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<mods:title id="E3FA56671A68F248FBCE77493714E510">Cephalops Fallén and Semicephalops De Meyer (Diptera: Pipunculidae) of Colombia, with description of five new species and an updated key to males of the Neotropical species</mods:title>
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<mods:namePart id="2178B284668F1F1C7EEC3EA1F77DA451">Marques, Dayse Willkenia A.</mods:namePart>
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<mods:affiliation id="614134042BC98B33F42FC39F52CADD9F">Instituto Nacional de Pesquisas da Amazônia, INPA, Manaus, Amazonas, Brazil. &amp; willkenia @ gmail. com; https: // orcid. org / 0000 - 0001 - 7260 - 5760</mods:affiliation>
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<mods:namePart id="5C6A65A59BA4F1661640FE22DDA0C87D">Rafael, José Albertino</mods:namePart>
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<mods:affiliation id="8C474DE38FBC05B142DED50CFB9D8815">Instituto Nacional de Pesquisas da Amazônia, INPA, Manaus, Amazonas, Brazil. &amp; jarafael @ inpa. gov. br; https: // orcid. org / 0000 - 0002 - 0170 - 0514</mods:affiliation>
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<paragraph id="4F44402AFFDA4847FF4ED255FDDB2FC5" blockId="17.[151,629,1645,1707]" box="[151,629,1645,1672]" pageId="17" pageNumber="218">
<heading id="140CF746FFDA4847FF4ED255FDDB2FC5" bold="true" box="[151,629,1645,1672]" fontSize="11" level="1" pageId="17" pageNumber="218" reason="1">
<taxonomicName id="88FB3BA9FFDA4847FF4ED255FDDB2FC5" ID-CoL="SDYC" authority="Rafael, 1990" authorityName="Rafael" authorityYear="1990" box="[151,629,1645,1672]" class="Insecta" family="Pipunculidae" genus="Semicephalops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="inpaganus">
<emphasis id="7D8F9C38FFDA4847FF4ED255FDDB2FC5" bold="true" box="[151,629,1645,1672]" pageId="17" pageNumber="218">
<emphasis id="7D8F9C38FFDA4847FF4ED255FE7C2FC5" bold="true" box="[151,466,1645,1672]" italics="true" pageId="17" pageNumber="218">Semicephalops inpaganus</emphasis>
<bibRefCitation id="2B6A3DDBFFDA4847FE03D255FDDB2FC5" author="Rafael, J. A." box="[474,629,1645,1671]" pageId="17" pageNumber="218" pagination="353 - 390" refId="ref19665" refString="Rafael, J. A. (1990) Revisao das especies neotropicais do genero Cephalops Fallen (Diptera: Pipunculidae). Acta Amazonica, 20, 353 - 390." type="journal article" year="1990">Rafael, 1990</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="07E113A1FFDA4847FF4ED2A9FEF42FE9" box="[151,346,1681,1707]" pageId="17" pageNumber="218" type="description">
<paragraph id="4F44402AFFDA4847FF4ED2A9FEF42FE9" blockId="17.[151,629,1645,1707]" box="[151,346,1681,1707]" pageId="17" pageNumber="218">
<figureCitation id="D7C05CAFFFDA4847FF4ED2A9FE8B2FE9" box="[151,293,1681,1707]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="17" pageNumber="218">Figs 94106</figureCitation>
,
<figureCitation id="D7C05CAFFFDA4847FEE9D2A9FEF42FE9" box="[304,346,1681,1707]" captionStart="FIGURE 108" captionStartId="20.[151,250,1701,1725]" captionTargetBox="[151,1436,669,1677]" captionTargetId="figure-187@20.[151,1436,669,1677]" captionTargetPageId="20" captionText="FIGURE 108. Geographical records of Semicephalops species in Colombia. Semicephalops folium sp. nov. and C. inpaganus." figureDoi="http://doi.org/10.5281/zenodo.6592688" httpUri="https://zenodo.org/record/6592688/files/figure.png" pageId="17" pageNumber="218">108</figureCitation>
</paragraph>
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<subSubSection id="07E113A1FFDA4847FF4ED2E8FABD2E67" pageId="17" pageNumber="218" type="reference_group">
<paragraph id="4F44402AFFDA4847FF4ED2E8FCDB2FAA" blockId="17.[151,1299,1744,1830]" box="[151,885,1744,1768]" pageId="17" pageNumber="218">
<treatmentCitationGroup id="6FEB6704FFDA4847FF4ED2E8FCDB2FAA" box="[151,885,1744,1768]" pageId="17" pageNumber="218">
<treatmentCitation id="CE5A663BFFDA4847FF4ED2E8FD062FAA" author="Hardy, D. E." box="[151,680,1744,1768]" page="20" pageId="17" pageNumber="218" year="1965">
<taxonomicName id="88FB3BA9FFDA4847FF4ED2E8FD062FAA" ID-CoL="8DDCB" authority="Hardy, 1965 b: 20" authorityName="Hardy" authorityPageNumber="20" authorityYear="1965" box="[151,680,1744,1768]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="paganus" subGenus="Pipunculus">
<emphasis id="7D8F9C38FFDA4847FF4ED2E8FEA72FAA" box="[151,265,1744,1768]" italics="true" pageId="17" pageNumber="218">Pipunculus</emphasis>
(
<emphasis id="7D8F9C38FFDA4847FEC1D2E8FE242FAA" box="[280,394,1744,1768]" italics="true" pageId="17" pageNumber="218">Pipunculus</emphasis>
)
<emphasis id="7D8F9C38FFDA4847FE40D2E9FE412FAA" box="[409,495,1745,1768]" italics="true" pageId="17" pageNumber="218">paganus</emphasis>
<bibRefCitation id="2B6A3DDBFFDA4847FE2CD2E8FD062FAA" author="Hardy, D. E." box="[501,680,1744,1768]" pageId="17" pageNumber="218" pagination="1 - 68" refId="ref18966" refString="Hardy, D. E. (1965 b) Neotropical Pipunculidae (Diptera) studies. Part Iv. Further studies of Brazilian species. Arquivos de Zoologia, 14, 1 - 68." type="journal article" year="1965">Hardy, 1965b: 20</bibRefCitation>
</taxonomicName>
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, figs 4 ae (partim).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4F44402AFFDA4847FF4ED2D7FC162E45" blockId="17.[151,1299,1744,1830]" box="[151,952,1775,1799]" pageId="17" pageNumber="218">
<treatmentCitationGroup id="6FEB6704FFDA4847FF4ED2D7FC162E45" box="[151,952,1775,1799]" pageId="17" pageNumber="218">
<treatmentCitation id="CE5A663BFFDA4847FF4ED2D7FD9F2E45" author="Rafael, J. A." box="[151,561,1775,1799]" page="362" pageId="17" pageNumber="218" year="1990">
<taxonomicName id="88FB3BA9FFDA4847FF4ED2D7FD9F2E45" ID-CoL="8BCT6" authority="Rafael, 1990: 362" authorityName="Rafael" authorityPageNumber="362" authorityYear="1990" box="[151,561,1775,1799]" class="Insecta" family="Pipunculidae" genus="Cephalops" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="inpaganus">
<emphasis id="7D8F9C38FFDA4847FF4ED2D7FEDA2E45" box="[151,372,1775,1799]" italics="true" pageId="17" pageNumber="218">Cephalops inpaganus</emphasis>
<bibRefCitation id="2B6A3DDBFFDA4847FEA3D2D7FD9F2E45" author="Rafael, J. A." box="[378,561,1775,1799]" pageId="17" pageNumber="218" pagination="353 - 390" refId="ref19665" refString="Rafael, J. A. (1990) Revisao das especies neotropicais do genero Cephalops Fallen (Diptera: Pipunculidae). Acta Amazonica, 20, 353 - 390." type="journal article" year="1990">Rafael, 1990: 362</bibRefCitation>
</taxonomicName>
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, figs 1724, 116;
<treatmentCitation id="CE5A663BFFDA4847FD3FD2D7FC1A2E45" author="De Meyer, M." box="[742,948,1775,1799]" page="98" pageId="17" pageNumber="218" year="1992">
<bibRefCitation id="2B6A3DDBFFDA4847FD3FD2D7FC1A2E45" author="De Meyer, M." box="[742,948,1775,1799]" pageId="17" pageNumber="218" pagination="93 - 99" refId="ref18490" refString="De Meyer, M. (1992) Description of new Cephalops species (Pipunculidae, Diptera) from the Oriental Region. Bulletin de l'Institut royal des Sciences naturelles de Belgique, 62, 93 - 99." type="journal article" year="1992">De Meyer, 1992: 98</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="4F44402AFFDA4847FF4ED335FABD2E67" blockId="17.[151,1299,1744,1830]" box="[151,1299,1805,1830]" pageId="17" pageNumber="218">
<treatmentCitationGroup id="6FEB6704FFDA4847FF4ED335FABD2E67" box="[151,1299,1805,1830]" pageId="17" pageNumber="218">
<treatmentCitation id="CE5A663BFFDA4847FF4ED335FD592E64" author="De Meyer, M." box="[151,759,1805,1830]" page="14" pageId="17" pageNumber="218" year="1994">
<taxonomicName id="88FB3BA9FFDA4847FF4ED335FD592E64" ID-CoL="8BCT6" authority="De Meyer, 1994: 14" authorityName="De Meyer" authorityPageNumber="14" authorityYear="1994" box="[151,759,1805,1830]" class="Insecta" family="Pipunculidae" genus="Cephalops" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="inpaganus" subGenus="Semicephalops">
<emphasis id="7D8F9C38FFDA4847FF4ED335FEAD2E67" box="[151,259,1805,1829]" italics="true" pageId="17" pageNumber="218">Cephalops</emphasis>
(
<emphasis id="7D8F9C38FFDA4847FECBD335FE042E67" box="[274,426,1805,1829]" italics="true" pageId="17" pageNumber="218">Semicephalops</emphasis>
)
<emphasis id="7D8F9C38FFDA4847FE60D336FD8D2E67" box="[441,547,1806,1829]" italics="true" pageId="17" pageNumber="218">inpaganus</emphasis>
<bibRefCitation id="2B6A3DDBFFDA4847FDF0D336FD592E64" author="De Meyer, M." box="[553,759,1806,1830]" pageId="17" pageNumber="218" pagination="7 - 18" refId="ref18581" refString="De Meyer, M. (1994) Phylogenetic relationship within the Cephalopsini (Diptera, Pipunculidae). Bulletin et annales de la Socieitei royale belge d'entomologie, 130, 7 - 18." type="journal article" year="1994">De Meyer, 1994: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="CE5A663BFFDA4847FCD8D336FC612E67" author="De Meyer, M." box="[769,975,1805,1830]" page="36" pageId="17" pageNumber="218" year="1996">
<bibRefCitation id="2B6A3DDBFFDA4847FCD8D336FC612E67" author="De Meyer, M." box="[769,975,1805,1830]" pageId="17" pageNumber="218" pagination="1 - 127" refId="ref18616" refString="De Meyer, M. (1996) World catalogue of Pipunculidae (Diptera). Documents de travail de l'Institut royal des Sciences naturelles de Belgique, 86, 1 - 127." type="journal article" year="1996">De Meyer, 1996: 36</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CE5A663BFFDA4847FC03D335FAA02E67" author="Rodriguez, H. C. &amp; Rafael, J. A." box="[986,1294,1805,1830]" page="16" pageId="17" pageNumber="218" year="2012">
<bibRefCitation id="2B6A3DDBFFDA4847FC03D335FAA02E67" author="Rodriguez, H. C. &amp; Rafael, J. A." box="[986,1294,1805,1830]" pageId="17" pageNumber="218" refId="ref19882" refString="Rodriguez, H. C. &amp; Rafael, J. A. (2012) Pipunculidae (Diptera) of the Latin America and the Caribbean: A catalog of Species with Notes on Biology and Pipunculid-Host Associations. Lambert Academic Publishing, Saarbrucken, 48 pp." type="book" year="2012">Rodriguez &amp; Rafael, 2012: 16</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="07E113A1FFDA4845FF4ED376FE182A8A" lastPageId="19" lastPageNumber="220" pageId="17" pageNumber="218" type="diagnosis">
<paragraph id="4F44402AFFDA4847FF4ED376FD872EBB" blockId="17.[151,1437,1870,2041]" pageId="17" pageNumber="218">
<emphasis id="7D8F9C38FFDA4847FF4ED376FEBC2E2A" bold="true" box="[151,274,1870,1896]" pageId="17" pageNumber="218">Diagnosis.</emphasis>
Pedicel brown to black, postpedicel yellow; coxae brown to black, wing brown infuscated, anal lobe narrower basally, abdomen with ground color dark brown, shiny, tergite 1 sparsely gray pruinose, tergite 2 brown to black pruinose basally; surstyli asymmetrical, both surstyli with a lobe on inner margin; apex of phallic guide short, with acute tip and distinct submedian lobe dorsally, backwards directed; phallus unbranched, with leaf-shaped membranous projections (
<figureCitation id="D7C05CAFFFDA4847FE62D3E6FDB32EBB" box="[443,541,2014,2041]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="17" pageNumber="218">Fig. 105</figureCitation>
).
</paragraph>
<caption id="1B8410A2FFD94844FF4ED349FBB22EB4" ID-DOI="http://doi.org/10.5281/zenodo.6592682" ID-Zenodo-Dep="6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="18" pageNumber="219" startId="18.[151,264,1905,1929]" targetBox="[181,1406,181,1882]" targetPageId="18">
<paragraph id="4F44402AFFD94844FF4ED349FBB22EB4" blockId="18.[151,1436,1905,2038]" pageId="18" pageNumber="219">
<emphasis id="7D8F9C38FFD94844FF4ED349FECF2EC8" bold="true" box="[151,353,1905,1930]" pageId="18" pageNumber="219">FIGURES 94106.</emphasis>
<taxonomicName id="88FB3BA9FFD94844FEBFD34AFDED2ECB" authorityName="De Meyer" authorityYear="1994" box="[358,579,1906,1930]" class="Insecta" family="Pipunculidae" genus="Cephalops" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="inpaganus">
<emphasis id="7D8F9C38FFD94844FEBFD34AFDED2ECB" box="[358,579,1906,1930]" italics="true" pageId="18" pageNumber="219">Cephalops inpaganus</emphasis>
</taxonomicName>
(IAvHM100905). Male.
<emphasis id="7D8F9C38FFD94844FC94D34AFCC82EC8" bold="true" box="[845,870,1906,1930]" pageId="18" pageNumber="219">94</emphasis>
, Habitus, left lateral view;
<emphasis id="7D8F9C38FFD94844FBA0D34AFB3C2EC8" bold="true" box="[1145,1170,1906,1930]" pageId="18" pageNumber="219">95</emphasis>
, Habitus, dorsal view;
<emphasis id="7D8F9C38FFD94844FAA4D34AFA382EC8" bold="true" box="[1405,1430,1906,1930]" pageId="18" pageNumber="219">96</emphasis>
, Antenna;
<emphasis id="7D8F9C38FFD94844FF22D3AEFEBA2EEC" bold="true" box="[251,276,1942,1966]" pageId="18" pageNumber="219">97</emphasis>
, Thorax, dorsal view;
<emphasis id="7D8F9C38FFD94844FE20D3AEFDBC2EEC" bold="true" box="[505,530,1942,1966]" pageId="18" pageNumber="219">98</emphasis>
, Wing;
<emphasis id="7D8F9C38FFD94844FDBDD3AEFDD32EEC" bold="true" box="[612,637,1942,1966]" pageId="18" pageNumber="219">99</emphasis>
, Abdomen, dorsal view;
<emphasis id="7D8F9C38FFD94844FCA4D3AEFC0D2EEC" bold="true" box="[893,931,1942,1966]" pageId="18" pageNumber="219">100</emphasis>
, Tergites and sternites 6 and 7, ventral view;
<emphasis id="7D8F9C38FFD94844FAA9D3AEFA382EEC" bold="true" box="[1392,1430,1942,1966]" pageId="18" pageNumber="219">101</emphasis>
, Terminalia, dorsal view;
<emphasis id="7D8F9C38FFD94844FE4ED382FE132E90" bold="true" box="[407,445,1978,2002]" pageId="18" pageNumber="219">102</emphasis>
, Left surstylus, lateral view;
<emphasis id="7D8F9C38FFD94844FD32D382FCBF2E90" bold="true" box="[747,785,1978,2002]" pageId="18" pageNumber="219">103</emphasis>
, Right surstylus, lateral view;
<emphasis id="7D8F9C38FFD94844FB95D382FBDC2E90" bold="true" box="[1100,1138,1978,2002]" pageId="18" pageNumber="219">104</emphasis>
, Hypandrium and gonopods, ventral view,
<emphasis id="7D8F9C38FFD94844FEF9D3E6FEE82EB4" bold="true" box="[288,326,2014,2038]" pageId="18" pageNumber="219">105</emphasis>
, Phallic guide and phallus, left lateral view;
<emphasis id="7D8F9C38FFD94844FCD2D3E6FC9F2EB4" bold="true" box="[779,817,2014,2038]" pageId="18" pageNumber="219">106</emphasis>
, Ejaculatory apodeme.
</paragraph>
</caption>
<paragraph id="4F44402AFFD84845FF1ED4AEFEA02BB3" blockId="19.[151,1437,150,968]" pageId="19" pageNumber="220">
<emphasis id="7D8F9C38FFD84845FF1ED4AEFE4629F2" bold="true" box="[199,488,150,176]" pageId="19" pageNumber="220">Intraspecific variability.</emphasis>
<bibRefCitation id="2B6A3DDBFFD84845FE36D4AEFD2529F2" author="Rafael, J. A." box="[495,651,150,176]" pageId="19" pageNumber="220" pagination="353 - 390" refId="ref19665" refString="Rafael, J. A. (1990) Revisao das especies neotropicais do genero Cephalops Fallen (Diptera: Pipunculidae). Acta Amazonica, 20, 353 - 390." type="journal article" year="1990">Rafael (1990)</bibRefCitation>
provided a complete description and illustrations of the Brazilian
<typeStatus id="9040FE88FFD84845FAAED4AEFF4F2996" pageId="19" pageNumber="220" type="holotype">holotype</typeStatus>
; however, we found variations in the Colombian material. Because of this, we provide a comparison of the variations between the Colombian and Brazilian specimens (between parenthesis, when comparable). MALE. Length with
<quantity id="8803EDCFFFD84845FEF4D53AFE28285F" box="[301,390,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="19" pageNumber="220" unit="mm" value="2.3">2.3 mm</quantity>
(
<emphasis id="7D8F9C38FFD84845FE40D53BFE4F285E" box="[409,481,259,284]" italics="true" pageId="19" pageNumber="220">versus</emphasis>
<quantity id="8803EDCFFFD84845FE33D53AFDEA285F" box="[490,580,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="19" pageNumber="220" unit="mm" value="3.1">3.1 mm</quantity>
in the
<typeStatus id="9040FE88FFD84845FD40D53AFCAE285F" box="[665,768,258,285]" pageId="19" pageNumber="220" type="holotype">holotype</typeStatus>
); postcranium gray pruinose laterally and brown pruinose dorsally (
<figureCitation id="D7C05CAFFFD84845FEDBD51EFE292803" box="[258,391,294,321]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Figs 9495</figureCitation>
); pedicel brown (
<figureCitation id="D7C05CAFFFD84845FD94D51EFD0E2803" box="[589,672,294,321]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 96</figureCitation>
) (
<emphasis id="7D8F9C38FFD84845FD63D51FFCAC2802" box="[698,770,295,320]" italics="true" pageId="19" pageNumber="220">versus</emphasis>
pedicel black); scutum and scutellum brown (
<figureCitation id="D7C05CAFFFD84845FAC8D51EFA3A2802" box="[1297,1428,294,321]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Figs 95, 97</figureCitation>
) (
<emphasis id="7D8F9C38FFD84845FF78D573FF472826" box="[161,233,331,356]" italics="true" pageId="19" pageNumber="220">versus</emphasis>
scutum and scutellum black); wing with membrane slightly brown infuscate (
<figureCitation id="D7C05CAFFFD84845FBBBD572FB192826" box="[1122,1207,330,356]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 98</figureCitation>
); coxae brown (
<figureCitation id="D7C05CAFFFD84845FAB6D572FF1A28CA" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 94</figureCitation>
) (
<emphasis id="7D8F9C38FFD84845FF16D557FEB928CA" box="[207,279,367,392]" italics="true" pageId="19" pageNumber="220">versus</emphasis>
coxae black); trochanters light brown (
<figureCitation id="D7C05CAFFFD84845FD00D556FC8228CA" box="[729,812,366,392]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 94</figureCitation>
) (
<emphasis id="7D8F9C38FFD84845FC9ED557FC2128CA" box="[839,911,367,392]" italics="true" pageId="19" pageNumber="220">versus</emphasis>
trochanters yellow); femora yellow, except in proximal third, light yellow, and short setae ventrally (
<figureCitation id="D7C05CAFFFD84845FD36D5AAFCC128EF" box="[751,879,402,429]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Figs 9495</figureCitation>
) (
<emphasis id="7D8F9C38FFD84845FC5CD5ABFC6328EE" box="[901,973,403,428]" italics="true" pageId="19" pageNumber="220">versus</emphasis>
femora entirely yellow, and without setae ventrally); tibiae light brown, with bases and apices light yellow, with a row of long and fine yellow setae posterolaterally (
<figureCitation id="D7C05CAFFFD84845FF3FD5E2FE9428B6" box="[230,314,474,500]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 94</figureCitation>
); tergites 35 dark brown, shiny, with inconspicuous scattered setae (
<figureCitation id="D7C05CAFFFD84845FBE2D5E2FB4A28B6" box="[1083,1252,474,501]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Figs 9495, 99</figureCitation>
) (
<emphasis id="7D8F9C38FFD84845FB25D5E3FAEA28B6" box="[1276,1348,475,500]" italics="true" pageId="19" pageNumber="220">versus</emphasis>
tergites 34 with narrow posterior band dark brown to black pruinose); tergites and sternites 6 and 7 as in
<figureCitation id="D7C05CAFFFD84845FB03D5C6FA942B5A" box="[1242,1338,510,536]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 100</figureCitation>
. Surstyli asymmetrical, shorter than epandrium, completely setose, both surstyli with a lobe on inner margin, left surstylus slightly shorter and thin (
<figureCitation id="D7C05CAFFFD84845FE6DD67EFDB82B22" box="[436,534,582,608]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 101</figureCitation>
); both surstyli with tips acute and downward directed when seen in lateral view (
<figureCitation id="D7C05CAFFFD84845FF46D652FE932BC6" box="[159,317,618,644]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Figs 102103</figureCitation>
); gonopods subsymmetrical, right gonopod slightly thinner than left (
<figureCitation id="D7C05CAFFFD84845FBE7D652FB0E2BC6" box="[1086,1184,618,644]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 104</figureCitation>
); apex of phallic guide short, with acute tip and distinct submedian lobe dorsally, backwards directed (
<figureCitation id="D7C05CAFFFD84845FBD3D6B6FBC22BEB" box="[1034,1132,654,681]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 105</figureCitation>
); ejaculatory apodeme pinshaped, with margins somewhat straight (
<figureCitation id="D7C05CAFFFD84845FDB1D68AFD652B8F" box="[616,715,690,717]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 106</figureCitation>
); phallus unbranched, with leaf-shaped membranous projections (
<figureCitation id="D7C05CAFFFD84845FF79D6EEFEAC2BB3" box="[160,258,726,753]" captionStart="FIGURES 94106" captionStartId="18.[151,264,1905,1929]" captionTargetBox="[181,1406,181,1882]" captionTargetId="figure-30@18.[181,1406,181,1882]" captionTargetPageId="18" captionText="FIGURES 94106. Cephalops inpaganus (IAvHM100905). Male. 94, Habitus, left lateral view; 95, Habitus, dorsal view; 96, Antenna; 97, Thorax, dorsal view; 98, Wing; 99, Abdomen, dorsal view; 100, Tergites and sternites 6 and 7, ventral view; 101, Terminalia, dorsal view; 102, Left surstylus, lateral view; 103, Right surstylus, lateral view; 104, Hypandrium and gonopods, ventral view, 105, Phallic guide and phallus, left lateral view; 106, Ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.6592682" httpUri="https://zenodo.org/record/6592682/files/figure.png" pageId="19" pageNumber="220">Fig. 105</figureCitation>
).
</paragraph>
<paragraph id="4F44402AFFD84845FF1ED6C2FD8F2A1F" blockId="19.[151,1437,150,968]" pageId="19" pageNumber="220">
<emphasis id="7D8F9C38FFD84845FF1ED6C2FE192A56" bold="true" box="[199,439,762,788]" pageId="19" pageNumber="220">Examined material.</emphasis>
<materialsCitation id="FF934A77FFD84845FE1BD6C2FDB32A1F" ID-GBIF-Occurrence="3797740311" collectionCode="CEUA" collectorName="J. Torres-Toro &amp; A. L Montoya" country="Colombia" county="Envigado" latitude="6.16913" location="Zona" longLatPrecision="1" longitude="-75.60201" municipality="Exito-Pirelli" pageId="19" pageNumber="220" specimenCount="1" specimenCount-male="1" stateProvince="Antioquia">
<collectingCountry id="37EC00BAFFD84845FE1BD6C2FDF42A56" box="[450,602,762,788]" name="Colombia" pageId="19" pageNumber="220">COLOMBIA</collectingCountry>
,
<collectingRegion id="8D3F8EC8FFD84845FDB1D6C2FD752A56" box="[616,731,762,788]" country="Colombia" name="Antioquia" pageId="19" pageNumber="220">Antioquia</collectingRegion>
,
<collectingCounty id="A62538A6FFD84845FD30D6C2FCF92A56" box="[745,855,762,788]" pageId="19" pageNumber="220">Envigado</collectingCounty>
, separador vial sector bodegas
<collectingMunicipality id="AF20DA50FFD84845FB1FD6C4FAFD2A55" box="[1222,1363,764,791]" pageId="19" pageNumber="220">Éxito-Pirelli</collectingMunicipality>
,
<location id="4A2416F1FFD84845FABAD6C3FA332A57" LSID="urn:lsid:plazi:treatment:C752F13CFFDA4845FF4ED255FE182A8A:4A2416F1FFD84845FABAD6C3FA332A57" box="[1379,1437,763,789]" country="Colombia" county="Envigado" latitude="6.16913" longLatPrecision="1" longitude="-75.60201" municipality="Exito-Pirelli" name="Zona" pageId="19" pageNumber="220" stateProvince="Antioquia">Zona</location>
industrial,
<location id="4A2416F1FFD84845FEC8D727FEE32A7B" LSID="urn:lsid:plazi:treatment:C752F13CFFDA4845FF4ED255FE182A8A:4A2416F1FFD84845FEC8D727FEE32A7B" box="[273,333,799,825]" country="Colombia" county="Envigado" latitude="6.16913" longLatPrecision="1" longitude="-75.60201" municipality="Exito-Pirelli" name="Jama" pageId="19" pageNumber="220" stateProvince="Antioquia">Jama</location>
,
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/
<geoCoordinate id="2ACF26EDFFD84845FE0BD726FDF82A7B" box="[466,598,798,825]" degrees="75,60201" direction="west" orientation="longitude" pageId="19" pageNumber="220" precision="1" value="-75.60201">75,60201W</geoCoordinate>
, 15.ago[viii]01.sep[ix].2017,
<collectorName id="E20E25FCFFD84845FC6AD727FBF82A7B" box="[947,1110,799,825]" pageId="19" pageNumber="220">J. Torres-Toro</collectorName>
,
<collectorName id="E20E25FCFFD84845FBB8D726FB552A7B" box="[1121,1275,798,825]" pageId="19" pageNumber="220">A.L Montoya</collectorName>
(
<specimenCount id="59FD8BA3FFD84845FAD6D726FA972A7B" box="[1295,1337,798,825]" pageId="19" pageNumber="220" type="male">1 ♂</specimenCount>
<collectionCode id="29EAD8EFFFD84845FA9BD726FA202A7A" box="[1346,1422,798,824]" pageId="19" pageNumber="220">CEUA</collectionCode>
100905) (photographed specimen)
</materialsCitation>
.
</paragraph>
<paragraph id="4F44402AFFD84845FF1ED75EFBC22AC2" blockId="19.[151,1437,150,968]" box="[199,1132,870,897]" pageId="19" pageNumber="220">
<emphasis id="7D8F9C38FFD84845FF1ED75EFDAB2AC2" bold="true" box="[199,517,870,896]" pageId="19" pageNumber="220">Geographical distribution.</emphasis>
<collectingCountry id="37EC00BAFFD84845FDD5D75EFDFA2AC2" box="[524,596,870,896]" name="Brazil" pageId="19" pageNumber="220">Brazil</collectingCountry>
;
<collectingCountry id="37EC00BAFFD84845FD86D75EFD632AC2" box="[607,717,870,896]" name="Colombia" pageId="19" pageNumber="220">Colombia</collectingCountry>
(
<collectingRegion id="8D3F8EC8FFD84845FD05D75EFCFE2AC3" box="[732,848,870,897]" country="Colombia" name="Antioquia" pageId="19" pageNumber="220">Antioquia</collectingRegion>
) (new record) (
<figureCitation id="D7C05CAFFFD84845FC24D75EFBF12AC2" box="[1021,1119,870,896]" captionStart="FIGURE 108" captionStartId="20.[151,250,1701,1725]" captionTargetBox="[151,1436,669,1677]" captionTargetId="figure-187@20.[151,1436,669,1677]" captionTargetPageId="20" captionText="FIGURE 108. Geographical records of Semicephalops species in Colombia. Semicephalops folium sp. nov. and C. inpaganus." figureDoi="http://doi.org/10.5281/zenodo.6592688" httpUri="https://zenodo.org/record/6592688/files/figure.png" pageId="19" pageNumber="220">Fig. 108</figureCitation>
).
</paragraph>
<paragraph id="4F44402AFFD84845FF1ED7B2FE182A8A" blockId="19.[151,1437,150,968]" pageId="19" pageNumber="220">
<emphasis id="7D8F9C38FFD84845FF1ED7B2FE842AE6" bold="true" box="[199,298,906,932]" pageId="19" pageNumber="220">Habitat.</emphasis>
The specimens were collected in urban areas at Envigado, in the department of
<collectingRegion id="8D3F8EC8FFD84845FB74D7B2FA8E2AE6" box="[1197,1312,906,932]" country="Colombia" name="Antioquia" pageId="19" pageNumber="220">Antioquia</collectingRegion>
, in the Andean region of
<collectingCountry id="37EC00BAFFD84845FE99D796FE1C2A8A" box="[320,434,942,968]" name="Colombia" pageId="19" pageNumber="220">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>