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<document id="C401DF8CB5294AD23A0896922ACE054E" ID-DOI="10.5281/zenodo.180453" ID-GBIF-Dataset="62cb965f-2144-4449-8276-e61141e7a041" ID-ISSN="1175-5326" ID-Zenodo-Dep="180453" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459807613697" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Ivanov, Boris G." docDate="2008" docId="03859630FFA11C0CCDF0566728F2FD5A" docLanguage="en" docName="zt01684p034.pdf" docOrigin="Zootaxa 1684" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Heptacarpus acuticarinatus Komai &amp; Ivanov, 2008, n. sp." docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FFBCEE48FFA91C02CD67505D2B7EFFB8" masterDocTitle="Identities of three taxa of the hippolytid shrimp genus Heptacarpus (Crustacea: Decapoda: Caridea), with description of a new species from East Asian waters" masterLastPageNumber="34" masterPageNumber="1" pageNumber="9" updateTime="1698220711174" updateUser="plazi">
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<mods:title id="A76D66FE8CD0A4C6340669C8518162B4">Identities of three taxa of the hippolytid shrimp genus Heptacarpus (Crustacea: Decapoda: Caridea), with description of a new species from East Asian waters</mods:title>
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<taxonomicName id="4C2C5CA5FFA11C0ACDF056672A98F9EC" box="[151,486,1594,1620]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="acuticarinatus" status="sp. nov.">
<emphasis id="B958FB34FFA11C0ACDF056672A98F9EC" bold="true" box="[151,486,1594,1620]" italics="true" pageId="8" pageNumber="9">Heptacarpus acuticarinatus</emphasis>
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(
<figureCitation id="13173BA3FFA11C0ACDF856002A77F9CF" box="[159,265,1629,1655]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStartId-0="9.[151,255,1717,1741]" captionStartId-1="10.[151,255,1890,1914]" captionStartId-2="11.[151,255,1824,1848]" captionStartId-3="12.[151,255,1874,1898]" captionStartId-4="13.[151,255,1826,1850]" captionTargetBox-0="[237,1348,1009,1692]" captionTargetBox-1="[186,1360,697,1889]" captionTargetBox-2="[186,1397,192,1823]" captionTargetBox-3="[179,1393,912,1873]" captionTargetBox-4="[151,1436,978,1825]" captionTargetId-0="figure@9.[151,1436,976,1710]" captionTargetId-1="figure@10.[151,1436,628,1924]" captionTargetId-2="figure@11.[168,1422,158,1881]" captionTargetId-3="figure@12.[144,1429,851,1951]" captionTargetId-4="figure@13.[151,1436,950,1852]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 5. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm." captionText-1="FIGURE 6. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for B D." captionText-2="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." captionText-3="FIGURE 8. Heptacarpus acuticarinatus n. sp. Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for A C; 0.5 mm for D, E." captionText-4="FIGURE 9. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view, showing coloration in life." httpUri-0="https://zenodo.org/record/180458/files/figure.png" httpUri-1="https://zenodo.org/record/180459/files/figure.png" httpUri-2="https://zenodo.org/record/180460/files/figure.png" httpUri-3="https://zenodo.org/record/180461/files/figure.png" httpUri-4="https://zenodo.org/record/180462/files/figure.png" pageId="8" pageNumber="9">Figs 59</figureCitation>
,
<figureCitation id="13173BA3FFA11C0ACC7256002A4BF9CF" box="[277,309,1629,1655]" captionStart="FIGURE 18" captionStartId="25.[151,255,1068,1092]" captionTargetBox="[170,1419,308,1053]" captionTargetId="figure@25.[162,1427,300,1059]" captionTargetPageId="25" captionText="FIGURE 18. Variation in the number of rostral ventral teeth of four Heptacarpus species." httpUri="https://zenodo.org/record/180471/files/figure.png" pageId="8" pageNumber="9">18</figureCitation>
,
<figureCitation id="13173BA3FFA11C0ACC2656002A21F9CF" box="[321,351,1629,1655]" captionStart="FIGURE 19" captionStartId="26.[151,255,1443,1467]" captionTargetBox="[157,1428,463,1421]" captionTargetId="figure@26.[151,1436,456,1426]" captionTargetPageId="26" captionText="FIGURE 19. Variation in the number of meral spines on the third pereopod of four Heptacarpus species." httpUri="https://zenodo.org/record/180472/files/figure.png" pageId="8" pageNumber="9">19</figureCitation>
)
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<taxonomicName id="4C2C5CA5FFA11C0ACDF056F02ACAF97C" box="[151,436,1709,1732]" class="Malacostraca" family="Hippolytidae" genus="Spirontocaris" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="camtchatica">
<emphasis id="B958FB34FFA11C0ACDF056F02ACAF97C" box="[151,436,1709,1732]" italics="true" pageId="8" pageNumber="9">Spirontocaris camtchatica</emphasis>
</taxonomicName>
.
<treatmentCitation id="0A8D0137FFA11C0ACC8756F029ECF97C" author="Balss" box="[480,658,1709,1732]" page="44" pageId="8" pageNumber="9" year="1914">
<bibRefCitation id="EFBD5AD7FFA11C0ACC8756F0291BF97C" author="Balss" box="[480,613,1709,1732]" pageId="8" pageNumber="31" refString="Balss, H. (1914) Ostasiatische Decapoden II: Die Natantia und Reptantia. In: Doflein, F. (Ed.), Beitrage zur Naturgeschichte Ostasiens, Dekapoden, part 7. Abhandlungen der Bayerischen Akademie der Wissenschaften, Munchen, 2 (Supplement), 10, 1 - 101, pl. 1." type="journal article" year="1914">Balss, 1914</bibRefCitation>
: 44
</treatmentCitation>
;
<treatmentCitation id="0A8D0137FFA11C0ACFC556F02829F97C" author="Parisi" box="[674,855,1709,1732]" page="47" pageId="8" pageNumber="9" year="1919">
<bibRefCitation id="EFBD5AD7FFA11C0ACFC556F02854F97C" author="Parisi" box="[674,810,1709,1732]" pageId="8" pageNumber="34" refString="Parisi, B. (1919) I Decapodi giapponesi del Museo di Milano. VII. Natantia. Atii della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 58, 59 - 99, pls 2 - 6." type="journal article" year="1919">Parisi, 1919</bibRefCitation>
: 47
</treatmentCitation>
;
<treatmentCitation id="0A8D0137FFA11C0ACE0056F02F4FF97C" author="Yokoya" box="[871,1073,1709,1732]" page="26" pageId="8" pageNumber="9" year="1933">
<bibRefCitation id="EFBD5AD7FFA11C0ACE0056F02F7AF97C" author="Yokoya" box="[871,1028,1709,1732]" pageId="8" pageNumber="34" refString="Yokoya, Y. (1933) On the distribution of decapod Crustacea inhabiting the continental shelf around Japan, chiefly based upon the materials collected by S. S. &quot; Soyo Maru &quot; during the years 1923 - 1930. Journal of the College of Agriculture, Tokyo Imperial University, 12: 1 - 236." type="journal article" year="1933">Yokoya, 1933</bibRefCitation>
: 26
</treatmentCitation>
. Not
<taxonomicName id="4C2C5CA5FFA11C0AC91356F02AFDF95C" authority="Stimpson, 1860" authorityName="Stimpson" authorityYear="1860" class="Malacostraca" family="Hippolytidae" genus="Spirontocaris" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="camtschatica">
<emphasis id="B958FB34FFA11C0AC91356F02EE5F97C" box="[1140,1435,1709,1732]" italics="true" pageId="8" pageNumber="9">Spirontocaris camtschatica</emphasis>
(
<bibRefCitation id="EFBD5AD7FFA11C0ACDA956902A05F95C" author="Stimpson" box="[206,379,1741,1764]" pageId="8" pageNumber="34" refString="Stimpson, W. (1860) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missam Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. Proceedings of the Academy of Natural Sciences, Philadelphia, 1860, 22 - 48 (91 - 117)." type="proceedings paper" year="1860">Stimpson, 1860</bibRefCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="AB3C0008FFA11C0ACDF056B029CBF8FC" pageId="8" pageNumber="9">
<taxonomicName id="4C2C5CA5FFA11C0ACDF056B02ABCF8BC" box="[151,450,1773,1796]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA11C0ACDF056B02ABCF8BC" box="[151,450,1773,1796]" italics="true" pageId="8" pageNumber="9">Heptacarpus camtschaticus</emphasis>
</taxonomicName>
.
<treatmentCitation id="0A8D0137FFA11C0ACC8D56B029ECF8BC" author="Liu" box="[490,658,1773,1796]" page="237" pageId="8" pageNumber="9" year="1963">
<bibRefCitation id="EFBD5AD7FFA11C0ACC8D56B02927F8BC" author="Liu" box="[490,601,1773,1796]" pageId="8" pageNumber="33" refString="Liu, J. Y. (1963) Zoogeographical Studies on the macrurous crustacean fauna of the Yellow Sea and the East China Sea. Oceanologia et Limnologia Sinica, 5 (3), 230 - 244. [in Chinese]" type="journal article" year="1963">Liu, 1963</bibRefCitation>
: 237
</treatmentCitation>
;
<treatmentCitation id="0A8D0137FFA11C0ACFC656B0288BF8BC" author="Hayashi" box="[673,1013,1773,1796]" page="134" pageId="8" pageNumber="9" year="1968">
<bibRefCitation id="EFBD5AD7FFA11C0ACFC656B028C3F8BC" author="Hayashi" box="[673,957,1773,1796]" pageId="8" pageNumber="32" refString="Hayashi, K. &amp; Miyake, S. (1968) Studies on the hippolytid shrimps of Japan, V. Hippolytid fauna of the sea around the Amakusa Marine Biological Laboratory. OHMU, 1, 121 - 163." type="journal article" year="1968">Hayashi &amp; Miyake, 1968</bibRefCitation>
: 134
</treatmentCitation>
, fig. 6;
<treatmentCitation id="0A8D0137FFA11C0AC93656B02EE6F8BC" author="Kikuchi" box="[1105,1432,1773,1796]" page="24" pageId="8" pageNumber="9" year="1978">
<bibRefCitation id="EFBD5AD7FFA11C0AC93656B02E12F8BC" author="Kikuchi" box="[1105,1388,1773,1796]" pageId="8" pageNumber="32" refString="Kikuchi, T. &amp; Miyake, S. (1978) Fauna and Flora of the Sea around the Amakusa Marine Biological Laboratory. Part II. Decapoda Crustacea (Revised Edition). Amakusa Marine Biological Laboratory, Tomioka, 52 pp." type="book" year="1978">Kikuchi &amp; Miyake, 1978</bibRefCitation>
: 24
</treatmentCitation>
;
<treatmentCitation id="0A8D0137FFA11C0ACDA057502AF0F89C" author="Hayashi" box="[199,398,1805,1828]" page="14" pageId="8" pageNumber="9" year="1979">
<bibRefCitation id="EFBD5AD7FFA11C0ACDA057502A18F89C" author="Hayashi" box="[199,358,1805,1828]" pageId="8" pageNumber="32" refString="Hayashi, K. (1979) Studies on hippolytid shrimps from Japan VII: The genus Heptacarpus Holmes. Journal of Shimonoseki University of Fisheries, 28, 11 - 32." type="journal article" year="1979">Hayashi, 1979</bibRefCitation>
: 14
</treatmentCitation>
; 1992: 180, figs 223a, 224a, 225a;
<treatmentCitation id="0A8D0137FFA11C0ACE6C57502F58F89C" author="Liu" box="[779,1062,1805,1828]" page="559" pageId="8" pageNumber="9" year="1994">
<bibRefCitation id="EFBD5AD7FFA11C0ACE6C5750288DF89C" author="Liu" box="[779,1011,1805,1828]" pageId="8" pageNumber="33" refString="Liu, R. (Liu, J. Y.) &amp; Zhong, Z. (1994) Decapoda. In: Huang, Z. (Ed.), Marine Species and Their Distributions in China's Seas. China Ocean Press, Beijing, pp. 545 - 568. [in Chinese]" type="book chapter" year="1994">Liu and Zhong, 1994</bibRefCitation>
: 559
</treatmentCitation>
(list);
<treatmentCitation id="0A8D0137FFA11C0AC90857502E3EF89C" author="Cha" box="[1135,1344,1805,1828]" page="90" pageId="8" pageNumber="9" year="2001">
<bibRefCitation id="EFBD5AD7FFA11C0AC90857502E69F89C" author="Cha" box="[1135,1303,1805,1828]" pageId="8" pageNumber="32" refString="Cha, H. K., Lee, J. U., Park, C. S., Baik, C. I., Hong, S. Y., Park, J. H., Lee, D. W., Choi, Y. M., Hwang, K., Kim, Z. G., Choi, K. H., Sohn, H., Sohn, M. H., Kim, D. H. &amp; Choi, J. H. (2001) Shrimps of the Korean Waters. National Fisheries Research and Development Institute, Pusan, 188 pp." type="book" year="2001">
Cha
<emphasis id="B958FB34FFA11C0AC9C657502FA6F89C" box="[1185,1240,1805,1828]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2001
</bibRefCitation>
: 90
</treatmentCitation>
91. Not
<taxonomicName id="4C2C5CA5FFA11C0ACDA0577029CFF8FC" authority="Stimpson, 1860" authorityName="Stimpson" authorityYear="1860" box="[199,689,1837,1860]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA11C0ACDA057702A90F8FC" box="[199,494,1837,1860]" italics="true" pageId="8" pageNumber="9">Heptacarpus camtschaticus</emphasis>
(
<bibRefCitation id="EFBD5AD7FFA11C0ACC9B577029D6F8FC" author="Stimpson" box="[508,680,1837,1860]" pageId="8" pageNumber="34" refString="Stimpson, W. (1860) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missam Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. Proceedings of the Academy of Natural Sciences, Philadelphia, 1860, 22 - 48 (91 - 117)." type="proceedings paper" year="1860">Stimpson, 1860</bibRefCitation>
)
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.
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<typeStatus id="54979984FFA11C0ACDF057282BACF837" box="[151,210,1909,1935]" pageId="8" pageNumber="9">Type</typeStatus>
material
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<typeStatus id="54979984FFA11C0ACC0157282AA9F837" box="[358,471,1909,1935]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
. CBM-ZC 8980, ovigerous female (cl
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), Sagami Bay,
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<geoCoordinate id="EE1841E1FFA11C0ACDF057C02A47F80F" box="[151,313,1949,1975]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="139.56163">139°33.698'E</geoCoordinate>
,
<quantity id="4CD48AC3FFA11C0ACC2257C02AB3F80F" box="[325,461,1949,1975]" metricMagnitude="2" metricUnit="m" metricValue="1.005" metricValueMax="1.01" metricValueMin="1.0" pageId="8" pageNumber="9" unit="m" value="100.5" valueMax="101.0" valueMin="100.0">100101 m</quantity>
, sand bottom,
<date id="FF9201E6FFA11C0ACF1057C02845F80F" box="[631,827,1949,1975]" pageId="8" pageNumber="9" value="2003-01-22">22 January 2003</date>
, RV
<emphasis id="B958FB34FFA11C0ACE1257C02F77F80E" box="[885,1033,1949,1974]" italics="true" pageId="8" pageNumber="9">Rinkai-maru</emphasis>
, dredge, coll. T. Komai.
</paragraph>
<paragraph id="8B932726FFA11C0BCDA057982F40FE72" blockId="8.[151,1436,1909,2015]" lastBlockId="9.[151,1437,152,938]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<typeStatus id="54979984FFA11C0ACDA057982A43F867" box="[199,317,1989,2015]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B958FB34FFA11C0ACC2F57982AEAF867" bold="true" box="[328,404,1989,2015]" pageId="8" pageNumber="9">
<collectingCountry id="F33B67B6FFA11C0ACC2F57982AEAF867" box="[328,404,1989,2015]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
</emphasis>
. CBM-ZC 3531,
<specimenCount id="9D2AECAFFFA11C0ACF0F579829B1F867" box="[616,719,1989,2015]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(cl 4.0 mm), off Kominato, Boso Peninsula,
<quantity id="4CD48AC3FFA11C0AC98E57982E0CF867" box="[1257,1394,1989,2015]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="8" pageNumber="9" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100150 m</quantity>
,
<date id="FF9201E6FFA11C0BC81857982A3BFF0A" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" value="1997-02-27">27 February 1997</date>
, gill net, coll. T. Komai; CBM-ZC 2695,
<specimenCount id="9D2AECAFFFA01C0BCE4E50C528E7FF0A" box="[809,921,152,178]" pageId="9" pageNumber="10" type="female">5 females</specimenCount>
(cl
<quantity id="4CD48AC3FFA01C0BCEA150C52F29FF0A" box="[966,1111,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.9" metricValueMin="2.6" pageId="9" pageNumber="10" unit="mm" value="3.25" valueMax="3.9" valueMin="2.6">2.63.9 mm</quantity>
), 1 ovigerous female (cl
<quantity id="4CD48AC3FFA01C0BC81F50C52BB7FF62" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="10" unit="mm" value="5.4">5.4 mm</quantity>
), off Shionomisaki, Kii Peninsula,
<quantity id="4CD48AC3FFA01C0BCF03509D29DCFF62" box="[612,674,192,218]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="10" unit="m" value="80.0">80 m</quantity>
,
<date id="FF9201E6FFA01C0BCFC9509D2836FF62" box="[686,840,192,218]" pageId="9" pageNumber="10" value="1991-07-25">25 July 1991</date>
, dredge, coll. S. Nagai; HUMZ-C 1174,
<specimenCount id="9D2AECAFFFA01C0BC841509D2E0BFF62" box="[1318,1397,192,218]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
(cl
<quantity id="4CD48AC3FFA01C0BCDF050B52B89FEBA" box="[151,247,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="10" unit="mm" value="3.7">3.7 mm</quantity>
), Kashima-nada Sea off Ibaraki Prefecture, depth unknown,
<date id="FF9201E6FFA01C0BCEA950B52FC7FEBA" box="[974,1209,232,258]" pageId="9" pageNumber="10" value="1989-09-25">25 September 1989</date>
, larva net accidentally on bottom, coll. D. Kitagawa; NSMT-Cr S 9, 1 ovigerous female (cl
<quantity id="4CD48AC3FFA01C0BCE9B514D2F24FE92" box="[1020,1114,272,298]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="10" unit="mm" value="4.9">4.9 mm</quantity>
), Sagami Bay,
<geoCoordinate id="EE1841E1FFA01C0BC86D514D2EE9FE92" box="[1290,1431,272,298]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="9" value="35.131668">35°07.90N</geoCoordinate>
,
<geoCoordinate id="EE1841E1FFA01C0BCDF051652A51FEEA" box="[151,303,312,338]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="9" value="139.57466">139°34.48E</geoCoordinate>
,
<quantity id="4CD48AC3FFA01C0BCC5951652AD2FEEA" box="[318,428,312,338]" metricMagnitude="1" metricUnit="m" metricValue="9.45" metricValueMax="9.5" metricValueMin="9.4" pageId="9" pageNumber="10" unit="m" value="94.5" valueMax="95.0" valueMin="94.0">9495 m</quantity>
, sand bottom,
<date id="FF9201E6FFA01C0BCF0651652842FEEA" box="[609,828,312,338]" pageId="9" pageNumber="10" value="2002-02-27">27 February 2002</date>
, RV
<emphasis id="B958FB34FFA01C0BCE1B51652F6EFEE9" box="[892,1040,312,337]" italics="true" pageId="9" pageNumber="10">Rinkai-maru</emphasis>
, dredge, coll. T. Komai.
<collectingCountry id="F33B67B6FFA01C0BC820516A2EEAFEE9" box="[1351,1428,311,337]" name="South Korea" pageId="9" pageNumber="10">
<emphasis id="B958FB34FFA01C0BC820516A2EEAFEE9" bold="true" box="[1351,1428,311,337]" pageId="9" pageNumber="10">Korea</emphasis>
</collectingCountry>
. NFRDI-Cr 20070417-1,
<specimenCount id="9D2AECAFFFA01C0BCCDD513D2952FEC2" box="[442,556,352,378]" pageId="9" pageNumber="10" type="female">2 females</specimenCount>
(cl 5.5,
<quantity id="4CD48AC3FFA01C0BCFE9513D2992FEC2" box="[654,748,352,378]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="9" pageNumber="10" unit="mm" value="5.6">5.6 mm</quantity>
),
<specimenCount id="9D2AECAFFFA01C0BCE66513D282EFEC2" box="[769,848,352,378]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
(cl
<quantity id="4CD48AC3FFA01C0BCE19513D28A3FEC2" box="[894,989,352,378]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="10" unit="mm" value="5.2">5.2 mm</quantity>
), Hansan Island,
<quantity id="4CD48AC3FFA01C0BC9CF513D2E6DFEC2" box="[1192,1299,352,378]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="9" pageNumber="10" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">2030 m</quantity>
,
<date id="FF9201E6FFA01C0BC878513D2BABFE1A" pageId="9" pageNumber="10" value="1998-09">September 1998</date>
; NFRDI-Cr 20070417-2, 5 ovigerous females (cl
<quantity id="4CD48AC3FFA01C0BCE7B51D528D3FE1A" box="[796,941,392,418]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.4" metricValueMin="5.9" pageId="9" pageNumber="10" unit="mm" value="6.15" valueMax="6.4" valueMin="5.9">5.96.4 mm</quantity>
), same locality,
<date id="FF9201E6FFA01C0BC90E51D52E6FFE1A" box="[1129,1297,392,418]" pageId="9" pageNumber="10" value="1999-04-20">20 April 1999</date>
; NFRDI-Cr 20070417-3, 5 ovigerous females (cl
<quantity id="4CD48AC3FFA01C0BCF2B51ED29A0FE72" box="[588,734,432,458]" metricMagnitude="-3" metricUnit="m" metricValue="6.449999999999999" metricValueMax="6.8" metricValueMin="6.1" pageId="9" pageNumber="10" unit="mm" value="6.449999999999999" valueMax="6.8" valueMin="6.1">6.16.8 mm</quantity>
), same locality,
<date id="FF9201E6FFA01C0BCEFD51ED2F44FE72" box="[922,1082,432,458]" pageId="9" pageNumber="10" value="1999-05-20">20 May 1999</date>
.
</paragraph>
<paragraph id="8B932726FFA01C0BCDA0518A2FA6FDFA" blockId="9.[151,1437,152,938]" pageId="9" pageNumber="10">
<emphasis id="B958FB34FFA01C0BCDA0518A2AFAFE49" bold="true" box="[199,388,471,497]" pageId="9" pageNumber="10">Other material</emphasis>
.
<emphasis id="B958FB34FFA01C0BCCF3518A2A9EFE49" bold="true" box="[404,480,471,497]" pageId="9" pageNumber="10">
<collectingCountry id="F33B67B6FFA01C0BCCF3518A2A9EFE49" box="[404,480,471,497]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
</emphasis>
. HUMZ-C 29,
<specimenCount id="9D2AECAFFFA01C0BCFFD51852872FE4A" box="[666,780,472,498]" pageId="9" pageNumber="10" type="female">2 females</specimenCount>
(cl 5.2,
<quantity id="4CD48AC3FFA01C0BCE17518528AEFE4A" box="[880,976,472,498]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="9" pageNumber="10" unit="mm" value="5.7">5.7 mm</quantity>
), off Irifune, Hakodate, Hakodate Bay, southern Hokkaido, ca.
<quantity id="4CD48AC3FFA01C0BCCD2525D2A89FDA2" box="[437,503,512,538]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="m" value="30.0">30 m</quantity>
,
<date id="FF9201E6FFA01C0BCF62525D299EFDA2" box="[517,736,512,538]" pageId="9" pageNumber="10" value="1986-11-09">9 November 1986</date>
, gill net, coll. T. Komai; HUMZ-C 1179,
<specimenCount id="9D2AECAFFFA01C0BC9BA525D2E3BFDA2" box="[1245,1349,512,538]" pageId="9" pageNumber="10" type="female">1 female</specimenCount>
(cl 5.0 mm), similar locality, depth unknown,
<date id="FF9201E6FFA01C0BCF395275283DFDFA" box="[606,835,552,578]" pageId="9" pageNumber="10" value="1990-11-19">19 November 1990</date>
, small beam trawl, coll. T. Komai.
</paragraph>
</subSubSection>
<subSubSection id="C33674ADFFA01C0FCDA052122FECF812" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10" type="description">
<paragraph id="8B932726FFA01C0BCDA052122F29FC12" blockId="9.[151,1437,152,938]" pageId="9" pageNumber="10">
<emphasis id="B958FB34FFA01C0BCDA052122AA4FDD1" bold="true" box="[199,474,591,617]" pageId="9" pageNumber="10">Description of female</emphasis>
. Body (
<figureCitation id="13173BA3FFA01C0BCF27520D29F4FDD2" box="[576,650,592,618]" captionStart="FIGURE 5" captionStartId="9.[151,255,1717,1741]" captionTargetBox="[237,1348,1009,1692]" captionTargetId="figure@9.[151,1436,976,1710]" captionTargetPageId="9" captionText="FIGURE 5. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/180458/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
) moderately robust for genus. Rostrum (
<figureCitation id="13173BA3FFA01C0BC9E6520D2FB2FDD2" box="[1153,1228,592,618]" captionStart="FIGURE 5" captionStartId="9.[151,255,1717,1741]" captionTargetBox="[237,1348,1009,1692]" captionTargetId="figure@9.[151,1436,976,1710]" captionTargetPageId="9" captionText="FIGURE 5. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/180458/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
,
<figureCitation id="13173BA3FFA01C0BC9BA520D2F92FDD2" box="[1245,1260,592,618]" captionStart="FIGURE 6" captionStartId="10.[151,255,1890,1914]" captionTargetBox="[186,1360,697,1889]" captionTargetId="figure@10.[151,1436,628,1924]" captionTargetPageId="10" captionText="FIGURE 6. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for B D." httpUri="https://zenodo.org/record/180459/files/figure.png" pageId="9" pageNumber="10">6</figureCitation>
A, B) straight, directed forward, slightly falling short of or reaching distal margin of antennal scale, 1.221.53 length of carapace; dorsal margin armed with 57 teeth including 46 on rostrum proper and 1 or 2 on carapace, posteriormost tooth arising from 0.130.17 length of carapace, distal 0.160.30 of dorsal margin unarmed; ventral blade relatively deep, deepest at slightly proximal to midlength of rostrum; ventral margin with 58 teeth (teeth slightly unequal or subequal in size except for minute distalmost tooth); lateral carina sharply defined. Carapace (
<figureCitation id="13173BA3FFA01C0BCC72531D2A22FCE2" box="[277,348,832,858]" captionStart="FIGURE 5" captionStartId="9.[151,255,1717,1741]" captionTargetBox="[237,1348,1009,1692]" captionTargetId="figure@9.[151,1436,976,1710]" captionTargetPageId="9" captionText="FIGURE 5. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/180458/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
,
<figureCitation id="13173BA3FFA01C0BCC0D531D2A07FCE2" box="[362,377,832,858]" captionStart="FIGURE 6" captionStartId="10.[151,255,1890,1914]" captionTargetBox="[186,1360,697,1889]" captionTargetId="figure@10.[151,1436,628,1924]" captionTargetPageId="10" captionText="FIGURE 6. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for B D." httpUri="https://zenodo.org/record/180459/files/figure.png" pageId="9" pageNumber="10">6</figureCitation>
A, B) with postorbital rostral ridge low, not extending to anterior 0.25 of carapace length; dorsal margin in lateral view nearly straight; suborbital lobe (
<figureCitation id="13173BA3FFA01C0BCEE6533528BDFC3A" box="[897,963,872,898]" captionStart="FIGURE 6" captionStartId="10.[151,255,1890,1914]" captionTargetBox="[186,1360,697,1889]" captionTargetId="figure@10.[151,1436,628,1924]" captionTargetPageId="10" captionText="FIGURE 6. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for B D." httpUri="https://zenodo.org/record/180459/files/figure.png" pageId="9" pageNumber="10">Fig 6</figureCitation>
A) rounded, constricted at base, falling short of or reaching antennal tooth; pterygostomial angle always with small tooth.
</paragraph>
<caption id="DF5377AEFFA01C0BCDF056E8293AF957" httpUri="https://zenodo.org/record/180458/files/figure.png" pageId="9" pageNumber="10" targetBox="[237,1348,1009,1692]" targetPageId="9">
<paragraph id="8B932726FFA01C0BCDF056E8293AF957" blockId="9.[151,1436,1717,1775]" pageId="9" pageNumber="10">
<emphasis id="B958FB34FFA01C0BCDF056E82A65F975" bold="true" box="[151,283,1717,1741]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
<taxonomicName id="4C2C5CA5FFA01C0BCC4456EB2931F975" box="[291,591,1718,1741]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="acuticarinatus" status="sp. nov.">
<emphasis id="B958FB34FFA01C0BCC4456EB2931F975" box="[291,591,1718,1741]" italics="true" pageId="9" pageNumber="10">Heptacarpus acuticarinatus</emphasis>
</taxonomicName>
<emphasis id="B958FB34FFA01C0BCF3056E829EBF975" bold="true" box="[599,661,1717,1741]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A26B464FFFA01C0BCF3056E829EBF975" box="[599,661,1717,1741]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="8B932726FFA01C08CDA0577E2FF2FF0A" blockId="9.[151,1437,1827,2013]" lastBlockId="10.[151,1437,152,618]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Pleon (
<figureCitation id="13173BA3FFA01C0BCC7D577E2A1EF885" box="[282,352,1827,1853]" captionStart="FIGURE 5" captionStartId="9.[151,255,1717,1741]" captionTargetBox="[237,1348,1009,1692]" captionTargetId="figure@9.[151,1436,976,1710]" captionTargetPageId="9" captionText="FIGURE 5. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/180458/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
) dorsally rounded, not gibbous. Second somite with faint transverse groove on tergite. Dorsal surface of third tergite evenly convex, posterodorsal margin somewhat produced. Pleura of anterior four somites broadly rounded; fifth pleuron with moderately large posteroventral tooth, posterolateral margin sinuous. Sixth somite 1.501.60 times longer than fifth and 1.902.00 times longer than high. Telson (
<figureCitation id="13173BA3FFA01C0BC87057C62E1CF80D" box="[1303,1378,1947,1973]" captionStart="FIGURE 5" captionStartId="9.[151,255,1717,1741]" captionTargetBox="[237,1348,1009,1692]" captionTargetId="figure@9.[151,1436,976,1710]" captionTargetPageId="9" captionText="FIGURE 5. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/180458/files/figure.png" pageId="9" pageNumber="10">Figs 5</figureCitation>
,
<figureCitation id="13173BA3FFA01C0BC81757C62E01F80D" box="[1392,1407,1947,1973]" captionStart="FIGURE 6" captionStartId="10.[151,255,1890,1914]" captionTargetBox="[186,1360,697,1889]" captionTargetId="figure@10.[151,1436,628,1924]" captionTargetPageId="10" captionText="FIGURE 6. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for B D." httpUri="https://zenodo.org/record/180459/files/figure.png" pageId="9" pageNumber="10">6</figureCitation>
C) 1.201.30 length of sixth somite, about 3.60 times longer than wide, armed with 5 or 6 dorsolateral spines on either side; posterior margin terminating in acute tooth, with 3 pairs of unequal spines.
</paragraph>
<paragraph id="8B932726FFA31C08CDA0509D2E24FEBA" blockId="10.[151,1437,152,618]" pageId="10" pageNumber="11">
Eye-stalk (
<figureCitation id="13173BA3FFA31C08CC20509D2AF0FF62" box="[327,398,192,218]" captionStart="FIGURE 6" captionStartId="10.[151,255,1890,1914]" captionTargetBox="[186,1360,697,1889]" captionTargetId="figure@10.[151,1436,628,1924]" captionTargetPageId="10" captionText="FIGURE 6. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for B D." httpUri="https://zenodo.org/record/180459/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
B) generally subpyriform; cornea slightly wider and longer than remaining part of eyestalk; ocellus distinct, showing as black spot; maximal diameter of cornea 0.200.22 of carapace length.
</paragraph>
<paragraph id="8B932726FFA31C08CDA0514D28D9FE72" blockId="10.[151,1437,152,618]" pageId="10" pageNumber="11">
Antennular peduncle (
<figureCitation id="13173BA3FFA31C08CCAC514D296EFE92" box="[459,528,272,298]" captionStart="FIGURE 6" captionStartId="10.[151,255,1890,1914]" captionTargetBox="[186,1360,697,1889]" captionTargetId="figure@10.[151,1436,628,1924]" captionTargetPageId="10" captionText="FIGURE 6. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for B D." httpUri="https://zenodo.org/record/180459/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
B) not reaching midlength of antennal scale. First segment unarmed on dorsodistal margin; stylocerite overreaching distal margin of first segment, acuminate, mesial margin convex or sinuous, closely in touch with first segment; second segment about 0.30 length of first segment, with small spine at dorsolateral distal angle; third segment short, with small spine on dorsodistal margin. Lateral flagellum with thickened aesthetasc-bearing portion 0.300.35 of carapace length.
</paragraph>
<paragraph id="8B932726FFA31C08CDA051852F24FDD2" blockId="10.[151,1437,152,618]" pageId="10" pageNumber="11">
Antenna (
<figureCitation id="13173BA3FFA31C08CC2751852AF3FE4A" box="[320,397,472,498]" captionStart="FIGURE 6" captionStartId="10.[151,255,1890,1914]" captionTargetBox="[186,1360,697,1889]" captionTargetId="figure@10.[151,1436,628,1924]" captionTargetPageId="10" captionText="FIGURE 6. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for B D." httpUri="https://zenodo.org/record/180459/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
B, D) with basicerite bearing moderately large ventrolateral distal tooth; carpocerite reaching 0.300.35 length of antennal scale or distal margin of second segment of antennular peduncle. Antennal scale 1.021.09 length of carapace and 3.303.70 times longer than wide; lateral margin straight; distal lamella rounded, moderately produced, exceeding beyond distolateral tooth.
</paragraph>
<caption id="DF5377AEFFA31C08CDF0573F2FCDF807" httpUri="https://zenodo.org/record/180459/files/figure.png" pageId="10" pageNumber="11" targetBox="[186,1360,697,1889]" targetPageId="10">
<paragraph id="8B932726FFA31C08CDF0573F2FCDF807" blockId="10.[151,1436,1890,1983]" pageId="10" pageNumber="11">
<emphasis id="B958FB34FFA31C08CDF0573F2A67F8C2" bold="true" box="[151,281,1890,1914]" pageId="10" pageNumber="11">FIGURE 6.</emphasis>
<taxonomicName id="4C2C5CA5FFA31C08CC78573E2936F8C2" box="[287,584,1891,1914]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acuticarinatus" status="sp. nov.">
<emphasis id="B958FB34FFA31C08CC78573E2936F8C2" box="[287,584,1891,1914]" italics="true" pageId="10" pageNumber="11">Heptacarpus acuticarinatus</emphasis>
</taxonomicName>
<emphasis id="B958FB34FFA31C08CF29573F29F4F8C2" bold="true" box="[590,650,1890,1914]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A26B464FFFA31C08CF29573F29F4F8C2" box="[590,650,1890,1914]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. A, rostrum and anterior part of carapace, lateral view; B, rostrum, anterior part of carapace and cephalic appendages, dorsal view; C, telson, dorsal view; D, left antenna, ventral view. Scale bars: 2 mm for A; 1 mm for BD.
</paragraph>
</caption>
<caption id="DF5377AEFFA21C09CDF0577D2E66F85D" httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="11" pageNumber="12" targetBox="[186,1397,192,1823]" targetPageId="11">
<paragraph id="8B932726FFA21C09CDF0577D2E66F85D" blockId="11.[151,1437,1824,2021]" pageId="11" pageNumber="12">
<emphasis id="B958FB34FFA21C09CDF0577D2A64F880" bold="true" box="[151,282,1824,1848]" pageId="11" pageNumber="12">FIGURE 7.</emphasis>
<taxonomicName id="4C2C5CA5FFA21C09CC44577C2931F880" box="[291,591,1825,1848]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acuticarinatus" status="sp. nov.">
<emphasis id="B958FB34FFA21C09CC44577C2931F880" box="[291,591,1825,1848]" italics="true" pageId="11" pageNumber="12">Heptacarpus acuticarinatus</emphasis>
</taxonomicName>
<emphasis id="B958FB34FFA21C09CF30577D29EBF880" bold="true" box="[599,661,1824,1848]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A26B464FFFA21C09CF30577D29EBF880" box="[599,661,1824,1848]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for CF, H, K.
</paragraph>
</caption>
<paragraph id="8B932726FFA51C0ECDA050C529F4FEBA" blockId="12.[151,1437,152,818]" pageId="12" pageNumber="13">
Third maxilliped (
<figureCitation id="13173BA3FFA51C0ECCC350C52A8BFF0A" box="[420,501,152,178]" captionStart="FIGURE 5" captionStartId="9.[151,255,1717,1741]" captionTargetBox="[237,1348,1009,1692]" captionTargetId="figure@9.[151,1436,976,1710]" captionTargetPageId="9" captionText="FIGURE 5. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/180458/files/figure.png" pageId="12" pageNumber="13">Figs 5</figureCitation>
,
<figureCitation id="13173BA3FFA51C0ECF6250C5296DFF0A" box="[517,531,152,178]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">7</figureCitation>
A) moderately stout, relatively long, reaching distal 0.150.30 of antennal scale; ultimate segment 2.202.40 length of carpus (= penultimate segment), tapering distally, with several darkly pigmented corneous spines distally.
</paragraph>
<paragraph id="8B932726FFA51C0ECDA0514D2E35FC8A" blockId="12.[151,1437,152,818]" pageId="12" pageNumber="13">
First pereopod (
<figureCitation id="13173BA3FFA51C0ECCE6514D2AB9FE92" box="[385,455,272,298]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
B) moderately stout, reaching nearly to midlength of antennal scale; chela (
<figureCitation id="13173BA3FFA51C0EC85E514D2E01FE92" box="[1337,1407,272,298]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
C) about 1.852.00 of carpal length and 3.503.80 times longer than wide; dactylus 0.500.60 length of palm, terminating in 2 darkly pigmented, strong corneous ungues (
<figureCitation id="13173BA3FFA51C0ECE12513D28C1FEC2" box="[885,959,352,378]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
D); fixed finger terminating 1 corneous unguis (
<figureCitation id="13173BA3FFA51C0ECD9251D52A42FE1A" box="[245,316,392,418]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
D); merus about 1.70 of carpal length and about 3.203.40 times longer than high; dorsolateral distal angle of ischium with small denticle. Second pereopods (
<figureCitation id="13173BA3FFA51C0ECEEF51ED28B0FE72" box="[904,974,432,458]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
E) equal, slightly falling short of distal margin of antennal scale; dactylus 0.700.75 of palm length (
<figureCitation id="13173BA3FFA51C0ECE1D518528BDFE4A" box="[890,963,472,498]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
F); carpus about 3.80 times longer than chela, divided in 7 unequal articles; ischium subequal in length to merus. Third to fifth pereopods relatively long, similar in structure. Third pereopod (
<figureCitation id="13173BA3FFA51C0ECFE0527529B5FDFA" box="[647,715,552,578]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
G) reaching or slightly overreaching distal margin of antennal scale; dactylus (
<figureCitation id="13173BA3FFA51C0ECC35520D2AE9FDD2" box="[338,407,592,618]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
H) 0.250.30 of propodal length, 4.505.00 times longer than deep, terminating in long, acute, pigmented unguis, armed with 5 or 6 accessory spinules on flexor margin; propodus with 2 rows of slender spinules on flexor margin (
<figureCitation id="13173BA3FFA51C0ECF2552FD29F5FD02" box="[578,651,672,698]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
H); carpus 0.400.45 of propodal length; merus 9.009.50 times longer than high, armed with 710 lateral spines; ischium unarmed. Fourth pereopod (
<figureCitation id="13173BA3FFA51C0EC9E852952FABFD5A" box="[1167,1237,712,738]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">
Fig.
<date id="FF9201E6FFA51C0EC9A152952FABFD5A" box="[1222,1237,712,738]" pageId="12" pageNumber="13">7</date>
</figureCitation>
I) reaching distal 0.700.80 of antennal scale; merus with 58 lateral spines. Fifth pereopod (
<figureCitation id="13173BA3FFA51C0EC94B52AD2F0AFCB2" box="[1068,1140,752,778]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
J) reaching midlength of antennal scale; propodus with tufts of grooming setae distally (
<figureCitation id="13173BA3FFA51C0ECE1F534528C0FC8A" box="[888,958,792,818]" captionStart="FIGURE 7" captionStartId="11.[151,255,1824,1848]" captionTargetBox="[186,1397,192,1823]" captionTargetId="figure@11.[168,1422,158,1881]" captionTargetPageId="11" captionText="FIGURE 7. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay. Left appendages (only fifth pereopod from right). A, third maxilliped, lateral view; B, first pereopod, lateral view; C, chela of first pereopod, dorsal (extensor) view; D, dactylus and fixed finger of first pereopod, oblique view; E, second pereopod, lateral view; F, chela of second pereopod; G, third pereopod, lateral view; H, dactylus and distal part of propodus of third pereopod, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view; K, dactylus and distal part of propodus of fifth pereopod, lateral view. Scale bars: 1 mm for A, B, E, G, I, J; 0.5 mm for C F, H, K." httpUri="https://zenodo.org/record/180460/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
K); merus with 36 lateral spines.
</paragraph>
<caption id="DF5377AEFFA51C0ECDF0570F294FF86A" httpUri="https://zenodo.org/record/180461/files/figure.png" pageId="12" pageNumber="13" targetBox="[179,1393,912,1873]" targetPageId="12">
<paragraph id="8B932726FFA51C0ECDF0570F294FF86A" blockId="12.[151,1436,1874,2002]" pageId="12" pageNumber="13">
<emphasis id="B958FB34FFA51C0ECDF0570F2A64F8D2" bold="true" box="[151,282,1874,1898]" pageId="12" pageNumber="13">FIGURE 8.</emphasis>
<taxonomicName id="4C2C5CA5FFA51C0ECC45570E2930F8D2" box="[290,590,1875,1898]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="acuticarinatus" status="sp. nov.">
<emphasis id="B958FB34FFA51C0ECC45570E2930F8D2" box="[290,590,1875,1898]" italics="true" pageId="12" pageNumber="13">Heptacarpus acuticarinatus</emphasis>
</taxonomicName>
<emphasis id="B958FB34FFA51C0ECF32570F29ECF8D2" bold="true" box="[597,658,1874,1898]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A26B464FFFA51C0ECF32570F29ECF8D2" box="[597,658,1874,1898]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for AC; 0.5 mm for D, E.
</paragraph>
</caption>
<paragraph id="8B932726FFA41C0FCDA050C52A3CFF62" blockId="13.[151,1436,152,938]" pageId="13" pageNumber="14">
Gill formula as in
<tableCitation id="C6AE129DFFA41C0FCCC250C52A80FF0A" box="[421,510,152,178]" captionStart="TABLE 1" captionStartId="6.[151,239,1271,1295]" captionTargetBox="[159,1359,1328,1649]" captionTargetPageId="6" captionText="TABLE 1. Branchial formula of the four Heptacarpus species treated in this study." httpUri="http://table.plazi.org/id/DF5377AEFFAF1C04CDF054AA2F72FAB7" pageId="13" pageNumber="14" tableUuid="DF5377AEFFAF1C04CDF054AA2F72FAB7">Table 1</tableCitation>
. Only third maxilliped with strap-like epipod corresponding to setobranch on first pereopod.
</paragraph>
<paragraph id="8B932726FFA41C0FCDA050B52F83FEBA" blockId="13.[151,1436,152,938]" box="[199,1277,232,258]" pageId="13" pageNumber="14">
Uropod (
<figureCitation id="13173BA3FFA41C0FCC5750B52A08FEBA" box="[304,374,232,258]" captionStart="FIGURE 5" captionStartId="9.[151,255,1717,1741]" captionTargetBox="[237,1348,1009,1692]" captionTargetId="figure@9.[151,1436,976,1710]" captionTargetPageId="9" captionText="FIGURE 5. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/180458/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
) with both rami reaching or slightly overreaching posterior margin of telson.
</paragraph>
<paragraph id="8B932726FFA41C0FCDA051522FBEFDD2" blockId="13.[151,1436,152,938]" pageId="13" pageNumber="14">
<emphasis id="B958FB34FFA41C0FCDA051522AB5FE91" bold="true" box="[199,459,271,297]" pageId="13" pageNumber="14">Description of males</emphasis>
. Body slightly more slender than in females (
<figureCitation id="13173BA3FFA41C0FCE9E514D2F3FFE92" box="[1017,1089,272,298]" captionStart="FIGURE 8" captionStartId="12.[151,255,1874,1898]" captionTargetBox="[179,1393,912,1873]" captionTargetId="figure@12.[144,1429,851,1951]" captionTargetPageId="12" captionText="FIGURE 8. Heptacarpus acuticarinatus n. sp. Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for A C; 0.5 mm for D, E." httpUri="https://zenodo.org/record/180461/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
A, C). Rostrum (
<figureCitation id="13173BA3FFA41C0FC868514D2E29FE92" box="[1295,1367,272,298]" captionStart="FIGURE 8" captionStartId="12.[151,255,1874,1898]" captionTargetBox="[179,1393,912,1873]" captionTargetId="figure@12.[144,1429,851,1951]" captionTargetPageId="12" captionText="FIGURE 8. Heptacarpus acuticarinatus n. sp. Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for A C; 0.5 mm for D, E." httpUri="https://zenodo.org/record/180461/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
A, B) 1.421.57 length of carapace, anterior 0.160.24 unarmed. Pleon (
<figureCitation id="13173BA3FFA41C0FCEC851652888FEEA" box="[943,1014,312,338]" captionStart="FIGURE 8" captionStartId="12.[151,255,1874,1898]" captionTargetBox="[179,1393,912,1873]" captionTargetId="figure@12.[144,1429,851,1951]" captionTargetPageId="12" captionText="FIGURE 8. Heptacarpus acuticarinatus n. sp. Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for A C; 0.5 mm for D, E." httpUri="https://zenodo.org/record/180461/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
C) weakly geniculate; third pleonal tergite evenly convex in posterior part. Corneal diameter about 0.200.22 of carapace length (
<figureCitation id="13173BA3FFA41C0FC982513D2E54FEC2" box="[1253,1322,352,378]" captionStart="FIGURE 8" captionStartId="12.[151,255,1874,1898]" captionTargetBox="[179,1393,912,1873]" captionTargetId="figure@12.[144,1429,851,1951]" captionTargetPageId="12" captionText="FIGURE 8. Heptacarpus acuticarinatus n. sp. Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for A C; 0.5 mm for D, E." httpUri="https://zenodo.org/record/180461/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
A). Outer flagellum of antennule larger than in females, thickened aesthetasc-bearing portion about 0.50 length of carapace (
<figureCitation id="13173BA3FFA41C0FCDBB51ED2A5FFE72" box="[220,289,432,458]" captionStart="FIGURE 8" captionStartId="12.[151,255,1874,1898]" captionTargetBox="[179,1393,912,1873]" captionTargetId="figure@12.[144,1429,851,1951]" captionTargetPageId="12" captionText="FIGURE 8. Heptacarpus acuticarinatus n. sp. Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for A C; 0.5 mm for D, E." httpUri="https://zenodo.org/record/180461/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
A). Antennal scale 1.141.25 times longer than carapace. Third to fifth pereopods similar to those of females. Endopod of first pleopod (
<figureCitation id="13173BA3FFA41C0FCF38518529DBFE4A" box="[607,677,472,498]" captionStart="FIGURE 8" captionStartId="12.[151,255,1874,1898]" captionTargetBox="[179,1393,912,1873]" captionTargetId="figure@12.[144,1429,851,1951]" captionTargetPageId="12" captionText="FIGURE 8. Heptacarpus acuticarinatus n. sp. Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for A C; 0.5 mm for D, E." httpUri="https://zenodo.org/record/180461/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
D) elongate subtriangular, with conspicuous appendix interna at terminal position; distolateral lobule delineated; mesial margin with row of small spiniform setae, lateral margin with row of long plumose setae. Second pleopod with appendix masculina slightly shorter than appendix interna, with numerous setae increasing in length distally on dorsal surface to tip (
<figureCitation id="13173BA3FFA41C0FC930520D2FE2FDD2" box="[1111,1180,592,618]" captionStart="FIGURE 8" captionStartId="12.[151,255,1874,1898]" captionTargetBox="[179,1393,912,1873]" captionTargetId="figure@12.[144,1429,851,1951]" captionTargetPageId="12" captionText="FIGURE 8. Heptacarpus acuticarinatus n. sp. Paratype, male (cl 3.7 mm; HUMZ-C 1174), Kashima-nada Sea, northeastern Honshu. A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view; C, pleon, lateral view; D, endopod of left first pleopod, ventral view; E, appendices interna and masculina of second pleopod, mesial view. Scale bars: 1 mm for A C; 0.5 mm for D, E." httpUri="https://zenodo.org/record/180461/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
E).
</paragraph>
<paragraph id="8B932726FFA41C0FCDA0522A2FF6FD2A" blockId="13.[151,1436,152,938]" box="[199,1160,631,658]" pageId="13" pageNumber="14">
<emphasis id="B958FB34FFA41C0FCDA0522A2B84FD29" bold="true" box="[199,250,631,657]" pageId="13" pageNumber="14">Size</emphasis>
. Females cl 3.0
<quantity id="4CD48AC3FFA41C0FCCA652252963FD2A" box="[449,541,632,658]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="13" pageNumber="14" unit="mm" value="6.8">6.8 mm</quantity>
, ovigerous females cl
<quantity id="4CD48AC3FFA41C0FCE45522528CDFD2A" box="[802,947,632,658]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="6.8" metricValueMin="4.4" pageId="13" pageNumber="14" unit="mm" value="5.6" valueMax="6.8" valueMin="4.4">4.46.8 mm</quantity>
; males cl
<quantity id="4CD48AC3FFA41C0FC94052252FFBFD2A" box="[1063,1157,632,658]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="13" pageNumber="14" unit="mm" value="5.2">5.2 mm</quantity>
.
</paragraph>
<paragraph id="8B932726FFA41C0FCDA052C22FEFFD5A" blockId="13.[151,1436,152,938]" pageId="13" pageNumber="14">
<emphasis id="B958FB34FFA41C0FCDA052C22A43FD01" bold="true" box="[199,317,671,697]" pageId="13" pageNumber="14">Variation</emphasis>
. A total of
<specimenCount id="9D2AECAFFFA41C0FCCAA52FD290CFD02" box="[461,626,672,698]" pageId="13" pageNumber="14" type="generic">25 specimens</specimenCount>
, including
<specimenCount id="9D2AECAFFFA41C0FCF9F52FD2805FD02" box="[760,891,672,698]" pageId="13" pageNumber="14" type="female">23 females</specimenCount>
and two males, were examined for assessing morphological variation in some characters possibly providing diagnostic significance.
</paragraph>
<paragraph id="8B932726FFA41C0FCDA052AD2F5DFCB2" blockId="13.[151,1436,152,938]" box="[199,1059,752,778]" pageId="13" pageNumber="14">
The number of the rostral ventral teeth varies from five to eight (
<figureCitation id="13173BA3FFA41C0FCED952AD2F6AFCB2" box="[958,1044,752,778]" captionStart="FIGURE 18" captionStartId="25.[151,255,1068,1092]" captionTargetBox="[170,1419,308,1053]" captionTargetId="figure@25.[162,1427,300,1059]" captionTargetPageId="25" captionText="FIGURE 18. Variation in the number of rostral ventral teeth of four Heptacarpus species." httpUri="https://zenodo.org/record/180471/files/figure.png" pageId="13" pageNumber="14">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8B932726FFA41C0FCDA053452E4FFC8A" blockId="13.[151,1436,152,938]" box="[199,1329,792,818]" pageId="13" pageNumber="14">In all the specimens examined, the carapace is provided with pterygostomial teeth on both sides.</paragraph>
<paragraph id="8B932726FFA41C0FCDA0531D2ABBFC12" blockId="13.[151,1436,152,938]" pageId="13" pageNumber="14">
The number of the meral spines on the third pereopod varies from seven to 10 (
<figureCitation id="13173BA3FFA41C0FC915531D2FB7FCE2" box="[1138,1225,832,858]" captionStart="FIGURE 19" captionStartId="26.[151,255,1443,1467]" captionTargetBox="[157,1428,463,1421]" captionTargetId="figure@26.[151,1436,456,1426]" captionTargetPageId="26" captionText="FIGURE 19. Variation in the number of meral spines on the third pereopod of four Heptacarpus species." httpUri="https://zenodo.org/record/180472/files/figure.png" pageId="13" pageNumber="14">Fig. 19</figureCitation>
), but the majority of the examined specimens (23 of
<specimenCount id="9D2AECAFFFA41C0FCF57533529ADFC3A" box="[560,723,872,898]" pageId="13" pageNumber="14" type="generic">25 specimens</specimenCount>
; 92 %) have seven to nine spines. Other two specimens (8.0 %) have ten meral spines.
</paragraph>
<caption id="DF5377AEFFA41C0FCDF0577F29D7F8E4" httpUri="https://zenodo.org/record/180462/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1436,978,1825]" targetPageId="13">
<paragraph id="8B932726FFA41C0FCDF0577F29D7F8E4" blockId="13.[151,1436,1826,1884]" pageId="13" pageNumber="14">
<emphasis id="B958FB34FFA41C0FCDF0577F2A65F882" bold="true" box="[151,283,1826,1850]" pageId="13" pageNumber="14">FIGURE 9.</emphasis>
<taxonomicName id="4C2C5CA5FFA41C0FCC44577E2931F882" box="[291,591,1827,1850]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="acuticarinatus" status="sp. nov.">
<emphasis id="B958FB34FFA41C0FCC44577E2931F882" box="[291,591,1827,1850]" italics="true" pageId="13" pageNumber="14">Heptacarpus acuticarinatus</emphasis>
</taxonomicName>
<emphasis id="B958FB34FFA41C0FCF30577F29EBF882" bold="true" box="[599,661,1826,1850]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A26B464FFFA41C0FCF30577F29EBF882" box="[599,661,1826,1850]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view, showing coloration in life.
</paragraph>
</caption>
<paragraph id="8B932726FFA41C0FCDA057CD2FECF812" blockId="13.[199,1228,1936,2002]" box="[199,1170,1936,1962]" pageId="13" pageNumber="14">
<emphasis id="B958FB34FFA41C0FCDA057CD2AE2F812" bold="true" box="[199,412,1936,1962]" pageId="13" pageNumber="14">Coloration in life</emphasis>
. Body and appendages generally pale pink; cornea gray (
<figureCitation id="13173BA3FFA41C0FC95B57CD2FFAF812" box="[1084,1156,1936,1962]" captionStart="FIGURE 9" captionStartId="13.[151,255,1826,1850]" captionTargetBox="[151,1436,978,1825]" captionTargetId="figure@13.[151,1436,950,1852]" captionTargetPageId="13" captionText="FIGURE 9. Heptacarpus acuticarinatus n. sp. Holotype, ovigerous female (cl 5.4 mm; CBM-ZC 8980), Sagami Bay, habitus in lateral view, showing coloration in life." httpUri="https://zenodo.org/record/180462/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33674ADFFA41C0FCDA057E52FB2F86A" box="[199,1228,1976,2002]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8B932726FFA41C0FCDA057E52FB2F86A" blockId="13.[199,1228,1936,2002]" box="[199,1228,1976,2002]" pageId="13" pageNumber="14">
<emphasis id="B958FB34FFA41C0FCDA057E52A1EF86A" bold="true" box="[199,352,1976,2002]" pageId="13" pageNumber="14">Distribution</emphasis>
. Southern Hokkaido to Kyushu,
<collectingCountry id="F33B67B6FFA41C0FCF8557E52856F86A" box="[738,808,1976,2002]" name="Japan" pageId="13" pageNumber="14">Japan</collectingCountry>
,
<collectingCountry id="F33B67B6FFA41C0FCE5357E52800F86A" box="[820,894,1976,2002]" name="South Korea" pageId="13" pageNumber="14">Korea</collectingCountry>
, and Yellow Sea,
<quantity id="4CD48AC3FFA41C0FC92857E52FB6F86A" box="[1103,1224,1976,2002]" metricMagnitude="1" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="13" pageNumber="14" unit="m" value="90.0" valueMax="150.0" valueMin="30.0">
30
<specimenCount id="9D2AECAFFFA41C0FC91B57E52FB6F86A" box="[1148,1224,1976,2002]" pageId="13" pageNumber="14" type="generic">150 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33674ADFFA71C0CCDA050CA29C1FD2A" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B932726FFA71C0CCDA050CA29C1FD2A" blockId="14.[151,1437,151,738]" pageId="14" pageNumber="15">
<emphasis id="B958FB34FFA71C0CCDA050CA2A47FF09" bold="true" box="[199,313,151,177]" pageId="14" pageNumber="15">Remarks</emphasis>
. The new species appears closest to
<taxonomicName id="4C2C5CA5FFA71C0CCF9150C52F43FF09" box="[758,1085,152,177]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA71C0CCF9150C52F43FF09" box="[758,1085,152,177]" italics="true" pageId="14" pageNumber="15">Heptacarpus camtschaticus</emphasis>
</taxonomicName>
, with which it has been confounded, but is readily distinguished from the latter by a number of characters, including the sharp lateral carina of the rostrum and the relatively longer pereopods with more numerous meral spines (see “Comparison” for details). The present study strongly suggests that
<taxonomicName id="4C2C5CA5FFA71C0CCE6A514D28A7FE91" box="[781,985,272,297]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA71C0CCE6A514D28A7FE91" box="[781,985,272,297]" italics="true" pageId="14" pageNumber="15">H. camtschaticus</emphasis>
</taxonomicName>
does not occur in the Pacific coast of Honshu to Kyushu,
<collectingCountry id="F33B67B6FFA71C0CCCE651652AB9FEEA" box="[385,455,312,338]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
. Thus, the records of
<taxonomicName id="4C2C5CA5FFA71C0CCFA3516528F3FEE9" box="[708,909,312,337]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA71C0CCFA3516528F3FEE9" box="[708,909,312,337]" italics="true" pageId="14" pageNumber="15">H. camtschaticus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFBD5AD7FFA71C0CCEDE51652F33FEEA" author="Balss" box="[953,1101,312,338]" pageId="14" pageNumber="31" refString="Balss, H. (1914) Ostasiatische Decapoden II: Die Natantia und Reptantia. In: Doflein, F. (Ed.), Beitrage zur Naturgeschichte Ostasiens, Dekapoden, part 7. Abhandlungen der Bayerischen Akademie der Wissenschaften, Munchen, 2 (Supplement), 10, 1 - 101, pl. 1." type="journal article" year="1914">Balss (1914)</bibRefCitation>
and
<bibRefCitation id="EFBD5AD7FFA71C0CC9E051652E60FEEA" author="Parisi" box="[1159,1310,312,338]" pageId="14" pageNumber="34" refString="Parisi, B. (1919) I Decapodi giapponesi del Museo di Milano. VII. Natantia. Atii della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 58, 59 - 99, pls 2 - 6." type="journal article" year="1919">Parisi (1919)</bibRefCitation>
from Sagami Bay and
<bibRefCitation id="EFBD5AD7FFA71C0CCC51513D2A99FEC2" box="[310,487,352,378]" pageId="14" pageNumber="34" refString="Yokoya, Y. (1933) On the distribution of decapod Crustacea inhabiting the continental shelf around Japan, chiefly based upon the materials collected by S. S. &quot; Soyo Maru &quot; during the years 1923 - 1930. Journal of the College of Agriculture, Tokyo Imperial University, 12: 1 - 236." type="journal article">Yokoya (1933)</bibRefCitation>
from Aichi Prefecture (all as
<taxonomicName id="4C2C5CA5FFA71C0CCE2B513D2894FEC1" box="[844,1002,352,377]" class="Malacostraca" family="Hippolytidae" genus="Spirontocaris" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B958FB34FFA71C0CCE2B513D2894FEC1" box="[844,1002,352,377]" italics="true" pageId="14" pageNumber="15">Spirontocaris</emphasis>
</taxonomicName>
) are referred to
<taxonomicName id="4C2C5CA5FFA71C0CC9CA513D2E02FEC1" box="[1197,1404,352,377]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="acuticarinatus" status="sp. nov.">
<emphasis id="B958FB34FFA71C0CC9CA513D2E02FEC1" box="[1197,1404,352,377]" italics="true" pageId="14" pageNumber="15">H. acuticarinatus</emphasis>
</taxonomicName>
<emphasis id="B958FB34FFA71C0CC8E351022BC4FE19" bold="true" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A26B464FFFA71C0CC8E351022BC4FE19" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
There is little doubt that the specimen from Amakusa, Kyushu, cited as
<taxonomicName id="4C2C5CA5FFA71C0CC94651D52F93FE19" box="[1057,1261,392,417]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA71C0CC94651D52F93FE19" box="[1057,1261,392,417]" italics="true" pageId="14" pageNumber="15">H. camtschaticus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFBD5AD7FFA71C0CC87B51D52A34FE72" author="Hayashi" pageId="14" pageNumber="32" refString="Hayashi, K. &amp; Miyake, S. (1968) Studies on the hippolytid shrimps of Japan, V. Hippolytid fauna of the sea around the Amakusa Marine Biological Laboratory. OHMU, 1, 121 - 163." type="journal article" year="1968">Hayashi &amp; Miyake (1968)</bibRefCitation>
,
<bibRefCitation id="EFBD5AD7FFA71C0CCC3051ED29EEFE72" author="Kikuchi" box="[343,656,432,458]" pageId="14" pageNumber="32" refString="Kikuchi, T. &amp; Miyake, S. (1978) Fauna and Flora of the Sea around the Amakusa Marine Biological Laboratory. Part II. Decapoda Crustacea (Revised Edition). Amakusa Marine Biological Laboratory, Tomioka, 52 pp." type="book" year="1978">Kikuchi &amp; Miyake (1978)</bibRefCitation>
,
<bibRefCitation id="EFBD5AD7FFA71C0CCFF951ED2833FE72" author="Hayashi" box="[670,845,432,458]" pageId="14" pageNumber="32" refString="Hayashi, K. (1979) Studies on hippolytid shrimps from Japan VII: The genus Heptacarpus Holmes. Journal of Shimonoseki University of Fisheries, 28, 11 - 32." type="journal article" year="1979">Hayashi (1979</bibRefCitation>
,
<bibRefCitation id="EFBD5AD7FFA71C0CCE3D51ED28E7FE72" author="Hayashi" box="[858,921,432,458]" pageId="14" pageNumber="32" refString="Hayashi, K. (1992) Prawns, shrimps and lobsters from Japan (63 - 68). Family Hippolytidae. Aquabiology, 14, 24 - 28, 108 - 112, 180 - 184, 270 - 274, 341 - 345, 436 - 439." type="journal article" year="1992">1992</bibRefCitation>
), is identical with the new species, because the morphological attributes described or shown in these references closely fit those of the new species. Similarly, the occurrence of
<taxonomicName id="4C2C5CA5FFA71C0CCCD2525D29FBFDA1" box="[437,645,512,537]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="acuticarinatus" status="sp. nov.">
<emphasis id="B958FB34FFA71C0CCCD2525D29FBFDA1" box="[437,645,512,537]" italics="true" pageId="14" pageNumber="15">H. acuticarinatus</emphasis>
</taxonomicName>
<emphasis id="B958FB34FFA71C0CCFE951A229ADFDA1" bold="true" box="[654,723,511,537]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A26B464FFFA71C0CCFE951A229ADFDA1" box="[654,723,511,537]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
instead of
<taxonomicName id="4C2C5CA5FFA71C0CCE3C525D2F59FDA1" box="[859,1063,512,537]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA71C0CCE3C525D2F59FDA1" box="[859,1063,512,537]" italics="true" pageId="14" pageNumber="15">H. camtschaticus</emphasis>
</taxonomicName>
has been confirmed in Korean waters. Therefore, Korean records of
<taxonomicName id="4C2C5CA5FFA71C0CCF3352752861FDF9" box="[596,799,552,577]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA71C0CCF3352752861FDF9" box="[596,799,552,577]" italics="true" pageId="14" pageNumber="15">H. camtschaticus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFBD5AD7FFA71C0CCE2B52752F6BFDFA" box="[844,1045,552,578]" pageId="14" pageNumber="32" refString="Cha, H. K., Lee, J. U., Park, C. S., Baik, C. I., Hong, S. Y., Park, J. H., Lee, D. W., Choi, Y. M., Hwang, K., Kim, Z. G., Choi, K. H., Sohn, H., Sohn, M. H., Kim, D. H. &amp; Choi, J. H. (2001) Shrimps of the Korean Waters. National Fisheries Research and Development Institute, Pusan, 188 pp." type="book">
Cha
<emphasis id="B958FB34FFA71C0CCEE3527528BEFDF9" box="[900,960,552,577]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
(2001)
</bibRefCitation>
are also referred to the new species. Regarding the geographical range, the record of
<taxonomicName id="4C2C5CA5FFA71C0CCE47520D2892FDD1" box="[800,1004,592,617]" class="Malacostraca" family="Hippolytidae" genus="Heptacarpus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camtschaticus">
<emphasis id="B958FB34FFA71C0CCE47520D2892FDD1" box="[800,1004,592,617]" italics="true" pageId="14" pageNumber="15">H. camtschaticus</emphasis>
</taxonomicName>
from northern
<collectingCountry id="F33B67B6FFA71C0CC9C1520D2F92FDD2" box="[1190,1260,592,618]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
(
<bibRefCitation id="EFBD5AD7FFA71C0CC998520D2E08FDD2" author="Liu" box="[1279,1398,592,618]" pageId="14" pageNumber="33" refString="Liu, J. Y. (1963) Zoogeographical Studies on the macrurous crustacean fauna of the Yellow Sea and the East China Sea. Oceanologia et Limnologia Sinica, 5 (3), 230 - 244. [in Chinese]" type="journal article" year="1963">Liu, 1963</bibRefCitation>
) is also most probably referred to the new species.
</paragraph>
</subSubSection>
<subSubSection id="C33674ADFFA71C0CCDA052C228F2FD5A" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8B932726FFA71C0CCDA052C228F2FD5A" blockId="14.[151,1437,151,738]" pageId="14" pageNumber="15">
<emphasis id="B958FB34FFA71C0CCDA052C22A32FD01" bold="true" box="[199,332,671,697]" pageId="14" pageNumber="15">Etymology</emphasis>
. The species name is a combination of the Latin
<emphasis id="B958FB34FFA71C0CCEF852FD2892FD01" box="[927,1004,672,697]" italics="true" pageId="14" pageNumber="15">acutus</emphasis>
(= sharp) and
<emphasis id="B958FB34FFA71C0CC9F952FD2E73FD01" box="[1182,1293,672,697]" italics="true" pageId="14" pageNumber="15">carinatus</emphasis>
(ridged), in reference to the characteristic sharp lateral carina of the rostrum.
</paragraph>
</subSubSection>
</treatment>
</document>