0.3" pageId="70" pageNumber="71" unit="m" value="765.0" valueMax="1500.0" valueMin="30.0">30–1500 m</quantity>
(
<bibRefCitation id="4E3185DCFD19FFC7ACDF1C1BFB68B906" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Cano, E. B." box="[1006,1238,1357,1383]" pageId="70" pageNumber="71" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. &amp; Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
Ratcliffe
<emphasis id="18D4243FFD19FFC7AB6B1C18FB2AB906" box="[1114,1172,1357,1383]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
2013
</bibRefCitation>
) in dry forests of Pacific coast of
<collectingCountry id="52B7B8BDFD19FFC7AE7A1C27FE1AB9EA" box="[331,420,1393,1419]" name="Mexico" pageId="70" pageNumber="71">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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